taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
03E587C7FF9CCD0FFF73FA7FFC47F834.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17323401/files/figure.png	https://doi.org/10.5281/zenodo.17323401	FIGURE 2. Representatives of the three European phenotypes of Natrix astreptophora astreptophora. Regular phenotype: (A) adult, Granada, (B) juvenile, Ávila, (C) hatchling, Navarra, (D) hatchling, Ronda, all Spain (photos: Gabriel Martínez del Mármol). Northern phenotype: (E) adult, Santander, (F) juvenile, north of Burgos (both photos: Gabriel Martínez del Mármol), (G) hatchling, Guitiriz, Lugo (photo: Rafael Vázquez Graña), (H) dark hatchling, west of Asturias (photo: Miguel Grazziani), all Spain. Western phenotype: (I) adult, Porto (photo:Alberto González Estremera), (J) young adult, Porto (photo: Gabriel Martínez del Mármol), (K) hatchling, Braga, all Portugal, (L) juvenile, Coiros, Spain (last two photos: Rafael Vázquez Graña).	FIGURE 2. Representatives of the three European phenotypes of Natrix astreptophora astreptophora. Regular phenotype: (A) adult, Granada, (B) juvenile, Ávila, (C) hatchling, Navarra, (D) hatchling, Ronda, all Spain (photos: Gabriel Martínez del Mármol). Northern phenotype: (E) adult, Santander, (F) juvenile, north of Burgos (both photos: Gabriel Martínez del Mármol), (G) hatchling, Guitiriz, Lugo (photo: Rafael Vázquez Graña), (H) dark hatchling, west of Asturias (photo: Miguel Grazziani), all Spain. Western phenotype: (I) adult, Porto (photo:Alberto González Estremera), (J) young adult, Porto (photo: Gabriel Martínez del Mármol), (K) hatchling, Braga, all Portugal, (L) juvenile, Coiros, Spain (last two photos: Rafael Vázquez Graña).	2025-09-16	Mármol, Gabriel Martínez Del;Geniez, Philippe;Neumann, Adrian;Glaw, Frank;Fritz, Uwe		Zenodo	biologists	Mármol, Gabriel Martínez Del;Geniez, Philippe;Neumann, Adrian;Glaw, Frank;Fritz, Uwe			
03E587C7FF9CCD0FFF73FA7FFC47F834.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17323403/files/figure.png	https://doi.org/10.5281/zenodo.17323403	FIGURE 3. Distribution of subspecies of Natrix astreptophora. (A) Approximate range of the regular phenotype of N. a. astreptophora; the broken lines indicate the approximate distribution borders for the northern (B) and western (C) phenotypes; in western Galicia all phenotypes co-occur.Asterisks are type localities (N. a. astreptophora has a ‘composed’ type locality of two sites, see Fritz & Schmidtler 2020; type locality of N. a. algerica is tentatively identified with Algiers). Data for nominotypical subspecies include 8299 records from GBIF (https://www.gbif.org) as available on 23 April 2025. Black circles represent melanistic individuals. Map base: Esri and OpenStreetMap contributors.	FIGURE 3. Distribution of subspecies of Natrix astreptophora. (A) Approximate range of the regular phenotype of N. a. astreptophora; the broken lines indicate the approximate distribution borders for the northern (B) and western (C) phenotypes; in western Galicia all phenotypes co-occur.Asterisks are type localities (N. a. astreptophora has a ‘composed’ type locality of two sites, see Fritz & Schmidtler 2020; type locality of N. a. algerica is tentatively identified with Algiers). Data for nominotypical subspecies include 8299 records from GBIF (https://www.gbif.org) as available on 23 April 2025. Black circles represent melanistic individuals. Map base: Esri and OpenStreetMap contributors.	2025-09-16	Mármol, Gabriel Martínez Del;Geniez, Philippe;Neumann, Adrian;Glaw, Frank;Fritz, Uwe		Zenodo	biologists	Mármol, Gabriel Martínez Del;Geniez, Philippe;Neumann, Adrian;Glaw, Frank;Fritz, Uwe			
03E587C7FF9CCD0FFF73FA7FFC47F834.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17323405/files/figure.png	https://doi.org/10.5281/zenodo.17323405	FIGURE 4. Dorsal pattern of different phenotypes of Natrix astreptophora. Nominotypical subspecies: (A) regular phenotype, Granada, Spain, (B) northern pale phenotype, Santander, Spain, (C) western phenotype, Porto, Portugal, (D) N. a. algerica, Cap Serrat, Tunisia, (E) N. a. soumiafahdae subsp. nov., Chefchaouen, Morocco (all photos: Gabriel Martínez del Mármol, except D: Martina Kiselová).	FIGURE 4. Dorsal pattern of different phenotypes of Natrix astreptophora. Nominotypical subspecies: (A) regular phenotype, Granada, Spain, (B) northern pale phenotype, Santander, Spain, (C) western phenotype, Porto, Portugal, (D) N. a. algerica, Cap Serrat, Tunisia, (E) N. a. soumiafahdae subsp. nov., Chefchaouen, Morocco (all photos: Gabriel Martínez del Mármol, except D: Martina Kiselová).	2025-09-16	Mármol, Gabriel Martínez Del;Geniez, Philippe;Neumann, Adrian;Glaw, Frank;Fritz, Uwe		Zenodo	biologists	Mármol, Gabriel Martínez Del;Geniez, Philippe;Neumann, Adrian;Glaw, Frank;Fritz, Uwe			
03E587C7FF9CCD0FFF73FA7FFC47F834.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17323409/files/figure.png	https://doi.org/10.5281/zenodo.17323409	FIGURE 5. Different stages of melanism in Natrix astreptophora astreptophora from Spain. (A) Adult with incipient melanism, note the marron iris, Padrenda, Ourense, (B) stronger melanistic juvenile, Ribeira, La Coruña, (C) greyish adult, Santa María de Oia, Pontevedra (photos A–C: Rafael Vázquez Graña), (D) hatchling, Baiona, Pontevedra (photo: Gabriel Martínez del Mármol), (E) nearly black adult with whitish elements on head and neck, Guipúzcoa, Basque Country (photo: Ander Izagirre Egaña), (F) completely black snake, Álava, Basque Country (photo: Alberto González Estremera).	FIGURE 5. Different stages of melanism in Natrix astreptophora astreptophora from Spain. (A) Adult with incipient melanism, note the marron iris, Padrenda, Ourense, (B) stronger melanistic juvenile, Ribeira, La Coruña, (C) greyish adult, Santa María de Oia, Pontevedra (photos A–C: Rafael Vázquez Graña), (D) hatchling, Baiona, Pontevedra (photo: Gabriel Martínez del Mármol), (E) nearly black adult with whitish elements on head and neck, Guipúzcoa, Basque Country (photo: Ander Izagirre Egaña), (F) completely black snake, Álava, Basque Country (photo: Alberto González Estremera).	2025-09-16	Mármol, Gabriel Martínez Del;Geniez, Philippe;Neumann, Adrian;Glaw, Frank;Fritz, Uwe		Zenodo	biologists	Mármol, Gabriel Martínez Del;Geniez, Philippe;Neumann, Adrian;Glaw, Frank;Fritz, Uwe			
03E587C7FF99CD00FF73FF3CFD9CFBAC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17323411/files/figure.png	https://doi.org/10.5281/zenodo.17323411	FIGURE 6. Natrix astreptophora algerica, (A) hatchling, Tabarka, Tunisia (photo: Mohsen Kalboussi), (B) hatchling, Djurdjura mountains, Algeria (photo: Olivier Peyre), (C) adult, Cap Serrat, Tunisia (photo: Martina Kiselová), (D) adult, El Milia, Algeria (photo: Krikra Oussama).	FIGURE 6. Natrix astreptophora algerica, (A) hatchling, Tabarka, Tunisia (photo: Mohsen Kalboussi), (B) hatchling, Djurdjura mountains, Algeria (photo: Olivier Peyre), (C) adult, Cap Serrat, Tunisia (photo: Martina Kiselová), (D) adult, El Milia, Algeria (photo: Krikra Oussama).	2025-09-16	Mármol, Gabriel Martínez Del;Geniez, Philippe;Neumann, Adrian;Glaw, Frank;Fritz, Uwe		Zenodo	biologists	Mármol, Gabriel Martínez Del;Geniez, Philippe;Neumann, Adrian;Glaw, Frank;Fritz, Uwe			
03E587C7FF99CD00FF73FF3CFD9CFBAC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17323413/files/figure.png	https://doi.org/10.5281/zenodo.17323413	FIGURE 7. Differences in hatchling coloration of the three Natrix astreptophora subspecies. (A–C) N. a. astreptophora, Cazorla, Spain (photos: Gabriel Martínez del Mármol), (D–F) N. a. algerica, Djurdjura mountains, Algeria (photos: Olivier Peyre), (G–I) N. a. soumiafahdae subsp. nov., Moulay Abdeslam, Morocco (photos: Gabriel Martínez del Mármol). Note that the light collar in N. a. soumiafahdae subsp. nov. is closed (continuous). Moreover, the upper postocular scale has the same color as the supraocular scale in N. a. astreptophora, and is whitish in the two African subspecies, i.e., it has the same color as the two other postocular scales.	FIGURE 7. Differences in hatchling coloration of the three Natrix astreptophora subspecies. (A–C) N. a. astreptophora, Cazorla, Spain (photos: Gabriel Martínez del Mármol), (D–F) N. a. algerica, Djurdjura mountains, Algeria (photos: Olivier Peyre), (G–I) N. a. soumiafahdae subsp. nov., Moulay Abdeslam, Morocco (photos: Gabriel Martínez del Mármol). Note that the light collar in N. a. soumiafahdae subsp. nov. is closed (continuous). Moreover, the upper postocular scale has the same color as the supraocular scale in N. a. astreptophora, and is whitish in the two African subspecies, i.e., it has the same color as the two other postocular scales.	2025-09-16	Mármol, Gabriel Martínez Del;Geniez, Philippe;Neumann, Adrian;Glaw, Frank;Fritz, Uwe		Zenodo	biologists	Mármol, Gabriel Martínez Del;Geniez, Philippe;Neumann, Adrian;Glaw, Frank;Fritz, Uwe			
03E587C7FF97CD07FF73F933FE7EF846.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17323415/files/figure.png	https://doi.org/10.5281/zenodo.17323415	FIGURE 8. Holotypes of (A) Tropidonotus natrixalgericus Hecht, 1930(ZMB 19636),“Südalgier” and(B) Natrix astreptophora soumiafahdae subsp. nov. (NMW 32908), Middle Atlas, Morocco (photos: Adrian Neumann). Note the dorsally interrupted vs. closed light collar and the differences in body pattern.	FIGURE 8. Holotypes of (A) Tropidonotus natrixalgericus Hecht, 1930(ZMB 19636),“Südalgier” and(B) Natrix astreptophora soumiafahdae subsp. nov. (NMW 32908), Middle Atlas, Morocco (photos: Adrian Neumann). Note the dorsally interrupted vs. closed light collar and the differences in body pattern.	2025-09-16	Mármol, Gabriel Martínez Del;Geniez, Philippe;Neumann, Adrian;Glaw, Frank;Fritz, Uwe		Zenodo	biologists	Mármol, Gabriel Martínez Del;Geniez, Philippe;Neumann, Adrian;Glaw, Frank;Fritz, Uwe			
03E587C7FF97CD07FF73F933FE7EF846.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17323417/files/figure.png	https://doi.org/10.5281/zenodo.17323417	FIGURE 9. Paratypes of Natrix astreptophora soumiafahdae subsp. nov., large adults. (A) Male BEV.6432 and (B) female BEV.6431, Oulmès, Morocco. Note that only the female (SVL 106 cm; tail incomplete, estimated TL 130 cm) has completely lost the light collar, as well as the dark nuchal and dorsolateral markings. The male (SVL 75 cm, TL 98 cm) still shows dark nuchal and dorsolateral markings and patterned supralabial scales; the faded collar is closed (photos: Jean Nicolas).	FIGURE 9. Paratypes of Natrix astreptophora soumiafahdae subsp. nov., large adults. (A) Male BEV.6432 and (B) female BEV.6431, Oulmès, Morocco. Note that only the female (SVL 106 cm; tail incomplete, estimated TL 130 cm) has completely lost the light collar, as well as the dark nuchal and dorsolateral markings. The male (SVL 75 cm, TL 98 cm) still shows dark nuchal and dorsolateral markings and patterned supralabial scales; the faded collar is closed (photos: Jean Nicolas).	2025-09-16	Mármol, Gabriel Martínez Del;Geniez, Philippe;Neumann, Adrian;Glaw, Frank;Fritz, Uwe		Zenodo	biologists	Mármol, Gabriel Martínez Del;Geniez, Philippe;Neumann, Adrian;Glaw, Frank;Fritz, Uwe			
03E587C7FF97CD07FF73F933FE7EF846.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17323419/files/figure.png	https://doi.org/10.5281/zenodo.17323419	FIGURE 10. Natrix astreptophora soumiafahdae subsp. nov., live individuals. (A) Young adult, Chefchaouen, NW Morocco (photo: Gabriel Martínez del Mármol). Note the whitish-yellow color of the closed collar and the contrasting black color of pileus, nuchal, dorsal, and lateral markings. (B, C) large megacephalic female, Souk El Arbaa Beni Hassan, Morocco, with uniform coloration (photos: Mario Schweiger).	FIGURE 10. Natrix astreptophora soumiafahdae subsp. nov., live individuals. (A) Young adult, Chefchaouen, NW Morocco (photo: Gabriel Martínez del Mármol). Note the whitish-yellow color of the closed collar and the contrasting black color of pileus, nuchal, dorsal, and lateral markings. (B, C) large megacephalic female, Souk El Arbaa Beni Hassan, Morocco, with uniform coloration (photos: Mario Schweiger).	2025-09-16	Mármol, Gabriel Martínez Del;Geniez, Philippe;Neumann, Adrian;Glaw, Frank;Fritz, Uwe		Zenodo	biologists	Mármol, Gabriel Martínez Del;Geniez, Philippe;Neumann, Adrian;Glaw, Frank;Fritz, Uwe			
03E587C7FF97CD07FF73F933FE7EF846.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17323421/files/figure.png	https://doi.org/10.5281/zenodo.17323421	FIGURE 11. Habitats of Natrix astreptophora soumiafahdae subsp. nov. Rif Mountains: (A) Bouhachem National Park, (B) Moulay Abdeslam, (C) Bab Taza; Middle Atlas: (D) Ifrane (photos A, B, D: Gabriel Martínez del Mármol; photo C: Frank Glaw).	FIGURE 11. Habitats of Natrix astreptophora soumiafahdae subsp. nov. Rif Mountains: (A) Bouhachem National Park, (B) Moulay Abdeslam, (C) Bab Taza; Middle Atlas: (D) Ifrane (photos A, B, D: Gabriel Martínez del Mármol; photo C: Frank Glaw).	2025-09-16	Mármol, Gabriel Martínez Del;Geniez, Philippe;Neumann, Adrian;Glaw, Frank;Fritz, Uwe		Zenodo	biologists	Mármol, Gabriel Martínez Del;Geniez, Philippe;Neumann, Adrian;Glaw, Frank;Fritz, Uwe			
03E587C7FF97CD07FF73F933FE7EF846.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17323403/files/figure.png	https://doi.org/10.5281/zenodo.17323403	FIGURE 3. Distribution of subspecies of Natrix astreptophora. (A) Approximate range of the regular phenotype of N. a. astreptophora; the broken lines indicate the approximate distribution borders for the northern (B) and western (C) phenotypes; in western Galicia all phenotypes co-occur.Asterisks are type localities (N. a. astreptophora has a ‘composed’ type locality of two sites, see Fritz & Schmidtler 2020; type locality of N. a. algerica is tentatively identified with Algiers). Data for nominotypical subspecies include 8299 records from GBIF (https://www.gbif.org) as available on 23 April 2025. Black circles represent melanistic individuals. Map base: Esri and OpenStreetMap contributors.	FIGURE 3. Distribution of subspecies of Natrix astreptophora. (A) Approximate range of the regular phenotype of N. a. astreptophora; the broken lines indicate the approximate distribution borders for the northern (B) and western (C) phenotypes; in western Galicia all phenotypes co-occur.Asterisks are type localities (N. a. astreptophora has a ‘composed’ type locality of two sites, see Fritz & Schmidtler 2020; type locality of N. a. algerica is tentatively identified with Algiers). Data for nominotypical subspecies include 8299 records from GBIF (https://www.gbif.org) as available on 23 April 2025. Black circles represent melanistic individuals. Map base: Esri and OpenStreetMap contributors.	2025-09-16	Mármol, Gabriel Martínez Del;Geniez, Philippe;Neumann, Adrian;Glaw, Frank;Fritz, Uwe		Zenodo	biologists	Mármol, Gabriel Martínez Del;Geniez, Philippe;Neumann, Adrian;Glaw, Frank;Fritz, Uwe			
03E587C7FF8ACD1DFF73FF3CFCB2FB17.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17323411/files/figure.png	https://doi.org/10.5281/zenodo.17323411	FIGURE 6. Natrix astreptophora algerica, (A) hatchling, Tabarka, Tunisia (photo: Mohsen Kalboussi), (B) hatchling, Djurdjura mountains, Algeria (photo: Olivier Peyre), (C) adult, Cap Serrat, Tunisia (photo: Martina Kiselová), (D) adult, El Milia, Algeria (photo: Krikra Oussama).	FIGURE 6. Natrix astreptophora algerica, (A) hatchling, Tabarka, Tunisia (photo: Mohsen Kalboussi), (B) hatchling, Djurdjura mountains, Algeria (photo: Olivier Peyre), (C) adult, Cap Serrat, Tunisia (photo: Martina Kiselová), (D) adult, El Milia, Algeria (photo: Krikra Oussama).	2025-09-16	Mármol, Gabriel Martínez Del;Geniez, Philippe;Neumann, Adrian;Glaw, Frank;Fritz, Uwe		Zenodo	biologists	Mármol, Gabriel Martínez Del;Geniez, Philippe;Neumann, Adrian;Glaw, Frank;Fritz, Uwe			
03E587C7FF8ACD1DFF73FF3CFCB2FB17.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17323413/files/figure.png	https://doi.org/10.5281/zenodo.17323413	FIGURE 7. Differences in hatchling coloration of the three Natrix astreptophora subspecies. (A–C) N. a. astreptophora, Cazorla, Spain (photos: Gabriel Martínez del Mármol), (D–F) N. a. algerica, Djurdjura mountains, Algeria (photos: Olivier Peyre), (G–I) N. a. soumiafahdae subsp. nov., Moulay Abdeslam, Morocco (photos: Gabriel Martínez del Mármol). Note that the light collar in N. a. soumiafahdae subsp. nov. is closed (continuous). Moreover, the upper postocular scale has the same color as the supraocular scale in N. a. astreptophora, and is whitish in the two African subspecies, i.e., it has the same color as the two other postocular scales.	FIGURE 7. Differences in hatchling coloration of the three Natrix astreptophora subspecies. (A–C) N. a. astreptophora, Cazorla, Spain (photos: Gabriel Martínez del Mármol), (D–F) N. a. algerica, Djurdjura mountains, Algeria (photos: Olivier Peyre), (G–I) N. a. soumiafahdae subsp. nov., Moulay Abdeslam, Morocco (photos: Gabriel Martínez del Mármol). Note that the light collar in N. a. soumiafahdae subsp. nov. is closed (continuous). Moreover, the upper postocular scale has the same color as the supraocular scale in N. a. astreptophora, and is whitish in the two African subspecies, i.e., it has the same color as the two other postocular scales.	2025-09-16	Mármol, Gabriel Martínez Del;Geniez, Philippe;Neumann, Adrian;Glaw, Frank;Fritz, Uwe		Zenodo	biologists	Mármol, Gabriel Martínez Del;Geniez, Philippe;Neumann, Adrian;Glaw, Frank;Fritz, Uwe			
03E587C7FF8ACD1DFF73FF3CFCB2FB17.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17323415/files/figure.png	https://doi.org/10.5281/zenodo.17323415	FIGURE 8. Holotypes of (A) Tropidonotus natrixalgericus Hecht, 1930(ZMB 19636),“Südalgier” and(B) Natrix astreptophora soumiafahdae subsp. nov. (NMW 32908), Middle Atlas, Morocco (photos: Adrian Neumann). Note the dorsally interrupted vs. closed light collar and the differences in body pattern.	FIGURE 8. Holotypes of (A) Tropidonotus natrixalgericus Hecht, 1930(ZMB 19636),“Südalgier” and(B) Natrix astreptophora soumiafahdae subsp. nov. (NMW 32908), Middle Atlas, Morocco (photos: Adrian Neumann). Note the dorsally interrupted vs. closed light collar and the differences in body pattern.	2025-09-16	Mármol, Gabriel Martínez Del;Geniez, Philippe;Neumann, Adrian;Glaw, Frank;Fritz, Uwe		Zenodo	biologists	Mármol, Gabriel Martínez Del;Geniez, Philippe;Neumann, Adrian;Glaw, Frank;Fritz, Uwe			
03E587C7FF8ACD1CFF73FB58FD34FE00.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17323419/files/figure.png	https://doi.org/10.5281/zenodo.17323419	FIGURE 10. Natrix astreptophora soumiafahdae subsp. nov., live individuals. (A) Young adult, Chefchaouen, NW Morocco (photo: Gabriel Martínez del Mármol). Note the whitish-yellow color of the closed collar and the contrasting black color of pileus, nuchal, dorsal, and lateral markings. (B, C) large megacephalic female, Souk El Arbaa Beni Hassan, Morocco, with uniform coloration (photos: Mario Schweiger).	FIGURE 10. Natrix astreptophora soumiafahdae subsp. nov., live individuals. (A) Young adult, Chefchaouen, NW Morocco (photo: Gabriel Martínez del Mármol). Note the whitish-yellow color of the closed collar and the contrasting black color of pileus, nuchal, dorsal, and lateral markings. (B, C) large megacephalic female, Souk El Arbaa Beni Hassan, Morocco, with uniform coloration (photos: Mario Schweiger).	2025-09-16	Mármol, Gabriel Martínez Del;Geniez, Philippe;Neumann, Adrian;Glaw, Frank;Fritz, Uwe		Zenodo	biologists	Mármol, Gabriel Martínez Del;Geniez, Philippe;Neumann, Adrian;Glaw, Frank;Fritz, Uwe			
03E587C7FF8ACD1CFF73FB58FD34FE00.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17323413/files/figure.png	https://doi.org/10.5281/zenodo.17323413	FIGURE 7. Differences in hatchling coloration of the three Natrix astreptophora subspecies. (A–C) N. a. astreptophora, Cazorla, Spain (photos: Gabriel Martínez del Mármol), (D–F) N. a. algerica, Djurdjura mountains, Algeria (photos: Olivier Peyre), (G–I) N. a. soumiafahdae subsp. nov., Moulay Abdeslam, Morocco (photos: Gabriel Martínez del Mármol). Note that the light collar in N. a. soumiafahdae subsp. nov. is closed (continuous). Moreover, the upper postocular scale has the same color as the supraocular scale in N. a. astreptophora, and is whitish in the two African subspecies, i.e., it has the same color as the two other postocular scales.	FIGURE 7. Differences in hatchling coloration of the three Natrix astreptophora subspecies. (A–C) N. a. astreptophora, Cazorla, Spain (photos: Gabriel Martínez del Mármol), (D–F) N. a. algerica, Djurdjura mountains, Algeria (photos: Olivier Peyre), (G–I) N. a. soumiafahdae subsp. nov., Moulay Abdeslam, Morocco (photos: Gabriel Martínez del Mármol). Note that the light collar in N. a. soumiafahdae subsp. nov. is closed (continuous). Moreover, the upper postocular scale has the same color as the supraocular scale in N. a. astreptophora, and is whitish in the two African subspecies, i.e., it has the same color as the two other postocular scales.	2025-09-16	Mármol, Gabriel Martínez Del;Geniez, Philippe;Neumann, Adrian;Glaw, Frank;Fritz, Uwe		Zenodo	biologists	Mármol, Gabriel Martínez Del;Geniez, Philippe;Neumann, Adrian;Glaw, Frank;Fritz, Uwe			
03E587C7FF8ACD1CFF73FB58FD34FE00.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17323399/files/figure.png	https://doi.org/10.5281/zenodo.17323399	FIGURE 1. Scalation and coloration traits on the pileus and body of a juvenile Natrix astreptophora from Moulay Abdeslam, Morocco, in dorsal (A) and lateral (B) view: (a) upper supraocular scale, (b) light spot on parietal scale, (c) light nuchal collar, (d) black nuchal marking, (e) supralabial scales, (f) loreal scale, (g) black dorsolateral spots, (h) black lateral and ventrolateral spots. Note the bicolored pileus (photos: Gabriel Martínez del Mármol).	FIGURE 1. Scalation and coloration traits on the pileus and body of a juvenile Natrix astreptophora from Moulay Abdeslam, Morocco, in dorsal (A) and lateral (B) view: (a) upper supraocular scale, (b) light spot on parietal scale, (c) light nuchal collar, (d) black nuchal marking, (e) supralabial scales, (f) loreal scale, (g) black dorsolateral spots, (h) black lateral and ventrolateral spots. Note the bicolored pileus (photos: Gabriel Martínez del Mármol).	2025-09-16	Mármol, Gabriel Martínez Del;Geniez, Philippe;Neumann, Adrian;Glaw, Frank;Fritz, Uwe		Zenodo	biologists	Mármol, Gabriel Martínez Del;Geniez, Philippe;Neumann, Adrian;Glaw, Frank;Fritz, Uwe			
03E587C7FF8ACD1CFF73FB58FD34FE00.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17323415/files/figure.png	https://doi.org/10.5281/zenodo.17323415	FIGURE 8. Holotypes of (A) Tropidonotus natrixalgericus Hecht, 1930(ZMB 19636),“Südalgier” and(B) Natrix astreptophora soumiafahdae subsp. nov. (NMW 32908), Middle Atlas, Morocco (photos: Adrian Neumann). Note the dorsally interrupted vs. closed light collar and the differences in body pattern.	FIGURE 8. Holotypes of (A) Tropidonotus natrixalgericus Hecht, 1930(ZMB 19636),“Südalgier” and(B) Natrix astreptophora soumiafahdae subsp. nov. (NMW 32908), Middle Atlas, Morocco (photos: Adrian Neumann). Note the dorsally interrupted vs. closed light collar and the differences in body pattern.	2025-09-16	Mármol, Gabriel Martínez Del;Geniez, Philippe;Neumann, Adrian;Glaw, Frank;Fritz, Uwe		Zenodo	biologists	Mármol, Gabriel Martínez Del;Geniez, Philippe;Neumann, Adrian;Glaw, Frank;Fritz, Uwe			
03E587C7FF8ACD1CFF73FB58FD34FE00.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17323417/files/figure.png	https://doi.org/10.5281/zenodo.17323417	FIGURE 9. Paratypes of Natrix astreptophora soumiafahdae subsp. nov., large adults. (A) Male BEV.6432 and (B) female BEV.6431, Oulmès, Morocco. Note that only the female (SVL 106 cm; tail incomplete, estimated TL 130 cm) has completely lost the light collar, as well as the dark nuchal and dorsolateral markings. The male (SVL 75 cm, TL 98 cm) still shows dark nuchal and dorsolateral markings and patterned supralabial scales; the faded collar is closed (photos: Jean Nicolas).	FIGURE 9. Paratypes of Natrix astreptophora soumiafahdae subsp. nov., large adults. (A) Male BEV.6432 and (B) female BEV.6431, Oulmès, Morocco. Note that only the female (SVL 106 cm; tail incomplete, estimated TL 130 cm) has completely lost the light collar, as well as the dark nuchal and dorsolateral markings. The male (SVL 75 cm, TL 98 cm) still shows dark nuchal and dorsolateral markings and patterned supralabial scales; the faded collar is closed (photos: Jean Nicolas).	2025-09-16	Mármol, Gabriel Martínez Del;Geniez, Philippe;Neumann, Adrian;Glaw, Frank;Fritz, Uwe		Zenodo	biologists	Mármol, Gabriel Martínez Del;Geniez, Philippe;Neumann, Adrian;Glaw, Frank;Fritz, Uwe			
03E587C7FF8BCD1FFF73FC93FC72FBE4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17323403/files/figure.png	https://doi.org/10.5281/zenodo.17323403	FIGURE 3. Distribution of subspecies of Natrix astreptophora. (A) Approximate range of the regular phenotype of N. a. astreptophora; the broken lines indicate the approximate distribution borders for the northern (B) and western (C) phenotypes; in western Galicia all phenotypes co-occur.Asterisks are type localities (N. a. astreptophora has a ‘composed’ type locality of two sites, see Fritz & Schmidtler 2020; type locality of N. a. algerica is tentatively identified with Algiers). Data for nominotypical subspecies include 8299 records from GBIF (https://www.gbif.org) as available on 23 April 2025. Black circles represent melanistic individuals. Map base: Esri and OpenStreetMap contributors.	FIGURE 3. Distribution of subspecies of Natrix astreptophora. (A) Approximate range of the regular phenotype of N. a. astreptophora; the broken lines indicate the approximate distribution borders for the northern (B) and western (C) phenotypes; in western Galicia all phenotypes co-occur.Asterisks are type localities (N. a. astreptophora has a ‘composed’ type locality of two sites, see Fritz & Schmidtler 2020; type locality of N. a. algerica is tentatively identified with Algiers). Data for nominotypical subspecies include 8299 records from GBIF (https://www.gbif.org) as available on 23 April 2025. Black circles represent melanistic individuals. Map base: Esri and OpenStreetMap contributors.	2025-09-16	Mármol, Gabriel Martínez Del;Geniez, Philippe;Neumann, Adrian;Glaw, Frank;Fritz, Uwe		Zenodo	biologists	Mármol, Gabriel Martínez Del;Geniez, Philippe;Neumann, Adrian;Glaw, Frank;Fritz, Uwe			
