taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
03EC0B35C511967EFF3B3BE0932CFC32.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16983920/files/figure.png	https://doi.org/10.5281/zenodo.16983920	FIGURE 2. Dilobitarsus lambirensis Arimoto, sp. nov. holotype, male (AA1214). A: Habitus, dorsal view; B: habitus, ventral view; C: scale-like setae on elytra; D: apical tarsomeres of left midleg (arrow: a thick seta at base of outsides of tarsal claws).	FIGURE 2. Dilobitarsus lambirensis Arimoto, sp. nov. holotype, male (AA1214). A: Habitus, dorsal view; B: habitus, ventral view; C: scale-like setae on elytra; D: apical tarsomeres of left midleg (arrow: a thick seta at base of outsides of tarsal claws).	2025-08-13	Arimoto, Kôichi;Kaliang, Clement Het;Mizuno, Takafumi;Meleng, Paulus;Kishimoto-Yamada, Keiko;Itioka, Takao		Zenodo	biologists	Arimoto, Kôichi;Kaliang, Clement Het;Mizuno, Takafumi;Meleng, Paulus;Kishimoto-Yamada, Keiko;Itioka, Takao			
03EC0B35C511967EFF3B3BE0932CFC32.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16983922/files/figure.png	https://doi.org/10.5281/zenodo.16983922	FIGURE 3. Dilobitarsus lambirensis haye, sp. nov., male. A: Head and anterior part of pronotum, dorsal view; B: head and anterior part of prothorax, anterior view (arrow: mandible bidentate); C: head and anterior part of prosternum, ventral view; D: head, prothorax and mesothorax, lateral view (white arrow: anterior tubercle, black arrow: posterior tubercle); E: right antenna, dorsal side; F: left maxillary palp, ventral view; G: habitus, dorsolateral view (white arrows:two anterior tubercles, black arrows: two posterior tubercles); H: head, prothorax and mesothorax, ventrolateral view (arrows: posterior edge of hypomeron with three notches mesally); I: scutellar shield from the direction perpendicular to dorsal surface; J: scutellum and elytra humerus, dorsolateral view (arrow: elytral humerus with a longitudinal carina); K: mesothorax and anterior part of metathorax, ventral view. A–E, G-J: Holotype (AA1214); F: paratype (AA1217); K: paratype (AA1220).	FIGURE 3. Dilobitarsus lambirensis haye, sp. nov., male. A: Head and anterior part of pronotum, dorsal view; B: head and anterior part of prothorax, anterior view (arrow: mandible bidentate); C: head and anterior part of prosternum, ventral view; D: head, prothorax and mesothorax, lateral view (white arrow: anterior tubercle, black arrow: posterior tubercle); E: right antenna, dorsal side; F: left maxillary palp, ventral view; G: habitus, dorsolateral view (white arrows:two anterior tubercles, black arrows: two posterior tubercles); H: head, prothorax and mesothorax, ventrolateral view (arrows: posterior edge of hypomeron with three notches mesally); I: scutellar shield from the direction perpendicular to dorsal surface; J: scutellum and elytra humerus, dorsolateral view (arrow: elytral humerus with a longitudinal carina); K: mesothorax and anterior part of metathorax, ventral view. A–E, G-J: Holotype (AA1214); F: paratype (AA1217); K: paratype (AA1220).	2025-08-13	Arimoto, Kôichi;Kaliang, Clement Het;Mizuno, Takafumi;Meleng, Paulus;Kishimoto-Yamada, Keiko;Itioka, Takao		Zenodo	biologists	Arimoto, Kôichi;Kaliang, Clement Het;Mizuno, Takafumi;Meleng, Paulus;Kishimoto-Yamada, Keiko;Itioka, Takao			
03EC0B35C511967EFF3B3BE0932CFC32.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16983924/files/figure.png	https://doi.org/10.5281/zenodo.16983924	FIGURE 4. Dilobitarsus lambirensis Arimoto, sp. nov., male. A: Left elytron, lateral view; B: left elytron, ventral view; C: left hind wing, dorsal view; D: left mid tarsus, lateral view (arrow: ventral lobe of tarsomere IV); E: tergite VIII, dorsal view; F: sternite VIII, ventral view; G: tergites IX–X, dorsal view; H: sternite IX, ventral view; I: aedeagus, dorsal view; J: aedeagus, ventral view; K: apical part of right paramere, dorsal view (arrows: proximal and distal apices); L: apical part of left paramere, ventral view (arrow: apex beyond lateral subapical barb narrowed in ventral side). A–C: paratype (AA1218); D: paratype (AA1216), except for tarsomere V and claw from holotype (AA1214); E–L holotype (AA1214).	FIGURE 4. Dilobitarsus lambirensis Arimoto, sp. nov., male. A: Left elytron, lateral view; B: left elytron, ventral view; C: left hind wing, dorsal view; D: left mid tarsus, lateral view (arrow: ventral lobe of tarsomere IV); E: tergite VIII, dorsal view; F: sternite VIII, ventral view; G: tergites IX–X, dorsal view; H: sternite IX, ventral view; I: aedeagus, dorsal view; J: aedeagus, ventral view; K: apical part of right paramere, dorsal view (arrows: proximal and distal apices); L: apical part of left paramere, ventral view (arrow: apex beyond lateral subapical barb narrowed in ventral side). A–C: paratype (AA1218); D: paratype (AA1216), except for tarsomere V and claw from holotype (AA1214); E–L holotype (AA1214).	2025-08-13	Arimoto, Kôichi;Kaliang, Clement Het;Mizuno, Takafumi;Meleng, Paulus;Kishimoto-Yamada, Keiko;Itioka, Takao		Zenodo	biologists	Arimoto, Kôichi;Kaliang, Clement Het;Mizuno, Takafumi;Meleng, Paulus;Kishimoto-Yamada, Keiko;Itioka, Takao			
03EC0B35C511967EFF3B3BE0932CFC32.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16983916/files/figure.png	https://doi.org/10.5281/zenodo.16983916	FIGURE 1. Collection sites of the southeast Asian Dilobitarsus species. Circles: D. pendleburyi Fleutiaux, 1934; Triangle: D. lambirensis Arimoto, sp. nov. A: Bettotan, Sabah, Malaysia (Fleutiaux, 1934); B: Fraser’s Hill, Pahang, Malaysia (Arimoto, 2016); C: Harau Valley, West Sumatra, Indonesia (Arimoto, 2016); D: Hutan Harapan, Jambi, Indonesia (Kasmiatun et al., 2020); E: Lambir Hills National Park, Miri, Sarawak, Malaysia (this study).	FIGURE 1. Collection sites of the southeast Asian Dilobitarsus species. Circles: D. pendleburyi Fleutiaux, 1934; Triangle: D. lambirensis Arimoto, sp. nov. A: Bettotan, Sabah, Malaysia (Fleutiaux, 1934); B: Fraser’s Hill, Pahang, Malaysia (Arimoto, 2016); C: Harau Valley, West Sumatra, Indonesia (Arimoto, 2016); D: Hutan Harapan, Jambi, Indonesia (Kasmiatun et al., 2020); E: Lambir Hills National Park, Miri, Sarawak, Malaysia (this study).	2025-08-13	Arimoto, Kôichi;Kaliang, Clement Het;Mizuno, Takafumi;Meleng, Paulus;Kishimoto-Yamada, Keiko;Itioka, Takao		Zenodo	biologists	Arimoto, Kôichi;Kaliang, Clement Het;Mizuno, Takafumi;Meleng, Paulus;Kishimoto-Yamada, Keiko;Itioka, Takao			
03EC0B35C5149671FF3B38809157FD3E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16983930/files/figure.png	https://doi.org/10.5281/zenodo.16983930	FIGURE 5. Dilobitarsus pendleburyi Fleutiaux, 1934. A: Male, dorsal view (DPF01); B: female, dorsal view (AA1223); C: female, ventral view (AA1223).	FIGURE 5. Dilobitarsus pendleburyi Fleutiaux, 1934. A: Male, dorsal view (DPF01); B: female, dorsal view (AA1223); C: female, ventral view (AA1223).	2025-08-13	Arimoto, Kôichi;Kaliang, Clement Het;Mizuno, Takafumi;Meleng, Paulus;Kishimoto-Yamada, Keiko;Itioka, Takao		Zenodo	biologists	Arimoto, Kôichi;Kaliang, Clement Het;Mizuno, Takafumi;Meleng, Paulus;Kishimoto-Yamada, Keiko;Itioka, Takao			
03EC0B35C5149671FF3B38809157FD3E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16983933/files/figure.png	https://doi.org/10.5281/zenodo.16983933	FIGURE 6. Dilobitarsus pendleburyi Fleutiaux, 1934.A: Head and anterior part of pronotum, dorsal view; B: head and anterior part of prothorax, anterior view; C: head and anterior part of prosternum, ventral view; D: head, prothorax and mesothorax, lateral view (white arrow: anterior tubercle, black arrows: posterior tubercles); E: right antenna, dorsal side; F: left antenna, dorsal side; G: left maxillary palp, dorsal view; H: habitus, dorsolateral view (white arrows: two anterior tubercles, black arrows: four posterior tubercles); I: head, prothorax and mesothorax, ventrolateral view (arrow: prosternal carina extending anterad of coxal cavities); J: scutellar shield from the direction perpendicular to dorsal surface; K: scutellum and elytra humerus, dorsolateral view; L: mesothorax and anterior part of metathorax, ventral view. A–D, F, G–K: female (AA1223); E, L: male (DPF01).	FIGURE 6. Dilobitarsus pendleburyi Fleutiaux, 1934.A: Head and anterior part of pronotum, dorsal view; B: head and anterior part of prothorax, anterior view; C: head and anterior part of prosternum, ventral view; D: head, prothorax and mesothorax, lateral view (white arrow: anterior tubercle, black arrows: posterior tubercles); E: right antenna, dorsal side; F: left antenna, dorsal side; G: left maxillary palp, dorsal view; H: habitus, dorsolateral view (white arrows: two anterior tubercles, black arrows: four posterior tubercles); I: head, prothorax and mesothorax, ventrolateral view (arrow: prosternal carina extending anterad of coxal cavities); J: scutellar shield from the direction perpendicular to dorsal surface; K: scutellum and elytra humerus, dorsolateral view; L: mesothorax and anterior part of metathorax, ventral view. A–D, F, G–K: female (AA1223); E, L: male (DPF01).	2025-08-13	Arimoto, Kôichi;Kaliang, Clement Het;Mizuno, Takafumi;Meleng, Paulus;Kishimoto-Yamada, Keiko;Itioka, Takao		Zenodo	biologists	Arimoto, Kôichi;Kaliang, Clement Het;Mizuno, Takafumi;Meleng, Paulus;Kishimoto-Yamada, Keiko;Itioka, Takao			
03EC0B35C5149671FF3B38809157FD3E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16983935/files/figure.png	https://doi.org/10.5281/zenodo.16983935	FIGURE 7. Dilobitarsus pendleburyi Fleutiaux, 1934. A: Left elytron, lateral view (arrow: elytron abruptly depressed after strongly elevated tubercle); B: left elytron, ventral view (arrow: underside of tubercle); C: left hind wing, dorsal view; D: left mid tarsus, lateral view (arrow: ventral lobe of tarsomere IV); E: tergite VIII, dorsal view; F: sternite VIII, ventral view; G: tergites IX–X, dorsal view; H: sternite IX, ventral view; I: aedeagus, dorsal view; J: aedeagus, ventral view; K: apical part of right paramere, dorsal view (arrows: proximal and distal apices); L: apical part of left paramere, ventral view (arrow: apex beyond lateral subapical barb narrowed in ventral side). A, B, D: female (AA1223); C, E–L male (DPF01).	FIGURE 7. Dilobitarsus pendleburyi Fleutiaux, 1934. A: Left elytron, lateral view (arrow: elytron abruptly depressed after strongly elevated tubercle); B: left elytron, ventral view (arrow: underside of tubercle); C: left hind wing, dorsal view; D: left mid tarsus, lateral view (arrow: ventral lobe of tarsomere IV); E: tergite VIII, dorsal view; F: sternite VIII, ventral view; G: tergites IX–X, dorsal view; H: sternite IX, ventral view; I: aedeagus, dorsal view; J: aedeagus, ventral view; K: apical part of right paramere, dorsal view (arrows: proximal and distal apices); L: apical part of left paramere, ventral view (arrow: apex beyond lateral subapical barb narrowed in ventral side). A, B, D: female (AA1223); C, E–L male (DPF01).	2025-08-13	Arimoto, Kôichi;Kaliang, Clement Het;Mizuno, Takafumi;Meleng, Paulus;Kishimoto-Yamada, Keiko;Itioka, Takao		Zenodo	biologists	Arimoto, Kôichi;Kaliang, Clement Het;Mizuno, Takafumi;Meleng, Paulus;Kishimoto-Yamada, Keiko;Itioka, Takao			
03EC0B35C5149671FF3B38809157FD3E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16983939/files/figure.png	https://doi.org/10.5281/zenodo.16983939	FIGURE 8. Dilobitarsus pendleburyi Fleutiaux, 1934, female (AA1223). A: Tergite VIII, dorsal view; B: sternite VIII, ventral view; C: sternite VIII and genitalia, dorsal view; D: apical part of ovipositor, ventral view; E: bursa copulatrix, dorsal view; F: bursa copulatrix, lateral view (arrows: colleterial glands, one of them is lost).	FIGURE 8. Dilobitarsus pendleburyi Fleutiaux, 1934, female (AA1223). A: Tergite VIII, dorsal view; B: sternite VIII, ventral view; C: sternite VIII and genitalia, dorsal view; D: apical part of ovipositor, ventral view; E: bursa copulatrix, dorsal view; F: bursa copulatrix, lateral view (arrows: colleterial glands, one of them is lost).	2025-08-13	Arimoto, Kôichi;Kaliang, Clement Het;Mizuno, Takafumi;Meleng, Paulus;Kishimoto-Yamada, Keiko;Itioka, Takao		Zenodo	biologists	Arimoto, Kôichi;Kaliang, Clement Het;Mizuno, Takafumi;Meleng, Paulus;Kishimoto-Yamada, Keiko;Itioka, Takao			
03EC0B35C5149671FF3B38809157FD3E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16983916/files/figure.png	https://doi.org/10.5281/zenodo.16983916	FIGURE 1. Collection sites of the southeast Asian Dilobitarsus species. Circles: D. pendleburyi Fleutiaux, 1934; Triangle: D. lambirensis Arimoto, sp. nov. A: Bettotan, Sabah, Malaysia (Fleutiaux, 1934); B: Fraser’s Hill, Pahang, Malaysia (Arimoto, 2016); C: Harau Valley, West Sumatra, Indonesia (Arimoto, 2016); D: Hutan Harapan, Jambi, Indonesia (Kasmiatun et al., 2020); E: Lambir Hills National Park, Miri, Sarawak, Malaysia (this study).	FIGURE 1. Collection sites of the southeast Asian Dilobitarsus species. Circles: D. pendleburyi Fleutiaux, 1934; Triangle: D. lambirensis Arimoto, sp. nov. A: Bettotan, Sabah, Malaysia (Fleutiaux, 1934); B: Fraser’s Hill, Pahang, Malaysia (Arimoto, 2016); C: Harau Valley, West Sumatra, Indonesia (Arimoto, 2016); D: Hutan Harapan, Jambi, Indonesia (Kasmiatun et al., 2020); E: Lambir Hills National Park, Miri, Sarawak, Malaysia (this study).	2025-08-13	Arimoto, Kôichi;Kaliang, Clement Het;Mizuno, Takafumi;Meleng, Paulus;Kishimoto-Yamada, Keiko;Itioka, Takao		Zenodo	biologists	Arimoto, Kôichi;Kaliang, Clement Het;Mizuno, Takafumi;Meleng, Paulus;Kishimoto-Yamada, Keiko;Itioka, Takao			
