Babina holsti ( Boulenger, 1892 )
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publication ID |
https://doi.org/ 10.26879/503 |
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persistent identifier |
https://treatment.plazi.org/id/0149542F-5C33-3526-5BF0-B1F0387E32BC |
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treatment provided by |
Felipe |
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scientific name |
Babina holsti ( Boulenger, 1892 ) |
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Babina holsti ( Boulenger, 1892) View in CoL —Holst’s Frog
Figure 3.5–3.11 View FIGURE 3
Occurrence (MNI). Minatogawa: 2; Sashiki (lower unit): 6.
Referred material. Minatogawa: 2 female humeri (right: YMHF-MA 002 ) and 2 ilia (right: YMHF-MA 003 ); Sashiki (lower unit): 6 female humeri (1 right and 5 left: RUMF-GF-04003), 2 male humeri (1 right and 1 left: RUMF-GF-04004), and 9 ilia (6 right and 3 left: RUMF-GF-04005) .
Humerus. Nokariya’s (1984) observations on the humerus of this large-sized frog note a laterally projected epicondylus radialis, a small and ridge-like crista lateralis (female only), and a proximally positioned waisted part of the shaft (female only). The other diagnostically useful character states of the humerus are: a robust and strongly waisted shaft; a high crista ventralis with a straight ventral edge that is parallel to the dorsal edge of the shaft (in mediolateral views); the distal edge of the crista ventralis that forms an oblique angle with the ventral outline of the shaft (in mediolateral views); the distance between the ventrodistal corner of the crista ventralis and the eminentia capitata, which is only 2.5 times longer than the length of the eminentia capitata; the crista paraventralis that extends proximally to the level of the ventrodistal corner of the crista ventralis; the proximal part of the epicondylus ulnaris that protrudes medially (more than the distal part in dorsoventral views); and a tapered and round outline of the proximal part of the olecranon scar (in dorsal view). The crista lateralis and the crista medialis of male humeri inflect dorsally, with the former extending laterally beyond the level of the lateral edge of the epicondylus radialis, and the latter extending medially beyond the level of the medial edge of the epicondylus ulnaris ( Figure 3.8–3.10 View FIGURE 3 ). These crests are ridge-like in the female ( Figure 3.5–3.7 View FIGURE 3 ) and extend proximally to the level of the crista ventralis forming angulated ridges of the shaft (as a consequence, the cross-section of the narrowest part of the shaft forms a semicircle). The fossula dividens scarcely concaves in the female and some males while it concaves deeply in some males; and the spina tuberculi medialis is weak in the female and some males, but developed in other males ( Figure 3.8–3.10 View FIGURE 3 ).
Ilium. Nokariya’s (1984) observations on the ilium of this species note a well-developed crista dorsalis and tuber superior. The other diagnostic combinations of the character states on the ilium ( Figure 3.11 View FIGURE 3 ) are: an elliptical and swell-like tuber superior without ridging in the margin; the tuber superior that is in contact with the acetabular margin (contra Nokariya, 1984); a thin and high (the height is higher than that of the acetabulum) crista dorsalis with a gently round posterodorsal corner; the pars ascendens with a posteriorly curved anterior edge; and a distinct supracetabular fossa.
Remarks. While the ilia and female humeri of this species are morphologically indistinguishable from those of Babina subaspera (the sibling species endemic to the Amami Island Group: Maeda and Matsui, 1999), fossils of these elements are referred on the basis of the species’ exclusive occurrence on the island. Male humeri of B. holsti differ from those of B. subaspera by having less-developed crista lateralis and crista medialis.
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