taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
03F6C023FFBD2C4BFC5D9D04E33EF9FB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15536145/files/figure.png	https://doi.org/10.5281/zenodo.15536145	Fig. 1. Map of the sampled localities in Thừa Thiên-Huế Province, Vietnam.	Fig. 1. Map of the sampled localities in Thừa Thiên-Huế Province, Vietnam.	2025-05-27	Constant, Jérôme;Pham, Hong Thai		Zenodo	biologists	Constant, Jérôme;Pham, Hong Thai			
03F6C023FFBD2C4BFC5D9D04E33EF9FB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15536147/files/figure.png	https://doi.org/10.5281/zenodo.15536147	Fig. 2. Bach Ma National Park, habitats. A. Detailed map of the sampling areas (1 = ‘Yes Hue Eco’; 2 = ‘pheasant trail’; 3 = ‘low altitude’; 4 = ‘roadside’; 5 = ‘summit’; 6 = ‘near ranger station’. B. Typical habitat at ‘Yes Hue Eco’ sampling area, 17 May 2023.	Fig. 2. Bach Ma National Park, habitats. A. Detailed map of the sampling areas (1 = ‘Yes Hue Eco’; 2 = ‘pheasant trail’; 3 = ‘low altitude’; 4 = ‘roadside’; 5 = ‘summit’; 6 = ‘near ranger station’. B. Typical habitat at ‘Yes Hue Eco’ sampling area, 17 May 2023.	2025-05-27	Constant, Jérôme;Pham, Hong Thai		Zenodo	biologists	Constant, Jérôme;Pham, Hong Thai			
03F6C023FFBD2C4BFC5D9D04E33EF9FB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15536149/files/figure.png	https://doi.org/10.5281/zenodo.15536149	Fig. 3. Bach Ma National Park, habitats. A. Typical habitat at ‘pheasant trail’ sampling area, 20 May 2023. B. Typical habitat at ‘low altitude’ sampling area, 15 May 2023.	Fig. 3. Bach Ma National Park, habitats. A. Typical habitat at ‘pheasant trail’ sampling area, 20 May 2023. B. Typical habitat at ‘low altitude’ sampling area, 15 May 2023.	2025-05-27	Constant, Jérôme;Pham, Hong Thai		Zenodo	biologists	Constant, Jérôme;Pham, Hong Thai			
03F6C023FFBD2C4BFC5D9D04E33EF9FB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15536151/files/figure.png	https://doi.org/10.5281/zenodo.15536151	Fig. 4. Bach Ma National Park, habitats. A. Typical habitat at ‘roadside’ sampling area, 21 May 2023. B. Typical habitat at ‘summit’ sampling area, 11 May 2023.	Fig. 4. Bach Ma National Park, habitats. A. Typical habitat at ‘roadside’ sampling area, 21 May 2023. B. Typical habitat at ‘summit’ sampling area, 11 May 2023.	2025-05-27	Constant, Jérôme;Pham, Hong Thai		Zenodo	biologists	Constant, Jérôme;Pham, Hong Thai			
03F6C023FFBD2C4BFC5D9D04E33EF9FB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15536153/files/figure.png	https://doi.org/10.5281/zenodo.15536153	Fig. 5. Bach Ma National Park and Phong Dien District, habitats. A. Typical habitat at ‘ranger station’ sampling area, 18 May 2023. B. Typical habitat at Phong Dien sampling area, 23 May 2023.	Fig. 5. Bach Ma National Park and Phong Dien District, habitats. A. Typical habitat at ‘ranger station’ sampling area, 18 May 2023. B. Typical habitat at Phong Dien sampling area, 23 May 2023.	2025-05-27	Constant, Jérôme;Pham, Hong Thai		Zenodo	biologists	Constant, Jérôme;Pham, Hong Thai			
03F6C023FFBD2C4BFC5D9D04E33EF9FB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15536155/files/figure.png	https://doi.org/10.5281/zenodo.15536155	Fig. 6. Ceratogergithus pictus sp. nov., dissected paratype, ♂ (RBINS). A. Habitus, dorsal view. B. Habitus, ventral view. C. Habitus, lateral view. D. Habitus, perpendicular view of frons. E. Habitus, anterolateral view. F. Right hind wing. G. Metatarsus and apex of metatibia, ventral view. H. Right hind leg, ventral view.	Fig. 6. Ceratogergithus pictus sp. nov., dissected paratype, ♂ (RBINS). A. Habitus, dorsal view. B. Habitus, ventral view. C. Habitus, lateral view. D. Habitus, perpendicular view of frons. E. Habitus, anterolateral view. F. Right hind wing. G. Metatarsus and apex of metatibia, ventral view. H. Right hind leg, ventral view.	2025-05-27	Constant, Jérôme;Pham, Hong Thai		Zenodo	biologists	Constant, Jérôme;Pham, Hong Thai			
03F6C023FFBD2C4BFC5D9D04E33EF9FB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15536157/files/figure.png	https://doi.org/10.5281/zenodo.15536157	Fig. 7. Ceratogergithus pictus sp. nov., holotype, ♂ (VNMN), terminalia. A–D. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Posterolateral view. C. Dorsal view. D. Caudal view. E–L. Aedeagus. E. Dorsal view. F. Anterodorsal view. G. Left lateral view. H. Posteroventral view. I. Ventral view. J. Left laterodorsal view. K. Left lateroventral view. L. Posterolateral view. Abbreviations: see Material and methods.	Fig. 7. Ceratogergithus pictus sp. nov., holotype, ♂ (VNMN), terminalia. A–D. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Posterolateral view. C. Dorsal view. D. Caudal view. E–L. Aedeagus. E. Dorsal view. F. Anterodorsal view. G. Left lateral view. H. Posteroventral view. I. Ventral view. J. Left laterodorsal view. K. Left lateroventral view. L. Posterolateral view. Abbreviations: see Material and methods.	2025-05-27	Constant, Jérôme;Pham, Hong Thai		Zenodo	biologists	Constant, Jérôme;Pham, Hong Thai			
03F6C023FFBD2C4BFC5D9D04E33EF9FB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15536159/files/figure.png	https://doi.org/10.5281/zenodo.15536159	Fig. 8. Ceratogergithus pictus sp. nov., live specimens in Bach Ma National Park. A–C. Pheasant trail, 12 May 2023.D. Near ranger station, 18 May 2023.E–F. Summit trail, 20 May 2023.G–H. Trophobiosis with ant, Camponotus sp., near ranger station, 17 May 2023.	Fig. 8. Ceratogergithus pictus sp. nov., live specimens in Bach Ma National Park. A–C. Pheasant trail, 12 May 2023.D. Near ranger station, 18 May 2023.E–F. Summit trail, 20 May 2023.G–H. Trophobiosis with ant, Camponotus sp., near ranger station, 17 May 2023.	2025-05-27	Constant, Jérôme;Pham, Hong Thai		Zenodo	biologists	Constant, Jérôme;Pham, Hong Thai			
03F6C023FFA42C52FC659DC2E5DAFD36.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15536145/files/figure.png	https://doi.org/10.5281/zenodo.15536145	Fig. 1. Map of the sampled localities in Thừa Thiên-Huế Province, Vietnam.	Fig. 1. Map of the sampled localities in Thừa Thiên-Huế Province, Vietnam.	2025-05-27	Constant, Jérôme;Pham, Hong Thai		Zenodo	biologists	Constant, Jérôme;Pham, Hong Thai			
03F6C023FFA42C52FC659DC2E5DAFD36.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15536147/files/figure.png	https://doi.org/10.5281/zenodo.15536147	Fig. 2. Bach Ma National Park, habitats. A. Detailed map of the sampling areas (1 = ‘Yes Hue Eco’; 2 = ‘pheasant trail’; 3 = ‘low altitude’; 4 = ‘roadside’; 5 = ‘summit’; 6 = ‘near ranger station’. B. Typical habitat at ‘Yes Hue Eco’ sampling area, 17 May 2023.	Fig. 2. Bach Ma National Park, habitats. A. Detailed map of the sampling areas (1 = ‘Yes Hue Eco’; 2 = ‘pheasant trail’; 3 = ‘low altitude’; 4 = ‘roadside’; 5 = ‘summit’; 6 = ‘near ranger station’. B. Typical habitat at ‘Yes Hue Eco’ sampling area, 17 May 2023.	2025-05-27	Constant, Jérôme;Pham, Hong Thai		Zenodo	biologists	Constant, Jérôme;Pham, Hong Thai			
03F6C023FFA42C52FC659DC2E5DAFD36.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15536151/files/figure.png	https://doi.org/10.5281/zenodo.15536151	Fig. 4. Bach Ma National Park, habitats. A. Typical habitat at ‘roadside’ sampling area, 21 May 2023. B. Typical habitat at ‘summit’ sampling area, 11 May 2023.	Fig. 4. Bach Ma National Park, habitats. A. Typical habitat at ‘roadside’ sampling area, 21 May 2023. B. Typical habitat at ‘summit’ sampling area, 11 May 2023.	2025-05-27	Constant, Jérôme;Pham, Hong Thai		Zenodo	biologists	Constant, Jérôme;Pham, Hong Thai			
03F6C023FFA42C52FC659DC2E5DAFD36.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15536161/files/figure.png	https://doi.org/10.5281/zenodo.15536161	Fig. 9. Gergithoides devyveri sp. nov., dissected paratype, ♂ (RBINS). A. Habitus, dorsal view. B. Habitus, ventral view. C. Habitus, lateral view. D. Habitus, perpendicular view of frons. E. Habitus, anterolateral view. F. Right hind wing. G. Right hind leg, ventral view. H. Metatarsus and apex of metatibia, ventral view.	Fig. 9. Gergithoides devyveri sp. nov., dissected paratype, ♂ (RBINS). A. Habitus, dorsal view. B. Habitus, ventral view. C. Habitus, lateral view. D. Habitus, perpendicular view of frons. E. Habitus, anterolateral view. F. Right hind wing. G. Right hind leg, ventral view. H. Metatarsus and apex of metatibia, ventral view.	2025-05-27	Constant, Jérôme;Pham, Hong Thai		Zenodo	biologists	Constant, Jérôme;Pham, Hong Thai			
03F6C023FFA42C52FC659DC2E5DAFD36.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15536165/files/figure.png	https://doi.org/10.5281/zenodo.15536165	Fig. 10. Gergithoides devyveri sp. nov., holotype, ♂ (VNMN), terminalia. A–D. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Caudal view. C. Posterolateral view. D. Dorsal view. E–N. Aedeagus. E. Posteroventral view. F. Anterodorsal view. G. Dorsal view. H. Left lateral view. I. Right lateral view. J. Left laterodorsal view. K. Right laterodorsal view. L. Ventral view. M. Left posterolateral view. N. Right posterolateral view. Abbreviations: see Material and methods.	Fig. 10. Gergithoides devyveri sp. nov., holotype, ♂ (VNMN), terminalia. A–D. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Caudal view. C. Posterolateral view. D. Dorsal view. E–N. Aedeagus. E. Posteroventral view. F. Anterodorsal view. G. Dorsal view. H. Left lateral view. I. Right lateral view. J. Left laterodorsal view. K. Right laterodorsal view. L. Ventral view. M. Left posterolateral view. N. Right posterolateral view. Abbreviations: see Material and methods.	2025-05-27	Constant, Jérôme;Pham, Hong Thai		Zenodo	biologists	Constant, Jérôme;Pham, Hong Thai			
03F6C023FFA42C52FC659DC2E5DAFD36.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15536167/files/figure.png	https://doi.org/10.5281/zenodo.15536167	Fig. 11. Gergithoides devyveri sp. nov., Bach Ma National Park. A–D. Summit trail, specimens, 11–13 May 2023. E–H. Rhododendron trail, 14 May 2023. E–G. Specimens. H. Microhabitat.	Fig. 11. Gergithoides devyveri sp. nov., Bach Ma National Park. A–D. Summit trail, specimens, 11–13 May 2023. E–H. Rhododendron trail, 14 May 2023. E–G. Specimens. H. Microhabitat.	2025-05-27	Constant, Jérôme;Pham, Hong Thai		Zenodo	biologists	Constant, Jérôme;Pham, Hong Thai			
03F6C023FFA22C64FC3D9E16E59BFE6C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15536147/files/figure.png	https://doi.org/10.5281/zenodo.15536147	Fig. 2. Bach Ma National Park, habitats. A. Detailed map of the sampling areas (1 = ‘Yes Hue Eco’; 2 = ‘pheasant trail’; 3 = ‘low altitude’; 4 = ‘roadside’; 5 = ‘summit’; 6 = ‘near ranger station’. B. Typical habitat at ‘Yes Hue Eco’ sampling area, 17 May 2023.	Fig. 2. Bach Ma National Park, habitats. A. Detailed map of the sampling areas (1 = ‘Yes Hue Eco’; 2 = ‘pheasant trail’; 3 = ‘low altitude’; 4 = ‘roadside’; 5 = ‘summit’; 6 = ‘near ranger station’. B. Typical habitat at ‘Yes Hue Eco’ sampling area, 17 May 2023.	2025-05-27	Constant, Jérôme;Pham, Hong Thai		Zenodo	biologists	Constant, Jérôme;Pham, Hong Thai			
03F6C023FFA22C64FC3D9E16E59BFE6C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15536149/files/figure.png	https://doi.org/10.5281/zenodo.15536149	Fig. 3. Bach Ma National Park, habitats. A. Typical habitat at ‘pheasant trail’ sampling area, 20 May 2023. B. Typical habitat at ‘low altitude’ sampling area, 15 May 2023.	Fig. 3. Bach Ma National Park, habitats. A. Typical habitat at ‘pheasant trail’ sampling area, 20 May 2023. B. Typical habitat at ‘low altitude’ sampling area, 15 May 2023.	2025-05-27	Constant, Jérôme;Pham, Hong Thai		Zenodo	biologists	Constant, Jérôme;Pham, Hong Thai			
03F6C023FFA22C64FC3D9E16E59BFE6C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15536151/files/figure.png	https://doi.org/10.5281/zenodo.15536151	Fig. 4. Bach Ma National Park, habitats. A. Typical habitat at ‘roadside’ sampling area, 21 May 2023. B. Typical habitat at ‘summit’ sampling area, 11 May 2023.	Fig. 4. Bach Ma National Park, habitats. A. Typical habitat at ‘roadside’ sampling area, 21 May 2023. B. Typical habitat at ‘summit’ sampling area, 11 May 2023.	2025-05-27	Constant, Jérôme;Pham, Hong Thai		Zenodo	biologists	Constant, Jérôme;Pham, Hong Thai			
03F6C023FFA22C64FC3D9E16E59BFE6C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15536153/files/figure.png	https://doi.org/10.5281/zenodo.15536153	Fig. 5. Bach Ma National Park and Phong Dien District, habitats. A. Typical habitat at ‘ranger station’ sampling area, 18 May 2023. B. Typical habitat at Phong Dien sampling area, 23 May 2023.	Fig. 5. Bach Ma National Park and Phong Dien District, habitats. A. Typical habitat at ‘ranger station’ sampling area, 18 May 2023. B. Typical habitat at Phong Dien sampling area, 23 May 2023.	2025-05-27	Constant, Jérôme;Pham, Hong Thai		Zenodo	biologists	Constant, Jérôme;Pham, Hong Thai			
03F6C023FFA22C64FC3D9E16E59BFE6C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15536169/files/figure.png	https://doi.org/10.5281/zenodo.15536169	Fig. 12. Hemisphaerius annamiticus sp. nov., dissected holotype, ♂ (VNMN). A. Habitus, dorsal view. B. Habitus, ventral view. C. Habitus, lateral view. D. Right hind wing. E. Habitus, perpendicular view of frons. F. Habitus, anterolateral view.	Fig. 12. Hemisphaerius annamiticus sp. nov., dissected holotype, ♂ (VNMN). A. Habitus, dorsal view. B. Habitus, ventral view. C. Habitus, lateral view. D. Right hind wing. E. Habitus, perpendicular view of frons. F. Habitus, anterolateral view.	2025-05-27	Constant, Jérôme;Pham, Hong Thai		Zenodo	biologists	Constant, Jérôme;Pham, Hong Thai			
03F6C023FFA22C64FC3D9E16E59BFE6C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15536172/files/figure.png	https://doi.org/10.5281/zenodo.15536172	Fig. 13. Hemisphaerius annamiticus sp. nov., dissected paratype, ♂ from Da Krong, Quang Tri Province (RBINS). A. Habitus, dorsal view. B. Habitus, ventral view. C. Habitus, lateral view. D. Habitus, perpendicular view of frons. E. Habitus, anterolateral view. F. Right hind wing.	Fig. 13. Hemisphaerius annamiticus sp. nov., dissected paratype, ♂ from Da Krong, Quang Tri Province (RBINS). A. Habitus, dorsal view. B. Habitus, ventral view. C. Habitus, lateral view. D. Habitus, perpendicular view of frons. E. Habitus, anterolateral view. F. Right hind wing.	2025-05-27	Constant, Jérôme;Pham, Hong Thai		Zenodo	biologists	Constant, Jérôme;Pham, Hong Thai			
03F6C023FFA22C64FC3D9E16E59BFE6C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15536174/files/figure.png	https://doi.org/10.5281/zenodo.15536174	Fig. 14. Hemisphaerius annamiticus sp. nov., dissected paratype, ♂ from Mang Kanh, Kon Tum Province (RBINS). A. Habitus, dorsal view. B. Habitus, ventral view. C. Habitus, lateral view. D. Habitus, perpendicular view of frons. E. Habitus, anterolateral view.	Fig. 14. Hemisphaerius annamiticus sp. nov., dissected paratype, ♂ from Mang Kanh, Kon Tum Province (RBINS). A. Habitus, dorsal view. B. Habitus, ventral view. C. Habitus, lateral view. D. Habitus, perpendicular view of frons. E. Habitus, anterolateral view.	2025-05-27	Constant, Jérôme;Pham, Hong Thai		Zenodo	biologists	Constant, Jérôme;Pham, Hong Thai			
03F6C023FFA22C64FC3D9E16E59BFE6C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15536176/files/figure.png	https://doi.org/10.5281/zenodo.15536176	Fig. 15. Hemisphaerius annamiticus sp. nov., holotype, ♂ (VNMN), terminalia. A–F. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Posterolateral view. C. Caudal view. D. Dorsal view. E. Ventral view. F. Posterodorsal view. G–R. Aedeagus.G. Caudal view. H. Left lateral view. I. Right lateral view. J. Posteroventral view. K. Left laterodorsal view. L. Right laterodorsal view. M. Left posterolateral view. N. Left lateroventral view. O. Right lateroventral view. P. Dorsal view. Q. Ventral view. R. Right posterolateral view. Abbreviations: see Material and methods.	Fig. 15. Hemisphaerius annamiticus sp. nov., holotype, ♂ (VNMN), terminalia. A–F. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Posterolateral view. C. Caudal view. D. Dorsal view. E. Ventral view. F. Posterodorsal view. G–R. Aedeagus.G. Caudal view. H. Left lateral view. I. Right lateral view. J. Posteroventral view. K. Left laterodorsal view. L. Right laterodorsal view. M. Left posterolateral view. N. Left lateroventral view. O. Right lateroventral view. P. Dorsal view. Q. Ventral view. R. Right posterolateral view. Abbreviations: see Material and methods.	2025-05-27	Constant, Jérôme;Pham, Hong Thai		Zenodo	biologists	Constant, Jérôme;Pham, Hong Thai			
03F6C023FFA22C64FC3D9E16E59BFE6C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15536180/files/figure.png	https://doi.org/10.5281/zenodo.15536180	Fig. 16. Hemisphaerius annamiticus sp. nov., live specimens in Bach Ma National Park. A–D. Pheasant trail. A–B. 12 May 2023. C–D. 20 May 2023. E–G. Low altitude location, 15 May 2023. H. Near ranger station, 18 May 2023.	Fig. 16. Hemisphaerius annamiticus sp. nov., live specimens in Bach Ma National Park. A–D. Pheasant trail. A–B. 12 May 2023. C–D. 20 May 2023. E–G. Low altitude location, 15 May 2023. H. Near ranger station, 18 May 2023.	2025-05-27	Constant, Jérôme;Pham, Hong Thai		Zenodo	biologists	Constant, Jérôme;Pham, Hong Thai			
03F6C023FFA22C64FC3D9E16E59BFE6C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15536182/files/figure.png	https://doi.org/10.5281/zenodo.15536182	Fig. 17. Hemisphaerius annamiticus sp. nov., distribution map.	Fig. 17. Hemisphaerius annamiticus sp. nov., distribution map.	2025-05-27	Constant, Jérôme;Pham, Hong Thai		Zenodo	biologists	Constant, Jérôme;Pham, Hong Thai			
03F6C023FF942C63FC4C9992E24EFA29.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15536147/files/figure.png	https://doi.org/10.5281/zenodo.15536147	Fig. 2. Bach Ma National Park, habitats. A. Detailed map of the sampling areas (1 = ‘Yes Hue Eco’; 2 = ‘pheasant trail’; 3 = ‘low altitude’; 4 = ‘roadside’; 5 = ‘summit’; 6 = ‘near ranger station’. B. Typical habitat at ‘Yes Hue Eco’ sampling area, 17 May 2023.	Fig. 2. Bach Ma National Park, habitats. A. Detailed map of the sampling areas (1 = ‘Yes Hue Eco’; 2 = ‘pheasant trail’; 3 = ‘low altitude’; 4 = ‘roadside’; 5 = ‘summit’; 6 = ‘near ranger station’. B. Typical habitat at ‘Yes Hue Eco’ sampling area, 17 May 2023.	2025-05-27	Constant, Jérôme;Pham, Hong Thai		Zenodo	biologists	Constant, Jérôme;Pham, Hong Thai			
03F6C023FF942C63FC4C9992E24EFA29.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15536149/files/figure.png	https://doi.org/10.5281/zenodo.15536149	Fig. 3. Bach Ma National Park, habitats. A. Typical habitat at ‘pheasant trail’ sampling area, 20 May 2023. B. Typical habitat at ‘low altitude’ sampling area, 15 May 2023.	Fig. 3. Bach Ma National Park, habitats. A. Typical habitat at ‘pheasant trail’ sampling area, 20 May 2023. B. Typical habitat at ‘low altitude’ sampling area, 15 May 2023.	2025-05-27	Constant, Jérôme;Pham, Hong Thai		Zenodo	biologists	Constant, Jérôme;Pham, Hong Thai			
03F6C023FF942C63FC4C9992E24EFA29.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15536185/files/figure.png	https://doi.org/10.5281/zenodo.15536185	Fig. 18. Hemisphaerius thaydoius sp. nov., dissected holotype, ♂ (VNMN). A. Habitus, dorsal view. B. Habitus, ventral view. C. Habitus, lateral view. D. Habitus, perpendicular view of frons. E. Habitus, anterolateral view. F. Right hind wing.	Fig. 18. Hemisphaerius thaydoius sp. nov., dissected holotype, ♂ (VNMN). A. Habitus, dorsal view. B. Habitus, ventral view. C. Habitus, lateral view. D. Habitus, perpendicular view of frons. E. Habitus, anterolateral view. F. Right hind wing.	2025-05-27	Constant, Jérôme;Pham, Hong Thai		Zenodo	biologists	Constant, Jérôme;Pham, Hong Thai			
03F6C023FF942C63FC4C9992E24EFA29.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15536187/files/figure.png	https://doi.org/10.5281/zenodo.15536187	Fig. 19. Hemisphaerius thaydoius sp. nov., dissected paratype, ♂, black morph, from Bach Ma National Park (RBINS). A. Habitus, dorsal view. B. Habitus, ventral view. C. Habitus, lateral view. D. Habitus, perpendicular view of frons. E. Habitus, anterolateral view. F. Right hind wing. G. Left hind leg, ventral view. H. Metatarsus and apex of metatibia, ventral view.	Fig. 19. Hemisphaerius thaydoius sp. nov., dissected paratype, ♂, black morph, from Bach Ma National Park (RBINS). A. Habitus, dorsal view. B. Habitus, ventral view. C. Habitus, lateral view. D. Habitus, perpendicular view of frons. E. Habitus, anterolateral view. F. Right hind wing. G. Left hind leg, ventral view. H. Metatarsus and apex of metatibia, ventral view.	2025-05-27	Constant, Jérôme;Pham, Hong Thai		Zenodo	biologists	Constant, Jérôme;Pham, Hong Thai			
03F6C023FF942C63FC4C9992E24EFA29.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15536189/files/figure.png	https://doi.org/10.5281/zenodo.15536189	Fig. 20. Hemisphaerius thaydoius sp. nov., dissected holotype, ♂ (VNMN), terminalia. A–D. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Posterolateral view. C. Caudal view. D. Dorsal view. E–N.Aedeagus.E. Left posterolateral view.F. Caudal view.G. Right posterolateral view.H.Anterodorsal view. I. Left lateral view. J. Right lateral view. K. Left laterodorsal view. L. Right laterodorsal view. M. Dorsal view. N. Ventral view. Abbreviations: see Material and methods.	Fig. 20. Hemisphaerius thaydoius sp. nov., dissected holotype, ♂ (VNMN), terminalia. A–D. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Posterolateral view. C. Caudal view. D. Dorsal view. E–N.Aedeagus.E. Left posterolateral view.F. Caudal view.G. Right posterolateral view.H.Anterodorsal view. I. Left lateral view. J. Right lateral view. K. Left laterodorsal view. L. Right laterodorsal view. M. Dorsal view. N. Ventral view. Abbreviations: see Material and methods.	2025-05-27	Constant, Jérôme;Pham, Hong Thai		Zenodo	biologists	Constant, Jérôme;Pham, Hong Thai			
03F6C023FF942C63FC4C9992E24EFA29.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15536193/files/figure.png	https://doi.org/10.5281/zenodo.15536193	Fig. 21. Hemisphaerius thaydoius sp. nov. A–B. Live specimen in Bach Ma National Park, pheasant trail, 12 May 2023.C–D. Bach Ma National Park, near ranger station. C. 18 May 2023. D. 19 Oct. 2024 (photographed in cage). E–H. Phong Dien District, 23 May 2023. E–G. Specimen. H. Microhabitat.	Fig. 21. Hemisphaerius thaydoius sp. nov. A–B. Live specimen in Bach Ma National Park, pheasant trail, 12 May 2023.C–D. Bach Ma National Park, near ranger station. C. 18 May 2023. D. 19 Oct. 2024 (photographed in cage). E–H. Phong Dien District, 23 May 2023. E–G. Specimen. H. Microhabitat.	2025-05-27	Constant, Jérôme;Pham, Hong Thai		Zenodo	biologists	Constant, Jérôme;Pham, Hong Thai			
03F6C023FF942C63FC4C9992E24EFA29.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15536197/files/figure.png	https://doi.org/10.5281/zenodo.15536197	Fig. 22. Hemisphaerius thaydoius sp. nov., distribution map.	Fig. 22. Hemisphaerius thaydoius sp. nov., distribution map.	2025-05-27	Constant, Jérôme;Pham, Hong Thai		Zenodo	biologists	Constant, Jérôme;Pham, Hong Thai			
03F6C023FF942C63FC4C9992E24EFA29.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15536153/files/figure.png	https://doi.org/10.5281/zenodo.15536153	Fig. 5. Bach Ma National Park and Phong Dien District, habitats. A. Typical habitat at ‘ranger station’ sampling area, 18 May 2023. B. Typical habitat at Phong Dien sampling area, 23 May 2023.	Fig. 5. Bach Ma National Park and Phong Dien District, habitats. A. Typical habitat at ‘ranger station’ sampling area, 18 May 2023. B. Typical habitat at Phong Dien sampling area, 23 May 2023.	2025-05-27	Constant, Jérôme;Pham, Hong Thai		Zenodo	biologists	Constant, Jérôme;Pham, Hong Thai			
03F6C023FF9C2C68FC619DFCE5C6F834.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15536145/files/figure.png	https://doi.org/10.5281/zenodo.15536145	Fig. 1. Map of the sampled localities in Thừa Thiên-Huế Province, Vietnam.	Fig. 1. Map of the sampled localities in Thừa Thiên-Huế Province, Vietnam.	2025-05-27	Constant, Jérôme;Pham, Hong Thai		Zenodo	biologists	Constant, Jérôme;Pham, Hong Thai			
03F6C023FF9C2C68FC619DFCE5C6F834.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15536147/files/figure.png	https://doi.org/10.5281/zenodo.15536147	Fig. 2. Bach Ma National Park, habitats. A. Detailed map of the sampling areas (1 = ‘Yes Hue Eco’; 2 = ‘pheasant trail’; 3 = ‘low altitude’; 4 = ‘roadside’; 5 = ‘summit’; 6 = ‘near ranger station’. B. Typical habitat at ‘Yes Hue Eco’ sampling area, 17 May 2023.	Fig. 2. Bach Ma National Park, habitats. A. Detailed map of the sampling areas (1 = ‘Yes Hue Eco’; 2 = ‘pheasant trail’; 3 = ‘low altitude’; 4 = ‘roadside’; 5 = ‘summit’; 6 = ‘near ranger station’. B. Typical habitat at ‘Yes Hue Eco’ sampling area, 17 May 2023.	2025-05-27	Constant, Jérôme;Pham, Hong Thai		Zenodo	biologists	Constant, Jérôme;Pham, Hong Thai			
03F6C023FF9C2C68FC619DFCE5C6F834.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15536151/files/figure.png	https://doi.org/10.5281/zenodo.15536151	Fig. 4. Bach Ma National Park, habitats. A. Typical habitat at ‘roadside’ sampling area, 21 May 2023. B. Typical habitat at ‘summit’ sampling area, 11 May 2023.	Fig. 4. Bach Ma National Park, habitats. A. Typical habitat at ‘roadside’ sampling area, 21 May 2023. B. Typical habitat at ‘summit’ sampling area, 11 May 2023.	2025-05-27	Constant, Jérôme;Pham, Hong Thai		Zenodo	biologists	Constant, Jérôme;Pham, Hong Thai			
03F6C023FF9C2C68FC619DFCE5C6F834.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15536201/files/figure.png	https://doi.org/10.5281/zenodo.15536201	Fig. 23. Ishiharanus dinhanus sp. nov., dissected holotype, ♂ (VNMN). A. Habitus, dorsal view. B. Habitus, ventral view. C. Right hind wing. D. Habitus, lateral view. E. Habitus, perpendicular view of frons. F. Habitus, anterolateral view.	Fig. 23. Ishiharanus dinhanus sp. nov., dissected holotype, ♂ (VNMN). A. Habitus, dorsal view. B. Habitus, ventral view. C. Right hind wing. D. Habitus, lateral view. E. Habitus, perpendicular view of frons. F. Habitus, anterolateral view.	2025-05-27	Constant, Jérôme;Pham, Hong Thai		Zenodo	biologists	Constant, Jérôme;Pham, Hong Thai			
03F6C023FF9C2C68FC619DFCE5C6F834.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15536203/files/figure.png	https://doi.org/10.5281/zenodo.15536203	Fig. 24. Ishiharanus dinhanus sp. nov., holotype, ♂ (VNMN), terminalia. A–F. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Posterolateral view. C. Caudal view. D. Dorsal view. E. Ventral view. F. Posterodorsal view. G–R. Aedeagus. G. Left lateral view. H. Right lateral view. I. Caudal view. J. Ventral view. K. Left laterodorsal view. L. Right laterodorsal view. M. Right posterolateral view. N. Left lateroventral view. O. Right lateroventral view. P. Dorsal view. Q. Anterodorsal view. R. Left posterolateral view. Abbreviations: see Material and methods.	Fig. 24. Ishiharanus dinhanus sp. nov., holotype, ♂ (VNMN), terminalia. A–F. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Posterolateral view. C. Caudal view. D. Dorsal view. E. Ventral view. F. Posterodorsal view. G–R. Aedeagus. G. Left lateral view. H. Right lateral view. I. Caudal view. J. Ventral view. K. Left laterodorsal view. L. Right laterodorsal view. M. Right posterolateral view. N. Left lateroventral view. O. Right lateroventral view. P. Dorsal view. Q. Anterodorsal view. R. Left posterolateral view. Abbreviations: see Material and methods.	2025-05-27	Constant, Jérôme;Pham, Hong Thai		Zenodo	biologists	Constant, Jérôme;Pham, Hong Thai			
03F6C023FF9C2C68FC619DFCE5C6F834.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15536205/files/figure.png	https://doi.org/10.5281/zenodo.15536205	Fig. 25. Ishiharanus dinhanus sp. nov., live specimen from Bach Ma National Park summit, 19 May 2023, photographed in cage. A. Dorsal view. B. Laterodorsal view.	Fig. 25. Ishiharanus dinhanus sp. nov., live specimen from Bach Ma National Park summit, 19 May 2023, photographed in cage. A. Dorsal view. B. Laterodorsal view.	2025-05-27	Constant, Jérôme;Pham, Hong Thai		Zenodo	biologists	Constant, Jérôme;Pham, Hong Thai			
03F6C023FF9C2C68FC619DFCE5C6F834.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15536221/files/figure.png	https://doi.org/10.5281/zenodo.15536221	Fig. 29. Ishiharanus pulchellus sp. nov., dissected holotype, ♂ (VNMN). A. Habitus, dorsal view. B. Habitus, ventral view. C. Habitus, lateral view. D. Habitus, perpendicular view of frons. E. Habitus, anterolateral view. F. Right hind wing. G. Right hind leg, ventral view. H. Metatarsus and apex of metatibia, ventral view.	Fig. 29. Ishiharanus pulchellus sp. nov., dissected holotype, ♂ (VNMN). A. Habitus, dorsal view. B. Habitus, ventral view. C. Habitus, lateral view. D. Habitus, perpendicular view of frons. E. Habitus, anterolateral view. F. Right hind wing. G. Right hind leg, ventral view. H. Metatarsus and apex of metatibia, ventral view.	2025-05-27	Constant, Jérôme;Pham, Hong Thai		Zenodo	biologists	Constant, Jérôme;Pham, Hong Thai			
03F6C023FF9C2C68FC619DFCE5C6F834.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15536223/files/figure.png	https://doi.org/10.5281/zenodo.15536223	Fig. 30. Ishiharanus pulchellus sp. nov., dissected paratype, ♂ (RBINS) from Bach Ma National Park, dark morph. A. Habitus, dorsal view. B. Habitus, ventral view. C. Habitus, lateral view. D. Habitus, perpendicular view of frons. E. Habitus, anterolateral view. F. Right hind wing.	Fig. 30. Ishiharanus pulchellus sp. nov., dissected paratype, ♂ (RBINS) from Bach Ma National Park, dark morph. A. Habitus, dorsal view. B. Habitus, ventral view. C. Habitus, lateral view. D. Habitus, perpendicular view of frons. E. Habitus, anterolateral view. F. Right hind wing.	2025-05-27	Constant, Jérôme;Pham, Hong Thai		Zenodo	biologists	Constant, Jérôme;Pham, Hong Thai			
03F6C023FF9C2C68FC619DFCE5C6F834.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15536225/files/figure.png	https://doi.org/10.5281/zenodo.15536225	Fig. 31. Ishiharanus pulchellus sp. nov., dissected paratype, ♂ (RBINS) from Kon Chu Rang National Park, pale morph. A. Habitus, dorsal view. B. Habitus, ventral view. C. Habitus, lateral view. D. Habitus, perpendicular view of frons. E. Habitus, anterolateral view. F. Right hind wing. G. Caudal view.	Fig. 31. Ishiharanus pulchellus sp. nov., dissected paratype, ♂ (RBINS) from Kon Chu Rang National Park, pale morph. A. Habitus, dorsal view. B. Habitus, ventral view. C. Habitus, lateral view. D. Habitus, perpendicular view of frons. E. Habitus, anterolateral view. F. Right hind wing. G. Caudal view.	2025-05-27	Constant, Jérôme;Pham, Hong Thai		Zenodo	biologists	Constant, Jérôme;Pham, Hong Thai			
03F6C023FF9C2C68FC619DFCE5C6F834.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15536227/files/figure.png	https://doi.org/10.5281/zenodo.15536227	Fig. 32. Ishiharanus pulchellus sp. nov., dissected holotype, ♂ (VNMN), terminalia. A–D. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Posterolateral view. C. Caudal view. D. Dorsal view. E–M. Aedeagus. E. Dorsal view. F. Ventral view. G. Left lateral view. H. Right lateral view. I. Left lateroventral view.J. Right lateroventral view. K. Left posterolateral view. L. Right posterolateral view. M. Caudal view. Abbreviations: see Material and methods.	Fig. 32. Ishiharanus pulchellus sp. nov., dissected holotype, ♂ (VNMN), terminalia. A–D. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Posterolateral view. C. Caudal view. D. Dorsal view. E–M. Aedeagus. E. Dorsal view. F. Ventral view. G. Left lateral view. H. Right lateral view. I. Left lateroventral view.J. Right lateroventral view. K. Left posterolateral view. L. Right posterolateral view. M. Caudal view. Abbreviations: see Material and methods.	2025-05-27	Constant, Jérôme;Pham, Hong Thai		Zenodo	biologists	Constant, Jérôme;Pham, Hong Thai			
03F6C023FF9C2C68FC619DFCE5C6F834.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15536230/files/figure.png	https://doi.org/10.5281/zenodo.15536230	Fig. 33. Ishiharanus pulchellus sp. nov., paratype, ♂ (RBINS) from Kon Chu Rang National Park, pale morph, terminalia. A–F. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Posterolateral view. C. Caudal view. D. Dorsal view. E. Ventral view. F. Posterodorsal view. G–S. Aedeagus. G. Caudal view. H. Left lateral view. I. Right lateral view. J. Posteroventral view. K. Left laterodorsal view. L. Right laterodorsal view. M. Right posterolateral view. N. Left lateroventral view. O. Right lateroventral view. P. Left posterolateral view. Q. Dorsal view. R. Posterodorsal view. S. Ventral view. Abbreviations: see Material and methods.	Fig. 33. Ishiharanus pulchellus sp. nov., paratype, ♂ (RBINS) from Kon Chu Rang National Park, pale morph, terminalia. A–F. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Posterolateral view. C. Caudal view. D. Dorsal view. E. Ventral view. F. Posterodorsal view. G–S. Aedeagus. G. Caudal view. H. Left lateral view. I. Right lateral view. J. Posteroventral view. K. Left laterodorsal view. L. Right laterodorsal view. M. Right posterolateral view. N. Left lateroventral view. O. Right lateroventral view. P. Left posterolateral view. Q. Dorsal view. R. Posterodorsal view. S. Ventral view. Abbreviations: see Material and methods.	2025-05-27	Constant, Jérôme;Pham, Hong Thai		Zenodo	biologists	Constant, Jérôme;Pham, Hong Thai			
03F6C023FF9A2C76FC2E9957E4EFF886.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15536145/files/figure.png	https://doi.org/10.5281/zenodo.15536145	Fig. 1. Map of the sampled localities in Thừa Thiên-Huế Province, Vietnam.	Fig. 1. Map of the sampled localities in Thừa Thiên-Huế Province, Vietnam.	2025-05-27	Constant, Jérôme;Pham, Hong Thai		Zenodo	biologists	Constant, Jérôme;Pham, Hong Thai			
03F6C023FF9A2C76FC2E9957E4EFF886.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15536147/files/figure.png	https://doi.org/10.5281/zenodo.15536147	Fig. 2. Bach Ma National Park, habitats. A. Detailed map of the sampling areas (1 = ‘Yes Hue Eco’; 2 = ‘pheasant trail’; 3 = ‘low altitude’; 4 = ‘roadside’; 5 = ‘summit’; 6 = ‘near ranger station’. B. Typical habitat at ‘Yes Hue Eco’ sampling area, 17 May 2023.	Fig. 2. Bach Ma National Park, habitats. A. Detailed map of the sampling areas (1 = ‘Yes Hue Eco’; 2 = ‘pheasant trail’; 3 = ‘low altitude’; 4 = ‘roadside’; 5 = ‘summit’; 6 = ‘near ranger station’. B. Typical habitat at ‘Yes Hue Eco’ sampling area, 17 May 2023.	2025-05-27	Constant, Jérôme;Pham, Hong Thai		Zenodo	biologists	Constant, Jérôme;Pham, Hong Thai			
03F6C023FF9A2C76FC2E9957E4EFF886.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15536151/files/figure.png	https://doi.org/10.5281/zenodo.15536151	Fig. 4. Bach Ma National Park, habitats. A. Typical habitat at ‘roadside’ sampling area, 21 May 2023. B. Typical habitat at ‘summit’ sampling area, 11 May 2023.	Fig. 4. Bach Ma National Park, habitats. A. Typical habitat at ‘roadside’ sampling area, 21 May 2023. B. Typical habitat at ‘summit’ sampling area, 11 May 2023.	2025-05-27	Constant, Jérôme;Pham, Hong Thai		Zenodo	biologists	Constant, Jérôme;Pham, Hong Thai			
03F6C023FF9A2C76FC2E9957E4EFF886.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15536207/files/figure.png	https://doi.org/10.5281/zenodo.15536207	Fig. 26. Ishiharanus iguchii (Matsumura, 1916). A–F. Dissected ♂ (RBINS) from Bach Ma. A. Habitus, dorsal view. B. Habitus, ventral view. C. Habitus, lateral view. D. Habitus, perpendicular view of frons. E. Habitus, anterolateral view. F. Right hind wing. G. Dissected ♂ from Cuc Phuong (RBINS), anterolateral view.	Fig. 26. Ishiharanus iguchii (Matsumura, 1916). A–F. Dissected ♂ (RBINS) from Bach Ma. A. Habitus, dorsal view. B. Habitus, ventral view. C. Habitus, lateral view. D. Habitus, perpendicular view of frons. E. Habitus, anterolateral view. F. Right hind wing. G. Dissected ♂ from Cuc Phuong (RBINS), anterolateral view.	2025-05-27	Constant, Jérôme;Pham, Hong Thai		Zenodo	biologists	Constant, Jérôme;Pham, Hong Thai			
03F6C023FF9A2C76FC2E9957E4EFF886.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15536211/files/figure.png	https://doi.org/10.5281/zenodo.15536211	Fig. 27. Ishiharanus iguchii (Matsumura, 1916), ♂ (RBINS) from Cuc Phuong National Park, terminalia. A–F. Pygofer, anal tube and gonostyli.A. Lateral view.B. Posterolateral view.C. Caudal view.D. Dorsal view. E. Ventral view. F. Posterodorsal view. G–S. Aedeagus.G. Left lateral view. H. Right lateral view. I. Caudal view. J. Caudoventral view. K. Right laterodorsal view. L. Left posterolateral view. M. Left laterocaudal view. N. Left lateroventral view. O. Right lateroventral view. P. Right laterocaudal view. Q. Anterodorsal view. R. Dorsal view. S. Ventral view. Abbreviations: see Material and methods.	Fig. 27. Ishiharanus iguchii (Matsumura, 1916), ♂ (RBINS) from Cuc Phuong National Park, terminalia. A–F. Pygofer, anal tube and gonostyli.A. Lateral view.B. Posterolateral view.C. Caudal view.D. Dorsal view. E. Ventral view. F. Posterodorsal view. G–S. Aedeagus.G. Left lateral view. H. Right lateral view. I. Caudal view. J. Caudoventral view. K. Right laterodorsal view. L. Left posterolateral view. M. Left laterocaudal view. N. Left lateroventral view. O. Right lateroventral view. P. Right laterocaudal view. Q. Anterodorsal view. R. Dorsal view. S. Ventral view. Abbreviations: see Material and methods.	2025-05-27	Constant, Jérôme;Pham, Hong Thai		Zenodo	biologists	Constant, Jérôme;Pham, Hong Thai			
03F6C023FF9A2C76FC2E9957E4EFF886.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15536217/files/figure.png	https://doi.org/10.5281/zenodo.15536217	Fig. 28. Ishiharanus iguchii (Matsumura, 1916). A. Live specimen, spotted morph, Bach Ma National Park, roadside near summit, 21 May 2023. B. Live specimen, dark brown morph, Pu Hu Nature Reserve, 5 Jul. 2023. C. Distribution in Vietnam.	Fig. 28. Ishiharanus iguchii (Matsumura, 1916). A. Live specimen, spotted morph, Bach Ma National Park, roadside near summit, 21 May 2023. B. Live specimen, dark brown morph, Pu Hu Nature Reserve, 5 Jul. 2023. C. Distribution in Vietnam.	2025-05-27	Constant, Jérôme;Pham, Hong Thai		Zenodo	biologists	Constant, Jérôme;Pham, Hong Thai			
03F6C023FF802C7FFC5F9957E5F9F82A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15536147/files/figure.png	https://doi.org/10.5281/zenodo.15536147	Fig. 2. Bach Ma National Park, habitats. A. Detailed map of the sampling areas (1 = ‘Yes Hue Eco’; 2 = ‘pheasant trail’; 3 = ‘low altitude’; 4 = ‘roadside’; 5 = ‘summit’; 6 = ‘near ranger station’. B. Typical habitat at ‘Yes Hue Eco’ sampling area, 17 May 2023.	Fig. 2. Bach Ma National Park, habitats. A. Detailed map of the sampling areas (1 = ‘Yes Hue Eco’; 2 = ‘pheasant trail’; 3 = ‘low altitude’; 4 = ‘roadside’; 5 = ‘summit’; 6 = ‘near ranger station’. B. Typical habitat at ‘Yes Hue Eco’ sampling area, 17 May 2023.	2025-05-27	Constant, Jérôme;Pham, Hong Thai		Zenodo	biologists	Constant, Jérôme;Pham, Hong Thai			
03F6C023FF802C7FFC5F9957E5F9F82A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15536149/files/figure.png	https://doi.org/10.5281/zenodo.15536149	Fig. 3. Bach Ma National Park, habitats. A. Typical habitat at ‘pheasant trail’ sampling area, 20 May 2023. B. Typical habitat at ‘low altitude’ sampling area, 15 May 2023.	Fig. 3. Bach Ma National Park, habitats. A. Typical habitat at ‘pheasant trail’ sampling area, 20 May 2023. B. Typical habitat at ‘low altitude’ sampling area, 15 May 2023.	2025-05-27	Constant, Jérôme;Pham, Hong Thai		Zenodo	biologists	Constant, Jérôme;Pham, Hong Thai			
03F6C023FF802C7FFC5F9957E5F9F82A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15536151/files/figure.png	https://doi.org/10.5281/zenodo.15536151	Fig. 4. Bach Ma National Park, habitats. A. Typical habitat at ‘roadside’ sampling area, 21 May 2023. B. Typical habitat at ‘summit’ sampling area, 11 May 2023.	Fig. 4. Bach Ma National Park, habitats. A. Typical habitat at ‘roadside’ sampling area, 21 May 2023. B. Typical habitat at ‘summit’ sampling area, 11 May 2023.	2025-05-27	Constant, Jérôme;Pham, Hong Thai		Zenodo	biologists	Constant, Jérôme;Pham, Hong Thai			
03F6C023FF802C7FFC5F9957E5F9F82A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15536153/files/figure.png	https://doi.org/10.5281/zenodo.15536153	Fig. 5. Bach Ma National Park and Phong Dien District, habitats. A. Typical habitat at ‘ranger station’ sampling area, 18 May 2023. B. Typical habitat at Phong Dien sampling area, 23 May 2023.	Fig. 5. Bach Ma National Park and Phong Dien District, habitats. A. Typical habitat at ‘ranger station’ sampling area, 18 May 2023. B. Typical habitat at Phong Dien sampling area, 23 May 2023.	2025-05-27	Constant, Jérôme;Pham, Hong Thai		Zenodo	biologists	Constant, Jérôme;Pham, Hong Thai			
03F6C023FF802C7FFC5F9957E5F9F82A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15536221/files/figure.png	https://doi.org/10.5281/zenodo.15536221	Fig. 29. Ishiharanus pulchellus sp. nov., dissected holotype, ♂ (VNMN). A. Habitus, dorsal view. B. Habitus, ventral view. C. Habitus, lateral view. D. Habitus, perpendicular view of frons. E. Habitus, anterolateral view. F. Right hind wing. G. Right hind leg, ventral view. H. Metatarsus and apex of metatibia, ventral view.	Fig. 29. Ishiharanus pulchellus sp. nov., dissected holotype, ♂ (VNMN). A. Habitus, dorsal view. B. Habitus, ventral view. C. Habitus, lateral view. D. Habitus, perpendicular view of frons. E. Habitus, anterolateral view. F. Right hind wing. G. Right hind leg, ventral view. H. Metatarsus and apex of metatibia, ventral view.	2025-05-27	Constant, Jérôme;Pham, Hong Thai		Zenodo	biologists	Constant, Jérôme;Pham, Hong Thai			
03F6C023FF802C7FFC5F9957E5F9F82A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15536223/files/figure.png	https://doi.org/10.5281/zenodo.15536223	Fig. 30. Ishiharanus pulchellus sp. nov., dissected paratype, ♂ (RBINS) from Bach Ma National Park, dark morph. A. Habitus, dorsal view. B. Habitus, ventral view. C. Habitus, lateral view. D. Habitus, perpendicular view of frons. E. Habitus, anterolateral view. F. Right hind wing.	Fig. 30. Ishiharanus pulchellus sp. nov., dissected paratype, ♂ (RBINS) from Bach Ma National Park, dark morph. A. Habitus, dorsal view. B. Habitus, ventral view. C. Habitus, lateral view. D. Habitus, perpendicular view of frons. E. Habitus, anterolateral view. F. Right hind wing.	2025-05-27	Constant, Jérôme;Pham, Hong Thai		Zenodo	biologists	Constant, Jérôme;Pham, Hong Thai			
03F6C023FF802C7FFC5F9957E5F9F82A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15536225/files/figure.png	https://doi.org/10.5281/zenodo.15536225	Fig. 31. Ishiharanus pulchellus sp. nov., dissected paratype, ♂ (RBINS) from Kon Chu Rang National Park, pale morph. A. Habitus, dorsal view. B. Habitus, ventral view. C. Habitus, lateral view. D. Habitus, perpendicular view of frons. E. Habitus, anterolateral view. F. Right hind wing. G. Caudal view.	Fig. 31. Ishiharanus pulchellus sp. nov., dissected paratype, ♂ (RBINS) from Kon Chu Rang National Park, pale morph. A. Habitus, dorsal view. B. Habitus, ventral view. C. Habitus, lateral view. D. Habitus, perpendicular view of frons. E. Habitus, anterolateral view. F. Right hind wing. G. Caudal view.	2025-05-27	Constant, Jérôme;Pham, Hong Thai		Zenodo	biologists	Constant, Jérôme;Pham, Hong Thai			
03F6C023FF802C7FFC5F9957E5F9F82A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15536227/files/figure.png	https://doi.org/10.5281/zenodo.15536227	Fig. 32. Ishiharanus pulchellus sp. nov., dissected holotype, ♂ (VNMN), terminalia. A–D. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Posterolateral view. C. Caudal view. D. Dorsal view. E–M. Aedeagus. E. Dorsal view. F. Ventral view. G. Left lateral view. H. Right lateral view. I. Left lateroventral view.J. Right lateroventral view. K. Left posterolateral view. L. Right posterolateral view. M. Caudal view. Abbreviations: see Material and methods.	Fig. 32. Ishiharanus pulchellus sp. nov., dissected holotype, ♂ (VNMN), terminalia. A–D. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Posterolateral view. C. Caudal view. D. Dorsal view. E–M. Aedeagus. E. Dorsal view. F. Ventral view. G. Left lateral view. H. Right lateral view. I. Left lateroventral view.J. Right lateroventral view. K. Left posterolateral view. L. Right posterolateral view. M. Caudal view. Abbreviations: see Material and methods.	2025-05-27	Constant, Jérôme;Pham, Hong Thai		Zenodo	biologists	Constant, Jérôme;Pham, Hong Thai			
03F6C023FF802C7FFC5F9957E5F9F82A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15536230/files/figure.png	https://doi.org/10.5281/zenodo.15536230	Fig. 33. Ishiharanus pulchellus sp. nov., paratype, ♂ (RBINS) from Kon Chu Rang National Park, pale morph, terminalia. A–F. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Posterolateral view. C. Caudal view. D. Dorsal view. E. Ventral view. F. Posterodorsal view. G–S. Aedeagus. G. Caudal view. H. Left lateral view. I. Right lateral view. J. Posteroventral view. K. Left laterodorsal view. L. Right laterodorsal view. M. Right posterolateral view. N. Left lateroventral view. O. Right lateroventral view. P. Left posterolateral view. Q. Dorsal view. R. Posterodorsal view. S. Ventral view. Abbreviations: see Material and methods.	Fig. 33. Ishiharanus pulchellus sp. nov., paratype, ♂ (RBINS) from Kon Chu Rang National Park, pale morph, terminalia. A–F. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Posterolateral view. C. Caudal view. D. Dorsal view. E. Ventral view. F. Posterodorsal view. G–S. Aedeagus. G. Caudal view. H. Left lateral view. I. Right lateral view. J. Posteroventral view. K. Left laterodorsal view. L. Right laterodorsal view. M. Right posterolateral view. N. Left lateroventral view. O. Right lateroventral view. P. Left posterolateral view. Q. Dorsal view. R. Posterodorsal view. S. Ventral view. Abbreviations: see Material and methods.	2025-05-27	Constant, Jérôme;Pham, Hong Thai		Zenodo	biologists	Constant, Jérôme;Pham, Hong Thai			
03F6C023FF802C7FFC5F9957E5F9F82A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15536232/files/figure.png	https://doi.org/10.5281/zenodo.15536232	Fig. 34. Ishiharanus pulchellus sp. nov., live specimens. A–D. In Bach Ma National Park, summit trail. A–B. 19 May 2023. C–D. 20 May 2023. E–H. Phong Dien District, 23 May 2023.	Fig. 34. Ishiharanus pulchellus sp. nov., live specimens. A–D. In Bach Ma National Park, summit trail. A–B. 19 May 2023. C–D. 20 May 2023. E–H. Phong Dien District, 23 May 2023.	2025-05-27	Constant, Jérôme;Pham, Hong Thai		Zenodo	biologists	Constant, Jérôme;Pham, Hong Thai			
03F6C023FF802C7FFC5F9957E5F9F82A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15536236/files/figure.png	https://doi.org/10.5281/zenodo.15536236	Fig. 35. Ishiharanus pulchellus sp. nov., distribution map.	Fig. 35. Ishiharanus pulchellus sp. nov., distribution map.	2025-05-27	Constant, Jérôme;Pham, Hong Thai		Zenodo	biologists	Constant, Jérôme;Pham, Hong Thai			
03F6C023FF802C7FFC5F9957E5F9F82A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15536207/files/figure.png	https://doi.org/10.5281/zenodo.15536207	Fig. 26. Ishiharanus iguchii (Matsumura, 1916). A–F. Dissected ♂ (RBINS) from Bach Ma. A. Habitus, dorsal view. B. Habitus, ventral view. C. Habitus, lateral view. D. Habitus, perpendicular view of frons. E. Habitus, anterolateral view. F. Right hind wing. G. Dissected ♂ from Cuc Phuong (RBINS), anterolateral view.	Fig. 26. Ishiharanus iguchii (Matsumura, 1916). A–F. Dissected ♂ (RBINS) from Bach Ma. A. Habitus, dorsal view. B. Habitus, ventral view. C. Habitus, lateral view. D. Habitus, perpendicular view of frons. E. Habitus, anterolateral view. F. Right hind wing. G. Dissected ♂ from Cuc Phuong (RBINS), anterolateral view.	2025-05-27	Constant, Jérôme;Pham, Hong Thai		Zenodo	biologists	Constant, Jérôme;Pham, Hong Thai			
03F6C023FF802C7FFC5F9957E5F9F82A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15536211/files/figure.png	https://doi.org/10.5281/zenodo.15536211	Fig. 27. Ishiharanus iguchii (Matsumura, 1916), ♂ (RBINS) from Cuc Phuong National Park, terminalia. A–F. Pygofer, anal tube and gonostyli.A. Lateral view.B. Posterolateral view.C. Caudal view.D. Dorsal view. E. Ventral view. F. Posterodorsal view. G–S. Aedeagus.G. Left lateral view. H. Right lateral view. I. Caudal view. J. Caudoventral view. K. Right laterodorsal view. L. Left posterolateral view. M. Left laterocaudal view. N. Left lateroventral view. O. Right lateroventral view. P. Right laterocaudal view. Q. Anterodorsal view. R. Dorsal view. S. Ventral view. Abbreviations: see Material and methods.	Fig. 27. Ishiharanus iguchii (Matsumura, 1916), ♂ (RBINS) from Cuc Phuong National Park, terminalia. A–F. Pygofer, anal tube and gonostyli.A. Lateral view.B. Posterolateral view.C. Caudal view.D. Dorsal view. E. Ventral view. F. Posterodorsal view. G–S. Aedeagus.G. Left lateral view. H. Right lateral view. I. Caudal view. J. Caudoventral view. K. Right laterodorsal view. L. Left posterolateral view. M. Left laterocaudal view. N. Left lateroventral view. O. Right lateroventral view. P. Right laterocaudal view. Q. Anterodorsal view. R. Dorsal view. S. Ventral view. Abbreviations: see Material and methods.	2025-05-27	Constant, Jérôme;Pham, Hong Thai		Zenodo	biologists	Constant, Jérôme;Pham, Hong Thai			
03F6C023FF8B2C02FC6A9DAAE5C6FDD4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15536145/files/figure.png	https://doi.org/10.5281/zenodo.15536145	Fig. 1. Map of the sampled localities in Thừa Thiên-Huế Province, Vietnam.	Fig. 1. Map of the sampled localities in Thừa Thiên-Huế Province, Vietnam.	2025-05-27	Constant, Jérôme;Pham, Hong Thai		Zenodo	biologists	Constant, Jérôme;Pham, Hong Thai			
03F6C023FF8B2C02FC6A9DAAE5C6FDD4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15536147/files/figure.png	https://doi.org/10.5281/zenodo.15536147	Fig. 2. Bach Ma National Park, habitats. A. Detailed map of the sampling areas (1 = ‘Yes Hue Eco’; 2 = ‘pheasant trail’; 3 = ‘low altitude’; 4 = ‘roadside’; 5 = ‘summit’; 6 = ‘near ranger station’. B. Typical habitat at ‘Yes Hue Eco’ sampling area, 17 May 2023.	Fig. 2. Bach Ma National Park, habitats. A. Detailed map of the sampling areas (1 = ‘Yes Hue Eco’; 2 = ‘pheasant trail’; 3 = ‘low altitude’; 4 = ‘roadside’; 5 = ‘summit’; 6 = ‘near ranger station’. B. Typical habitat at ‘Yes Hue Eco’ sampling area, 17 May 2023.	2025-05-27	Constant, Jérôme;Pham, Hong Thai		Zenodo	biologists	Constant, Jérôme;Pham, Hong Thai			
03F6C023FF8B2C02FC6A9DAAE5C6FDD4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15536149/files/figure.png	https://doi.org/10.5281/zenodo.15536149	Fig. 3. Bach Ma National Park, habitats. A. Typical habitat at ‘pheasant trail’ sampling area, 20 May 2023. B. Typical habitat at ‘low altitude’ sampling area, 15 May 2023.	Fig. 3. Bach Ma National Park, habitats. A. Typical habitat at ‘pheasant trail’ sampling area, 20 May 2023. B. Typical habitat at ‘low altitude’ sampling area, 15 May 2023.	2025-05-27	Constant, Jérôme;Pham, Hong Thai		Zenodo	biologists	Constant, Jérôme;Pham, Hong Thai			
03F6C023FF8B2C02FC6A9DAAE5C6FDD4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15536151/files/figure.png	https://doi.org/10.5281/zenodo.15536151	Fig. 4. Bach Ma National Park, habitats. A. Typical habitat at ‘roadside’ sampling area, 21 May 2023. B. Typical habitat at ‘summit’ sampling area, 11 May 2023.	Fig. 4. Bach Ma National Park, habitats. A. Typical habitat at ‘roadside’ sampling area, 21 May 2023. B. Typical habitat at ‘summit’ sampling area, 11 May 2023.	2025-05-27	Constant, Jérôme;Pham, Hong Thai		Zenodo	biologists	Constant, Jérôme;Pham, Hong Thai			
03F6C023FF8B2C02FC6A9DAAE5C6FDD4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15536238/files/figure.png	https://doi.org/10.5281/zenodo.15536238	Fig. 36. Varisphaerius hoaiae sp. nov., dissected paratype, ♂ (RBINS) spotted morph. A. Habitus, dorsal view. B. Habitus, ventral view. C. Habitus, lateral view. D. Habitus, perpendicular view of frons. E. Habitus, anterolateral view. F. Left hind leg, ventral view. G. Metatarsus and apex of metatibia, ventral view. H. Right hind wing.	Fig. 36. Varisphaerius hoaiae sp. nov., dissected paratype, ♂ (RBINS) spotted morph. A. Habitus, dorsal view. B. Habitus, ventral view. C. Habitus, lateral view. D. Habitus, perpendicular view of frons. E. Habitus, anterolateral view. F. Left hind leg, ventral view. G. Metatarsus and apex of metatibia, ventral view. H. Right hind wing.	2025-05-27	Constant, Jérôme;Pham, Hong Thai		Zenodo	biologists	Constant, Jérôme;Pham, Hong Thai			
03F6C023FF8B2C02FC6A9DAAE5C6FDD4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15536240/files/figure.png	https://doi.org/10.5281/zenodo.15536240	Fig. 37. Varisphaerius hoaiae sp. nov., dissected paratype, ♂ (RBINS) reticulated morph. A. Habitus, dorsal view. B. Habitus, ventral view. C. Habitus, lateral view. D. Habitus, perpendicular view of frons. E. Habitus, anterolateral view. F. Right hind wing.	Fig. 37. Varisphaerius hoaiae sp. nov., dissected paratype, ♂ (RBINS) reticulated morph. A. Habitus, dorsal view. B. Habitus, ventral view. C. Habitus, lateral view. D. Habitus, perpendicular view of frons. E. Habitus, anterolateral view. F. Right hind wing.	2025-05-27	Constant, Jérôme;Pham, Hong Thai		Zenodo	biologists	Constant, Jérôme;Pham, Hong Thai			
03F6C023FF8B2C02FC6A9DAAE5C6FDD4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15536242/files/figure.png	https://doi.org/10.5281/zenodo.15536242	Fig. 38. Varisphaerius hoaiae sp. nov., dissected holotype, ♂ (VNMN), terminalia. A–D. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Posterolateral view. C. Caudal view. D. Dorsal view. E–P. Aedeagus. E. Dorsal view. F. Anterolateral view. G. Ventral view. H. Caudal view. I. Left lateral view. J. Right lateral view. K. Left posterolateral view. L. Left laterodorsal view. M. Right laterodorsal view. N. Right posterolateral view.O. Left lateroventral view.P. Right lateroventral view.Abbreviations: see Material and methods.	Fig. 38. Varisphaerius hoaiae sp. nov., dissected holotype, ♂ (VNMN), terminalia. A–D. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Posterolateral view. C. Caudal view. D. Dorsal view. E–P. Aedeagus. E. Dorsal view. F. Anterolateral view. G. Ventral view. H. Caudal view. I. Left lateral view. J. Right lateral view. K. Left posterolateral view. L. Left laterodorsal view. M. Right laterodorsal view. N. Right posterolateral view.O. Left lateroventral view.P. Right lateroventral view.Abbreviations: see Material and methods.	2025-05-27	Constant, Jérôme;Pham, Hong Thai		Zenodo	biologists	Constant, Jérôme;Pham, Hong Thai			
03F6C023FF8B2C02FC6A9DAAE5C6FDD4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15536244/files/figure.png	https://doi.org/10.5281/zenodo.15536244	Fig. 39. Varisphaerius hoaiae sp. nov., live specimens in Bach Ma National Park, pheasant trail. A. 17 Jul. 2011. B. 12 May 2023. C. 21 May 2023. D–F. 15 May 2023. G. 21 May 2023. H. 20 May 2023.	Fig. 39. Varisphaerius hoaiae sp. nov., live specimens in Bach Ma National Park, pheasant trail. A. 17 Jul. 2011. B. 12 May 2023. C. 21 May 2023. D–F. 15 May 2023. G. 21 May 2023. H. 20 May 2023.	2025-05-27	Constant, Jérôme;Pham, Hong Thai		Zenodo	biologists	Constant, Jérôme;Pham, Hong Thai			
