Estoloides chiapensis, Santos-Silva & Wappes & Galileo, 2018

Santos-Silva, Antonio, Wappes, James E. & Galileo, Maria Helena M., 2018, Descriptions and synonymies in American Desmiphorini (Coleoptera, Cerambycidae, Lamiinae), Zootaxa 4375 (4), pp. 451-501 : 466-467

Estoloides chiapensis sp. nov.

( Figs 88–92 )

Description. Male. Integument mostly dark brown; apex of palpomeres yellowish brown; elytra dark reddish brown with black areas (dark areas slightly obscured by pubescence, typically forming irregular bands).

Head. Frons and area between antennal tubercles moderately coarsely, abundantly punctate; remaining surface of vertex with sparser punctures, especially close to prothoracic margin; dorsal surface with yellowish brown pubescence obscuring integument, but exposing punctures, interspersed with short, curved yellow setae throughout, and a few long, erect brown setae close to lower eye lobes and yellow close to upper eye lobes. Area behind upper eye lobes impunctate; with yellowish brown pubescence obscuring integument. Area behind lower eye lobes, close to eye, tumid; moderately coarsely, sparsely punctate on tumid area toward gena, minutely, moderately sparsely punctate on remaining surface, sparser toward gena; tumid area with yellowish brown pubescence, close to eye, obscuring integument (this area widened toward gena), interspersed with long, erect, sparse yellow and yellowish brown setae; with longitudinal band with yellowish brown pubescence adjacent to tumid area, sparser than on area close to eye (this band fused with pubescence behind upper eye lobes); remaining surface glabrous. Antennal tubercles densely micropunctate; pubescence as on frons with a few long, erect yellowish brown setae interspersed. Median groove distinct from clypeus to prothoracic margin. Genae finely, abundantly transversely striate on basal 2/3, with a few coarse punctures interspersed, micropunctate on distal third; with yellowish brown pubescence not obscuring integument, slightly lighter close to eyes, with a few long, erect brown setae interspersed. Gulamentum smooth, shining, glabrous except narrow distal area moderately coarsely, transversely sulcate, with sparse yellowish brown pubescence. Labrum convex on basal 2/3, inclined on distal third; minutely abundantly punctate (punctures slightly coarser adjacent to inclined area); with short and long, spare yellowish brown setae, fringe of golden setae at distal margin, and a few long, thick dark setae laterally on convex area. Distance between upper eye lobes 0.65 times length of scape (4.3 times width of one lobe); in frontal view, distance between lower eye lobes equal to length of scape. Antennae 2.00 times elytral length, reaching elytral apex at distal third of antennomere VII. Scape gradually widened at inner side; densely micropunctate; with yellowish brown pubescence partially obscuring integument, with long, erect setae interspersed ventrally on distal third. Antennomeres IV–XI with yellowish brown pubescence except narrow basal ring with white pubescence (slightly so also at apex of IV–X); antennomeres III–XI with long, erect setae ventrally, gradually shorter and sparser toward XI. Antennal formula (ratio) based on length of antennomere III: scape = 0.76; pedicel = 0.16; IV = 1.08; V = 0.85; VI = 0.80; VII = 0.75; VIII = 0.65; IX = 0.62; X = 0.51; XI = 0.50.

Thorax. Prothorax, maximum width 1.3 times wider than long (including lateral tubercles); lateral tubercles large, placed at middle, with apex acute. Pronotum with distinct, elongate gibbosity centrally on basal half; coarsely, moderately abundantly punctate, denser on each side of smooth gibbosity; with yellowish brown pubescence nearly obscuring integument, but exposing punctures, with a few long, erect setae interspersed on basal third. Sides of prothorax coarsely, moderately sparsely punctate on wide central area, impunctate on base and on wide, oblique area from procoxal cavity to distal margin; with yellowish brown pubescence partially obscuring integument. Prosternum finely, sparsely punctate on narrow area close to procoxal cavities and prosternal process, smooth on remaining surface; with yellowish brown pubescence not obscuring integument. Prosternal process slightly depressed on wide central area, coarsely, somewhat rugose-punctate; with yellowish brown pubescence not obscuring integument, bristly toward apex. Mesoventrite with yellowish brown pubescence exposing integument centrally, distinctly denser laterally. Mesanepisternum, mesepimeron, metanepisternum and metaventrite (except nearly glabrous central area) with yellowish brown pubescence partially obscuring integument. Scutellum with dense yellow pubescence, except nearly glabrous central area on base. Elytra. Moderately coarsely and abundantly punctate on basal quarter, sparser on remaining surface (punctures subaligned except on basal quarter); with yellowish brown pubescence obscuring integument, with short, moderately abundant, erect yellow setae interspersed; elytral apices individually rounded. Legs. Femora with yellowish brown pubescence partially obscuring integument.

Abdomen. Ventrites with yellowish brown pubescence partially obscuring integument (more yellowish on narrow distal area of ventrites I–IV), except for small glabrous, circular areas (more distinct on ventrites II–V). Distal margin of ventrite V truncate.

Female. Differs from male by the shorter antennae (1.55 times elytral length, reaching elytral apex near apex of antennomere IX).

Variation. Distance between upper eye lobes in male from 0.65 to 0.70 times length of scape; antennae in male reaching elytral apex at middle of antennomere VIII, in female, at about middle of X.

Dimensions (mm), holotype male/ paratype male/ paratypes female. Total length, 11.16/9.22/11.61–11.88; prothoracic length, 2.16/1.80/2.20–2.25; basal prothoracic width 2.22/1.80/2.34–2.43; distal prothoracic width, 1.84/1.65/2.11–2.16; maximum prothoracic width (between apices of lateral tubercles), 2.83/2.29/2.97–3.06; humeral width, 3.28/2.61/3.46–3.47; elytral length, 7.60/6.34/7.87–8.28.

Type material. Holotype male from MEXICO, Chiapas: 17 km W. Tutl. Gtz. (3300’), 1–8.VII.1986, Wappes col. ( FSCA) . Paratypes – MEXICO, Chiapas: 1 male, 2 females, same data as holotype (1 male, 1 female, ACMT, 1 female MZSP)) ; El Aguacero (16 km N Ocozocoautla ), 1 female, 16–23.X.1988, Giesbert col. ( FSCA) . Guerrero: 10.5 km N Iguala , 1 female, 12.X.1994, Giesbert col. ( FSCA) .

Etymology. Named for Chiapas, the Mexican State where the holotype and three of the paratypes were collected.

Remarks. Estoloides chiapensis ( Figs 88, 92 ) is similar to E. noguerai sp. nov., but has a more slender body and scape (which is more gradually widened internally toward apex), and finer, sparser and less conspicuous elytral punctation. E. noguerai ( Figs 77 , 81 ), has a wider body and scape (which is more distinctly widened internally), and denser, more conspicuous elytral punctation. It differs from E. chamelae ( Figs 21 , 22 ) by the slender body, and punctation on dorsal surface of body less obliterated by pubescence.

Figures

FIGURES 88–98. 88–92, Estoloides chiapensis: 88, holotype male, dorsal habitus; 89, holotype male, ventral habitus; 90, holotype male, lateral habitus; 91, holotype male, head, frontal view; 92, paratype female, dorsal habitus. 93–97, Estoloides pubescens, holotype female:93, head, frontal view; 94, mesoventral process; 95, dorsal habitus; 96, ventral habitus; 97, lateral habitus.98, Estoloides andresi, holotype female, mesoventral process.

FIGURES 72–79. 72–75, Estoloides nayeliae, holotype female: 72, dorsal habitus; 73, ventral habitus; 74, lateral habitus; 75, head, frontal view. 76–79, Estoloides noguerai, holotype male: 76, head, frontal view; 77, dorsal habitus; 78, ventral habitus; 79, lateral habitus.

FIGURES 80–87. 80–82, Estoloides noguerai: 80, paratype male from Nicaragua, dorsal habitus; 81, paratype female, dorsal habitus; 82, paratype female, ventral habitus. 83–86, Estoloides schusteri, holotype male: 83, head, frontal view; 84, dorsal habitus; 85, ventral habitus; 86, lateral habitus. 87, Estoloides leucosticta, holotype male, dorsal habitus.

FIGURES 13–21. 13, Estoloides alboscutellaris, holotype female, dorsal habitus.14, E. annulicornis, holotype female, dorsal habitus. 15, E. reflexa, holotype, dorsal habitus. 16–18, E. prolongata, syntype: 16, lateral habitus; 17, labels, 18, dorsal habitus.19–20, E.scabracaulis, dorsal habitus:19, male; 20, female.21, E.chamelae, male, dorsal habitus.

FIGURES 22–28. 22–24, Estoloides chamelae, dorsal habitus: 22, female; 23, holotype male; 24, paratype female; 25, paratype male. 26–28, E. modica: 26, paratype male, dorsal habitus; 27, holotype male, dorsal habitus; 28, holotype male, lateral habitus.