identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03D42163EC29BF6DFF7FFBB8FD1AF53F.text	03D42163EC29BF6DFF7FFBB8FD1AF53F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Microdera abdullohi N. Bekchanov et Nabozhenko 2025	<div><p>Microdera abdullohi N. Bekchanov et Nabozhenko,  sp. nov.</p><p>(Figs 1, 2)</p><p>Microdera globulicollis: Pirnazarov, 1970: 27 (partly, record from “Sultanuizdag” = Sulton-Uvays Mts.)</p><p>Microdera convexa: Bekchanov et al., 2023: 152 (part, only one specimen from Beruni Distr.).</p><p>Type material.  Holotype, ♂ (ZIN) and   paratypes, 2♂, 4♀ (ZIN, PCMN), 1♀ (PCNB), Uzbekistan, Karakalpakistan, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=60.44278&amp;materialsCitation.latitude=42.024166" title="Search Plazi for locations around (long 60.44278/lat 42.024166)">lava fields in the foothills of Sulton-Uvays Mts opposite Lower Amudarya State Biosphere Reserve</a>, 42°01ʹ49ʺN / 60°24ʹ25ʺE, h = 100 m, 24.04.2024 (M.V. and S.V. Nabozhenko, N.Kh. Bekchanov, U.E. Duschanov);   2♀ (ZIN), Uzbekistan, Karakalpakstan, Beruni District, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=60.44278&amp;materialsCitation.latitude=42.024166" title="Search Plazi for locations around (long 60.44278/lat 42.024166)">turning from the road to Lower Amudarya State Biosphere Reserve</a>, 42°01ʹ27ʺN / 60°26ʹ34ʺE, 24– 25.04.2023 (M.V. and S.V. Nabozhenko, N.Kh. and Kh.U. Bekchanov);  1♂ (ZIN), 1♂, 3♀ (PCNB), the same locality, 19.03.2024 (N.Kh. Bekchanov, Kh.U. Bekchanov and M.Kh. Bekchanova).</p><p>Description. Male. Body slender, convex, shiny, black, with long and slender antennae and legs, distal parts of legs and antennae brown. Measurements: Body length 10 mm, width 3.6 mm; PH w = 1.3; P 1 P w =1.13; E l E w =1.55; EH w =1.85; EP w = 1.42; EP l =2.47.</p><p>Head (Fig. 1D, E) dorsally widest at eye level, where 1.03 times as wide as at genal level and 1.3 times as wide as interocular space. Eyes comparatively large (in length dorsally), moderately convex. Temple not protruding and not forms angle with posterior edge of eye (lateral edge of head not angular at level of temple and eye), visual area of eye lateral. Supraorbital keels slightly sinuous, long. Prong of anterior margin of epistoma slightly asymmetric, rounded, not separated as a subparallel small process; margins of prong not emarginated (Fig. 1E). Puncturation of head coarse and dense dorsally: punctures sparser in anterior half of head than on frons; punctures smaller and denser along margin of head (especially prong and margin of gena). Puncturation of head ventrally sparse; mentum with small and round punctures; temples ventrally with sparse punctures, every puncture has a small seta. Antennae long, with 3 distal antennomeres extending beyond base of pronotum. Antennomere 3 longest (length to width ratio: 3.6–3.7), 1.95 times as long as 2nd antennomere and 1.7 times as long as 4th antennomere; antennomeres 1 and 8 widest (1.2 times as long as wide for the latter). Proximal antennomeres moderately coarsely and densely punctured, while three distal antennomeres with very fine and sparse punctures.</p><p>Prothorax (Fig. 1D, F, G). Pronotum transverse, trapezoidal, widest at anterior fourth. Anterior margin almost straight, lateral margins very weakly rounded or almost straight in basal two thirds and strongly rounded in apical third, base evenly weakly rounded. Antero-lateral corners widely rounded; postero-lateral corners obtuse. Anterior edge margined only on sides, while lateral margins and base completely margined. Base much wider margined. Disc of pronotum strongly evenly convex, sparsely punctured by round moderately coarse punctures (inperpuncture distance 3–4 time wider than punctures). Prosternum sparsely but distinctly punctured by fine punctures, smoothly wrinkled on sides. Prothoracic hypomera with moderately coarse distinct sparse puncturation (Fig. 1G) or sparse and finer puncturation and smooth wrinkles (Fig. 1F), some specimens have smaller or almost invisible and sparser punctures than on prosternum. Prosternal process with sharp bend between dorsal and vertical portions (Fig. 1F, G).</p><p>Pterothorax (Fig. 1A, C). Elytra oval, widest at middle. Puncturation of elytra very fine and sparse, much sparser and finer than on pronotum. Anterior half of mesoventrite with transverse rasp-like bracket-like foveae or roughly merged foveae (one male specimen has rough lines); posterior half with thin long transverse grooves; intercoxal process of mesoventrite convex, widely longitudinally depressed in middle, finely and sparsely punctured. Mesepisterna, mesepimera and metepisterna sparsely and finely punctured; metepimera not punctured. Metaventrite with sparse rasp-like larger punctures at anterior half and very sparse and almost invisible punctures at posterior half.</p><p>Abdomen (Fig. 1C, H, I). Abdominal ventrites evenly punctured with sparse and fine punctures. Intercoxal process of abdominal ventrite 1 not margined at apex. Abdominal ventrite 5 widely roundely emarginated at apex, without vertical plate in middle (Fig. 1H, I). Male terminalia and genitalia (Fig. 2A–F). Inner tergite VIII shortly emarginated at apex, while inner sternite VIII with deep triangle emargination in middle and acute apices. Inner sternite VIII more sclerotized in basal sides (Fig. 2E). Spiculum gastrale with transverse triangular blades having long lateral processes (Fig. 2F). Apical piece of tegmen unevenly narrowed from base to apex, bisinuate, lateral margins shortly emarginated in basal fourths and widely emarginated in middle, widest at base, apex acute; basal piece 1.3 times as wide and 1.1 times as short as apical piece, widest at basal third (Fig. 2A, B). Median lobe has the same length as tegmen of aedeagus, narrow, widest at basal stem (Fig. 2C, D). Apical piece of tegmen and median lobe laterally bent (Fig. 2B).</p><p>Legs long and slender. Femora dorsally with the extensor side punctured sparsely and setated with short setae, ventral surface of femora additionally with sparse and short spines. Metatibia on the extensor side shiny, with fine and sparse puncturation (Fig. 1J). Metatibiae can be curved (Fig. 1J) or straight. Ratio of length of pro-, meso- and metatibiae: 1: 1.1: 1.2. Ratio of length of protarsomeres: 1.65: 1: 1: 1: 3.3; mesotarsomeres: 1.55: 1: 1: 1: 2.1; metatarsomeres: 2.1: 1: 1: 1.8.</p><p>Female (Fig. 1B). Body length 8.5–10.5, width 3.2–3.8. Tarsi and antennae shorter than in male. Pronotum widest at anterior third (widest at anterior fourth in male), 1.23 times as wide as long, 1.3 times as wide as head. Elytra 1.35–1.4 times as wide as pronotum and 1.9–2 times as wide as head.</p><p>Ovipositor (Fig. 2G). Paraproct moderately long, 2.2 times as long as coxite; coxite lobe I large, with transverse baculi, coxite lobes II–IV merged; lobe IV with strongly sclerotized slightly curved, rounded at apex apical processes (blades).</p><p>Etymology. The species is named in honour of Abdulloh Khudayberganov, the beloved son of the first co-author and the grandson of the third co-author, who, despite his small preschool age, assists us collect beetles.</p><p>Comparative diagnosis. The new species differs from all known  Microdera by various characters, especially the long antennae with 3 (male) (Fig. 1A) or 2 (female) (Fig. 1B) antennomeres extending beyond the base of the pronotum. It can be compared with the taxa without vertical plate at the apex of abdominal ventrite 5 and the non-depressed elytra. Externally  M. abdullohi sp. nov. is slightly similar to  M. convexa (Tauscher, 1812) in the strongly convex pronotum, but differs in the much longer and slender legs (the metatarsi half or less shorter than metatibiae (Fig. 1A), while in  M. convexa metatarsi are shorter than half length of the metatibiae (Fig. 4A)) and the antennae (not reaching the base of the pronotum in  M. convexa (Fig. 4A)), the shiny body (the elytra also entirely shiny vs opaque elytral lateral sides and the apical part with disappearing puncturation in  M. convexa) and the shiny smooth outer (extensor) side of metatibiae with fine sparse punctures (Fig. 1J) (with dense and coarse confluent puncturation, appear rough at  M. convexa (Fig. 4G)). The new species differs (in addition to the long and slender antennae and legs) from  M. campestris and  M. minax in the strongly convex pronotum vs slightly convex in the two mentioned species (Fig. 3C, 13B).</p><p>Distribution. Uzbekistan: Sulton-Uvays Mts.</p></div>	https://treatment.plazi.org/id/03D42163EC29BF6DFF7FFBB8FD1AF53F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Bekchanov, Norbek Kh.;Nabozhenko, Maxim V.;Bekchanov, Khudaybergan U.;Bekchanova, Mokhira Kh.	Bekchanov, Norbek Kh., Nabozhenko, Maxim V., Bekchanov, Khudaybergan U., Bekchanova, Mokhira Kh. (2025): A review of the genus Microdera Eschscholtz, 1831 (Coleoptera: Tenebrionidae) of Uzbekistan. Zootaxa 5590 (4): 451-480, DOI: 10.11646/zootaxa.5590.4.1, URL: https://doi.org/10.11646/zootaxa.5590.4.1
03D42163EC2EBF6DFF7FFB50FC1AF64C.text	03D42163EC2EBF6DFF7FFB50FC1AF64C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Microdera campestris (Steven 1828)	<div><p>Microdera campestris (Steven, 1828)</p><p>(Fig. 3)</p><p>Tentyria campestris Steven 1828: 93 .</p><p>=  Microdera gracilis Eschscholtz 1831: 6; Pirnazarov, 1973: 15.</p><p>= Rostrax menetriesii Fischer von Waldheim 1844: 68.</p><p>=  Microdera macrops Reitter 1915: 93 .</p><p>=  Microdera campestris karanogaica Abdurakhmanov &amp; Nabozhenko 2011: 183 .</p><p>Type material examined. See in Nabozhenko &amp; Kalashian (2022).</p><p>Notes. We didn’t collect this species. Our images are based on the specimens from supralittoral sand of the Aral Sea on the former Barsakelmes Island in Kazakhstan (very close area to the western coast of former Aral). The synonymy was proposed and argued in details by Nabozhenko &amp; Kalashian (2022).</p><p>Distribution. European part of Russia: Caspian depression (Nabozhenko &amp; Kalashian 2022), Kazakhstan (from the Caspian Depression across Aral sands to middle reaches of the Syr Darya: Kyzylorda), Uzbekistan: sandy supralittoral zone of the western coast of the Aral Sea (Pirnazarov 1973).</p></div>	https://treatment.plazi.org/id/03D42163EC2EBF6DFF7FFB50FC1AF64C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Bekchanov, Norbek Kh.;Nabozhenko, Maxim V.;Bekchanov, Khudaybergan U.;Bekchanova, Mokhira Kh.	Bekchanov, Norbek Kh., Nabozhenko, Maxim V., Bekchanov, Khudaybergan U., Bekchanova, Mokhira Kh. (2025): A review of the genus Microdera Eschscholtz, 1831 (Coleoptera: Tenebrionidae) of Uzbekistan. Zootaxa 5590 (4): 451-480, DOI: 10.11646/zootaxa.5590.4.1, URL: https://doi.org/10.11646/zootaxa.5590.4.1
03D42163EC2DBF61FF7FFAA8FAB4F35B.text	03D42163EC2DBF61FF7FFAA8FAB4F35B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Microdera convexa (Tauscher 1812)	<div><p>Microdera convexa (Tauscher, 1812)</p><p>(Fig. 4)</p><p>Tentyria convexa Tauscher, 1812: 39; Pirnazarov, 1973: 15.</p><p>Microdera transversicollis v. parvicollis Reitter, 1897: 235 (nom. praeocc. nec.  Microdera parvicollis Bates, 1879); synonymized by Nabozhenko &amp; Kalashian (2022).</p><p>=  Microdera reitteri Kaszab, 1966: 280 (nom. n. nec.  Microdera parvicollis Bates, 1879); synonymized by Nabozhenko &amp; Kalashian (2022).</p><p>Material examined.  25 specimens (ZIN),   Karakalpakstan,  Western coast of Aral sea, 12 km E of Komsomolsk-on-Ustyurt (now Qubla-ustyurt), 11.vi.1972 (G.S. Medvedev) ;  5 specimens (ZIN),   Karakalpakstan,  Western coast of Aral sea, 30 km N of Komsomolsk-on-Ustyurt (now Qubla-ustyurt), 30.v.1978 (G.S. Medvedev) ;  4 specimens (ZIN),   Karakalpakstan, Karateren Lake (43°13ʹN / 60°23ʹE), 24.v.1972 (G.S. Medvedev) ;  3 specimens (ZIN),   Navoiy Region, Kyzylkum desert, Bukantau Mts., N Kulkuduk, 13.05.1965 (E.L. Gur’eva) ;  5 specimens (ZIN),   Navoiy Region, Kyzylkum desert, Bukantau Mts., 6.05.1966 (G.S. Medvedev) ;  18 specimens (PCNB),  21 specimens (PCMN):   Uzbekistan, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=58.72278&amp;materialsCitation.latitude=43.02861" title="Search Plazi for locations around (long 58.72278/lat 43.02861)">Karakalpakstan</a>, Kungrad Distr., near the mausoleum “Daut ata”, 43°01ʹ43ʺN / 58°43ʹ22ʺE, h= 60m, 26.04.2024 (M.V. and S.V. Nabozhenko, N. Kh. and Kh. U. Bekchanov, U.E. Duschanov) ;  14 specimens (PCNB),  10 specimens (PCMN):   Uzbekistan, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=58.014446&amp;materialsCitation.latitude=43.379166" title="Search Plazi for locations around (long 58.014446/lat 43.379166)">Karakalpakstan</a>, Kungrad Distr., 6.3 km NW “Barsakelmes” railway stn., NE coast of Barsakelmes dried salty lake, 43°22ʹ45ʺN / 58°00ʹ52ʺE, h= 40-80m, 27– 28.04.2024 (M.V. and S.V. Nabozhenko, N. Kh. and Kh. U. Bekchanov, Y.R. Tokhirov)  .</p><p>Type material examined.  Microdera kaszabi Skopin, 1961 Holotype, ♀ (ZIN): plate with glued abdominal ventrite 5 and inner tergite VII, “Ср. Или, Илийск, полЫнники на песках 3.05.1958 ” (Sr. Ili, Iliysk, polynniki na peskakh 3.05.1958 N. Skopin leg. ( Russian Cyrillic label), now Kazakhstan, middle  Ili River, Iliysk [flooded by Kapchagay reservoir in 1970], 43°53ʹN, 77°07ʹE,  Artemisia on the sands), “ Holotypus  Microdera kaszabi N. Skopin, 1961 ”.</p><p>The holotype of  Microdera reitteri was examined previously (Nabozhenko &amp; Kalashian 2022)</p><p>Notes. Skopin (1968) noted wide intraspecific variability of  M. convexa and identified three subspecies and five varieties of the nomynotypical subspecies:  M. convexa convexa ( M. convexa typical variety,  M. convexa var. fortesulpta Skopin, 1968,  M. convexa var. mixta Skopin, 1968,  M. covexa var. subglobosa Kraatz, 1865,  M. convexa var. laevipeda Skopin, 1968),  M. convexa kaszabi Skopin, 1961 and  M. convexa globulicollis Ménétriés, 1849 . He also provided a table with the frequency of occurrence of these varieties and subspecies in different regions of Kazakhstan.</p><p>The situation with  M. globulicollis is clear: Skopin did not study syntypes of this species and his determinations are incorrect. Erroneous interpretations of the taxa  M. campestris and  M. globulicollis by different authors, as well as the correct synonymy, are explained in detail in the paper of Nabozhenko &amp; Kalashian (2022).</p><p>No one accepted the synonymy of  M. subglobosa (synonymized by Skopin (1968) after transfer of this taxon to a status of a variety:  M. convexa var. subglobosa). The status of this taxon needs to be revised or confirmed after examination of type specimens.</p><p>The species  M. kaszabi Skopin, 1961 was later interpreted as a subspecies of  M. convexa (Skopin, 1968), that is reflected in the catalogue (Iwan et al. 2020). We examined the holotype of this taxon (Fig. 5) and again return this subspecies to the status of a species:  Microdera kaszabi Skopin, 1961, stat. resurr. This species is distinctly different from  M. convexa in many characters: the shiny body, the elytra entirely with coarser distinct puncturation (the puncturation is much finer and sparser and disappears on the opaque lateral sides and the elytral apex in  M. convexa); the lateral margins of gena converging to the epistome and the head is shorter in front of the eyes (the genae strongly diverging from eyes to widest portion and then converging to epistome in  M. convexa), the metatibiae shiny and smooth on the extensor side, with fine and sparse puncturation (opaque and appear wrinkled due to the merging of coarse, elongated punctures or at least coarsely and densely punctured in  M. convexa).</p><p>Thus, Skopin interpreted the variability very broadly and underestimated some characters, such as the shape of the head and genae, the puncturation and shine of the elytra, and the sculpture of the metatibiae.</p><p>The most variable characters in  M. convexa are the puncturation of the prohypomera and the thickness (and height) of the border of the pronotal base. The puncturation and roughness of the metatibia are also slightly variable, but punctures are always elongated, coarse and dense, often merging. Here we illustrate only Uzbek populations from Ustyurt. The specimens from the former southern Aral coast and Bukantau in Kyzylkum do not differ from the East Ustyurt specimens, only in the population from Bukantau the punctation of the pronotum is slightly finer and sparser.</p><p>Distribution. European part of Russia (Caspian depression), Azerbaijan (Apsheron Peninsula) (Nabozhenko &amp; Kalashian 2022); Kazakhstan (western regions, Ustyurt, Mangyshlak, Aral sea region, middle reaches of Syrdarya, Muyunkum desert); Uzbekistan (Ustyurt, Amudarya delta and Aral sea coast, Central Kyzylkum—Bukantau).</p></div>	https://treatment.plazi.org/id/03D42163EC2DBF61FF7FFAA8FAB4F35B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Bekchanov, Norbek Kh.;Nabozhenko, Maxim V.;Bekchanov, Khudaybergan U.;Bekchanova, Mokhira Kh.	Bekchanov, Norbek Kh., Nabozhenko, Maxim V., Bekchanov, Khudaybergan U., Bekchanova, Mokhira Kh. (2025): A review of the genus Microdera Eschscholtz, 1831 (Coleoptera: Tenebrionidae) of Uzbekistan. Zootaxa 5590 (4): 451-480, DOI: 10.11646/zootaxa.5590.4.1, URL: https://doi.org/10.11646/zootaxa.5590.4.1
03D42163EC22BF62FF7FFDC0FD72F323.text	03D42163EC22BF62FF7FFDC0FD72F323.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Microdera deserta subsp. deserta (Tauscher 1812)	<div><p>Microdera deserta deserta (Tauscher, 1812)</p><p>(Fig. 6)</p><p>Tentyria deserta Tauscher, 1812: 38 .</p><p>=  Tentyria deplanata Gebler, 1841: 373 .</p><p>Type material examined.  Tentyria deplanata (Fig. 6A, E, F). Lectotype (ZIN), ♂, hereby designated: goldish square, “Camp. Kirg or.” (the first two words are printed, the third is handwritten; east of Kirghiz steppes = central and eastern Kazakhstan),  “ deplanata Gebl. St. Kirg. or.” (handwritten), “ Lectotypus  Tentyria deplanata Gebler, 1841 des. M. Nabozhenko, 2024”. Paralectotype (ZIN), sex unknown: goldish square, “Camp. Kirg. or.”. The second paralectotype (as a syntype) from Zoological Museum of Moscow State University was mentioned by Abdurakhmanov &amp; Nabozhenko (2011). References and synonymy were indicated in Nabozhenko &amp; Kalashian (2022).</p><p>Material examined.   1 ♀ (ZIN):  Tashkent, 5.vi.1915 (T.E.S. = Turkestan Entomological Station) (Fig. 6B–D)  .</p><p>Distribution. European part of Russia: Caspian depression (Nabozhenko &amp; Kalashian 2022), Kazakhstan: from Caspian Depression across Aral sands to Ili valley and Alakol Lake (Skopin 1961a), Uzbekistan: Tashkent (new record for the country), China: Ili valley (Skopin 1961a). The second subspecies  Microdera deserta skopini Kaszab, 1966 was described from SE Kazakhstan and it is probably a good species, because distinctly differs from  M. deserta s. str. in the shape of the pronotum.</p></div>	https://treatment.plazi.org/id/03D42163EC22BF62FF7FFDC0FD72F323	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Bekchanov, Norbek Kh.;Nabozhenko, Maxim V.;Bekchanov, Khudaybergan U.;Bekchanova, Mokhira Kh.	Bekchanov, Norbek Kh., Nabozhenko, Maxim V., Bekchanov, Khudaybergan U., Bekchanova, Mokhira Kh. (2025): A review of the genus Microdera Eschscholtz, 1831 (Coleoptera: Tenebrionidae) of Uzbekistan. Zootaxa 5590 (4): 451-480, DOI: 10.11646/zootaxa.5590.4.1, URL: https://doi.org/10.11646/zootaxa.5590.4.1
03D42163EC21BF62FF7FFD10FBDFF7B8.text	03D42163EC21BF62FF7FFD10FBDFF7B8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Microdera ferghanensis Kaszab 1957	<div><p>Microdera ferghanensis Kaszab, 1957, sp. resurr.</p><p>(Figs 7, 8)</p><p>Microdera ferghanensis Kaszab, 1957: 290, 1966: 289.</p><p>Microdera ferganensis (sic): Nabozhenko &amp; Kalashian, 2022: 1370 (as a junior synonym of  M. globulicollis).</p><p>Type material examined.   Holotype (NHMB), ♂: “ Kokand Fergana ”, “  Sammlung Adr. Schuster”, “Staud.”, “Museum Frey München”, “HOLOTYPUS  Microdera ferghanensis m. det. Dr. Kaszab, 1957 ”  .  3 paratypes with the same label but “Paratypus ...” are also deposited in NHMB . The paratype from HNHM was indicated by Nabozhenko &amp; Kalashian (2022).</p><p>Material examined.  12 specimens (PCNB),  10 specimens (PCMN, ZIN):   Uzbekistan, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=70.806114&amp;materialsCitation.latitude=40.28583" title="Search Plazi for locations around (long 70.806114/lat 40.28583)">Fergana Region</a>, Sho‘rsuv, 40°17ʹ09ʺN / 70°48ʹ22ʺE, h= 630m, 16.04.2024 (M.V. and S.V. Nabozhenko, N.Kh. and Kh.U. Bekchanov) ;  3 specimens (PCNB),  2 specimens (PCMN):   Uzbekistan, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=70.966385&amp;materialsCitation.latitude=40.32" title="Search Plazi for locations around (long 70.966385/lat 40.32)">Fergana Region</a>, O‘qchi env., backward of garden, 40°19ʹ12ʺN / 70°57ʹ59ʺE, h= 680m, 16.04.2024 (M.V. and S.V. Nabozhenko, N.Kh. and Kh.U. Bekchanov)  .</p><p>Notes. The species was unfairly synonymized with  M. globulicollis by Nabozhenko &amp; Kalashian (2022). Their opinion was based on the examination of one paratype from HNHM. This paratype has the least expressed species characteristics, such as very fine puncturation of the prothoracic hypomera only near coxae, the puncturation of head basally is slightly elongated and with almost the same density as on the other surface of frons. The authors attributed this to the variability of  M. globulicollis, which also occurs in Fergana valley. After examination of the holotype (Fig. 7A–C) and a series of beetles from the Qampirqoq Mountains (S of Kokand, Fergana valley) (Fig. 7D–F, 8), the independence of this species is confirmed.  Microdera ferghanensis is clearly distinguished from  M. globulicollis by the presence of the puncturation on the entire surface of the hypomera or at least near the procoxal area (Fig. 7F), slightly elongated puncturation of the pronotum on sides (Fig. 7A, E), denser and elongated puncturation at the base of the head (Fig. 7E), as well as the angularly rounded mesoventrite between mesocoxae (Fig. 7B, 8A) (not evenly, as in  M. globulicollis (Fig. 10A)). The tarsi of  M. ferghanensis (Fig. 8F–H) are thinner than those of  M. globulicollis (Fig. 10B, C). Both species also differ in the shape of the aedeagus (compare Figs 8I and 10E). Thus, we resurrect this species:  Microdera ferghanensis Kaszab, 1957, sp. resurr. The name of the species was misspelled as  M. ferganensis by Skopin (1968), Iwan et al. (2020) and Nabozhenko &amp; Kalashian (2022).</p><p>Distribution. Uzbekistan (SW areas of Fergana Region: Qampirqoq Mts). Skopin (1968) recorded this species for central and southern Kazakhstan from Arys Lake to Syr-Darya valley and Muyunkum desert. We didn’t find any Skopin’s specimens of  M. ferghanensis in ZIN; these records need verification.</p></div>	https://treatment.plazi.org/id/03D42163EC21BF62FF7FFD10FBDFF7B8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Bekchanov, Norbek Kh.;Nabozhenko, Maxim V.;Bekchanov, Khudaybergan U.;Bekchanova, Mokhira Kh.	Bekchanov, Norbek Kh., Nabozhenko, Maxim V., Bekchanov, Khudaybergan U., Bekchanova, Mokhira Kh. (2025): A review of the genus Microdera Eschscholtz, 1831 (Coleoptera: Tenebrionidae) of Uzbekistan. Zootaxa 5590 (4): 451-480, DOI: 10.11646/zootaxa.5590.4.1, URL: https://doi.org/10.11646/zootaxa.5590.4.1
03D42163EC26BF67FF7FFF35FBFFF553.text	03D42163EC26BF67FF7FFF35FBFFF553.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Microdera globulicollis Menetries 1849	<div><p>Microdera globulicollis Ménétriés, 1849</p><p>(Figs 9, 10)</p><p>Microdera globulicollis Ménétriés, 1849: 230; Jachontov et al., 1962: 85; Davletshina, 1967: 525; Davletshina et al., 1979: 106.</p><p>=  Microdera margelanica Kraatz in Heyden, 1882: 81;</p><p>=  Microdera extabesa Skopin, 1961b: 393; Kaszab, 1966: 288; Skopin, 1968: 107, syn. nov.</p><p>Type material examined. Type material for  M. globulicollis and  M. margelanica see in Nabozhenko &amp; Kalashian (2022).</p><p>Microdera extabesa (Fig. 9A, B):   Holotype, ♂ (ZIN): “ Yu. Kizil-Kum, Ayak-Agitma 6.08.1936 N. Skopin leg.” ( Russian Cyrillic; now: Uzbekistan, Bukhara Region, 40°39ʹN, 64°29ʹE), “ ♂ ” (square), “Holotypus  Microdera extabesa N. Skopin, 1960 ”.</p><p>Paratypes. 11 specimens (ZIN): “Golodnaya step Mussa-Kuduk 25.06.1959 leg. A. Andreeva Coll. N. Skopin ” (Russian Cyrillic), “Paratypus  Microdera extabesa N. Skopin, 1960 ” ;   1♂ (ZIN): “Termez 22.09.1938 N. Skopin leg.” (Russian Cyrillic), “ Paratypus  Microdera extabesa N. Skopin, 1960 ” ;   5 specimens (ZIN): “Yu Kyzyl-Kum, Ayak-Agitma, 6.08.1936 N. Skopin leg.” (Cyrillic Russian label), “ Paratypus  Microdera extabesa N. Skopin, 1960 ”  .</p><p>Material examined.   4 specimens (ZIN):  Navoiy Region, Kyzylkum desert, Ayakguzhumdy, 21.iii.1961, 13.v.1961 (B.M. Mamaev) ;   1 specimen (ZIN):  Kuldzhuktau, Ayakguzhumdy, 2.iv.1966 (G.S. Medvedev) ;   1 specimen (ZIN):  Navoiy Region, 15 km SSW Tamdy (= Tamdybulak), 2.iv.1966 (G.S. Medvedev) ;   2 specimens (ZIN):  Navoiy Region, Aznek, 70 km W Tamdy (= Tamdybulak), 1.v.1965 (L.V. Arnoldi) ;   2 specimens (ZIN):  Navoiy Region, Kyzylkum, Auminzatau, 10.5.1966 (G.S. Medvedev) ;   2 specimens (ZIN):  Navoiy Region, Kyzylkum, sands 45 km NW Dzhing. [Dzhingildy], 13.06.1965 (L.V. Arnoldi) ;   2 specimens, 40 km W Bukhara,  Kyzylkum, 15.v.1966 (G.S. Medvedev) ;  2 specimens (ZIN): SO Kyzylkum, on clayey soil, 11.vi.1960 (P. Ler);   1 specimen (ZIN):  Bukhara Region, Karaulbazar, 10.iv.1927 (V.V. Yakhontov) ;   15 specimens (ZIN):  Tashkent Region, Bekabad, 17.v.1966 (N.G. Skopin) ;   1 ♂ (ZIN): Uzbekistan, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=66.73583&amp;materialsCitation.latitude=37.5325" title="Search Plazi for locations around (long 66.73583/lat 37.5325)">Surkhandarya Region</a>, 7 km E Aktash, 37°31ʹ57ʺN / 66°44ʹ09ʺE, h= 450m, 13.04.2024 (M.V. and S.V. Nabozhenko, N.Kh. and Kh.U. Bekchanov, D.A. Yavkachev)  .</p><p>Notes. We examined all type series of  M. extabesa and didn’t find any differences with  M. globulicollis . Skopin mentioned some differences in the shape of “thorax” in  M. extabesa and  M. globulicollis and more rounded postero-lateral corners of pronotum in  M. extabesa . We did not find these differences. The holotype of  M. extabesa completely corresponds to the lectotype of  M. globulicollis from Turkmenistan. So the following new synonymy is established:  Microdera globulicollis Ménétriés, 1849 =  Microdera extabesa Skopin, 1961: 393,  syn. nov.</p><p>We don’t know, which species mentioned Pirnazarov (1973) under the name “  Microdera globulicollis ” for Karakalpakstan (Uzbekistan).</p><p>Distribution. Turkmenistan (Mashhad-i Misrian, foothills of Kopetdag, south Turkmenistan, Badkhyz, Kugitangtau), Uzbekistan (Kyzylkum desert, Kattakum desert, Golodnaya (Hungry) steppe, Fergana valley), Kazakhstan (Eastern Kyzylkum, Josaly, Yanykurgan, Turkestan) (ZIN collection).</p></div>	https://treatment.plazi.org/id/03D42163EC26BF67FF7FFF35FBFFF553	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Bekchanov, Norbek Kh.;Nabozhenko, Maxim V.;Bekchanov, Khudaybergan U.;Bekchanova, Mokhira Kh.	Bekchanov, Norbek Kh., Nabozhenko, Maxim V., Bekchanov, Khudaybergan U., Bekchanova, Mokhira Kh. (2025): A review of the genus Microdera Eschscholtz, 1831 (Coleoptera: Tenebrionidae) of Uzbekistan. Zootaxa 5590 (4): 451-480, DOI: 10.11646/zootaxa.5590.4.1, URL: https://doi.org/10.11646/zootaxa.5590.4.1
03D42163EC24BF67FF7FFBD4FEFAF71D.text	03D42163EC24BF67FF7FFBD4FEFAF71D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Microdera heydeni Kraatz 1882	<div><p>Microdera heydeni Kraatz, 1882</p><p>(Figs 11, 12)</p><p>Microdera heydeni Kraatz, 1882: 82; Kaszab, 1966: 288.</p><p>Material examined.  1 specimen (ZIN):   Namangan env., 1903 (I. Yankovsky) ;  1 specimen (ZIN):   Fergana region, sands N Yaz’yavan, 6.v.1961 (V.A. Zaslavsky) ;  5 specimens (PCNB),  5 specimens (PCMN, ZIN):   Uzbekistan, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=71.52806&amp;materialsCitation.latitude=40.605553" title="Search Plazi for locations around (long 71.52806/lat 40.605553)">Fergana Region</a>, Yozyovon sands, 40°36ʹ20ʺN / 71°31ʹ41ʺE, h = 420 m, 17.04.2024 (M.V. and S.V. Nabozhenko, N.Kh. and Kh.U. Bekchanov) ;  11 specimens (PCNB),  8 specimens (PCMN, ZIN):   Uzbekistan, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=70.86778&amp;materialsCitation.latitude=40.64306" title="Search Plazi for locations around (long 70.86778/lat 40.64306)">Fergana Region</a>, Kokand sands, 40°38ʹ35ʺN / 70°52ʹ04ʺE, h = 360 m, 15.04.2024 (M.V. and S.V. Nabozhenko, N.Kh. and Kh.U. Bekchanov)  .</p><p>Distribution. S Kazakhstan (Skopin 1968) and Tajikistan (middle reaches of the Syr Darya), Uzbekistan (Fergana valley: west and centre) (Kaszab 1966, ZIN collection). Erroneously recorded for Turkmenistan by Iwan et al. (2020).</p></div>	https://treatment.plazi.org/id/03D42163EC24BF67FF7FFBD4FEFAF71D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Bekchanov, Norbek Kh.;Nabozhenko, Maxim V.;Bekchanov, Khudaybergan U.;Bekchanova, Mokhira Kh.	Bekchanov, Norbek Kh., Nabozhenko, Maxim V., Bekchanov, Khudaybergan U., Bekchanova, Mokhira Kh. (2025): A review of the genus Microdera Eschscholtz, 1831 (Coleoptera: Tenebrionidae) of Uzbekistan. Zootaxa 5590 (4): 451-480, DOI: 10.11646/zootaxa.5590.4.1, URL: https://doi.org/10.11646/zootaxa.5590.4.1
03D42163EC3ABF7AFF7FFF35FB51F6BA.text	03D42163EC3ABF7AFF7FFF35FB51F6BA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Microdera minax Reitter 1897	<div><p>Microdera minax Reitter, 1897</p><p>(Figs 13, 14)</p><p>Microdera minax Reitter, 1897: 233; Kaszab, 1966: 287; Pirnazarov, 1970: 27, 1973: 15; Davletshina et al., 1979: 106; Medvedev &amp; Nepesova, 1985a: 67.</p><p>Material examined.  8 specimens (ZIN):   Karakalpakstan, Kungrad District, Kyzylzhar, 9–10.v.1939 (P.A. Veltishchev) ;  1 specimen (ZIN):   Karakalpakstan, Amudarya delta, Kubetau Mt., 17.vi.1911 (K.A. Molchanov) ;  1 specimen (ZIN):   Karakalpakstan, Amudarya delta, vii (without year) (Kazakov) ;  1 specimen (ZIN):   Karakalpakstan, 70 km SO Nukus, 24.viii.1968 (B.P. Pirnazarov) ;  4 specimens (ZIN):   Karakalpakstan, 30 km S Takhta-Kupyr, 7.ix.1968 (B.P. Pirnazarov) ;  1 specimen (ZIN):   Karakalpakstan, 1 km S Nukus, 24.v.1972 (G.S. Medvedev) ;  1 specimen (ZIN):   Khorezm Region, Khiva, 22.03.1927 (L. Zimin) ;  6 specimens (ZIN):   Khorezm Region, Khiva District, Ravat, 21.iv.1927 (V. Gussakovsky) ;  6 specimens (ZIN):  the same locality, 27.iv.1927, 2.ix.1927 (L. Zimin);  1 specimen (ZIN):   Khorezm Region, Khiva District, Daniyar, 4.iv.1927 (V. Gussakovsky) ;  3 specimens (ZIN):   Khorezm Region, 6 km SSW Khiva, 31.v.1972 (G.S. Medvedev) ;  1 specimen (ZIN):   Khorezm Region, SO Meshekli, Kulatau tugay, Amudarya, 10.iv.1966 (G.S. Medvedev) ;  1 specimen (ZIN):   Bukhara Region, Karabulak, 14.v.1911 (A.K. Golbek) ;  1 specimen (ZIN):   90–100 km W Gazli, 27.v.1965 (L.V. Arnoldi) ;  3 specimens (ZIN):   Bukhara Region, 80 km NW Gazli, 12.iv.1966 (G.S. Medvedev) ;  18 specimens (ZIN):   Navoiy Region, Kuldzhuktau, 20 km S Ayakguzhumdy, 5.iv.1966 (G.S. Medvedev) ;  1 specimens (ZIN):   Navoy Region, Tamdytau, 8 km SO from Aktau Mt., 4.v.1965 (E.L. Gur’eva) ;  2 specimens (ZIN):   Navoy Region, Tamdybulak, 7.v.1965 (L.V. Arnoldi) ;  3 specimens (PCNB),  8 specimens (PCMN):   Uzbekistan, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=64.02945&amp;materialsCitation.latitude=39.171112" title="Search Plazi for locations around (long 64.02945/lat 39.171112)">Bukhara Region</a>, SW coast of Dengizkul lake, 39°10ʹ16ʺN / 64°01ʹ46ʺE, h= 180m, 11– 12.04.2024 (M.V. and S.V. Nabozhenko, N.Kh. and Kh.U. Bekchanov, D.A. Yavkachev) ;  4 specimens (PCNB),  3 specimens (PCMN):   Uzbekistan, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=59.095554&amp;materialsCitation.latitude=43.130833" title="Search Plazi for locations around (long 59.095554/lat 43.130833)">Karakalpakistan</a>, Kungrad Distr., Kyzylzhar env.,43°06ʹ58ʺN / 59°03ʹ39ʺE, h=70m and 43°07ʹ51ʺN / 59°05ʹ44ʺE, h= 80–100m, 25.04.2024 (M.V. and S.V. Nabozhenko, N.Kh. and Kh.U. Bekchanov, U.E. Duschanov) ;  9 specimens (PCNB):   Uzbekistan, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=60.65364&amp;materialsCitation.latitude=41.9" title="Search Plazi for locations around (long 60.65364/lat 41.9)">Karakalpakistan</a>, Beruniy Distr., 41°53ʹ60ʺN / 60°39ʹ13.1ʺE, 19.03.2024 (N.Kh.Bekchanov, Kh. U. Bekchanov and M.Kh.Bekchanova)  .</p><p>Variability. Disc of the pronotum can be with elongate puncturation on sides.</p><p>Distribution. Turkmenistan (Karakum desert), Uzbekistan (Karakum and Kyzylkum deserts, sands of Amudarya delta) (Medvedev &amp; Nepesova 1985a; Bekchanov et al., 2023; ZIN collection), NW Afghanistan (Kushka) (Gridelli 1954, Kaszab 1959, 1966, ZIN collection), Kazakhstan (Kyzylkum, Muyunkum, Ili River) (ZIN collection). The record of this species for Tien Shan (Kyrgyzstan: Naryn) (Gridelli 1954) is very doubtful.</p></div>	https://treatment.plazi.org/id/03D42163EC3ABF7AFF7FFF35FB51F6BA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Bekchanov, Norbek Kh.;Nabozhenko, Maxim V.;Bekchanov, Khudaybergan U.;Bekchanova, Mokhira Kh.	Bekchanov, Norbek Kh., Nabozhenko, Maxim V., Bekchanov, Khudaybergan U., Bekchanova, Mokhira Kh. (2025): A review of the genus Microdera Eschscholtz, 1831 (Coleoptera: Tenebrionidae) of Uzbekistan. Zootaxa 5590 (4): 451-480, DOI: 10.11646/zootaxa.5590.4.1, URL: https://doi.org/10.11646/zootaxa.5590.4.1
03D42163EC38BF7CFF7FFF35FB79F77F.text	03D42163EC38BF7CFF7FFF35FB79F77F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Microdera semenoviana Bogatchev 1947	<div><p>Microdera semenoviana Bogatchev, 1947</p><p>(Figs 15, 16)</p><p>=  Microdera semenovi Bogatchev, 1946: 210 (nom. praeocc. nec.  Microdera semenovi Reitter, 1897).</p><p>Microdera semenoviana Bogatchev, 1947: 463 (nom. n.); Medvedev &amp; Nepesova, 1985a: 68.</p><p>Type material examined.   Lectotype (Fig. 15A, B), ♂ (ZIN) hereby designated: “Пархар на р. ПЯндж. Таджикист. Луппова 9.VIII.934 ” (Russian Cyryllic: Parkhar na r. Pyandzh.Tadzhikist. Luppova 9.VIII.934 = Tajikistan,  Parkhar on Pyanj River), “LECTOTYPUS  Microdera semenoviana Bogatchev, 1947 des. M. Nabozhenko et N. Bekchanov”  .   Paralectotypes: 1 specimen (ZIN): “  Parkhar na r. Pyandzh. Tadzhikist . Luppova 9.VIII.934 ” (Russian Cyrillic), “PARALECTOTYPUS  Microdera semenoviana Bogatchev, 1947 det. M. Nabozhenko ” ;  1 specimen (ZIN): the same locality, but 10.viii.1934 and the same type label;   1 specimen (ZIN): “КЫЗЫл-Кала, р. Вахш. Тадж. 9.VI.934 Гуссаковский” (Russian Cyrillic:  Kyzyl-Kala, r. Vakhsh. Tadzh . 9.VI.1934 Gussakovsky = Tajikistan, Vakhsh River, Kyzyl-Kala) and the same type label ;   1 specimen (ZIN): “Кабадиан, Таджикист. 10.VII.934 Гуссаковский” (Russian Cyrillic:  Kabadian, Tadzhikist. 10.VII.1934, Gussakovsky = Tajikistan, Qubodiyon) and the same type label  .</p><p>Material examined.   2 specimens (ZIN collection): Uzbekistan /Tajikistan, S Babatag, between  Dzharkurgan [Uzbekistan] and Shaartuz [Tajikistan], 19.iv.1962 (E.L. Gur’eva)  .</p><p>Distribution. SE Turkmenistan (Kugitangtau Mts: Garlyk), Uzbekistan (Babatag Mts), SW Tajikistan, Afghanistan (Bogatchev 1946, Kaszab 1968, Medvedev &amp; Nepesova 1985a, ZIN collection).</p></div>	https://treatment.plazi.org/id/03D42163EC38BF7CFF7FFF35FB79F77F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Bekchanov, Norbek Kh.;Nabozhenko, Maxim V.;Bekchanov, Khudaybergan U.;Bekchanova, Mokhira Kh.	Bekchanov, Norbek Kh., Nabozhenko, Maxim V., Bekchanov, Khudaybergan U., Bekchanova, Mokhira Kh. (2025): A review of the genus Microdera Eschscholtz, 1831 (Coleoptera: Tenebrionidae) of Uzbekistan. Zootaxa 5590 (4): 451-480, DOI: 10.11646/zootaxa.5590.4.1, URL: https://doi.org/10.11646/zootaxa.5590.4.1
03D42163EC3FBF7EFF7FF92DFDC4F587.text	03D42163EC3FBF7EFF7FF92DFDC4F587.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Microdera shasenema G. S. Medvedev et Nepesova 1985	<div><p>Microdera shasenema G.S. Medvedev et Nepesova, 1985</p><p>(Fig. 17)</p><p>Microdera shasenema Medvedev &amp; Nepesova, 1985b: 359, Fig. 3–6, 360; Medvedev &amp; Nepesova, 1985a: 68.</p><p>Type material examined.   Holotype (Fig. 17H, I), ♂ (ZIN): “ЮЗ Турк. Мешед-Мессериан Г.Медведев12 М 984” (YuZ Turk. Meshed-Messerian G. Medvedev 12.V 984) (Russian Cyrillic label (now SW Turkmenistan, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=54.624443&amp;materialsCitation.latitude=38.27" title="Search Plazi for locations around (long 54.624443/lat 38.27)">Mashhadi Misrian</a> (=Dehistan), 38°16′12″N, 54°37′28″E)  .   Paratypes. 21 paratypes (ZIN): “  Shasenem, sev. Karakumy Kir’yakova 7/VI 953” (Russian Cyrillic label) (N Turkmenistan, Shasenem railway station, 41°39ʹN, 59°23ʹE), “na svet” (at light; Russian Cyrillic label), “Paratypus  Microdera shasenema sp. nov. Medv. et Nepes.” (the same label for all paratypes) ;   6 paratypes (ZIN): “YuZ Turk.  Meshed-Messerian G. Medvedev 12.V 984 ” (Russian Cyrillic label) (SW Turkmenistan, Mashhad-i Misrian) ;   2 paratypes (ZIN): “  Krasnovodsk Zakasp. obl. 28.III.916 B. Il’in” (Russian Cyrillic label) (now Turkmenistan, Turkmenbashi);  8 paratypes (ZIN): 5 km Z Khodzheyli G. Medvedev 4 VI 972” (Russian Cyrillic label (now in Uzbekistan, Karakalpakstan);   1 paratypes (ZIN): “  N Shakhi-burun B. Balkhany Turkm . V. Popov 16 VI 934 ” (Russian Cyrillic label) ;   6 paratypes (ZIN): “  Beltau, oz. Karateren G. Medvedev 24.V 972 ” (Russian Cyrillic label) ;   2 paratypes (ZIN): “  Zap. ber. Aral m. 12 km V Komsomol’ska G. Medvedev II VI 972” (Russian Cyrillic label) (western coast of the Aral sea, 12 km W of Komsomolsk-on-Ustyurt = Qubla ustyurt) ;   2 paratypes (ZIN): “  Karakalpak 40 km W Nukus 14-VII-1966 Pirnazarov ” (Russian Cyrillic label) ;   1 paratype (ZIN): “  Karakalpak Nukus 20-V-1969 Pirnazarov ” (Russian Cyrillic label), “N 21” ;   1 paratype (ZIN) “fl.  Amudarya ” ;   6 paratypes (ZIN):  Karakalpakstan, Beltau, Karateren Lake, 24.v.1972 (G.S. Medvedev) (Russian Cyrillic label)  .</p><p>Material examined.   1 specimen (ZIN): Uzbekistan,  Karakalpakstan, Beltau, Karateren Lake, 24.v.1972 (G.S. Medvedev) ;   1 ♂ (PCMN): Uzbekistan, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=59.095554&amp;materialsCitation.latitude=43.130833" title="Search Plazi for locations around (long 59.095554/lat 43.130833)">Karakalpakstan</a>, Kungrad Distr., Kyzylzhar env., 43°07ʹ51ʺN / 59°05ʹ44ʺE, h = 80–100 m, 25.04.2024 (M.V. and S.V. Nabozhenko, N.Kh. and Kh.U. Bekchanov, U.E. Duschanov)  .</p><p>Additional material examined.   4 specimens (ZIN): Tajikistan, Khatlon Region,  Chiluchorchashma W of Shaartuz, 20.iv.1962 (collector unknown)  .</p><p>Notes. Specimens from the Amudarya delta (Fig. 17A–G, J–L), the coast of the former Aral Sea and NE Karakum (Turkmenistan: Shasenem) differ from those in SW Turkmenistan, having more slender body, less transverse pronotum, more elongate tarsi, lengths of metatarsomeres 1 and 4 are subequal (Fig. 17D) (metatarsomere 1 distinctly shorter than 4 in populations from SW Turkmenistan (Fig. 17I)). The mentioned characters are slightly variable in the population from Uzbekistan and  northern Karakum (Turkmenistan: Shasenem). The population from SW Tajikistan more similar to those from SW Turkmenistan, but beetles also slightly differ from other two populations. Medvedev in the original description used characters of the north populations for proportions of metatarsi, while the holotype and specimens from the type locality have other proportions. The status of three disjunctive populations of  M. shasenema can be established after examination of a larger number of specimens, especially from the area of the middle reaches of Amudarya (in case the species occurs there). We use characters of the north population (from Uzbekistan and NE Turkmenistan) in a key below.</p><p>Distribution. Turkmenistan (Karakum desert), Uzbekistan (Karakalpakstan, lower reaches of Amudarya, Aral coast) (Medvedev &amp; Nepesova 1985b; ZIN collection), Tadjikistan (valley of Kofarnihon, tributary of Amudarya river; new record for the country).</p></div>	https://treatment.plazi.org/id/03D42163EC3FBF7EFF7FF92DFDC4F587	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Bekchanov, Norbek Kh.;Nabozhenko, Maxim V.;Bekchanov, Khudaybergan U.;Bekchanova, Mokhira Kh.	Bekchanov, Norbek Kh., Nabozhenko, Maxim V., Bekchanov, Khudaybergan U., Bekchanova, Mokhira Kh. (2025): A review of the genus Microdera Eschscholtz, 1831 (Coleoptera: Tenebrionidae) of Uzbekistan. Zootaxa 5590 (4): 451-480, DOI: 10.11646/zootaxa.5590.4.1, URL: https://doi.org/10.11646/zootaxa.5590.4.1
03D42163EC3DBF7EFF7FFAB4FD50F6F1.text	03D42163EC3DBF7EFF7FFAB4FD50F6F1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Microdera transversicollis Reitter 1887	<div><p>Microdera transversicollis Reitter, 1887</p><p>(Fig. 18)</p><p>Microdera transversicollis Reitter, 1887: 517; Medvedev &amp; Nepesova, 1985a: 68.</p><p>Type material examined.   Syntype, sex unknown (ZIN): goldish circle, “Tekke Radde Reitter” (handwritten), “к. Г. Сиверса” (Russian Cyrillic; collection of G.I. Sivers), “  Microdera transversicollis n. sp. ” (old label, handwritten). Gridelli indicated another syntype of this species deposited in  Natural History Museum Budapest with labels: “Turcmenien. Leder. Reitter”  .</p><p>Material examined.   2 specimens (ZIN):  Bukhara Region, Kyzylkum, 40 km W Bukhara, 15.v.1966 (G.S. Medvedev) ;   1 specimen (ZIN): “Tashk. U….. V.4.12” ( Tashkent District, 4.v.1912) ;   5 specimens (ZIN): S Kyzylkum,  Basilbek, 11.vi.1936 (N.G. Skopin) ;   6 specimens (ZIN):  Navoyi Region, NW Bukantau Mts., S Kyzylkum, Tortkuduk, 28.vii.1936 (N.G. Skopin) ;   1 specimen (ZIN):  Navoiy Region, Susyz-Kara-Tau (= Kuldzhuktau Mts.), 20.vi.1936 (N.G. Skopin) ;   1 specimen (ZIN):  Navoyi Region, Soz-batyr, 19.vi.1936 (N.G. Skopin) ;   1 specimen (ZIN):  Navoyi Region, Dzhetymtau, 20.vi.1936 (N.G. Skopin) ;   4 specimens (ZIN):  Navoyi Region, Manamdzhan, 10.vi.1936  (N.G. Skopin).</p><p>Distribution. Kazakhstan (Kyzylkum desert), Turkmenistan (Karakum desert), Uzbekistan (Kyzylkum desert) (ZIN collection), Afghanistan (ZIN collection; Gridelli 1954, Kaszab 1959, 1966). The species was erroneously recorded for Armenia by Iwan et al. (2020).</p></div>	https://treatment.plazi.org/id/03D42163EC3DBF7EFF7FFAB4FD50F6F1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Bekchanov, Norbek Kh.;Nabozhenko, Maxim V.;Bekchanov, Khudaybergan U.;Bekchanova, Mokhira Kh.	Bekchanov, Norbek Kh., Nabozhenko, Maxim V., Bekchanov, Khudaybergan U., Bekchanova, Mokhira Kh. (2025): A review of the genus Microdera Eschscholtz, 1831 (Coleoptera: Tenebrionidae) of Uzbekistan. Zootaxa 5590 (4): 451-480, DOI: 10.11646/zootaxa.5590.4.1, URL: https://doi.org/10.11646/zootaxa.5590.4.1
03D42163EC33BF71FF7FFF35FC33F60F.text	03D42163EC33BF71FF7FFF35FC33F60F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Microdera uzbekistanica N. Bekchanov et Nabozhenko 2025	<div><p>Microdera uzbekistanica N. Bekchanov et Nabozhenko,  sp. nov.</p><p>(Figs 19, 20)</p><p>Type material (ZIN).   Holotype, ♂: «АЯкгужумдЫ Кульджуктау Г. Медведев 1.IV.966» (Uzbekistan, Bukhara / Navoi regions, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=63.7625&amp;materialsCitation.latitude=40.75028" title="Search Plazi for locations around (long 63.7625/lat 40.75028)">Kuldzhuktau Mts, Ayakguzhumdy</a>, 40°45ʹ1ʺN / 63°45ʹ45ʺE, 1.iv.1966, leg. G.S. Medvedev)  .  Paratypes. 20 specimens (unsexed): the same locality and collector, dates from 1 to 5.iii.1966;   4 specimens:  Navoi / Bukhara regions, Kuldzhuktau, Mts., 15.03.1961 (leg. B. Mamaev) ;  4 specimens: the same locality and collector, but 17.03.1961;  1 specimen: the same locality, 12.v.1966 (leg. Pastukhov);   1♂, 2 specimens (unsexed): Navoi Region,  Kudzhuktau Mts, Sultanbibi draw-well, 24.iii.1961 (leg. B. Mamaev, collection of N. Skopin);   Bukhara Region,  Zhamansay natural boundary, 140 km NW Shafirkan (20–25 km NE Ayakguzhumdy), 27.x.1969 (leg. M. Falkovitsh)  .</p><p>Description. Male. Body moderately slender, convex, opaque, black or dark-brown, legs brown to black. Measurements: PH w —1.38, P 1 P w —1.13, E l E w —1.5–1.52, EH w —1.8–1.83, EP w —1.34–1.36, EP l —2.24–2.25. Body length 7.5–10 mm, width 2.5 mm</p><p>Head (Fig. 19D) dorsally widest at eye level, where 1.03 times as wide as at genal level and 1.35 times as wide as interocular space. Eyes comparatively large (in length dorsally), moderately convex. Temple not protruding and not forms angle with posterior edge of eye (lateral edge of head not angular at level of temple and eye), visual area of eye lateral. Supraorbital keels slightly sinuous, long. Prong of anterior margin of epistome slightly asymmetric and slightly projected, apex narrowly rounded; margins of epistome on sides of prong emarginated (left weak, right stronger) (Fig. 20A). Head with regular fine and sparse puncturation of round punctures, interpuncture space 1.5–2 times as long as puncture diameter. Puncturation of head ventrally very sparse and very fine, epicranium at middle almost without punctures. Antennae moderately long, reaching base of pronotum.Antennomere 3 longest, 1.7 times as long as 2nd antennomere and 1.5 times as long as 4th antennomere; antennomeres 1 and 9 widest (1.2 times as long as wide for the latter). Five proximal antennomeres coarser and denser punctured that rest.</p><p>Prothorax (Fig. 19D–F). Pronotum slightly transverse, widest slightly ahead of middle. Anterior margin almost straight, lateral margins slightly rounded in basal half and strongly rounded in apical half. Base strongly evenly rounded. Anterior margin with widely interrupted in middle sulcus, lateral margins finally margined, base widely margined. Antero-lateral corners widely rounded; postero-lateral corners obtuse. Disc of pronotum strongly convex, flattened along base in middle (Fig. 19 A–B), so disc outline in lateral view emarginate in the basal portion (Fig. 19F). Puncturation of disc fine and sparse, often finer and sparser than on head; puncturation in basal portion coarser than on other surface. Prosternum very finely and sparsely punctured, smoothly wrinkled on sides. Prothoracic hypomera sparsely punctured only near procoxae, other surface without punctures, only with very small and short setae (Fig. 19E). Prosternal process declivious rounded.</p><p>Pterothorax (Fig. 19A–C). Elytra oval, not elongated apically. Surface of elytra slightly shiny in middle and opaque at apex and on lateral sides. Puncturation very fine and sparse (finer and sparser than on pronotum), disappearing apically. Anterior half of mesoventrite with transverse bracket-like foveae on sides and transverse oval punctures in middle; posterior half with thin long transverse grooves; intercoxal process of mesoventrite convex (evenly rounded in lateral view), widely longitudinally depressed in middle, finely and sparsely punctured, sides wrinkled. Mesepisterna, mesepimera and metepisterna sparsely and finely punctured; metepimera not punctured. Metaventrite with sparse rasp-like larger punctures along mesocoxal cavities and very sparse and almost invisible punctures on rest surface.</p><p>Abdomen (Fig. 19C, 20B, C). Abdominal ventrites very sparsely and finely punctured. Intercoxal process of abdominal ventrite 1 not bordered at the apex. Abdominal ventrite 5 with rectangular emargination at apex, without small but distinct vertical plate in middle; this plate with small emargination in middle. Male aedeagus (Fig. 20D–F) almost the same as in  M. abdullohi sp. nov. and  M. minax, with bisinuate lateral margins; basal piece 1.13 times as short as apical piece, widest at basal third. Spiculum gastrale with slightly oval blades, without lateral processes (Fig. 20G).</p><p>Legs moderately long. Femora setated with short setae, additionally with sparse and short reddish spines. Metatibiae straight, opaque on extensor side, with very coarse and dense merged puncturation, look rough (Fig. 19G).</p><p>Female usually with slightly more robust body and slightly wider pronotum (Fig. 19B). Body length 8–10.5 mm, width 2.7–4 mm. Small males and females are almost indistinguishable.</p><p>Etymology. The name is derived from the country Uzbekistan.</p><p>Comparative diagnosis. The new species is externally very similar to  M. convexa by the body shape, the opaque elytra with disappearing puncturation apically and laterally and coarsely rough metatibiae on the extensor side but differs from the latter in the very fine and sparse puncturation of the head and pronotum, the non-punctured prohypomera and presence of the vertical plate at rectangularly emarginated apex of the abdominal ventrite 5 (Fig. 20C). The new species differs from all known  Microdera spp. by the flattened pronotal base in middle (the outline of the disc appears as emarginated basally in lateral view) (Fig. 19A, B, F).</p></div>	https://treatment.plazi.org/id/03D42163EC33BF71FF7FFF35FC33F60F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Bekchanov, Norbek Kh.;Nabozhenko, Maxim V.;Bekchanov, Khudaybergan U.;Bekchanova, Mokhira Kh.	Bekchanov, Norbek Kh., Nabozhenko, Maxim V., Bekchanov, Khudaybergan U., Bekchanova, Mokhira Kh. (2025): A review of the genus Microdera Eschscholtz, 1831 (Coleoptera: Tenebrionidae) of Uzbekistan. Zootaxa 5590 (4): 451-480, DOI: 10.11646/zootaxa.5590.4.1, URL: https://doi.org/10.11646/zootaxa.5590.4.1
03D42163EC31BF73FF7FF9A7FA4FF2C8.text	03D42163EC31BF73FF7FF9A7FA4FF2C8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Microdera Eschscholtz 1831	<div><p>Key to species of the genus  Microdera of Uzbekistan</p><p>1. Apex of abdominal ventrite 5 with vertical plate, that can have deep or shallow median notch (Figs 8E, 9G, 11F, 15G, 20C). 2</p><p>- Apex of abdominal ventrite 5 without vertical plate (Figs 1I, 3G, 4F, 13F)........................................ 6</p><p>2. Disc of pronotum flattened medio- basally (Fig. 19A, B); disc outline in lateral view emarginated in the basal portion (Fig. 19F). Elytra with opaque lateral sides and apical portion, where puncturation disappears. Metatibiae opaque from extensor side, with very coarse and dense merged puncturation; appear rough (Fig. 19G).........................  M. uzbekistanica sp. nov.</p><p>- Disc of pronotum evenly convex. Elytra without opaque areas, puncturation fine and sparse, but distinct on all surface. Metatibiae shiny from extensor side, with fine and sparse punctures (Figs 8B, 10D, 12A, 15I)......................... 3</p><p>3. Prothoracic hypomera with large and coarse transverse wrinkles (Fig. 15H). Lateral margin of head (dorsal view) protruded and angular at level of eye-temple connection (Fig. 15C).............................................  M. semenoviana</p><p>- Prothoracic hypomera not wrinkled (Figs 7F, 10A, 11D). Lateral margin of eye together with temple rounded............ 4</p><p>4. Eyes small (Fig. 11C), margins of epistomal prong not emarginated; prong widely rounded with not strongly projected apex (11B). Pronotum slightly convex, more transverse (1.23–1.25 times as wide as long)........................  M. heydeni</p><p>- Eyes large (Figs 7E, 9E), margins of epistomal prong widely emarginated; prong projected (Figs 8C, 9D). Pronotum strongly convex, less transverse (1.1–1.17)........................................................................ 5</p><p>5. Head basally with coarser and denser elongate puncturation (Fig. 7E). Pronotum with slightly elongate puncturation on lateral sides (Fig. 7A, E). Prothoracic hypomera with sparse and fine, but distinct puncturation at least in basal half (Fig. 7F). Intercoxal process of mesoventrite angularly rounded (lateral view) (Figs 7B, 8A)..............................  M. ferghanensis</p><p>- Head basally with the same sparse and fine puncturation as remaining parts of dorsum (Fig. 9E). Pronotum entirely with simple round puncturation (Fig. 9E). Prothoracic hypomera not punctured (Fig. 10A). Intercoxal process of mesoventrite, strongly but not angularly rounded (lateral view) (Fig. 10A).................................................  M. globulicollis</p><p>6. Antennae long, with two (female) (Fig. 1B) or three (male) (Fig. 1A, C) apical antennomeres exceeding beyond base of pronotum. Prosternal process sharply bends in middle between horizontal and vertical portions (Fig. 1F, G)..................................................................................................  M. abdullohi sp. nov.</p><p>- Antennae shorter, reaching or not base of pronotum (Figs 3A, 4A, 6B, 13A, 17A, 18H). Prosternal process declivious rounded (Figs 3D, 4D, 13G).................................................................................... 7</p><p>7. Prothoracic hypomera smooth, without visible puncturation (Figs 17E, 18E)...................................... 8</p><p>- Prothoracic hypomera moderately or coarsely punctured (Figs 3D, 4D, 13G)...................................... 9</p><p>8. Epistomal prong sharply separated from epistome, with subparallel sides (Fig. 18B). 1 st metatarsomere shorter than 4 th one (Fig. 18D).................................................................................  M. transversicollis</p><p>- Epistome with simple triangular non-separated prong (Fig. 17B). Metatarsomeres 1 and 4 subequal in length (only specimens from Uzbekistan and NE Turkmenistan) (Fig. 17D)................................................  M. shasenema</p><p>9. Prothoracic hypomera with very coarse and dense sharp puncturation (Fig. 13G).............................  M. minax</p><p>- Prothoracic hypomera with disappearing, fine or moderately coarse and sparse puncturation (Figs 3D, 4D)............. 10</p><p>10. Eyes small and slightly convex (Fig. 6C). Epistomal prong almost not expressed, epistome widely rounded in middle (Fig. 6D). Elytra widely longitudinally depressed in middle (Fig. 6A, B)...........................................  M. deserta</p><p>- Eyes large and moderately convex (Figs 3C, 4C). Epistome with triangle prong in middle (Figs 3B, 4B). Elytra not depressed (Figs 3A, 4A)....................................................................................... 11</p><p>11. Pronotum slightly convex (Fig. 3A, C). Elytra not opaque laterally and apically, entirely with distinct moderately dense puncturation (Fig. 3A).......................................................................  M. campestris</p><p>- Pronotum strongly convex (Fig. 4A, C). Elytra opaque laterally and apically, with disappearing puncturation on opaque areas (Fig. 4A)....................................................................................  M. convexa</p></div>	https://treatment.plazi.org/id/03D42163EC31BF73FF7FF9A7FA4FF2C8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Bekchanov, Norbek Kh.;Nabozhenko, Maxim V.;Bekchanov, Khudaybergan U.;Bekchanova, Mokhira Kh.	Bekchanov, Norbek Kh., Nabozhenko, Maxim V., Bekchanov, Khudaybergan U., Bekchanova, Mokhira Kh. (2025): A review of the genus Microdera Eschscholtz, 1831 (Coleoptera: Tenebrionidae) of Uzbekistan. Zootaxa 5590 (4): 451-480, DOI: 10.11646/zootaxa.5590.4.1, URL: https://doi.org/10.11646/zootaxa.5590.4.1
