taxonID	type	description	language	source
03D42163EC29BF6DFF7FFBB8FD1AF53F.taxon	description	(Figs 1, 2)	en	Bekchanov, Norbek Kh., Nabozhenko, Maxim V., Bekchanov, Khudaybergan U., Bekchanova, Mokhira Kh. (2025): A review of the genus Microdera Eschscholtz, 1831 (Coleoptera: Tenebrionidae) of Uzbekistan. Zootaxa 5590 (4): 451-480, DOI: 10.11646/zootaxa.5590.4.1, URL: https://doi.org/10.11646/zootaxa.5590.4.1
03D42163EC29BF6DFF7FFBB8FD1AF53F.taxon	materials_examined	Type material. Holotype, ♂ (ZIN) and paratypes, 2 ♂, 4 ♀ (ZIN, PCMN), 1 ♀ (PCNB), Uzbekistan, Karakalpakistan, lava fields in the foothills of Sulton-Uvays Mts opposite Lower Amudarya State Biosphere Reserve, 42 ° 01 ʹ 49 ʺN / 60 ° 24 ʹ 25 ʺE, h = 100 m, 24.04.2024 (M. V. and S. V. Nabozhenko, N. Kh. Bekchanov, U. E. Duschanov); 2 ♀ (ZIN), Uzbekistan, Karakalpakstan, Beruni District, turning from the road to Lower Amudarya State Biosphere Reserve, 42 ° 01 ʹ 27 ʺN / 60 ° 26 ʹ 34 ʺE, 24 – 25.04.2023 (M. V. and S. V. Nabozhenko, N. Kh. and Kh. U. Bekchanov); 1 ♂ (ZIN), 1 ♂, 3 ♀ (PCNB), the same locality, 19.03.2024 (N. Kh. Bekchanov, Kh. U. Bekchanov and M. Kh. Bekchanova).	en	Bekchanov, Norbek Kh., Nabozhenko, Maxim V., Bekchanov, Khudaybergan U., Bekchanova, Mokhira Kh. (2025): A review of the genus Microdera Eschscholtz, 1831 (Coleoptera: Tenebrionidae) of Uzbekistan. Zootaxa 5590 (4): 451-480, DOI: 10.11646/zootaxa.5590.4.1, URL: https://doi.org/10.11646/zootaxa.5590.4.1
03D42163EC29BF6DFF7FFBB8FD1AF53F.taxon	description	Description. Male. Body slender, convex, shiny, black, with long and slender antennae and legs, distal parts of legs and antennae brown. Measurements: Body length 10 mm, width 3.6 mm; PH w = 1.3; P 1 P w = 1.13; E l E w = 1.55; EH w = 1.85; EP w = 1.42; EP l = 2.47. Head (Fig. 1 D, E) dorsally widest at eye level, where 1.03 times as wide as at genal level and 1.3 times as wide as interocular space. Eyes comparatively large (in length dorsally), moderately convex. Temple not protruding and not forms angle with posterior edge of eye (lateral edge of head not angular at level of temple and eye), visual area of eye lateral. Supraorbital keels slightly sinuous, long. Prong of anterior margin of epistoma slightly asymmetric, rounded, not separated as a subparallel small process; margins of prong not emarginated (Fig. 1 E). Puncturation of head coarse and dense dorsally: punctures sparser in anterior half of head than on frons; punctures smaller and denser along margin of head (especially prong and margin of gena). Puncturation of head ventrally sparse; mentum with small and round punctures; temples ventrally with sparse punctures, every puncture has a small seta. Antennae long, with 3 distal antennomeres extending beyond base of pronotum. Antennomere 3 longest (length to width ratio: 3.6 – 3.7), 1.95 times as long as 2 nd antennomere and 1.7 times as long as 4 th antennomere; antennomeres 1 and 8 widest (1.2 times as long as wide for the latter). Proximal antennomeres moderately coarsely and densely punctured, while three distal antennomeres with very fine and sparse punctures. Prothorax (Fig. 1 D, F, G). Pronotum transverse, trapezoidal, widest at anterior fourth. Anterior margin almost straight, lateral margins very weakly rounded or almost straight in basal two thirds and strongly rounded in apical third, base evenly weakly rounded. Antero-lateral corners widely rounded; postero-lateral corners obtuse. Anterior edge margined only on sides, while lateral margins and base completely margined. Base much wider margined. Disc of pronotum strongly evenly convex, sparsely punctured by round moderately coarse punctures (inperpuncture distance 3 – 4 time wider than punctures). Prosternum sparsely but distinctly punctured by fine punctures, smoothly wrinkled on sides. Prothoracic hypomera with moderately coarse distinct sparse puncturation (Fig. 1 G) or sparse and finer puncturation and smooth wrinkles (Fig. 1 F), some specimens have smaller or almost invisible and sparser punctures than on prosternum. Prosternal process with sharp bend between dorsal and vertical portions (Fig. 1 F, G). Pterothorax (Fig. 1 A, C). Elytra oval, widest at middle. Puncturation of elytra very fine and sparse, much sparser and finer than on pronotum. Anterior half of mesoventrite with transverse rasp-like bracket-like foveae or roughly merged foveae (one male specimen has rough lines); posterior half with thin long transverse grooves; intercoxal process of mesoventrite convex, widely longitudinally depressed in middle, finely and sparsely punctured. Mesepisterna, mesepimera and metepisterna sparsely and finely punctured; metepimera not punctured. Metaventrite with sparse rasp-like larger punctures at anterior half and very sparse and almost invisible punctures at posterior half. Abdomen (Fig. 1 C, H, I). Abdominal ventrites evenly punctured with sparse and fine punctures. Intercoxal process of abdominal ventrite 1 not margined at apex. Abdominal ventrite 5 widely roundely emarginated at apex, without vertical plate in middle (Fig. 1 H, I). Male terminalia and genitalia (Fig. 2 A – F). Inner tergite VIII shortly emarginated at apex, while inner sternite VIII with deep triangle emargination in middle and acute apices. Inner sternite VIII more sclerotized in basal sides (Fig. 2 E). Spiculum gastrale with transverse triangular blades having long lateral processes (Fig. 2 F). Apical piece of tegmen unevenly narrowed from base to apex, bisinuate, lateral margins shortly emarginated in basal fourths and widely emarginated in middle, widest at base, apex acute; basal piece 1.3 times as wide and 1.1 times as short as apical piece, widest at basal third (Fig. 2 A, B). Median lobe has the same length as tegmen of aedeagus, narrow, widest at basal stem (Fig. 2 C, D). Apical piece of tegmen and median lobe laterally bent (Fig. 2 B). Legs long and slender. Femora dorsally with the extensor side punctured sparsely and setated with short setae, ventral surface of femora additionally with sparse and short spines. Metatibia on the extensor side shiny, with fine and sparse puncturation (Fig. 1 J). Metatibiae can be curved (Fig. 1 J) or straight. Ratio of length of pro-, meso- and metatibiae: 1: 1.1: 1.2. Ratio of length of protarsomeres: 1.65: 1: 1: 1: 3.3; mesotarsomeres: 1.55: 1: 1: 1: 2.1; metatarsomeres: 2.1: 1: 1: 1.8. Female (Fig. 1 B). Body length 8.5 – 10.5, width 3.2 – 3.8. Tarsi and antennae shorter than in male. Pronotum widest at anterior third (widest at anterior fourth in male), 1.23 times as wide as long, 1.3 times as wide as head. Elytra 1.35 – 1.4 times as wide as pronotum and 1.9 – 2 times as wide as head. Ovipositor (Fig. 2 G). Paraproct moderately long, 2.2 times as long as coxite; coxite lobe I large, with transverse baculi, coxite lobes II – IV merged; lobe IV with strongly sclerotized slightly curved, rounded at apex apical processes (blades).	en	Bekchanov, Norbek Kh., Nabozhenko, Maxim V., Bekchanov, Khudaybergan U., Bekchanova, Mokhira Kh. (2025): A review of the genus Microdera Eschscholtz, 1831 (Coleoptera: Tenebrionidae) of Uzbekistan. Zootaxa 5590 (4): 451-480, DOI: 10.11646/zootaxa.5590.4.1, URL: https://doi.org/10.11646/zootaxa.5590.4.1
03D42163EC29BF6DFF7FFBB8FD1AF53F.taxon	etymology	Etymology. The species is named in honour of Abdulloh Khudayberganov, the beloved son of the first co-author and the grandson of the third co-author, who, despite his small preschool age, assists us collect beetles.	en	Bekchanov, Norbek Kh., Nabozhenko, Maxim V., Bekchanov, Khudaybergan U., Bekchanova, Mokhira Kh. (2025): A review of the genus Microdera Eschscholtz, 1831 (Coleoptera: Tenebrionidae) of Uzbekistan. Zootaxa 5590 (4): 451-480, DOI: 10.11646/zootaxa.5590.4.1, URL: https://doi.org/10.11646/zootaxa.5590.4.1
03D42163EC29BF6DFF7FFBB8FD1AF53F.taxon	diagnosis	Comparative diagnosis. The new species differs from all known Microdera by various characters, especially the long antennae with 3 (male) (Fig. 1 A) or 2 (female) (Fig. 1 B) antennomeres extending beyond the base of the pronotum. It can be compared with the taxa without vertical plate at the apex of abdominal ventrite 5 and the non-depressed elytra. Externally M. abdullohi sp. nov. is slightly similar to M. convexa (Tauscher, 1812) in the strongly convex pronotum, but differs in the much longer and slender legs (the metatarsi half or less shorter than metatibiae (Fig. 1 A), while in M. convexa metatarsi are shorter than half length of the metatibiae (Fig. 4 A )) and the antennae (not reaching the base of the pronotum in M. convexa (Fig. 4 A )), the shiny body (the elytra also entirely shiny vs opaque elytral lateral sides and the apical part with disappearing puncturation in M. convexa) and the shiny smooth outer (extensor) side of metatibiae with fine sparse punctures (Fig. 1 J) (with dense and coarse confluent puncturation, appear rough at M. convexa (Fig. 4 G )). The new species differs (in addition to the long and slender antennae and legs) from M. campestris and M. minax in the strongly convex pronotum vs slightly convex in the two mentioned species (Fig. 3 C, 13 B).	en	Bekchanov, Norbek Kh., Nabozhenko, Maxim V., Bekchanov, Khudaybergan U., Bekchanova, Mokhira Kh. (2025): A review of the genus Microdera Eschscholtz, 1831 (Coleoptera: Tenebrionidae) of Uzbekistan. Zootaxa 5590 (4): 451-480, DOI: 10.11646/zootaxa.5590.4.1, URL: https://doi.org/10.11646/zootaxa.5590.4.1
03D42163EC29BF6DFF7FFBB8FD1AF53F.taxon	distribution	Distribution. Uzbekistan: Sulton-Uvays Mts.	en	Bekchanov, Norbek Kh., Nabozhenko, Maxim V., Bekchanov, Khudaybergan U., Bekchanova, Mokhira Kh. (2025): A review of the genus Microdera Eschscholtz, 1831 (Coleoptera: Tenebrionidae) of Uzbekistan. Zootaxa 5590 (4): 451-480, DOI: 10.11646/zootaxa.5590.4.1, URL: https://doi.org/10.11646/zootaxa.5590.4.1
03D42163EC2EBF6DFF7FFB50FC1AF64C.taxon	description	(Fig. 3)	en	Bekchanov, Norbek Kh., Nabozhenko, Maxim V., Bekchanov, Khudaybergan U., Bekchanova, Mokhira Kh. (2025): A review of the genus Microdera Eschscholtz, 1831 (Coleoptera: Tenebrionidae) of Uzbekistan. Zootaxa 5590 (4): 451-480, DOI: 10.11646/zootaxa.5590.4.1, URL: https://doi.org/10.11646/zootaxa.5590.4.1
03D42163EC2EBF6DFF7FFB50FC1AF64C.taxon	materials_examined	Type material examined. See in Nabozhenko & Kalashian (2022).	en	Bekchanov, Norbek Kh., Nabozhenko, Maxim V., Bekchanov, Khudaybergan U., Bekchanova, Mokhira Kh. (2025): A review of the genus Microdera Eschscholtz, 1831 (Coleoptera: Tenebrionidae) of Uzbekistan. Zootaxa 5590 (4): 451-480, DOI: 10.11646/zootaxa.5590.4.1, URL: https://doi.org/10.11646/zootaxa.5590.4.1
03D42163EC2EBF6DFF7FFB50FC1AF64C.taxon	discussion	Notes. We didn’t collect this species. Our images are based on the specimens from supralittoral sand of the Aral Sea on the former Barsakelmes Island in Kazakhstan (very close area to the western coast of former Aral). The synonymy was proposed and argued in details by Nabozhenko & Kalashian (2022).	en	Bekchanov, Norbek Kh., Nabozhenko, Maxim V., Bekchanov, Khudaybergan U., Bekchanova, Mokhira Kh. (2025): A review of the genus Microdera Eschscholtz, 1831 (Coleoptera: Tenebrionidae) of Uzbekistan. Zootaxa 5590 (4): 451-480, DOI: 10.11646/zootaxa.5590.4.1, URL: https://doi.org/10.11646/zootaxa.5590.4.1
03D42163EC2EBF6DFF7FFB50FC1AF64C.taxon	distribution	Distribution. European part of Russia: Caspian depression (Nabozhenko & Kalashian 2022), Kazakhstan (from the Caspian Depression across Aral sands to middle reaches of the Syr Darya: Kyzylorda), Uzbekistan: sandy supralittoral zone of the western coast of the Aral Sea (Pirnazarov 1973).	en	Bekchanov, Norbek Kh., Nabozhenko, Maxim V., Bekchanov, Khudaybergan U., Bekchanova, Mokhira Kh. (2025): A review of the genus Microdera Eschscholtz, 1831 (Coleoptera: Tenebrionidae) of Uzbekistan. Zootaxa 5590 (4): 451-480, DOI: 10.11646/zootaxa.5590.4.1, URL: https://doi.org/10.11646/zootaxa.5590.4.1
03D42163EC2DBF61FF7FFAA8FAB4F35B.taxon	description	(Fig. 4)	en	Bekchanov, Norbek Kh., Nabozhenko, Maxim V., Bekchanov, Khudaybergan U., Bekchanova, Mokhira Kh. (2025): A review of the genus Microdera Eschscholtz, 1831 (Coleoptera: Tenebrionidae) of Uzbekistan. Zootaxa 5590 (4): 451-480, DOI: 10.11646/zootaxa.5590.4.1, URL: https://doi.org/10.11646/zootaxa.5590.4.1
03D42163EC2DBF61FF7FFAA8FAB4F35B.taxon	materials_examined	Material examined. 25 specimens (ZIN), Karakalpakstan, Western coast of Aral sea, 12 km E of Komsomolsk-on-Ustyurt (now Qubla-ustyurt), 11. vi. 1972 (G. S. Medvedev); 5 specimens (ZIN), Karakalpakstan, Western coast of Aral sea, 30 km N of Komsomolsk-on-Ustyurt (now Qubla-ustyurt), 30. v. 1978 (G. S. Medvedev); 4 specimens (ZIN), Karakalpakstan, Karateren Lake (43 ° 13 ʹN / 60 ° 23 ʹE), 24. v. 1972 (G. S. Medvedev); 3 specimens (ZIN), Navoiy Region, Kyzylkum desert, Bukantau Mts., N Kulkuduk, 13.05.1965 (E. L. Gur’eva); 5 specimens (ZIN), Navoiy Region, Kyzylkum desert, Bukantau Mts., 6.05.1966 (G. S. Medvedev); 18 specimens (PCNB), 21 specimens (PCMN): Uzbekistan, Karakalpakstan, Kungrad Distr., near the mausoleum “ Daut ata ”, 43 ° 01 ʹ 43 ʺN / 58 ° 43 ʹ 22 ʺE, h = 60 m, 26.04.2024 (M. V. and S. V. Nabozhenko, N. Kh. and Kh. U. Bekchanov, U. E. Duschanov); 14 specimens (PCNB), 10 specimens (PCMN): Uzbekistan, Karakalpakstan, Kungrad Distr., 6.3 km NW “ Barsakelmes ” railway stn., NE coast of Barsakelmes dried salty lake, 43 ° 22 ʹ 45 ʺN / 58 ° 00 ʹ 52 ʺE, h = 40 - 80 m, 27 – 28.04.2024 (M. V. and S. V. Nabozhenko, N. Kh. and Kh. U. Bekchanov, Y. R. Tokhirov). Type material examined. Microdera kaszabi Skopin, 1961 Holotype, ♀ (ZIN): plate with glued abdominal ventrite 5 and inner tergite VII, “ Ср. Или, Илийск, полЫнники на песках 3.05.1958 ” (Sr. Ili, Iliysk, polynniki na peskakh 3.05.1958 N. Skopin leg. (Russian Cyrillic label), now Kazakhstan, middle Ili River, Iliysk [flooded by Kapchagay reservoir in 1970], 43 ° 53 ʹN, 77 ° 07 ʹE, Artemisia on the sands), “ Holotypus Microdera kaszabi N. Skopin, 1961 ”. The holotype of Microdera reitteri was examined previously (Nabozhenko & Kalashian 2022)	en	Bekchanov, Norbek Kh., Nabozhenko, Maxim V., Bekchanov, Khudaybergan U., Bekchanova, Mokhira Kh. (2025): A review of the genus Microdera Eschscholtz, 1831 (Coleoptera: Tenebrionidae) of Uzbekistan. Zootaxa 5590 (4): 451-480, DOI: 10.11646/zootaxa.5590.4.1, URL: https://doi.org/10.11646/zootaxa.5590.4.1
03D42163EC2DBF61FF7FFAA8FAB4F35B.taxon	discussion	Notes. Skopin (1968) noted wide intraspecific variability of M. convexa and identified three subspecies and five varieties of the nomynotypical subspecies: M. convexa convexa (M. convexa typical variety, M. convexa var. fortesulpta Skopin, 1968, M. convexa var. mixta Skopin, 1968, M. covexa var. subglobosa Kraatz, 1865, M. convexa var. laevipeda Skopin, 1968), M. convexa kaszabi Skopin, 1961 and M. convexa globulicollis Ménétriés, 1849. He also provided a table with the frequency of occurrence of these varieties and subspecies in different regions of Kazakhstan. The situation with M. globulicollis is clear: Skopin did not study syntypes of this species and his determinations are incorrect. Erroneous interpretations of the taxa M. campestris and M. globulicollis by different authors, as well as the correct synonymy, are explained in detail in the paper of Nabozhenko & Kalashian (2022). No one accepted the synonymy of M. subglobosa (synonymized by Skopin (1968) after transfer of this taxon to a status of a variety: M. convexa var. subglobosa). The status of this taxon needs to be revised or confirmed after examination of type specimens. The species M. kaszabi Skopin, 1961 was later interpreted as a subspecies of M. convexa (Skopin, 1968), that is reflected in the catalogue (Iwan et al. 2020). We examined the holotype of this taxon (Fig. 5) and again return this subspecies to the status of a species: Microdera kaszabi Skopin, 1961, stat. resurr. This species is distinctly different from M. convexa in many characters: the shiny body, the elytra entirely with coarser distinct puncturation (the puncturation is much finer and sparser and disappears on the opaque lateral sides and the elytral apex in M. convexa); the lateral margins of gena converging to the epistome and the head is shorter in front of the eyes (the genae strongly diverging from eyes to widest portion and then converging to epistome in M. convexa), the metatibiae shiny and smooth on the extensor side, with fine and sparse puncturation (opaque and appear wrinkled due to the merging of coarse, elongated punctures or at least coarsely and densely punctured in M. convexa). Thus, Skopin interpreted the variability very broadly and underestimated some characters, such as the shape of the head and genae, the puncturation and shine of the elytra, and the sculpture of the metatibiae. The most variable characters in M. convexa are the puncturation of the prohypomera and the thickness (and height) of the border of the pronotal base. The puncturation and roughness of the metatibia are also slightly variable, but punctures are always elongated, coarse and dense, often merging. Here we illustrate only Uzbek populations from Ustyurt. The specimens from the former southern Aral coast and Bukantau in Kyzylkum do not differ from the East Ustyurt specimens, only in the population from Bukantau the punctation of the pronotum is slightly finer and sparser.	en	Bekchanov, Norbek Kh., Nabozhenko, Maxim V., Bekchanov, Khudaybergan U., Bekchanova, Mokhira Kh. (2025): A review of the genus Microdera Eschscholtz, 1831 (Coleoptera: Tenebrionidae) of Uzbekistan. Zootaxa 5590 (4): 451-480, DOI: 10.11646/zootaxa.5590.4.1, URL: https://doi.org/10.11646/zootaxa.5590.4.1
03D42163EC2DBF61FF7FFAA8FAB4F35B.taxon	distribution	Distribution. European part of Russia (Caspian depression), Azerbaijan (Apsheron Peninsula) (Nabozhenko & Kalashian 2022); Kazakhstan (western regions, Ustyurt, Mangyshlak, Aral sea region, middle reaches of Syrdarya, Muyunkum desert); Uzbekistan (Ustyurt, Amudarya delta and Aral sea coast, Central Kyzylkum — Bukantau).	en	Bekchanov, Norbek Kh., Nabozhenko, Maxim V., Bekchanov, Khudaybergan U., Bekchanova, Mokhira Kh. (2025): A review of the genus Microdera Eschscholtz, 1831 (Coleoptera: Tenebrionidae) of Uzbekistan. Zootaxa 5590 (4): 451-480, DOI: 10.11646/zootaxa.5590.4.1, URL: https://doi.org/10.11646/zootaxa.5590.4.1
03D42163EC22BF62FF7FFDC0FD72F323.taxon	description	(Fig. 6)	en	Bekchanov, Norbek Kh., Nabozhenko, Maxim V., Bekchanov, Khudaybergan U., Bekchanova, Mokhira Kh. (2025): A review of the genus Microdera Eschscholtz, 1831 (Coleoptera: Tenebrionidae) of Uzbekistan. Zootaxa 5590 (4): 451-480, DOI: 10.11646/zootaxa.5590.4.1, URL: https://doi.org/10.11646/zootaxa.5590.4.1
03D42163EC22BF62FF7FFDC0FD72F323.taxon	materials_examined	Type material examined. Tentyria deplanata (Fig. 6 A, E, F). Lectotype (ZIN), ♂, hereby designated: goldish square, “ Camp. Kirg or. ” (the first two words are printed, the third is handwritten; east of Kirghiz steppes = central and eastern Kazakhstan), “ deplanata Gebl. St. Kirg. or. ” (handwritten), “ Lectotypus Tentyria deplanata Gebler, 1841 des. M. Nabozhenko, 2024 ”. Paralectotype (ZIN), sex unknown: goldish square, “ Camp. Kirg. or. ”. The second paralectotype (as a syntype) from Zoological Museum of Moscow State University was mentioned by Abdurakhmanov & Nabozhenko (2011). References and synonymy were indicated in Nabozhenko & Kalashian (2022). Material examined. 1 ♀ (ZIN): Tashkent, 5. vi. 1915 (T. E. S. = Turkestan Entomological Station) (Fig. 6 B – D).	en	Bekchanov, Norbek Kh., Nabozhenko, Maxim V., Bekchanov, Khudaybergan U., Bekchanova, Mokhira Kh. (2025): A review of the genus Microdera Eschscholtz, 1831 (Coleoptera: Tenebrionidae) of Uzbekistan. Zootaxa 5590 (4): 451-480, DOI: 10.11646/zootaxa.5590.4.1, URL: https://doi.org/10.11646/zootaxa.5590.4.1
03D42163EC22BF62FF7FFDC0FD72F323.taxon	distribution	Distribution. European part of Russia: Caspian depression (Nabozhenko & Kalashian 2022), Kazakhstan: from Caspian Depression across Aral sands to Ili valley and Alakol Lake (Skopin 1961 a), Uzbekistan: Tashkent (new record for the country), China: Ili valley (Skopin 1961 a). The second subspecies Microdera deserta skopini Kaszab, 1966 was described from SE Kazakhstan and it is probably a good species, because distinctly differs from M. deserta s. str. in the shape of the pronotum.	en	Bekchanov, Norbek Kh., Nabozhenko, Maxim V., Bekchanov, Khudaybergan U., Bekchanova, Mokhira Kh. (2025): A review of the genus Microdera Eschscholtz, 1831 (Coleoptera: Tenebrionidae) of Uzbekistan. Zootaxa 5590 (4): 451-480, DOI: 10.11646/zootaxa.5590.4.1, URL: https://doi.org/10.11646/zootaxa.5590.4.1
03D42163EC21BF62FF7FFD10FBDFF7B8.taxon	description	(Figs 7, 8)	en	Bekchanov, Norbek Kh., Nabozhenko, Maxim V., Bekchanov, Khudaybergan U., Bekchanova, Mokhira Kh. (2025): A review of the genus Microdera Eschscholtz, 1831 (Coleoptera: Tenebrionidae) of Uzbekistan. Zootaxa 5590 (4): 451-480, DOI: 10.11646/zootaxa.5590.4.1, URL: https://doi.org/10.11646/zootaxa.5590.4.1
03D42163EC21BF62FF7FFD10FBDFF7B8.taxon	materials_examined	Type material examined. Holotype (NHMB), ♂: “ Kokand Fergana ”, “ Sammlung Adr. Schuster ”, “ Staud. ”, “ Museum Frey München ”, “ HOLOTYPUS Microdera ferghanensis m. det. Dr. Kaszab, 1957 ”. 3 paratypes with the same label but “ Paratypus ... ” are also deposited in NHMB. The paratype from HNHM was indicated by Nabozhenko & Kalashian (2022). Material examined. 12 specimens (PCNB), 10 specimens (PCMN, ZIN): Uzbekistan, Fergana Region, Sho‘rsuv, 40 ° 17 ʹ 09 ʺN / 70 ° 48 ʹ 22 ʺE, h = 630 m, 16.04.2024 (M. V. and S. V. Nabozhenko, N. Kh. and Kh. U. Bekchanov); 3 specimens (PCNB), 2 specimens (PCMN): Uzbekistan, Fergana Region, O‘qchi env., backward of garden, 40 ° 19 ʹ 12 ʺN / 70 ° 57 ʹ 59 ʺE, h = 680 m, 16.04.2024 (M. V. and S. V. Nabozhenko, N. Kh. and Kh. U. Bekchanov). Notes. The species was unfairly synonymized with M. globulicollis by Nabozhenko & Kalashian (2022). Their opinion was based on the examination of one paratype from HNHM. This paratype has the least expressed species characteristics, such as very fine puncturation of the prothoracic hypomera only near coxae, the puncturation of head basally is slightly elongated and with almost the same density as on the other surface of frons. The authors attributed this to the variability of M. globulicollis, which also occurs in Fergana valley. After examination of the holotype (Fig. 7 A – C) and a series of beetles from the Qampirqoq Mountains (S of Kokand, Fergana valley) (Fig. 7 D – F, 8), the independence of this species is confirmed. Microdera ferghanensis is clearly distinguished from M. globulicollis by the presence of the puncturation on the entire surface of the hypomera or at least near the procoxal area (Fig. 7 F), slightly elongated puncturation of the pronotum on sides (Fig. 7 A, E), denser and elongated puncturation at the base of the head (Fig. 7 E), as well as the angularly rounded mesoventrite between mesocoxae (Fig. 7 B, 8 A) (not evenly, as in M. globulicollis (Fig. 10 A )). The tarsi of M. ferghanensis (Fig. 8 F – H) are thinner than those of M. globulicollis (Fig. 10 B, C). Both species also differ in the shape of the aedeagus (compare Figs 8 I and 10 E). Thus, we resurrect this species: Microdera ferghanensis Kaszab, 1957, sp. resurr. The name of the species was misspelled as M. ferganensis by Skopin (1968), Iwan et al. (2020) and Nabozhenko & Kalashian (2022).	en	Bekchanov, Norbek Kh., Nabozhenko, Maxim V., Bekchanov, Khudaybergan U., Bekchanova, Mokhira Kh. (2025): A review of the genus Microdera Eschscholtz, 1831 (Coleoptera: Tenebrionidae) of Uzbekistan. Zootaxa 5590 (4): 451-480, DOI: 10.11646/zootaxa.5590.4.1, URL: https://doi.org/10.11646/zootaxa.5590.4.1
03D42163EC21BF62FF7FFD10FBDFF7B8.taxon	distribution	Distribution. Uzbekistan (SW areas of Fergana Region: Qampirqoq Mts). Skopin (1968) recorded this species for central and southern Kazakhstan from Arys Lake to Syr-Darya valley and Muyunkum desert. We didn’t find any Skopin’s specimens of M. ferghanensis in ZIN; these records need verification.	en	Bekchanov, Norbek Kh., Nabozhenko, Maxim V., Bekchanov, Khudaybergan U., Bekchanova, Mokhira Kh. (2025): A review of the genus Microdera Eschscholtz, 1831 (Coleoptera: Tenebrionidae) of Uzbekistan. Zootaxa 5590 (4): 451-480, DOI: 10.11646/zootaxa.5590.4.1, URL: https://doi.org/10.11646/zootaxa.5590.4.1
03D42163EC26BF67FF7FFF35FBFFF553.taxon	description	(Figs 9, 10)	en	Bekchanov, Norbek Kh., Nabozhenko, Maxim V., Bekchanov, Khudaybergan U., Bekchanova, Mokhira Kh. (2025): A review of the genus Microdera Eschscholtz, 1831 (Coleoptera: Tenebrionidae) of Uzbekistan. Zootaxa 5590 (4): 451-480, DOI: 10.11646/zootaxa.5590.4.1, URL: https://doi.org/10.11646/zootaxa.5590.4.1
03D42163EC26BF67FF7FFF35FBFFF553.taxon	materials_examined	Type material examined. Type material for M. globulicollis and M. margelanica see in Nabozhenko & Kalashian (2022). Microdera extabesa (Fig. 9 A, B): Holotype, ♂ (ZIN): “ Yu. Kizil-Kum, Ayak-Agitma 6.08.1936 N. Skopin leg. ” (Russian Cyrillic; now: Uzbekistan, Bukhara Region, 40 ° 39 ʹN, 64 ° 29 ʹE), “ ♂ ” (square), “ Holotypus Microdera extabesa N. Skopin, 1960 ”. Paratypes. 11 specimens (ZIN): “ Golodnaya step Mussa-Kuduk 25.06.1959 leg. A. Andreeva Coll. N. Skopin ” (Russian Cyrillic), “ Paratypus Microdera extabesa N. Skopin, 1960 ”; 1 ♂ (ZIN): “ Termez 22.09.1938 N. Skopin leg. ” (Russian Cyrillic), “ Paratypus Microdera extabesa N. Skopin, 1960 ”; 5 specimens (ZIN): “ Yu Kyzyl-Kum, Ayak-Agitma, 6.08.1936 N. Skopin leg. ” (Cyrillic Russian label), “ Paratypus Microdera extabesa N. Skopin, 1960 ”. Material examined. 4 specimens (ZIN): Navoiy Region, Kyzylkum desert, Ayakguzhumdy, 21. iii. 1961, 13. v. 1961 (B. M. Mamaev); 1 specimen (ZIN): Kuldzhuktau, Ayakguzhumdy, 2. iv. 1966 (G. S. Medvedev); 1 specimen (ZIN): Navoiy Region, 15 km SSW Tamdy (= Tamdybulak), 2. iv. 1966 (G. S. Medvedev); 2 specimens (ZIN): Navoiy Region, Aznek, 70 km W Tamdy (= Tamdybulak), 1. v. 1965 (L. V. Arnoldi); 2 specimens (ZIN): Navoiy Region, Kyzylkum, Auminzatau, 10.5.1966 (G. S. Medvedev); 2 specimens (ZIN): Navoiy Region, Kyzylkum, sands 45 km NW Dzhing. [Dzhingildy], 13.06.1965 (L. V. Arnoldi); 2 specimens, 40 km W Bukhara, Kyzylkum, 15. v. 1966 (G. S. Medvedev); 2 specimens (ZIN): SO Kyzylkum, on clayey soil, 11. vi. 1960 (P. Ler); 1 specimen (ZIN): Bukhara Region, Karaulbazar, 10. iv. 1927 (V. V. Yakhontov); 15 specimens (ZIN): Tashkent Region, Bekabad, 17. v. 1966 (N. G. Skopin); 1 ♂ (ZIN): Uzbekistan, Surkhandarya Region, 7 km E Aktash, 37 ° 31 ʹ 57 ʺN / 66 ° 44 ʹ 09 ʺE, h = 450 m, 13.04.2024 (M. V. and S. V. Nabozhenko, N. Kh. and Kh. U. Bekchanov, D. A. Yavkachev).	en	Bekchanov, Norbek Kh., Nabozhenko, Maxim V., Bekchanov, Khudaybergan U., Bekchanova, Mokhira Kh. (2025): A review of the genus Microdera Eschscholtz, 1831 (Coleoptera: Tenebrionidae) of Uzbekistan. Zootaxa 5590 (4): 451-480, DOI: 10.11646/zootaxa.5590.4.1, URL: https://doi.org/10.11646/zootaxa.5590.4.1
03D42163EC26BF67FF7FFF35FBFFF553.taxon	discussion	Notes. We examined all type series of M. extabesa and didn’t find any differences with M. globulicollis. Skopin mentioned some differences in the shape of “ thorax ” in M. extabesa and M. globulicollis and more rounded postero-lateral corners of pronotum in M. extabesa. We did not find these differences. The holotype of M. extabesa completely corresponds to the lectotype of M. globulicollis from Turkmenistan. So the following new synonymy is established: Microdera globulicollis Ménétriés, 1849 = Microdera extabesa Skopin, 1961: 393, syn. nov. We don’t know, which species mentioned Pirnazarov (1973) under the name “ Microdera globulicollis ” for Karakalpakstan (Uzbekistan).	en	Bekchanov, Norbek Kh., Nabozhenko, Maxim V., Bekchanov, Khudaybergan U., Bekchanova, Mokhira Kh. (2025): A review of the genus Microdera Eschscholtz, 1831 (Coleoptera: Tenebrionidae) of Uzbekistan. Zootaxa 5590 (4): 451-480, DOI: 10.11646/zootaxa.5590.4.1, URL: https://doi.org/10.11646/zootaxa.5590.4.1
03D42163EC26BF67FF7FFF35FBFFF553.taxon	distribution	Distribution. Turkmenistan (Mashhad-i Misrian, foothills of Kopetdag, south Turkmenistan, Badkhyz, Kugitangtau), Uzbekistan (Kyzylkum desert, Kattakum desert, Golodnaya (Hungry) steppe, Fergana valley), Kazakhstan (Eastern Kyzylkum, Josaly, Yanykurgan, Turkestan) (ZIN collection).	en	Bekchanov, Norbek Kh., Nabozhenko, Maxim V., Bekchanov, Khudaybergan U., Bekchanova, Mokhira Kh. (2025): A review of the genus Microdera Eschscholtz, 1831 (Coleoptera: Tenebrionidae) of Uzbekistan. Zootaxa 5590 (4): 451-480, DOI: 10.11646/zootaxa.5590.4.1, URL: https://doi.org/10.11646/zootaxa.5590.4.1
03D42163EC24BF67FF7FFBD4FEFAF71D.taxon	description	(Figs 11, 12)	en	Bekchanov, Norbek Kh., Nabozhenko, Maxim V., Bekchanov, Khudaybergan U., Bekchanova, Mokhira Kh. (2025): A review of the genus Microdera Eschscholtz, 1831 (Coleoptera: Tenebrionidae) of Uzbekistan. Zootaxa 5590 (4): 451-480, DOI: 10.11646/zootaxa.5590.4.1, URL: https://doi.org/10.11646/zootaxa.5590.4.1
03D42163EC24BF67FF7FFBD4FEFAF71D.taxon	materials_examined	Material examined. 1 specimen (ZIN): Namangan env., 1903 (I. Yankovsky); 1 specimen (ZIN): Fergana region, sands N Yaz’yavan, 6. v. 1961 (V. A. Zaslavsky); 5 specimens (PCNB), 5 specimens (PCMN, ZIN): Uzbekistan, Fergana Region, Yozyovon sands, 40 ° 36 ʹ 20 ʺN / 71 ° 31 ʹ 41 ʺE, h = 420 m, 17.04.2024 (M. V. and S. V. Nabozhenko, N. Kh. and Kh. U. Bekchanov); 11 specimens (PCNB), 8 specimens (PCMN, ZIN): Uzbekistan, Fergana Region, Kokand sands, 40 ° 38 ʹ 35 ʺN / 70 ° 52 ʹ 04 ʺE, h = 360 m, 15.04.2024 (M. V. and S. V. Nabozhenko, N. Kh. and Kh. U. Bekchanov).	en	Bekchanov, Norbek Kh., Nabozhenko, Maxim V., Bekchanov, Khudaybergan U., Bekchanova, Mokhira Kh. (2025): A review of the genus Microdera Eschscholtz, 1831 (Coleoptera: Tenebrionidae) of Uzbekistan. Zootaxa 5590 (4): 451-480, DOI: 10.11646/zootaxa.5590.4.1, URL: https://doi.org/10.11646/zootaxa.5590.4.1
03D42163EC24BF67FF7FFBD4FEFAF71D.taxon	distribution	Distribution. S Kazakhstan (Skopin 1968) and Tajikistan (middle reaches of the Syr Darya), Uzbekistan (Fergana valley: west and centre) (Kaszab 1966, ZIN collection). Erroneously recorded for Turkmenistan by Iwan et al. (2020).	en	Bekchanov, Norbek Kh., Nabozhenko, Maxim V., Bekchanov, Khudaybergan U., Bekchanova, Mokhira Kh. (2025): A review of the genus Microdera Eschscholtz, 1831 (Coleoptera: Tenebrionidae) of Uzbekistan. Zootaxa 5590 (4): 451-480, DOI: 10.11646/zootaxa.5590.4.1, URL: https://doi.org/10.11646/zootaxa.5590.4.1
03D42163EC3ABF7AFF7FFF35FB51F6BA.taxon	description	(Figs 13, 14)	en	Bekchanov, Norbek Kh., Nabozhenko, Maxim V., Bekchanov, Khudaybergan U., Bekchanova, Mokhira Kh. (2025): A review of the genus Microdera Eschscholtz, 1831 (Coleoptera: Tenebrionidae) of Uzbekistan. Zootaxa 5590 (4): 451-480, DOI: 10.11646/zootaxa.5590.4.1, URL: https://doi.org/10.11646/zootaxa.5590.4.1
03D42163EC3ABF7AFF7FFF35FB51F6BA.taxon	materials_examined	Material examined. 8 specimens (ZIN): Karakalpakstan, Kungrad District, Kyzylzhar, 9 – 10. v. 1939 (P. A. Veltishchev); 1 specimen (ZIN): Karakalpakstan, Amudarya delta, Kubetau Mt., 17. vi. 1911 (K. A. Molchanov); 1 specimen (ZIN): Karakalpakstan, Amudarya delta, vii (without year) (Kazakov); 1 specimen (ZIN): Karakalpakstan, 70 km SO Nukus, 24. viii. 1968 (B. P. Pirnazarov); 4 specimens (ZIN): Karakalpakstan, 30 km S Takhta-Kupyr, 7. ix. 1968 (B. P. Pirnazarov); 1 specimen (ZIN): Karakalpakstan, 1 km S Nukus, 24. v. 1972 (G. S. Medvedev); 1 specimen (ZIN): Khorezm Region, Khiva, 22.03.1927 (L. Zimin); 6 specimens (ZIN): Khorezm Region, Khiva District, Ravat, 21. iv. 1927 (V. Gussakovsky); 6 specimens (ZIN): the same locality, 27. iv. 1927, 2. ix. 1927 (L. Zimin); 1 specimen (ZIN): Khorezm Region, Khiva District, Daniyar, 4. iv. 1927 (V. Gussakovsky); 3 specimens (ZIN): Khorezm Region, 6 km SSW Khiva, 31. v. 1972 (G. S. Medvedev); 1 specimen (ZIN): Khorezm Region, SO Meshekli, Kulatau tugay, Amudarya, 10. iv. 1966 (G. S. Medvedev); 1 specimen (ZIN): Bukhara Region, Karabulak, 14. v. 1911 (A. K. Golbek); 1 specimen (ZIN): 90 – 100 km W Gazli, 27. v. 1965 (L. V. Arnoldi); 3 specimens (ZIN): Bukhara Region, 80 km NW Gazli, 12. iv. 1966 (G. S. Medvedev); 18 specimens (ZIN): Navoiy Region, Kuldzhuktau, 20 km S Ayakguzhumdy, 5. iv. 1966 (G. S. Medvedev); 1 specimens (ZIN): Navoy Region, Tamdytau, 8 km SO from Aktau Mt., 4. v. 1965 (E. L. Gur’eva); 2 specimens (ZIN): Navoy Region, Tamdybulak, 7. v. 1965 (L. V. Arnoldi); 3 specimens (PCNB), 8 specimens (PCMN): Uzbekistan, Bukhara Region, SW coast of Dengizkul lake, 39 ° 10 ʹ 16 ʺN / 64 ° 01 ʹ 46 ʺE, h = 180 m, 11 – 12.04.2024 (M. V. and S. V. Nabozhenko, N. Kh. and Kh. U. Bekchanov, D. A. Yavkachev); 4 specimens (PCNB), 3 specimens (PCMN): Uzbekistan, Karakalpakistan, Kungrad Distr., Kyzylzhar env., 43 ° 06 ʹ 58 ʺN / 59 ° 03 ʹ 39 ʺE, h = 70 m and 43 ° 07 ʹ 51 ʺN / 59 ° 05 ʹ 44 ʺE, h = 80 – 100 m, 25.04.2024 (M. V. and S. V. Nabozhenko, N. Kh. and Kh. U. Bekchanov, U. E. Duschanov); 9 specimens (PCNB): Uzbekistan, Karakalpakistan, Beruniy Distr., 41 ° 53 ʹ 60 ʺN / 60 ° 39 ʹ 13.1 ʺE, 19.03.2024 (N. Kh. Bekchanov, Kh. U. Bekchanov and M. Kh. Bekchanova).	en	Bekchanov, Norbek Kh., Nabozhenko, Maxim V., Bekchanov, Khudaybergan U., Bekchanova, Mokhira Kh. (2025): A review of the genus Microdera Eschscholtz, 1831 (Coleoptera: Tenebrionidae) of Uzbekistan. Zootaxa 5590 (4): 451-480, DOI: 10.11646/zootaxa.5590.4.1, URL: https://doi.org/10.11646/zootaxa.5590.4.1
03D42163EC3ABF7AFF7FFF35FB51F6BA.taxon	description	Variability. Disc of the pronotum can be with elongate puncturation on sides.	en	Bekchanov, Norbek Kh., Nabozhenko, Maxim V., Bekchanov, Khudaybergan U., Bekchanova, Mokhira Kh. (2025): A review of the genus Microdera Eschscholtz, 1831 (Coleoptera: Tenebrionidae) of Uzbekistan. Zootaxa 5590 (4): 451-480, DOI: 10.11646/zootaxa.5590.4.1, URL: https://doi.org/10.11646/zootaxa.5590.4.1
03D42163EC3ABF7AFF7FFF35FB51F6BA.taxon	distribution	Distribution. Turkmenistan (Karakum desert), Uzbekistan (Karakum and Kyzylkum deserts, sands of Amudarya delta) (Medvedev & Nepesova 1985 a; Bekchanov et al., 2023; ZIN collection), NW Afghanistan (Kushka) (Gridelli 1954, Kaszab 1959, 1966, ZIN collection), Kazakhstan (Kyzylkum, Muyunkum, Ili River) (ZIN collection). The record of this species for Tien Shan (Kyrgyzstan: Naryn) (Gridelli 1954) is very doubtful.	en	Bekchanov, Norbek Kh., Nabozhenko, Maxim V., Bekchanov, Khudaybergan U., Bekchanova, Mokhira Kh. (2025): A review of the genus Microdera Eschscholtz, 1831 (Coleoptera: Tenebrionidae) of Uzbekistan. Zootaxa 5590 (4): 451-480, DOI: 10.11646/zootaxa.5590.4.1, URL: https://doi.org/10.11646/zootaxa.5590.4.1
03D42163EC38BF7CFF7FFF35FB79F77F.taxon	description	(Figs 15, 16)	en	Bekchanov, Norbek Kh., Nabozhenko, Maxim V., Bekchanov, Khudaybergan U., Bekchanova, Mokhira Kh. (2025): A review of the genus Microdera Eschscholtz, 1831 (Coleoptera: Tenebrionidae) of Uzbekistan. Zootaxa 5590 (4): 451-480, DOI: 10.11646/zootaxa.5590.4.1, URL: https://doi.org/10.11646/zootaxa.5590.4.1
03D42163EC38BF7CFF7FFF35FB79F77F.taxon	materials_examined	Type material examined. Lectotype (Fig. 15 A, B), ♂ (ZIN) hereby designated: “ Пархар на р. ПЯндж. Таджикист. Луппова 9. VIII. 934 ” (Russian Cyryllic: Parkhar na r. Pyandzh. Tadzhikist. Luppova 9. VIII. 934 = Tajikistan, Parkhar on Pyanj River), “ LECTOTYPUS Microdera semenoviana Bogatchev, 1947 des. M. Nabozhenko et N. Bekchanov ”. Paralectotypes: 1 specimen (ZIN): “ Parkhar na r. Pyandzh. Tadzhikist. Luppova 9. VIII. 934 ” (Russian Cyrillic), “ PARALECTOTYPUS Microdera semenoviana Bogatchev, 1947 det. M. Nabozhenko ”; 1 specimen (ZIN): the same locality, but 10. viii. 1934 and the same type label; 1 specimen (ZIN): “ КЫЗЫл-Кала, р. Вахш. Тадж. 9. VI. 934 Гуссаковский ” (Russian Cyrillic: Kyzyl-Kala, r. Vakhsh. Tadzh. 9. VI. 1934 Gussakovsky = Tajikistan, Vakhsh River, Kyzyl-Kala) and the same type label; 1 specimen (ZIN): “ Кабадиан, Таджикист. 10. VII. 934 Гуссаковский ” (Russian Cyrillic: Kabadian, Tadzhikist. 10. VII. 1934, Gussakovsky = Tajikistan, Qubodiyon) and the same type label. Material examined. 2 specimens (ZIN collection): Uzbekistan / Tajikistan, S Babatag, between Dzharkurgan [Uzbekistan] and Shaartuz [Tajikistan], 19. iv. 1962 (E. L. Gur’eva).	en	Bekchanov, Norbek Kh., Nabozhenko, Maxim V., Bekchanov, Khudaybergan U., Bekchanova, Mokhira Kh. (2025): A review of the genus Microdera Eschscholtz, 1831 (Coleoptera: Tenebrionidae) of Uzbekistan. Zootaxa 5590 (4): 451-480, DOI: 10.11646/zootaxa.5590.4.1, URL: https://doi.org/10.11646/zootaxa.5590.4.1
03D42163EC38BF7CFF7FFF35FB79F77F.taxon	distribution	Distribution. SE Turkmenistan (Kugitangtau Mts: Garlyk), Uzbekistan (Babatag Mts), SW Tajikistan, Afghanistan (Bogatchev 1946, Kaszab 1968, Medvedev & Nepesova 1985 a, ZIN collection).	en	Bekchanov, Norbek Kh., Nabozhenko, Maxim V., Bekchanov, Khudaybergan U., Bekchanova, Mokhira Kh. (2025): A review of the genus Microdera Eschscholtz, 1831 (Coleoptera: Tenebrionidae) of Uzbekistan. Zootaxa 5590 (4): 451-480, DOI: 10.11646/zootaxa.5590.4.1, URL: https://doi.org/10.11646/zootaxa.5590.4.1
03D42163EC3FBF7EFF7FF92DFDC4F587.taxon	description	(Fig. 17)	en	Bekchanov, Norbek Kh., Nabozhenko, Maxim V., Bekchanov, Khudaybergan U., Bekchanova, Mokhira Kh. (2025): A review of the genus Microdera Eschscholtz, 1831 (Coleoptera: Tenebrionidae) of Uzbekistan. Zootaxa 5590 (4): 451-480, DOI: 10.11646/zootaxa.5590.4.1, URL: https://doi.org/10.11646/zootaxa.5590.4.1
03D42163EC3FBF7EFF7FF92DFDC4F587.taxon	materials_examined	Type material examined. Holotype (Fig. 17 H, I), ♂ (ZIN): “ ЮЗ Турк. Мешед-Мессериан Г. Медведев 12 М 984 ” (YuZ Turk. Meshed-Messerian G. Medvedev 12. V 984) (Russian Cyrillic label (now SW Turkmenistan, Mashhadi Misrian (= Dehistan), 38 ° 16 ′ 12 ″ N, 54 ° 37 ′ 28 ″ E). Paratypes. 21 paratypes (ZIN): “ Shasenem, sev. Karakumy Kir’yakova 7 / VI 953 ” (Russian Cyrillic label) (N Turkmenistan, Shasenem railway station, 41 ° 39 ʹN, 59 ° 23 ʹE), “ na svet ” (at light; Russian Cyrillic label), “ Paratypus Microdera shasenema sp. nov. Medv. et Nepes. ” (the same label for all paratypes); 6 paratypes (ZIN): “ YuZ Turk. Meshed-Messerian G. Medvedev 12. V 984 ” (Russian Cyrillic label) (SW Turkmenistan, Mashhad-i Misrian); 2 paratypes (ZIN): “ Krasnovodsk Zakasp. obl. 28. III. 916 B. Il’in ” (Russian Cyrillic label) (now Turkmenistan, Turkmenbashi); 8 paratypes (ZIN): 5 km Z Khodzheyli G. Medvedev 4 VI 972 ” (Russian Cyrillic label (now in Uzbekistan, Karakalpakstan); 1 paratypes (ZIN): “ N Shakhi-burun B. Balkhany Turkm. V. Popov 16 VI 934 ” (Russian Cyrillic label); 6 paratypes (ZIN): “ Beltau, oz. Karateren G. Medvedev 24. V 972 ” (Russian Cyrillic label); 2 paratypes (ZIN): “ Zap. ber. Aral m. 12 km V Komsomol’ska G. Medvedev II VI 972 ” (Russian Cyrillic label) (western coast of the Aral sea, 12 km W of Komsomolsk-on-Ustyurt = Qubla ustyurt); 2 paratypes (ZIN): “ Karakalpak 40 km W Nukus 14 - VII- 1966 Pirnazarov ” (Russian Cyrillic label); 1 paratype (ZIN): “ Karakalpak Nukus 20 - V- 1969 Pirnazarov ” (Russian Cyrillic label), “ N 21 ”; 1 paratype (ZIN) “ fl. Amudarya ”; 6 paratypes (ZIN): Karakalpakstan, Beltau, Karateren Lake, 24. v. 1972 (G. S. Medvedev) (Russian Cyrillic label).	en	Bekchanov, Norbek Kh., Nabozhenko, Maxim V., Bekchanov, Khudaybergan U., Bekchanova, Mokhira Kh. (2025): A review of the genus Microdera Eschscholtz, 1831 (Coleoptera: Tenebrionidae) of Uzbekistan. Zootaxa 5590 (4): 451-480, DOI: 10.11646/zootaxa.5590.4.1, URL: https://doi.org/10.11646/zootaxa.5590.4.1
03D42163EC3FBF7EFF7FF92DFDC4F587.taxon	discussion	Notes. Specimens from the Amudarya delta (Fig. 17 A – G, J – L), the coast of the former Aral Sea and NE Karakum (Turkmenistan: Shasenem) differ from those in SW Turkmenistan, having more slender body, less transverse pronotum, more elongate tarsi, lengths of metatarsomeres 1 and 4 are subequal (Fig. 17 D) (metatarsomere 1 distinctly shorter than 4 in populations from SW Turkmenistan (Fig. 17 I )). The mentioned characters are slightly variable in the population from Uzbekistan and northern Karakum (Turkmenistan: Shasenem). The population from SW Tajikistan more similar to those from SW Turkmenistan, but beetles also slightly differ from other two populations. Medvedev in the original description used characters of the north populations for proportions of metatarsi, while the holotype and specimens from the type locality have other proportions. The status of three disjunctive populations of M. shasenema can be established after examination of a larger number of specimens, especially from the area of the middle reaches of Amudarya (in case the species occurs there). We use characters of the north population (from Uzbekistan and NE Turkmenistan) in a key below.	en	Bekchanov, Norbek Kh., Nabozhenko, Maxim V., Bekchanov, Khudaybergan U., Bekchanova, Mokhira Kh. (2025): A review of the genus Microdera Eschscholtz, 1831 (Coleoptera: Tenebrionidae) of Uzbekistan. Zootaxa 5590 (4): 451-480, DOI: 10.11646/zootaxa.5590.4.1, URL: https://doi.org/10.11646/zootaxa.5590.4.1
03D42163EC3FBF7EFF7FF92DFDC4F587.taxon	distribution	Distribution. Turkmenistan (Karakum desert), Uzbekistan (Karakalpakstan, lower reaches of Amudarya, Aral coast) (Medvedev & Nepesova 1985 b; ZIN collection), Tadjikistan (valley of Kofarnihon, tributary of Amudarya river; new record for the country).	en	Bekchanov, Norbek Kh., Nabozhenko, Maxim V., Bekchanov, Khudaybergan U., Bekchanova, Mokhira Kh. (2025): A review of the genus Microdera Eschscholtz, 1831 (Coleoptera: Tenebrionidae) of Uzbekistan. Zootaxa 5590 (4): 451-480, DOI: 10.11646/zootaxa.5590.4.1, URL: https://doi.org/10.11646/zootaxa.5590.4.1
03D42163EC3DBF7EFF7FFAB4FD50F6F1.taxon	description	(Fig. 18)	en	Bekchanov, Norbek Kh., Nabozhenko, Maxim V., Bekchanov, Khudaybergan U., Bekchanova, Mokhira Kh. (2025): A review of the genus Microdera Eschscholtz, 1831 (Coleoptera: Tenebrionidae) of Uzbekistan. Zootaxa 5590 (4): 451-480, DOI: 10.11646/zootaxa.5590.4.1, URL: https://doi.org/10.11646/zootaxa.5590.4.1
03D42163EC3DBF7EFF7FFAB4FD50F6F1.taxon	materials_examined	Type material examined. Syntype, sex unknown (ZIN): goldish circle, “ Tekke Radde Reitter ” (handwritten), “ к. Г. Сиверса ” (Russian Cyrillic; collection of G. I. Sivers), “ Microdera transversicollis n. sp. ” (old label, handwritten). Gridelli indicated another syntype of this species deposited in Natural History Museum Budapest with labels: “ Turcmenien. Leder. Reitter ”. Material examined. 2 specimens (ZIN): Bukhara Region, Kyzylkum, 40 km W Bukhara, 15. v. 1966 (G. S. Medvedev); 1 specimen (ZIN): “ Tashk. U … .. V. 4.12 ” (Tashkent District, 4. v. 1912); 5 specimens (ZIN): S Kyzylkum, Basilbek, 11. vi. 1936 (N. G. Skopin); 6 specimens (ZIN): Navoyi Region, NW Bukantau Mts., S Kyzylkum, Tortkuduk, 28. vii. 1936 (N. G. Skopin); 1 specimen (ZIN): Navoiy Region, Susyz-Kara-Tau (= Kuldzhuktau Mts.), 20. vi. 1936 (N. G. Skopin); 1 specimen (ZIN): Navoyi Region, Soz-batyr, 19. vi. 1936 (N. G. Skopin); 1 specimen (ZIN): Navoyi Region, Dzhetymtau, 20. vi. 1936 (N. G. Skopin); 4 specimens (ZIN): Navoyi Region, Manamdzhan, 10. vi. 1936 (N. G. Skopin).	en	Bekchanov, Norbek Kh., Nabozhenko, Maxim V., Bekchanov, Khudaybergan U., Bekchanova, Mokhira Kh. (2025): A review of the genus Microdera Eschscholtz, 1831 (Coleoptera: Tenebrionidae) of Uzbekistan. Zootaxa 5590 (4): 451-480, DOI: 10.11646/zootaxa.5590.4.1, URL: https://doi.org/10.11646/zootaxa.5590.4.1
03D42163EC3DBF7EFF7FFAB4FD50F6F1.taxon	distribution	Distribution. Kazakhstan (Kyzylkum desert), Turkmenistan (Karakum desert), Uzbekistan (Kyzylkum desert) (ZIN collection), Afghanistan (ZIN collection; Gridelli 1954, Kaszab 1959, 1966). The species was erroneously recorded for Armenia by Iwan et al. (2020).	en	Bekchanov, Norbek Kh., Nabozhenko, Maxim V., Bekchanov, Khudaybergan U., Bekchanova, Mokhira Kh. (2025): A review of the genus Microdera Eschscholtz, 1831 (Coleoptera: Tenebrionidae) of Uzbekistan. Zootaxa 5590 (4): 451-480, DOI: 10.11646/zootaxa.5590.4.1, URL: https://doi.org/10.11646/zootaxa.5590.4.1
03D42163EC33BF71FF7FFF35FC33F60F.taxon	description	(Figs 19, 20)	en	Bekchanov, Norbek Kh., Nabozhenko, Maxim V., Bekchanov, Khudaybergan U., Bekchanova, Mokhira Kh. (2025): A review of the genus Microdera Eschscholtz, 1831 (Coleoptera: Tenebrionidae) of Uzbekistan. Zootaxa 5590 (4): 451-480, DOI: 10.11646/zootaxa.5590.4.1, URL: https://doi.org/10.11646/zootaxa.5590.4.1
03D42163EC33BF71FF7FFF35FC33F60F.taxon	materials_examined	Type material (ZIN). Holotype, ♂: « АЯкгужумдЫ Кульджуктау Г. Медведев 1. IV. 966 » (Uzbekistan, Bukhara / Navoi regions, Kuldzhuktau Mts, Ayakguzhumdy, 40 ° 45 ʹ 1 ʺN / 63 ° 45 ʹ 45 ʺE, 1. iv. 1966, leg. G. S. Medvedev). Paratypes. 20 specimens (unsexed): the same locality and collector, dates from 1 to 5. iii. 1966; 4 specimens: Navoi / Bukhara regions, Kuldzhuktau, Mts., 15.03.1961 (leg. B. Mamaev); 4 specimens: the same locality and collector, but 17.03.1961; 1 specimen: the same locality, 12. v. 1966 (leg. Pastukhov); 1 ♂, 2 specimens (unsexed): Navoi Region, Kudzhuktau Mts, Sultanbibi draw-well, 24. iii. 1961 (leg. B. Mamaev, collection of N. Skopin); Bukhara Region, Zhamansay natural boundary, 140 km NW Shafirkan (20 – 25 km NE Ayakguzhumdy), 27. x. 1969 (leg. M. Falkovitsh).	en	Bekchanov, Norbek Kh., Nabozhenko, Maxim V., Bekchanov, Khudaybergan U., Bekchanova, Mokhira Kh. (2025): A review of the genus Microdera Eschscholtz, 1831 (Coleoptera: Tenebrionidae) of Uzbekistan. Zootaxa 5590 (4): 451-480, DOI: 10.11646/zootaxa.5590.4.1, URL: https://doi.org/10.11646/zootaxa.5590.4.1
03D42163EC33BF71FF7FFF35FC33F60F.taxon	description	Description. Male. Body moderately slender, convex, opaque, black or dark-brown, legs brown to black. Measurements: PH w — 1.38, P 1 P w — 1.13, E l E w — 1.5 – 1.52, EH w — 1.8 – 1.83, EP w — 1.34 – 1.36, EP l — 2.24 – 2.25. Body length 7.5 – 10 mm, width 2.5 mm Head (Fig. 19 D) dorsally widest at eye level, where 1.03 times as wide as at genal level and 1.35 times as wide as interocular space. Eyes comparatively large (in length dorsally), moderately convex. Temple not protruding and not forms angle with posterior edge of eye (lateral edge of head not angular at level of temple and eye), visual area of eye lateral. Supraorbital keels slightly sinuous, long. Prong of anterior margin of epistome slightly asymmetric and slightly projected, apex narrowly rounded; margins of epistome on sides of prong emarginated (left weak, right stronger) (Fig. 20 A). Head with regular fine and sparse puncturation of round punctures, interpuncture space 1.5 – 2 times as long as puncture diameter. Puncturation of head ventrally very sparse and very fine, epicranium at middle almost without punctures. Antennae moderately long, reaching base of pronotum. Antennomere 3 longest, 1.7 times as long as 2 nd antennomere and 1.5 times as long as 4 th antennomere; antennomeres 1 and 9 widest (1.2 times as long as wide for the latter). Five proximal antennomeres coarser and denser punctured that rest. Prothorax (Fig. 19 D – F). Pronotum slightly transverse, widest slightly ahead of middle. Anterior margin almost straight, lateral margins slightly rounded in basal half and strongly rounded in apical half. Base strongly evenly rounded. Anterior margin with widely interrupted in middle sulcus, lateral margins finally margined, base widely margined. Antero-lateral corners widely rounded; postero-lateral corners obtuse. Disc of pronotum strongly convex, flattened along base in middle (Fig. 19 A – B), so disc outline in lateral view emarginate in the basal portion (Fig. 19 F). Puncturation of disc fine and sparse, often finer and sparser than on head; puncturation in basal portion coarser than on other surface. Prosternum very finely and sparsely punctured, smoothly wrinkled on sides. Prothoracic hypomera sparsely punctured only near procoxae, other surface without punctures, only with very small and short setae (Fig. 19 E). Prosternal process declivious rounded. Pterothorax (Fig. 19 A – C). Elytra oval, not elongated apically. Surface of elytra slightly shiny in middle and opaque at apex and on lateral sides. Puncturation very fine and sparse (finer and sparser than on pronotum), disappearing apically. Anterior half of mesoventrite with transverse bracket-like foveae on sides and transverse oval punctures in middle; posterior half with thin long transverse grooves; intercoxal process of mesoventrite convex (evenly rounded in lateral view), widely longitudinally depressed in middle, finely and sparsely punctured, sides wrinkled. Mesepisterna, mesepimera and metepisterna sparsely and finely punctured; metepimera not punctured. Metaventrite with sparse rasp-like larger punctures along mesocoxal cavities and very sparse and almost invisible punctures on rest surface. Abdomen (Fig. 19 C, 20 B, C). Abdominal ventrites very sparsely and finely punctured. Intercoxal process of abdominal ventrite 1 not bordered at the apex. Abdominal ventrite 5 with rectangular emargination at apex, without small but distinct vertical plate in middle; this plate with small emargination in middle. Male aedeagus (Fig. 20 D – F) almost the same as in M. abdullohi sp. nov. and M. minax, with bisinuate lateral margins; basal piece 1.13 times as short as apical piece, widest at basal third. Spiculum gastrale with slightly oval blades, without lateral processes (Fig. 20 G). Legs moderately long. Femora setated with short setae, additionally with sparse and short reddish spines. Metatibiae straight, opaque on extensor side, with very coarse and dense merged puncturation, look rough (Fig. 19 G). Female usually with slightly more robust body and slightly wider pronotum (Fig. 19 B). Body length 8 – 10.5 mm, width 2.7 – 4 mm. Small males and females are almost indistinguishable.	en	Bekchanov, Norbek Kh., Nabozhenko, Maxim V., Bekchanov, Khudaybergan U., Bekchanova, Mokhira Kh. (2025): A review of the genus Microdera Eschscholtz, 1831 (Coleoptera: Tenebrionidae) of Uzbekistan. Zootaxa 5590 (4): 451-480, DOI: 10.11646/zootaxa.5590.4.1, URL: https://doi.org/10.11646/zootaxa.5590.4.1
03D42163EC33BF71FF7FFF35FC33F60F.taxon	etymology	Etymology. The name is derived from the country Uzbekistan.	en	Bekchanov, Norbek Kh., Nabozhenko, Maxim V., Bekchanov, Khudaybergan U., Bekchanova, Mokhira Kh. (2025): A review of the genus Microdera Eschscholtz, 1831 (Coleoptera: Tenebrionidae) of Uzbekistan. Zootaxa 5590 (4): 451-480, DOI: 10.11646/zootaxa.5590.4.1, URL: https://doi.org/10.11646/zootaxa.5590.4.1
03D42163EC33BF71FF7FFF35FC33F60F.taxon	diagnosis	Comparative diagnosis. The new species is externally very similar to M. convexa by the body shape, the opaque elytra with disappearing puncturation apically and laterally and coarsely rough metatibiae on the extensor side but differs from the latter in the very fine and sparse puncturation of the head and pronotum, the non-punctured prohypomera and presence of the vertical plate at rectangularly emarginated apex of the abdominal ventrite 5 (Fig. 20 C). The new species differs from all known Microdera spp. by the flattened pronotal base in middle (the outline of the disc appears as emarginated basally in lateral view) (Fig. 19 A, B, F).	en	Bekchanov, Norbek Kh., Nabozhenko, Maxim V., Bekchanov, Khudaybergan U., Bekchanova, Mokhira Kh. (2025): A review of the genus Microdera Eschscholtz, 1831 (Coleoptera: Tenebrionidae) of Uzbekistan. Zootaxa 5590 (4): 451-480, DOI: 10.11646/zootaxa.5590.4.1, URL: https://doi.org/10.11646/zootaxa.5590.4.1
