taxonID	type	description	language	source
03EEAE6293785E67FF54C25EFD815CE8.taxon	materials_examined	Distribution Portugal. Mainland: Aveiro (iNaturalist, Appendix 1).	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE6293785E67FF54C25EFD815CE8.taxon	biology_ecology	Host plants Portugal. No information.	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE6293785E67FF54C25EFD815CE8.taxon	discussion	Comments. New for Portugal.	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE6293785E67FF54C306FD815E2C.taxon	materials_examined	Material examined. Faro: 11 ♂, 14 ♀, Cerca dos Pomares, 37 ° 19 ' 23.5 " N 8 ° 46 ' 03.7 " W, 11. ix. 2024, Tamarix africana (leg. D. Burckhardt, B. Duarte & D. L. Queiroz) (BDC, NHMB, dry mounted and 70 % ethanol); 2 ♀, Várzea de Aljezur, 37 ° 18 ' 19.8 " N 8 ° 47 ' 52.1 " W, 12. ix. 2024, T. africana (leg. B. Duarte) (BDC, 70 % ethanol). Distribution Portugal. Mainland: Faro (this study).	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE6293785E67FF54C306FD815E2C.taxon	biology_ecology	Host plants Portugal. No information.	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE6293785E67FF54C306FD815E2C.taxon	discussion	Comments. New for Portugal.	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE6293785E67FF54C041FD315F70.taxon	description	(Fig. 1)	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE6293785E67FF54C041FD315F70.taxon	materials_examined	Distribution Portugal. Madeira: Madeira Island (Loginova 1972, as Rhodochlanis parvipunctatus; Aguiar & Martin 1999), Porto Santo (Loginova 1972; Aguiar & Martin 1999), Selvagens Islands (Aguiar et al. 2019). Mainland: Faro (iNaturalist, Appendix 1).	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE6293785E67FF54C041FD315F70.taxon	biology_ecology	Host plants Portugal. Amaranthaceae: Suaeda vera Forssk. ex J. F. Gmel. (Aguiar & Martin 1999).	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE6293785E67FF54C041FD315F70.taxon	discussion	Comments. New for mainland Portugal.	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE62937F5E60FF54C36AFC505E8E.taxon	description	(Fig. 2)	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE62937F5E60FF54C36AFC505E8E.taxon	materials_examined	Material examined. Faro: 1 ♀, Aljezur, 37 ° 17 ' 04.2 " N 8 ° 48 ' 20.2 " W, 2. v. 2021, Daphne gnidium (leg. B. Duarte) (BDC, dry mounted); 1 ♀, Faro, 37 ° 05 ' 02.9 " N 7 ° 53 ' 29.4 " W, 9. iv. 2021, Grevillea robusta (leg. B. Duarte) (BDC, dry mounted); 2 ♀, 2 immatures, Olhão, 37 ° 03 ' 39.7 " N 7 ° 50 ' 36.6 " W, 21. iii. 2021, Pistacia lentiscus (photo B. Duarte); 3 ♂, 2 ♀, São Brás de Alportel, 4 km N São Brás, 16. vi. 1977, Pistacia sp. (leg. D. Hollis) (MHNG, dry mounted). Distribution Portugal. No details provided (Burckhardt & Lauterer 1989). Mainland: Faro (this study; iNaturalist, Appendix 1).	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE62937F5E60FF54C36AFC505E8E.taxon	biology_ecology	Host plants Portugal. Anacardiaceae: Pistacia lentiscus L. (this study).	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE62937F5E61FF54C0DEFBCD5A18.taxon	description	(Fig. 3)	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE62937F5E61FF54C0DEFBCD5A18.taxon	materials_examined	Material examined. Faro: 3 ♂, 2 ♀, 1 immature, Faro, 37 ° 02 ' 38.6 " N 7 ° 58 ' 30.8 " W, 29. vi. 2021, Pistacia lentiscus (leg. B. Duarte) (BDC, 70 % ethanol); 2 ♂, 1 ♀, Faro, 37 ° 02 ' 15.9 " N 7 ° 58 ' 41.2 " W, vii. 2020, P. lentiscus (leg. L. Neto) (BDC, dry mounted); 5 ♂, 2 ♀, Tavira, 37 ° 06 ' 15.6 " N 7 ° 43 ' 00.1 " W, 3. iii. 2021, P. lentiscus (leg. B. Duarte) (BDC, 70 % ethanol); 2 ♂, 2 ♀, 1 immature, Tavira, 37 ° 09 ' 49.8 " N 7 ° 32 ' 13.2 " W, 24. iii. 2021, P. lentiscus (leg. B. Duarte) (BDC, 70 % ethanol); 2 ♀, Portimão, Praia da Rocha, 10. ix. 1979 (leg. C. Lienhard) (MHNG, dry mounted); 1 ♀, 4 immatures, Fonte Benémola, 37 ° 12 ' 21.2 " N 8 ° 00 ' 11.5 " W, 12. v. 2024, P. lentiscus (leg. B. Duarte) (BDC, 70 % ethanol); 3 ♂, 2 ♀, Praia do Barril, 37 ° 05 ' 21.8 " N 7 ° 40 ' 12.7 " W, 8. ix. 2024, P. lentiscus (leg. D. Burckhardt, B. Duarte & D. L. Queiroz) (BDC, NHMB, dry mounted and 70 % ethanol); 1 ♂, 1 ♀, Cerca dos Pomares, 37 ° 19 ' 23.5 " N 8 ° 46 ' 03.7 " W, 11. ix. 2024, P. lentiscus (leg. D. Burckhardt & D. L. Queiroz) (NHMB, dry mounted); 1 ♂, 1 ♀, Ribat da Atalaia, 37 ° 19 ' 08.4 " N 8 ° 52 ' 28.9 " W, 12. ix. 2024, P. lentiscus (leg. D. Burckhardt & D. L. Queiroz) (NHMB, 70 % ethanol); 2 ♂, 1 ♀, Vale da Telha, 37 ° 18 ' 43.9 " N 8 ° 51 ' 54.0 " W, 12. ix. 2024, P. lentiscus (leg. D. Burckhardt & D. L. Queiroz) (NHMB, 70 % ethanol). Setúbal: 1 ♂, Capuchos, 26. vii. 1980, Acacia sp. (leg. A. van Harten) (MHNG, dry mounted). Distribution Portugal. Azores: No details provided (Hodkinson & Hollis 1981; Burckhardt & Lauterer 1989; Borges 2005). Mainland: Faro (this study, iNaturalist, Appendix 1); Setúbal (this study).	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE62937F5E61FF54C0DEFBCD5A18.taxon	biology_ecology	Host plants Portugal. Anacardiaceae: Pistacia lentiscus L. (this study). Parasitoids. Hymenoptera: Encyrtidae were obtained from mummified immatures.	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE62937E5E61FF54C3F5FB525E50.taxon	description	(Fig. 4)	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE62937E5E61FF54C3F5FB525E50.taxon	materials_examined	Distribution Portugal. Mainland: Faro (Pérez-Otero et al. 2011; iNaturalist, Appendix 1).	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE62937E5E61FF54C3F5FB525E50.taxon	biology_ecology	Host plants Portugal. Myrtaceae: Eucalyptus camaldulensis Dehnh. (Pérez-Otero et al. 2011).	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE62937E5E61FF54C454FAFD5D30.taxon	materials_examined	Material examined. Faro: 2 ♂, 2 ♀, 3 immatures (immatures not preserved), Alcaria, Aljezur, 37 ° 17 ' 04.2 " N 8 ° 48 ' 20.2 " W, 2. v. 2021, Cistus albidus (leg. B. Duarte) (BDC, dry mounted); 2 ♂, 3 ♀, Médo, Aljezur, 37 ° 19 ' 56.9 " N 8 ° 51 ' 19.7 " W, 2. v. 2021, C. halimifolius (leg. B. Duarte) (BDC, 70 % ethanol); 1 ♀, Médo, Aljezur, 37 ° 19 ' 56.9 " N 8 ° 51 ' 19.7 " W, 2. v. 2021, C. halimifolius (leg. B. Duarte) (BDC, dry mounted); 1 ♀, 7 immatures, Castro Marim, 37 ° 11 ' 52.5 " N 7 ° 28 ' 13.0 " W, 24. iii. 2021, C. salviifolius (leg. B. Duarte) (BDC, 70 % ethanol); 2 ♂, 1 ♀, Faro, 37 ° 02 ' 38.6 " N 7 ° 58 ' 30.8 " W, 23. iv. 2021, C. salviifolius (leg. B. Duarte) (BDC, dry mounted); 1 immature, Faro, 37 ° 02 ' 15.9 " N 7 ° 58 ' 41.2 " W, 17. iii. 2021, C. calycinus (leg. B. Duarte) (BDC, dry mounted); 1 ♂, 4 ♀, 1 immature, Faro, 37 ° 02 ' 15.9 " N 7 ° 58 ' 41.2 " W, 21. iv. 2021, C. calycinus (leg. B. Duarte) (BDC, dry mounted); 11 immature, Faro, 37 ° 02 ' 15.9 " N 7 ° 58 ' 41.2 " W, 08. iv. 2021, C. halimifolius (leg. B. Duarte) (BDC, dry mounted and 70 % ethanol); 3 ♂, 4 ♀, Fonte Benémola, 37 ° 12 ' 21.2 " N 8 ° 00 ' 11.5 " W, 12. v. 2024, C. albidus (leg. B. Duarte) (BDC, 70 % ethanol). Distribution Portugal. Mainland: Faro (Hodkinson & Hollis 1981, as Rhachistoneura varicicosta; this study), Leiria (Hodkinson & Hollis 1981, as R. varicicosta); Tavares (1905 a) reported an unidentified psyllid from Castelo Branco which might have concerned this species.	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE62937E5E61FF54C454FAFD5D30.taxon	biology_ecology	Host plants Portugal. Cistaceae: Cistus albidus L., C. calycinus L., C. halimifolius L., C. salviifolius L. (this study). Tavares (1905 a) reported an unidentified psyllid from C. ladanifer (as C. ladaniferus) L., which is a likely host. Hodkinson & Hollis (1981) reported material from Portugal from Ericaceae: Calluna vulgaris (L.) Hull which they considered to be a probable host. Based on the presence of immatures and adults, Rapisarda (1987) showed that this is not the case, and that Cistus spp. are hosts of this species. Cistus albidus, C. calycinus and C. halimifolius are reported here the first time as hosts of L. varicicosta.	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE62937E5E62FF54C0A3FF415A3C.taxon	materials_examined	Material examined. Madeira: 7 ♀, 1 immature, near airport, route to Machico, 60 m, 19. xi. 1992 (leg. C. Lienhard) (MHNG, dry mounted); 2 ♂, 3 ♀, SW of Porto Moniz, between Santa Maria Madalena and Achadas da Cruz, 700 m, 23. xi. 1992 (leg. C. Lienhard) (MHNG, dry mounted); 2 ♂, 1 ♀, Calheta, 950 m, 24. xi. 1992 (leg. C. Lienhard) (MHNG, dry mounted); 4 ♂, 4 ♀, 1 immature, Prazeres, 200 m, 14. v. 1998, mixed forest with Eucalyptus, Acer, etc. (leg. C. Lienhard) (MHNG, NHMB, dry mounted). Faro: 1 ♀, 2 ♂, 10 immatures, Aljezur, 37 ° 14 ' 29.3 " N 8 ° 48 ' 15.8 " W, 3. v. 2021, Eucalyptus globulus (leg. B. Duarte) (BDC, 70 % ethanol and dry mounted); 1 ♂, Aljezur, 37 ° 14 ' 29.3 " N 8 ° 48 ' 15.8 " W, 3. v. 2021, E. camaldulensis (leg. B. Duarte) (BDC, 70 % ethanol); 1 ♀, Faro, 37 ° 02 ' 45.7 " N 7 ° 57 ' 00.9 " W, 16. ii. 2022, Citrus sinensis (leg. B. Duarte) (BDC, dry mounted); 1 ♀, Tavira, 37 ° 12 ' 11.8 " N 7 ° 30 ' 17.6 " W, 27. i. 2021, E. globulus (leg. B. Duarte) (BDC, dry mounted); 1 ♂, Tavira, 37 ° 09 ' 49.8 " N 7 ° 32 ' 13.2 " W, 02. vii. 2020, Cerantonia siliqua (leg. L. Neto) (BDC, dry mounted); 3 ♂, 1 ♀, Pontal, Montenegro, 37 ° 02 ' 08.2 " N 7 ° 58 ' 54.1 " W, 7. ix. 2024, E. globulus (leg. D. Burckhardt & D. L. Queiroz) (NHMB, dry mounted); 2 ♀, Fóia, Serra de Monchique, 37 ° 18 ' 42.1 " N 8 ° 35 ' 15.4 " W, 8. ix. 2024, E. globulus (leg. D. Burckhardt & D. L. Queiroz) (NHMB, 70 % ethanol); 1 ♂, Miradouro da Fóia, Serra de Monchique, 37 ° 18 ' 58.7 " N 8 ° 35 ' 32.6 " W, 9. ix. 2024, parking lot with planted pine trees, Mediterranean scrub (leg. D. Burckhardt & D. L. Queiroz) (NHMB, dry mounted); 5 ♂, Bagagem, Aljezur, 37 ° 17 ' 58.9 " N 8 ° 48 ' 41.8 " W, 12. ix. 2024, E. globulus (leg. D. Burckhardt, B. Duarte & D. L. Queiroz) (BDC, NHMB, 70 % ethanol). Distribution Portugal. Azores: São Miguel (Hodkinson 1999, 2007); unknown location (Borges 2005). Madeira: Madeira Island (Burckhardt 1998; Aguiar & Martin 1999; Labina et al. 2007; this study). Mainland: unknown location (Nogueira 1971, as Psyllidae; Azevedo & Figo 1979); Aveiro (iNaturalist, Appendix 1); Faro (this study); Leiria, Lisbon (iNaturalist, Appendix 1).	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE62937E5E62FF54C0A3FF415A3C.taxon	biology_ecology	Host plants Portugal. Myrtaceae: Eucalyptus globulus Labill. (Azevedo & Figo 1979; Aguiar & Martin 1999).	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE62937D5E62FF54C4B1FDC35BC8.taxon	materials_examined	Material examined. Faro: 1 ♂, Aljezur, 37 ° 14 ' 29.3 " N 8 ° 48 ' 15.8 " W, 3. v. 2021, Eucalyptus camaldulensis (leg. B. Duarte) (BDC, dry mounted); 1 ♀, Aljezur, 37 ° 14 ' 29.3 " N 8 ° 48 ' 15.8 " W, 3. v. 2021, E. globulus (leg. B. Duarte) (BDC, dry mounted); 1 ♂, Faro, 37 ° 02 ' 45.7 " N 7 ° 57 ' 00.9 " W, 27. iv. 2021, Erica sp. (leg. B. Duarte) (BDC, dry mounted); 2 ♂, Balsinha, Aljezur, 37 ° 15 ' 46.8 " N 8 ° 46 ' 16.2 " W, 2. v. 2021, Fabaceae (leg. B. Duarte) (BDC, 70 % ethanol). Distribution Portugal. Mainland: “ all over ” (Valente et al. 2004); Aveiro (iNaturalist, Appendix 1); Faro (this study); Leiria, Porto (iNaturalist, Appendix 1).	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE62937D5E62FF54C4B1FDC35BC8.taxon	biology_ecology	Host plants Portugal. Myrtaceae: Eucalyptus dalrympleana Maiden, E. maidenii F. Muell., E. nitens (H. Deane & Maiden) Maiden (Valente et al. 2004).	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE62937D5E63FF54C225FC2358D4.taxon	materials_examined	Material examined. Beja: 1 ♂, Beringel, 38 ° 05 ' 11.2 " N 8 ° 01 ' 28.8 " W, 28. vii. 2020, Citrus sp. (leg. B. Duarte) (BDC, dry mounted); Faro: 1 ♂, 8 immatures, Aljezur, 37 ° 14 ' 29.3 " N 8 ° 48 ' 15.8 " W, 3. v. 2021, Eucalyptus camaldulensis (leg. B. Duarte) (BDC, dry mounted and 70 % ethanol); 1 ♀, Tavira, 37 ° 12 ' 11.8 " N 7 ° 30 ' 17.6 " W, 27. i. 2021, E. globulus (leg. B. Duarte) (BDC, dry mounted); 1 ♂, 1 ♀, Tavira, 37 ° 09 ' 49.8 " N 7 ° 32 ' 13.2 " W, Persea americana (leg. B. Duarte) (BDC, dry mounted); 25 ♂, 19 ♀, 5 immatures, Pontal, Montengro, 37 ° 02 ' 08.2 " N 7 ° 58 ' 54.1 " W, 7. ix. 2024, E. globulus (leg. D. Burckhardt, B. Duarte & D. L. Queiroz) (BDC, NHMB, dry mounted and 70 % ethanol). Lisbon: 8 ♂, 17 ♀, Lisbon, Tapada da Ajuda, 18. ix. 2008, E. camaldulensis (leg. C. Bovida) (NHMB, dry mounted). Distribution Portugal. Mainland: Beja (Valente & Hodkinson 2009; Bella 2013; this study, iNaturalist, Appendix 1); Bragança (iNaturalist, Appendix 1); Castelo Branco (Valente & Hodkinson 2009; iNaturalist, Appendix 1); Coimbra (iNaturalist, Appendix 1); Évora (Valente & Hodkinson 2009); Faro (Valente & Hodkinson 2009; this study; iNaturalist, Appendix 1); Lisbon (Moreira 2009; this study; iNaturalist, Appendix 1); Guarda, Portalegre (Valente & Hodkinson 2009); Santarém (Valente & Hodkinson 2009; iNaturalist, Appendix 1); Setúbal (iNaturalist, Appendix 1).	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE62937D5E63FF54C225FC2358D4.taxon	biology_ecology	Host plants Portugal. Myrtaceae: Eucalyptus camaldulensis Dehnh. (Valente & Hodkinson 2009; Bella 2013; Dhahri et al. 2014), E. cypellocarpa F. Muell., E. grandis W. Hill, E. macarthurii H. Deane & Maiden, E. nicholii Maiden & Blakely, E. nitens (H. Deane & Maiden) Maiden, E. ovata Labill., E. pauciflora Sieber, E. pilularis Sm., E. propinqua Deane & Maiden, E. pulverulenta Sims, E. regnans F. Muell., E. resinifera Sm., E. rudis Endl., E. sideroxylon A. Cunn. ex Woolls, E. tereticornis Smith, E. viminalis Labill. (Dhahri et al. 2014). Dhahri et al. (2014) evaluated the susceptibility of various Eucalyptus spp. to attack by G. brimblecombei in a survey carried out in a eucalypt arboretum in Lisbon in 2012. The authors showed that E. camaldulensis, E. ovata and E. tereticornis were highly susceptible (with 40.4 % to 68.4 % of leaves attacked) while E. cypellocarpa, E. grandis, E. marcarthurii, E. nicholii, E. nitens, E. pauciflora, E. pilularis, E. propinqua, E. pulverulenta, E. regnans, E. resinifera, E. rudis, E. sideroxylon and E. viminalis were considered moderately to barely susceptible (ranging from 26.9 % to 0.6 % attacked leaves). Another 13 species (E. botryoides, E. cinerea, E. citriodora, E. creba, E. globulus, E. goniocalyx, E. maculata, E. meliodora, E. occidentalis, E. perriniana, E. polyanthemos, E. robusta and E. saligna) were resistant to attack by Glyscaspis brimblecombei. Parasitoids. Dhahri et al. (2014) reported the presence of the parasitoid Psyllaephagus bliteus Riek (Hymenoptera: Encyrtidae) for the first time in mainland Portugal.	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE62937D5E63FF54C225FC2358D4.taxon	description	(Hymenoptera: Encyrtidae) for the first time in mainland Portugal.	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE62937C5E63FF54C789FF7B5CC4.taxon	materials_examined	Material examined. Faro: several adults and immatures, Aljezur, 37 ° 19 ' 00.9 " N 8 ° 48 ' 11.5 " W, 09. x. 2021, Schinus areira (leg. B. Duarte) (BDC, dry mounted and 70 % ethanol); several adults and immatures, Faro, 37 ° 01 ' 25.2 " N 7 ° 55 ' 11.3 " W, 25. iii. 2021, S. areira (leg. B. Duarte) (BDC, dry mounted and 70 % ethanol); several adults and immatures, Olhão, 37 ° 02 ' 06.0 " N 7 ° 49 ' 56.4 " W, 25. iii. 2021, S. areira (leg. B. Duarte) (BDC, dry mounted and 70 % ethanol); several adults and immatures, Tavira, 37 ° 10 ' 31.8 " N 7 ° 32 ' 07.8 " W, 24. iii. 2021, S. areira (leg. B. Duarte) (BDC, dry mounted and 70 % ethanol); 3 ♂, 2 ♀, Várzea de Aljezur, 37 ° 18 ' 19.8 " N 8 ° 47 ' 52.1 " W, 9 – 12. ix. 2024, S. areira (leg. D. Burckhardt & D. L. Queiroz) (NHMB, dry mounted and 70 % ethanol). Distribution Portugal. Madeira: Madeira Island (iNaturalist, Appendix 1). Mainland: Coimbra (iNaturalist, Appendix 1); Faro (this study); Lisbon (Zina et al. 2012; Bella 2013; iNaturalist, Appendix 1); Porto (iNaturalist, Appendix 1).	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE62937C5E63FF54C789FF7B5CC4.taxon	biology_ecology	Host plants Portugal. Anacardiaceae: Schinus areira L. (this study). The report from S. molle L. (Zina et al. 2012) probably also concerns S. areira; until recently, these two species have been regarded as synonyms (Zapater et al. 2018). All previously published host records of C. schini that we could check concerned S. areira, not S. molle (D. Burckhardt, unpubl. data). The latter is probably not used as a host by C. schini. Parasitoid. Immatures of C. schini parasitised by Tamarixia sp. (Hymenoptera: Eulophidae) were found in the district of Faro.	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE62937C5E63FF54C789FF7B5CC4.taxon	discussion	Comments. In a faunistic survey in several localities of the districts of Faro (Tavira, Patacão) and Santarém (Azambuja) in 2011, C. schini was not found (Zina et al. 2012). Here we report the species for the first time from south Portugal, where it is now widely distributed in the district of Faro. Also, entries in iNaturalist suggest that it is widely distributed throughout mainland Portugal, reflecting the popular use of the host as an ornamental tree in cities.	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE62937C5E6CFF54C398FA805F44.taxon	description	(Fig. 5)	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE62937C5E6CFF54C398FA805F44.taxon	materials_examined	Material examined. Aveiro: 2 ♂, 1 ♀, Luso, Buçaco Forest, 19. ix. 1979 (leg. C. Lienhard) (NHMB, dry mounted). Faro: 7 immatures, Fonte Benémola, Querença, 37 ° 12 ' 21.2 " N 8 ° 00 ' 11.5 " W, 12. v. 2024, Ficus carica (leg. B. Duarte) (BDC, 70 % ethanol); 2 ♂, 4 ♀, Fonte Benémola, Querença, 37 ° 12 ' 21.2 " N 8 ° 00 ' 11.5 " W, 12. v. 2024, F. carica (leg. B. Duarte) (BDC, 70 % ethanol); 1 ♂, between Casais and Cercada, 37 ° 17 ' 18.2 " N 8 ° 35 ' 57.5 " W, 9. ix. 2024, F. carica (leg. D. Burckhardt & D. L. Queiroz) (NHMB, dry mounted). Distribution Portugal. Mainland: No details provided (Burckhardt 1989); Aveiro (this study); Castelo Branco, Coimbra (iNaturalist, Appendix 1); Faro (this study; iNaturalist, Appendix 1).	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE62937C5E6CFF54C398FA805F44.taxon	biology_ecology	Host plants Portugal. Moraceae: Ficus carica L. (this study). ⁂ ‡ Macrohomotoma gladiata Kuwayama, 1908 (Figs 6 – 8)	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE62937C5E6CFF54C398FA805F44.taxon	materials_examined	Material examined. Faro: 3 ♂, 4 ♀, 2 immatures, Olhão, 37.023329 ° N 7.846906 ° W, 19. ix. 2024, Ficus microcarpa (leg. B. Duarte) (BDC, 70 % ethanol). Distribution Portugal. Mainland: Faro (this study).	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE62937C5E6CFF54C398FA805F44.taxon	biology_ecology	Host plants Portugal. Moraceae: Ficus microcarpa L. f. (this study).	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE62937C5E6CFF54C398FA805F44.taxon	discussion	Comments. New for Portugal. Parasitoids. Confirmed in the laboratory, but emerged hymenopteran parasitoids not yet identified.	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE6293725E6DFF54C718FB0A5988.taxon	materials_examined	Distribution Portugal. Madeira: Madeira Island (Aguiar & Martin 1999).	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE6293725E6DFF54C718FB0A5988.taxon	biology_ecology	Host plants Portugal. Oleaceae: Picconia excelsa (Aiton) DC. (Aguiar & Martin 1999).	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE6293725E6DFF54C7E4FD4A5D9C.taxon	description	(Fig. 9)	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE6293725E6DFF54C7E4FD4A5D9C.taxon	materials_examined	Material examined. Faro: 3 ♂, 9 ♀, Citriaroeira, Tavira, 37 ° 12 ' 11.8 " N 7 ° 30 ' 17.6 " W, 03. ix. 2020, Olea europaea subsp. europaea (leg. B. Duarte) (BDC, 70 % ethanol); 1 ♂, 1 ♀, 1 immature, Gambelas, Faro, 37 ° 02 ' 38.6 " N 7 ° 58 ' 30.8 " W, 01. iii. 2021, O. europaea subsp. europaea (leg. B. Duarte) (BDC, 70 % ethanol); 2 ♂, 3 ♀, Tavira, 37 ° 12 ' 11.8 " N 7 ° 30 ' 17.6 " W, 10. iii. 2021, O. europaea subsp. europaea (leg. B. Duarte) (BDC, 70 % ethanol); 2 ♂, 1 ♀, Gambelas, Faro, 37 ° 02 ' 15.9 " N 7 ° 58 ' 41.2 " W, 17. iii. 2021, O. europaea subp. europaea (leg. B. Duarte) (BDC, 70 % ethanol); 2 ♂, 5 ♀, Gambelas, Faro, 37 ° 02 ' 38.6 " N 7 ° 58 ' 30.8 " W, 1. iv. 2021, Phillyrea angustifolia (leg. B. Duarte) (BDC, dry mounted); 2 ♂, 2 ♀, 1 immature, Faro, 37 ° 02 ' 38.6 " N 7 ° 58 ' 30.8 " W, 1. vii. 2021, P. angustifolia (leg. B. Duarte) (BDC, dry mounted); 2 ♂, 3 ♀, Fonte de Benémola, Querença, 37 ° 12 ' 21.2 " N 8 ° 00 ' 11.5 " W, 6. ix. 2024, O. europaea (leg. D. Burckhardt & D. L. Queiroz) (NHMB, dry mounted); 2 ♂, 7 ♀, Mata de Gambelas, Montenegro, 37 ° 02 ' 30.5 " N 7 ° 58 ' 29.6 " W, 7. ix. 2024, O. europaea (leg. D. Burckhardt & D. L. Queiroz) (NHMB, 70 % ethanol); 4 ♂, between Aljezur and Bagagem, 37 ° 18 ' 41.8 " N 8 ° 48 ' 44.3 " W, 10. ix. 2024, O. europaea (leg. D. Burckhardt & D. L. Queiroz) (NHMB, 70 % ethanol). Distribution Portugal. Azores: Terceira (Lopes et al. 2010). Madeira: Madeira Island (Aguiar 2009); Porto Santo (Aguiar et al. 2019). Mainland: Aveiro (iNaturalist, Appendix 1); Faro (this study; iNaturalist, Appendix 1); Leiria, Portalegre (iNaturalist, Appendix 1).	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE6293725E6DFF54C7E4FD4A5D9C.taxon	biology_ecology	Host plants Portugal. Oleaceae: Olea europaea subsp. europaea L., Phillyrea angustifolia L. (this study); O. europaea L., O. maderensis (Lowe) Rivas Mart. & del Arco (Aguiar 2009); adults were collected on Picconia excelsa (Aiton) DC., which is a potential host (Aguiar et al. 2019). Parasitoids. Hymenopteran parasitoids of three genera emerged from mummies in the laboratory: Psyllaephagus (Encyrtidae), Pachyneuron (Pteromalidae) and Apocharips (Figitidae). The encyrtid is probably Psyllaephagus euphyllurae (Masi), the most important parasitoid of E. olivina in the western Mediterranean Basin (Triapitsyn et al. 2014). Triapitsyn et al. (2014) considered Pachyneuron and Apocharips to be hyperparasitoids of P. euphyllurae.	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE6293725E6DFF54C7E4FD4A5D9C.taxon	discussion	Comments. Immature E. olivina secrete white waxy flocculence resembling cotton; the secretions are usually found on the inflorescences or in the axils of small branches. The secretions by E. olivina on Olea europaea subsp. europaea and Phillyrea angustifolia appear identical.	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE6293725E6DFF54C3D0FD815F4C.taxon	materials_examined	Distribution Portugal. Mainland: Castelo Branco (Tavares 1905 a); Lisbon (Tavares 1905 a); Portalegre (Tavares 1905 a).	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE6293725E6DFF54C3D0FD815F4C.taxon	biology_ecology	Host plants Portugal. Oleaceae: Fraxinus angustifolia Vahl (Tavares 1905 a). ⁂ Psyllopsis fraxinicola (Foerster, 1848)	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE6293725E6DFF54C3D0FD815F4C.taxon	materials_examined	Distribution Portugal. Mainland: Faro (iNaturalist, Appendix 1).	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE6293725E6DFF54C3D0FD815F4C.taxon	biology_ecology	Host plants Portugal. No information.	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE6293725E6DFF54C3D0FD815F4C.taxon	discussion	Comments. New for Portugal.	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE6293715E6FFF54C0DCFE7558D4.taxon	materials_examined	Material examined. Madeira: 1 ♂, 2 ♀, Calheta, 950 m, 24. xi. 1992 (leg. C. Lienhard) (MHNG, dry mounted); 5 ♂, 1 ♀, near Poiso, road to Pico de Arieiro, 1420 m, 20. xi. 1992 (leg. C. Lienhard) (MHNG, dry mounted); 1 ♀, SW of Porto Moniz, between Santa and Achadas da Cruz, 700 m, 23. xi. 1992 (leg. C. Lienhard) (MHNG, dry mounted); 1 ♀, Santo da Serra, road to Poiso, 850 m, 20. xi. 1992 (leg. C. Lienhard) (MHNG, dry mounted). Distribution Portugal. Madeira: Madeira Island (Loginova 1976; Hodkinson 1981 a; Aguiar & Martin 1999; Labina et al. 2007; this study).	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE6293715E6FFF54C0DCFE7558D4.taxon	biology_ecology	Host plants Portugal. Ericaceae: Erica arborea L. (Aguiar & Martin 1999); E. scoparia L. (Loginova 1976; Aguiar & Martin 1999). Loginova (1976) listed a single male from E. “ zucinulus ”, a probable error referring to E. scoparia (Hodkinson 1982).	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE6293705E6FFF54C719FE925BC8.taxon	description	(Figs 10 – 15)	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE6293705E6FFF54C719FE925BC8.taxon	materials_examined	Material examined. Faro: 9 ♂, 12 ♀, Faro, 37 ° 02 ' 15.9 " N 7 ° 58 ' 41.2 " W, 17. iii. 2021, Erica umbellata (leg. B. Duarte) (BDC, dry mounted); 3 ♂, 8 ♀, Faro, 37 ° 02 ' 15.9 " N 7 ° 58 ' 41.2 " W, 10. iv. 2021, E. umbellata (leg. B. Duarte) (BDC, dry mounted); 9 ♂, 14 ♀, 1 immature, Faro, 37 ° 02 ' 15.9 " N 7 ° 58 ' 41.2 " W, 10. iv. 2021, Erica spp. (leg. B. Duarte) (BDC, 70 % ethanol); 1 ♂, Faro, 37 ° 02 ' 15.9 " N 7 ° 58 ' 41.2 " W, 15. iv. 2021, E. umbellata (leg. B. Duarte) (BDC, 70 % ethanol); 1 ♀, 1 immature, Aljezur, Canal, 37 ° 16 ' 36.0 " N 8 ° 50 ' 09.2 " W, 2. v. 2021, E. umbellata (leg. B. Duarte) (BDC, 70 % ethanol); 1 ♂, Aljezur, Mourão, 37 ° 16 ' 19.1 " N 8 ° 45 ' 36.7 " W, 2. v. 2021, Erica spp. (leg. B. Duarte) (BDC, dry mounted). Distribution Portugal. Mainland: Faro (Hodkinson 1981 a, 1982; this study).	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE6293705E6FFF54C719FE925BC8.taxon	biology_ecology	Host plants Portugal. Ericaceae: Erica australis L. (Hodkinson 1981 a, 1982); E. umbellata L. (this work).	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE6293705E6FFF54C719FE925BC8.taxon	discussion	Comments. Strophingia australis was previously known only from a single male and female from São Brás (Faro) (Hodkinson 1981 a). The species displays sexual dimorphism in the forewing size and shape (Figs 10, 11), and the coronal suture is reduced in basal half (Fig. 12). Strophingia australis is characterised by the paramere, the distal segment of the aedeagus and the female terminalia. The paramere, in lateral view, is lanceolate, lacks a posterior lobe and bears a small sclerotised apical tooth (Fig. 13); and the distal segment of the aedeagus bears a weakly curved, apically rounded apical dilation (Fig. 14). The female terminalia, in lateral view, are elongate (Fig. 15); the dorsal margin of the proctiger is weakly undulate and the ventral margin of the subgenital plate is angled in the middle.	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE6293705E6FFF54C225FDF95E2C.taxon	materials_examined	Material examined. Faro. 1 ♂, Aljezur, 37 ° 15 ' 46.8 " N 8 ° 46 ' 16.2 " W, 2. v. 2021, unidentified Fabaceae (leg. B. Duarte) (BDC, 70 % ethanol); 1 ♂, 10 ♀, Padescas, 37 ° 18 ' 10.1 " N 8 ° 44 ' 33.0 " W, 9. ix. 2024, Erica arborea (leg. D. Burckhardt & D. L. Queiroz) (NHMB, 70 % ethanol); 1 ♂, Miradouro da Fóia, Serra de Monchique, 37 ° 18 ' 58.7 " N 8 ° 35 ' 32.6 " W, 9. ix. 2024, parking lot with planted pine trees, Mediterranean scrub, Calluna vulgaris (leg. D. Burckhardt & D. L. Queiroz) (NHMB, dry mounted); 14 ♂, 56 ♀, between Aljezur and Bagagem, 37 ° 18 ' 41.8 " N 8 ° 48 ' 44.3 " W, 10. ix. 2024, Mediterranean scrub with pines, E. arborea (leg. D. Burckhardt & D. L. Queiroz) (NHMB, dry mounted and 70 % ethanol); 1 ♀, Cerca dos Pomares, Aljezur, 37 ° 19 ' 23.5 " N 8 ° 46 ' 03.7 " W, 11. ix. 2024, vegetation along dry riverbed, E. arborea (leg. D. Burckhardt & D. L. Queiroz) (NHMB, dry mounted); 3 ♀, Vale das Amoreiras, Carrascalinho, 37 ° 20 ' 12.5 " N 8 ° 46 ' 41.2 " W, 11. ix. 2024, oak forest, E. arborea (leg. D. Burckhardt & D. L. Queiroz) (NHMB, 70 % ethanol). Distribution Portugal. Mainland: no details provided (Hodkinson 1981 a); Faro (this study; iNaturalist, Appendix 1).	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE6293705E6FFF54C225FDF95E2C.taxon	biology_ecology	Host plants Portugal. Ericaceae: Erica lusitanica Rudolphi (Hodkinson 1981 a); adults were collected also on E. arborea, a confirmed host elsewhere.	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE6293705E6FFF54C041FBF65F28.taxon	materials_examined	Material examined. Faro. 2 ♀, Faro, 37 ° 02 ' 24.4 " N 7 ° 58 ' 37.7 " W, 10. iv. 2021, Erica spp. (leg. B. Duarte) (BDC, 70 % ethanol). Distribution Portugal. Mainland: no details provided (Hodkinson 1981 a); Faro (this study).	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE6293705E6FFF54C041FBF65F28.taxon	biology_ecology	Host plants Portugal. Ericaceae: Calluna vulgaris (L.) Hull (Hodkinson 1981 a).	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE6293705E68FF54C14CFE0F58D4.taxon	materials_examined	Material examined. Madeira: 2 ♂, 2 ♀, Machico, 160 m, 21. xi. 1992 (leg. C. Lienhard) (MHNG, dry mounted). Distribution Portugal. Madeira: Madeira Island (Loginova 1976; Hodkinson 1981 a; Aguiar & Martin 1999; Labina et al. 2007; this study).	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE6293705E68FF54C14CFE0F58D4.taxon	biology_ecology	Host plants Portugal. Ericaceae: Erica arborea L. (Hodkinson 1981 a; Aguiar & Martin 1999); E. scoparia L. (Aguiar & Martin 1999).	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE6293775E68FF54C25CFD2E5D0C.taxon	materials_examined	Distribution Portugal. Mainland: No details provided (Nogueira 1971, as Rhinocola speciosa).	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE6293775E68FF54C25CFD2E5D0C.taxon	biology_ecology	Host plants Portugal. Salicaceae: Populus nigra L. (Nogueira 1971).	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE6293775E68FF54C25CFD2E5D0C.taxon	discussion	Comments. According to Nogueira (1971), C. speciosa was the commonest psyllid on forest trees in Portugal, before the discovery of the eucalypt psyllids.	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE6293775E68FF54C360FD815FDC.taxon	materials_examined	Distribution Portugal. Mainland: Castelo Branco (Tavares 1905 a, as Livia juncorum; iNaturalist, Appendix 1); Coimbra (iNaturalist, Appendix 1); Beja (Jarzembowski et al. 2013); Leiria (iNaturalist, Appendix 1), Lisbon (Tavares 1901, 1905 a, as L. juncorum); Porto (Tavares 1905 a, as L. juncorum); Vila Real (iNaturalist, Appendix 1).	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE6293775E68FF54C360FD815FDC.taxon	biology_ecology	Host plants Portugal. Juncaceae: Juncus fontanesii J. Gay ex Laharpe (Jarzembowski et al. 2013), J. articulatus subsp. articulatus (Tavares 1901, 1905 a, as J. lamprocarpus), and J. supinus (Tavares 1905 a). ⁂ Livia mediterranea Loginova, 1974 (Fig. 16)	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE6293775E68FF54C360FD815FDC.taxon	materials_examined	Material examined. Faro: 5 ♂, 7 ♀, Praia do Barril, Santa Luzia, 37 ° 05 ' 21.8 " N 7 ° 40 ' 12.7 " W, 8. ix. 2024, Pinus pinea (leg. D. Burckhardt, B. Duarte & D. L. Queiroz) (BDC, NHMB, dry mounted and 70 % ethanol); 1 ♀, Miradouro da Fóia, Serra de Monchique, 37 ° 18 ' 58.7 " N 8 ° 35 ' 32.6 " W, 9. ix. 2024, P. pinea (leg. D. Burckhardt & D. L. Queiroz) (NHMB, dry mounted). Distribution Portugal. Mainland: Faro (this study; iNaturalist, Appendix 1).	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE6293775E68FF54C360FD815FDC.taxon	biology_ecology	Host plants Portugal. No information.	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE6293775E68FF54C360FD815FDC.taxon	discussion	Comments. New for Portugal.	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE6293775E68FF54C719FB2C5A60.taxon	materials_examined	Material examined. Azores: 1 ♂, Faial, Capelo Parque Florestal, 240 m, 17. xi. 1992 (leg. C. Lienhard) (MHNG, dry mounted); 2 ♂, Faial, near Areeiro, 300 – 500 m, 17. xi. 1992 (leg. C. Lienhard) (MHNG, dry mounted); 2 ♂, Faial, road to Ribeira Funda and Largo Jaime de Melo, 400 m, 17. xi. 1992 (leg. C. Lienhard) (MHNG, dry mounted); Faial, road Horta to Caldeira, 16. xi. 1992 (leg. C. Lienhard) (MHNG, dry mounted). Distribution Portugal. Azores: Faial (Borges 2005; this study); Flores (Hodkinson 1981 a; Borges 2005); Pico (Borges 2005); Santa Maria (Borges 2005); São Jorge (Borges 2005); São Miguel (Hodkinson 1981 a); Terceira (Borges 2005); unknown location (Gaspar et al. 2008).	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE6293775E68FF54C719FB2C5A60.taxon	biology_ecology	Host plants Portugal. Ericaceae: Erica azorica Hochst. ex Seub. (Hodkinson 1981 a).	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE6293775E68FF54C48DFD985BC8.taxon	materials_examined	Material examined. Faro: 1 ♂, 1 ♀, between Aljezur and Bagagem, 37 ° 18 ' 41.8 " N 8 ° 48 ' 44.3 " W, 10. ix. 2024, Mediterranean scrub with pines, Erica arborea (leg. D. Burckhardt & D. L. Queiroz) (NHMB, dry mounted); 2 ♂, 4 ♀, Vale da Telha, Arrifana, 37 ° 18 ' 43.9 " N 8 ° 51 ' 54.0 " W, maquis shrubland between garden and houses, E. arborea (leg. D. Burckhardt & D. L. Queiroz) (NHMB, dry mounted). Distribution Portugal. Mainland: Faro (this study); Leiria (Hodkinson 1981 a).	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE6293775E68FF54C48DFD985BC8.taxon	biology_ecology	Host plants Portugal. Ericaceae: Erica sp. Tourn. ex L. (Hodkinson 1981 a); adults were collected on E. arborea, a confirmed host elsewhere.	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE6293765E69FF54C719FCE25A3C.taxon	materials_examined	Material examined. Faro: 1 ♂, 13 immatures, Silves, 37 ° 20 ' 47.3 " N 8 ° 22 ' 24.8 " W, 8. x. 2021, Acacia dealbata (leg. B. Duarte) (BDC, 70 % ethanol); 1 ♂, Monchique, 37 ° 18 ' 42.1 " N 8 ° 35 ' 15.4 " W, 9. ix. 2024, eucalypt plantation with Mediterranean understorey vegetation (leg. D. Burckhardt & D. L. Queiroz) (NHMB, 70 % ethanol and dry mounted). Distribution Portugal. Mainland: Faro (this study).	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE6293765E69FF54C719FCE25A3C.taxon	biology_ecology	Host plants Portugal. Fabaceae: Acacia dealbata Link (Fabaceae) (this study).	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE6293765E69FF54C719FCE25A3C.taxon	discussion	Comments. New for Portugal and the Iberian Peninsula.	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE6293765E69FF54C4B0FB215B38.taxon	materials_examined	Material examined. Faro: 2 immatures, Vila Real de Santo António, 37 ° 11 ' 48.7 " N 7 ° 25 ' 32.5 " W, 10. vi. 2022, Albizia julibrissin (leg. B. Duarte) (BDC, 70 % ethanol). Distribution Portugal. Mainland: Coimbra (Bella 2013); Faro (this study).	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE6293765E69FF54C4B0FB215B38.taxon	biology_ecology	Host plants Portugal. Fabaceae: Albizia julibrissin Durazz. (Bella 2013; this study).	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE6293765E69FF54C5B4FA365E98.taxon	materials_examined	Material examined. Faro: 42 immatures, Aljezur, 37 ° 16 ' 19.1 " N 8 ° 45 ' 36.7 " W, 2. v. 2021, Acacia sp. (leg. B. Duarte) (BDC, dry mounted and 70 % ethanol); 1 ♂, 3 ♀, 2 immatures, Silves, 37 ° 20 ' 47.3 " N 8 ° 22 ' 24.8 " W, 8. x. 2021, A. dealbata (leg. B. Duarte) (BDC, dry mounted); 1 ♂, Vila Real de Santo António, 37 ° 11 ' 48.7 " N 7 ° 25 ' 32.5 " W, 10. vi. 2022, Albizia sp. (leg. B. Duarte) (BDC, 70 % ethanol); 1 ♂, Faro, 37 ° 02 ' 38.6 " N 7 ° 58 ' 30.8 " W, 29. vi. 2020, Pistacia lentiscus (leg. B. Duarte) (BDC, dry mounted); 1 ♀, Aljezur, Moinho da berberia, 37 ° 18 ' 39.0 " N 8 ° 47 ' 11.8 " W, 01. v. 2021, Juniperus communis (leg. B. Duarte) (BDC, dry mounted); 2 ♂, 2 ♀, 4 immatures, Mata da Conceição, Santa Rita, 37 ° 11 ' 23.9 " N 7 ° 34 ' 14.6 " W, 4. v. 2024, Acacia cf. pycnantha (leg. B. Duarte) (BDC, 70 % ethanol); 1 ♀, Praia do Barril, 37 ° 05 ' 21.8 " N 7 ° 40 ' 12.7 " W, 8. ix. 2024, A. saligna (leg. D. Burckhardt & D. L. Queiroz) (NHMB, dry mounted); 44 ♂, 25 ♀, 3 immatures, Monchique, 37 ° 18 ' 42.1 " N 8 ° 35 ' 15.4 " W, 9. ix. 2024, A. saligna (leg. D. Burckhardt & D. L. Queiroz) (NHMB, 70 % ethanol and dry mounted); 18 ♂, 9 ♀, Várzea de Aljezur, 37 ° 18 ' 19.8 " N 8 ° 47 ' 52.1 " W, 9 – 12. ix. 2024, A. saligna (leg. D. Burckhardt, B. Duarte & D. L. Queiroz) (BDC, NHMB, 70 % ethanol); 46 ♂, 34 ♀, 8 immatures, between Aljezur and Bagagem, 37 ° 18 ' 41.8 " N 8 ° 48 ' 44.3 " W, 10. ix. 2024, A. saligna (leg. D. Burckhardt, B. Duarte & D. L. Queiroz) (BDC, NHMB, 70 % ethanol); 5 ♂, 8 ♀, Vale da Telha, 37 ° 18 ' 43.9 " N 8 ° 51 ' 54.0 " W, 12. ix. 2024, A. saligna (leg. D. Burckhardt & D. L. Queiroz) (NHMB, 70 % ethanol). Lisbon: 7 ♂, 3 ♀, Sintra, between Sintra and Parque Monserrate, 13. xi. 1992 (leg. C. Lienhard) (MHNG, dry mounted). Madeira: 1 ♂, Porto Santo, Pico do Castelo, 300 m, 22. xi. 1992 (leg. C. Lienhard) (MHNG, dry mounted). Distribution Portugal. Azores: Pico, Terceira (Borges 2005); no details provided (Gaspar et al. 2008). Madeira: Madeira Island (Aguiar et al. 2019). Mainland: no details provided (Hodkinson & Hollis 1987); Faro, Lisbon (this study; iNaturalist, Appendix 1); Porto (iNaturalist, Appendix 1).	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE6293765E69FF54C5B4FA365E98.taxon	biology_ecology	Host plants Portugal. Fabaceae: Acacia baileyana F. Muell., A. longifolia (Andrews) Willd., A. mearnsii De Wild. (Aguiar et al. 2019); A. dealbata Link, A. cf. pycnantha Benth., A. saligna (Labill.) H. L. Wendl. (this study).	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE6293765E6AFF54C10CFE3359AC.taxon	description	(Fig. 17)	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE6293765E6AFF54C10CFE3359AC.taxon	materials_examined	Material examined. Faro: 3 ♂, 2 ♀, 3 immatures, Faro, 37 ° 02 ' 15.9 " N 7 ° 58 ' 41.2 " W, 9. x. 2020, Tipuana tipu (leg. B. Duarte) (BDC, 70 % ethanol); 1 ♂, 1 ♀, 4 immatures, Faro, 37 ° 02 ' 15.9 " N 7 ° 58 ' 41.2 " W, 2. iii. 2021, T. tipu (leg. B. Duarte) (BDC, dry mounted); 5 immatures, Olhão, 37 ° 01 ' 52.3 " N 7 ° 50 ' 10.3 " W, 16. iii. 2021, T. tipu (leg. B. Duarte) (BDC, 70 % ethanol); 8 ♂, 6 ♀, 1 immature, Faro, 37 ° 01 ' 34.3 " N 7 ° 56 ' 09.0 " W, 8. vii. 2021, T. tipu (leg. B. Duarte) (BDC, 70 % ethanol). Distribution Portugal. Madeira: Madeira Island (Aguiar et al. 2019; Bella & Aguiar 2020; iNaturalist, Appendix 1); Porto Santo (Aguiar et al. 2019). Mainland: Aveiro (iNaturalist, Appendix 1); Coimbra (Bella 2013); Faro (Sánchez 2011; this study; iNaturalist, Appendix 1); Lisbon (Bella 2013); Santarém, Setúbal (iNaturalist, Appendix 1).	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE6293765E6AFF54C10CFE3359AC.taxon	biology_ecology	Host plants Portugal. Fabaceae: Tipuana tipu (Benth.) Kuntze (Sánchez 2011; Bella 2013; Aguiar et al. 2019; Bella & Aguiar 2020).	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE6293755E6BFF54C3F0FAFD5988.taxon	description	(Fig. 18)	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE6293755E6BFF54C3F0FAFD5988.taxon	materials_examined	Material examined. Coimbra: 1 ♂, 1 ♀, between the cities of Lousã and Coimbra, 24. v. 1979 (leg. C. Lienhard) (MHNG, dry mounted). Faro: 1 ♀, between Casais and Cercada, 37 ° 17 ' 18.2 " N 8 ° 35 ' 57.5 " W, 9. ix. 2024, Adenocarpus hispanicus subsp. hispanicus (leg. D. Burckhardt & D. L. Queiroz) (NHMB, dry mounted); 4 ♂, 3 ♀, Cerca dos Pomares, 37 ° 17 ' 18.2 " N 8 ° 35 ' 57.5 " W, 11. ix. 2024, A. anisochilus (leg. D. Burckhardt, B. Duarte & D. L. Queiroz) (BDC, NHMB, dry mounted and 70 % ethanol). Distribution Portugal. No details provided (Hodkinson & Hollis 1987). Mainland: Coimbra (this study); Faro (this study; iNaturalist, Appendix 1); Guarda, Porto (iNaturalist, Appendix 1).	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE6293755E6BFF54C3F0FAFD5988.taxon	biology_ecology	Host plants Portugal. Adults were collected on Fabaceae: Adenocarpus complicatus (L.) J. Gay and A. hispanicus subsp. hispanicus (Lam.) DC., which are likely hosts. ⁂ Arytaina genistae (Latreille, 1804)	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE6293755E6BFF54C3F0FAFD5988.taxon	materials_examined	Material examined. Lisbon: 2 ♂, 8 ♀, Colares, Serra de Sintra, 16. ix. 1979, Cytisus sp. (leg. C. Lienhard) (MHNG, dry mounted). Distribution Portugal. Mainland: Bragança, Faro, Guarda (iNaturalist, Appendix 1); Lisbon (this study); Porto (iNaturalist, Appendix 1).	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE6293755E6BFF54C3F0FAFD5988.taxon	biology_ecology	Host plants Portugal. No information.	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE6293755E6BFF54C3F0FAFD5988.taxon	discussion	Comments. New for Portugal. The mention by Aulmann (1913, as Psylla spartii) is erroneous (Table 2).	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE6293745E6BFF54C063FD235EBF.taxon	materials_examined	Distribution Portugal. Mainland: near Mondego River (Hodkinson & Hollis 1987).	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE6293745E6BFF54C063FD235EBF.taxon	biology_ecology	Host plants Portugal. No information.	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE6293745E74FF54C12FFD815964.taxon	materials_examined	Distribution Portugal. No details provided (Hodkinson & Hollis 1987).	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE6293745E74FF54C12FFD815964.taxon	biology_ecology	Host plants Portugal. No information. ⁂ Arytainilla delarbrei (Puton, 1873)	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE6293745E74FF54C12FFD815964.taxon	materials_examined	Material examined. Faro: 1 ♀, Aljezur, Monte da Cruz, 37 ° 18 ' 46.4 " N 8 ° 47 ' 30.1 " W, 2. v. 2021, Melaleuca sp. (leg. B. Duarte) (BDC, dry mounted). Distribution Portugal. Mainland: Faro (this study).	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE6293745E74FF54C12FFD815964.taxon	biology_ecology	Host plants Portugal. No information.	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE6293745E74FF54C12FFD815964.taxon	discussion	Comments. New for Portugal.	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE62936B5E74FF54C788FACF5B14.taxon	materials_examined	Material examined. Faro: 1 ♀, Mafra, 38 ° 57 ' 25.7 " N 9 ° 22 ' 04.9 " W, 1. iii. 2021, Acacia sp. (leg. B. Duarte) (BDC, dry mounted); 16 ♂, 22 ♀, 6 immatures, Faro, 37 ° 02 ' 38.6 " N 7 ° 58 ' 30.8 " W, 1. iii. 2021, Cytisus grandiflorus (leg. B. Duarte) (BDC, dry mounted and 70 % ethanol); 3 ♂, 40 immatures, Faro, Mata Gambelas, 37 ° 02 ' 15.9 " N 7 ° 58 ' 41.2 " W, 1. iii. 2021, C. grandiflorus (leg. B. Duarte) (BDC, 70 % ethanol); 1 ♀, Faro, 37 ° 02 ' 15.9 " N 7 ° 58 ' 41.2 " W, 10. iv. 2021, Genista hirsuta (leg. B. Duarte) (BDC, dry mounted); 1 ♂, Faro, 37 ° 02 ' 15.9 " N 7 ° 58 ' 41.2 " W, 10. iv. 2021, Phillyrea angustifolia (leg. B. Duarte) (BDC, dry mounted). Distribution Portugal. No details provided (Hodkinson & Hollis 1987). Mainland: Faro (this study), Guarda (iNaturalist, Appendix 1), Lisbon (this study).	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE62936B5E74FF54C788FACF5B14.taxon	biology_ecology	Host plants Portugal. Fabaceae: Cytisus grandiflorus DC. (this study), a new host record for this species.	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE62936B5E74FF54C558FF415BEC.taxon	materials_examined	Distribution Portugal. Madeira: Madeira Island (Loginova 1976; Aguiar & Martin 1999; Percy 2003).	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE62936B5E74FF54C558FF415BEC.taxon	biology_ecology	Host plants Portugal. Fabaceae: Genista maderensis (Webb & Berthel.) Lowe (Aguiar & Martin 1999; Percy 2003).	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE62936B5E74FF54C200FAEC5E50.taxon	materials_examined	Distribution Portugal. Madeira: Madeira Island (Loginova 1976; Aguiar & Martin 1999; Percy 2003).	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE62936B5E74FF54C200FAEC5E50.taxon	biology_ecology	Host plants Portugal. Fabaceae: Genista tenera (Jacq. ex Murray) Kuntze (Percy 2003). ⁂ Cacopsylla alaterni (Foerster, 1848)	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE62936B5E74FF54C200FAEC5E50.taxon	materials_examined	Material examined. Faro: 2 ♀, 1 immature, Faro, Patacão, 37 ° 02 ' 45.7 " N 7 ° 57 ' 00.9 " W, 27. iv. 2021, Rhamnus alaternus (leg. B. Duarte) (BDC, dry mounted); 3 ♀, Cerca dos Pomares, Aljezur, 37 ° 19 ' 23.5 " N 8 ° 46 ' 03.7 " W, 11. ix. 2024, R. alaternus (leg. D. Burckhardt, B. Duarte & D. L. Queiroz) (BDC, NHMB, dry mounted and 70 % ethanol); 2 ♂, 12 ♀, Vale das Amoreiras, Carrascalinho, 37 ° 20 ' 12.5 " N 8 ° 46 ' 41.2 " W, 11. ix. 2024, R. alaternus (leg. D. Burckhardt, B. Duarte & D. L. Queiroz) (BDC, NHMB, dry mounted and 70 % ethanol). Distribution Portugal. Mainland: Faro (this study; iNaturalist, Appendix 1).	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE62936B5E74FF54C200FAEC5E50.taxon	biology_ecology	Host plants Portugal. Rhamnaceae: Rhamnus alaternus L. (this study).	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE62936B5E74FF54C200FAEC5E50.taxon	discussion	Comments. This species is reported here for the first time from Portugal. As Rhamnus alaternus is very common in mainland Portugal (Flora-On 2024), C. alaterni is probably also widely distributed in mainland Portugal.	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE62936B5E74FF54C09CFEF15F94.taxon	materials_examined	Material examined. Madeira: 2 ♂, 2 ♀, below Santana, road to Achada do Teixeira, 800 m, 18. v. 1998, eucalypt forest with shrubs and Juniperus (leg. C. Lienhard) (MHNG, NHMB, dry mounted). Distribution Portugal. Madeira: Madeira Island (Loginova 1976, as Psylla atlantica; Aguiar & Martin 1999, as P. atlantica; this study).	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE62936B5E74FF54C09CFEF15F94.taxon	biology_ecology	Host plants Portugal. Salicaceae: Salix pedicellata subsp. canariensis (C. Sm. ex Link) A. K. Skvortsov (Aguiar & Martin 1999).	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE62936A5E75FF54C6A8FAC65964.taxon	materials_examined	Material examined. Madeira: 4 ♂, 5 ♀, Jardim da Serra, below Estreito de Câmara de Lobos, near forest house, 1150 – 1250 m, 13. v. 1998, conifers and various shrubs (leg. C. Lienhard) (MHNG, NHMB, dry mounted). Distribution Portugal. Madeira: Madeira Island (Aguiar & Martin 1999, as Psylla exima; this study). Host plants Portugal. Rhamnaceae: Rhamnus glandulosa Aiton (Aguiar & Martin 1999). Comments. Rhamnus glandulosa is endemic to Madeira and the Canary Islands (Aguiar & Martin 1999).	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE62936A5E75FF54C788FB6B5A60.taxon	materials_examined	Distribution Portugal. Madeira: Madeira Island (Aguiar et al. 2019; Bella & Aguiar 2020). Mainland: Porto (iNaturalist, Appendix 1).	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE62936A5E75FF54C788FB6B5A60.taxon	biology_ecology	Host plants Portugal. Elaeagnaceae: Elaeagnus x submacrophylla Servett. (Bella & Aguiar 2020, as Elaeagnus x ebbingei Boom.); E. pungens Thunb. (Aguiar et al. 2019, as E. pungens var. variegata Bean.).	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE62936A5E75FF54C48CFC905CC7.taxon	materials_examined	Material examined. Faro: 13 ♂, 20 ♀, 36 immatures, Faro, 37 ° 02 ' 24.4 " N 7 ° 58 ' 37.7 " W, 21. iv. 2021, Rhamnus lycioides subsp. oleoides (L. Neto) (BDC, dry mounted and 70 % ethanol). Distribution Portugal. Mainland: Faro (this study; iNaturalist, Appendix 1).	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE62936A5E75FF54C48CFC905CC7.taxon	biology_ecology	Host plants Portugal. Rhamnaceae: Rhamnus lycioides subsp. oleoides (L.) Jahand. & Maire (this study). Parasitoids. A specimen of Hymenoptera: Encyrtidae emerged from immatures kept in the laboratory.	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE62936A5E75FF54C48CFC905CC7.taxon	materials_examined	Material examined. Faro: 1 ♂, Faro, 37 ° 02 ' 45.7 " N 7 ° 57 ' 00.9 " W, 19. xi. 2021, Citrus sinensis (leg. B. Duarte) (BDC, dry mounted). Distribution Portugal. Mainland: Faro (this study).	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE62936A5E75FF54C48CFC905CC7.taxon	biology_ecology	Host plants Portugal. No information.	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE62936A5E75FF54C48CFC905CC7.taxon	discussion	Comments. New for Portugal. This species has been reported only from France (Lauterer & Burckhardt 1997), where adults were collected on Salix cinerea and S. caprea.	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE62936A5E75FF54C328FF5F5DE4.taxon	materials_examined	Distribution Portugal. Mainland: Guarda, Vila Real, Viseu (Sauvion et al. 2021).	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE62936A5E75FF54C328FF5F5DE4.taxon	biology_ecology	Host plants Portugal. Rosaceae: Prunus mahaleb L., P. armeniaca L. (Sauvion et al. 2021).	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE62936A5E75FF54C328FF5F5DE4.taxon	discussion	Comments. Cacopsylla pruni is a complex of two cryptic species distinguishable by molecular markers only (Sauvion et al. 2007; Peccoud et al. 2013). According to Sauvion et al. (2021), only ‘ species A’ is present in Portugal.	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE62936A5E75FF54C008FBD45F28.taxon	materials_examined	Material examined. Faro: 11 ♂, 25 ♀, 6 immatures, Faro, 37 ° 02 ' 45.7 " N 7 ° 57 ' 00.9 " W, 14. iv. 2022, Cercis siliquastrum (leg. B. Duarte) (BDC, 70 % ethanol). Distribution Portugal. Azores: Pico (Borges 2005), no details provided (Gaspar et al. 2008). Mainland: Aveiro (iNaturalist, Appendix 1); Faro (Sánchez 2011, this study); Lisbon (Sánchez 2011; iNaturalist, Appendix 1); Porto (iNaturalist, Appendix 1); Setúbal (iNaturalist, Appendix 1).	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE62936A5E75FF54C008FBD45F28.taxon	biology_ecology	Host plants Portugal. Fabaceae: Cercis siliquastrum L. (Sánchez 2011; this study).	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE62936A5E76FF54C144FD1459AC.taxon	materials_examined	Distribution Portugal. Mainland: no details provided (Coelho 1993; Torres et al. 2014); Leiria, Lisbon (Marques 2011). Madeira: Madeira Island (Aguiar & Martin 1999).	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE62936A5E76FF54C144FD1459AC.taxon	biology_ecology	Host plants Portugal. Rosaceae: Pyrus communis L. (Matias 1983; Aguiar & Martin 1999; Torres et al. 2014).	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE62936A5E76FF54C144FD1459AC.taxon	discussion	Comments. According to Matias (1983), C. pyri has been present in Portugal for a long time but emerged as a pest only in 1977 – 1978. He also suggested that C. pyrisuga is much less frequent in Portugal than C. pyri, and he drew attention to the possible occurrence of C. pyricola in Portugal. We think that the occurrence of C. bidens is more likely than that of C. pyricola. Matias (1983) described plants with symptoms associated with “ pear decline ”; the presence of ‘ Candidatus Phytoplasma pyri’ was, however, confirmed only much later in mainland Portugal (Marques 2011; Torres et al. 2014; CABI 2021).	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE6293695E76FF54C7C1FBCF5A60.taxon	materials_examined	Distribution Portugal. Mainland: Castelo Branco (Tavares 1901, as Psylla pyrisuga).	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE6293695E76FF54C7C1FBCF5A60.taxon	biology_ecology	Host plants Portugal. Rosaceae: Pyrus communis L. (Tavares 1901, Matias 1983).	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE6293695E76FF54C48DFEFC5B80.taxon	materials_examined	Material examined. Faro: 1 ♀, 1 ♀, between Casais and Cercada, 37 ° 17 ' 18.2 " N 8 ° 35 ' 57.5 " W, 9. ix. 2024, Adenocarpus hispanicus subsp. hispanicus (leg. D. Burckhardt & D. L. Queiroz) (NHMB, dry mounted). Distribution Portugal. Mainland: Faro (this study); Portalegre (Hodkinson & Hollis 1987).	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE6293695E76FF54C48DFEFC5B80.taxon	biology_ecology	Host plants Portugal. Adults were collected on Adenocarpus hispanicus subsp. hispanicus (Lam.) DC., which is a possible host.	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE6293695E76FF54C5EDFB515D0C.taxon	description	(Fig. 19)	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE6293695E76FF54C5EDFB515D0C.taxon	materials_examined	Material examined. Faro: 1 ♀, Fonte Benémola, Querença, 37 ° 12 ' 21.2 " N 8 ° 00 ' 11.5 " W, 12. v. 2024, Pyrus communis subsp. communis (leg. B. Duarte) (BDC, 70 % ethanol); 13 ♂, 10 ♀, Fonte Benémola, Querença, 37 ° 12 ' 21.2 " N 8 ° 00 ' 11.5 " W, 6. ix. 2024, Genista hirsuta (leg. D. Burckhardt, B. Duarte & D. L. Queiroz) (BDC, NHMB, dry mounted and 70 % ethanol). Distribution Portugal. No details provided (Hodkinson & Hollis 1987); Faro (iNaturalist, Appendix 1, this study); Mainland: Leiria, Porto (iNaturalist, Appendix 1).	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE6293695E76FF54C5EDFB515D0C.taxon	biology_ecology	Host plants Portugal. Adults were collected on Genista hirsuta Vahl, which is a possible host.	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE6293695E76FF54C361FBDE5DC0.taxon	materials_examined	Distribution Portugal. Mainland: Guarda (Percy 2002).	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE6293695E76FF54C361FBDE5DC0.taxon	biology_ecology	Host plants Portugal. Fabaceae: Adenocarpus complicatus (L.) J. Gay (Percy 2002).	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE6293695E77FF54C02DFD815940.taxon	materials_examined	Material examined. Faro: 2 ♂, 3 ♀, Padescas, 37 ° 18 ' 10.1 " N 8 ° 44 ' 33.0 " W, 9. ix. 2024, Genista hirsuta (leg. D. Burckhardt, B. Duarte & D. L. Queiroz) (BDC, NHMB, 70 % ethanol); 11 ♂, 15 ♀, between Aljezur and Bagagem, 37 ° 18 ' 41.8 " N 8 ° 48 ' 44.3 " W, 10. ix. 2024, G. hirsuta (leg. D. Burckhardt, B. Duarte & D. L. Queiroz) (BDC, NHMB, dry mounted and 70 % ethanol); 1 ♂, Cerca dos Pomares, Aljezur, 37 ° 19 ' 23.5 " N 8 ° 46 ' 03.7 " W, 11. ix. 2024, G. hirsuta (leg. D. Burckhardt & D. L. Queiroz) (NHMB, dry mounted). Distribution Portugal. Mainland: Coimbra (Hodkinson & Hollis 1987); Faro (this study).	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE6293695E77FF54C02DFD815940.taxon	biology_ecology	Host plants Portugal. Fabaceae: Genista triacanthos Brot. (Hodkinson & Hollis 1987); adults were also collected on G. hirsuta Vahl, a likely host. ⁂ Livilla maculipennis Hodkinson & Hollis, 1987	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE6293695E77FF54C02DFD815940.taxon	materials_examined	Material examined. Faro: 6 ♂, 13 ♀, Vale da Telha, 37 ° 18 ' 43.9 " N 8 ° 51 ' 54.0 " W, 12. ix. 2024, Genista triacanthos (leg. D. Burckhardt, B. Duarte & D. L. Queiroz) (BDC, NHMB, dry mounted and 70 % ethanol). Leiria: 1 ♂, 1 ♀, Pinhal de Leiria, Estremadura, ix. 1966 (leg. N. L. H. Krauss) (NHMB, dry mounted). Distribution Portugal. Mainland: Extremadura (this study).	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE6293695E77FF54C02DFD815940.taxon	biology_ecology	Host plants Portugal. Adults were collected on Genista triacanthos Brot., a potential host.	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE6293695E77FF54C02DFD815940.taxon	discussion	Comments. New for Portugal.	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE6293685E77FF54C7ADFD815C10.taxon	materials_examined	Material examined. Faro: 3 ♂, 3 ♀, 62 immatures, Tavira, 37 ° 09 ' 49.8 " N 7 ° 32 ' 13.2 " W, 24. iii. 2021, Retama monosperma (leg. B. Duarte) (BDC, 70 % ethanol); 4 ♂, Mata das Gambelas, 37 ° 02 ' 30.5 " N 7 ° 58 ' 29.6 " W, 7. ix. 2024, R. monosperma (leg. D. Burckhardt & D. L. Queiroz) (NHMB, 70 % ethanol); 17 ♂, 20 ♀, 2 immatures, Praia do Barril, Santa Luzia, 8. ix. 2024, R. monosperma (leg. D. Burckhardt & D. L. Queiroz) (NHMB, dry mounted and 70 % ethanol). Distribution Portugal. Madeira: no details provided (Labina et al. 2007); Mainland: No details provided (Hodkinson & Hollis, 1987); Faro (this study; iNaturalist, Appendix 1).	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE6293685E77FF54C7ADFD815C10.taxon	biology_ecology	Host plants Portugal. Retama monosperma (L.) Boiss. (this study). ⁂ Psylla foersteri (Flor, 1861) (Fig. 20)	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE6293685E77FF54C7ADFD815C10.taxon	materials_examined	Material examined. Faro: 1 ♂, 3 ♀, Várzea de Aljezur, 37 ° 18 ' 19.8 " N 8 ° 47 ' 52.1 " W, 9 – 12. ix. 2024, Alnus glutinosa (leg. D. Burckhardt, B. Duarte & D. L. Queiroz) (NHMB, BDC, dry mounted and 70 % ethanol). Distribution Portugal. Mainland: Faro (this study); Guarda, Porto (iNaturalist, Appendix 1).	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE6293685E77FF54C7ADFD815C10.taxon	biology_ecology	Host plants Portugal. No information.	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE6293685E77FF54C7ADFD815C10.taxon	discussion	Comments. New for Portugal.	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE6293685E77FF54C25CFC5C5CC4.taxon	materials_examined	Distribution Portugal. Mainland: Castelo Branco (Tavares 1901, 1905 a, as Psylla buxi).	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE6293685E77FF54C25CFC5C5CC4.taxon	biology_ecology	Host plants Portugal. Buxaceae: Buxus sempervirens L. (Tavares 1901).	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE6293685E77FF54C329FD815F94.taxon	materials_examined	Material examined. Faro: 1 ♂, 1 ♀, Várzea de Aljezur, 37 ° 18 ' 19.8 " N 8 ° 47 ' 52.1 " W, 9 – 12. ix. 2024, Solanum tuberosum (leg. D. Burckhardt & D. L. Queiroz) (NHMB, dry mounted). Distribution Portugal. Mainland: Faro (this study).	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE6293685E77FF54C329FD815F94.taxon	biology_ecology	Host plants Portugal. No information.	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE6293685E77FF54C329FD815F94.taxon	discussion	Comments. New for Portugal. ⁂ Bactericera substriola Ossiannilsson, 1992	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE6293685E77FF54C329FD815F94.taxon	materials_examined	Material examined. Faro: 4 ♂, 5 ♀, Várzea de Aljezur, 37 ° 18 ' 19.8 " N 8 ° 47 ' 52.1 " W, 9 – 12. ix. 2024, Salix eleagnos (leg. D. Burckhardt, B. Duarte & D. L. Queiroz) (BDC, NHMB, dry mounted and 70 % ethanol); 5 ♂, 7 ♀, Cerca dos Pomares, Aljezur, 37 ° 19 ' 23.5 " N 8 ° 46 ' 03.7 " W, 11. ix. 2024, S. eleagnos (leg. D. Burckhardt, B. Duarte & D. L. Queiroz) (BDC, NHMB, 70 % ethanol). Distribution Portugal. Mainland: Faro (this study; iNaturalist, Appendix 1).	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE6293685E77FF54C329FD815F94.taxon	biology_ecology	Host plants Portugal. Adults were collected on Salix eleagnos Scop., a confirmed host elsewhere.	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE6293685E77FF54C329FD815F94.taxon	discussion	Comments. New for Portugal.	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE62936F5E70FF54C6A8FB925CB4.taxon	materials_examined	Distribution Portugal. Mainland: Faro, Setúbal (Hodkinson 1981 b).	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE62936F5E70FF54C6A8FB925CB4.taxon	biology_ecology	Host plants Portugal. Apiaceae: Daucus carota L. (Hodkinson 1981 b). ⁂ Bactericera trisignata (Löw, 1886), comb. rev.	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE62936F5E70FF54C6A8FB925CB4.taxon	materials_examined	Distribution Portugal. Mainland: Leiria, Porto (iNaturalist, Appendix 1, as Trioza trisignata).	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE62936F5E70FF54C6A8FB925CB4.taxon	biology_ecology	Host plants Portugal. No information.	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE62936F5E70FF54C6A8FB925CB4.taxon	discussion	Comments. New for Portugal. Burckhardt & Lauterer (1997), following Conci & Tamanini (1984), used Phylloplecta to include Trioza trisignata and another six holarctic species, forming a putatively monophyletic group associated with Rubus spp. However, in the molecular phylogeny of Percy et al. (2018), Phylloplecta trisignata groups within Bactericera. To avoid rendering Bactericera paraphyletic by keeping Phylloplecta as a separate genus, we propose here the following synonymies: Bactericera Puton, 1876, = Phylloplecta Riley, 1884, syn. nov., = Sinitrioza Yang & Li, in Li & Yang, 1984, syn. nov. These synonymies require the following new or revived combinations: Bactericera chunghsingica Lauterer, Yang & Fang, 1988, comb. rev. (from Phylloplecta) (= replacement name for Bactericera nigra Yang, 1984 nec Bactericera nigra (Crawford, 1910); secondary homonym); B. occidentalis (Tuthill, 1939) comb. nov. (from Trioza); B. rubicola (Tuthill, 1943) comb. nov. (from Trioza); B. rubisuga (Yang & Li, in Li & Yang, 1984) comb. nov. (from Sinitrioza); B. stylifera (Patch, 1912) comb. nov. (from Trioza); B. tripunctata (Fitch, 1851) comb. nov. (from Psylla); and B. trisignata (Löw, 1886) comb. rev. (from Trioza). Klimaszewski (1973) included Trioza trisignata in Bactericera, though without providing any justification. Li’s (2011) concept of Phylloplecta contains, in addition to two species associated with Rubus, two additional species developing on Euphorbiaceae and one on Amaranthaceae. The last three species are not closely related to Bactericera and are not further discussed here. Incidentally, according to Bliven’s (1955, 1956) descriptions, the Californian Choricymoza sequoiae Bliven, 1955, seems to be a synonym of P. rubicola from Washington (Hodkinson 1988). However, Bliven (1955, 1956) published his papers privately and there remains some doubt as to whether the distribution of his papers was sufficiently wide to signify ‘ publication’ in the sense of the International Code of Zoological Nomenclature (Hodkinson 1988).	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE62936F5E70FF54C338FBD85D68.taxon	materials_examined	Distribution Portugal. Madeira: no details provided (Aguiar 2001).	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE62936F5E70FF54C338FBD85D68.taxon	biology_ecology	Host plants Portugal. Pittosporaceae: Pittosporum coriaceum Aiton (Aguiar 2001).	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE62936F5E70FF54C384FBD85E1C.taxon	materials_examined	Distribution Portugal. Madeira: no details provided (Aguiar 2001); Madeira Island (iNaturalist, Appendix 1). Host plants Portugal. Pittosporaceae: Pittosporum coriaceum Aiton (Aguiar 2001).	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE62936F5E71FF54C050FB6258D4.taxon	description	(Fig. 21)	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE62936F5E71FF54C050FB6258D4.taxon	materials_examined	Material examined. Faro: 1 ♂, Faro, 37 ° 02 ' 38.6 " N 7 ° 58 ' 30.8 " W, 15. iv. 2021, trap (leg. L. Neto) (BDC, dry mounted); 26 ♂, 53 ♀, 2 immatures, Várzea de Aljezur, 37 ° 18 ' 19.8 " N 8 ° 47 ' 52.1 " W, 9 – 12. ix. 2024, Atriplex prostata (leg. D. Burckhardt, B. Duarte & D. L. Queiroz) (BDC, NHMB, dry mounted and 70 % ethanol). Madeira: 1 ♀, Santo da Serra, 680 m, 25. x. 1992 (leg. C. Lienhard) (MHNG, dry mounted). Distribution Portugal. Madeira: Madeira Island (Loginova 1976, as Trioza chenopodii; this study); Porto Santo (Loginova 1976); Selvagens Islands (Aguiar et al. 2019). Mainland: Aveiro (iNaturalist, Appendix 1); Faro (this study).	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE62936F5E71FF54C050FB6258D4.taxon	biology_ecology	Host plants Portugal. Amaranthaceae: Atriplex prostrata Boucher ex DC., Chenopodium album L. (Loginova 1976), C. murale (L.) S. Fuentes, Uotila & Borsch, Suaeda vera Forssk. ex J. F. Gmel. (Aguiar et al. 2019).	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE62936F5E71FF54C050FB6258D4.taxon	discussion	Comments. Loginova (1976) cited the genus Atriplex as a host plant for this insect in north Madeira. According to Aguiar & Martin (1999), A. prostrata is the only species of this genus present at this locality.	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE62936E5E72FF54C17FFBE35989.taxon	materials_examined	Material examined. Faro: 1 ♂, 4 ♀, 2 immatures, Olhão, 37 ° 05 ' 03.5 " N 7 ° 47 ' 16.7 " W, 15. viii. 2020, Laurus nobilis (leg. L. Neto) (BDC, dry mounted); 1 ♂, Tavira, Campina, 37 ° 06 ' 15.6 " N 7 ° 43 ' 00.1 " W, 16. xii. 2020, Laurus nobilis (leg. B. Duarte) (BDC, 70 % ethanol); 2 ♀, between Casais and Cercada, 37 ° 17 ' 18.2 " N 8 ° 35 ' 57.5 " W, 9. ix. 2024, Laurus nobilis (leg. D. Burckhardt & D. L. Queiroz) (NHMB, dry mounted). Distribution Portugal. Azores: São Miguel (iNaturalist, Appendix 1). Mainland: Aveiro, Beja (iNaturalist, Appendix 1); Braga (Tavares 1905 a, as Trioza alacris; iNaturalist, Appendix 1); Bragança (iNaturalist, Appendix 1); Castelo Branco (Tavares 1901, as T. alacris); Coimbra (Tavares 1901, as T. alacris; iNaturalist, Appendix 1); Évora (iNaturalist, Appendix 1); Faro (this study; iNaturalist, Appendix 1); Leiria (Tavares 1901, as T. alacris; iNaturalist, Appendix 1); Lisbon (iNaturalist, Appendix 1); Portalegre (Tavares 1905 a, as T. alacris); Porto, Santarém (iNaturalist, Appendix 1); Setúbal (Tavares 1901, as T. alacris; iNaturalist, Appendix 1); Vila Real (iNaturalist, Appendix 1).	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE62936E5E72FF54C17FFBE35989.taxon	biology_ecology	Host plants Portugal. Lauraceae: Laurus nobilis L. (Tavares 1901; this study).	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE62936D5E72FF54C7E5FF545AA9.taxon	materials_examined	Distribution Portugal. Azores: Flores, Faial, Pico, Graciosa, São Jorge, Terceira, São Miguel, Santa Maria (Aguiar, 2010). Madeira: Madeira Island (Tavares 1905 b, as Trioza alacris; Aguiar & Martin 1999).	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE62936D5E72FF54C7E5FF545AA9.taxon	biology_ecology	Host plants Portugal. Lauraceae: Laurus azorica (Seub.) Franco (Aguiar & Martin 1999; Tsafack et al. 2022), L. novocanariensis Rivas Mart., Lousã, Fern. Prieto, E. Días, J. C. Costa & C. Aguiar (Tavares 1905 b, as Trioza alacris).	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE62936D5E72FF54C4C5FD815BED.taxon	materials_examined	Material examined. Faro: 4 ♀, 2 immatures (immatures lost) Faro, 37 ° 02 ' 45.7 " N 7 ° 57 ' 00.9 " W, 27. iv. 2021, Valeriana rubra (leg. B. Duarte) (BDC, dry mounted). Distribution Portugal. Mainland: Bragança (iNaturalist, Appendix 1); Faro (this study).	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE62936D5E72FF54C4C5FD815BED.taxon	biology_ecology	Host plants Portugal. Caprifoliaceae: Valeriana rubra L. (this study). Parasitoids. A parasitised immature was found from which a Hymenoptera: Encyrtidae emerged (this study).	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE62936D5E72FF54C4C5FD815BED.taxon	discussion	Comments. New for Portugal.	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE62936D5E7CFF54C201FC395AF0.taxon	materials_examined	Distribution Portugal. Mainland: Aveiro (DGAV 2024; iNaturalist, Appendix 1); Beja (DGAV 2024; Duarte et al. 2024); Braga (DGAV 2024); Coimbra (DGAV 2024; iNaturalist, Appendix 1); Faro (DGAV 2024; Duarte et al. 2024); Leiria (DGAV 2024); Lisbon (DGAV 2024; iNaturalist, Appendix 1); Porto, Santarém (DGAV 2024); Setúbal (DGAV 2024; Duarte et al. 2024; iNaturalist, Appendix 1); Viana do Castelo, Vila Real, Viseu (DGAV 2024). Madeira: Madeira Island (Aguiar et al. 2019; iNaturalist, Appendix 1); Porto Santo (Aguiar et al. 2019).	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE62936D5E7CFF54C201FC395AF0.taxon	biology_ecology	Host plants Portugal. Rutaceae: Citrus × aurantium L., Citrus × aurantium f. aurantium L., Citrus × aurantium f. deliciosa (Ten.) M. Hiroe, Citrus × limon (L.) Osbeck, Citrus trifoliata L. (Duarte 2022; Duarte et al. 2024).	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE62936D5E7CFF54C201FC395AF0.taxon	discussion	Comments. Most of the records were obtained through symptom visualisation on citrus trees as they are easy to detect. Although present in most districts of the Portuguese mainland, the distribution of T. erytreae is restricted to the western coastline. North of Lisbon it was found up to 85 km from the sea. South of Lisbon, the maximum distance from the sea was only about 15 km. This distribution is regularly updated by the Portuguese agencies of agriculture in DGAV (2024). Now, owing to successful control by the parasitoid Tamarixia dryi (Waterston) (Hymenoptera: Eulophidae), T. erytreae is hard to find in northern Portugal and has not been detected in the south since October 2021 (DGAV 2024; Duarte et al. 2024). ⁂ ‡ Trioza eugeniae Froggatt, 1901 (Figs 22 – 24, 27, 29, 30)	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE62936D5E7CFF54C201FC395AF0.taxon	materials_examined	Material examined. Faro: 2 ♂, 2 ♀, 4 immatures, Aljezur, 37 ° 18 ' 46.4 " N 8 ° 47 ' 30.1 " W, 2. v. 2021, Melaleuca sp. (leg. B. Duarte) (BDC, dry mounted); 1 ♀, Tavira, Brejo, 37 ° 06 ' 13.8 " N 7 ° 44 ' 15.2 " W, 28. iv. 2021, Citrus sinensis (leg. B. Duarte) (BDC, dry mounted); 1 ♀, Praia do Barril, Santa Luzia, 8. ix. 2024, M. armillaris (leg. B. Duarte) (BDC, 70 % ethanol); 2 ♂, 1 ♀, between Casais and Cercada, 37 ° 17 ' 18.2 " N 8 ° 35 ' 57.5 " W, garden, hedges, vegetation along unsurfaced road, Quercus suber plantation, M. armillaris (leg. D. Burckhardt & D. L. Queiroz) (NHMB, dry mounted); 3 ♂, 5 ♀, Várzea de Aljezur, 37 ° 18 ' 19.8 " N 8 ° 47 ' 52.1 " W, 9 – 12. ix. 2024, M. armillaris (leg. D. Burckhardt & D. L. Queiroz) (NHMB, 70 % ethanol); 37 ♂, 33 ♀, 8 immatures, Vale da Telha, Arrifana, 37 ° 18 ' 43.9 " N 8 ° 51 ' 54.0 " W, 12. ix. 2024, maquis shrubland between garden and houses, M. armillaris (leg. D. Burckhardt, B. Duarte & D. L. Queiroz) (BDC, NHMB, dry mounted and 70 % ethanol). Distribution Portugal. Mainland: Faro (this study); Santarém (iNaturalist, Appendix 1).	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE62936D5E7CFF54C201FC395AF0.taxon	biology_ecology	Host plants Portugal. Myrtaceae: Melaleuca armillaris (Sol. ex Gaertn.) Sm. (this study) (Figs 25, 26). Parasitoids. Some parasitised immatures were observed (Duarte 2022) but no hymenopterans emerged for obtaining an identification.	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE62936D5E7CFF54C201FC395AF0.taxon	discussion	Comments. New for Europe and Portugal. Trioza eugeniae is an Australian species that was confused with T. adventicia Tuthill, 1952 prior to the study by Taylor & Martoni (2019). The latter species, also of Australian origin, was described from New Zealand (Tuthill 1952) and has been reported from the USA (as T. eugeniae). Taylor & Martoni (2019) redescribed the two species and suggested that T. curta (Ferris & Klyver, 1932), another Myrtaceae-inhabiting species described from New Zealand, is morphologically identical to T. eugeniae but differs slightly at the molecular level. Here we list some additional characters by which T. eugeniae can be separated from T. adventicia: male proctiger massive (Fig. 27) (versus relatively slender, Fig. 28); male subgenital plate, in lateral view, subglobular (Fig. 27) (versus triangular, Fig. 28); paramere with a long slender apex (Figs 27, 29) (versus short and blunt, Figs 28, 31); proximal segment of aedeagus wide in distal three-quarters (Fig. 27) (versus narrow, Fig. 28); and distal segment of aedeagus slender apically (Fig. 30) (versus inflated, Fig. 32). The characters given here for T. eugeniae fit also the description of T. curta by Ferris & Klyver (1932), confirming the conclusion by Taylor & Martoni (2019) that the two species are morphologically almost identical. Taylor & Martoni (2019) listed Syzygium smithii (Poir.) Nied. (Syzygieae) (type series) and Backhousia myrtifolia Hook. & Harv. (Backhousieae) as confirmed hosts of T. eugeniae. Here we add Melaleuca armillaris (Myrtaceae). The three host taxa are not closely related, making T. eugeniae a widely oligophagous species, a feature also present in the closely related T. adventicia (Taylor & Martoni 2019). Trioza eugeniae has been observed in Portugal for about four or five years (B. Z. Duarte, G. Duarte, pers. obs.), where it was found to be always associated with Melaleuca armillaris, not inducing any galls, never with any other Myrtaceae. Melaleuca armillaris originates from Australia (New South Wales, South Australia, Tasmania, Victoria) and was introduced into continental Portugal as an ornamental plant.	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE6293635E7CFF54C57CFE955D54.taxon	materials_examined	Material examined. Faro: Galls on leaves, Aljezur, 37 ° 17 ' 04.2 " N 8 ° 48 ' 20.2 " W, 2. v. 2021, Rhamnus alaternus (leg. B. Duarte) (BDC, 70 % ethanol); 8 immatures, 2 skins, several galls on leaves, Fonte de Benémola, Querença, 37 ° 12 ' 21.2 " N 8 ° 00 ' 11.5 " W, 6. ix. 2024, R. alaternus (leg. D. Burckhardt, B. Duarte & D. L. Queiroz) (BDC, NHMB, dry mounted and 70 % ethanol); 5 immatures, 3 skins, several galls in leaves, between Casais and Cercada, 37 ° 17 ' 18.2 " N 8 ° 35 ' 57.5 " W, 9. ix. 2024, garden, hedges, vegetation along unsurfaced road, Quercus suber plantation, R. lycioides (leg. D. Burckhardt, B. Duarte & D. L. Queiroz) BDC, (NHMB, dry mounted and 70 % ethanol). Distribution Portugal. Mainland: Aveiro, Castelo Branco, Coimbra (iNaturalist, Appendix 1); Faro (this study; iNaturalist, Appendix 1); Leiria (Tavares 1905 a, as Asterolecanium rhamni; iNaturalist, Appendix 1); Lisbon (Tavares 1905 a; iNaturalist, Appendix 1); Portalegre, Santarém (iNaturalist, Appendix 1); Setúbal (Tavares 1905 a; iNaturalist, Appendix 1).	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE6293635E7CFF54C57CFE955D54.taxon	biology_ecology	Host plants Portugal. Rhamnaceae: Rhamnus alaternus L. (Tavares 1905 a; this study; iNaturalist, Appendix 1); R. lycioides L. (this study). Parasitoids. Confirmed in the laboratory, but emerged hymenopteran parasitoids are still in need of identification.	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE6293635E7CFF54C398FD4B5F70.taxon	materials_examined	Material examined. Faro: 1 ♂, Tavira, 37 ° 09 ' 49.8 " N 7 ° 32 ' 13.2 " W, 24. iii. 2021, Myoporum acuminatum (leg. B. Duarte) (BDC, dry mounted); 1 ♀, Tavira, Buraco, 37 ° 09 ' 49.8 " N 7 ° 32 ' 13.2 " W, 24. iii. 2021, Rosa sp. (leg. B. Duarte) (BDC, dry mounted); 1 ♀, Tavira, Buraco, 37 ° 09 ' 49.8 " N 7 ° 32 ' 13.2 " W, 24. iii. 2021, Morus nigra (leg. B. Duarte) (BDC, dry mounted); 1 ♂, 1 ♀, Várzea de Aljezur, 37 ° 18 ' 19.8 " N 8 ° 47 ' 52.1 " W, 9 – 12. ix. 2024, Urtica dioica (leg. D. Burckhardt & D. L. Queiroz) (NHMB, 70 % ethanol). Distribution Portugal. Madeira: Madeira Island (Tavares 1903 a, 1905 b; Loginova 1976). Mainland: Aveiro (iNaturalist, Appendix 1); Castelo Branco (Tavares 1907); Coimbra (iNaturalist, Appendix 1); Faro (this study); Guarda, Porto (iNaturalist, Appendix 1).	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE6293635E7CFF54C398FD4B5F70.taxon	biology_ecology	Host plants Portugal. Urticaceae: Urtica dioica L. (Tavares 1907), U. membranacea Poir. ex Savigny (Tavares 1903 a, 1905 b).	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
03EEAE6293635E7CFF54C398FD4B5F70.taxon	discussion	Comments. Both host plant species are widely distributed on the Portuguese mainland (Flora-On 2024), suggesting that this psyllid is widely distributed also.	en	Duarte, Beatriz Zarcos, Burckhardt, Daniel, Ouvrard, David, Queiroz, Dalva L., Valkenburg, Thijs, Neto, Luís (2025): The Psylloidea (Hemiptera) of Portugal (mainland, Azores and Madeira): checklist and new distributional and host records. Zootaxa 5642 (4): 343-385, DOI: 10.11646/zootaxa.5642.4.4, URL: https://doi.org/10.11646/zootaxa.5642.4.4
