taxonID	type	description	language	source
03E68793FF93FFFB5FB219A47300FA2E.taxon	description	Fig 1 A – E, 2 A – J, 3	en	Uchima-Taborda, Diego, Hortúa, Andrés Lugo De La, Vega-Badillo, Viridiana, Fernández, Fernando, Zaragoza-Caballero, Santiago (2025): Platysognathus gen. nov. a new genus of railroad-worm beetle (Coleoptera: Phengodidae, Mastinocerinae) from Colombian Amazonía. Zootaxa 5636 (3): 577-586, DOI: 10.11646/zootaxa.5636.3.11, URL: https://doi.org/10.11646/zootaxa.5636.3.11
03E68793FF93FFFB5FB219A47300FA2E.taxon	diagnosis	Diagnosis: Head wider than long, feebly dorsoventrally flattened (Fig. 1 C – E). Mandibles large and anteriorly projected, with a wide medial irregularly rhomboid tooth (Fig. 2 A). In ventral view submentum expanded and projected, buccal parts such as lacinia and the insertion of the maxillary palps, visible in dorsal view (Fig. 1 E). Maxillary palps four segmented, labial three segmented (Fig 2 B). Antennae with 11 antennomeres, biflabellate from IV to X, and antennomere XI filiform (Fig 1 C). First pro- and mesotarsomere with a ventral tarsal comb, tarsal claws falcated and with basal spines (Fig. 2 C – D). Elytra short and lanceolate (Fig. 2 E). Genitalia slightly asymmetrical, parameres with apical teeth, flagellum edge serrated (Fig. 2 G, I – J).	en	Uchima-Taborda, Diego, Hortúa, Andrés Lugo De La, Vega-Badillo, Viridiana, Fernández, Fernando, Zaragoza-Caballero, Santiago (2025): Platysognathus gen. nov. a new genus of railroad-worm beetle (Coleoptera: Phengodidae, Mastinocerinae) from Colombian Amazonía. Zootaxa 5636 (3): 577-586, DOI: 10.11646/zootaxa.5636.3.11, URL: https://doi.org/10.11646/zootaxa.5636.3.11
03E68793FF93FFFB5FB219A47300FA2E.taxon	discussion	Remarks: This genus differs from the rest of Phengodidae with 11 antennomeres by the presence of this characteristic irregularly rhomboid mandibular tooth. Until now, only Euryognathus, Wittmer 1976 had been described with both 11 antennomeres and mandibular tooth; however, in Euryognathus the mandibular tooth is rhomboid while in Platysognathus is irregularly rhomboid with an acute projection in the middle, giving the appearance of a wide apex tooth due to the transect between its anterior emergence to its apex. Additionally, in Euryognathus the antennal flabellae gradually fuse towards the last antennomere, while in Platysognathus are elongated, simple and non-fused to the antennomere.	en	Uchima-Taborda, Diego, Hortúa, Andrés Lugo De La, Vega-Badillo, Viridiana, Fernández, Fernando, Zaragoza-Caballero, Santiago (2025): Platysognathus gen. nov. a new genus of railroad-worm beetle (Coleoptera: Phengodidae, Mastinocerinae) from Colombian Amazonía. Zootaxa 5636 (3): 577-586, DOI: 10.11646/zootaxa.5636.3.11, URL: https://doi.org/10.11646/zootaxa.5636.3.11
03E68793FF93FFFB5FB219A47300FA2E.taxon	description	Description: Male. Head: Wider than long, longer than pronotum, and dorsoventrally flattened (Fig. 1 A, C). In lateral view, eyes semispherical and reduced in relation to temples (Fig. 1 B, D). In both dorsal and ventral view eye is smaller than interocular distance (Fig. 1 C, E). Interantennal distance wider than scape length and wider than antenal socket width (Fig. 1 C – D). Clypeus fused to frons, in dorsal view labrum bilobed surpassing clypeus; maxillary and labial palps insertion, as well as lacinia visible between the mandibles (Fig. 1 C, E). Mandibles large, not occluding, in middle with pronounced mandibular tooth which is irregularly rhomboid (Fig. 1 C, 2 A). In ventral view, with single tentorial fossa, submentum spatulated, anteriorly projected; mandibular insertions large and depressed, strongly marked (Fig. 1 E). Galea densely setose, maxillary palps four-segmented, labial palps three-segmented (Fig. 2 B). Antennae 11 segmented, antennomeres 4 to 10 biflabellate, flabellae branching from base of antennomere, flabellae and last antennomere filiform (Fig. 1 C). Thorax: Pronotum transverse (Fig. 1 C – D). Elytra short, slightly surpassing first abdominal segment and longer than pronotum (Fig. 1 A, 2 E). Hind wings with anal and radial cells both closed; r 4 present and r 3 absent, AA 3 absent fused to CAS (Fig. 2 F). Tarsal combs on first pro- and mesotarsomere, tarsal claws falcated (Fig. 2 C – D). Abdomen: Abdominal segments simple, without expansions. The aedeagus slightly asymmetrical, central lobe wedge-shaped; paramere apex with truncated tooth (Fig. 2 H – I). Genital flagellum longer than aedeagus length, border of flagellum serrated (Fig. 2 J). Female and larvae unknown.	en	Uchima-Taborda, Diego, Hortúa, Andrés Lugo De La, Vega-Badillo, Viridiana, Fernández, Fernando, Zaragoza-Caballero, Santiago (2025): Platysognathus gen. nov. a new genus of railroad-worm beetle (Coleoptera: Phengodidae, Mastinocerinae) from Colombian Amazonía. Zootaxa 5636 (3): 577-586, DOI: 10.11646/zootaxa.5636.3.11, URL: https://doi.org/10.11646/zootaxa.5636.3.11
03E68793FF93FFFB5FB219A47300FA2E.taxon	etymology	Etymology: This name is a combination of the words “ Platys- ” meaning broad, wide and “ - gnathus ” meaning mandible. This name refers to the wide tooth in the mandible.	en	Uchima-Taborda, Diego, Hortúa, Andrés Lugo De La, Vega-Badillo, Viridiana, Fernández, Fernando, Zaragoza-Caballero, Santiago (2025): Platysognathus gen. nov. a new genus of railroad-worm beetle (Coleoptera: Phengodidae, Mastinocerinae) from Colombian Amazonía. Zootaxa 5636 (3): 577-586, DOI: 10.11646/zootaxa.5636.3.11, URL: https://doi.org/10.11646/zootaxa.5636.3.11
03E68793FF93FFFD5FB21F2074D9F857.taxon	description	Fig. 1 A – E, 2 A – J, 3	en	Uchima-Taborda, Diego, Hortúa, Andrés Lugo De La, Vega-Badillo, Viridiana, Fernández, Fernando, Zaragoza-Caballero, Santiago (2025): Platysognathus gen. nov. a new genus of railroad-worm beetle (Coleoptera: Phengodidae, Mastinocerinae) from Colombian Amazonía. Zootaxa 5636 (3): 577-586, DOI: 10.11646/zootaxa.5636.3.11, URL: https://doi.org/10.11646/zootaxa.5636.3.11
03E68793FF93FFFD5FB21F2074D9F857.taxon	materials_examined	Examined material: Holotype: Colombia: Vaupés: Estación Biológica Mosiro-Itajura (Caparú) - Centro Ambiental, 1 ° 4 ’ S - 69 ° 31 ’ W, 20 / 01 - 1 / 02 / 2003, 60 msnm, Trampa Malaise, M. Sharkey, IAvH-E- 262320 (Pinned) (1 Male). Paratypes: Colombia: Caquetá: PNN Serranía de Chiribiquete - Cuñaré-Amú, Bosque tierra firme, 0 ° 13 ’ N - 72 ° 25 ’ W, 19 – 22 / 02 / 2001, 300 msnm, Trampa Malaise, E. González, IAvH-E- 262323 (Alcohol) (1 Male), Ídem except for IAvH-E- 262324 (Alcohol) (1 Male). Ídem except for 0 ° 30 ’ N - 72 ° 38 ’ W, 1 – 5 / 11 / 2000, Trampa Malaise, E. González, IAvH-E- 262325 (Pinned) (1 Male). Vaupés: Estación Biológica Mosiro-Itajura (Caparú) - Antigua Cabaña, 1 ° 4 ’ S - 69 ° 31 ’ W, 9 – 14 / 04 / 2004, 60 msnm, Trampa Malaise, J. Pinzón, IAvH-E- 262318 (Alcohol) (1 Male), IAvH-E- 262321 (Alcohol) (1 Male). Ídem except for 21 – 28 / 04 / 2004, 60 msnm, Trampa Malaise, J. Pinzón, IAvH- E- 262322 (Alcohol) (1 Male). Ídem except for Estación Biológica Mosiro-Itajura (Caparú) - Centro Ambiental, 1 ° 4 ’ S - 69 ° 31 ’ W, 20 / 01 – 01 / 02 / 2003, M. Sharkey, IAvH-E- 262319 (Pinned) (1 Male).	en	Uchima-Taborda, Diego, Hortúa, Andrés Lugo De La, Vega-Badillo, Viridiana, Fernández, Fernando, Zaragoza-Caballero, Santiago (2025): Platysognathus gen. nov. a new genus of railroad-worm beetle (Coleoptera: Phengodidae, Mastinocerinae) from Colombian Amazonía. Zootaxa 5636 (3): 577-586, DOI: 10.11646/zootaxa.5636.3.11, URL: https://doi.org/10.11646/zootaxa.5636.3.11
03E68793FF93FFFD5FB21F2074D9F857.taxon	diagnosis	Diagnosis: Eyes small, semispherical, in dorsal view interocular distance five times eye width, in ventral view 4.7 – 5 times (Fig. 1 C – E). Mandibular tooth irregularly rhomboid with a medial acute projection (Fig. 2 A). Maxillary palps around three times longer than labial palps, maxillary palpomere IV sub equal to II + III and obovate; labial palpomere III longer than I + II, fusiform and thickened in the middle (Fig 2 B). Scape longer than antennomeres II + III, flabellae short and slightly widened towards the apex (Fig 1 C). Pronotum transverse, quadrangular, anterior border straight, posterior border, sides and corners rounded (Fig 1 C). Elytra short, surpassing the second abdominal segment (Fig. 1 A, 2 E). Genitalia asymmetrical, parameres with an apical tooth (Fig. 2 G – I) and genital flagellum with a strongly serrated edge (Fig. 2 J).	en	Uchima-Taborda, Diego, Hortúa, Andrés Lugo De La, Vega-Badillo, Viridiana, Fernández, Fernando, Zaragoza-Caballero, Santiago (2025): Platysognathus gen. nov. a new genus of railroad-worm beetle (Coleoptera: Phengodidae, Mastinocerinae) from Colombian Amazonía. Zootaxa 5636 (3): 577-586, DOI: 10.11646/zootaxa.5636.3.11, URL: https://doi.org/10.11646/zootaxa.5636.3.11
03E68793FF93FFFD5FB21F2074D9F857.taxon	description	Description: Male. Color: Body in general orange yellow, with exception of antennae slightly clearer, elytra brown darkened at middle and lighter towards apex, legs lighter than body (Fig 1 A – B). Total length 8.6, width at shoulders high 1.66. Head: Head wider (1.78) than long (1.40), 1.4 times as wide as pronotum (Fig. 1 A, C). Eyes small and reduced (0.48) when compared to the temples (0.56), latter straight in its basal half, abruptly expanding near middle, then convex to reach eye (Fig. 1 C, E). Eyes semispherical, posterior border oblique near straight, in lateral view taller (0.48) than long (0.46); interocular distance in dorsal view wide (1.29), five times eye width (0.27) in ventral view 4.7 times (Fig. 1 C – D). Antennal insertions covered with disperse punctuation with foveae smaller than its lumen, punctuation similar in vertex. Interantennal distance wide (0.68) 1.8 to 2 times scape length (0.37) (Fig 1 C). Frontoclypeus anterior border undulated, concave, weakly projected medially, this projection apically truncated, with sparse and scarce punctuations. In dorsal view, membranous and bilobed labrum surpassing frontoclypeus and lacinia; insertion of maxillary and labial palps visible between mandibles (Fig 1 C). Mandibles prominent (1.05) mandibular tooth irregularly rhomboid with medial acute projection (0.27), tooth angles feebly projected giving a bilobed tooth appearance, in general both teeth asymmetrical, left convex, right sinuated (Fig. 1 C, 2 A). Mentum spatulated, anteriorly projected; mandibular insertion large, depressed, strongly pronounced (Fig. 1 E). Maxillary palps three times longer (0.8) than labial palps (0.24); maxillary palps with thickened insertion, palpomere I small, 2.5 times shorter than II, II longer than III, both fusiform, IV sub equal to II + III and obovate; first labial palpomere shorter than the others, II shorter than III, III longer than I + II; I – II cylindrical, III fusiform medially thickened (Fig. 2 B). Antennae (2.48) feebly surpassing pronotum posterior border, flabellae short and expanding towards apex, flabellae of antennomeres IV – VI from 2.5 to 2.9 times longer than its respective antennomere. Scape (0.37) longer than II + III (0.23), last antennomere S-shaped (Fig 1 C – E). Thorax: Pronotum rectangular, wider (1.56) than long (1.23), posterior border straight feebly undulated, lateral borders basally straight, curving from last quarter to apex; posterior corners right-angled, not expanded, posterior border and corners curved, lateral borders not expanded. Disc punctuation superficial, lumen feebly chagrined and greater than puncture diameter. Prosternum posterior border and procoxa external border straight (Fig. 1 D – E). Scutellum nearly as wide (0.26) as long (0.28), posterior border convex. Elytra short, subtriangular, length (2.19) 2.73 times its width at shoulder height (0.80), 1.7 times pronotum length, surpassing first abdominal segment; covered by disperse punctation, lumen subequal to puncture diameter, punctuation denser towards apex (Fig 1 A, 2 E). Hind wing length (4.3) 2.38 times width (1.8), radial cell closed, r 4 complete reaching radial cell and RP, r 3 absent, reduced RP not reaching half of MP 1 + 2, midfield with five main veins MSP, MP 3, MP 4, CuA 2, CAS; CuA 1, AA 4 and CuP reduced and fused, AA 3 + 4 straight, AA 3 fused to CAS, AA 4 absent, AP well marked, AP 3 + 4 long (Fig. 2 F). Mesosclerite and metasclerite sparsely punctuated, puncture diameter smaller than lumen. Tibiae 1.2 to 1.5 times longer than tarsi; tarsal combs on first pro- and mesotarsomere, with seven and nine denticles respectively, first tarsomere nearly as large as 2 nd and 3 rd together, fourth smaller than the rest and one quarter of the length of the first, tarsal claws falcate with two spines at base, claws separated in straight angle (Fig. 2 B – C). Abdomen: Tergites and ventrites surface lustrous and covered by superficial punctuation. Genitalia with central lobe wedge-shaped; left paramere smaller than right, paramere apex with one truncated flattened tooth, covered with long and clear setae. Left paramere curves and anchors slightly behind the central lobe, while the right paramere protrudes anteriorly (Fig. 2 G – I). Genital flagellum 1.5 times aedeagus length, edge of this flagellum strongly serrated (Fig. 2 J).	en	Uchima-Taborda, Diego, Hortúa, Andrés Lugo De La, Vega-Badillo, Viridiana, Fernández, Fernando, Zaragoza-Caballero, Santiago (2025): Platysognathus gen. nov. a new genus of railroad-worm beetle (Coleoptera: Phengodidae, Mastinocerinae) from Colombian Amazonía. Zootaxa 5636 (3): 577-586, DOI: 10.11646/zootaxa.5636.3.11, URL: https://doi.org/10.11646/zootaxa.5636.3.11
03E68793FF93FFFD5FB21F2074D9F857.taxon	distribution	Distribution: Amazonic ecosystems, in the biogeographical province Imeri (Morrone et al. 2022), known to occur in the PNN “ Serranía del Chibiriquete ” and in the biological station “ Mosiro Itajurá-Caparú ” (Fig. 3).	en	Uchima-Taborda, Diego, Hortúa, Andrés Lugo De La, Vega-Badillo, Viridiana, Fernández, Fernando, Zaragoza-Caballero, Santiago (2025): Platysognathus gen. nov. a new genus of railroad-worm beetle (Coleoptera: Phengodidae, Mastinocerinae) from Colombian Amazonía. Zootaxa 5636 (3): 577-586, DOI: 10.11646/zootaxa.5636.3.11, URL: https://doi.org/10.11646/zootaxa.5636.3.11
03E68793FF93FFFD5FB21F2074D9F857.taxon	etymology	Etymology: the species name “ amazonicus ” refers to the amazonic ecosystems where the species is distributed, the PNN “ Serranía del Chibiriquete ” and in the biological station “ Mosiro Itajurá-Caparú ”.	en	Uchima-Taborda, Diego, Hortúa, Andrés Lugo De La, Vega-Badillo, Viridiana, Fernández, Fernando, Zaragoza-Caballero, Santiago (2025): Platysognathus gen. nov. a new genus of railroad-worm beetle (Coleoptera: Phengodidae, Mastinocerinae) from Colombian Amazonía. Zootaxa 5636 (3): 577-586, DOI: 10.11646/zootaxa.5636.3.11, URL: https://doi.org/10.11646/zootaxa.5636.3.11
