taxonID	type	description	language	source
03ED5C1AFFCAFA5894CFDD06FA0954B5.taxon	discussion	We define here the serraticornis group as those species of Lycoderides in which the scutellum is concealed by the pronotum (i. e., not externally visible), and the forewings are at least basally sclerotized and punctuated, with the remaining portion partially transparent hyaline and speckled. This group includes L. abditus, L. aburraensis sp. nov., L. phi, L. santainensis sp. nov., and L. serraticornis. Lycoderides serraticornis, L. santainensis, and L. abditus exhibit strong sexual dimorphism, with females bearing an anterior pronotal process, while males lack this process and instead present a dorsal elevation. In contrast, both males and females of Lycoderides phi share a similar morphology, each possessing an anterior process resembling that of the females of the aforementioned species (Lapèze & Lopez-Vaamonde 2024). L. aburraensis is currently known only from a male, which exhibits a morphology consistent with the males of the other species in this group (except L. phi).	en	Flórez-V, Camilo, Lapèze, Jérémie, Urban, Julie M. (2025): Taxonomic and ecological notes on Lycoderides Sakakibara (Hemiptera: Membracidae), including two new species from the highlands of the Colombian Andes. Zootaxa 5665 (2): 151-186, DOI: 10.11646/zootaxa.5665.2.1, URL: https://doi.org/10.11646/zootaxa.5665.2.1
03ED5C1AFFCBFA5A94CFDD66FBC856E0.taxon	description	(Fig. 2 A – B, 4 A – B, 5 A – B, 7)	en	Flórez-V, Camilo, Lapèze, Jérémie, Urban, Julie M. (2025): Taxonomic and ecological notes on Lycoderides Sakakibara (Hemiptera: Membracidae), including two new species from the highlands of the Colombian Andes. Zootaxa 5665 (2): 151-186, DOI: 10.11646/zootaxa.5665.2.1, URL: https://doi.org/10.11646/zootaxa.5665.2.1
03ED5C1AFFCBFA5A94CFDD66FBC856E0.taxon	discussion	Remarks: Lycoderides abditus was described based on a male specimen from French Guiana (Fig. 7), collected using an interception trap (vitre) at Mont Itoupé (570 masl) (Sakakibara, 2013). Recent sampling and research in French Guiana by JL led to the collection of additional male specimens of L. abditus, as well as other specimens that were presumed to be females based on morphological similarities with males (i. e., scutellum covered by pronotum, dorsal contour, and forewing coloration), and their resemblance to females of L. serraticornis and L. santainensis sp. nov. (i. e., anterior process shape). Subsequent DNA barcoding analyses confirmed that these females belong to L. abditus (see Fig. 1 & Table 1 in Lapèze & Lopez-Vaamonde 2024). Five of these presumed L. abditus females were barcoded using the “ single-molecule real-time sequencing ” method. All sequences obtained matched the Barcode Index Number of the male L. abditus: BOLD: AEC 2962 (see Fig. 1 & Table 1 in Lapèze & Lopez-Vaamonde 2024). DNA sequence data and images are available in BOLD (dx. doi. org / 10.5883 / DS-LYCODER). The females of L. abditus closely resemble those of L. santainensis sp. nov. and L. serraticornis, just as the males of L. abditus are similar to those of L. aburraensis sp. nov., L. santainensis sp. nov., and L. serraticornis. Known localities are situated between 0 and 350 masl; most specimens were captured using light traps (mercury lamps), but no in situ behavioral observations could be made, and the natural history of this species remains unknown. Additionally, Sakakibara (2013) indicated the holotype repository is MNHN (Muséum National d’Histoire Naturelle, Paris, France), but it is currently housed in DZUP, Curitiba, Brazil.	en	Flórez-V, Camilo, Lapèze, Jérémie, Urban, Julie M. (2025): Taxonomic and ecological notes on Lycoderides Sakakibara (Hemiptera: Membracidae), including two new species from the highlands of the Colombian Andes. Zootaxa 5665 (2): 151-186, DOI: 10.11646/zootaxa.5665.2.1, URL: https://doi.org/10.11646/zootaxa.5665.2.1
03ED5C1AFFCBFA5A94CFDD66FBC856E0.taxon	diagnosis	Diagnosis: Male (Fig. 2 A, 4 A, 5 A, 7). Pronotum elevated above the head, 1.5 x head length in frontal view, descending behind the humeral angles; suprahumeral horns small, wart-like; without anteroventral carina below the suprahumeral horns; metopidium slightly reclined anteriorly; pronotum with a wax-like line adjacent to the supraocular callosity. Forewing sparsely speckled at the apical third, with veins lacking warts. Female. Variegated light brown to dark brown. Waxy pubescence present on the pronotum, extending from the anterior margin, crossing posterior to the supraocular callosities, and reaching the posterior area of the metopidium. Pronotum elevated above the head, 2 x head length in frontal view; pronotal anterior process 0.5 x pronotum height, with a sinuous posterior margin. In lateral view, the subapical crest of the posterior process is well elevated. Forewing veins lacking warts.	en	Flórez-V, Camilo, Lapèze, Jérémie, Urban, Julie M. (2025): Taxonomic and ecological notes on Lycoderides Sakakibara (Hemiptera: Membracidae), including two new species from the highlands of the Colombian Andes. Zootaxa 5665 (2): 151-186, DOI: 10.11646/zootaxa.5665.2.1, URL: https://doi.org/10.11646/zootaxa.5665.2.1
03ED5C1AFFCBFA5A94CFDD66FBC856E0.taxon	description	Description: Female (Fig. 2 B, 4 B, 5 B): Color: Variegated light brown to dark brown. Post-clypeus and contiguous area on vertex and supra-antennal ledges dark-brown, posterior half of vertex light brown. Eyes and ocelli brownish gray. Pronotum dark brown, with warts, suprahumeral horns carinae and dorso-medial carina yellow; supraocullar callosities brown; humeral angles and adjacent area over basal ¼ of forewing clavus, as well as the subapical depression and tip of the posterior pronotal process, light brown to brown. Forewing sclerotized and punctuated up to half its length; basal ¼ brown, followed by a transversal dark brown band extending through the basal 1 / 3, with a yellow triangular area posteriorly; apical half of the forewing membrane is opaque hyaline, sparsely speckled with brown; veins brown. Legs light brown; femora and tarsomeres brown; coxae brown. Sculpture: densely and coarsely punctuated, giving an overall opaque appearance. Light brown pubescence, sparsely distributed on the pronotum, sclerotized areas of the forewing, dorsal surface of the tibiae, and tarsomeres. Waxy pubescence present on pronotum, extending from anterior margin, crossing posterior to the supraocular callosities, and reaching the posterior area of the metopidium, as well as along the mesothorax margin, femora, and tibiae. Wart-like protuberances scattered across the anterior pronotal process, metopidium, and the area surrounding the humeral angles. Thorax: Pronotum in frontal view (Fig. 2 B) more or less triangular, with an anterior process elevated 1 x head length above the metopidium; suprahumeral horns triangular and carinate, each carina surrounded by warts; dorso-median carina extending distinctly from the anterior margin of the metopidium to the posterior process; in lateral view (Fig. 3 B), pronotal horn directed obliquely antero-dorsally, with sinuous dorsal margin descending to the posterior third, then forming a small crest before reaching the posterior margin of the posterior process, which extends above the claval apex; in dorsal view (Fig. 4 B), suprahumeral horns small and triangular. Scutellum completely covered. Tibiae with flattened dorsal surface. Measurements: male / female (mm): Body length: 5.8 / 6.85; pronotum length: 4.55 / 5.13; pronotum height: 1.83 / 2.74; forewing length: 4.91 / 5.89; width between humeral angles: 2.05 / 2.23; head width: 1.81 / 1.99; vertex length: 0.64 / 0.74; vertex width: 1.21 / 1.37.	en	Flórez-V, Camilo, Lapèze, Jérémie, Urban, Julie M. (2025): Taxonomic and ecological notes on Lycoderides Sakakibara (Hemiptera: Membracidae), including two new species from the highlands of the Colombian Andes. Zootaxa 5665 (2): 151-186, DOI: 10.11646/zootaxa.5665.2.1, URL: https://doi.org/10.11646/zootaxa.5665.2.1
03ED5C1AFFCBFA5A94CFDD66FBC856E0.taxon	materials_examined	Examined material: FRENCH GUIANA: Holotype male in DZUP: “ GUYANE FRANC ̧ AISE | Itoupe ́ — DZ 570 m | N 03 º 01 ’ W 053 º 05 ’ | 24 / 05 / 2010 | (vitre) — S. E. A. G. ”. Other specimens examined: “ 11 / 10 / 2018; Piège Lumineux; Montagne de Kaw, Route de Kaw, Carrefour Fourgassier, Guyane Française, J. Lapèze rec ” (1 male in J. Lapèze collection); “ 20 / 06 / 2014; GF [Guyane française]. Montagne de Kaw, Route de Kaw, Carrefour Fourgassier; PL [piège lumineux]; F. Robin rec ” (1 female in J. Lapèze collection); “ 09 / 06 / 2021; GF [Guyane française]. Montagne de Kaw, Route de Kaw, Carrefour Fourgassier; PL [piège lumineux]; J. Lapèze rec ” (1 female in J. Lapèze collection); “ 27 / 05 / 2020; GF [Guyane française]. Roura, Montagne de Kaw, Réserve Naturelle Trésor, parking; piège lumineux; J. Lapèze leg. ” (1 male in J. Lapèze collection); “ 04 / 09 / 2015; Montagne de Kaw, Route de Kaw, Carrefour Fourgassier, Guyane Française; PL [piège lumineux]; F. Sonzogni rec ” (1 female in J. Lapèze collection); “ 03 / 09 / 2021; GF [Guyane française]. Réserve Naturelle de La Trinité, Mont Tabulaire, DZ 300 m; PL [piège lumineux]; J. Lapèze leg. (SEAG) ” (1 female in J. Lapèze collection); “ 20 / 05 / 2023; GF [Guyane française]. Papaïchton, Crique Banba; PL [piège lumineux]; J. Lapèze & N. Giraud-Audine leg. ” (1 female in J. Lapèze collection); “ 17 / 09 / 2023; GF [Guyane française]. Réserve Naturelle de La Trinité, Mont Tabulaire, DZ 300 m; PL [piège lumineux]; SEAG leg. ” (1 male and 1 female in J. Lapèze collection); “ 30 / 06 / 2000, Montagne de Kaw, Route de Kaw pk 37,5, Guyane Française ” (2 females in D. Faure collection); “ 28 / 01 / 1989; Piste Bélizon pk 22; Guyane Française ” (5 females in M. Duranton collection); “ 23 / 09 / 1989; Piste Saint-Elie; Guyane Française ” (1 female in M. Duranton collection).	en	Flórez-V, Camilo, Lapèze, Jérémie, Urban, Julie M. (2025): Taxonomic and ecological notes on Lycoderides Sakakibara (Hemiptera: Membracidae), including two new species from the highlands of the Colombian Andes. Zootaxa 5665 (2): 151-186, DOI: 10.11646/zootaxa.5665.2.1, URL: https://doi.org/10.11646/zootaxa.5665.2.1
03ED5C1AFFCBFA5A94CFDD66FBC856E0.taxon	distribution	Distribution: FRENCH GUIANA: Mont Itoupé (loc. type; 3.022222 ºN, 53.094722 ºW, 600 m), Montagne de Kaw (4.642848 ºN, 52.299772 ºW, 243 m), Forêt de Coralie (4.495463 ºN, 52.395143 ºW, 50 m), Piste Bélizon (4.244238 ºN, 52.397345 ºW, 120 m), Crique Banba (3.793853 ºN, 54.110389 ºW, 120 m), Mont Tabulaire de La Trinité (4.610861 ºN, 53.358896 ºW, 300 m), Piste de Saint-Elie (5.297495 ºN, 53.052031 ºW, 45 m). Flórez-V et al. (2015) erroneously recorded this species for Colombia (see below in L. serraticornis).	en	Flórez-V, Camilo, Lapèze, Jérémie, Urban, Julie M. (2025): Taxonomic and ecological notes on Lycoderides Sakakibara (Hemiptera: Membracidae), including two new species from the highlands of the Colombian Andes. Zootaxa 5665 (2): 151-186, DOI: 10.11646/zootaxa.5665.2.1, URL: https://doi.org/10.11646/zootaxa.5665.2.1
03ED5C1AFFC9FA5C94CFDCD2FD525648.taxon	description	urn: lsid: zoobank. org: act: F 3 BBE 447 - C 3 C 3 - 4 F 46 - B 92 A-C 6 A 66 E 82 A 4 C 1 (Fig. 3 C, 4 G, 5 G, 6 E, 12 F – J, 16 A)	en	Flórez-V, Camilo, Lapèze, Jérémie, Urban, Julie M. (2025): Taxonomic and ecological notes on Lycoderides Sakakibara (Hemiptera: Membracidae), including two new species from the highlands of the Colombian Andes. Zootaxa 5665 (2): 151-186, DOI: 10.11646/zootaxa.5665.2.1, URL: https://doi.org/10.11646/zootaxa.5665.2.1
03ED5C1AFFC9FA5C94CFDCD2FD525648.taxon	diagnosis	Diagnosis: The male of this species resembles those of L. abditus, L. santainensis sp. nov., and L. serraticornis but can be distinguished by its short pubescence on the body, head 2.5 × wider than long, and pronotum higher than wide, elevated 2 × head length above the head; suprahumeral horns are larger, the anterior process is more dorsally directed, and the apical crest of the posterior process is more elevated than in other Lycoderides species. Femora and tibiae are entirely light brown. Forewing hyaline area is small. Male genitalia resemble those of L. santainensis and L. serraticornis.	en	Flórez-V, Camilo, Lapèze, Jérémie, Urban, Julie M. (2025): Taxonomic and ecological notes on Lycoderides Sakakibara (Hemiptera: Membracidae), including two new species from the highlands of the Colombian Andes. Zootaxa 5665 (2): 151-186, DOI: 10.11646/zootaxa.5665.2.1, URL: https://doi.org/10.11646/zootaxa.5665.2.1
03ED5C1AFFC9FA5C94CFDCD2FD525648.taxon	description	Description: Holotype male. Color: General color reddish brown. Head dark brown, lighter along superior margin around ocelli, yellowish brown near eyes; eyes greyish brown, ocelli pink. Pronotum and warts reddish brown, supraocular callosities pinkish. Dorso-medial carina above metopidium and posterior process, suprahumeral horn edges yellow. Forewing sclerotized, punctuated, reddish dark brown until half-length; forewing membrane opaque brown on apical half, one hyaline and speckled spot on apex of R basal cell, R discoidal cell, and adjacent area of M basal cell; reddish hyaline area on subapical region of M and Cu basal cell. Hindwings hyaline. Coxae dark brown with lighter edges. Femora and tibiae light brown. Abdominal segments orange-brown. Sculpture: Densely and coarsely punctuated, appearing opaque, Head, pronotum, sclerotized area of forewing, forewing veins, dorsal surface of tibiae and tarsomeres covered with short light brown pubescence. Wart-like protuberances scattered over pronotum, forewing veins, and sclerotized, punctuated area of membrane. Head (Fig. 3 C): 2.5 x wider than long; eyes hemispheric; ocelli above transocular line, closer to posterior margin, positioned closer to eyes than to each other; supra-antennal ledges arched; frontoclypeus slightly emarginate above supra-antennal ledges. Beak extending to metacoxae. Thorax: Finger-shaped process on ventral base of mesepimeron. Pronotum in frontal view (Fig. 3 C), higher than wide, elevated above head almost 2 x head length; humeral angles triangular; anterior process with two triangular suprahumeral horns; in lateral view (Fig. 4 G), metopidium vertical, truncate at level of suprahumeral horns, summit behind suprahumeral horns, descending abruptly in sinuous line until above M and Cu bifurcation, forming small crest before reaching posterior margin of posterior process; posterior process reaching before claval suture of forewing; scutellum completely covered; in dorsal view (Fig. 5 G), suprahumeral horns slightly longer than wide, dorso-medial carinae irregular forming zig-zag-line. Tibia with flattened dorsal surface, metatibia with row II of cucullate setae. Abdomen (Fig. 12 F – J): Male genitalia. Lateral plate totally fused to pygofer. Style attached to subgenital plate on basal 1 / 3 of subgenital plate. Subgenital plate bilobed, divided almost from base, attached with VIII abdominal sternite. Styles mesally expanded, apex hook-shaped; tooth obliquely directed lateral- and ventrally. Aedeagus with anterior arm reduced, posterior arm in lateral view, basally expanded, then slightly narrow until apex; anterior and antero-lateral surface of apical 1 / 3 of posterior arm with denticles; in posterior view, subcylindrical. Female and late-instar nymph unknown. Measurements: Holotype male (mm): Body length: 7.58; pronotum length: 4.63; pronotum height: 1.80; forewing length: 6.43; width between humeral angles: 2.52; head width: 2.32; vertex length: 0.86; vertex width: 1.52.	en	Flórez-V, Camilo, Lapèze, Jérémie, Urban, Julie M. (2025): Taxonomic and ecological notes on Lycoderides Sakakibara (Hemiptera: Membracidae), including two new species from the highlands of the Colombian Andes. Zootaxa 5665 (2): 151-186, DOI: 10.11646/zootaxa.5665.2.1, URL: https://doi.org/10.11646/zootaxa.5665.2.1
03ED5C1AFFC9FA5C94CFDCD2FD525648.taxon	biology_ecology	Biology: One male was found in an Alto-Andean forest on a Quercus humboldtii (Fagaceae) leaf (Fig. 16 A). However, additional records are needed to confirm if Q. humboldtii is its host plant.	en	Flórez-V, Camilo, Lapèze, Jérémie, Urban, Julie M. (2025): Taxonomic and ecological notes on Lycoderides Sakakibara (Hemiptera: Membracidae), including two new species from the highlands of the Colombian Andes. Zootaxa 5665 (2): 151-186, DOI: 10.11646/zootaxa.5665.2.1, URL: https://doi.org/10.11646/zootaxa.5665.2.1
03ED5C1AFFC9FA5C94CFDCD2FD525648.taxon	materials_examined	Examined material: Holotype male in CBUCES. COLOMBIA: Antioquia: El Retiro: “ COLOMBIA. Antioquia, El Retiro, Reserva Natural San Sebastián de la Castellana, 6.109670 ° N, 75.546140 ° W, 2500 – 2800 msnm, sobre hoja de Quercus humboldtii, Mar. 4 / 2018, leg. Semillero Entomología 2018 - I (C. Flórez-V), CBUCES-F 8029 ” (1 male in CBUCES). Holotype minuten mounted, in excellent state of preservation with dissected abdomen placed in vials with glycerin pinned with specimens.	en	Flórez-V, Camilo, Lapèze, Jérémie, Urban, Julie M. (2025): Taxonomic and ecological notes on Lycoderides Sakakibara (Hemiptera: Membracidae), including two new species from the highlands of the Colombian Andes. Zootaxa 5665 (2): 151-186, DOI: 10.11646/zootaxa.5665.2.1, URL: https://doi.org/10.11646/zootaxa.5665.2.1
03ED5C1AFFC9FA5C94CFDCD2FD525648.taxon	distribution	Distribution: COLOMBIA: Antioquia: El Retiro (Reserva Natural San Sebastián de la Castellana, 6.109670 ° N, 75.546140 ° W, 2800 masl, Fig. 1).	en	Flórez-V, Camilo, Lapèze, Jérémie, Urban, Julie M. (2025): Taxonomic and ecological notes on Lycoderides Sakakibara (Hemiptera: Membracidae), including two new species from the highlands of the Colombian Andes. Zootaxa 5665 (2): 151-186, DOI: 10.11646/zootaxa.5665.2.1, URL: https://doi.org/10.11646/zootaxa.5665.2.1
03ED5C1AFFC9FA5C94CFDCD2FD525648.taxon	etymology	Etymology: The specific epithet ‘ aburraensis’ refers to the Valle de Aburrá, the type locality of this species. The name derives from the ‘ Aburráes’, the Indigenous people who originally inhabited the region.	en	Flórez-V, Camilo, Lapèze, Jérémie, Urban, Julie M. (2025): Taxonomic and ecological notes on Lycoderides Sakakibara (Hemiptera: Membracidae), including two new species from the highlands of the Colombian Andes. Zootaxa 5665 (2): 151-186, DOI: 10.11646/zootaxa.5665.2.1, URL: https://doi.org/10.11646/zootaxa.5665.2.1
03ED5C1AFFC9FA5C94CFDCD2FD525648.taxon	discussion	Remarks: This species was found in a locality close to those of L. serraticornis and L. santainensis sp. nov., on the slopes of the Valle de Aburrá. L. aburraensis sp. nov. can be distinguished from L. santainensis sp. nov. and L. serraticornis by: head 2.5 × wider than long; pronotum higher than wide, elevated 2 × head length above head, suprahumeral horns larger, anterior process more dorsally directed and apical crest of posterior process more elevated than the other Lycoderides species. Male genitalia of these species are very similar between, and they are difficult to distinguish based only on these structures (Fig. 12). Females of this species are unknown, but the description of the female of L. abditus, L. santainensis sp. nov. and L. serraticornis give us insights about the morphology of females of L. aburraensis sp. nov.	en	Flórez-V, Camilo, Lapèze, Jérémie, Urban, Julie M. (2025): Taxonomic and ecological notes on Lycoderides Sakakibara (Hemiptera: Membracidae), including two new species from the highlands of the Colombian Andes. Zootaxa 5665 (2): 151-186, DOI: 10.11646/zootaxa.5665.2.1, URL: https://doi.org/10.11646/zootaxa.5665.2.1
03ED5C1AFFCFFA4394CFDC75FAB054E8.taxon	description	(Fig. 3 A – B, 4 E – F, 5 E – F, 6 C – D, 10 D, 10 H, 11 D, 11 H, 12 K – O, 14 J – M, 15)	en	Flórez-V, Camilo, Lapèze, Jérémie, Urban, Julie M. (2025): Taxonomic and ecological notes on Lycoderides Sakakibara (Hemiptera: Membracidae), including two new species from the highlands of the Colombian Andes. Zootaxa 5665 (2): 151-186, DOI: 10.11646/zootaxa.5665.2.1, URL: https://doi.org/10.11646/zootaxa.5665.2.1
03ED5C1AFFCFFA4394CFDC75FAB054E8.taxon	diagnosis	Diagnosis: Sexually dimorphic. Warts distinct in color from surrounding body area. Forewing veins with wart-like protuberances. Male. General color reddish dark brown to dark brown. Metopidium vertical; pronotum elevated above head 1 × head length, descending almost from above humeral angles; suprahumeral horns reduced to white wart-like protuberances. Forewing veins with warts; hyaline area from mid-length to apex, densely speckled around veins at third apical; veins lighter on non-speckled areas; sclerotized and punctuated until behind half-length. Female. General color light grizzled brown to brown. Metopidium vertical, with one pronotal anterior process obliquely directed, bearing two triangular, carinate suprahumeral horns. Forewing opaque hyaline from apex of basal cell M to apex; brownish on fourth and fifth apical cells and apical area of second and third apical cells; sparsely speckled with brown; densely speckled with dark brown spots around R 2 + 3, M 1 + 2, M 3 + 4, and m-cu.	en	Flórez-V, Camilo, Lapèze, Jérémie, Urban, Julie M. (2025): Taxonomic and ecological notes on Lycoderides Sakakibara (Hemiptera: Membracidae), including two new species from the highlands of the Colombian Andes. Zootaxa 5665 (2): 151-186, DOI: 10.11646/zootaxa.5665.2.1, URL: https://doi.org/10.11646/zootaxa.5665.2.1
03ED5C1AFFCFFA4394CFDC75FAB054E8.taxon	description	Description. Holotype male (Fig. 3 A, 4 E, 5 E, 6 C). Color: General color reddish dark brown to dark brown. Head light brown, with one dark brown spot on coronal suture above frontoclypeus and one pair of dark brown spots oblique to ocelli; supra-antennal ledges lighter. Metopidium, supraocular callosities, and postocular lobes light brown; dorsal and posterior surface dark brown with warts; suprahumeral horns and lowest area of dorsal margin white. Forewing sclerotized, punctuated, reddish dark brown until behind mid-length; membrane from mid-length to apex hyaline, densely speckled around veins at third apical; veins lighter in non-speckled areas. Hindwings hyaline. Femora and tibiae light brown, tibiae with two darker bands; coxae dark brown. Abdominal segments III – VI orange, segments VII – IX dark brown. Sculpture: Head, pronotum, sclerotized area of forewing, forewing veins, dorsal surface of tibiae and tarsomeres with light brown pubescence; surface densely and coarsely punctuated (opaque appearance); scattered wart-like protuberances around pronotal median carina and forewing veins. Head: Hexagonal, twice as wide as long; eyes hemispheric; ocelli above transocular line, closer to posterior margin and to eyes than to each other; supra-antennal ledges arched; frontoclypeus slightly emarginate above supra-antennal ledges; in lateral view, head slightly projected posteriorly; beak extending to metacoxae. Thorax: Ventral base of mesepimeron with finger-shaped process. Pronotum in frontal view (Fig. 3 A), more or less triangular, elevated above head almost 1.5 x head length; humeral angles triangular; anterior process with two suprahumeral horns reduced to white wart-like protuberances; in lateral view (Fig. 4 E), metopidium vertical, truncate at level of suprahumeral horns; summit behind suprahumeral horns, descending more or less straight until above forewing vein A 3, then forming small crest before reaching posterior margin of posterior process, terminating before claval suture of forewing; scutellum completely covered; in dorsal view (Fig. 5 E), dorso-medial carinae irregular, zig-zag-line. Tibia with flattened dorsal surface, metatibia with row II of cucullate setae. Abdomen (Fig. 12 K – O): Male genitalia. Lateral plate totally fused to pygofer. Stlyes attached to subgenital plate on basal 1 / 3 of subgenital plate. Subgenital plate bilobed, divided almost from base, attached with VIII abdominal sternite. Styles in lateral view, mesally expanded, apex hook-shaped; tooth obliquely directed lateral- and ventrally. Aedeagus with anterior arm reduced, posterior arm in lateral view, basally expanded, then slightly narrow until apex; anterior and antero-lateral surface of apical 1 / 3 of posterior arm with denticles; in posterior view, subcylindrical. Female (Fig. 3 B, 4 F, 5 F, 6 D). Color: Variegated light grizzled brown to brown. Post-clypeus dark brown. Pronotal posterior process tip light brown; warts dark brown. Forewing sclerotized, punctuated, and light grizzled brown until behind half-length; membrane from apex of basal cell M to apex opaque hyaline, sparsely speckled with brown; fourth and fifth apical cells and apical area of second and third apical cells brownish; veins R 2 + 3, M 1 + 2, M 3 + 4, m-cu densely speckled with dark brown spots. Legs light grizzled brown; tibiae with two darker bands; femora and tarsomeres brown; coxae dark brown. Sculpture: Surface densely and coarsely punctuated (opaque appearance). Light brown pubescence, sparsely distributed on head, pronotum, sclerotized area of forewing, forewing veins, dorsal surface of tibiae, tarsomeres. Waxy pubescence present on gena, post-ocular lobes, thoracic pleurites and sternites, coxae (leaving variable smooth spots), trochanter, ventral and dorsal surface of femora (leaving triangular smooth bands on anterior and posterior surface), and abdominal sternites. Wart-like protuberances scattered around dorso-median carina and forewings. Thorax: Mesepimeron lacking finger shape process on ventral base (different to males). Pronotum in frontal view (Fig. 3 B), more or less triangular; with pronotal anterior process, elevated 1 × head length above metopidium, bearing two triangular, carinate suprahumeral horns, each carina surrounded by warts, dorso-median carina from anterior margin of metopidium; in lateral view (Fig. 4 F), pronotal horn directed obliquely forward and upward, 0.5 × length of remainder pronotum length; dorsal margin descending almost straight until tip of pronotal horn to posterior third, then forming a small crest before reach posterior margin of posterior process; scutellum completely covered; in dorsal view (Fig. 5 F), suprahumeral horns triangular. Left forewing with two additional r-m cross-veins, and on right forewing with one complete and one partial additional m-cu crossvein. Abdomen (Fig. 14 J – M): Female genitalia. Terga with sclerotized areas and pits ventrally. Sternites, pleurites, ventral margin of pygofer and gonoplac with thick and rolled setae. Genitalia: Second valvulae blade-shaped, basal half narrower than apical half; apical half separated, dorsal margin of apical half with sub-quadrangular teeth (TE); ramus extended to apical portion; pores over ramus and below dorsal margin of apical half and ventral margin of 1 / 3 apical area. Variations: There is no difference in color between live and pinned specimens. Late instar nymph (Fig. 10 D, 10 H, 11 D, 11 H): Dorsoventrally compressed. Color: brownish orange. Sculpture: Dorsal tegument sparsely covered with chalazal setae and plant indument, moss, and lichen. Head: small subtriangular processes above ventro-anterior margin; in frontal view; in ventral and dorsal view, ventro-anterior margin forming a continuous and widely arched margin with ventrolateral lobes. In lateral view, projected obliquely forward and downward; in dorsal view, more or less five times wider than long, rectangular. Thorax: Pronotum with metopidium obliquely directed dorso-posteriorly; dorso-median margin forming a sinuous plateau with two rows of short chalazae; two humps (suprahumeral horns) at each side of anterior plateau. Abdomen: In dorsal view, tergal segments IV – VIII with large subtriangular lateral lamellae, directed postero-laterally, smaller on tergum IV; terga III – VIII with three pairs of longitudinal rows of short tuberculate chalazae: one pair near the midline, one above the lateral lamellae, and one between the previously mentioned pairs. Live specimens: Densely covered by plant indument, moss, and lichen. Color varies from green to brownish orange, seemingly influenced by location on host plant. Measurements: Holotype male / paratype female (mm): Body length: 7.47 / 8.71; pronotum length: 4.57 / 5.15; pronotum height: 1.73 / 2.55; forewing length: 6.28 / 7.42; width between humeral angles: 2.50 / 2.94; head width: 2.29 / 2.61; vertex length: 0.85 / 0.86; vertex width: 1.49 / 1.76.	en	Flórez-V, Camilo, Lapèze, Jérémie, Urban, Julie M. (2025): Taxonomic and ecological notes on Lycoderides Sakakibara (Hemiptera: Membracidae), including two new species from the highlands of the Colombian Andes. Zootaxa 5665 (2): 151-186, DOI: 10.11646/zootaxa.5665.2.1, URL: https://doi.org/10.11646/zootaxa.5665.2.1
03ED5C1AFFCFFA4394CFDC75FAB054E8.taxon	biology_ecology	Biology: The nymphs of this species are highly camouflaged against their host plant, Quercus humboldtii (Fagaceae) (Fig. 15 A, C). They are covered with the plant’s indument and moss and are commonly found in leaf axils, often beneath the stipules (Fig. 15 C). Nymphs do not form aggregations, but they are often found in close proximity, with up to three individuals near each other. They were always associated with Crematogaster sp. (Myrmicinae) (Fig. 15 A). This ant species is usually present in small numbers (1 – 3 ants per nymph). Frequently, when the nymphs or the nearby branch are disturbed, the ants raise their abdomens and move them as if vibrating, eventually secreting a white substance. However, some ants flee when the nymphs are disturbed. On one occasion, three ants were observed feeding on a late-instar nymph (Fig. 15 B). Observations were made during four field trips to the ‘ Páramo de Santa Inés’ (3250 masl), the first in February of 2015, the second in August of 2015, the third in February 2016 and the last in December 2016. The first time, nymphs were found on a seedling of Q. humboldtii, tended by Crematogaster sp. In this trip, nymphs or adults were not found on other nearby individuals of Q. humboldtii. During the second field trip, nymphs were found with ants and a male in the same individual of Quercus (Fig. 15 D). Also, on the upper part of this plant, an aggregation of nymphs and a female of Metcalfiella sp. (Membracinae) were observed (Fig. 15 G). Ants were not observed tending these treehoppers, but ants were feeding on honeydew drops that fell on a nearby leaf. On another individual Quercus plant, one male was found. During the third field trip, nymphs were once again observed on the same Quercus individual, as well as on four additional plants. On one of these plants, a female was spotted, well-camouflaged and mimicking the color of the oak stem (Fig. 15 E – F). Thus, nymphs, males, and females exhibit mimicry with the plant’s indument, though in different ways. Additionally, on the same plants, two different species of Metcalfiella and a female of Aetalion nigromarginatum (Aetalionidae) were observed sitting on eggs (Fig. 15 H). During the last field trip, only nymphs were found on the same Quercus individuals.	en	Flórez-V, Camilo, Lapèze, Jérémie, Urban, Julie M. (2025): Taxonomic and ecological notes on Lycoderides Sakakibara (Hemiptera: Membracidae), including two new species from the highlands of the Colombian Andes. Zootaxa 5665 (2): 151-186, DOI: 10.11646/zootaxa.5665.2.1, URL: https://doi.org/10.11646/zootaxa.5665.2.1
03ED5C1AFFCFFA4394CFDC75FAB054E8.taxon	materials_examined	Examined material: Holotype male in CBUCES. COLOMBIA: Antioquia: Belmira: “ COLOMBIA. Antioquia, Belmira, Páramo de Santa Inés, por el lado de la Laguna en Sabanas, 6.615899 ° N, 75.648855 ° W, 2800 - 3250 msnm, manual, en Quercus humboldtii, Ago. 16 / 2015, leg. J. Cardona & C. Flórez-V, CBUCES-F 8030 ” (CBUCES).	en	Flórez-V, Camilo, Lapèze, Jérémie, Urban, Julie M. (2025): Taxonomic and ecological notes on Lycoderides Sakakibara (Hemiptera: Membracidae), including two new species from the highlands of the Colombian Andes. Zootaxa 5665 (2): 151-186, DOI: 10.11646/zootaxa.5665.2.1, URL: https://doi.org/10.11646/zootaxa.5665.2.1
03ED5C1AFFCFFA4394CFDC75FAB054E8.taxon	materials_examined	Paratypes in CBUCES: “ COLOMBIA. Antioquia, Belmira, Páramo de Santa Inés por el lado de la Laguna en Sabanas, 6.615899 ° N, 75.648855 ° W, 2800 – 3250 msnm, manual, en Quercus humboldtii, Feb. 14 / 2016, leg. C. Flórez-V, CBUCES-F 8031 ” (1 female in CBUCES); “ COLOMBIA. Antioquia, Belmira, Páramo de Santa Inés, Sector Sabanas, cerca de la Laguna, 6.615899 ° N, 75.648855 ° W, 2800 – 3250 msnm, manual, en Quercus humboldtii, Feb. 4 – 6 / 2015, leg. N. Arcila, C. Flórez-V, I. Olivares \ CBUCES-F 8024 ” (1 nymph in CBUCES); “ COLOMBIA. Antioquia, Belmira, Páramo de Santa Inés, Sector Sabanas, cerca de la Laguna, 6.615899 ° N, 75.648855 ° W, 3200 – 3250 msnm, manual, en Quercus humboldtii, Ago. 16 / 2015, leg. J. Cardona, V. Correa, C. Flórez-V, H. Londoño, CBUCES-F 447 ” (1 male in CBUCES); “ COLOMBIA. Antioquia, Belmira, cerca del Páramo de Santa Inés, robledal subiendo a Sector Sabanas, cerca de la Laguna, 6.615899 ° N, 75.648855 ° W, 2900 msnm, manual, en Quercus humboldtii, Sep. 2 – 7 / 2017, leg. V. Correa, C. Flórez-V, A. Ospina, D. Taborda, CBUCES-F 1708 ” (1 female in CBUCES). Holotype minuten mounted, in excellent state of preservation. Paratypes adults minuten mounted. One male and one female paratype with dissected abdomens placed in vials with glycerin pinned with specimens. Two nymphs and 1 male in 75 % ethanol.	en	Flórez-V, Camilo, Lapèze, Jérémie, Urban, Julie M. (2025): Taxonomic and ecological notes on Lycoderides Sakakibara (Hemiptera: Membracidae), including two new species from the highlands of the Colombian Andes. Zootaxa 5665 (2): 151-186, DOI: 10.11646/zootaxa.5665.2.1, URL: https://doi.org/10.11646/zootaxa.5665.2.1
03ED5C1AFFCFFA4394CFDC75FAB054E8.taxon	distribution	Distribution: COLOMBIA: Antioquia: Belmira (Páramo de Santa Inés: 6.621161 ° N, 75.645648 ° W; 2800 – 3250 masl) (Fig. 1).	en	Flórez-V, Camilo, Lapèze, Jérémie, Urban, Julie M. (2025): Taxonomic and ecological notes on Lycoderides Sakakibara (Hemiptera: Membracidae), including two new species from the highlands of the Colombian Andes. Zootaxa 5665 (2): 151-186, DOI: 10.11646/zootaxa.5665.2.1, URL: https://doi.org/10.11646/zootaxa.5665.2.1
03ED5C1AFFCFFA4394CFDC75FAB054E8.taxon	etymology	Etymology: The name refers to the type locality in ‘ Páramo de Santa Inés’, located in the northern part of the Cordillera Central in Antioquia, Colombia. This remarkable and ecologically significant páramo is one of the main water sources for Medellín and 11 municipalities in central Antioquia.	en	Flórez-V, Camilo, Lapèze, Jérémie, Urban, Julie M. (2025): Taxonomic and ecological notes on Lycoderides Sakakibara (Hemiptera: Membracidae), including two new species from the highlands of the Colombian Andes. Zootaxa 5665 (2): 151-186, DOI: 10.11646/zootaxa.5665.2.1, URL: https://doi.org/10.11646/zootaxa.5665.2.1
03ED5C1AFFCFFA4394CFDC75FAB054E8.taxon	discussion	Remarks: Males of L. santainensis sp. nov. and L. abditus are similar. However, males of L. santainensis sp. nov. can be distinguished by their larger size, darker and more reddish coloration, vertical metopidium (as opposed to the forward-reclined metopidium in L. abditus), suprahumeral horns reduced to small warts, forewing veins with warts, and relatively longer forewings in proportion to pronotal length. Additionally, the tibiae are more foliaceous. Males of L. santainensis also resemble those of L. serraticornis and L. aburraensis, but they can be distinguished by their larger size, pronotal shape, smaller suprahumeral horns, forewing sclerotization, coloration, and wart distribution, as well as the male genitalia, where the aedeagus, which has a more prominent hump at the base of the posterior arm (Fig. 12 N), as well as by its styles, which are broader along their extension in both lateral and dorsal views (Fig. 12 L – M). Males of these four species share a distinct pronotal shape compared to other Lycoderides, lacking a well-developed anterior process and having reduced suprahumeral horns. The female of L. santainensis differs from other Lycoderides species and can be distinguished by its pronotum, which has an anterior horn elevated to a height equal to the head length above the metopidium, the presence of warts throughout the pronotal horn and the dorso-medial carinae, and an elevated posterior region of the posterior pronotal process. Additionally, the thick and rolled setae on the sternites, pleurites, pygofer, and gonoplac have not been observed in any of the other species examined in this study (Fig. 14 M). The female of this species is morphologically similar to those of L. abditus and L. serraticornis, sharing features such as a pronotum with an anterior horn, a scutellum concealed by the pronotum, and partially hyaline and speckled forewings. This represents the first record of a Stegaspidinae species occurring above 3000 masl, as well as the first record of the genus Lycoderides on Fagaceae, specifically on the genus Quercus. Both adults and nymphs camouflage themselves in different ways with their host plant. The dorsoventrally flattened shape of the nymphs resembles that of other groups of Membracidae in which nymphs are camouflaged when lying on the twigs of their host plant, such as certain species of Nicomiinae (e. g., Nicomia, Flórez-V pers. obs.) and other Stegaspidinae (e. g., Microcentrini, Bocydium, Stylocentrus, Problematode; Flórez-V & Evangelista 2017, Flórez-V 2019). On the other hand, males and females mimic the plant bark in different ways: females resemble the stem bark in more sclerotized areas, while males resemble the axillary buds. The accumulation of moss or plant indument in nymphs may be facilitated by the presence of chalazae and lateral abdominal lamellae, as well as by the flattened body shape and possibly by specific behaviors that allow individuals to actively collect these materials, since this external covering would be lost with each molt.	en	Flórez-V, Camilo, Lapèze, Jérémie, Urban, Julie M. (2025): Taxonomic and ecological notes on Lycoderides Sakakibara (Hemiptera: Membracidae), including two new species from the highlands of the Colombian Andes. Zootaxa 5665 (2): 151-186, DOI: 10.11646/zootaxa.5665.2.1, URL: https://doi.org/10.11646/zootaxa.5665.2.1
03ED5C1AFFD0FA4494CFDEDAFBEB54CC.taxon	description	(Figs. 2 C – D, 4 C – D, 5 C – D, 6 A – B, 12 A – E, 14 A – C, 16 B)	en	Flórez-V, Camilo, Lapèze, Jérémie, Urban, Julie M. (2025): Taxonomic and ecological notes on Lycoderides Sakakibara (Hemiptera: Membracidae), including two new species from the highlands of the Colombian Andes. Zootaxa 5665 (2): 151-186, DOI: 10.11646/zootaxa.5665.2.1, URL: https://doi.org/10.11646/zootaxa.5665.2.1
03ED5C1AFFD0FA4494CFDEDAFBEB54CC.taxon	discussion	Remarks: In recent expeditions, we collected several females and one male of L. serraticornis in Antioquia. Based on these specimens, we determined that Flórez-V et al. (2015) mistakenly recorded Lycoderides abditus from Colombia. The specimen examined in Flórez-V et al. (2015) actually belongs to a male of L. serraticornis. The male genitalia of L. serraticornis, L. aburraensis sp. nov., and L. santainensis sp. nov. share similarities but exhibit key differences. In L. serraticornis, the aedeagus has a smaller hump at the base of the posterior arm (Fig. 12 D), and the styles are slender in lateral and dorsal views (Fig. 12 B – C), distinguishing it from L. santainensis sp. nov. Externally, males of L. serraticornis differ from those of L. abditus, L. aburraensis sp. nov., and L. santainensis sp. nov. in several ways, including a more elevated pronotum that descends from above the humeral angles, larger and more triangular suprahumeral horns, the presence of a pair of anteroventral carinae below the suprahumeral horns, a pair of wart-like protuberances on the dorsal margin of the head, and differences in forewing coloration and sclerotization. The male of L. serraticornis is more similar to L. abditus than to L. santainensis sp. nov. The warts match the surrounding body coloration (Fig. 4 C), as in L. abditus, but different from L. santainensis sp. nov. (Fig. 4 E); additionally, L. serraticornis lacks warts on the forewing veins (Fig. 4 C), unlike L. santainensis. Females of L. serraticornis can be distinguished from those of L. abditus and L. santainensis sp. nov. by the following features: a longer and more elevated pronotal horn with an almost straight posterior margin, well-developed suprahumeral horns, a lower subapical elevation on the posterior process, and a brownish-hyaline forewing membrane densely speckled in the apical third.	en	Flórez-V, Camilo, Lapèze, Jérémie, Urban, Julie M. (2025): Taxonomic and ecological notes on Lycoderides Sakakibara (Hemiptera: Membracidae), including two new species from the highlands of the Colombian Andes. Zootaxa 5665 (2): 151-186, DOI: 10.11646/zootaxa.5665.2.1, URL: https://doi.org/10.11646/zootaxa.5665.2.1
03ED5C1AFFD0FA4494CFDEDAFBEB54CC.taxon	diagnosis	Diagnosis: Male. General color brown to dark brown. Forewings hyaline, sparsely speckled from basal third to apical quarter, densely speckled apically; sclerotized area scattered with small wart-like protuberances; veins lacking warts; warts same color as surrounding body area. Head with one pair of wart-like protuberances on dorsal margin. Metopidium slightly inclined forward; pronotum elevated above head 2 × head length, descending almost above humeral angles; suprahumeral horns triangular; one pair of anteroventral carinae below suprahumeral horns. Female. Pronotum elevated above head 2.5 × head length in frontal view; pronotal anterior process almost as long as pronotal height; in lateral view, subapical crest of posterior process slightly elevated. Forewing membrane brownish-hyaline, densely speckled at apical third; veins lacking warts.	en	Flórez-V, Camilo, Lapèze, Jérémie, Urban, Julie M. (2025): Taxonomic and ecological notes on Lycoderides Sakakibara (Hemiptera: Membracidae), including two new species from the highlands of the Colombian Andes. Zootaxa 5665 (2): 151-186, DOI: 10.11646/zootaxa.5665.2.1, URL: https://doi.org/10.11646/zootaxa.5665.2.1
03ED5C1AFFD0FA4494CFDEDAFBEB54CC.taxon	description	Description: Male (Fig. 2 C – D, 4 C – D, 5 C – D, 6 A). Color: General color brown. Head dark brown, lighter around dorso-lateral margin of vertex; eyes variegated red, brown, and dark brown. Metopidium around median carina, suprahumeral horns, summit, and crest at posterior pronotal process lighter brown. Forewing sclerotized, punctuated, and brown on basal 1 / 3; forewing membrane hyaline from basal 2 / 3 to apex, speckled, with apical 1 / 4 densely speckled; veins lighter on non-speckled areas. Hindwings hyaline. Legs brown. Warts same color as surrounding body area. Sculpture: Pronotum and sclerotized forewing areas with scattered wart-like protuberances, denser on metopidium and around dorso-median carina. Head, pronotum, sclerotized forewing areas, forewing veins, dorsal surface of tibiae, and tarsomeres with light brown pubescence. Body densely and coarsely punctuated (appearing opaque). Head (Fig. 2 C): Hexagonal, twice as wide as long; eyes hemispheric; ocelli located above transocular line, closer to posterior margin and to eyes than to each other; one pair of wart-like protuberances above ocelli, on dorsal margin of vertex; supra-antennal ledges arched; frontoclypeus slightly emarginate above supra-antennal ledges; area of vertex on transocular line and frontoclypeus convex. In lateral view (Fig. 4 C), head slightly projected posteriorly, beak reaching metacoxae. Thorax: Mesepimeron with one finger shape process on ventral base. Pronotum in frontal view (Fig. 2 C), more or less triangular, elevated above head 2 x head length; suprahumeral horns triangular, with carinae along edges; humeral angles triangular, weakly produced; in lateral view (Fig. 4 C), metopidium slightly inclined anteriorly, forming an obtuse angle with dorsal margin; one pair of short ventrolateral carinae below suprahumeral horns, each 1 x eye length; dorsal margin slightly arched after suprahumeral horns, strongly descending above humeral angles until A 2 forewing vein, then forming a small crest before reaching posterior margin of posterior process, extending near claval suture of forewing; in dorsal view (Fig. 5 C), dorso-median carinae irregular, forming a zig-zag-line. Scutellum completely covered by pronotum. Tibia slightly spatulate on dorsal surface, metatibia with row II of cucullate setae. Abdomen (Fig. 12 A – E): Male genitalia. Lateral plate totally fused to pygofer. Styles attached to subgenital plate on basal 1 / 3 of subgenital plate. Subgenital plate bilobed, divided almost from base, attached with VIII abdominal sternite. Styles in lateral view, with broad throughout until apical 1 / 3, with one tooth directed laterally; in ventral view, mesally expanded, apex hook shaped. Aedeagus with anterior arm reduced, posterior arm in lateral view, broad basally, then slightly narrow until apex; anterior and antero-lateral surface of apical 1 / 3 of posterior arm with denticles; in posterior view, subrectangular, slightly narrow at 1 / 2. Female. Genitalia (Fig. 14 A – C): Second valvulae blade-shaped, basal third narrower than apical 2 / 3; apical 2 / 3 separated, wider toward subapex, apex acute; dorsal margin of apical 2 / 3 with sub-quadrangular teeth (TE); ramus extended to apical portion; pores over ramus and below dorsal margin of apical half and ventral margin of 1 / 3 apical area. Late instar nymph (based on exuviae): Dorsoventrally flattened. Color: brown. Surface: Dorsal tegument densely covered with short chalazal setae and by indument of the plant, moss and lichen. Head: Ventrolateral lobes extended until external eye margin; small subtriangular processes above ventro-anterior margin; in ventral and dorsal view, ventro-anterior margin forming a continuous and widely arched margin with ventrolateral lobes. In lateral view, projected obliquely forward and downward; in dorsal view, about four times wider than long, rectangular. Thorax: Pronotum with metopidium vertical, with a short conical dorso-anteriorly projected process; an oblique carina extending from apex to humeral angles; dorso-median and oblique carinae with chalazae. Abdomen: In dorsal view, tergal segments IV – VIII with sub-triangular lateral lamellae, directed posterolaterally, smaller on tergum IV; terga III – VIII with three pairs of longitudinal rows of short tuberculate chalazae: one pair near the midline, one above the lateral lamellae, and one between the previously mentioned pairs. Measurements: Male (mm): Body length: 6.50; pronotum length: 4.45; pronotum height: 2.06; forewing length: 5.28; width between humeral angles: 1.96; head width: 1.81; vertex length: 0.79; vertex width: 1.11.	en	Flórez-V, Camilo, Lapèze, Jérémie, Urban, Julie M. (2025): Taxonomic and ecological notes on Lycoderides Sakakibara (Hemiptera: Membracidae), including two new species from the highlands of the Colombian Andes. Zootaxa 5665 (2): 151-186, DOI: 10.11646/zootaxa.5665.2.1, URL: https://doi.org/10.11646/zootaxa.5665.2.1
03ED5C1AFFD0FA4494CFDEDAFBEB54CC.taxon	biology_ecology	Biology: In Colombia, adults and exuviae were recorded on Psidium guajava (Myrtaceae) at 2000 masl (Fig. 16 B). The integument of both males and females closely matched the color and texture of the bark of their host plants. On these plants, we also observed aggregations or females guarding their eggs or nymphs of a new species of Alchisme. Additionally, Richter (1942) recorded this species on Bellucia sp. (Melastomataceae) in Chocó. Kenji Nishida (pers. comm.) recorded in Costa Rica adults of a Lycoderides species similar to L. serraticornis on Conostegia xalapensis (Melastomataceae), as well as both adults and nymphs on Terminalia amazonia (Combretaceae). On the latter plant species, he observed nymphs inside shelters built on stems by Azteca sp. ants. However, we could not unequivocally identify this Lycoderides species as L. serraticornis. Given the scarcity of biological records for Lycoderides, we include this observation here for reference.	en	Flórez-V, Camilo, Lapèze, Jérémie, Urban, Julie M. (2025): Taxonomic and ecological notes on Lycoderides Sakakibara (Hemiptera: Membracidae), including two new species from the highlands of the Colombian Andes. Zootaxa 5665 (2): 151-186, DOI: 10.11646/zootaxa.5665.2.1, URL: https://doi.org/10.11646/zootaxa.5665.2.1
03ED5C1AFFD0FA4494CFDEDAFBEB54CC.taxon	materials_examined	Examined material and distribution: COLOMBIA: Antioquia: Caldas (Vereda La Clara, Refugio Natural Alto de San Miguel — 1900 – 2100 masl); Cauca: Timbío (Vereda La Rivera — 1600 masl).	en	Flórez-V, Camilo, Lapèze, Jérémie, Urban, Julie M. (2025): Taxonomic and ecological notes on Lycoderides Sakakibara (Hemiptera: Membracidae), including two new species from the highlands of the Colombian Andes. Zootaxa 5665 (2): 151-186, DOI: 10.11646/zootaxa.5665.2.1, URL: https://doi.org/10.11646/zootaxa.5665.2.1
03ED5C1AFFD7FA4694CFDD06FE165048.taxon	description	(Fig. 8 A, 10 A – B, 10 E – F, 11 A – B, 11 E – F, 13 F – J, 14 D – F, 16 C) Description: Male. Genitalia (Fig. 13 F – J): Lateral plate totally fused to pygofer. Styles attached to subgenital plate on basal 1 / 3 of subgenital plate. Subgenital plate bilobed, divided on apical 1 / 3, attached with VIII abdominal sternite. Styles with apex hook-shaped, widest area subapically and then abruptly narrow; tooth directed laterally. Aedeagus with anterior arm reduced, posterior arm in lateral view slender throughout, narrow toward apex, apex tongue-shaped; anterior and antero-lateral surface of apical 1 / 3 of posterior arm with denticles; in posterior view, subcylindrical. Female. Genitalia (Fig. 14 D – F): Second valvulae blade-shaped, basal half narrower than apical half; dorsal margin of apical half with sub-quadrangular teeth (TE); ramus extended to apical portion; pores over ramus and below dorsal margin of apical half and ventral margin of 1 / 3 apical area. Late instar nymph (Fig. 10 A – B, 10 E – F, 11 A – B, 11 E – F): Overall color light brown to green; tibiae, lamellae, and anal tube reddish. Sculpture: Dorsal tegument glabrous (Fig. 10 F) or densely covered (Fig. 10 E) with chalazal setae. Head: Ventrolateral lobes extended laterally to the external eye margin and anteriorly 1.5 × eye length; dorsal processes obsolete; in frontal view, extending ventrally and anteriorly, vertex forming two wide arches from the middle to the lateral margin of the ventrolateral lobes; in lateral view, projected obliquely forward and downward; in dorsal view, approximately 3 × wider than long. Thorax: Pronotum with metopidium obliquely directed dorsally and posteriorly, then dorso-median margin forming a plateau; without humps (suprahumeral horns) at each side of the anterior area of the plateau. Abdomen: In dorsal view, tergal segments IV – VIII with large lateral finger-shaped lamellae, directed posterolaterally.	en	Flórez-V, Camilo, Lapèze, Jérémie, Urban, Julie M. (2025): Taxonomic and ecological notes on Lycoderides Sakakibara (Hemiptera: Membracidae), including two new species from the highlands of the Colombian Andes. Zootaxa 5665 (2): 151-186, DOI: 10.11646/zootaxa.5665.2.1, URL: https://doi.org/10.11646/zootaxa.5665.2.1
03ED5C1AFFD7FA4694CFDD06FE165048.taxon	biology_ecology	Biology: This species has been found exclusively on Melastomataceae. Adults tend to be solitary, whereas nymphs often form small aggregations and are tended by ants. In Antioquia, at elevations between 1900 and 2400 masl, nymphs were observed on Andesanthus lepidota (Melastomataceae), primarily at the base of branches and in leaf axils near the apex, where they blended well with the plant’s texture (Fig. 16 C). Adults were also found, though always solitary, on leaves and in axils near the apex (Fig. 16 C). In one locality at the north of the Valle de Aburrá (2400 masl), nymphs were tended by Camponotus sp. and Myrmelachista sp. (Formicinae) on A. lepidota, while in the southern part of the Valle de Aburrá (1900 – 2100 masl), they were tended by Linepithema sp. (Dolichoderinae), also on A. lepidota. To the north of the Valle de Aburrá, at elevations between 600 and 1700 masl, adults were found solitary on seedlings and trees of Miconia spp., near the apex. In these plants, pairs or small groups of up to three nymphs or adults were occasionally observed being tended by Crematogaster sp. (Myrmicinae). In the Magdalena Medio region, small aggregations of up to five nymphs were observed on Bellucia axinanthera and Miconia sp. (Melastomataceae), associated with Ectatomma tuberculatum, E. ruidum and Camponotus coruscus. Nymphs and adults were also observed sharing the same plant with Aphetea sp. (Polyglyptini), cf. Hebeticoides (Darnini), and Stilbophora (Tragopini). In one locality (Yarumal), nymphs and adults were found along an elevational gradient between 1400 and 2000 masl. At elevations between 1400 and 1700 masl, they were observed on Miconia spp., while at 1900 – 2000 masl, they were found on A. lepidota.	en	Flórez-V, Camilo, Lapèze, Jérémie, Urban, Julie M. (2025): Taxonomic and ecological notes on Lycoderides Sakakibara (Hemiptera: Membracidae), including two new species from the highlands of the Colombian Andes. Zootaxa 5665 (2): 151-186, DOI: 10.11646/zootaxa.5665.2.1, URL: https://doi.org/10.11646/zootaxa.5665.2.1
03ED5C1AFFD7FA4694CFDD06FE165048.taxon	materials_examined	Examined material: COLOMBIA: Antioquia: Caldas (Reserva Natural Alto de San Miguel — 1900 – 2000 masl); Gómez Plata (Finca Vegas de la Clara — 800 masl); Remedios (Finca La Brillantina — 400 – 500 masl); San Carlos (vereda Santa Bárbara — 700 – 900 masl); San Luís (corregimiento El Prodigio — 700 – 900 masl); San Vicente (Finca La Mosca — 2400 masl); Yarumal (Reserva Los Magnolios — 1400 – 2000 masl). Caldas: Pensilvania (1900 – 2300 masl). Valle del Cauca: Buenaventura (PNN Farallones de Cali — 600 masl).	en	Flórez-V, Camilo, Lapèze, Jérémie, Urban, Julie M. (2025): Taxonomic and ecological notes on Lycoderides Sakakibara (Hemiptera: Membracidae), including two new species from the highlands of the Colombian Andes. Zootaxa 5665 (2): 151-186, DOI: 10.11646/zootaxa.5665.2.1, URL: https://doi.org/10.11646/zootaxa.5665.2.1
03ED5C1AFFD7FA4694CFDD06FE165048.taxon	discussion	Remarks: The integument and coloration of nymphs in this species vary according to the host plant. Nymphs found on Andesanthus lepidota are glabrous (lacking chalazae) (Fig. 11 F), resembling the smooth scales of the young stems (Fig. 16 C), whereas those on Miconia spp. are densely covered with short chalazae (Fig. 11 E), mimicking the plant’s indument. The coloration of live specimens ranges from green to brownish-orange, depending on the specific part of the plant where the nymph is located. While the general body shape remains unchanged, some nymphs, even on the same host plant, appear more dorsoventrally flattened. These nymphs resemble those of L. phasianus more than any other Lycoderides species examined in this study. In particular, the integument and coloration of nymphs on Miconia spp. are more similar to those of L. phasianus than those on A. lepidota. A useful diagnostic feature for L. luteus nymphs is the more pronounced anterior projection of the head (Figs 10 E, F) compared to other Lycoderides species, forming two broad arches. Adults from both Miconia spp. and A. lepidota are morphologically identical, including male genitalia, although individuals appear to be more gregarious in lowland habitats. This is one of the first records documenting morphological variation in membracid nymphs depending on the host plant. However, further evidence (e. g. molecular data) is needed to determine whether individuals from different host plants in L. luteus belong to the same species or represent a species complex. Interestingly, a similar pattern has been observed in the temperate North American Enchenopa binotata complex, where several biologically distinct species have nearly identical adults but nymphs that differ depending on their host plant (Pratt and Wood 1992). As previously noted for Lycoderes argutus and Lycoderides fernandezi by Lapèze and Lopez-Vaamonde (2024), L. luteus exhibits variation in forewing venation. The fourth apical cell can be either petiolate (with the r-m crossvein located basad to the M fork) or non-petiolate (with r-m located distad to the M fork). In the holotype of L. luteus, the fourth apical cell is non-petiolate, although the r-m crossvein is positioned very close to the M fork. Additionally, Sakakibara (2013) proposed that beyond the presence of a petiolate fourth apical cell, the apical cells in Lycoderides are arranged obliquely. However, this character appears to be highly variable and seems to form a continuum across species of Lycoderes and Lycoderides. Aside from the characters mentioned by Sakakibara (1972, 2013), we did not find additional features that reliably differentiate the two genera. Nevertheless, a detailed taxonomic revision and a phylogenetic analysis of the species in both genera are necessary to confidently determine whether they should be treated as synonyms.	en	Flórez-V, Camilo, Lapèze, Jérémie, Urban, Julie M. (2025): Taxonomic and ecological notes on Lycoderides Sakakibara (Hemiptera: Membracidae), including two new species from the highlands of the Colombian Andes. Zootaxa 5665 (2): 151-186, DOI: 10.11646/zootaxa.5665.2.1, URL: https://doi.org/10.11646/zootaxa.5665.2.1
03ED5C1AFFD5FA4994CFDA7AFA995010.taxon	description	(Fig. 8 B, 10 C, 10 G, 11 C, 11 G, 13 A – E, 14 G – I, 16 D – H) Description: Male. Genitalia (Fig. 13 A – E): Lateral plate totally fused to pygofer. Styles attached to subgenital plate on basal 1 / 3 of subgenital plate. Subgenital plate bilobed, divided on apical 1 / 3, attached with VIII abdominal sternite. Styles with widest area subapically and then abruptly narrow; apex bilobed, one tooth directed obliquely dorsal- and laterally, and second tooth ventral directed. Aedeagus with anterior arm reduced, posterior arm in lateral view slender throughout, narrow toward apex, apex finger-shaped; anterior and lateral surface of apical 1 / 3 of posterior arm with denticles; in posterior view, subcylindrical. Female. Genitalia (Fig. 14 G – I): Second valvulae blade-shaped, basal half narrower than apical half; apical half separated, dorsal margin of apical half with sub-quadrangular teeth; ramus extended to apical portion; pores over ramus and below dorsal margin of apical half and ventral margin of 1 / 3 apical area. Late instar nymph (Fig. 10 C, 10 G, 11 C, 11 G): Dorsoventrally flattened. Color: brown with reddish spots throughout body. Surface: Dorsal tegument densely covered with short chalazal setae. Head: Ventrolateral lobes extended anteriorly 1 × eye length; dorsal processes obsolete; in frontal view, ventro-anterior margin carinate, extended ventral and anteriorly, forming one wide arch. In lateral view, projected obliquely forward and downward; in dorsal view, more or less 4 × wider than long. Thorax: Pronotum with dorsal margin conical in lateral view, metopidium obliquely directed dorsal- and posteriorly, with a dorso-anterior process, with small humps (suprahumeral horns) at each side of anterior of dorso-anterior process. Abdomen: In dorsal view, tergal segments IV – VIII with large lateral conical lamellae directed posterolaterally, with anterior margin widely arched posterad. Live specimens: Color could vary from green to brownish orange, seemingly by the place of the plant where is located the nymph.	en	Flórez-V, Camilo, Lapèze, Jérémie, Urban, Julie M. (2025): Taxonomic and ecological notes on Lycoderides Sakakibara (Hemiptera: Membracidae), including two new species from the highlands of the Colombian Andes. Zootaxa 5665 (2): 151-186, DOI: 10.11646/zootaxa.5665.2.1, URL: https://doi.org/10.11646/zootaxa.5665.2.1
03ED5C1AFFD5FA4994CFDA7AFA995010.taxon	biology_ecology	Biology: Adults of this species are more gregarious, and nymphs form larger aggregations than those of other Lycoderides species examined in this study. The nymphs were tended by at least eight morphospecies of ants belonging to the genera Camponotus, Crematogaster, Dolichoderus, Ectatomma, Linepithema, and several unidentified myrmicines. This species was found on at least ten different morphospecies of Melastomataceae, all belonging or related to the genus Miconia, at various altitudes ranging from 200 to 1700 masl. It was also frequently observed forming mixed-species aggregations tended by ants in lowland areas between 200 and 700 masl (Fig. 16 E). Adults were found either solitary or in groups of up to nine individuals, usually on apical leaves or axils. Nymphs were often encountered in pairs or small groups on plants also hosting adults, but they could also form larger aggregations of up to 15 individuals (Fig. 16 D). As L. luteus, nymphs of this species were highly camouflaged with their host plants. In PNN Tatamá, above 1100 masl, nymphs were found in smaller numbers and formed smaller aggregations, always located in axils. In contrast, at lower elevations (below 1100 masl), nymphs formed larger aggregations and were found on apical branches, though not exclusively in axils. This species frequently co-occurred in mixed-species aggregations with Erosne bracteata, Bolbonota sp., Micrutalis sp., and Stegaspis fronditia. In PNN Farallones de Cali, this species was commonly observed (n> 30) forming mixed-species aggregations with Bolbonota (Bolbonota) sp. and Micrutalis sp., and occasionally (n = 10) with a new species of Endoiastus Fowler, Bocydium sakakibarai, and / or Stylocentrus championi. Additionally, one aggregation of nymphs was observed living inside ant shelters alongside adults and nymphs of Bolbonota sp. (Fig. 16 F). In Nuquí (Chocó), a solitary female was found on Cerro Jánano, a mountain near the Pacific coast, on an unidentified Melastomataceae at 600 masl. In Mocoa (Putumayo), a solitary female was recorded in the Amazonian foothills on an unidentified species of Miconia. Additionally, in PNN Farallones de Cali and Leticia, some adults were found parasitized by entomophagous fungi (Fig. 16 G – H).	en	Flórez-V, Camilo, Lapèze, Jérémie, Urban, Julie M. (2025): Taxonomic and ecological notes on Lycoderides Sakakibara (Hemiptera: Membracidae), including two new species from the highlands of the Colombian Andes. Zootaxa 5665 (2): 151-186, DOI: 10.11646/zootaxa.5665.2.1, URL: https://doi.org/10.11646/zootaxa.5665.2.1
03ED5C1AFFD5FA4994CFDA7AFA995010.taxon	materials_examined	Examined material and distribution: COLOMBIA: Chocó: Nuquí (Cerro Jánano - 400 masl). Meta: San Martín de los Llanos (Reserva Rey Zamuro — Matarredonda). Putumayo: Mocoa (400 masl). Risaralda: Pueblo Rico (Corregimiento Santa Cecilia — 300 – 600 masl; PNN Tatamá, vereda Montebello — 1200 – 1700 masl). Valle del Cauca: Dagua (PNN Farallones de Cali, Alto Anchicayá — 600 masl) (Fig. 1).	en	Flórez-V, Camilo, Lapèze, Jérémie, Urban, Julie M. (2025): Taxonomic and ecological notes on Lycoderides Sakakibara (Hemiptera: Membracidae), including two new species from the highlands of the Colombian Andes. Zootaxa 5665 (2): 151-186, DOI: 10.11646/zootaxa.5665.2.1, URL: https://doi.org/10.11646/zootaxa.5665.2.1
03ED5C1AFFD5FA4994CFDA7AFA995010.taxon	discussion	Remarks: The lateral lamellae in nymphs of L. phasianus are consistently broader than in other Lycoderides species, thus lateral lamellae seem much more closed (Fig. 11 G). In general shape, these nymphs closely resemble those of L. luteus. However, unlike L. luteus, no variation in nymphal morphology related to host plants has been observed. Nevertheless, a systematic collection of L. phasianus nymphs from different host plants is necessary to determine whether nymphal variation, similar to that observed in L. luteus, also occurs in this species.	en	Flórez-V, Camilo, Lapèze, Jérémie, Urban, Julie M. (2025): Taxonomic and ecological notes on Lycoderides Sakakibara (Hemiptera: Membracidae), including two new species from the highlands of the Colombian Andes. Zootaxa 5665 (2): 151-186, DOI: 10.11646/zootaxa.5665.2.1, URL: https://doi.org/10.11646/zootaxa.5665.2.1
03ED5C1AFFDAFA4994CFDA82FC2252D0.taxon	description	(Fig. 8 C – D, 9 A) This species was found on Melastomataceae in Leticia and Mitú (Colombia) (Fig. 17 A – C), where it was tended by ants of the genera Azteca and Camponotus, and the species Crematogaster cf. levior and Crematogaster cf. brasiliensis. Lapèze and Lopez-Vaamonde (2024) discussed the potential synonymy between Lycoderes argutus and Lycoderides fernandezi. Individuals from the presumably same population exhibit a high degree of variation in forewing venation, particularly in the position of the r-m crossvein relative to the M division, which varies continuously from distal to basal. These individuals have been found on the same species of Melastomataceae, suggesting that this variation may not correspond to distinct species but rather to intraspecific variability.	en	Flórez-V, Camilo, Lapèze, Jérémie, Urban, Julie M. (2025): Taxonomic and ecological notes on Lycoderides Sakakibara (Hemiptera: Membracidae), including two new species from the highlands of the Colombian Andes. Zootaxa 5665 (2): 151-186, DOI: 10.11646/zootaxa.5665.2.1, URL: https://doi.org/10.11646/zootaxa.5665.2.1
03ED5C1AFFDAFA4994CFDA82FC2252D0.taxon	materials_examined	The specimen with the following information was collected: COLOMBIA. Amazonas: Leticia, Antigua Estación Biológica El Zafire, 4 ° 0 ’ 10.296 ’’ S, 69 ° 53 ’ 44.087 ’’ W, 100 – 150 masl, July 14 / 2022, leg. C. Flórez-V, M. Idárraga, S. Quintero-M, M. Arcángel, CBUCES-F; Vaupés: Mitú, Comunidad Cucura, 1.164157 ºN, 70.169093 ºW, 240 masl, February 20 / 2024, leg. C. Flórez-V, S. Quintero-Montoya, CBUCES-F.	en	Flórez-V, Camilo, Lapèze, Jérémie, Urban, Julie M. (2025): Taxonomic and ecological notes on Lycoderides Sakakibara (Hemiptera: Membracidae), including two new species from the highlands of the Colombian Andes. Zootaxa 5665 (2): 151-186, DOI: 10.11646/zootaxa.5665.2.1, URL: https://doi.org/10.11646/zootaxa.5665.2.1
03ED5C1AFFDEFA4D94CFD9A7FD1252A7.taxon	description	(Fig. 8 E, 9 B, 17 D) A solitary adult was found on Piper sp. (Piperaceae) (Fig. 17 D). On the same plant, there were other treehopper species, two of them associated with ants (Stilbophora sp., Amastris sp. and Cladonota sp.).	en	Flórez-V, Camilo, Lapèze, Jérémie, Urban, Julie M. (2025): Taxonomic and ecological notes on Lycoderides Sakakibara (Hemiptera: Membracidae), including two new species from the highlands of the Colombian Andes. Zootaxa 5665 (2): 151-186, DOI: 10.11646/zootaxa.5665.2.1, URL: https://doi.org/10.11646/zootaxa.5665.2.1
03ED5C1AFFDEFA4D94CFD9A7FD1252A7.taxon	materials_examined	The specimens with the following information were collected: COLOMBIA. Meta: San Martín de los Llanos, Reserva Rey Zamuro Matarredonda, 3.549699 ° N, 73.404248 ° W, 250 m, February 2 / 2024, leg. C. Flórez-V, S. Quintero-Montoya, CBUCES-F 21036 (1 female in CBUCES); January 20 / 2024, CBUCES-F 21015 (1 female); COLOMBIA: Vaupés: Mitú, reserva Chundú, leg. C. Flórez-V, S. Quintero-Montoya, CBUCES-F 20894 (1 female in CBUCES). This is the first record of this species for Colombia.	en	Flórez-V, Camilo, Lapèze, Jérémie, Urban, Julie M. (2025): Taxonomic and ecological notes on Lycoderides Sakakibara (Hemiptera: Membracidae), including two new species from the highlands of the Colombian Andes. Zootaxa 5665 (2): 151-186, DOI: 10.11646/zootaxa.5665.2.1, URL: https://doi.org/10.11646/zootaxa.5665.2.1
03ED5C1AFFDFFA4C94CFDBA0FC42504F.taxon	materials_examined	Here, we record for Colombia for the first time. The specimens with the following label were examined: “ COLOMBIA. Meta. Bajo Menegua, El Lagunazo. 290 mts IV- 12 - 1984 ” (ICN: 1 female).	en	Flórez-V, Camilo, Lapèze, Jérémie, Urban, Julie M. (2025): Taxonomic and ecological notes on Lycoderides Sakakibara (Hemiptera: Membracidae), including two new species from the highlands of the Colombian Andes. Zootaxa 5665 (2): 151-186, DOI: 10.11646/zootaxa.5665.2.1, URL: https://doi.org/10.11646/zootaxa.5665.2.1
03ED5C1AFFDFFA4C94CFDA78FDB753D3.taxon	description	(Fig. 8 F, 9 C) A solitary adult was found on Clusia sp. (Clusiaceae) on a white sand amazonian forest. Some Cephalotes atratus ants were found tending nymphs on the axils of the plant. Adults and nymphs of a species of Darnini were also found on the same plant, also tended by ants (Camponotus cf. femoratus).	en	Flórez-V, Camilo, Lapèze, Jérémie, Urban, Julie M. (2025): Taxonomic and ecological notes on Lycoderides Sakakibara (Hemiptera: Membracidae), including two new species from the highlands of the Colombian Andes. Zootaxa 5665 (2): 151-186, DOI: 10.11646/zootaxa.5665.2.1, URL: https://doi.org/10.11646/zootaxa.5665.2.1
03ED5C1AFFDFFA4C94CFDA78FDB753D3.taxon	materials_examined	The specimen with the following information was collected: COLOMBIA: Vaupés: Mitú, Comunidad Cucura, 1.164157 ºN, 70.169093 ºW, 240 masl, March 8 / 2024, leg. C. Flórez-V, S. Quintero-Montoya, CBUCES-F 20715 (1 female in CBUCES). This is the first record of this species group for Colombia.	en	Flórez-V, Camilo, Lapèze, Jérémie, Urban, Julie M. (2025): Taxonomic and ecological notes on Lycoderides Sakakibara (Hemiptera: Membracidae), including two new species from the highlands of the Colombian Andes. Zootaxa 5665 (2): 151-186, DOI: 10.11646/zootaxa.5665.2.1, URL: https://doi.org/10.11646/zootaxa.5665.2.1
03ED5C1AFFDFFA4C94CFD9CCFC9C52F3.taxon	description	(Fig. 8 G, 9 D) Here, we record for Colombia for the first time. The specimens with the following data were collected: COLOMBIA. Meta: San Martín de los Llanos, reserva Rey Zamuro-Matarredonda, manual, January 20 / 2024, leg. Camilo FlórezV, Santiago Quintero-M, CBUCES-F 21015 (1 female).	en	Flórez-V, Camilo, Lapèze, Jérémie, Urban, Julie M. (2025): Taxonomic and ecological notes on Lycoderides Sakakibara (Hemiptera: Membracidae), including two new species from the highlands of the Colombian Andes. Zootaxa 5665 (2): 151-186, DOI: 10.11646/zootaxa.5665.2.1, URL: https://doi.org/10.11646/zootaxa.5665.2.1
03ED5C1AFFE3FA7094CFDA28FAA153F6.taxon	description	(Fig. 8 H, 9 E) We found solitary adults under leaves of Melastomataceae, on plants with parabiotic ants (Camponotus femoratus and Crematogaster cf. limata) and other treehoppers (Dioclophara sp., Tropidolomia sp., Anobilia aff. guianae). In MPUJ, specimens were labeled as collected on Melastomataceae. This is the first record of this species for Colombia.	en	Flórez-V, Camilo, Lapèze, Jérémie, Urban, Julie M. (2025): Taxonomic and ecological notes on Lycoderides Sakakibara (Hemiptera: Membracidae), including two new species from the highlands of the Colombian Andes. Zootaxa 5665 (2): 151-186, DOI: 10.11646/zootaxa.5665.2.1, URL: https://doi.org/10.11646/zootaxa.5665.2.1
03ED5C1AFFE3FA7094CFDA28FAA153F6.taxon	materials_examined	The specimens with the following labels were examined: “ COLOMBIA Casanare, Taura- / mena, Kiosco Verde,. 5 km / SW de Tauramena, 526 m, / 5.00385 ° N 72.77376 ° W, 8 – 12 / Sep 2014, D. Forero ”, “ Ex: Melastomataceae / nocturna ” (3 females, 1 male in MPUJ). “ COLOMBIA. Vaupés, Mitú, Carretera a la comunidad Tayazu, 1.0975 ° N, 70.0461111 ° W, 171 m, manual, en Melastomataceae, Mar. 3 / 2024, leg. Camilo Flórez-V, Santiago Quintero-Montoya, CBUCES-F 20988 ” (1 male in CBUCES); “ COLOMBIA. Vaupés, Mitú, reserva Chundú, manual, feb. 18 / 2024, leg. Camilo Flórez-V, Santiago Quintero-Montoya, CBUCES-F 20883 ” (1 female in CBUCES).	en	Flórez-V, Camilo, Lapèze, Jérémie, Urban, Julie M. (2025): Taxonomic and ecological notes on Lycoderides Sakakibara (Hemiptera: Membracidae), including two new species from the highlands of the Colombian Andes. Zootaxa 5665 (2): 151-186, DOI: 10.11646/zootaxa.5665.2.1, URL: https://doi.org/10.11646/zootaxa.5665.2.1
