identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03E7726733384828FF39994B0BA78298.text	03E7726733384828FF39994B0BA78298.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cyamophila Loginova 1976	<div><p>Genus Cyamophila Loginova, 1976</p><p>Cyamophila Loginova, 1976: 596 . Type species: Psylla fabra Loginova, 1964, by original designation.</p></div>	https://treatment.plazi.org/id/03E7726733384828FF39994B0BA78298	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Inoue, Hiromitsu	Inoue, Hiromitsu (2025): Taxonomy and DNA barcoding of Cyamophila (Hemiptera: Psyllidae) from Japan, with the description of a new species. Zootaxa 5727 (1): 148-169, DOI: 10.11646/zootaxa.5727.1.10, URL: https://doi.org/10.11646/zootaxa.5727.1.10
03E77267333B482CFF399C370A2B86AE.text	03E77267333B482CFF399C370A2B86AE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cyamophila hexastigma (Horvath 1899)	<div><p>Cyamophila hexastigma (Horváth, 1899)</p><p>(Japanese name: Mutsuboshi-kijirami)</p><p>(Figs 1–6, 7–10, 26, 29, 32, 35–36, 41–42)</p><p>Psylla hexastigma Horváth, 1899: 373 . Type locality: Japan (Hokkaidô).</p><p>Cyamophila hexastigma: Loginova (1977: 582).</p><p>Cyamophila floribundae Cho &amp; Burckhardt in Cho et al. (2017: 551), syn. nov.</p><p>Description. Adult. Colouration. General colour (Figs 35, 41) light yellowish green with obscure light yellowish brown markings and longitudinal stripes on thorax; in overwintered individuals (Fig. 36), body colour chestnut brown to dark brown in general, with many cream-yellow coloured fine markings and stripes on head and thorax. Antenna yellow; apices of segments IV–VI and most of segment VII dark brown; segments VIII–X entirely dark brown to black. Forewing membrane transparent, colourless in general but faintly yellowish along veins in the apical half of forewing in fully matured adults, with a prominent dark brown spots at the location of the radular spinules in cells m 1, m 2, and cu 1 (Fig. 3); veins yellow in summer adults, becoming uniformly dark brown in overwintered adults. Apical tooth of paramere black.</p><p>Structure. Head (Fig. 1) strongly inclined downwards, 85–90° from longitudinal body axis in profile, slightly wider than thorax. Vertex nearly half as long as wide or slightly shorter. Genal processes conical, about 0.8 times as long as vertex, slightly divergent and subacute apically. Antenna long, 2.2–2.4 times as long as head width; longer terminal seta of segment X about twice as long as shorter seta, about 0.6 times as long as segment X (Fig. 2).</p><p>Forewing (Fig. 3) oblong oval, 2.3–2.4 times as long as wide, widest at around 2/3 from the base; membrane with dense surface spinules in all cells; spinule-free bands along veins rather broad; fields of radular spinules as in Fig. 3; pterostigma well developed, more than 1/3 of the forewing length; Rs slightly sinuate, slightly curved towards costal margin apically; M 1+2 rather strongly arched at around the basal 1/3 and nearly straight thereafter, very slightly curved towards costal margin apically; Cu 1a strongly arched around the basal 1/4 and nearly straight thereafter. Meracanthus moderate in size, subacute and slightly curved downwards apically; metatibia with prominent genual spine, with five apical sclerotised spurs arranged in 1+3+1; basal segment of metatarsus with a pair of sclerotised lateral spurs.</p><p>Male terminalia (Fig. 4) moderate in size. Proctiger slender, slightly curved caudad apically. Paramere stout, 0.7 times as long as proctiger, strongly constricted in the middle, strongly widened towards the tip, weakly produced cephalad at anteroapical corner, and strongly produced caudad at posteroapical corner; inner surface (Fig. 5) with many retrorse setae; inner apical tooth prominent, acute, and projected cephalad. Distal aedeagal segment slightly longer than paramere; apex round and thickened, strongly hooked.</p><p>Female terminalia (Fig. 6) rather stout. Proctiger curved and slightly sinuate at dorsal margin, slightly upturned apically, and slightly truncated at apex (Fig. 32). Subgenital plate with sparse setae, acute at apex.</p><p>Fifth instar immature. Body (Fig. 42) light green, gently swollen dorsally. Antenna slender, 2.1–2.2 times as long as forewing pad, seven-segmented, with one apical rhinarium each on segments III and V, and two on the middle of segment VII, with rather long capitate setae on the middle of segment III and apices of segments III–V (Fig. 26). Forewing pad oblong oval; outer margin with 8–9 long capitate setae and short simple setae present in between (Fig. 29). Hindwing pad with two long capitate setae apically. Legs long, hairy, with many short simple setae and long capitate setae. Abdomen rounded apically, with many long capitate setae dorsally and many long simple setae ventrally. Caudal plate with many long capitate setae on dorsum and margin, with 4+4 truncated sectasetae on posterior margin. Anus located on ventral side. Outer circumanal pore ring relatively small, heart-shaped, strongly curved in front, comprising a single row of elongated pores. Inner circumanal pore ring comprising a single row of small elongated pores.</p><p>Measurements (in mm): Adult (n = 5 males, 5 females): BL: 3.13–3.80; WL: 2.59–3.15; WW: 1.06–1.35; AL: 1.90–2.33; HW: 0.82–0.96; VW: 0.46–0.58; VL: 0.22–0.27; GL: 0.18–0.23; MP: 0.36–0.43; PL: 0.27–0.31; FP: 0.68–0.83. Fifth instar immature (n = 5): BL: 2.28–2.60; BW: 1.23–1.50; AL: 1.38–1.63; WPL: 0.65–0.78; and CRW: 0.09–0.10.</p><p>Material examined. Hokkaidô: 5 males, 3 females, 2 immatures, Sapporo-shi, Minami-ku, Jôzankei, 300 m, 9.vii.2001, on Maackia amurensis, H. Inoue (dry- and slide-mounted; HIC) ; 13 males, 12 females, Sapporo-shi, Minami-ku, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=141.159&amp;materialsCitation.latitude=42.982" title="Search Plazi for locations around (long 141.159/lat 42.982)">Jôzankei</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=141.159&amp;materialsCitation.latitude=42.982" title="Search Plazi for locations around (long 141.159/lat 42.982)">Jôzankei Dam</a>, 42.982º N, 141.159º E, 300 m, 19–20.vii.2012, on M. amurensis, H. Inoue (dry- and slide-mounted; HIC) ; 18 males, 24 females, same data, 12.vi.2013 (dry- and slide-mounted; HIC); 9 males, 8 females, 23 immatures, Kamikawa-gun, Shimizu-chô, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=142.905&amp;materialsCitation.latitude=42.957" title="Search Plazi for locations around (long 142.905/lat 42.957)">Haobi</a>, 42.957º N, 142.905º E, 190 m, 30.vi.2015, on M. amurensis, H. Inoue (dry- and slide-mounted, 99.5% and 70% ethanol; HIC) ; 3 males, 1 females, Sarugun, Hidaka-chô, Chiroro-rindô, 2.vii.1998, N. Takahashi (dry-mounted; HIC) ; 1 female, Akan-gun, Akan-chô, Meakan-dake, 29.vi.1998, N. Takahashi (dry-mounted; HIC) ; Honshû: 8 males, 8 females, 4 immatures, Aomori-ken, Sannohe-gun, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=141.131&amp;materialsCitation.latitude=40.444" title="Search Plazi for locations around (long 141.131/lat 40.444)">Shingô-mura</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=141.131&amp;materialsCitation.latitude=40.444" title="Search Plazi for locations around (long 141.131/lat 40.444)">Herai</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=141.131&amp;materialsCitation.latitude=40.444" title="Search Plazi for locations around (long 141.131/lat 40.444)">Kosaka</a>, 40.444º N, 141.131º E, 170 m, 2.vii.2015, on M. amurensis, H. Inoue (dry- and slide-mounted, 99.5% ethanol; HIC) ; 3 males, 5 females, Aomori-ken, Sannohe-gun, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=141.149&amp;materialsCitation.latitude=40.454" title="Search Plazi for locations around (long 141.149/lat 40.454)">Shingô-mura</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=141.149&amp;materialsCitation.latitude=40.454" title="Search Plazi for locations around (long 141.149/lat 40.454)">Herai</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=141.149&amp;materialsCitation.latitude=40.454" title="Search Plazi for locations around (long 141.149/lat 40.454)">Sawaguchi</a>, 40.454º N, 141.149º E, 160 m, 2.vii.2015, on M. amurensis, H. Inoue (dry- and slide-mounted; HIC) ; Kyûshû: 2 females, Fukuoka-ken, Miyako-gun, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=130.97&amp;materialsCitation.latitude=33.502" title="Search Plazi for locations around (long 130.97/lat 33.502)">Miyako-machi</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=130.97&amp;materialsCitation.latitude=33.502" title="Search Plazi for locations around (long 130.97/lat 33.502)">Saigawahobashira</a>, near <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=130.97&amp;materialsCitation.latitude=33.502" title="Search Plazi for locations around (long 130.97/lat 33.502)">Notôge</a>, 33.502º N, 130.970º E, 750–930 m, 9.vi.2014, H. Inoue (slide-mounted; HIC) ; 19 males, 26 females, 10 immatures, Nagasakiken, Unzen-shi, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=130.27&amp;materialsCitation.latitude=32.745" title="Search Plazi for locations around (long 130.27/lat 32.745)">Obama-chô</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=130.27&amp;materialsCitation.latitude=32.745" title="Search Plazi for locations around (long 130.27/lat 32.745)">Unzen</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=130.27&amp;materialsCitation.latitude=32.745" title="Search Plazi for locations around (long 130.27/lat 32.745)">Ikenohara</a>, 32.745º N, 130.270º E, 750 m, 27.v.2010, on Maackia floribunda, H. Inoue (dry- and slide-mounted; HIC) ; 136 males, 140 females, 4 immatures, same data, 4.vi.2010 (dry- and slide-mounted, 99.5% ethanol; HIC); 2 males, 5 females, same data, 14.iv.2013 (dry-mounted; HIC); 38 males, 50 females, same data, 13.vi.2014 (dry-mounted, 99.5% ethanol; HIC) .</p><p>Distribution. Japan (Hokkaidô, Honshû, Shikoku, Kyûshû, Tsushima); China (Beijing, Jilin, Liaoning, Shanxi), South Korea, Russian Far East (Khabarovsk Territory, Primorsky Territory) (Wu 1932, as P. hexastigma; Sasaki 1954, as P. hexastigma; Miyatake 1963, as P. hexastigma; Kuwayama &amp; Miyatake 1971, as P. hexastigma; Miyatake 1976, as P. hexastigma; Park et al. 1979, as P. hexastigma; Konovalova 1988; Li 2011; Cho et al. 2017, as C. floribundae; Kwon &amp; Kwon 2020).</p><p>Host plant. Maackia amurensis Rupr. (Miyatake 1976); Maackia floribunda (Miq.) Takeda ( Fabales: Fabaceae) (new host plant record). These host plants were confirmed by the presence of immatures.</p><p>Styphnolobium japonicum (= Sophora japonica) ( Fabaceae) recorded by Kwon (1983) is the host for Cyamophila willieti (Cho et al. 2022) . Kuwayama (1932) referred to Hydrangea ( Cornales: Hydrangeaceae) as follows: ‘In Hokkaidô, adults of Psylla hexastigma are abundant on Saxifragaceae plants, such as Hydrangea paniculata and Hydrangea serrata, around August. This species likely feeds on these plants’; however, these plants are not hosts on which the immatures develop. Hydrangea has been cited as a host for this psyllid species in subsequent studies, such as in Sasaki (1954), Miyatake (1963), Klimaszewski (1973), Kwon (1983), and Kwon &amp; Kwon (2020), and is therefore formally eliminated here as the host of C. hexastigma .</p><p>Biology. Univoltine. Adults emerge from late May (Kyûshû in southwest Japan) to early July (Hokkaidô in northern Japan) and leave the host plant within a short period. Dispersed adults are thought to stay on the evergreen shelter plants until spring.</p><p>Comments. Cyamophila floribundae Cho &amp; Burckhardt, which feeds on Maackia floribunda, was described from Jeju Island, in the southernmost part of South Korea (Cho et al. 2017). In my field survey, a Cyamophila population feeding on M. floribunda was confirmed from Kyûshû, Japan; this population cannot be distinguished from C. hexastigma populations feeding on M. amurensis from Hokkaidô, from which this psyllid species was originally described, through morphology as well as DNA barcoding (Fig. 48). According to Cho et al. (2017), C. floribundae differs from C. hexastigma in the following aspects: 1) denser surface spinules of forewing membrane; 2) slightly widened parameres towards the apex; and 3) host plant. However, 1) some variation was observed in the density of the forewing surface spinules in C. hexastigma, ranging from sparse (Figs 7, 9) to dense (Figs 8, 10), even within the same locality, although the spinules in M. floribunda -feeders tended to be somewhat denser. 2) No stable morphological differences were observed in paramere between the populations of different host plants. These slight differences were likely within the range of intraspecific variation. 3) Notably, Japanese C. hexastigma also feeds on M. floribunda, implying that these two nominal psyllid species cannot be distinguished from each other by their host plants. Additionally, for the Korean C. hexastigma and C. floribundae molecular data on the GenBank database used in the analysis of Cho et al. (2019), no divergence was observed in the 575 bp of the COI-tRNAleu- COII region, and only 1.45% (p -distance) or 1.46% (K2P distance) divergence was observed in&gt;690 bp of the mitochondrial 12S and 16S ribosomal RNA data. Therefore, I conclude that these two taxa are conspecific and can be synonymised as Cyamophila hexastigma (Horváth, 1899) = Cyamophila floribundae Cho &amp; Burckhardt, 2017, syn. nov.</p></div>	https://treatment.plazi.org/id/03E77267333B482CFF399C370A2B86AE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Inoue, Hiromitsu	Inoue, Hiromitsu (2025): Taxonomy and DNA barcoding of Cyamophila (Hemiptera: Psyllidae) from Japan, with the description of a new species. Zootaxa 5727 (1): 148-169, DOI: 10.11646/zootaxa.5727.1.10, URL: https://doi.org/10.11646/zootaxa.5727.1.10
03E77267333C4820FF399AD70E3387E2.text	03E77267333C4820FF399AD70E3387E2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cyamophila burckhardti Inoue 2025	<div><p>Cyamophila burckhardti sp. nov.</p><p>urn:lsid:zoobank.org:act: AD760C92-FC32-4DD8-8413-EED0415FA7FF</p><p>(Japanese name: Usuiro-mutsuboshi-kijirami)</p><p>(Figs 11–19, 27, 30, 33, 37–38, 43–44)</p><p>Cyamophila hexastigma sensu Inoue &amp; Yamauchi (2001: 55), nec Horváth (1899: 373).</p><p>Cyamophila sp.: Hayashi et al. (2011: 218); Hayashi &amp; Miyatake (2012: 28); Nozawa (2022: 90).</p><p>Description. Adult. Colouration. General colour (Figs 37, 43) light yellowish green with obscure light yellowish brown markings and longitudinal stripes on thorax; in overwintered individuals (Fig. 38), body colour chestnut brown to dark brown in general, with many cream-yellow coloured fine markings and stripes on head and thorax. Antenna yellow; apices of segments IV–VI and most of segment VII dark brown; segments VIII–X dark brown to black. Forewing membrane transparent in general, faintly yellowish along veins in the apical half of forewing in younger adults, strongly yellowish throughout the apical half of forewing in fully matured adults, with reduced and obscure pale brown spots at the location of the radular spinules in cells m 1, m 2, and cu 1 (Fig. 13); veins yellow in summer adults, becoming uniformly dark brown in overwintered adults. Apical tooth of paramere black.</p><p>Structure. Head (Fig. 11) strongly inclined downwards, 85–90° from longitudinal body axis in profile, slightly wider than thorax. Vertex nearly half as long as wide or slightly shorter. Genal processes conical, about 0.7 times as long as vertex, slightly divergent and subacute apically. Antenna long, 2.3–2.6 times as long as head width; longer terminal seta of segment X about 1.8–2.0 times as long as shorter seta, 0.5–0.6 times as long as segment X (Fig. 12).</p><p>Forewing (Fig. 13) oblong oval, about 2.2–2.3 times as long as wide, widest at around 2/3 from the base; membrane with dense surface spinules in all cells; spinule-free bands along veins rather narrow, gradually becoming narrower apically; fields of radular spinules as in Fig. 13; pterostigma well developed, more than 1/3 of the forewing length; Rs slightly sinuate, not curved towards costal margin apically; M 1+2 rather strongly arched at around the basal 1/3 and nearly straight thereafter, mostly very slightly curved towards costal margin apically; Cu 1a strongly arched around the basal 1/3–1/4 and nearly straight thereafter. Meracanthus moderate in size, subacute and slightly curved downwards apically; metatibia with prominent genual spine, with five apical sclerotised spurs arranged in 1+3+1; basal segment of metatarsus with a pair of sclerotised lateral spurs.</p><p>Male terminalia (Fig. 14) moderate in size. Proctiger slender, slightly curved caudad apically. Paramere stout, 0.8 times as long as proctiger, narrowest at base, gradually becoming wider and curved caudad towards the tip, truncated apically; anterior margin roundly projected cephalad around the apical 1/3; posteroapical corner strongly projected caudad; inner surface (Fig. 15) with many retrorse setae; inner apical tooth prominent, acute and projected cephalad. Distal aedeagal segment slightly longer than paramere; apex round and thickened, strongly hooked.</p><p>Female terminalia (Fig. 16) rather slender. Proctiger almost looks straight and very slightly curved at dorsal margin, not upturned apically, round at apex (Fig. 33). Subgenital plate with rather dense setae, acute at apex.</p><p>Fifth instar immature. Body (Figs 17, 44) light green, gently swollen dorsally. Antenna slender, 2.1–2.6 times as long as forewing pad, seven-segmented, with one apical rhinarium each on segments III and V, two on the middle of segment VII, with rather short capitate setae on the middle of segment III and apices of segments III–V (Fig. 27). Forewing pad oblong oval; outer margin with 8–9 long capitate setae and short simple setae in between (Fig. 30). Hindwing pad with two long capitate setae apically. Legs long, hairy, with many short simple setae and long capitate setae; tarsal arolia petiolate and fan-shaped (Fig. 18). Abdomen rounded apically, with many long capitate setae dorsally and many long simple setae ventrally. Caudal plate with many long capitate setae on dorsum and margin, with 4+4 truncated sectasetae on posterior margin. Anus located on ventral side. Outer circumanal pore ring (Fig. 19) relatively small, heart-shaped, strongly curved in front, comprising a single row of elongated pores; caudal margin of outer circumanal pore ring away from abdominal margin. Inner circumanal pore ring comprising a single row of small elongated pores.</p><p>Measurements (in mm): Adult (n = 5 males, 5 females): BL: 3.62–3.92; WL: 3.02–3.29; WW: 1.19–1.37; AL: 2.18–2.68; HW: 0.96–1.05; VW: 0.56–0.61; VL: 0.26–0.28; GL: 0.22–0.27; MP: 0.40–0.47; PL: 0.34–0.38; FP: 0.85–0.99. Fifth instar immature (n = 5): BL: 2.50–2.83; BW: 1.28–1.50; AL: 1.78–1.85; WPL: 0.70–0.85; CRW: 0.08–0.09.</p><p>Material examined. HOLOTYPE: male (dry-mounted; NARO), Japan, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=132.083&amp;materialsCitation.latitude=34.492" title="Search Plazi for locations around (long 132.083/lat 34.492)">Honshû</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=132.083&amp;materialsCitation.latitude=34.492" title="Search Plazi for locations around (long 132.083/lat 34.492)">Hiroshima-ken</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=132.083&amp;materialsCitation.latitude=34.492" title="Search Plazi for locations around (long 132.083/lat 34.492)">Hatsukaichishi</a>, Yoshiwa, Nakatsuya, 34.492º N, 132.083º E, 750 m, 3.vi.2022,on Cladrastis shikokiana, H. Inoue . PARATYPES: Honshû: 2 males, 1 female, Gunma-ken, Tone-gun, Kawaba-mura, Kawaba-yubara, Akakura-keikoku, 6.v.2004, N. Takahashi (dry-mounted; HIC) ; 7 males, 7 females, Nagano-ken, Matsumoto-shi, Azumi, Shimashimadani, 4.vi.2009, N. Takahashi (dry-mounted; HIC) ; 9 males, 7 females, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=137.744&amp;materialsCitation.latitude=36.145" title="Search Plazi for locations around (long 137.744/lat 36.145)">Nagano-ken</a>, Matsumoto-shi, Azuminagawado, 36.145º N, 137.744º E, 934 m, 26.vi.2021, on C. shikokiana, T. Matsuda (dry- and slide-mounted; HIC) ; 1 male, 6 females, 1 immature, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=138.499&amp;materialsCitation.latitude=36.978" title="Search Plazi for locations around (long 138.499/lat 36.978)">Nagano-ken</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=138.499&amp;materialsCitation.latitude=36.978" title="Search Plazi for locations around (long 138.499/lat 36.978)">Shimominochi-gun</a>, Sakae-mura, Shiratori, 36.978º N, 138.499º E, 287 m, 13.vi.2021, on Platyosprion platycarpum, T. Matsuda (dry- and slide-mounted; HIC) ; 1 male, Hiroshima-ken, Shôbara-shi, Takano-chô, Kenashi-yama, 950 m, 7.viii.2004, at light trap, T. Yamauchi (dry-mounted; HIC) ; 1 male, 1 female, Hiroshima-ken, Fukuyama-shi, Ryûzu-kyô, 13.v.2009, N. Takahashi (dry-mounted; HIC) ; 77 males, 87 females, 97 immatures, same data as holotype (dry- and slide-mounted, 99.5% and 70% ethanol; HIC, OMNH); 10 males, 11 females, same locality, 22.vii.2000, light trap, T. Yamauchi (dry- and slide-mounted; HIC); Kyûshû: 1 female, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=130.97&amp;materialsCitation.latitude=33.502" title="Search Plazi for locations around (long 130.97/lat 33.502)">Fukuoka-ken</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=130.97&amp;materialsCitation.latitude=33.502" title="Search Plazi for locations around (long 130.97/lat 33.502)">Miyako-gun</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=130.97&amp;materialsCitation.latitude=33.502" title="Search Plazi for locations around (long 130.97/lat 33.502)">Miyako-machi</a>, Saigawahobashira, near Notôge, 33.502º N, 130.970º E, 750–930 m, 18.xi.2000, H. Inoue, on Tsuga sieboldii (dry-mounted; HIC) ; 13 males, 12 females, same locality, 1.vi.2001, H. Inoue, on Aria japonica (dry- and slide-mounted; HIC) ; 5 males, 3 females, same data, 3.vi.2001 (dry-mounted; HIC); 2 males, 3 females, same data, 12.vi.2001 (dry- and slide-mounted, 99.5% ethanol; HIC); 1 male, same data, 21.vi.2001 (dry-mounted; HIC); 1 female, same locality, 7.iv.2012, N. Takahashi (dry-mounted; HIC); 3 females, same data, 7.v.2012 (dry-mounted; HIC); 1 female, same data, 4.vi.2012 (dry-mounted; HIC); 45 males, 47 females, same locality, 9.vi.2014, H. Inoue (dry- and slide-mounted, 99.5% ethanol; HIC); 1 female, same data, 20.v.2015 (dry-mounted; HIC); 80 males, 77 females, 4 immatures, Kumamoto-ken, Kamimashiki-gun, Yabemachi, Naidaijin-rindô, 7.vi.2001, H. Inoue (dry- and slide-mounted, 99.5% ethanol; HIC) ; 4 males, 5 females, Miyazaki-ken, Higashiusuki-gun, Shiiba-son, Shiiya-tôge, 7.vi.2001, H. Inoue (dry-mounted; HIC) .</p><p>Distribution. Japan (Honshû, Kyûshû).</p><p>Host plant. Cladrastis shikokiana (Makino) Makino; Platyosprion platycarpum (Maxim.) Maxim. (= Cladrastis platycarpa (Maxim.) Makino) ( Fabales: Fabaceae). These host plants were confirmed by the presence of immatures.</p><p>Etymology. This species is dedicated to and named after our most famous psyllidologist, Daniel Burckhardt, who is still very active at over 70 years of age.</p><p>Biology. Univoltine. The adults emerge in early June and leave the host plant within a short period. In summer and autumn, the adults are often observed on non-host plants, such as Aria japonica Decne. ( Rosales: Rosaceae) and Tsuga sieboldii Carrière ( Pinales: Pinaceae), which grow on the ridges of mountainous areas. Presumably, adults overwinter on evergreen shelter plants, such as T. sieboldii .</p><p>Comments. The diagnoses for the identification of Japanese Cyamophila species are presented in Table 2. The population corresponding to the new species was first collected using a light trap at the type locality (Fig. 47), as reported by Inoue &amp; Yamauchi (2001). The material was provisionally recorded as ‘ C. hexastigma ’, but the absence of prominent spots on the forewing posterior margin required reconsideration for identification. This species was therefore subsequently reported by several authors as ‘ Cyamophila sp. ’ (see synonymic list).</p><p>Whereas C. shikokiana is endemic to Japan, P. platycarpum is found also in China (eFloras 2008). Therefore, C. burckhardti may also occur in China; however, psyllids corresponding to or similar to this species have not yet been recorded.</p></div>	https://treatment.plazi.org/id/03E77267333C4820FF399AD70E3387E2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Inoue, Hiromitsu	Inoue, Hiromitsu (2025): Taxonomy and DNA barcoding of Cyamophila (Hemiptera: Psyllidae) from Japan, with the description of a new species. Zootaxa 5727 (1): 148-169, DOI: 10.11646/zootaxa.5727.1.10, URL: https://doi.org/10.11646/zootaxa.5727.1.10
03E7726733304827FF399B070C0A8238.text	03E7726733304827FF399B070C0A8238.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cyamophila willieti (Wu 1932)	<div><p>Cyamophila willieti (Wu, 1932)</p><p>(Japanese name: Yatsuboshi-kijirami)</p><p>(Figs 20–25, 28, 31, 34, 39–40, 45–46)</p><p>Psylla willieti Wu, 1932: 71 . Type locality: China (Beijing).</p><p>Cyamophila willieti; Conci &amp; Tamanini (1988: 171).</p><p>Description. Adult. Colouration. General colour (Figs 39, 45) light yellowish green with obscure light yellowish brown markings on thorax; in overwintered individuals (Fig. 40), body colour chestnut brown to dark brown in general, with many cream-yellow coloured fine markings and stripes on head and thorax. Antenna yellow; apices of segments IV and V, apical half of segment VI and most of segment VII dark brown; segments VIII–X entirely dark brown to black. Forewing membrane transparent, colourless, with prominent dark brown spots at the location of the radular spinules in cells r 2, m 1, m 2, and cu 1 (Fig. 22); veins yellow in summer adults, becoming uniformly dark brown in overwintered adults. Apical tooth of paramere black.</p><p>Structure. Head (Fig. 20) strongly inclined downwards, 85–90° from longitudinal body axis in profile, slightly wider than thorax. Vertex shorter than the half width. Genal processes conical, about 0.8 times as long as vertex, slightly divergent, subacute or slightly rounded apically. Antenna long, 2.1–2.2 times as long as head width; longer terminal seta of segment X 1.8–2.2 times as long as shorter seta, about half as long as segment X (Fig. 21).</p><p>Forewing (Fig. 22) oblong oval, about 2.2–2.3 times as long as wide, widest at around 2/3 from the base; membrane with dense surface spinules in all cells; spinule-free bands along veins broad; fields of radular spinules as in Fig. 22; pterostigma broad, more than 1/3 of the forewing length; Rs rather strongly sinuate, slightly curved towards costal margin apically; M 1+2 strongly arched at around the basal 1/3 and slightly or rather strongly curved towards costal margin apically; Cu 1a strongly arched at approximately basal 1/4 and nearly straight thereafter. Meracanthus moderate in size, subacute and slightly curved downwards apically; metatibia with prominent genual spine, with five apical sclerotised spurs arranged in 1+3+1; basal segment of metatarsus with a pair of sclerotised lateral spurs.</p><p>Male terminalia (Fig. 23) moderate in size. Proctiger slender, slightly curved caudad apically. Paramere slender, 0.8 times as long as proctiger, almost constant in width throughout, truncated apically; anterior margin angularly projected cephalad around the apical 1/4; posteroapical corner weakly projected caudad; inner surface (Fig. 24) with many retrorse setae; inner apical tooth rather prominent, acute and projected cephalad. Distal aedeagal segment slightly longer than paramere; apex round and thickened, strongly hooked.</p><p>Female terminalia (Fig. 25) stout. Proctiger stout, gently curved and slightly sinuate at dorsal margin, very slightly upturned apically, round at apex (Fig. 34). Subgenital plate with sparse setae, acute at apex.</p><p>Fifth instar immature. Body (Fig. 46) light green, gently swollen dorsally. Antenna slender, 2.0–2.2 times as long as forewing pad, seven-segmented, with one apical rhinarium on each of segments III and V, two on the middle of segment VII, with rather short capitate setae on the middle of segment III and apices of segments III–V (Fig. 28). Forewing pad oblong oval; outer margin with 8–9 long capitate setae and short capitate setae in between (Fig. 31). Hindwing pad with two long capitate setae apically. Legs long, hairy, with many short simple setae and long capitate setae. Abdomen rounded apically, with many long capitate setae dorsally and many long simple setae ventrally. Caudal plate with many long capitate setae on dorsum and margin, with 4+4 truncated sectasetae on posterior margin. Anus located on ventral side. Outer circumanal pore ring relatively small, heart-shaped, strongly curved in front, comprising a single row of elongated pores. Inner circumanal pore ring comprising a single row of small elongated pores.</p><p>Measurements (in mm): Adult (n = 5 males, 5 females): BL: 3.62–4.16; WL: 2.99–3.40; WW: 1.24–1.44; AL: 2.10–2.90; HW: 0.97–1.06; VW: 0.58–0.63; VL: 0.27–0.28; GL: 0.19–0.23; MP: 0.43–0.45; PL: 0.33–0.36; FP: 0.83–0.93. Fifth instar immature (n = 5): BL: 2.05–2.90; BW: 1.30–1.60; AL: 1.43–1.63; WPL: 0.73–0.83; CRW: 0.10–0.11.</p><p>Material examined. Honshû: 3 males, 5 females, Ibaraki-ken, Tsukuba-shi, Fujimoto, 36.051º N, 140.100º E, 20 m, 30.v.2003, on Styphnolobium japonicum, H. Inoue (slide-mounted, 99.5% ethanol; HIC) ; 25 males, 36 females, same data, 2.vi.2003 (dry-mounted; HIC, OMNH); 2 females, same data, 3.vi.2003 (dry-mounted; HIC); 4 males, 9 females, same data, 15.vi.2003 (dry-mounted; HIC); 3 males, 9 females, 10 immatures, same data, 15.iv.2004 (dry- and slide-mounted; HIC) .</p><p>Distribution. Japan (Honshû; new distributional record); China, Korea (Wu 1932, as P. willieti; Kwon 1983, as C. hexastigma)</p><p>Host plant. Styphnolobium japonicum (L.) Schott (= Sophora japonica L.) ( Fabales: Fabaceae) (Conci &amp; Tamanini 1988). The host plant was confirmed by the presence of immatures.</p><p>Biology. Probably univoltine. In April, overwintered adults gather on host plants to mate and lay eggs on their shoots. The new adults emerge in June. Adults are thought to leave the host plant after the emergence and hibernate on evergreen shelter plants; however, the details of where they stay throughout the winter are unclear.</p><p>Comments. This species is newly recorded from Japan. The host plant, the Japanese pagoda tree, S. japonicum is native to China and was introduced into Japan, where it has been planted as a garden and roadside tree (Ohashi 2016). Therefore, C. willieti may be a pest of S. japonicum in Japan. No other species of Styphnolobium are native to Japan, C. willieti is also likely an introduced species.</p></div>	https://treatment.plazi.org/id/03E7726733304827FF399B070C0A8238	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Inoue, Hiromitsu	Inoue, Hiromitsu (2025): Taxonomy and DNA barcoding of Cyamophila (Hemiptera: Psyllidae) from Japan, with the description of a new species. Zootaxa 5727 (1): 148-169, DOI: 10.11646/zootaxa.5727.1.10, URL: https://doi.org/10.11646/zootaxa.5727.1.10
