identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03CA87E35C48FFAB6F99FDC0FBBEFCB9.text	03CA87E35C48FFAB6F99FDC0FBBEFCB9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Coleoptera Linnaeus 1758	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> (  COLEOPTERA , SCRABAEIDAE) </p>
            <p>S. A. Shabalin 1*), L. A. Akhmetova 2), A. V. Frolov 2)</p>
            <p>1) Federal Scientific Center of the East Asia Terrestrial Biodiversity, Far East Branch of the Russian Academy of Sciences, Vladivostok, 690022, Russia. * Corresponding author. Email: oxecetonia@mail.ru</p>
            <p>2) Zoological Institute, Russian Academy of Sciences, St. Petersburg, 199034, Russia. Email: Lilia.Akhmetova@zin.ru, afrolov@zin.ru</p>
            <p> Summary. The dung-beetle species  Aphodius botulus Balthasar, 1945 was recorded in Russia for the first time based on the specimens collected from Primorskii krai. Photographs of the imago habitus, male genitalia, and a map of collecting localities in Russia are provided. </p>
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	https://treatment.plazi.org/id/03CA87E35C48FFAB6F99FDC0FBBEFCB9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Shabalin, S. A.;Akhmetova, L. A.;Frolov, A. V.	Shabalin, S. A., Akhmetova, L. A., Frolov, A. V. (2024): NEW TO RUSSIA SPECIES OF THE GENUS APHODIUS (COLEOPTERA, SCRABAEIDAE). Far Eastern Entomologist 509: 7-10, DOI: 10.25221/fee.509.2, URL: https://doi.org/10.25221/fee.509.2
03CA87E35C49FFA96EADFEF4FBA0FEF0.text	03CA87E35C49FFA96EADFEF4FBA0FEF0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Aphodius (Phalacronothus) botulus Balthasar 1945	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Aphodius (Phalacronothus) botulus Balthasar, 1945</p>
            <p>Figs 1–4</p>
            <p> Aphodius (Paragolius) botulus Balthasar, 1945: 43 . Type locality “Mandschukuo, circum oppidium Charbin” [= China: Heilongjiang]; Balthasar, 1964: 281. </p>
            <p> Aphodius (Phalacronothus) botulus : Kim, 1989: 3; Takahashi, 1999: 34; Dellacasa &amp; Dellacasa, 2006: 134; Murata et al., 2011: 84; Kim, 2012: 126. </p>
            <p> Phalacronothus botulus : Dellacasa &amp; Dellacasa, 2016: 145. </p>
            <p> Aphodius (Orodalus) naraensis Nakane, 1956: 120 . Type locality “ Nara, Yamato, Honshu” [= Japan: Honshu]; Balthasar, 1964: 201; Stebnicka, 1980: 239. Synonymized by Stebnicka, 1982: 78. </p>
            <p> Aphodius (Amidorus) manschuriensis Petrovitz, 1958: 138 . Type locality ”Manschurei: Charbin” [= China: Heilongjiang]. Synonymized by Balthasar, 1964: 281. </p>
            <p> Liothorax plagiatus auct.: Shabalin, 2020: 188; Shabalin, 2021: 625; Shabalin, 2022: 477; Shabalin, 2023: 20; Shabalin, 2024: 256. </p>
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                  MATERIAL. Russia: Primorskii krai: valley of  Ryazanovka River (42°82' N, 131°23' E), 29. V 2018, 19 specimens in sheep dung, leg. S. Shabalin  ;  same locality, 7. VI 2018, 9 specimens in sheep and cow dung, leg. S. Shabalin ;  same locality, 15. VI 2018, 5 specimens in sheep and cow dung, leg. S. Shabalin ;  same locality, 6.VII 2018, 1 specimen in sheep dung, leg. S. Shabalin ;   
                <a title="Search Plazi for locations around (long 132.66667/lat 43.533333)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=132.66667&amp;materialsCitation.latitude=43.533333">Safari-Park</a>
                 (43°32' N, 132°40' E), 8. VI 2020, 3 specimens in deer dung, leg. S. Shabalin  ;   5 km E  
                <a title="Search Plazi for locations around (long 132.86667/lat 43.383335)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=132.86667&amp;materialsCitation.latitude=43.383335">Romanovka</a>
                 (43°23' N, 132°52' E) 15. V 2019, 6 specimens in cow dung, leg. S. Shabalin  ;   vicinity  
                <a title="Search Plazi for locations around (long 133.23334/lat 43.0)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=133.23334&amp;materialsCitation.latitude=43.0">Peretino</a>
                 (43°00' N, 133°14' E), 15. V 2019, 1 specimen in cow dung, leg. S. Shabalin  ;   16 km SSE  
                <a title="Search Plazi for locations around (long 133.3/lat 43.1)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=133.3&amp;materialsCitation.latitude=43.1">Partizansk</a>
                 (43°06' N, 133°18' E), 15. V 2019, 19 specimens in sheep dung, leg. S. Shabalin  ;  same locality, 24. V 2019, 11 specimens in sheep dung, leg. S. Shabalin ;  same locality, 3. VI 2019, 8 specimens in sheep dung, leg. S. Shabalin ;  same locality, 12. VI 2019, 2 specimens in sheep dung, leg. S. Shabalin ;   15 km NNE  
                <a title="Search Plazi for locations around (long 133.28334/lat 43.433334)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=133.28334&amp;materialsCitation.latitude=43.433334">Partizansk</a>
                 (43°26' N, 133°17' E), 15. V 2019, 202 specimens in horse and cow dung, leg. S. Shabalin  ;  same locality, 24. V 2019, 8 specimens in horse and cow dung, leg. S. Shabalin ;  same locality, 3. VI 2019, 7 specimens in horse and cow dung, leg. S. Shabalin ;   
                <a title="Search Plazi for locations around (long 133.23334/lat 44.05)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=133.23334&amp;materialsCitation.latitude=44.05">Novogordeevka</a>
                 (44°03' N, 133°14' E), 15. V 2020, 1 specimen in horse dung, leg. S. Shabalin  ;  same locality, 23. V 2020, 1 specimen in horse dung, leg. S. Shabalin ;   vicinity of  Kornilovka , 15. V 2020, 2 specimens in cow dung, leg. S. Shabalin  ;  same locality, 23. V 2020, 1 specimen cow dung, leg. S. Shabalin.  Four specimens from the valley of Ryazanovka River are deposited in the collection of the Zoological Institute (Saint Petersburg), but the rest material deposited in the collection of the Federal Scientific Center of the  East Asia Terrestrial Biodiversity (Vladivostok)  . 
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            <p>DISTRIBUTION. Russia (first record, Primorskii krai), Japan (Honshu, Kyushu), North Korea, China (Heilongjiang, Jilin).</p>
            <p> NOTES. Adults of  Aphodius botulus can be separated from other species of subgenus  Phalacronothus recorded from Russia, namely  A. biguttatus Germar, 1824 ,  A. citellorum Semenov et Medvedev, 1928 , and  A. quadrimaculatus , by denser and coarser punctation of head, pronotum and elytra, and by the shape of the parameres. In Primorskii krai,  A. botulus occurs along with the more abundant  A. pusillus , and can be confused with it. It differs from the latter, apart from the characters mentioned above, also in having a hooked protibial spur in males and in a smaller average size. In North Korea, the beetles were found in sheep and cow dung (Stebnicka, 1980). In addition to the dung of abovementioned animals, in Russia the beetles were also found in horse and deer dung. Our data suggest that  A. botulus is a generalist coprophagous species with imago active mostly in late spring and early summer. </p>
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	https://treatment.plazi.org/id/03CA87E35C49FFA96EADFEF4FBA0FEF0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Shabalin, S. A.;Akhmetova, L. A.;Frolov, A. V.	Shabalin, S. A., Akhmetova, L. A., Frolov, A. V. (2024): NEW TO RUSSIA SPECIES OF THE GENUS APHODIUS (COLEOPTERA, SCRABAEIDAE). Far Eastern Entomologist 509: 7-10, DOI: 10.25221/fee.509.2, URL: https://doi.org/10.25221/fee.509.2
