taxonID	type	description	language	source
03D71761B676FFE8FC95F987114BF917.taxon	type_taxon	TYPE SPECIES. — Glyphiulus cavernicolus Silvestri, 1923, by monotypy [type locality: Bidi Caves, Sarawak, Borneo].	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, Den Spiegel, Didier Van (2009): Review of the millipede genusPlusioglyphiulusSilvestri, 1923, with descriptions of new species from southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Zoosystema 31 (1): 71-116, DOI: 10.5252/z2009n1a5, URL: http://dx.doi.org/10.5252/z2009n1a5
03D71761B676FFE8FC95F987114BF917.taxon	diagnosis	DIAGNOSIS. — 1. Male legs 1 generally strongly reduced, but remaining parts enlarged; sternite with a single, high, median process, strongly unciform in adults, directed frontally and usually supplied with several strong setae at the base on frontal face; coxites usually subconical and setose; telopodites completely reduced, only rarely visible as a minute knob distomedially. This is one of the basic autapomorphies of the genus. 2. Male legs 2 usually strongly enlarged, flattened sagittally, densely hairy on caudal face; tarsus stout, shortened; penes slender, subconical, devoid of setae. This is another strong autapomorphy of the genus. 3. Male legs 3 usually with elongated coxae, but with shortened and compact telopodites. This is a weaker autapomorphy of the genus, since certain species in some other genera of Cambalopsidae show similar conditions. 4. Both collum and segment 2 often very strongly enlarged, as wide as segments of posterior third of body or nearly so, thus making postcollar constriction remarkably narrow. This is a strong autapomorphy of the genus. 5. Pleural regions of segments 2 - 4 conspicuously expanded ventrad. This is a weak autapomorphy, because several species from other genera of Cambalopsidae show similar conditions. 6. Ventral extensions of male segment 7 especially prominent, usually with ends turned abruptly caudad. This is a weak autapomorphy, because several species from other genera of Cambalopsidae show similar conditions. 7. Carinotaxy patterns of collum highly variable. Crests proper virtually never obliterated or reduced, very often subdivided into 2 - 4 parts, often with 1 - 3 intercalary tubercles or short crests; usually 10 crests reaching anterior margin. This is a strong autapomorphy of the genus. 8. The typical carinotaxy pattern of the metaterga is 3 / 3 + I / i + 3 / 3 (4) / 3 + m / (m) / m + 3 / 3 (4) / 3 + I / i + 3 / 3, thus reflecting the fact that all or most of the dorsal crests or tubercles are arranged in three transverse rows (except P. bedosae n. sp. and P. pallidior n. sp.). The middle transverse row in adults is sometimes not in line with the two other rows, being intercalary; this row seems to only be attained at or towards the onset of adulthood. The median crest is usually lower than others, never doubled anteriorly. This is a relatively weak autapomorphy, because some species from other genera (e. g., Glyphiulus bedosae Golovatch, Geoffroy, Mauriès & Van den Spiegel, 2007) also show three transverse rows of tubercles on the metaterga, whereas two rows seem to regularly occur in immature stadia of some Plusioglyphiulus species. 9. The anterior gonopods usually have a plate-like sternum or coxosternum, normally without a median outgrowth; coxites separate or contiguous medially, but never fused, each with one or two distinct processes; telopodites moveable, elongate, subcylindrical, lateral to caudolateral in position, 1 - segmented, often curved and setose. This is a rather strong, but not absolute autapomorphy of the genus, because in many Cambalopsidae representing various genera the anterior gonopods look alike. 10. Posterior gonopods smaller than anterior ones, but usually elongated, not compressed; coxite normally showing an arm or its traces, a shelf-like structure on frontal face and a fovea subapically, devoid of the often plumose flagella that are so characteristic of Glyphiulus species. This seems to be a weak autapomorphy, since the variation range in posterior gonopod conformation in Plusioglyphiulus is unusually wide. 11. Clypeus with three teeth anteromedially, only seldom variable (P. pallidior n. sp.). This is a typical condition occurring throughout Juliformia. 12. Promentum always well separated from eumentum.	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, Den Spiegel, Didier Van (2009): Review of the millipede genusPlusioglyphiulusSilvestri, 1923, with descriptions of new species from southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Zoosystema 31 (1): 71-116, DOI: 10.5252/z2009n1a5, URL: http://dx.doi.org/10.5252/z2009n1a5
03D71761B670FFE4FD7DF9ED1278FAC3.taxon	description	REDESCRIPTION Male about 32 mm long and 1.5 mm wide, with 58 p + 3 a + T, collum being broadest; females about 35 and 38 mm long, 1.6 and 1.9 mm wide, with 63 p + 3 a + T and 78 p + 2 a + T, respectively, segments of posterior third of body being broadest. Coloration, including that of collum and segment 2, uniformly grey-brown to brown, dorsal crests and ozoporiferous tubercles usually dark brown. Surface smooth and polished on frons, very delicately shagreened above. Antennae short and clavate (Fig. 3 A), antennomeres 6 and 7 each with a large group of bacilliform sensilla distodorsally (Fig. 2 C). Ocellaria transversely subreniform, with 9 - 12 blackish ocelli in 5 or 6 longitudinal rows, but without adjacent darker spots dorsally. Gnathochilarium oligotrichous, each lamella lingualis with 3 or 4 setae. Postcollar constriction very evident, due to especially enlarged collum and segment 2 (Fig. 1 B). Carinotaxy formula of collum: 1 p / t + 2 p / t + 3 p / t + (ta) / (t) + 4 p / t / t + (ta) / (t) + 5 p / t / t + (ta) / (t) + pp / t / t + ma / t (Fig. 1 A, B). Carinotaxy of metatergum 2, 7 / 7 + m / m + 7 / 7; of metaterga 3 and 4, 7 / 7 + M (m / m) + 7 / 7; of metatergum 5 and subsequent metaterga, except last few, 3 / 3 + I / i + 3 / 3 + m / m + 3 / 3 + I / i + 3 / 3; of apodous segments, usually 7 + m + 7 (Fig. 1); all crests and tubercles, including poriferous cones, rather low; poriferous tubercles (I) only slightly higher, broader than high, but median ones (m) slightly lower than others (Figs 1 C, D; 2 B). Dorsal crests on several posteriormost segments considerably higher than others (Fig. 1 E). Midbody segments circular in cross-section (Fig. 2 B). Pleural regions of segments 2 - 4 conspicuously expanded, flap-shaped, especially so on segment 3 (Fig. 1 A). An evident, transverse pleural ridge behind gonopod aperture on male segment 7, with rounded flaps bent abruptly caudad. Tegument of metazona delicately alveolate-areolate and dull throughout, stricture between pro- and metazona striolate longitudinally, other parts of prozona very finely shagreened. Metatergal setae absent. Limbus extremely finely and more or less regularly denticulate. Epiproct broadly rounded and cariniform apically, with 2 + m + 2 tubercles dorsally at midway, of which the lateralmost and, especially, the median can be very low; paraprocts regularly convex, devoid of an elevation at midline; hypoproct emarginate at caudal margin (Fig. 2 A). Legs rather short, about 2 / 3 as long as body diameter (Fig. 2 B), claws with an evident spine near base (Figs 2 D; 3 B). Tarsi very delicately fringed apically; most of distal setae with scattered denticles. Male legs 1 with elongate, conical, setose coxites and a single, very large, falcate, median sternal process with several long setae at base on frontal face. Male legs 2 evidently enlarged, coxae deeply emarginate distally on caudal face, telopodite densely hirsute on frontal face; penes conical, elongated, bare, fused at base. Male legs 3 with particularly elongate and slender coxae, but with compact telopodites. Anterior gonopods (Fig. 3 C, D) with a long, slender, median sternal process (st); coxite with a paramedian pair of low inner processes (cxp 1) and another pair of long, slender, outer, distally beak-shaped processes (cxp 2) with a short uncus at base; telopodites (te) slender, curved, attached to coxal region caudolaterally, capable of movement, almost non-setose even subapically. Posterior gonopods small, rather simple, coxites well separated from sternum, at least on caudal face (Fig. 3 E), each with a spiniform structure medio-subapically and a complex fringed apex.	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, Den Spiegel, Didier Van (2009): Review of the millipede genusPlusioglyphiulusSilvestri, 1923, with descriptions of new species from southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Zoosystema 31 (1): 71-116, DOI: 10.5252/z2009n1a5, URL: http://dx.doi.org/10.5252/z2009n1a5
03D71761B670FFE4FD7DF9ED1278FAC3.taxon	discussion	REMARKS Hoffman (1977) described this species from Batu Caves, so the above material can be considered as near-topotypic. It agrees well with the original description, but differs slightly in showing only 7 / 7 + m / m + 7 / 7, not 8 / 8 + m / m + 8 / 8, tubercles on segments 2 - 4, sometimes a lower number of dorsal tubercles (4 or 5, not 7) on the epiproct, and the presence on the legs of a strong spine at the base of the claw. Any obvious troglomorphic traits in this species seem to be absent.	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, Den Spiegel, Didier Van (2009): Review of the millipede genusPlusioglyphiulusSilvestri, 1923, with descriptions of new species from southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Zoosystema 31 (1): 71-116, DOI: 10.5252/z2009n1a5, URL: http://dx.doi.org/10.5252/z2009n1a5
03D71761B67CFFE0FF73FA7E10B7FA26.taxon	description	(Figs 4 - 7)	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, Den Spiegel, Didier Van (2009): Review of the millipede genusPlusioglyphiulusSilvestri, 1923, with descriptions of new species from southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Zoosystema 31 (1): 71-116, DOI: 10.5252/z2009n1a5, URL: http://dx.doi.org/10.5252/z2009n1a5
03D71761B67CFFE0FF73FA7E10B7FA26.taxon	materials_examined	TYPE MATERIAL. — Vietnam. Dong Nai Prov., Dinh Quan, 11.27 ° N, 107.22 ° E, lava tube 1, 23. XII. 1994, leg. L. Deharveng, Truong Quang Tam & A. Bedos (VIET- 048), holotype ♂ (MNHN GA 055); 1 ♂ paratype (MNHN GA 055); 1 ♂ paratype (SEM); 1 ♂ paratype (ZMUM).	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, Den Spiegel, Didier Van (2009): Review of the millipede genusPlusioglyphiulusSilvestri, 1923, with descriptions of new species from southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Zoosystema 31 (1): 71-116, DOI: 10.5252/z2009n1a5, URL: http://dx.doi.org/10.5252/z2009n1a5
03D71761B67CFFE0FF73FA7E10B7FA26.taxon	etymology	ETYMOLOGY. — To emphasize the ampulliform distal part of the posterior gonopods.	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, Den Spiegel, Didier Van (2009): Review of the millipede genusPlusioglyphiulusSilvestri, 1923, with descriptions of new species from southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Zoosystema 31 (1): 71-116, DOI: 10.5252/z2009n1a5, URL: http://dx.doi.org/10.5252/z2009n1a5
03D71761B67CFFE0FF73FA7E10B7FA26.taxon	diagnosis	DIAGNOSIS. — Differs from congeners by certain details of gonopod structure (very characteristic shapes of the various outgrowths and processes), coupled with the isostichic carinotaxy of the collum and of other body segments, the flattened paraprocts, the broadly rounded epiproct and the presence of spots dorsad of the ocellaria.	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, Den Spiegel, Didier Van (2009): Review of the millipede genusPlusioglyphiulusSilvestri, 1923, with descriptions of new species from southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Zoosystema 31 (1): 71-116, DOI: 10.5252/z2009n1a5, URL: http://dx.doi.org/10.5252/z2009n1a5
03D71761B67CFFE0FF73FA7E10B7FA26.taxon	description	DESCRIPTION Length 30 - 35 mm, width 2.0 - 2.2 mm, collum being broadest. Coloration mainly dirty brown to red-brown, only collum and, in part, head and segment 2 contrastingly yellowish; metatergal crests / tubercles usually dark brown to blackish, due to heavy sclerotization; ocellaria black, head dorsad of ocellaria with 1 + 1, nearly contiguous, marbled brown spots; antennomeres 5 (distally) to 7 (entirely) usually dark brown; both a narrow axial line and a broader stripe level to ozopores grey-brown, usually distinct; legs and venter yellow-brown, only distal podomeres slightly reddish. Body with 55 - 72 p + 2 a + T. Length of holotype about 30 mm, width 2.0 mm, with 58 p + 2 a + T. Antennae relatively short and clavate (Fig. 7 A), antennomeres 6 and 7 each with a small group of bacilliform sensilla distodorsally (Fig. 7 A). Ocellaria black, ovoid in shape, with 18 - 20 ocelli in 5 or 6 longitudinal rows. Gnathochilarium (Fig. 4 C) oligotrichous. Postcollar constriction very evident, due to especially enlarged collum and segment 2 (Fig. 4 B). Carinotaxy formula of collum: I + t / t + 3 p / t + IV + V (a) + VI + 7 a + P + ma (Fig. 4 A, B). Carinotaxy of metatergum 2, 7 / 7 + m / m + 7 / 7, anterior parts of all crests except “ m ” transverse, those of “ m ” longitudinal; formula of metaterga 3 and 4, 4 / 4 + 3 + m + 3 + 4 / 4, of which 3 + m + 3 rounded and placed transversely; formula of metatergum 5, 3 / 3 + I / i + 3 / 3 + m + 3 / 3 + I / i + 3 / 3; formula of subsequent metaterga, except last few, 3 / 3 / 3 + I / i + 3 / 3 / 3 + m / m + 3 / 3 / 3 + I / i + 3 / 3 / 3; formula of last ozoporiferous metatergum, 3 / 3 + I / i + 3 / 3 + m / m + 3 / 3 + I / i + 3 / 3; that of apodous segments, 2 + 5 / 5 + m / m + 5 / 5 + 2 (Fig. 4 A-E); all crests and tubercles, including poriferous cones, rather low. Dorsal crests on several posteriormost segments considerably higher than others (Fig. 4 E). Midbody segments circular in cross-section (Fig. 5 B). Pleural regions of segments 2 - 4 conspicuously expanded, flap-shaped, especially so on segment 3 (Fig. 4 A). An evident, transverse pleural ridge behind gonopod aperture on male segment 7, with rounded flaps bent abruptly caudad (Fig. 5 A). Tegument and limbus as in P. grandicollis. Epiproct broadly rounded apically, with 1 + 1 paramedian tubercles at midway; paraprocts evidently flattened, without an elevation at midline; hypoproct emarginate at caudal margin (Fig. 4 E, F). Legs short, about half as long as body diameter (Fig. 5 B), claws with an evident spine near base (Fig. 7 F). Tarsi and tarsal setae as in P. grandicollis. Male legs 1 - 3 as in P. grandicollis. Male legs 1 with elongate, setose coxites and a single, very large, falcate, median sternal process with several long setae at base on frontal face (Figs 5 C; 7 B, C). Male legs 2 evidently enlarged, telopodite hirsute on frontal face; penes subtriangular, fused at base (Figs 5 D; 7 D). Male legs 3 with particularly elongate and slender coxae, and shortened telopodites (Fig. 7 E). Anterior gonopods with a paramedian pair of high, slender, apically curved, contiguous but free coxosternal processes (cxp 1) anteriorly, each with several lateral setae in basal half; coxal region produced mesad into a slender sigmoid process (cxp 2); telopodites (te) elongate, attached to coxal region on caudal face, probably capable of movement, carrying a few setae subapically and tridentate apically (Figs 6 A, B; 7 G). Posterior gonopods small, complex, coxites well separated from sternum, at least on caudal face (Figs 6 C, D; 7 H); frontal face of each coxite with a strong, dentate arm (d) at midway and a complex ampulla-like structure with several teeth nearby.	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, Den Spiegel, Didier Van (2009): Review of the millipede genusPlusioglyphiulusSilvestri, 1923, with descriptions of new species from southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Zoosystema 31 (1): 71-116, DOI: 10.5252/z2009n1a5, URL: http://dx.doi.org/10.5252/z2009n1a5
03D71761B67CFFE0FF73FA7E10B7FA26.taxon	discussion	REMARKS Any obvious troglomorphic traits in this species seem to be absent.	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, Den Spiegel, Didier Van (2009): Review of the millipede genusPlusioglyphiulusSilvestri, 1923, with descriptions of new species from southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Zoosystema 31 (1): 71-116, DOI: 10.5252/z2009n1a5, URL: http://dx.doi.org/10.5252/z2009n1a5
03D71761B678FFE3FCB3FA061740FB64.taxon	description	(Figs 8 - 10)	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, Den Spiegel, Didier Van (2009): Review of the millipede genusPlusioglyphiulusSilvestri, 1923, with descriptions of new species from southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Zoosystema 31 (1): 71-116, DOI: 10.5252/z2009n1a5, URL: http://dx.doi.org/10.5252/z2009n1a5
03D71761B678FFE3FCB3FA061740FB64.taxon	materials_examined	TYPE MATERIAL. — Laos. Luang Prabang Prov., 19.88 ° N, 102.14 ° E, W of Tham, 3. VIII. 1992, leg. J. - P. Besson, holotype ♂ (MNHN GA 056); 1 ♀ paratype (MNHN GA 056); 1 ♂ paratype (SEM).	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, Den Spiegel, Didier Van (2009): Review of the millipede genusPlusioglyphiulusSilvestri, 1923, with descriptions of new species from southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Zoosystema 31 (1): 71-116, DOI: 10.5252/z2009n1a5, URL: http://dx.doi.org/10.5252/z2009n1a5
03D71761B678FFE3FCB3FA061740FB64.taxon	etymology	ETYMOLOGY. — To emphasize the foveate apex of the posterior gonopods.	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, Den Spiegel, Didier Van (2009): Review of the millipede genusPlusioglyphiulusSilvestri, 1923, with descriptions of new species from southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Zoosystema 31 (1): 71-116, DOI: 10.5252/z2009n1a5, URL: http://dx.doi.org/10.5252/z2009n1a5
03D71761B678FFE3FCB3FA061740FB64.taxon	diagnosis	DIAGNOSIS. — Differs from congeners by certain details of gonopod structure (characteristic shapes of the different outgrowths and processes), coupled with the mixostichic carinotaxy of the collum, the isostichic carinotaxy of other body segments, the convex paraprocts, the broadly rounded epiproct and the presence of spots dorsad of the ocellaria.	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, Den Spiegel, Didier Van (2009): Review of the millipede genusPlusioglyphiulusSilvestri, 1923, with descriptions of new species from southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Zoosystema 31 (1): 71-116, DOI: 10.5252/z2009n1a5, URL: http://dx.doi.org/10.5252/z2009n1a5
03D71761B678FFE3FCB3FA061740FB64.taxon	description	DESCRIPTION Length 28 - 35 mm, width 1.3 - 1.9 mm, collum and midbody segments being equally broadest. Coloration whitish to yellow-brown; collum and, partly, head and segment 2 entirely pale; tips of most of tergal crests / tubercles, as well as a narrow axial line and a slightly broader lateral stripe level to ozopores pale grey-brown to brown; ocellaria dark brown with pale to dark brown, widely separated spots dorsally. Body with 49 - 66 p + 4 - 1 a + T. Length of holotype about 30 mm, width 1.3 mm, with 58 p + 2 a + T. Antennae rather short and clavate (Fig. 10 A), antennomeres 6 and 7 each with a considerable group of bacilliform sensilla distodorsally; similar but minute sensilla present also on antennomere 7 (Fig. 9 C). Ocellaria black, ovoid in shape, with 10 - 13 ocelli in 5 - 7 longitudinal rows. Gnathochilarium oligotrichous. Postcollar constriction evident, but collum and segment 2 relatively modestly enlarged (Fig. 8 B). Carinotaxy formula of collum: ta / t + 2 p / t + 3 p / t + 4 p / t / t + ta / 5 / t + (ta) / t + VI (p) / t + pp / t / t + ma / t (Fig. 8 A, B). Carinotaxy of metaterga 2 - 4, 7 / 7 + m / m + 7 / 7, all crests being longitudinal; formula of metaterga 5 to 15, 3 / 3 + I / i + 3 / 3 / 3 + m / m + 3 / 3 / 3 + I / i + 3 / 3; formula of subsequent metaterga usually 3 / 3 + I / i + 3 / 3 / 3 + m / (m) / m + 3 / 3 / 3 + I / i + 3 / 3; that of apodous segments, 3 + 4 / 4 + m / m + 4 / 4 + 3 (Fig. 8); all crests and tubercles rather high, poriferous cones (I) much higher, nearly as high as broad, but median ones (m) very low (Fig. 9 B). Dorsal crests on several posteriormost segments considerably higher than others (Fig. 8 E). Midbody segments circular in cross-section (Fig. 9 B). Pleural regions of segments 2 - 4 conspicuously expanded, flap-shaped, especially so on segment 3 (Fig. 8 A). An evident, transverse pleural ridge behind gonopod aperture on male segment 7, with rounded flaps bent abruptly caudad. Tegument and limbus as in P. grandicollis or P. ampullifer n. sp. Epiproct broadly rounded and cariniform apically, with 2 + 2 paramedian tubercles at midway; paraprocts regularly convex, not flattened; hypoproct emarginate at caudal margin (Fig. 9 A). Legs short, on midbody segments about ⅔ as long as segment diameter (Fig. 9 B). Claw at base with a strong accessory spine almost half as long as claw itself (Figs 9 D; 10 E). Tarsi, tarsal setae and male legs 1 - 3 (Fig. 10 B-D) as in P. grandicollis. Anterior gonopods (Fig. 10 F) much like in P. ampullifer n. sp., but processes cxp 1 large, non-contiguous, flattened sagittally and each carrying a short medial uncus apically; processes cxp 2 slightly curved, slender and bifid; telopodites (te) simple, movable, elongate-subconical, setose laterally in distal half. Posterior gonopods (Fig. 10 G) relatively simple, with a pair of strongly reduced telopodites (te) laterally at base; coxite with short denticles (d) instead of arms, fovea-shaped apically.	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, Den Spiegel, Didier Van (2009): Review of the millipede genusPlusioglyphiulusSilvestri, 1923, with descriptions of new species from southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Zoosystema 31 (1): 71-116, DOI: 10.5252/z2009n1a5, URL: http://dx.doi.org/10.5252/z2009n1a5
03D71761B67BFFFDFCADFAC4127AF961.taxon	description	(Figs 11 - 13)	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, Den Spiegel, Didier Van (2009): Review of the millipede genusPlusioglyphiulusSilvestri, 1923, with descriptions of new species from southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Zoosystema 31 (1): 71-116, DOI: 10.5252/z2009n1a5, URL: http://dx.doi.org/10.5252/z2009n1a5
03D71761B67BFFFDFCADFAC4127AF961.taxon	materials_examined	TYPE MATERIAL. — Thailand. Nan, 18.776 ° N, 100.783 ° E, Cave Th am Pha Tup (Tham Phra), 10 - 11. VIII. 1992, leg. J. - P. Besson, holotype ♂ (MNHN GA 057); 2 ♂♂ paratypes, 1 ♀ paratype (MNHN GA 057); 1 ♀ paratype (ZMUM); 1 ♀ paratype (SEM).	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, Den Spiegel, Didier Van (2009): Review of the millipede genusPlusioglyphiulusSilvestri, 1923, with descriptions of new species from southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Zoosystema 31 (1): 71-116, DOI: 10.5252/z2009n1a5, URL: http://dx.doi.org/10.5252/z2009n1a5
03D71761B67BFFFDFCADFAC4127AF961.taxon	etymology	ETYMOLOGY. — To honour Jean-Pierre Besson, who collected this material.	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, Den Spiegel, Didier Van (2009): Review of the millipede genusPlusioglyphiulusSilvestri, 1923, with descriptions of new species from southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Zoosystema 31 (1): 71-116, DOI: 10.5252/z2009n1a5, URL: http://dx.doi.org/10.5252/z2009n1a5
03D71761B67BFFFDFCADFAC4127AF961.taxon	diagnosis	DIAGNOSIS. — Differs from congeners by certain details of gonopod structure (complicated anterior gonopods versus strongly simplified posterior ones), coupled with the mixostichic carinotaxy of the collum, the isostichic carinotaxy of other body segments, the convex paraprocts, the subtrapeziform epiproct and the absence of spots dorsad of the ocellaria.	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, Den Spiegel, Didier Van (2009): Review of the millipede genusPlusioglyphiulusSilvestri, 1923, with descriptions of new species from southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Zoosystema 31 (1): 71-116, DOI: 10.5252/z2009n1a5, URL: http://dx.doi.org/10.5252/z2009n1a5
03D71761B67BFFFDFCADFAC4127AF961.taxon	description	DESCRIPTION Length 25 - 43 mm, width 1.7 - 2.0 mm, collum and midbody segments being equally broadest. Coloration usually uniformly grey-brown, sometimes with a paramedian pair of more or less wide, grey-yellowish stripes; collum and, in part, head and segment 2 entirely pale; tips of tergal crests / tubercles usually dark brown; ocellaria dark brown to blackish, devoid of adjacent spots dorsally. Body with 56 - 75 p + 4 - 1 a + T. Length of holotype about 34 mm, width 2.0 mm, with 61 p + 4 a + T. Antennae more elongate (Fig. 13 A), antennomeres 6 and 7 each with a small group of bacilliform sensilla distodorsally. Ocellaria transverse and ribbonshaped, with 10 - 13 ocelli in 5 - 7 longitudinal rows. Gnathochilarium oligotrichous. Postcollar constriction evident, but collum and segment 2 relatively modestly enlarged (Fig. 11 A). Carinotaxy formula of collum: ta / t + 2 p / t + // (t) + 3 p / t / t + 4 p / t / t + (ta) / (t) / (t) + 5 p / t / t + (ta) / t + pp / t + ma / t (Fig. 11 A). Carinotaxy of metaterga 2 and 3, 8 / 8 + m / m + 8 / 8, all crests being longitudinal; formula of metatergum 4, 2 + 6 / 6 + m / m + 6 / 6 + 2; formula of metaterga 5 - 7, 3 / 3 + I / i + 3 / 3 / 3 + m / m + 3 / 3 / 3 + I / i + 3 / 3; formula of subsequent metaterga, 3 / 3 + I / i + 3 / 3 / 3 + m / m / m + 3 / 3 / 3 + I / i + 3 / 3; that of apodous segments, 3 (4) + 4 (3) / 4 (3) + M (m / m) + 4 (3) / 4 (3) + 3 (4) (Fig. 11 A, C-F); all crests and tubercles rather high, poriferous cones (I) only slightly higher, and median ones (m) slightly lower, than others (Fig. 11 B-E). Dorsal crests on several posteriormost segments considerably higher than others (Fig. 11 E). Midbody segments circular in cross-section (Fig. 11 B). Pleural regions of segments 2 - 4 conspicuously expanded, flap-shaped, especially so on segment 3 (Fig. 11 A). Pleural ridge behind gonopod aperture on male segment 7, tegument and an extremely finely and more or less regularly denticulate limbus (Fig. 12 C) as in other congeners. Epiproct relatively long, produced caudally and broadly subtrapeziform apically, with 1 + 1 paramedian tubercles at midway; paraprocts regularly convex, not flattened; hypoproct emarginate at caudal margin (Figs 11 E; 12 A). Legs short, on midbody segments about half as long as segment diameter (Fig. 13 E). Claw at base with a strong accessory spine almost half as long as claw itself (Figs 12 B, D; 13 E). Tarsi, tarsal setae and male legs 1 - 3 (Fig. 13 B-D) as in P. grandicollis. Anterior gonopods (Fig. 13 F) much like in P. ampullifer n. sp., but processes cxp 1 spiniform and non-contiguous; processes cxp 2 slightly curved, broad, each with a flange on frontal face; telopodites (te) simple, lateral in position, movable, subcylindrical, setose laterally in distal half. Posterior gonopods (Fig. 13 G) relatively simple, without traces of telopodites; each coxite with a short sacciform structure (d) instead of arms, fovea-shaped apically and supplied with a sublateral spinicle.	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, Den Spiegel, Didier Van (2009): Review of the millipede genusPlusioglyphiulusSilvestri, 1923, with descriptions of new species from southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Zoosystema 31 (1): 71-116, DOI: 10.5252/z2009n1a5, URL: http://dx.doi.org/10.5252/z2009n1a5
03D71761B67BFFFDFCADFAC4127AF961.taxon	discussion	REMARKS Any obvious troglomorphic traits in this species seem to be absent.	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, Den Spiegel, Didier Van (2009): Review of the millipede genusPlusioglyphiulusSilvestri, 1923, with descriptions of new species from southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Zoosystema 31 (1): 71-116, DOI: 10.5252/z2009n1a5, URL: http://dx.doi.org/10.5252/z2009n1a5
03D71761B665FFFAFCAFFF2A127AFA47.taxon	description	(Figs 14 - 17)	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, Den Spiegel, Didier Van (2009): Review of the millipede genusPlusioglyphiulusSilvestri, 1923, with descriptions of new species from southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Zoosystema 31 (1): 71-116, DOI: 10.5252/z2009n1a5, URL: http://dx.doi.org/10.5252/z2009n1a5
03D71761B665FFFAFCAFFF2A127AFA47.taxon	materials_examined	TYPE MATERIAL. — Laos. Khammouan Prov., Gnommalat, 17 ° 37.907 ’ N, 105 ° 07.461 ’ E, Cave Tham Kamuk, guano, 19. II. 2003, leg. H. Steiner, holotype ♂ (VMNH); 2 ♂♂ paratypes, 1 fragment paratype, 2 juveniles paratypes (VMNH). — Same locality, 20. II. 2003, 1 ♂, 1 ♀ paratypes (VMNH); 2 ♂♂, 1 ♀ paratypes (MNHN GA 058); 1 ♂ paratype (ZMUM); 1 ♂ paratype (ZMUC); 1 ♂ paratype (SEM).	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, Den Spiegel, Didier Van (2009): Review of the millipede genusPlusioglyphiulusSilvestri, 1923, with descriptions of new species from southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Zoosystema 31 (1): 71-116, DOI: 10.5252/z2009n1a5, URL: http://dx.doi.org/10.5252/z2009n1a5
03D71761B665FFFAFCAFFF2A127AFA47.taxon	etymology	ETYMOLOGY. — To honour Helmut Steiner, who collected this material.	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, Den Spiegel, Didier Van (2009): Review of the millipede genusPlusioglyphiulusSilvestri, 1923, with descriptions of new species from southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Zoosystema 31 (1): 71-116, DOI: 10.5252/z2009n1a5, URL: http://dx.doi.org/10.5252/z2009n1a5
03D71761B665FFFAFCAFFF2A127AFA47.taxon	diagnosis	DIAGNOSIS. — Differs from congeners by the especially complex gonopod structure, coupled with the isostichic carinotaxy of the collum and of other body segments, the convex paraprocts, the subtrapeziform epiproct and the absence of spots dorsad of the ocellaria.	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, Den Spiegel, Didier Van (2009): Review of the millipede genusPlusioglyphiulusSilvestri, 1923, with descriptions of new species from southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Zoosystema 31 (1): 71-116, DOI: 10.5252/z2009n1a5, URL: http://dx.doi.org/10.5252/z2009n1a5
03D71761B665FFFAFCAFFF2A127AFA47.taxon	description	DESCRIPTION Length of adults 27 - 35 mm, width 1.8 - 2.0 mm, collum and segments of posterior third of body being equally broadest. Coloration usually uniformly grey-brown, sometimes with a paramedian pair of more or less wide, grey-yellowish stripes; collum, head and segment 2 mostly pale; tips of tergal crests / tubercles usually dark brown; ocellaria dark brown to blackish, devoid of adjacent spots dorsally. Body with 56 - 69 p + 2 - 1 a + T. Length of holotype about 27 mm, width 1.8 mm, with 56 p + 1 a + T. Antennae rather elongate (Figs 14 A; 17 A), antennomeres 6 and 7 each with a considerable group of bacilliform sensilla distodorsally; similar but minute sensilla present also on antennomere 7. Ocellaria transversely subreniform, with 9 - 12 ocelli in 5 - 7 longitudinal rows. Gnathochilarium oligotrichous (Fig. 17 B). Postcollar constriction very evident, collum and segment 2 being strongly enlarged (Fig. 14 B). Carinotaxy formula of collum: I + // t + 2 p / t + 3 p / t + 4 p / t + ta / t + 6 p / t / t + 7 a / t + pp / t + ma / t (Fig. 14 A, B). Carinotaxy of metatergum 2, 8 / 8 + m / m + 8 / 8; formula of metaterga 3 and 4, 7 / 7 + M (m / m) + 7 / 7, dorsal 3 + 3 tubercles in anterior row being subtransverse; formula of metatergum 5, 3 / 3 + I / i + 3 / 3 + m / m + 3 / 3 + I / i + 3 / 3; formula of subsequent metaterga, 3 / 3 + I / i + 3 / 3 / 3 + m / m (m / m / m) + 3 / 3 / 3 + I / i + 3 / 3; that of apodous segments, 7 / 7 + m / m + 7 / 7 (Fig. 14); all crests and tubercles rather high, poriferous cones (I) only slightly higher, and median ones (m) slightly lower, than others (Figs 14 A-E; 15 B). Dorsal crests on several posteriormost segments considerably higher than others (Fig. 14 E). Midbody segments circular in cross-section (Fig. 15 B). Pleural regions of segments 2 - 4 conspicuously expanded, flap-shaped, especially so on segment 3 (Fig. 14 A). Pleural ridge behind gonopod aperture on male segment 7 (Fig. 14 D), tegument and limbus as in above congeners. Epiproct regularly and broadly rounded apically, with a small median knob and 1 + 1 unusually high, paramedian, subtransverse tubercles at midway; paraprocts flattened; hypoproct emarginate at caudal margin (Figs 14 E, F; 15 A). Legs short, on midbody segments about half as long as body diameter (Fig. 15 B). Claw at base with a strong but relatively short accessory spine about ¼ as long as claw itself (Fig. 15 F). Tarsi, tarsal setae and male legs 1 - 3 (Figs 15 C-E; 17 C-E) as in P. grandicollis, but a rudimentary telopodite of male leg 1 persisting as a knob. Anterior gonopods (Figs 16 A, B; 17 F) much like in P. ampullifer n. sp., but processes cxp 1 non-contiguous, very slender and high, with short pilosity laterally near base, subsecuriform apically; processes cxp 2 very slightly curved, similarly slender and high; telopodites (te) simple, laterocaudal in position, movable, high, subcylindrical, setose in distal third, apically curved mesad. Posterior gonopods (Figs 16 C, D; 17 G) small, complex, resembling a pair of curved forceps, without traces of telopodites; each coxite crescent-shaped, attenuating distad, with a rather long and flagelliform arm (d) at about basal third mesally, a shelf-like structure and two dentiform flaps (x and y) on frontal face.	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, Den Spiegel, Didier Van (2009): Review of the millipede genusPlusioglyphiulusSilvestri, 1923, with descriptions of new species from southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Zoosystema 31 (1): 71-116, DOI: 10.5252/z2009n1a5, URL: http://dx.doi.org/10.5252/z2009n1a5
03D71761B665FFFAFCAFFF2A127AFA47.taxon	discussion	REMARKS Any obvious troglomorphic traits in this species seem to be absent.	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, Den Spiegel, Didier Van (2009): Review of the millipede genusPlusioglyphiulusSilvestri, 1923, with descriptions of new species from southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Zoosystema 31 (1): 71-116, DOI: 10.5252/z2009n1a5, URL: http://dx.doi.org/10.5252/z2009n1a5
03D71761B662FFF6FF74F9E7127AFB44.taxon	materials_examined	TYPE MATERIAL. — Laos. Luang Prabang Prov., 19.88 ° N, 102.14 ° E, Xieng Maen, Cave Vat Tham, by hand, 30. XII. 1999, leg. L. Deharveng & A. Bedos (LAO- 106), holotype ♂ (MNHN GA 059); 1 ♂, 1 ♀ paratypes (MNHN GA 059); 1 ♀ paratype (ZMUM); 1 ♀ paratype (SEM).	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, Den Spiegel, Didier Van (2009): Review of the millipede genusPlusioglyphiulusSilvestri, 1923, with descriptions of new species from southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Zoosystema 31 (1): 71-116, DOI: 10.5252/z2009n1a5, URL: http://dx.doi.org/10.5252/z2009n1a5
03D71761B662FFF6FF74F9E7127AFB44.taxon	etymology	ETYMOLOGY. — To honour Louis Deharveng, one of the collectors and donators of this material.	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, Den Spiegel, Didier Van (2009): Review of the millipede genusPlusioglyphiulusSilvestri, 1923, with descriptions of new species from southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Zoosystema 31 (1): 71-116, DOI: 10.5252/z2009n1a5, URL: http://dx.doi.org/10.5252/z2009n1a5
03D71761B662FFF6FF74F9E7127AFB44.taxon	diagnosis	DIAGNOSIS. — Differs from congeners by certain details of gonopod structure (complex anterior gonopods versus very simple posterior ones), coupled with the mixostichic carinotaxy of the collum, the isostichic carinotaxy of other body segments, the flattened paraprocts, the elongated and subtrapeziform epiproct and the presence of spots dorsad of the ocellaria.	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, Den Spiegel, Didier Van (2009): Review of the millipede genusPlusioglyphiulusSilvestri, 1923, with descriptions of new species from southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Zoosystema 31 (1): 71-116, DOI: 10.5252/z2009n1a5, URL: http://dx.doi.org/10.5252/z2009n1a5
03D71761B662FFF6FF74F9E7127AFB44.taxon	description	DESCRIPTION Length of adults 20 - 33 mm, width 1.3 - 1.9 mm, collum and segments of posterior third of body being equally broadest. Coloration usually uniformly yellow-brown, sometimes with a paramedian pair of more or less wide, grey-yellowish stripes; collum, head and segment 2 mostly pale; tips of tergal crests / tubercles usually dark brown; ocellaria brown to blackish, with a pair of adjacent, light brown, well-separated spots dorsally; antennae and distal podomeres light brownish. Body with 48 - 59 p + 4 - 2 a + T. Length of holotype about 33 mm, width 1.9 mm, with 59 p + 2 a + T. Antennae short and clavate (Fig. 20 A), antennomeres 6 and 7 each with a modest group of bacilliform sensilla distodorsally. Ocellaria transversely subreniform, with 8 - 11 ocelli in 5 - 7 longitudinal rows. Gnathochilarium oligotrichous, each lamella lingualis with three setae distally. Postcollar constriction evident, but collum and segment 2 not particularly strongly enlarged (Fig. 18 B). Carinotaxy formula of collum slightly variable: ta / t + / (t) / + 2 p / t + (ta) / (t) + 3 p / t + 4 p + ta / t / t / t + 6 p / t / t + ta / t + pp / t / t + ma / t (usually), ta / t + 2 p / t + 3 p / t / t + ta / t + 5 p + ta / t + 7 p / t / t + ta / t + pp / t / t + ma / t or ta / t + 2 p / t + (ta) / (t) + 3 p / t + (ta) / (t) + 4 p + ta / t / t / t + 6 p / t / t + ta / t + pp / t / t + ma / t (more rarely) (Fig. 18 A, B). Carinotaxy of metatergum 2, 8 / 8 + m / m + 8 / 8; formula of metaterga 3 and 4, 7 / 7 + m / m + 7 / 7; formula of metaterga 5 to 16 and of several segments before apodous, 3 / 3 + I / i + 3 / 3 / 3 + m / m + 3 / 3 / 3 + I / i + 3 / 3; formula of midbody metaterga, 3 / 3 + I / i + 3 / 3 / 3 + m / m / m + 3 / 3 / 3 + I / i + 3 / 3; that of apodous segments, 4 + 3 / 3 + m / m + 3 / 3 + 4 (Figs 18; 19 A); all crests and tubercles rather low, poriferous cones (I) considerably higher, about as high as broad, while median ones (m) lower than others (Figs 18; 19 C). Dorsal crests on several posteriormost segments considerably higher than others (Fig. 19 A). Midbody segments circular in cross-section (Fig. 19 C). Pleural regions of segments 2 - 4 conspicuously expanded, flap-shaped, especially so on segment 3 (Fig. 18 A). Pleural ridge behind gonopod aperture on male segment 7, tegument and limbus as in above congeners. Epiproct elongate and subtrapeziform apically, with a paramedian pair of high longitudinal crests / tubercles flanked on each side by 1 or 2 small knobs at about midway; paraprocts flattened; hypoproct emarginate at caudal margin (Fig. 19 A, B). Legs short and rather stout, on midbody segments about half as long as body diameter (Figs 19 C; 20 E). Claw at base with a strong but relatively short accessory spine about ¼ as long as claw itself (Fig. 19 D). Tarsi, their distal setae and male legs 1 - 3 (Fig. 20 B-D) as in P. grandicollis. Anterior gonopods (Fig. 20 F) much like in P. ampullifer n. sp., but processes cxp 1 non-contiguous, lamelliform, broad and high, with a few setae laterally near base, subsecuriform, terminating in a minute uncus apico-medially and a long, digitiform, slightly curved outgrowth apico-laterally; processes cxp 2 of similar shape and length, but retrorse at end; telopodites (te) simple, laterocaudal in position, movable, rather slender and high, subcylindrical, setose in distal half laterally. Posterior gonopods (Fig. 20 G) small, simple, without traces of telopodites; each coxite with a rather long, spiniform arm (d) at about midway on frontal face, distal half lamelliform, parallel-sided, with a deep fovea and a modest emargination apically.	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, Den Spiegel, Didier Van (2009): Review of the millipede genusPlusioglyphiulusSilvestri, 1923, with descriptions of new species from southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Zoosystema 31 (1): 71-116, DOI: 10.5252/z2009n1a5, URL: http://dx.doi.org/10.5252/z2009n1a5
03D71761B662FFF6FF74F9E7127AFB44.taxon	discussion	REMARKS Any obvious troglomorphic traits in this species seem to be absent.	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, Den Spiegel, Didier Van (2009): Review of the millipede genusPlusioglyphiulusSilvestri, 1923, with descriptions of new species from southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Zoosystema 31 (1): 71-116, DOI: 10.5252/z2009n1a5, URL: http://dx.doi.org/10.5252/z2009n1a5
03D71761B66EFFF7FF63FAE510B5FD9D.taxon	description	(Figs 21 - 23)	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, Den Spiegel, Didier Van (2009): Review of the millipede genusPlusioglyphiulusSilvestri, 1923, with descriptions of new species from southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Zoosystema 31 (1): 71-116, DOI: 10.5252/z2009n1a5, URL: http://dx.doi.org/10.5252/z2009n1a5
03D71761B66EFFF7FF63FAE510B5FD9D.taxon	materials_examined	TYPE MATERIAL. — Indonesia. Borneo, Kalimantan Timur, KutaiTimur, Sangkulirang, Baai, 01.332 ° N, 117.397 ° E, Cave Gua Ambulabung, by hand, 15. VIII. 2004, leg. A. Bedos & L. Deharveng (KAL- 071), holotype ♂ (MZB); 1 ♀ paratype (MNHN GA 060), 1 juvenile ♂ paratype (SEM).	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, Den Spiegel, Didier Van (2009): Review of the millipede genusPlusioglyphiulusSilvestri, 1923, with descriptions of new species from southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Zoosystema 31 (1): 71-116, DOI: 10.5252/z2009n1a5, URL: http://dx.doi.org/10.5252/z2009n1a5
03D71761B66EFFF7FF63FAE510B5FD9D.taxon	etymology	ETYMOLOGY. — To honour Anne Bedos, one of the collectors of this material.	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, Den Spiegel, Didier Van (2009): Review of the millipede genusPlusioglyphiulusSilvestri, 1923, with descriptions of new species from southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Zoosystema 31 (1): 71-116, DOI: 10.5252/z2009n1a5, URL: http://dx.doi.org/10.5252/z2009n1a5
03D71761B66EFFF7FF63FAE510B5FD9D.taxon	diagnosis	DIAGNOSIS. — Differs from congeners in the small size, the particular carinotaxy formulae, coupled with the peculiar structure of male legs 1 - 3, as well as of the gonopods.	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, Den Spiegel, Didier Van (2009): Review of the millipede genusPlusioglyphiulusSilvestri, 1923, with descriptions of new species from southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Zoosystema 31 (1): 71-116, DOI: 10.5252/z2009n1a5, URL: http://dx.doi.org/10.5252/z2009n1a5
03D71761B66EFFF7FF63FAE510B5FD9D.taxon	description	DESCRIPTION Length of adults about 22 mm, width 1.3 mm, segments of posterior third of body being broadest. Coloration generally marbled grey-brown, with two broad, paramedian, grey-yellowish, more or less distinct stripes dorsally and with brown prozona, thus providing a cingulate pattern; collum and segment 2 mostly yellow to light brown; head mostly pale brown to brown, at least dorsad of interantennal isthmus, with a yellow clypeus; ocellaria blackish; antennae dark brown, distal podomeres light brown to brown. Body of adults with 52 p + 2 a + T (holotype) or 51 p + 3 a + T (paratype). Antennae short and clavate (Figs 21 A; 23 A), antennomeres 5 and 6 each with a small group of bacilliform sensilla distodorsally. Ocellaria transverse, with 11 - 13, mostly isolated ocelli in 5 or 6 longitudinal rows. Gnathochilarium oligotrichous. Postcollar constriction evident, but collum and segment 2 not particularly strongly enlarged (Fig. 21 A, B). Carinotaxy formula of collum: I + // (t) + / II + III + IV + 5 a + VI + 7 a + P + ma (Fig. 21 A, B). Carinotaxy of metatergum 2, 4 / 4 + 4 + M + 4 + 4 / 4; formula of metaterga 3 and 4, 8 / 8 + m / m + 8 / 8; formula of metaterga 5 to several before apodous, 4 / 4 + I / i + 3 / 3 / 3 + m / m + 3 / 3 / 3 + I / i + 4 / 4; formula of several caudal metaterga in front of apodous, 4 / 4 + I / i + 3 / 3 + m / m + 3 / 3 + I / i + 4 / 4; that of apodous segments, 3 + 1 / 1 + 3 + M + 3 + 1 / 1 + 3 (Fig. 21 A-E); all crests and tubercles rather low, poriferous cones (I) slightly higher, broader than high, while median ones (m) slightly lower than others (Fig. 22 A). Dorsal crests on several posteriormost segments considerably higher than others (Fig. 21 E). Midbody segments circular in cross-section (Fig. 22 A). Pleural regions of segments 2 - 4 conspicuously expanded, flap-shaped, especially so on segment 3 (Fig. 21 A). Pleural ridge behind gonopod aperture on male segment 7, tegument and limbus as in above congeners. Epiproct regularly rounded apically, with a paramedian pair of evident, caudally converging tubercles at about midway; paraprocts flattened; hypoproct emarginate at caudal margin (Fig. 21 E, F). Legs short, rather stout, on midbody segments about half as long as body diameter (Figs 22 A; 23 E). Claw at base with a strong accessory spine about half as long as claw itself (Figs 22 B, C; 23 E). Tarsi and their distal setae as in other congeners. Male coxae 1 (Fig. 23 B) subconical, strongly reduced and shorter than central sternal uncus. Male telopodites 2 (Fig. 23 C) not particularly incrassate, with an elongated tarsus. Male telopodites 3 (Fig. 23 D) only slightly reduced compared to elongated coxae. Anterior gonopods (Fig. 23 F, G) much like in P. ampullifer n. sp., processes cxp 1 contiguous medially, lamellate, digitiform, very finely fringed apically, shorter than processes cxp 2; latter also lamellate, elongate and subacuminate, almost in touch near apex; telopodites (te) simple, laterocaudal in position, movable, rather slender and high, subcylindrical, setose on caudal face. Posterior gonopods (Fig. 23 H) small, simple, without traces of telopodites; each coxite with a rather long, rod-shaped, distally micropilose arm (d) at basal third on frontal face, distal part finely fringed apically, lamelliform, nearly parallel-sided, with long lateral and mesal flaps, latter flap being surmounted with a short process. This is a rather atypical member of Plusioglyphiulus. It deviates chiefly in the somewhat more plesiomorphic characters of both male legs 2 and 3, but, on account of the increased number of metatergal tubercles and collar crests, as well as the overall gonopod conformation, its assignment to this genus seems preferable for the moment. Any obvious troglomorphic traits in this species seem to be absent.	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, Den Spiegel, Didier Van (2009): Review of the millipede genusPlusioglyphiulusSilvestri, 1923, with descriptions of new species from southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Zoosystema 31 (1): 71-116, DOI: 10.5252/z2009n1a5, URL: http://dx.doi.org/10.5252/z2009n1a5
03D71761B66FFFF2FCB4FDAE175AF961.taxon	description	(Figs 24 - 26)	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, Den Spiegel, Didier Van (2009): Review of the millipede genusPlusioglyphiulusSilvestri, 1923, with descriptions of new species from southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Zoosystema 31 (1): 71-116, DOI: 10.5252/z2009n1a5, URL: http://dx.doi.org/10.5252/z2009n1a5
03D71761B66FFFF2FCB4FDAE175AF961.taxon	materials_examined	TYPE MATERIAL. — Indonesia. Borneo, Kalimantan Timur, Kutai Timur, Sangkulirang, Baai, 01.332 ° N, 117.397 ° E, Cave Gua Ambulabung, by hand, 14. VIII. 2004, leg. A. Bedos & L. Deharveng (KAL- 068), holotype ♂ (MZB); 2 subadult ♂♂ paratypes (MZB); 1 ♂, 1 ♀ paratypes (MNHN GA 061). — Same cave, 15. VIII. 2004, leg. A. Bedos & L. Deharveng (KAL- 071), 1 ♂ paratype (ZMUM). — Deep part of the same cave, by hand, 16. VIII. 2004, leg. A. Bedos & L. Deharveng (KAL- 110), 1 subadult ♀ paratype (MNHN GA 061); 1 juvenile ♀ paratype (SEM).	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, Den Spiegel, Didier Van (2009): Review of the millipede genusPlusioglyphiulusSilvestri, 1923, with descriptions of new species from southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Zoosystema 31 (1): 71-116, DOI: 10.5252/z2009n1a5, URL: http://dx.doi.org/10.5252/z2009n1a5
03D71761B66FFFF2FCB4FDAE175AF961.taxon	etymology	ETYMOLOGY. — To emphasize the lighter general coloration compared to the sympatric and even syntopic P. bedosae n. sp.	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, Den Spiegel, Didier Van (2009): Review of the millipede genusPlusioglyphiulusSilvestri, 1923, with descriptions of new species from southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Zoosystema 31 (1): 71-116, DOI: 10.5252/z2009n1a5, URL: http://dx.doi.org/10.5252/z2009n1a5
03D71761B66FFFF2FCB4FDAE175AF961.taxon	diagnosis	DIAGNOSIS. — Differs from congeners by certain details of gonopod structure (unusually enlarged anterior gonopods versus rather slender posterior ones), coupled with the mixostichic carinotaxy of the collum and of other body segments, the presence of a marked depression on the collum and of only two rows of bituberculate crests below the poriferous cones on most of the body segments, the flattened paraprocts, the short and broadly rounded epiproct and the absence of spots dorsad of the ocellaria.	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, Den Spiegel, Didier Van (2009): Review of the millipede genusPlusioglyphiulusSilvestri, 1923, with descriptions of new species from southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Zoosystema 31 (1): 71-116, DOI: 10.5252/z2009n1a5, URL: http://dx.doi.org/10.5252/z2009n1a5
03D71761B66FFFF2FCB4FDAE175AF961.taxon	description	DESCRIPTION Length of adults 43 - 50 mm, width 2.0 - 2.1 mm, segments of posterior third of body being broadest. Coloration uniformly light yellow-brown; head (except for blackish ocelli), collum and segments 2 - 7 entirely yellow. Body of adults with 64 - 70 p + 2 - 1 a + T. Length of holotype about 50 mm, width 2.1 mm (width of collum, 2.0 mm) with 70 p + 2 a + T. Clypeus with 3 (n = 2) or 5 teeth (n = 1) anteromedially. Antennae short and clavate (Fig. 26 A), antennomeres 6 and 7 each with a very considerable group of bacilliform sensilla distodorsally. Ocellaria transversely subtriangular, with 8 - 11 ocelli in 5 - 7 longitudinal rows. Gnathochilarium oligotrichous (Fig. 26 B). Postcollar constriction very evident, collum and segment 2 particularly strongly enlarged (Fig. 24 A). Carinotaxy formula of collum variable: ta / t + 2 p / t + 3 p / t / t / t + /// (t) + 4 p / t / t / t + 5 p / t / t / t + pp / t / t / t + // ma / t or // t + 2 p / t / t + 3 p / t / t + 4 p / t / t / t + // (ta) / (t) + 5 p / t / t / t + pp / t / t / t + ma / t (Fig. 24 A). Collum strongly depressed near anterior third (Fig. 24 A). Carinotaxy of metatergum 2, 6 / 6 + m / m + 6 / 6; formula of metaterga 3 and 4, 3 / 3 + 3 + M + 3 + 3 / 3; formula of metaterga 5 - 7 and of several segments before apodous, 2 / 2 + I / i + 3 / 3 + m / m + 3 / 3 + I / i + 2 / 2; formula of midbody metaterga, 2 / 2 + I / i + 3 / 4 / 3 + m // m + 3 / 4 / 3 + I / i + 2 / 2 in adults, 2 / 2 + I / i + 3 / 3 + m / m + 3 / 4 / 3 + I / i + 2 / 2 in subadults; formula of apodous segments, 4 + 0 + 4 (Figs 24; 25 A); all crests and tubercles rather high, poriferous cones (I) considerably higher, about as high as broad, while median ones (m) slightly lower than others (Fig. 24 B). All tergal tubercles microgranular (Fig. 25 C). Dorsal crests on several posteriormost segments considerably higher than others (Fig. 25 A). Midbody segments circular in cross-section (Fig. 24 B). Pleural regions of segments 2 - 4 conspicuously expanded, flap-shaped, especially so on segment 3 (Fig. 24 A). Pleural ridge behind gonopod aperture on male segment 7, tegument and limbus as in above congeners. Epiproct short, regularly and broadly rounded apically, with a paramedian pair of high but short crests / tubercles sometimes flanked on each side by 1 or 2 small knobs at about midway; paraprocts fl attened; hypoproct emarginate at caudal margin (Fig. 25 A, B). Legs relatively long, on midbody segments about ¾ as long as body diameter (Fig. 24 B). Claw at base with a very short accessory spine (Fig. 25 D). Tarsi and their distal setae as in other congeners. Male legs 1 - 3 (Fig. 26 C-E) more or less as in P. grandicollis, but male coxa 1 relatively short, while male telopodite 3 not so compact. Anterior gonopods (Fig. 26 F) generally like in P. ampullifer n. sp., but coxites swollen and stout, non-contiguous medially; processes cxp 1 very short, beak-shaped, finely ribbed apically; processes cxp 2 high, hyaline, subspatuliform, setose at base on frontal face; telopodites (te) simple, lateral in position, movable, rather stout, subcylindrical with tip curved mesad, setose at about midway in distal half laterally. Posterior gonopods (Fig. 26 G) small, relatively simple, without traces of telopodites; each coxite with a long, spiniform arm (d) laterally and a shelf-like structure at about midway on frontal face; distal half lamellose, fringed and with a deep fovea apically. Subadults about 26 mm long, 1.3 mm wide, pallid, with 53 - 55 p + 4 a + T and 3 - 5 blackish ocelli in 2 - 4 longitudinal rows. Male legs 1 similar to adult condition, but central sternal hook nearly straight, non-unciform. Male legs 2 and 3 slightly less strongly modified than in adults.	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, Den Spiegel, Didier Van (2009): Review of the millipede genusPlusioglyphiulusSilvestri, 1923, with descriptions of new species from southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Zoosystema 31 (1): 71-116, DOI: 10.5252/z2009n1a5, URL: http://dx.doi.org/10.5252/z2009n1a5
03D71761B66FFFF2FCB4FDAE175AF961.taxon	discussion	REMARKS The most striking observation obtained from the above material is that an additional, middle row of intercalary tubercles on the metaterga (like in Fig. 27 C, D) seems to appear only at the onset of adulthood. Any obvious troglomorphic traits in this species seem to be absent. The size differences between P. bedosae n. sp. and P. pallidior n. sp. are so extreme that niche segregation is strongly suggested, such is the case in some other soil- or cave-dwelling millipedes (Enghoff 1992; Enghoff et al. 1997). This is so far the only locality known that supports two species of Plusioglyphiulus. That we deal with two different species is beyond any doubt. Even the subadults of P. pallidior n. sp. are considerably larger than the adults of P. bedosae n. sp. In addition, they differ in gonopod structure so drastically that one (P. pallidior n. sp.) could hardly be seen as an ontogenetic derivative of the other (P. bedosae n. sp.).	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, Den Spiegel, Didier Van (2009): Review of the millipede genusPlusioglyphiulusSilvestri, 1923, with descriptions of new species from southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Zoosystema 31 (1): 71-116, DOI: 10.5252/z2009n1a5, URL: http://dx.doi.org/10.5252/z2009n1a5
03D71761B654FFCFFE9DF9DA127AF961.taxon	description	(Figs 27 - 30)	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, Den Spiegel, Didier Van (2009): Review of the millipede genusPlusioglyphiulusSilvestri, 1923, with descriptions of new species from southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Zoosystema 31 (1): 71-116, DOI: 10.5252/z2009n1a5, URL: http://dx.doi.org/10.5252/z2009n1a5
03D71761B654FFCFFE9DF9DA127AF961.taxon	materials_examined	TYPE MATERIAL. — Indonesia. Borneo, KalimantanTimur, Pengadan, 01.257 ° N, 117.706 ° E, Cave Gua Ampanas, by hand, 18. VIII. 2004, leg. A. Bedos & L. Deharveng (KAL- 113), holotype ♂ (MZB); 1 ♂, 1 ♀ paratypes (MNHN GA 062); 1 ♂ paratype (SEM). — Same cave, 18. VIII. 2004, leg. Y. R. Suhardjono & C. Rahmadi (KAL-YC 04 - 09), 1 ♀ paratype (MZB); 1 ♂, 1 ♀ paratypes (ZMUM); 1 ♀ paratype (ZMUC); 1 ♀ paratype (VMNH).	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, Den Spiegel, Didier Van (2009): Review of the millipede genusPlusioglyphiulusSilvestri, 1923, with descriptions of new species from southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Zoosystema 31 (1): 71-116, DOI: 10.5252/z2009n1a5, URL: http://dx.doi.org/10.5252/z2009n1a5
03D71761B654FFCFFE9DF9DA127AF961.taxon	etymology	ETYMOLOGY. — To emphasize the strong similarities to P. pallidior n. sp.	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, Den Spiegel, Didier Van (2009): Review of the millipede genusPlusioglyphiulusSilvestri, 1923, with descriptions of new species from southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Zoosystema 31 (1): 71-116, DOI: 10.5252/z2009n1a5, URL: http://dx.doi.org/10.5252/z2009n1a5
03D71761B654FFCFFE9DF9DA127AF961.taxon	diagnosis	DIAGNOSIS. — Differs from the most similar P. pallidior n. sp. in the smaller size, the presence of three rows of bituberculate crests below the poriferous cones on most of the body segments and the somewhat longer legs.	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, Den Spiegel, Didier Van (2009): Review of the millipede genusPlusioglyphiulusSilvestri, 1923, with descriptions of new species from southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Zoosystema 31 (1): 71-116, DOI: 10.5252/z2009n1a5, URL: http://dx.doi.org/10.5252/z2009n1a5
03D71761B654FFCFFE9DF9DA127AF961.taxon	description	DESCRIPTION Length 17 - 27 mm, width 1.0 - 1.6 mm, segments of posterior third of body being broadest. Coloration marbled brown, sometimes with broad, paramedian, yellow stripes dorsally; head (except for blackish ocelli), collum and segment 2 mostly pale yellow; tips of metatergal tubercles usually brown, those of poriferous tubercles dark brown; antennae and legs light brown. Body with 42 - 50 p + 3 - 1 a + T. Length of holotype about 26 mm, width 1.6 mm, with 50 p + 1 a + T. Antennae rather short and clavate (Figs 27 A; 30 A), antennomeres 6 and 7 each with a small group of bacilliform sensilla distodorsally. Ocellaria transversely subtriangular, with 8 - 11 ocelli in 5 - 7 longitudinal rows. Gnathochilarium slightly more densely setose due to a few smal setae on promentum (Fig. 30 B). Postcollar constriction very evident, collum and segment 2 particularly strongly enlarged (Fig. 27 A, B). Carinotaxy formula of collum: 1 p / t + 2 p / t + 3 p / t / t + // (ta) / (t) + 4 p / t / t + 5 p / t / t / t + // (ta) / (t) + pp / t / t / t + // ma / t (Fig. 27 B). Collum slightly depressed near anterior third. Carinotaxy of metatergum 2, 7 / 7 + m / m + 7 / 7; formula of metaterga 3 and 4, 4 / 4 + 3 + M + 3 + 4 / 4; formula of metaterga 5 - 7 and of several segments before apodous, 3 / 3 + I / i + 3 / 3 + m / m + 3 / 3 + I / i + 2 / 2; formula of midbody metaterga, 3 / 3 + I / i + 3 / 4 (5) / 3 + m // m + 3 / 4 (5) / 3 + I / i + 3 / 3; that of apodous segments, 4 / 4 + m / m + 4 / 4 (Fig. 27 A-E); all crests and tubercles rather low, poriferous cones (I) considerably higher, but still broader than high, while median ones (m) as high as others (Figs 27 A-E; 28 A). Dorsal crests on several posteriormost segments considerably higher than others (Fig. 27 E). Midbody segments circular in cross-section (Fig. 28 A). Pleural regions of segments 2 - 4 conspicuously expanded, flap-shaped, especially so on segment 3 (Fig. 27 A). Pleural ridge behind gonopod aperture on male segment 7, tegument and limbus as in above congeners. Epiproct short, regularly and broadly rounded apically, with a paramedian pair of high but short crests / tubercles sometimes flanked on each side by a small knob at about midway; paraprocts more or less regularly convex, not flattened; hypoproct only slightly emarginate at caudal margin (Fig. 27 E, F). Legs relatively long, on midbody segments about as long as body diameter (Figs 28 A; 30 F). Claw at base with a short accessory spine (Fig. 28 B). Tarsi and their distal setae, as well as male legs 1 - 3 (Figs 28 C, D; 29 A; 30 C-E) typical. Both anterior and posterior gonopods (Figs 29 B-D; 30 G-I) very much like in P. pallidior n. sp., but both coxal processes cxp 1 and cxp 2, and arm (d), slightly different in shape.	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, Den Spiegel, Didier Van (2009): Review of the millipede genusPlusioglyphiulusSilvestri, 1923, with descriptions of new species from southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Zoosystema 31 (1): 71-116, DOI: 10.5252/z2009n1a5, URL: http://dx.doi.org/10.5252/z2009n1a5
03D71761B654FFCFFE9DF9DA127AF961.taxon	discussion	REMARKS Any obvious troglomorphic traits in this species seem to be absent.	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, Den Spiegel, Didier Van (2009): Review of the millipede genusPlusioglyphiulusSilvestri, 1923, with descriptions of new species from southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Zoosystema 31 (1): 71-116, DOI: 10.5252/z2009n1a5, URL: http://dx.doi.org/10.5252/z2009n1a5
03D71761B657FFCBFCB9FF2A10B7FA85.taxon	description	(Figs 31 - 33)	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, Den Spiegel, Didier Van (2009): Review of the millipede genusPlusioglyphiulusSilvestri, 1923, with descriptions of new species from southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Zoosystema 31 (1): 71-116, DOI: 10.5252/z2009n1a5, URL: http://dx.doi.org/10.5252/z2009n1a5
03D71761B657FFCBFCB9FF2A10B7FA85.taxon	materials_examined	TYPE MATERIAL. — Malaysia. Borneo, Sarawak, Gunung Mulu National Park, 4.05 ° N, 114.93 ° E, Deer Water Cave (Luang Sungai Payau), halfway, in guano, 25. IV. 1978, leg. P. J. Chapman, holotype ♂ (VMNH); 3 ♂♂, 5 ♀♀, 1 subadult ♂, 1 subadult ♀ paratypes (VMNH); 1 ♂, 1 ♀ paratypes (MNHN GA 063); 1 ♂, 1 ♀ paratypes (ZMUM); 1 ♀ paratype (ZMUC); 1 ♀ paratype (SEM).	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, Den Spiegel, Didier Van (2009): Review of the millipede genusPlusioglyphiulusSilvestri, 1923, with descriptions of new species from southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Zoosystema 31 (1): 71-116, DOI: 10.5252/z2009n1a5, URL: http://dx.doi.org/10.5252/z2009n1a5
03D71761B657FFCBFCB9FF2A10B7FA85.taxon	etymology	ETYMOLOGY. — To honour Richard L. Hoffman (VMNH), a prominent specialist in the systematics of Diplopoda, who provided several samples for this study.	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, Den Spiegel, Didier Van (2009): Review of the millipede genusPlusioglyphiulusSilvestri, 1923, with descriptions of new species from southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Zoosystema 31 (1): 71-116, DOI: 10.5252/z2009n1a5, URL: http://dx.doi.org/10.5252/z2009n1a5
03D71761B657FFCBFCB9FF2A10B7FA85.taxon	diagnosis	DIAGNOSIS. — Differs from the most similar P. macfarlanei Mauriès, 1983 in minor details of gonopod structure (cxp 1 of the anterior gonopod is not extended into a flat lobe, te of the same gonopod is recurved and unciform, the apex of the posterior gonopod is less elaborate), the short epiproct, the regularly convex paraprocts and the absence of complete crests on the collum.	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, Den Spiegel, Didier Van (2009): Review of the millipede genusPlusioglyphiulusSilvestri, 1923, with descriptions of new species from southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Zoosystema 31 (1): 71-116, DOI: 10.5252/z2009n1a5, URL: http://dx.doi.org/10.5252/z2009n1a5
03D71761B657FFCBFCB9FF2A10B7FA85.taxon	description	DESCRIPTION Length 19 - 30 mm, width 1.0 - 1.6 mm, collum and segments of posterior third of body being equally broadest. Coloration uniformly light yellow-brown to light brown, sometimes with infuscated sides and tips of poriferous tubercles; head (except for brown-blackish ocelli and a paramedian pair of separated, light brown spots dorsad of ocellaria), collum and segment 2 mostly pale yellow; antennae and legs light brown. Body with 45 - 53 p + 3 - 1 a + T. Length of holotype about 24 mm, width 1.5 mm, with 52 p + 2 a + T. Antennae short and clavate (Figs 31 A; 33 A), antennomeres 6 and 7 each with a considerable group of bacilliform sensilla distodorsally; similar but minute sensilla present also on antennomere 7 (Fig. 32 C). Ocellaria transversely subtriangular, with 5 - 9 ocelli in 4 - 6 longitudinal rows. Gnathochilarium oligotrichous. Postcollar constriction very evident, collum and segment 2 particularly strongly enlarged (Fig. 31 A, B). Carinotaxy formula of collum: ta / t + 2 p / t + 3 p / t + // ta / t + / (ta) / (t) / (t) + 5 p / t / t + 5 p / t / t + // ta / t + pp / t / t / t + // ma / t (Fig. 31 A, B). Carinotaxy of metatergum 2, 8 / 8 + m / m + 8 / 8; formula of metaterga 3 and 4, 3 / 3 + 4 + M + 4 + 3 / 3; formula of metaterga 5 and 6, and of several segments before apodous, 3 / 3 + I / i + 3 / 3 + m / m + 3 / 3 + I / i + 3 / 3; formula of midbody metaterga, 3 / 3 + I / i + 3 / 4 / 3 + m // m + 3 / 4 / 3 + I / i + 3 / 3; that of apodous segments, 5 + (m) + 5 (Figs 31; 32 B); all crests and tubercles rather low, poriferous cones (I) considerably higher, but still broader than high, while median ones (m) as high as others (Figs 31 C, D; 32 B). Dorsal crests on several posteriormost segments considerably higher than others (Fig. 31 E). Midbody segments circular in cross-section (Fig. 32 B). Pleural regions of segments 2 - 4 conspicuously expanded, flap-shaped, especially so on segment 3 (Fig. 31 A). An unusually high, transverse pleural ridge behind gonopod aperture on male segment 7, with very short rounded flaps bent caudad at apex. Tegument and limbus typical. Epiproct very short, subtrapeziform, broadly rounded apically, with a paramedian pair of well-separated, high but short crests / tubercles at about midway; paraprocts regularly convex, not flattened; hypoproct only slightly emarginate at caudal margin (Figs 31 F; 32 A). Legs relatively long, on midbody segments about as long as body diameter (Figs 31 C; 33 B). Claw at base with a short accessory spine (Figs 32 D; 33 B). Tarsi and their distal setae, as well as male legs 1 - 3 as in P. grandicollis, P. ampullifer n. sp. or P. pallidior n. sp. Anterior gonopods (Fig. 33 C) very much like in P. macfarlanei in coxites being contiguous but not fused medially, each coxite bearing only a single, laterally serrate, digitiform process (cxp 1); telopodites (te) simple, lateral in position, movable, long, likewise digitiform, devoid of setation, with tip unciform and directed laterad. Posterior gonopods (Fig. 33 D) small, relatively simple, without traces of telopodites; each coxite with a very short, rudimentary arm (d) distomedially near a shelf-like structure; a lobiform tip and a subterminal lateral setoid laterally. Subadults about 16 mm long, 0.9 mm wide, pallid, with 41 - 42 p + 4 a + T and three blackish ocelli in two longitudinal rows. Tubercles of middle, intercalary row on midbody segments very small, rudimentary. Male legs 1 similar to adult condition, but central sternal hook nearly straight, non-unciform. Male legs 2 and 3 slightly less strongly modified than in adults.	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, Den Spiegel, Didier Van (2009): Review of the millipede genusPlusioglyphiulusSilvestri, 1923, with descriptions of new species from southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Zoosystema 31 (1): 71-116, DOI: 10.5252/z2009n1a5, URL: http://dx.doi.org/10.5252/z2009n1a5
03D71761B657FFCBFCB9FF2A10B7FA85.taxon	discussion	REMARKS Unlike P. pallidior n. sp., in which the subadults show no traces of a row of intercalary tubercles on the midbody segments, the subadults of P. hoffmani n. sp. do have such tubercles, albeit strongly reduced in size. In this case these tubercles can be presumed as appearing not at, but towards the onset of adulthood. Any obvious troglomorphic traits in this species seem to be absent. This contradicts the observations of Chapman (1984), who referred to an eyeless, white, troglobitic species belonging to a new genus close to Plusioglyphiulus as one of the inhabitants of this cave (det. R. L. Hoffman). Because the above material comes from the original sample collected by Chapman and provisionally identified by Hoffman, Chapman (1984) was certainly referring to P. hoffmani n. sp. The diplopod fauna of the caves of Gunung Mulu National Park is known to be rather rich and diverse, containing seven genera of four orders. Most, if not all, of the species have been presumed to be troglobites (Chapman 1984). Based on geographical reasons alone, when the occurrence of two congeners in the same cave is exceptional, the recent records of P. cavernicolus in several caves of Sarawak other than Bidi Caves, including a few caves from Gunung Mulu National Park (Decu et al. 2001), are most likely to represent misidentifications.	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, Den Spiegel, Didier Van (2009): Review of the millipede genusPlusioglyphiulusSilvestri, 1923, with descriptions of new species from southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Zoosystema 31 (1): 71-116, DOI: 10.5252/z2009n1a5, URL: http://dx.doi.org/10.5252/z2009n1a5
