taxonID	type	description	language	source
03CA87C3FF97B01DFFC9F954FE1D2C2C.taxon	type_taxon	Type species. Macrotoma minor Lubbock, 1862	en	Mandal, Guru Pada, Kumari, Simran, Mandal, Pritha, Roy, Koushik Kumar, Suman, Kusumendra Kumar, Bhattacharya, Kaushik Kumar (2024): Description of a new species with three new records of Tomocerus Nicolet (Collembola: Tomoceridae) from India. Journal of Insect Biodiversity and Systematics 10 (4): 755-768, DOI: 10.61186/jibs.10.4.755, URL: https://doi.org/10.61186/jibs.10.4.755
03CA87C3FF97B01DFFC9F954FE1D2C2C.taxon	diagnosis	Diagnosis. Average adult body length ranges 3 mm or more, moderate-sized Tomocerinae, body colour pale grey to deep brown with deep purple pigment patches and covered with shiny deep blue or brown scales. Eyes 6 + 6, PAO sometimes present; trochanteral organ reduced to a trochantero-femoral organ with 1, 1 pointed slender seta; dens basal portion with many simple or complex spines and distal portion with ciliated, fathered chaetae; mucro elongated with numerous chaetae of elongated sockets, paired basal teeth with corner toothlet present, outer lamella with a varying (usually from 3 – 11) number of intermediate teeth.	en	Mandal, Guru Pada, Kumari, Simran, Mandal, Pritha, Roy, Koushik Kumar, Suman, Kusumendra Kumar, Bhattacharya, Kaushik Kumar (2024): Description of a new species with three new records of Tomocerus Nicolet (Collembola: Tomoceridae) from India. Journal of Insect Biodiversity and Systematics 10 (4): 755-768, DOI: 10.61186/jibs.10.4.755, URL: https://doi.org/10.61186/jibs.10.4.755
03CA87C3FF94B019FF38FF5EFC7D2213.taxon	materials_examined	Materials examined. Holotype: 1 ♀ on slide, INDIA, Sikkim, Gangtok district, Lachen, 27 ° 42 ' 34 " N, 88 ° 33 ' 43 " E, 3248 m a. s. l., 9. ix. 2022, G. P. Mandal leg., Reg. no – 3399 / H 14. Paratypes: 1 ♀ on slide, 12 specimens in alcohol, same data as holotype. Reg. no – 3585 / H 14; all materials are deposited at NZC, ZSI.	en	Mandal, Guru Pada, Kumari, Simran, Mandal, Pritha, Roy, Koushik Kumar, Suman, Kusumendra Kumar, Bhattacharya, Kaushik Kumar (2024): Description of a new species with three new records of Tomocerus Nicolet (Collembola: Tomoceridae) from India. Journal of Insect Biodiversity and Systematics 10 (4): 755-768, DOI: 10.61186/jibs.10.4.755, URL: https://doi.org/10.61186/jibs.10.4.755
03CA87C3FF94B019FF38FF5EFC7D2213.taxon	etymology	Etymology. The species name is given on its collection state name.	en	Mandal, Guru Pada, Kumari, Simran, Mandal, Pritha, Roy, Koushik Kumar, Suman, Kusumendra Kumar, Bhattacharya, Kaushik Kumar (2024): Description of a new species with three new records of Tomocerus Nicolet (Collembola: Tomoceridae) from India. Journal of Insect Biodiversity and Systematics 10 (4): 755-768, DOI: 10.61186/jibs.10.4.755, URL: https://doi.org/10.61186/jibs.10.4.755
03CA87C3FF94B019FF38FF5EFC7D2213.taxon	description	Description. Average adult body length about 4.1 mm (without appendages). Body with brown scales, Ant. I – II all over-scaled, no scales on Ant. III – IV. Ant. III – IV with distinct annulations. Colouration. (Fig. 1 A – B) Body ground colour pale yellow / gray, anterior part of the mouth with purple pigmentation. Ant. I – II distally with a slight purple patch; pigment gradually darker from Ant. III – IV. Eyes in dark purple patches. Diffuse pigment patches present in lateral side of head and abdomen; legs coloured up to claw. Head. Head with 6 + 6 eyes, PAO absent. Antennae: Cephalic diagonal ratio = 3.3: 1. Ant. I: II: III: IV ratio as 1: 1.7: 9.7: 2.7. Cephalic dorsal macrochaetotaxy as Fig. 2 A, anterior area each side 3, 3; interocular area each side 2, 4; post ocular area 2 + 2; posterior area 2 + 2. Posterior margin with numerous small chaetae. Labral formula 4 / 5, 5, 4; Md. with four apical and five basal teeth. Mx. general type (Fig. 2 C) Outer lobe of maxillary palp with 1 large, 1 median and 2 small basal chaetae (Fig. 2 D). Lateral process of labial palp not reaching the base. Thorax. Th. II bordered with a row of dense mac and few antero-lateral mac. Five central mac arranged in a triangle on Th. II (a 2, m 2, m 3, m 4, a 5) and two bothriotricha (m 6, a 6) present on each side. 3 + 3 posterior mac present on both Th. II (p 2 – p 4) and Th. III (p 1, p 3, p 5) (Fig. 2 B). Trochantero-femoral organ with 1, 1 slender chaeta. Ventral strong chaetae formula of Tita. I – III as 6 – 7, 5, 6 – 7. Unguis slender with baso-internal ridging visible, pointed paired lateral teeth present. Inner edge of unguis with basal tooth and 6 inner distal teeth, distal ones smaller. Unguiculus lanceolate with a single inner tooth. Tenent hair strong, clavate and slightly longer than inner edge of unguis; base of unguis with two-minute accessory chaetae; two thin and long guard chaetae (Figs 2 E, 4 A). Abdomen. Pattern of body chaetotaxy as in Fig. 2 B. Two median mac on Abd III (m 3, m 6) and Abd. IV with one medio-lateral mac (m 6). Abd. I – V posterior row mac formula- 3 (m 2 – m 4), 3 (m 2 – m 4), 4 (p 1, p 3, p 6, p 7), 2 (p 6, p 7), 4 (m 2, m 3, m 5, m 6). Abd. VI with numerous moderate-size chaetae, most of them on the lateral and posterior side. Formula of midline pseudopores of terga as 1, 1, 1, 1, 0, 0. Abd. III and Abd. IV with one and two bothriotricha respectively. Ventral tube scaled with numerous posterior and 57 – 60 lateral flap chaetae (Fig. 3 A). Rami of tenaculum with 4 + 4 teeth, anterior face of corpus with 5 chaetae (Fig. 3 B). Ratio of manubrium: dens: mucro 3.5: 4.2: 1. Manubrium ventrally scaled, lateral side with 10 – 11 strong spinous chaetae (Fig. 3 C). Dental spines with numerous denticles, total 19 spines arranged as 2, 5, 5 / 5 (6), II (Figs 4 B, 5 A – B); proximal part with three distinct rows (Figs 4 C, 5 C). Mucro elongated with paired basal and 4 intermediate teeth; numerous smooth chaetae with elongated sockets present (Figs 4 D, 5 D).	en	Mandal, Guru Pada, Kumari, Simran, Mandal, Pritha, Roy, Koushik Kumar, Suman, Kusumendra Kumar, Bhattacharya, Kaushik Kumar (2024): Description of a new species with three new records of Tomocerus Nicolet (Collembola: Tomoceridae) from India. Journal of Insect Biodiversity and Systematics 10 (4): 755-768, DOI: 10.61186/jibs.10.4.755, URL: https://doi.org/10.61186/jibs.10.4.755
03CA87C3FF94B019FF38FF5EFC7D2213.taxon	discussion	Remarks. Tomocerous (s. str.) sikkimensis sp. nov. is characterised by the presence of three rows of proximal dental spines and two large distal dental spines, 4 mucronal teeth, scaled ventral tube, 6 unguis teeth, this species is closely related to the species Tomocerus (s. str.) conagensis Sun, Liang & Huang, 2006. The new species also has a similarity with T. (s. str.) nepalicus Yosii, 1971 in dental basal spine arrangement but dissimilar in mucro teeth number. Details of other differential characters of the new species with other related species are reflected in Table 1.	en	Mandal, Guru Pada, Kumari, Simran, Mandal, Pritha, Roy, Koushik Kumar, Suman, Kusumendra Kumar, Bhattacharya, Kaushik Kumar (2024): Description of a new species with three new records of Tomocerus Nicolet (Collembola: Tomoceridae) from India. Journal of Insect Biodiversity and Systematics 10 (4): 755-768, DOI: 10.61186/jibs.10.4.755, URL: https://doi.org/10.61186/jibs.10.4.755
03CA87C3FF93B01AFFC9F9E3FB142C28.taxon	materials_examined	Material examined. 2 specimens, INDIA, West Bengal, Darjeeling district, Singalila National park, 1 km North-East from Gairibas WB Govt. Rest House, 27 ° 03 ' 04.54 '' N, 88 ° 01 ' 59.91 E '', 2577 m a. s. l., 28. ii. 2021, K. K. Suman leg., Reg no. – 3056 / H 14.	en	Mandal, Guru Pada, Kumari, Simran, Mandal, Pritha, Roy, Koushik Kumar, Suman, Kusumendra Kumar, Bhattacharya, Kaushik Kumar (2024): Description of a new species with three new records of Tomocerus Nicolet (Collembola: Tomoceridae) from India. Journal of Insect Biodiversity and Systematics 10 (4): 755-768, DOI: 10.61186/jibs.10.4.755, URL: https://doi.org/10.61186/jibs.10.4.755
03CA87C3FF93B01AFFC9F9E3FB142C28.taxon	description	Description. Ground colour of body pale yellow with antennal joining pigmented and gradually darken from Ant. III – Ant. IV. Purple patches extended on lateral side of Th. II – Abd. II, coxa and femur of foreleg sometimes with dark purple patch (Fig. 6 A). Ant. I: II: III ratio as 1: 1.9: 7.6, Ant. IV missing. Antenna: head diagonal ratio 3.18: 1. Ocelli 6 + 6 dark purple coloured, PAO absent. Prelabral chaetae 4, labral chaetae formula 5, 5, 4; all smooth. Md. with four apical and seven basal teeth. Labial palp with four chaetae, apex of lateral process not reaching the base. Brown-coloured scales present on all over the body. Trochanteral-femoral organ with 1, 1 chaeta. Strong chaetae formula of fore to hind tita as 6, 6, 7. Tenet hair clavate, small accessory chaetae, and guard chaetae almost equal lengths of tenent hair. Unguis slender with distinct basal paired, 6 distal pointed inner and paired lateral teeth present. Unguiculus lanceolate with one-minute inner tooth (Fig. 6 B). Corpus of tenaculum with 11 chaetae and rami with 4 teeth, scales absent. Ratio of manubrium: dens: mucro 10.7: 14: 1. Manubrium laterally with large round scales, 1 – 2 large chaetae on proximal side and 9 spinous chaetae. Dorsal dental basal part with a prominent pointed dorsal chaetae and simple and plumose chaetae. 12 tridentate dental spines arranged as 5 / 3 – 4, I, 1 – 2, I (Fig. 6 C). Mucro with paired basal tooth with toothlet, outer lamella with 6 intermediate teeth and small apical and subapical teeth (Fig. 6 D).	en	Mandal, Guru Pada, Kumari, Simran, Mandal, Pritha, Roy, Koushik Kumar, Suman, Kusumendra Kumar, Bhattacharya, Kaushik Kumar (2024): Description of a new species with three new records of Tomocerus Nicolet (Collembola: Tomoceridae) from India. Journal of Insect Biodiversity and Systematics 10 (4): 755-768, DOI: 10.61186/jibs.10.4.755, URL: https://doi.org/10.61186/jibs.10.4.755
03CA87C3FF93B01AFFC9F9E3FB142C28.taxon	diagnosis	Differential diagnosis. Tomocerus minor is characterised by tridentate denticles of dental spine arranged in a single row with two large distal spines with 1 – 2 intermediate small spines. Habitat and distribution. Found under moss vegetation surrounding leaf litter on the way to Gairibas Hill Stream, Singalila National Park, India. The species is common in all parts of Europe.	en	Mandal, Guru Pada, Kumari, Simran, Mandal, Pritha, Roy, Koushik Kumar, Suman, Kusumendra Kumar, Bhattacharya, Kaushik Kumar (2024): Description of a new species with three new records of Tomocerus Nicolet (Collembola: Tomoceridae) from India. Journal of Insect Biodiversity and Systematics 10 (4): 755-768, DOI: 10.61186/jibs.10.4.755, URL: https://doi.org/10.61186/jibs.10.4.755
03CA87C3FF91B014FFC9FA90FB8427F6.taxon	materials_examined	Material examined. 1 specimen on slide, INDIA, West Bengal, Darjeeling district, Singalila National Park, Gairibas, Red Panda Site, collected from leaf litter, 27 ° 09 ' 3.6 " N, 87 ° 13 ' 58.8 " E, 909 m a. s. l., 28. ii. 2021, K. K. Suman leg., reg. no. - 3101 / H 14.	en	Mandal, Guru Pada, Kumari, Simran, Mandal, Pritha, Roy, Koushik Kumar, Suman, Kusumendra Kumar, Bhattacharya, Kaushik Kumar (2024): Description of a new species with three new records of Tomocerus Nicolet (Collembola: Tomoceridae) from India. Journal of Insect Biodiversity and Systematics 10 (4): 755-768, DOI: 10.61186/jibs.10.4.755, URL: https://doi.org/10.61186/jibs.10.4.755
03CA87C3FF91B014FFC9FA90FB8427F6.taxon	description	Description. Adult body length about 1.12 mm. Ground colour pale yellow to light brown (in alcohol), Ant. I – II distally purple pigmented, Ant. III – IV missing. Eye patches black and frontal area of head with purple pigment. Rest of body without any pigment. Body covered with distinctly striated rounded or truncated scales. Ratio of Ant. I: II: III – 1: 1.19: 5.23. Eyes 6 + 6, head diagonal: body length – 1: 4.47. Head chaetotaxy same as per Gong et al., 2018. Labral formula as 4 / 5, 5, 4; all smooth, anterior portion with four curved denticles. Labial palp with 4 chaetae, tip of lateral process not reaching the base. Unguis with basal paired, two lateral, one external and 4 – 5 internal teeth. Unguiculus lanceolate with an inner tooth, sometimes absent on fore claw (Fig. 7 A). Trochantero-femoral organ with 1 + 1 chaetae. Tita I – III with 8, 8, 7 spine-like chaetae respectively on its ventral side. Tenent hair clavate. Mac and bothriotricha of abdomen not very different from the original description. Retinaculum quadridentate with 19 chaetae on corpus. Manubrium: dens: mucro ratio as 2.87: 4.11: 1. Manubrium with large corner chaetae and 10 spines like chaetae laterally. Dental spine formula – 5 / 6, I; serrated spines with evenly distributed small denticles, basal spines gradually increase in length. Mucro setaceous with paired basal, 6 – 7 intermediate, large apical and slightly small subapical tooth (Fig. 7 B).	en	Mandal, Guru Pada, Kumari, Simran, Mandal, Pritha, Roy, Koushik Kumar, Suman, Kusumendra Kumar, Bhattacharya, Kaushik Kumar (2024): Description of a new species with three new records of Tomocerus Nicolet (Collembola: Tomoceridae) from India. Journal of Insect Biodiversity and Systematics 10 (4): 755-768, DOI: 10.61186/jibs.10.4.755, URL: https://doi.org/10.61186/jibs.10.4.755
03CA87C3FF91B014FFC9FA90FB8427F6.taxon	distribution	Distribution in India. West Bengal (Darjeeling). General distribution. Sino-Japanese region.	en	Mandal, Guru Pada, Kumari, Simran, Mandal, Pritha, Roy, Koushik Kumar, Suman, Kusumendra Kumar, Bhattacharya, Kaushik Kumar (2024): Description of a new species with three new records of Tomocerus Nicolet (Collembola: Tomoceridae) from India. Journal of Insect Biodiversity and Systematics 10 (4): 755-768, DOI: 10.61186/jibs.10.4.755, URL: https://doi.org/10.61186/jibs.10.4.755
03CA87C3FF91B014FFC9FA90FB8427F6.taxon	discussion	Remarks. The species is first time reported from India as well as the state of West Bengal. T. paraspinulus has close similarity with T. spinulus Chen & Christiansen, 1998 and T. pseudospinulus, though differs from them in having a blunt prominent chaeta on manubrium.	en	Mandal, Guru Pada, Kumari, Simran, Mandal, Pritha, Roy, Koushik Kumar, Suman, Kusumendra Kumar, Bhattacharya, Kaushik Kumar (2024): Description of a new species with three new records of Tomocerus Nicolet (Collembola: Tomoceridae) from India. Journal of Insect Biodiversity and Systematics 10 (4): 755-768, DOI: 10.61186/jibs.10.4.755, URL: https://doi.org/10.61186/jibs.10.4.755
03CA87C3FF9EB014FF39FB93FC1D2C08.taxon	materials_examined	Material examined. 12 specimens, INDIA, Arunachal Pradesh, Tawang district, Tawang, collected from moss and leaf litter surrounding snowflakes. 27 ° 36.918 ' N, 91 ° 51.913 ' E, 3417 m a. s. l., 01. iv. 2023, G. P. Mandal leg., Reg no. 3536 / H 14, deposited in NZC of ZSI.	en	Mandal, Guru Pada, Kumari, Simran, Mandal, Pritha, Roy, Koushik Kumar, Suman, Kusumendra Kumar, Bhattacharya, Kaushik Kumar (2024): Description of a new species with three new records of Tomocerus Nicolet (Collembola: Tomoceridae) from India. Journal of Insect Biodiversity and Systematics 10 (4): 755-768, DOI: 10.61186/jibs.10.4.755, URL: https://doi.org/10.61186/jibs.10.4.755
03CA87C3FF9EB014FF39FB93FC1D2C08.taxon	description	Description. Body length about 3.15 mm. Body scaled. Ground colour yellow with diffused light purple pigmentation on the anterior part of the head. Ant. I – II have slight purple infusion, Ant. III is gradually darker from base to tip continuing to Ant. IV. Eyes in purple patches. Th. II and Th. III with purple pigment on the lateral side (Fig. 8 A). PAO absent, 6 + 6 eyes present. Ant. I: II: III: IV ratio as 1: 1.4: 8.5. Ant. I and Ant. II with scales. Ant. III – IV with annulations instead of scales. Antenna: cephalic diagonal ratio 3.9: 1. Labral chaetae formula as 4 / 5, 5, 4. Dorsal cephalic chaetotaxy – anterior area: 2, 4; interocular area: 2, 7 and with central chaeta; postocular area: 2 + 2; posterior area; 2 + 2. Numerous small chaetae present on posterior head margin. Bothrioticha arrangement from Th. II – Abd V: 2, 1, 0, 0, 1, 2, 0. Body macrochaetae arrangement formula as Th. II – Abd V: 3, 3, 3, 3, 4, 2, 4. Th. II – Abd V each segment with one pseudopore. Trochantero-femoral organ with 1, 1 slender chaetae. Strong ventral chaetae formula of tita I – III as 6, 6, 7. Tenent hair clavate; 2 minute accessory chaetae, 2 thin and long guard chaetae present. Unguis with a basal toothlet, 5 inner distal teeth. One internal tooth present on unguiculus. VT scaled with numerous chaetae. Tenaculum quadridentate, anterior face of corpus with 5 – 6 chaetae. Ratio of Manubrium: dens: mucro – 2.95: 4.05: 1. Manubrium with large round scales and 9 – 10 stout spinous chaetae. Dental spine formula – 5 / 4, II; moderate size numerous denticles present, distal spine strongest (Fig. 8 B). Outer lamella of mucro with with 4 – 5 intermediate teeth (Fig. 8 D).	en	Mandal, Guru Pada, Kumari, Simran, Mandal, Pritha, Roy, Koushik Kumar, Suman, Kusumendra Kumar, Bhattacharya, Kaushik Kumar (2024): Description of a new species with three new records of Tomocerus Nicolet (Collembola: Tomoceridae) from India. Journal of Insect Biodiversity and Systematics 10 (4): 755-768, DOI: 10.61186/jibs.10.4.755, URL: https://doi.org/10.61186/jibs.10.4.755
03CA87C3FF9EB014FF39FB93FC1D2C08.taxon	discussion	Remarks. In our collection, one specimen has one side of the dens with a malfunctioned arrangement (Fig. 8 C) of dental spines in a zig-zag row. The species has similarity with T. ocreatus Denis, 1948, T. leyensis Yu & Deharveng, 2018, T. virgatus Yu, Qin, Ding, Hu, Zhang & Liu, 2018, T. yueluensis Yu, Qin, Ding, Hu, Zhang & Liu, 2018, in shape of dental spines, but differs from them in number of dental spines, mucronal intermediate teeth and unguis inner teeth number.	en	Mandal, Guru Pada, Kumari, Simran, Mandal, Pritha, Roy, Koushik Kumar, Suman, Kusumendra Kumar, Bhattacharya, Kaushik Kumar (2024): Description of a new species with three new records of Tomocerus Nicolet (Collembola: Tomoceridae) from India. Journal of Insect Biodiversity and Systematics 10 (4): 755-768, DOI: 10.61186/jibs.10.4.755, URL: https://doi.org/10.61186/jibs.10.4.755
