Gondwanodromia mutabilis (Collin) Sinclair & Saigusa, 2022

Sinclair, Bradley J. & Saigusa, Toyohei, 2022, A New Aquatic Associated Genus of Trichopezinae from the Southern Hemisphere (Diptera: Empidoidea: Brachystomatidae), Records of the Australian Museum (Rec. Aust. Mus.) 74 (3), pp. 75-98 : 88-89

publication ID	https://doi.org/ 10.3853/j.2201-4349.74.2022.1797
publication LSID	lsid:zoobank.org:pub:58350A47-1E4C-4B31-A9C1-91159F363D35
persistent identifier	https://treatment.plazi.org/id/021487E7-3D20-FF93-0DEA-FF3909FEFA2B
treatment provided by	Felipe
scientific name	Gondwanodromia mutabilis (Collin)
status	comb. nov.

Treatment

Gondwanodromia mutabilis (Collin) comb. nov.

Figs 34–36 View Figures 34–36 , 37 View Figures 37–40 , 51 View Figures 50–52

Dipsomyia mutabilis Collin, 1928: 60 ; Smith, 1967: 44. Adipsomyia mutabilis .— Yang et al., 2007: 21, 52. “Undescribed genus A” mutabilis .— Sinclair, 2011: 31.

Holotype ♀, labelled: “ TYPE / ♀ [circle label with red border]”; “ New Zealand:/ Glenorchy [–44.849 ° 168.384 °]/ 3.i.1923 / C.C. Fenwick. / B.M.1923 – 150”; “ Dipsomyia / mutabilis [dark circle label”; “ Dipsomyia / mutabilis/ TYPE ♀ Collin”; “Wing of ♀ TYPE / figured in text of/ author’s description”[dissected] ( BMNH).

Additional material examined. New Zealand. WD: 4 ♂, 1 ♀, Haast River / Harris Ck / Greenstone Ck., Glitterburn [43°56'S 169°07'E], 3.xi.1994, J.B. Ward ( CMNZ, CNC) GoogleMaps .

Diagnosis. The species is readily distinguished by eyes with sparse ommatrichia, scutum clothed in greyish pruinescence ( Fig. 37 View Figures 37–40 ), lateral scutal setae spine-like; female cercus slender, short subequal to length of tergite 10; male terminalia with epandrium prolonged beyond surstylus.

Description. Wing length 4–4.2 mm. Male. Head. Eye with sparse ommatrichia; face with bluish pruinescence, wider than antennal sockets. Ocellar setae long and divergent, inserted anteromedial to lateral ocelli; 2 pairs of shorter posterior setae; 2 vertical setae, outer seta arched over margin of eye, longer than ocellar seta; occipital setae shorter and thinner than vertical setae. Antennae inserted above middle of head; scape slightly longer than globular pedicel; postpedicel with broad base, gradually curved and tapered; stylus three-quarters length of postpedicel, small basal segment and short apical peg-like sensillum. Palpus very long and slender, three-quarters height of head, bearing scattered setae; labrum greatly lengthened, longer than height of head, slightly incurved; labium apparently slightly longer than palpus.

Thorax. Pleura, coxae, postpronotal lobe and notopleuron with pale blue pruinescence. Scutum brown with faint vittae between rows of acrostichals and dorsocentrals. Antepronotum with several pairs of short setulae.Acrostichals uniserial in front, slightly longer and biserial behind, extending to prescutellar depression; 6 dorsocentral setae, intermixed with long and short, slender setae; lateral setae spine-like: 1–3 postpronotal setae; 1 presutural supra-alar seta, with 2–3 short setae; 2 notopleural setae, lower slender and short; 2 postsutural supra-alar setae with 2–3 short setae; 1 postalar seta; 2 pairs of scutellar setae, outer pair slender. Laterotergite with patch of pale brown setae. Wing. Broad, infuscate (see Collin, 1928, fig. 17); basal costal seta long and slender; costa with erect dorsal setae beyond R 1; R 4 slightly sinuous; halter pale brown. Legs. Coxae with pale golden setae. Femora elongate, narrowed subapically. Fore femur with anteroventral and posteroventral rows of short slender, shorter than width of femur. Mid femur with anteroventral and posteroventral rows of stout, black setae, mostly shorter than width of femur; apical half with 3–4 stout anterior and posterior setae. Mid tibia with row of short erect stout setae beneath, with several stronger preapical setae. Hind femur straight with long, erect pale dorsal and lateral setae from base to beyond middle. Tarsomere 1 slender with dense ventral setae on all legs.

Abdomen. Brown, sparsely setose with setae mostly confined to posterior margins. Segment 7 narrow posteriorly; sternite 7 with short, triangular posterolateral extension. Terminalia ( Fig. 34 View Figures 34–36 ). Lying dorsally over segments 7 and 8. Hypandrium round apically, encircling phallus (likely postgonites fused with hypandrium at apex). Phallus with cap-like apex; ejaculatory apodeme small and narrow. Epandrium narrow, not expanded with round posterodorsal corner; fused to hypandrium anteroventrally. Surstylus digitiform with short setae along posterior margin and medial face. Cercus membranous, clothed in microtrichia; several long, stout setae on anterior face.

Female. Similar to male except as follows: Fore femur with two rows of short, stout setae on inner face, from middle to near apex. Terminalia ( Fig. 35 View Figures 34–36 ). Tergite 8 with small membranous mid-dorsal cleft; sclerite bare, except for transverse row of small setae increasing in size towards lateral margin; sternite 8 broad, extending beneath segment 10. Syntergite 9+10 divided mid-dorsally into two rectangular sclerites; each sclerite with mid-dorsal row of 4–5 short setae. Sternite 10 apparently absent. Cercus narrow, arched dorsally, clothed in setulae. Spermatheca ( Fig. 36 View Figures 34–36 ) small, narrow, at apex of long duct arising from small genital fork.

Distribution. This species is currently known from two localities on South Island, New Zealand ( Fig. 51 View Figures 50–52 ).

Remarks. In a note added in proof, Sinclair (1995) recognized that Dipsomyia mutabilis was not congeneric with the type species, D. spinifera Bezzi and belonged to a new genus. Yang et al. (2007) transferred mutabilis to their new genus, Adipsomyia and later the species was removed and transferred to “Undescribed genus A” by Sinclair (2011) in a key to New Zealand Empidoidea.

The male of this species is described herein for the first time. The association of these new specimens is based in part upon the female terminalia.