taxonID	type	description	language	source
03F487971E7488684CCDFEE0FECCFB17.taxon	type_taxon	Type species. Punkochyzeria minaevi sp. nov.	en	Kolesnikov, Vasiliy B., Turbanov, Ilya S., Vorontsov, Dmitry D. (2025): Punk’s not dead? Punkochyzeria gen. nov. (Acari: Chyzeriidae) from Cretaceous Myanmar amber. Zootaxa 5686 (4): 451-484, DOI: 10.11646/zootaxa.5686.4.1, URL: https://doi.org/10.11646/zootaxa.5686.4.1
03F487971E7488684CCDFEE0FECCFB17.taxon	diagnosis	Diagnosis. Palps slender. Palptibia with one large claw (odontus), accessory claw absent. Dorsal row of spinose setae (ctenidia) present in some species, ventral row of spinose setae (basidonts) absent. Crista metopica with two sensillary areas located next to each other. Anterior sensillary area (ASA) extended into anterior, triangle shaped process. Posterior sensillary area (PSA) oval in outline, with indistinct or long distinct posterior process. Bothridia of posterior sensilla (PSens) are larger than those of anterior sensilla (ASens). Eyes located laterally to crista, at level of posterior sensilla, anterior lens slightly larger than posterior lens. Idiosoma markedly widened behind aspidosoma / opisthosoma border and surrounded by cuticular rim. Four bundles of long dorsal opisthosomal setae located on four rounded projections; two kinds of setae originate from these projections: 1) densely ciliated or plumose and 2) rigid, spine-like, without cilia and with small irregularities. Moderately long setae (no more than 1 / 2 the length of idiosoma), ciliated or plumose, cover opisthosoma posteriorly and project backwards. Other idiosomal setae moderately short, ciliated or plumose, densely covering idiosoma ventrally and laterally. Dorsal surface of opisthosoma posterior to four bunches of long setae without dense setation.	en	Kolesnikov, Vasiliy B., Turbanov, Ilya S., Vorontsov, Dmitry D. (2025): Punk’s not dead? Punkochyzeria gen. nov. (Acari: Chyzeriidae) from Cretaceous Myanmar amber. Zootaxa 5686 (4): 451-484, DOI: 10.11646/zootaxa.5686.4.1, URL: https://doi.org/10.11646/zootaxa.5686.4.1
03F487971E7488684CCDFEE0FECCFB17.taxon	discussion	Remarks. The new genus, like Parachyzeria, has four tufts of long setae originating from short dorsal opisthosomal projections. It differs from Parachyzeria in the armament of the palptibia: the new genus lacks a developed accessory claw (paradont) or ventral spinose setae (basidonts) (vs. paradont and up to 7 ventral spinose setae (basidonts) present in Parachyzeria). It should be noted that the sensillary area and bothridia were clearly resolved only in one specimen of this study (the holotype of P. khoyi sp. nov.) and less clearly in specimen IM- 1227 a, not attributed to a species.	en	Kolesnikov, Vasiliy B., Turbanov, Ilya S., Vorontsov, Dmitry D. (2025): Punk’s not dead? Punkochyzeria gen. nov. (Acari: Chyzeriidae) from Cretaceous Myanmar amber. Zootaxa 5686 (4): 451-484, DOI: 10.11646/zootaxa.5686.4.1, URL: https://doi.org/10.11646/zootaxa.5686.4.1
03F487971E7488684CCDFEE0FECCFB17.taxon	etymology	Etymology. The prefix punk is derived from the transliteration of the English word “ punk ”, a subculture, one of the characteristic features of which is a hairstyle reminiscent of the arrangement of long dorsal opisthosomal setae in the new genus.	en	Kolesnikov, Vasiliy B., Turbanov, Ilya S., Vorontsov, Dmitry D. (2025): Punk’s not dead? Punkochyzeria gen. nov. (Acari: Chyzeriidae) from Cretaceous Myanmar amber. Zootaxa 5686 (4): 451-484, DOI: 10.11646/zootaxa.5686.4.1, URL: https://doi.org/10.11646/zootaxa.5686.4.1
03F487971E7488604CCDFA88FC5BFEB4.taxon	description	(Figs. 1 – 6, 17 A – C, 18 A – E, Table 2)	en	Kolesnikov, Vasiliy B., Turbanov, Ilya S., Vorontsov, Dmitry D. (2025): Punk’s not dead? Punkochyzeria gen. nov. (Acari: Chyzeriidae) from Cretaceous Myanmar amber. Zootaxa 5686 (4): 451-484, DOI: 10.11646/zootaxa.5686.4.1, URL: https://doi.org/10.11646/zootaxa.5686.4.1
03F487971E7488604CCDFA88FC5BFEB4.taxon	diagnosis	Diagnosis. Long dorsal setae of two kinds: 1) slender, very densely ciliated or plumose, length of cilia at least twice the width of setal shaft (Fig. 17 A – C), density of cilia decreases to base of seta; and 2) slightly shorter, thicker, rigid, spine-like (Fig. 17 A, B). Posterior perimeter of opisthosoma with long ciliated or plumose setae (Figs. 1 E, I, 5 D, 6 C). Other dorsal opisthosomal setae moderately short, ciliated or plumose (Fig. 5 E). Odontus strong, its base in lateral projection equal to or wider than width of palptarsus (Figs. 4 C, D, 18 A – E). Palp tarsus extends beyond termination of odontus, shorter than twice the length of odontus (Figs. 3 E, F, 4 C, D, 18 A – E). Dorsal ctenidium of 1 – 2 spinose setae (Figs. 4 C, D, 18 A – E). Dorsal integument of opisthosoma reticulated (Fig. 1 H). Dorsal surface of opisthosoma without dense setation, with rare setae forming medial longitudinal row, with at least two setae laterad to medial row (Fig. 1 E, F).	en	Kolesnikov, Vasiliy B., Turbanov, Ilya S., Vorontsov, Dmitry D. (2025): Punk’s not dead? Punkochyzeria gen. nov. (Acari: Chyzeriidae) from Cretaceous Myanmar amber. Zootaxa 5686 (4): 451-484, DOI: 10.11646/zootaxa.5686.4.1, URL: https://doi.org/10.11646/zootaxa.5686.4.1
03F487971E7488604CCDFA88FC5BFEB4.taxon	materials_examined	Type material. Holotype PIN- 5608 / 343 a, paratype IM- 1177 a. Occurrence and geological age. Mid-Cretaceous amber, Hukawng Valley, Kachin State, northern Myanmar. Type deposition. PIN 5608 / 343 a — Borissiak Paleontological Institute, Russian Academy of Sciences; IM- 1177 a — private collection of Ilya G. Minaev (Ryazan, Russia).	en	Kolesnikov, Vasiliy B., Turbanov, Ilya S., Vorontsov, Dmitry D. (2025): Punk’s not dead? Punkochyzeria gen. nov. (Acari: Chyzeriidae) from Cretaceous Myanmar amber. Zootaxa 5686 (4): 451-484, DOI: 10.11646/zootaxa.5686.4.1, URL: https://doi.org/10.11646/zootaxa.5686.4.1
03F487971E7488604CCDFA88FC5BFEB4.taxon	description	Description Holotype (PIN 5608 / 343 a, ex. IM- 870). Postlarval stage, complete, well-preserved specimen, visible in dorsal, ventral and lateral aspects (Figs. 1 – 4). Idiosoma (length 1050, width 570) markedly widened behind aspidosoma / opisthosoma border and surrounded by cuticular rim, with four dorsal short rounded projections each bearing a tuft of long setae (Figs. 1 C, E, 3 A). Chelicerae not visible, hidden under hypostome. Hypostome long, narrow (Fig. 3 B, C), with two ventral rows of 8 – 9 setae (setae not longer than twice the width of hypostome), and 1 – 2 lateral setae at apex, all setae setiform, barbed. Palps relatively slender (Fig. 2 B, C). Palp tibia with one strong claw (odontus), accessory claw (paradont) not discerned; dorsal ctenidium represented by one spinose seta (Figs. 4 C, D, 18 A, B). Distally palp with numerous setiform setae (Figs. 4 C, D, 18 A, B), not aligned into rows (combs) dorsally or ventrally. Proximal part of palpfemur bears barbed setae (Fig. 3 D). Palp tarsus oval, elongate, with numerous short setae, extending beyond termination of odontus, shorter than twice the length of odontus (Figs. 3 E, F, 4 C, D, 18 A, B). Aspidosoma clearly delimited, triangular in outline (Fig. 2 B, D). Naso, sensillary area, crista metopica and eyes not discerned, obscured by long dorsal setae located above them. Long dorsal setae originating from four dorsal projections of two types: 1) numerous very long, slender, very densely ciliated or plumose, length of cilia at least twice the width of setal shaft (Fig. 17 A, C), varying in length, longest 940, ratio to idiosomal length 0.84, density of cilia decreases to base of seta; 2) slightly shorter, thicker, rigid, spine-like, without cilia, with small irregularities (Fig. 17 A, C). Plumose setae tend to appear distally near borders of tuft. Other idiosomal setae: 3) numerous long (130 – 200), densely ciliated or plumose, located on posterior part of idiosoma (Fig. 1 E, I) and in anterior part of aspidosoma; 4) numerous moderately long, 2 – 3 times shorter than (3), ciliated, located in lateral and ventral parts of opisthosoma and on aspidosoma, absent on dorsal surface of opisthosoma (Fig. 1 E, F); 5) rare moderately long (about two times shorter than (3), without cilia and with short spines, located in longitudinal rows dorso-medially on opisthosoma (Fig. 1 F, G). Dorsal integument of opisthosoma distinctly reticulated with irregular cells of polygonal shape (Fig. 1 H). Genital opening located behind level of coxae IV, surrounded by paired sclerites (valves) (Fig. 2 B); each valve covered with setae. Anus surrounded with sclerotized valves and situated approx. half way between posterior margin of genital opening and termination of idiosoma (Fig. 2 B). Legs moderately robust. Legs I and IV longer than II and III, and a little longer than idiosoma (Fig. 3 A). Legs I with slightly wider segments than other legs. Leg segments covered with numerous setae of different kinds: ciliated, barbed or smooth (Fig. 3 G – K). Tarsi II – IV long, each terminates with paired claws of similar size and shape (Fig. 3 I – K), tarsus I slightly shorter, with pair of shorter claws (Fig. 3 G, H), empodium not resolved. Paratype (IM- 1177 a). Postlarval stage, complete, well-preserved specimen (Figs. 5, 6), located at border between layers of amber and compressed in lateral direction (Fig. 6 A, B); due to optical distortions, resolution in dorsal and ventral aspects limited. Left part of idiosoma dorsally obscured by bubble (Figs. 5 A, B, 6 A). Specimen mostly visible in lateral aspects (Fig. 5). Idiosomal length 1040, width 470 (but significantly underestimated due to lateral distortion). Types of idiosomal setation similar to holotype specimen, some details of setation better resolved in lateral aspects. Longest dorsal setae 740, ratio to idiosomal length 0.71. Dorsal surface of opisthosoma not resolved, obscured by long setae. Setation best resolved in lateral aspect (Fig. 5 D) Chelicerae not visible, hidden under hypostome. Hypostome long, narrow, with two ventral rows of at least 9 barbed setae, and 1 – 2 lateral setae at apex (setae not longer than double width of hypostome) (Fig. 6 D). Palps relatively slender (Fig. 6 C), details well visible in lateral aspect (Fig. 18 C – E). Left palp viewed in medio-lateral aspect shows a structure that can be interpreted either as a weakly developed paradont or as a single spinose seta of ctenidium (Fig. 18 C, D, arrow). Setae on palp smooth, barbed and ciliated (in proximal part of palp). Palp tarsus oval, elongate, slightly longer than odontus (Fig. 18 C, D). Crista metopica with two sensillary areas located next to each other, PSA with indistinct posterior process (Fig. 5 C). Naso not discerned, obscured by legs. Front legs are raised in antenna-like fashion (Fig. 5 A, B, D). Larva: unknown.	en	Kolesnikov, Vasiliy B., Turbanov, Ilya S., Vorontsov, Dmitry D. (2025): Punk’s not dead? Punkochyzeria gen. nov. (Acari: Chyzeriidae) from Cretaceous Myanmar amber. Zootaxa 5686 (4): 451-484, DOI: 10.11646/zootaxa.5686.4.1, URL: https://doi.org/10.11646/zootaxa.5686.4.1
03F487971E7488604CCDFA88FC5BFEB4.taxon	etymology	Etymology. Punkochyzeria minaevi sp. nov. is named after Ilya G. Minaev (Ryazan, Russia), the owner of the studied specimens. Ilya has provided the amber pieces for our study and has kindly donated three holotype specimens for permanent deposition in the museum collection.	en	Kolesnikov, Vasiliy B., Turbanov, Ilya S., Vorontsov, Dmitry D. (2025): Punk’s not dead? Punkochyzeria gen. nov. (Acari: Chyzeriidae) from Cretaceous Myanmar amber. Zootaxa 5686 (4): 451-484, DOI: 10.11646/zootaxa.5686.4.1, URL: https://doi.org/10.11646/zootaxa.5686.4.1
03F487971E7488604CCDFA88FC5BFEB4.taxon	discussion	Remarks. Although not all characters are resolved in both studied specimens, we consider them conspecific based on a number of similarities: details of long dorsal setae (Fig. 17 A – C), of other idiosomal setae and morphology of palps (Fig. 18 A – E). The reticulations of integument of the dorsal opisthosoma (Fig. 1 H) could be confused with an artifact either remaining after the loss of numerous setae, usually covering the body in Pteridopodinae (Mayoral et al. 2018) or resulting from the incorrect focusing at setal bases or terminations. However, the imaging of the same surface with CLSM clearly shows that most of the dorsal opisthosoma lacks setation, with only a central longitudinal row of setae and a couple of setae lateral to it clearly fluorescent against the darker body cuticle (Fig. 1 E, F). In addition, the extracted fragment of dorsal opisthosomal cuticle of another species of Punkochyzeria gen. nov. (P. khoyi sp. nov.) was imaged using different methods, including SEM (see below), which confirmed the presence of the polygonal structure of the dorsal integument. In both specimens setation was better resolved by CLSM (compare Figs. 2 B and D, 6 B and C). A single strong seta above the base of odontus (Fig. 18 C, D, arrow) could be initially considered a weakly developed paradont. However, the other two species of Punkochyzeria gen. nov. possess similarly shaped spinose setae aligned in a row of three or five (= ctenidium) (Fig. 18 F, I, arrows). Given this, we consider this seta in P. minaevi sp. nov. as representing a ctenidium. Despite that, in P. minaevi sp. nov., the number of spinose dorsal setae near the base of odontus is uncertain (one or two, Fig. 18 A, B), we consider this number as one of the species-specific characters. The lifetime loss of the ctenidium setae is unlikely, as both studied specimens show similar morphology of each of the palps.	en	Kolesnikov, Vasiliy B., Turbanov, Ilya S., Vorontsov, Dmitry D. (2025): Punk’s not dead? Punkochyzeria gen. nov. (Acari: Chyzeriidae) from Cretaceous Myanmar amber. Zootaxa 5686 (4): 451-484, DOI: 10.11646/zootaxa.5686.4.1, URL: https://doi.org/10.11646/zootaxa.5686.4.1
03F487971E7B887B4CCDFD62FED5FD94.taxon	description	(Figs. 7 – 9, 16 D, E, 17 D, E, Table 2)	en	Kolesnikov, Vasiliy B., Turbanov, Ilya S., Vorontsov, Dmitry D. (2025): Punk’s not dead? Punkochyzeria gen. nov. (Acari: Chyzeriidae) from Cretaceous Myanmar amber. Zootaxa 5686 (4): 451-484, DOI: 10.11646/zootaxa.5686.4.1, URL: https://doi.org/10.11646/zootaxa.5686.4.1
03F487971E7B887B4CCDFD62FED5FD94.taxon	diagnosis	Diagnosis. Long dorsal setae of two kinds: 1) ciliated, moderately rigid, length of cilia at least three times the width of setal shaft (Fig. 17 D, E), uniform throughout entire length of seta and 2) thick, rigid, spine-like (Fig. 17 E). Posterior part of opisthosoma with long ciliated setae (Fig. 8 C). Odontus relatively slender (its base in lateral projection smaller than width of palptarsus), on short tibial projection (Figs. 9 B, 18 F, G). Palp tarsus twice longer than odontus (Fig. 18 F, G). Ctenidium of 3 spinose setae (Fig. 18 F, G).	en	Kolesnikov, Vasiliy B., Turbanov, Ilya S., Vorontsov, Dmitry D. (2025): Punk’s not dead? Punkochyzeria gen. nov. (Acari: Chyzeriidae) from Cretaceous Myanmar amber. Zootaxa 5686 (4): 451-484, DOI: 10.11646/zootaxa.5686.4.1, URL: https://doi.org/10.11646/zootaxa.5686.4.1
03F487971E7B887B4CCDFD62FED5FD94.taxon	materials_examined	Type material. Holotype PIN 5608 / 344 a. Occurrence and geological age. Mid-Cretaceous amber, Hukawng Valley, Kachin State, northern Myanmar. Type deposition: Borissiak Paleontological Institute, Russian Academy of Sciences.	en	Kolesnikov, Vasiliy B., Turbanov, Ilya S., Vorontsov, Dmitry D. (2025): Punk’s not dead? Punkochyzeria gen. nov. (Acari: Chyzeriidae) from Cretaceous Myanmar amber. Zootaxa 5686 (4): 451-484, DOI: 10.11646/zootaxa.5686.4.1, URL: https://doi.org/10.11646/zootaxa.5686.4.1
03F487971E7B887B4CCDFD62FED5FD94.taxon	description	Description Holotype, PIN 5608 / 344 a (ex. IM- 851 a). Postlarval stage, incomplete (lacks distal segments of legs on left side) but otherwise well-preserved specimen (Figs. 7 – 9). Amber filled with microscopic debris and bubbles. Idiosoma (1050, 600) markedly widened behind aspidosoma / opisthosoma border and surrounded with cuticular rim, dorsally with four short rounded projections, each holding a tuft of long setae (Fig. 7 B, C). Chelicera not visible, hidden under hypostome. Hypostome long, narrow (Fig. 8 B, D), chaetotaxy not resolved, setae not longer than twice the width of hypostome. Palps relatively slender (Figs. 7 E, 8 B, C). Palp tibia with odontus on short tibial projection, without paradont (Figs. 8 E, 18 F, G). Row of three spinose setae, interpreted as ctenidia, dorsally on palptibia (Fig. 18 F, G, arrows). Palp tarsus oval, elongate, with numerous short setae, ca. twice the length of odontus (Figs. 8 E, 18 F, G). Setae on palp strong with short sparse barbs or some simple setiform setae on palptibia. Aspidosoma clearly delimited, triangular in outline. Naso, sensillary area, crista metopica and eyes not discerned, obscured by long dorsal setae above them. Long dorsal setae originating from four dorsal projections of two types: 1) numerous very long, ciliated, moderately rigid, length of cilia at least three times the width of setal shaft (uniform throughout entire length of seta) (Fig. 17 D, E), varying in length, longest 1100, ratio to idiosomal length 0.9; 2) slightly shorter, thicker, rigid, spine-like, without cilia, with small irregularities (Fig. 17 E). Other idiosomal setae: 3) numerous long (270 – 370), densely ciliated, located in posterior part of opisthosoma (Fig. 8 C) and in anterior part of aspidosoma (Fig. 7 E); 4) numerous moderately long, 2 – 3 times shorter than (3), ciliated, located in lateral and ventral parts of opisthosoma and on aspidosoma (Fig. 8 C), absent on dorsal surface of opisthosoma. Genital opening located behind level of coxae IV, surrounded by paired sclerites (valves); each valve covered with setae (Fig. 8 C). Legs partially preserved, some distal segments of left legs missing (Fig. 8 A). Setae on legs obscured by numerous small bubbles. Legs moderately robust. Legs I and IV longer than II and III, and a little longer than idiosoma. Legs I with slightly wider segments than other legs. Leg segments covered with numerous ciliated setae and rare simple setiform setae (Fig. 8 G). Tarsus I densely covered with short, ciliated setae, significantly obscured by small bubbles. Tarsi II – IV terminated in paired claws of similar size and shape, tarsus I with a pair of shorter claws, empodium not resolved. Larva: unknown.	en	Kolesnikov, Vasiliy B., Turbanov, Ilya S., Vorontsov, Dmitry D. (2025): Punk’s not dead? Punkochyzeria gen. nov. (Acari: Chyzeriidae) from Cretaceous Myanmar amber. Zootaxa 5686 (4): 451-484, DOI: 10.11646/zootaxa.5686.4.1, URL: https://doi.org/10.11646/zootaxa.5686.4.1
03F487971E7B887B4CCDFD62FED5FD94.taxon	etymology	Etymology. Punkochyzeria makolae sp. nov. is named after Joanna Mąkol (Wrocław University of Environmental and Life Sciences, Poland), a renowned acarologist and a specialist in Parasitengona mites.	en	Kolesnikov, Vasiliy B., Turbanov, Ilya S., Vorontsov, Dmitry D. (2025): Punk’s not dead? Punkochyzeria gen. nov. (Acari: Chyzeriidae) from Cretaceous Myanmar amber. Zootaxa 5686 (4): 451-484, DOI: 10.11646/zootaxa.5686.4.1, URL: https://doi.org/10.11646/zootaxa.5686.4.1
03F487971E7B887B4CCDFD62FED5FD94.taxon	discussion	Remarks. The holotype specimen (PIN 5608 / 344 a) is better viewable in dorsal and ventral aspects; it is obscured by optical distortions at the borders between the layers of amber in frontal and lateral aspects (Fig. 8 B, E), as the mite is located at such border. Long dorsal setae overhang the opisthosoma and do not allow to resolve most of its surface neither in reflected light (Fig. 7 C) nor with CLSM (not shown). Palpi are better resolved in ventral aspect, where it was possible to grind the amber closer to the mite; in dorsal and lateral aspects the minimal dimensions of the prepared amber piece were determined by the long projecting setae. Palps are oriented fortunate for observation, as one of them (right) is axially rotated by almost 90 degrees and can thus be viewed in lateral aspect, the other palp (left) is in normal orientation (ventral aspect), with palptarsus bent ventrally (otherwise it would obscure the claw when viewed in transmitted light) (Figs. 8 E, 18 F, G). Long dorsal setae are mostly projected anteriorly, posteriorly and laterally (Figs. 7 B, 8 F). Among the long dorsal setae on the left side of the mite there is a syninclusion (PIN 5608 / 344 f, Figs. 7 B, C, arrows): another mite, which was tentatively determined as Tenuipalpidae (Acari: Tetranychoidea) by Alexander A. Khaustov (Tyumen, Russia). Here we show as much as could be assessed with our optics through the thickness of amber (Fig. 19 A – D), dictated by the length of dorsal setae of Punkochyzeria makolae sp. nov. The mite possesses a number of very long (longer than its body length) slightly barbed dorsal setae (Fig. 19 C, D), entangled among the rich dorsal setation of Punkochyzeria.	en	Kolesnikov, Vasiliy B., Turbanov, Ilya S., Vorontsov, Dmitry D. (2025): Punk’s not dead? Punkochyzeria gen. nov. (Acari: Chyzeriidae) from Cretaceous Myanmar amber. Zootaxa 5686 (4): 451-484, DOI: 10.11646/zootaxa.5686.4.1, URL: https://doi.org/10.11646/zootaxa.5686.4.1
03F487971E6788784CCDFD0DFC9DFCBC.taxon	description	(Figs. 10, 11, 17 J, 18 I, Table 2)	en	Kolesnikov, Vasiliy B., Turbanov, Ilya S., Vorontsov, Dmitry D. (2025): Punk’s not dead? Punkochyzeria gen. nov. (Acari: Chyzeriidae) from Cretaceous Myanmar amber. Zootaxa 5686 (4): 451-484, DOI: 10.11646/zootaxa.5686.4.1, URL: https://doi.org/10.11646/zootaxa.5686.4.1
03F487971E6788784CCDFD0DFC9DFCBC.taxon	diagnosis	Diagnosis. Long dorsal setae of two kinds: 1) ciliated, moderately rigid, with length of cilia at least three times the width of setal shaft (Fig. 17 J), uniform throughout entire length of seta and 2) thick, rigid, spine-like. Posterior part of opisthosoma with long ciliated setae (Figs. 10 A, B). Odontus relatively slender (its base smaller than width of palptarsus), on tibial projection (Figs. 10 I, 18 I). Palp tarsus approx. twice longer than odontus (Figs. 10 I, 18 I). Ctenidium of 5 spinose setae (Fig. 18 I). Dorsal integument of opisthosoma reticulated (Fig. 11).	en	Kolesnikov, Vasiliy B., Turbanov, Ilya S., Vorontsov, Dmitry D. (2025): Punk’s not dead? Punkochyzeria gen. nov. (Acari: Chyzeriidae) from Cretaceous Myanmar amber. Zootaxa 5686 (4): 451-484, DOI: 10.11646/zootaxa.5686.4.1, URL: https://doi.org/10.11646/zootaxa.5686.4.1
03F487971E6788784CCDFD0DFC9DFCBC.taxon	materials_examined	Type material. Holotype PIN 5608 / 345 a. Occurrence and geological age. Mid-Cretaceous amber, Hukawng Valley, Kachin State, northern Myanmar. Type deposition: Borissiak Paleontological Institute, Russian Academy of Sciences.	en	Kolesnikov, Vasiliy B., Turbanov, Ilya S., Vorontsov, Dmitry D. (2025): Punk’s not dead? Punkochyzeria gen. nov. (Acari: Chyzeriidae) from Cretaceous Myanmar amber. Zootaxa 5686 (4): 451-484, DOI: 10.11646/zootaxa.5686.4.1, URL: https://doi.org/10.11646/zootaxa.5686.4.1
03F487971E6788784CCDFD0DFC9DFCBC.taxon	description	Description Holotype, PIN 5608 / 345 a (ex. IM- 1310). Postlarval stage, incomplete (lacks ventral surface of opisthosoma) but otherwise well-preserved specimen, almost fully impregnated with fossil resin and transparent (Fig. 10), with large air bubble obscuring idiosoma ventrally (but opened and cleared during preparation) and a few smaller air bubbles at bases of legs. Idiosoma (length 2150, width 1200) markedly widened behind aspidosoma / opisthosoma border and surrounded with cuticular rim, dorsally with four short rounded projections, each holding a tuft of long setae (Fig. 10 A). All four tufts of long setae pressed to idiosoma dorsum and projected posteriorly. Chelicera not visible, hidden under hypostome. Hypostome long, narrow (Fig. 10 I, J), chaetotaxy not resolved, setae not longer than twice the width of hypostome. Palps relatively slender (Figs. 10 I, J, 18 I). Palp tibia with odontus on long tibial projection, without paradont. Dorsally on palptibia row of five spinose setae (ctenidia) (Fig. 18 I, arrows). Palp tarsus oval, elongate, with numerous short setae, ca. twice the length of odontus. Setae on palp strong with short sparse barbs or some simple setiform setae on palptibia. Aspidosoma clearly delimited, triangular in outline (Fig. 10 D). Crista metopica with two pairs of trichobothria (Fig. 10 G). ASA extended into anterior, triangular process. PSA oval in outline, with indistinct posterior process. Bothridia of posterior sensilla (PSens) larger than anterior sensilla (ASens). Eyes located laterally to crista metopica, at level of posterior sensilla, anterior lens slightly larger than posterior one (Fig. 10 E). Long dorsal setae originating from four dorsal projections of two types: 1) numerous very long, ciliated, moderately rigid, length of cilia at least three times the width of setal shaft (uniform throughout entire length of seta) (Fig. 17 J), varying in length, longest 1900, ratio to idiosomal length 0.83; 2) slightly shorter, thicker, rigid, spine-like, without cilia, with small irregularities. Other idiosomal setae: 3) numerous long (680 – 740), densely ciliated, located in posterior part of opisthosoma (Fig. 10 A, B) and in anterior part of aspidosoma (Fig. 10 D); 4) numerous moderately long, 3 – 7 times shorter than (3), ciliated, located in lateral part of opisthosoma (Fig. 10 F) and on aspidosoma, absent on dorsal surface of opisthosoma. Dorsal integument of opisthosoma distinctly reticulated with irregular cells of polygonal shape and lacks dense setation (Fig. 11). Legs moderately robust. Legs I and IV longer than II and III, and a little longer than idiosoma (Fig. 10 A, B). Legs I with slightly wider segments than other legs. Leg segments covered with numerous setae of different kinds: ciliated, barbed or smooth (Figs. 10 K – M). Tarsi II – IV long, each terminates with paired claws of similar size and shape (Figs. 10 L, M), tarsus I slightly shorter, with pair of shorter claws (Fig. 10 K), empodium not resolved. Larva: unknown.	en	Kolesnikov, Vasiliy B., Turbanov, Ilya S., Vorontsov, Dmitry D. (2025): Punk’s not dead? Punkochyzeria gen. nov. (Acari: Chyzeriidae) from Cretaceous Myanmar amber. Zootaxa 5686 (4): 451-484, DOI: 10.11646/zootaxa.5686.4.1, URL: https://doi.org/10.11646/zootaxa.5686.4.1
03F487971E6788784CCDFD0DFC9DFCBC.taxon	etymology	Etymology. Punkochyzeria khoyi sp. nov. is named after Yuri N. Klinskikh (1964 – 2000), also known as “ Yuri Khoy ”, a Russian musician, singer and a founder of the punk-rock band “ Sektor Gaza ”.	en	Kolesnikov, Vasiliy B., Turbanov, Ilya S., Vorontsov, Dmitry D. (2025): Punk’s not dead? Punkochyzeria gen. nov. (Acari: Chyzeriidae) from Cretaceous Myanmar amber. Zootaxa 5686 (4): 451-484, DOI: 10.11646/zootaxa.5686.4.1, URL: https://doi.org/10.11646/zootaxa.5686.4.1
03F487971E6788784CCDFD0DFC9DFCBC.taxon	discussion	Remarks. The absence of the ventral portion of the opisthosoma, in conjunction with the bubble's interaction with its dorsal aspect from within the mite, engendered a distinctive opportunity to examine the dorsal cuticle, which would have otherwise remained concealed beneath the tufts of protracted dorsal setae. Upon opening the bubble during the preparation of the amber specimen, it was observed that portions of the dorsal cuticular wall of the opisthosoma had exfoliated from the amber. These fragments were meticulously extracted and mounted individually, with the external side exposed for subsequent analysis using SEM and CLSM. Imaging of the extracted cuticular fragments in reflected and transmitted light (Figs. 11 A, B) confirmed the lack of dorsal setation and the presence of polygonal reticulation of the cuticle, similar to that of P. minaevi (Fig. 1 H). Imaging with SEM revealed the presence of longitudinal grooves on the external surface (Figs. 11 C, E), which can be the result of fossilization. In addition, the inner layer of the cuticle contained thick mesh (Figs. 11 I, J). This mesh, however, cannot be responsible for the polygonal reticulation visible in transmitted light, as these structures are of different scale (compare the same fragment imaged in Figs. 11 G – J), but it still can be resolved in transmitted light at higher magnification. Rotating one of the fragments and imaging its internal surface (relative to the mite) revealed arrays of regular pits with probable deeper orifices at their bottoms (Figs. 11 D, F) similar in size to the pattern of the cuticle seen in transmitted light (compare Figs. 11 B, D).	en	Kolesnikov, Vasiliy B., Turbanov, Ilya S., Vorontsov, Dmitry D. (2025): Punk’s not dead? Punkochyzeria gen. nov. (Acari: Chyzeriidae) from Cretaceous Myanmar amber. Zootaxa 5686 (4): 451-484, DOI: 10.11646/zootaxa.5686.4.1, URL: https://doi.org/10.11646/zootaxa.5686.4.1
