Cladobradus Pic, 1918

Ivie, Michael A. & Pollock, Darren A., 2012, The Familial Placement ofCladobradusPic (Coleoptera: Ptilodactylidae, Dascillidae, Mycteridae), The Coleopterists Bulletin 66 (1), pp. 7-13 : 12

Cladobradus Pic, 1918

Type species: Cladobradus atritarsis Pic, 1918: 3 , by monotypy. Lectotype female (here designated) in MNHN.

Diagnosis. The unique habitus ( Figs. 1, 2 ) will immediately distinguish this species from all other known Mycteridae . The grooves on the face ( Fig. 5 ), dark hind wing ( Fig. 12 ), posteriorly broad, laterally upturned, and medially grooved pronotum ( Figs. 8, 9 ), and corrugated elytra ( Fig. 1 ) will confirm the identity of this genus and species.

Description. Body ( Figs. 1, 2 ) yellow-orange except for eye, antennae, tarsomeres (except last), and hind wing infuscate to piceous; most of body covered with moderately dense, suberect, golden setae that do not obscure surface. Shape elongate, slightly broader apically; [distortion of the specimen makes the following measurements approximate] body estimated at 3.1X longer than greatest width; total length 12 mm. Head ( Figs. 3, 4 5 ): Relatively short, slightly narrowed behind eyes; epistoma trapezoidal, frontoclypeal suture indicated frontolaterally by deep notch at juncture with anterolateral lobe frons, from which extends deep, arcuate groove; frons with pair of strong arcuate grooves continuous from sides of epistoma to nearly level of posterior margin of eyes; anterior margin of epistoma with unsclerotized strip of cuticle; labrum transverse, anterior margin weakly emarginate, crenulate; antennal insertions not concealed dorsally. Antennae ( Fig. 7 ): Moderate in length, antennomeres 3–10 strongly serrate, 11 distinctly narrowed and produced in apical ¼. Eyes ( Figs. 3–5 ): Moderate, hemispherical, widely separated; facets moderately fine, with short, fine, erect intrafacetal setae. Mandibles ( Figs. 3, 5 ): Strongly curved; acute; apices bidentate. Maxilla ( Fig. 3 ): Palp elongate, apical palpomere slightly securiform. Labium ( Fig. 3 ): Mentum transverse; ligula transverse, broadened and extended significantly anterad base of palpi; labial palpi short, distal palpomere blunt apically. Gula ( Fig. 3 ): Lyriform, sutures convergent anteriorly, surface indistinctly convex. Prothorax ( Figs. 8–11 ): Pronotum broadly campanulate, 1.5X wider than long (greatest pronotal width/pronotal length 1.55); narrowest at anterior margin, widest just anterad hind angles; lateral margins curved, continuous with anterior margin, ending at right hind angles; without lateral carina; disc upturned laterally in basal half, with large transverse depression on each side, medially with longitudinal narrow, glabrous groove; posterior pits present, openings incised, foveate, joined transversely by narrow, bisinuate groove; subrugosely punctate; prosternal process lamellate between procoxae, extending slightly behind procoxae, ending in a knob; coxal cavities open externally, closed internally; protrochantin hidden. Mesothorax ( Fig. 10 ): Mesoventrite triangular, lateral margins straight; mesepisterna narrowly separated anteriorly, mesocoxal cavities closed, with hidden trochantins; mesoventral process elongate, slender, touching short metaventral process. Metaventrite ( Figs. 2 , 14 ): Convex, with distinct, nearly complete discrimen which ends at level of posterior edge of mesocoxae. Metendosternite: Unstudied. Elytra: Elongate, 1.9X longer than greatest combined width (condition of the specimen makes this figure approximate), moderately convex; surface with 3 longitudinally rounded ridges, rendering the surface corrugated; setae arranged in a herringbone pattern, directed 45° off posterior from bottom of grooves onto ridges from each side; epipleuron distinct, visible to near end of ventrite 4. Legs: Slen- der, all similar in size and shape; femora broad, tibiae relatively straight and slender; tibial spurs similar in size and shape, short, not extending beyond setal fringe; tarsal formula 5-5-4, pro- and mesotarsomeres 1-4 of decreasing length and increasing width, 3 and 4 short dorsally but 4 strongly lobed ventrally; tarsomere 5 elongate, distally widened slightly; metatarsus similar, but in this case only 3 is short and lobed; tarsal claws with strong quadrate tooth reaching halfway to acute apex. Abdomen: First 3 ventrites connate. Genitalia: Unstudied.

Geographical Distribution. Known only from a single specimen, labeled Bogota, taken to be Bogotá, Colombia .

Figures

Figs. 1–2. Cladobradus atritarsis. 1) Dorsal habitus; 2) Ventral habitus.

Figs. 3–7. Cladobradus atritarsis. 3) Head, ventral view; 4) Head, lateral view; 5) Head, frontal view. 6) Type labels; 7) Antenna.

Figs. 12–14. Cladobradus atritarsis. 12) Metathoracic wing; 13) Elytral apices; 14) Abdomen.

Figs. 8–11. Cladobradus atritarsis. 8) Pronotum, dorsal view; 9) Pronotum, oblique anterior view; 10) Pro- and mesosterna; 11) Internal closure of procoxae.