identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03D887FDFFDC6D7DFF180F58FE64CBCC.text	03D887FDFFDC6D7DFF180F58FE64CBCC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Boaedon broadleyi Hallermann & Hawlitschek 2025	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Boaedon broadleyi sp. nov.</p>
            <p>(Figs. 3, 4)</p>
            <p> Lamprophis fuliginosus Broadley Doria &amp; Wigge 2003: 236 Photo 81 from Addis Ababa </p>
            <p> Lamprophis fuliginosus Largen &amp; Spawls 2010: 469 Fig. 300 </p>
            <p> Boaedon fuliginosus Spawls et al. 2023: 442 , 443 from Lake Langano </p>
            <p> Holotype: ZMH R08485 (DNA voucher JH 10): an adult female from near Addis Ababa (Abada) (approx. 09° 01’ 48” N 38° 04’ 24” E, 2552 m), Oromia Region, collected by Dr. Peter Angenstein during the end of the 1980s, kept in captivity until 2006 by Ralf Hörold and donated to the Zoological Museum Hamburg (ZMH) in Dec. 2006 (Fig. 3)</p>
            <p>
                  Paratypes (13):  All from Ethiopia: MNHN 4342, an adult female from Ethiopia (Abyssinie) with no specific locality, received by natural produce dealer Deyrolle (Henry or  Emile ) and collected by A. M. J. Raffray in 1874  ;   ZFMK 16231-32, adult females from Negele (approx. 07° 41’ 00” N 39° 15’ 00” E, 1575 m),  
                <a title="Search Plazi for locations around (long 39.25/lat 7.6833334)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=39.25&amp;materialsCitation.latitude=7.6833334">Sidamo Province</a>
                 , Oromia Region, collected by H. Rupp, 28. Feb. 1975  ;   ZFMK 16293, an adult female from  
                <a title="Search Plazi for locations around (long 39.083332/lat 8.391944)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=39.083332&amp;materialsCitation.latitude=8.391944">Lake Koka</a>
                 (approx. 08° 23’ 31” N 39° 05’ 00” E, 1595 m) Oromia Region, collected by H. Rupp, Sept. 1973  ;   ZFMK 16294-95, adult female and juvenile respectively from Arba Minch (approx. 06° 02’ 00” N 37° 33’ 00” E, 1285 m) Southern Nations, Nationalities, and  
                <a title="Search Plazi for locations around (long 37.55/lat 6.0333333)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=37.55&amp;materialsCitation.latitude=6.0333333">Peoples Region</a>
                 , collected by H. Rupp, 26. Mar. 1975  ;   ZFMK 16799, an adult male from Kaki (approx. 08° 56’ 00” N 35° 03’ 00” E, 3000 m)  
                <a title="Search Plazi for locations around (long 35.05/lat 8.933333)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=35.05&amp;materialsCitation.latitude=8.933333">Tulu Welel mountains</a>
                 , Oromia Region, collected by G. Nikolaus, 20.Nov. 1975  ;   ZFMK 19159, an adult female from Chifra (  
                <a title="Search Plazi for locations around (long 40.20996/lat 11.603945)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=40.20996&amp;materialsCitation.latitude=11.603945">Sifra</a>
                 ) (approx. 11° 36’ 14.2” N 40° 12’ 35.86” E, 825 m) Afar Region  ;   ZFMK 84994 a juvenile from  
                <a title="Search Plazi for locations around (long 38.75/lat 9.666667)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=38.75&amp;materialsCitation.latitude=9.666667">Jijiga</a>
                 (approx. 9° 40’ 00” N 38° 45’ 00” E, 1634 m) Somali Region  ;  ZMH R08486, an adult male, same data as holotype ,  ZMH R11333 (DNA voucher JH 59), an adult female same data as holotype ,  ZMH R11341-42, an adult male and female respectively, same data as holotype but donated to the ZMH in 2019 . 
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            <p> Diagnosis.  Boaedon broadleyi sp. nov. can be distinguished from most congeners by a beige to light brown to reddish ground colour, with two short white stripes on the sides of the head. The upper stripe begins just behind the eye on the upper postocular, runs through the lower part of the parietal and the upper first temporal and terminates on the second temporal; the lower stripe passes from the lower postocular through the first temporal and terminates on the last supralabial. Frequently three supralabials contact the eye and two preoculars, the upper one much larger than the lower one touching the frontal. PAR is equal to the distance between the frontal and the rostral scale. Middorsal scale rows varies between 27 and 33 (mean 29.1), ventrals are 223–240 in females and 191–224 in males and subcaudals 45–55 and 59–63 respectively. (Table 3) </p>
            <p> Within Ethiopian  Boaedon ,  Boaedon broadley sp. nov. differs from Ethiopian B. sp. (black form) and Somalian  B. subniger sp. nov. by the combination of its reddish, olive-brown or light brown body colouration with two short head stripes (versus blackish-brown or dark brown body colouration and head stripes missing) with the first two body scale rows uncoloured (versus no dorsals uncoloured) and a higher average of mid-dorsal scale counts of 29.1 (versus 27.0 in  B. subniger sp. nov. ).  Boaedon maculatus has an additional row of scales between the supralabials and loreal between nasal and preocular scale (versus no row of scales) and 10–11 supralabials (versus 8). Extralimital species:  Boaedon montanus (Trape et al. 2022) differs genetically from  B. broadleyi sp. nov. There is no differential diagnostic character to distinguish the species of  B. montanus in its hole distribution area from  B. broadleyi because all meristic characters vary in different populations of this species. Within the type series of  B. montanus one character is different: no contact of the upper preocular with the frontal (versus contact) and PAR longer than distance of frontal to rostral (versus PAR equal to distance from frontal to rostral). Specimens of the Uganda clade (  B. montanus ) and from South Sudan (  cf. montanus ) predominantly have a dark brown coloration (versus light brown in  broadleyi ) and, variably, the preocular is separate from the frontal or in contact with it, predominantly two supralabials touching the eye (91%) (versus 55% in  broadleyi sp. nov. ) and frequently a lower mean of MSR of 28.0– 28.8 in all populations (versus 29.1in  B. broadleyi sp. nov. ) (table 2).  B. perisilvestris with dark brown colouration and no clear head or body lines (versus headlines present and light brown or olive brown colour in  B. broadleyi sp. nov. ) has PAR longer than distance of margin of frontal to end of snout (versus PAR equal in  B. broadleyi sp. nov. ).  B. paralineatus differs in having clearly marginated light headlines anterior and posterior to eye and on anterior part of body (versus short head stripes not well contrasted and no body line in  broadleyi sp. nov. );  B. capensis species complex differs in having loreal length/high ratio mainly 2 or lower (versus 2 and higher) and a dorsal scale count mean of 27.0 (versus 29.1 in  B. broadleyi sp. nov. ). </p>
            <p>Description of holotype: Adult female, 1196 mm SVL; head sub-triangular, slightly distinct from the neck, HL 3.3 % of SVL (39.6 mm); interocular distance 13.7 mm, pupil elliptical, eye diameter 3.2 mm; loreal rectangular, about 2.8 x as long (5 mm) as high (1.8 mm); body cylindrical; tail moderately short (10.3 % of SVL). Supralabials 8/8, 3rd, 4th and 5th on both sides contacting orbit; infralabials 9/10, 1st on each side in contact behind mental, first three on both sides in contact with anterior chin shields and 4th on both sides in contact with posterior chin shields; 2 preoculars on both sides, upper one more than two times larger than lower and touching frontal on the left side, on the right head side it is separated from the frontal by the contact of prefrontal with supraocular; 2 postoculars in contact with the lower part of parietal scale caudally, upper postocular in contact with supraocular anterior-dorsally. Lower postocular in contact with 5 th and 6 th supralabial (both sides of head) ventrally; temporals 1+2+3 on both sides; two internasals; nasal divided; frontal longer (8.3 mm) than wide (4.4 mm); on anterior left border of frontal a small scale is intercalated between frontal and upper preocular; dorsal scales smooth, 22 scale rows one head-length posterior to jaw rictus, 29 scale rows at midbody, 21 scale rows one head length anterior to vent; ventrals 243; anal plate entire; 53 subcaudals, all paired.</p>
            <p>Coloration of holotype in life. The coloration is light olive with a creamy venter, first dorsal scale row creamy. Two short white strips on side of head continuing only on head to the end of supralabials and temporals respectively. In preservative as above but body and head light brown above, first lower dorsal scale row creamy as venter.</p>
            <p> Variation. Variation in measurements and scalation of the paratypes of  B. broadleyi sp. nov. is presented in Table 3. The ground colour of the paratypes in preservative is predominantly light brown in 12 of 13 specimens, one is greyish. Lower 1–3 dorsal scale rows have the same colouration as venter, creamy or whitish. All paratypes have two short white head lines, the extension of the upper head line varies a little, in some specimens it begins on prefrontals runs through preoculars, supraocular, upper postocular, lower part of parietals and terminate on second or third temporals. In others, it begins on upper postocular and runs until first or third temporals. In 5 of 13 paratypes there are three supralabials in contact with the eye. In others only two supralabials are in contact. In all paratypes the upper preocular is in contact with the frontal and two postoculars are in contact with the descending part of the parietal scale. There exist some photo vouchers by Vladimír Tralin from Langano and Filtu which seems to belong to the new species. The colour in life varies from beige, light brown to almost brown. The brown coloured specimen (TMHC 2021.05.976) with two short light headlines from Yabello (DNA voucher TMHC 2021.05.976) belongs definitively to  B. broadleyi sp. nov. (Fig 1). </p>
            <p>Distribution. The species is currently known from different localities in elevations from approx. 750 m up to 3000 m asl in Ethiopian Regional States in Afar, Oromia, Somali and Southern Nations, Nationalities, and Peoples’ National Regional State and most probably in Northern Somalia (Somaliland) (Fig. 2).</p>
            <p>Habitat and Natural History notes. The specimens were found in savannas, grasslands and woodlands, absent in arid or semi-arid habitats.</p>
            <p> Etymology. The specific epithet is a patronym in the masculine genitive singular name of Donald George (Don) Broadley, a British born Herpetologist and emeritus curator of the Natural History Museum of Zimbabwe in Bulawayo. We honour his extensive work on African reptiles especially on  Boaedon , which he could not publish before he died in 2016. </p>
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	https://treatment.plazi.org/id/03D887FDFFDC6D7DFF180F58FE64CBCC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hallermann, Jakob;Hawlitschek, Oliver	Hallermann, Jakob, Hawlitschek, Oliver (2025): Two new species of Boaedon from Ethiopia and Somalia, with a review of the species of East Africa. Zootaxa 5569 (1): 119-137, DOI: 10.11646/zootaxa.5569.1.4, URL: https://doi.org/10.11646/zootaxa.5569.1.4
03D887FDFFD36D7DFF180D59FA75C8E7.text	03D887FDFFD36D7DFF180D59FA75C8E7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Boaedon broadleyi Hallermann & Hawlitschek 2025	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Boaedon broadleyi sp. nov.</p>
            <p> Ethiopia: Oromia region: MNHN 1905.0194 (juvenile) Mega (former Abysinie, Mego), (approx. 04° 03’ N 38° 18’ E) ;  TMHC 2021.05.976 (adult, collector Vladimír Trailin DNA voucher), Yabello (04° 53’ N 38° 05’ E, 1906 m) . </p>
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	https://treatment.plazi.org/id/03D887FDFFD36D7DFF180D59FA75C8E7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hallermann, Jakob;Hawlitschek, Oliver	Hallermann, Jakob, Hawlitschek, Oliver (2025): Two new species of Boaedon from Ethiopia and Somalia, with a review of the species of East Africa. Zootaxa 5569 (1): 119-137, DOI: 10.11646/zootaxa.5569.1.4, URL: https://doi.org/10.11646/zootaxa.5569.1.4
03D887FDFFD36D78FF180DA4FD19CC26.text	03D887FDFFD36D78FF180DA4FD19CC26.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Boaedon subniger Hallermann & Hawlitschek 2025	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Boaedon subniger sp. nov.</p>
            <p>(Figs. 5, 6)</p>
            <p> Boaedon fuliginosus Spawls, Mazuch &amp; Mohamad 2023: 443 from Erigavo (Somaliland) </p>
            <p>
                  Holotype: ZMH R22350 (DNA voucher TMHC 2023.01.1018): a juvenile female from 8.5 km south of  
                <a title="Search Plazi for locations around (long 47.182777/lat 10.3)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=47.182777&amp;materialsCitation.latitude=10.3">Yuffleh</a>
                 (Yufle, Jooflie) (approx. 10° 18’ 00” N 47° 10’ 58” E, 1758 m), Sanaag region, Somaliland collected by Tomáš Mazuch, 31 August 2017, and donated to the Zoological Museum Hamburg (ZMH) in September 2024 (Figs. 5, 6) 
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            <p> Diagnosis.  Boaedon subniger sp. nov. can be distinguished from most congeners by its dark brown to blackbrown dorsal colour, without any light head stripes, two supralabials (4 th &amp; 5 th) touching on eye and one preocular, touching the frontal. PAR is longer than the distance between the frontal and the rostral scale. Venter whitish (for more characters see table 4). </p>
            <p> Boaedon subniger sp. nov. differs from  B. broadleyi sp. nov. by having fewer MSR (, X 27.0, versus 27–33, X 29.1, only one preocular (versus two) and its blackish brown or dark brown body colouration without light coloured head stripes (versus reddish, olive-brown or light brown body colouration with two short white head stripes and lips whitish coloured), with no dorsals uncoloured (versus first two rows uncoloured in  broadleyi sp. nov. ).  Boaedon maculatus has an additional row of scales between the supralabials and loreal between nasal and preocular scale, (versus no row of scales) and 10–11 supralabials (versus 8). Extralimital species:  Boaedon montanus in its whole distribution area differs in having two at least short head lines and whitish upper and lower lips (versus no headlines and no whitish lips),  Boaedon perisilvestris with similar colouration differs in having more MSR (29–31, X 29.4 (versus 27 X 27.0 in  B. subniger sp. nov. ) and frequently three supralabial scales touching eye (versus two in  B. subniger sp. nov. ) and a genetic interspecific distances of 0.024).  B. paralineatus and  B. capensis species complex differs in having clear head and/or body lines (versus no head and body lines in  B. subniger sp. nov. ) and a brown or reddish colouration (versus blackish brown colouration in  B. subniger sp. nov. ) </p>
            <p>Description of holotype: Juvenile female, 222 mm SVL; head sub-triangular, slightly distinct from the neck, HL 11.4 mm (5.1 % of SVL (222 mm); interocular distance 4.0 mm, pupil elliptical, eye diameter 2.3 mm; loreal rectangular, about 2.1 x as long (1.5 mm) as high (0.7 mm); body cylindrical; tail moderately short (8.5 % of SVL). Supralabials 8/8, 3th, 4th and 5th on left side of head and 4 th and 5 th on right side of head contacting orbit; infralabials 9/9, 1st on each side in contact behind mental, first three on both sides in contact with anterior chin shields and 4th on both sides in contact with posterior chin shields; one preocular on both sides touching; two postoculars in contact with the lower part of parietal scale caudally, upper postocular in contact with supraocular anterior-dorsally. Lower postocular in contact with 5th and 6th supralabial (both sides of head) ventrally; temporals 1+2+3 on both sides; two internasals; nasal divided; frontal longer (3.5 mm) than wide (1.9 mm); Parietal length (3.8 mm) longer than distance from frontal to rostral (2.3 mm); dorsal scales smooth, 22 SR one head-length posterior to jaw rictus, 27 MSR at midbody, 21 SR one head length anterior to vent; ventrals 225; anal plate entire; 43 subcaudals, all paired.</p>
            <p>Colouration of holotype in ethanol: dorsal body colouration dark brown without any light head stripes. Venter whitish, no dorsal scales light coloured, upper and lower labials same colour as dorsum.</p>
            <p>Colouration in life (Fig. 6): as above, but colour almost black.</p>
            <p>Variation: see Table 4.</p>
            <p>Distribution: the species is found only in Northern Somalia (Sanaag region, Somaliland (1760–1780 m asl) (Fig. 2).</p>
            <p>Habitat and Natural History notes: The specimens were found in high-altitude semiarid or arid habitats from Somaliland (Sanaag region).</p>
            <p> Etymology: The species is named after the Latin word  “subniger ” for blackish, somewhat dark, from its blackish body coloration without any stripes on head or body. </p>
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	https://treatment.plazi.org/id/03D887FDFFD36D78FF180DA4FD19CC26	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hallermann, Jakob;Hawlitschek, Oliver	Hallermann, Jakob, Hawlitschek, Oliver (2025): Two new species of Boaedon from Ethiopia and Somalia, with a review of the species of East Africa. Zootaxa 5569 (1): 119-137, DOI: 10.11646/zootaxa.5569.1.4, URL: https://doi.org/10.11646/zootaxa.5569.1.4
