Gondwanodromia wardi, Sinclair & Saigusa, 2022

Sinclair, Bradley J. & Saigusa, Toyohei, 2022, A New Aquatic Associated Genus of Trichopezinae from the Southern Hemisphere (Diptera: Empidoidea: Brachystomatidae), Records of the Australian Museum (Rec. Aust. Mus.) 74 (3), pp. 75-98 : 94-95

Gondwanodromia wardi sp. nov.

urn:lsid:zoobank.org:act:1A1E47BA-06AF-4790-AD9A-36689F91DBAB

Figs 46, 47 , 49 , 51

Holotype ♂, labelled: “ NEW ZEALAND:/ Haast R.| Harris Ck./ Greenstone Ck.|/ Glitterburn [43°56'S 169°07'E], 3.xi.1994 / J.B. Ward ”; “ HOLOTYPE / Gondwanodromia / wardi/ Sinclair & Saigusa [red label]” [dissected] ( CMNZ) . Paratype: 1 ♀, same data as holotype ( CMNZ) .

Diagnosis. The species is readily distinguished by the scutum with shiny vittae between setal rows, lateral scutal setae spine-like, and male terminalia with epandrium prolonged beyond base of surstylus in lateral view.

Description. Wing length 3.8– 4 mm. Male. Head. Eye bare; widely separated on frons and face, wider than antennal sockets. Ocellar setae long and divergent, inserted anteromedial to lateral ocelli; 2 pairs of shorter posterior setae; 2 vertical setae, outer seta arched over eye, longer than ocellar setae. Antennae ( Fig. 46 ) inserted above middle of head; scape slightly longer than globular pedicel; postpedicel with broad base, gradually curved and tapered; stylus threequarters length of postpedicel, small basal segment and short apical peg-like sensillum. Palpus long and slender, twothirds height of head, bearing scattered setae; labrum greatly lengthened, longer than height of head, slightly incurved; labium subequal in length to labrum.

Thorax. Pleura, coxae, pprn and lower margin of npl with pale silvery-grey pruinescence ( Fig. 46 ). Scutum brown with shiny vittae between rows of acrostichals and dorsocentrals ( Fig. 47 ). Antepronotum with several pairs of short setulae. Acrostichals uniserial in front, slightly longer and biserial behind, extending to prescutellar depression; 6 dorsocentral setae, intermixed with long and short, slender setae; lateral setae spine-like: 1 postpronotal seta; 1 presutural supraalar seta; 2 notopleural setae, lower slender and short; 3 postsutural supra-alar setae with 1–2 short setae; 1 postalar seta; 2–3 pairs scutellar setae and outer pairs shorter than median pair. Laterotergite with patch of dark brown setae Wing. Broad, infuscate; basal costal seta long and slender; costa with erect dorsal setae beyond R 1; R 4 slightly sinuous; halter pale yellowish-brown. Legs. Coxae with pale golden setae; knees yellowish-brown. Fore femur with anteroventral and posteroventral row of short, slender setae, shorter than width of femur except basal posteroventral setae longer than width; 3 stout dorsal setae on basal half, subequal to width of femur. Mid femur with anteroventral and posteroventral rows of stout, black setae, posteroventral setae mostly longer than width of femur; apical half with 3–4 stout anterior setae. Mid tibia lacking erect ventral setae. Hind femur sinuous with long, erect black dorsal and anteroventral setae from base to beyond middle. Hind tibia with stout ventral and dorsal setae, shorter than width of tibia.

Abdomen. Brown, sparsely setose with setae mostly confined to the posterior margins. Segment 7 narrow posteriorly; sternite 7 with short, triangular posterolateral extension. Terminalia ( Fig. 49 ). Lying dorsally over segments 7 and 8. Hypandrium round apically, encircling phallus (likely postgonites fused with hypandrium at apex). Phallus with broad membranous cap; ejaculatory apodeme broad and tapered ventrally. Epandrium broad, with narrowed and tapered posterodorsal corner. Surstylus subtriangular with short setae on apex and medial face. Cercus membranous, clothed in microtrichia; several long, setae on anterior and posterior faces.

Female. Similar to male except as follows: Abdominal sternites 3–6 with lateral marginal tuft of long setae; sternite 7 with two pairs of closely approximated ventral setae, length two-thirds width of segment. Terminalia [undissected]. Cercus elongate, shiny, well sclerotized, projecting horizontally, longer than segment 8.

Etymology. This species is named after the collector of the type series, John Ward, who collected many valuable species of Empidoidea.

Distribution. This species is currently known from the type locality on South Island, New Zealand ( Fig. 51 ).

Figures

Figures 43–47. New Zealand species of Gondwanodromia: (43) G. tongariro sp. nov, holotype, male terminalia, scale bar = 0.25 mm; (44) G. femorata sp. nov., female midleg, scale bar = 0.5 mm (courtesy Canterbury Museum); (45) G. femorata sp. nov., female abdomen, scale bar = 0.5 mm (courtesy Canterbury Museum); (46) G. wardi sp. nov., holotype (abdomen dissected), lateral view, scale bar = 0.5 mm (courtesy Canterbury Museum); (47) G. wardi sp. nov., holotype (abdomen dissected), dorsal view, scale bar = 0.5 mm (courtesy Canterbury Museum). Abbreviations: epand lb – epandrial lobe; sur – surstylus.

Figures 48–49. Male terminalia ofAustralian and New Zealand species of Gondwanodromia, lateral view: (48) G. tonnoiri sp. nov.; (49) G. wardi sp. nov. Scale bar = 0.1 mm. Abbreviations: a sur – anterior surstylus; cerc –cercus; epand – epandrium; hypd – hypandrium; p sur – posterior surstylus; ph – phallus; sur – surstylus.

Figures 50–52. Distribution of Gondwanodromia: (50) Australian species; (51) New Zealand species; (52) South American species.