taxonID	type	description	language	source
03F487F9731FFFA453D4A53AFC3EF9F4.taxon	description	(Figs 3, 38, 73, 105, 106)	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F9731FFFA453D4A53AFC3EF9F4.taxon	materials_examined	Type material. Lectotype by present designation: ♀, with labels and data: white, black printed “ Kina Balu Borneo ” / white, black printed “ Bought from Staudinger & Bang-Haas, 1930 ” / red, black printed “ Type ” / white, black printed and handwritten in black ink “ Clivina monticola Andr. Type H. E. Andrewes det. ” / white, black printed “ H. E. Andrewes Coll. B. M. 1945 - 97. ” (NHMUK). Paralectotype by present designation: 1 ♂, with labels and data: white, black printed “ Kina Balu Borneo ” / white, black printed and handwritten in black ink “ Kauf 8. 1936 von Staudinger ” / red, handwritten in black ink “ Paratypus monticola Andr. ” / white, black printed and handwritten in black ink “ Clivina monticola Andr. Cotype H. E. Andrewes det. ” / white, handwritten in black ink “ monticola Andr. ” / white, black printed “ F. van Emden Bequest. B. M. 1960 - 129. ” (NHMUK). Remark. The available type material consists of two conspecific specimens, labelled as ‘ Type’, and ‘ Cotype’, and´Paratype´. Andrewes (1931) mentioned six specimens. For reasons of unambiguousness, a lectotype and paralectotype is designated. The species is described by Andrewes (1931). Additional material. 1 ♂, Kina Balu Borneo / Bought from Staudinger & Bang-Haas, 1930 / H. E. Andrewes Coll. B. M. 1945 - 97. (NHMUK); 6 ♂♂, 7 ♀♀, BRUNNEI – Temburong Belalong Field Res. C. Mix. Dipterocarp For. ~ 100 m, 2 - 8.5.95 Heiss (CADW); 1 ♂, BORNEO: SABAH, Bingkor N Keningau, 400 - 500 m, at light; 20. XI. 1996, leg. D. Grimm / Clivina cf. monticola ANDR. det. Balkenohl, 1998 (SMNS); 5 specs, BORNEO, BRUNEI 14. - 15. ii. 2010 Temburong Prov. Kuala Belalong F. stn., at light, BMNH (E) 2010 - 15 (NHMUK). Compendious description. A large sized piceous species with the following measurements: Body length 7.24 – 8.96 mm (ẍ = 8.02 mm *), width 2.03 – 2.51 mm (ẍ = 2.26 mm *), ratio L / W of pronotum 0.89 – 0.93 (ẍ = 0.91 *), ratio L / W of elytra 1.89 – 1.99 (ẍ = 1.93 *); (* n = 10). Head with labrum 7 - setose, slightly trilobed anteriorly. Head somewhat rugose, frons smooth. Median part of clypeus separated from clypeal-wing by distinct notch, clypeal-wings slightly projecting. Eye distinctly convex, projecting laterally, gena small. Clypeo-frontal furrow distinct, rugose. Neck constriction developed as distinct step. Frons with circular impression at middle, with another wider depression anteriorly, facial carina distinct. Pronotum (Fig. 38) square, wider than long, distinctly convex in frontal view, slightly convex in lateral view, with lateral margin nearly parallel, widest at middle, anterior angles rounded off, posterior angles sharp, conspicuously projecting laterally, lateral margin slightly crenate, surface bilaterally with line-like impression, with two groups of punctures anterior to impressions, glossy on disc, with finely scattered punctures, with isodiametric reticulation at base; basal part of lateral margin from posterior angle to flange of base slanted with angle of around 20 °, basal channel and carina of flange narrow. Proepisternum with distinct isodiametric reticulation, laterally with fine transverse wrinkles, medially with fine punctures. Elytron (Fig. 3) subcylindrical, lateral margin crenate in anterior quarter, striae one to four ending free at base, interval three with four setigerous punctures, stria one with small apical setigerous puncture. Intervals moderately convex, intervals seven and eight as well as base and apex isodiametrically reticulated, interval eight carinate in apical quarter, seventh with very short carina at apex, third at base with small tubercle. Lower surface with abdomen with isodiametric reticulation, two apical setigerous punctures widely separated. Hind wing fully developed. Intermediate tibia with nine tubercles with setae above the spur at the lateral side. Male genitalia (Fig. 73). Median lobe distinctly and regularly arcuate at middle, laterally nearly straight, with moderately acuminated and rounded apex. Parameres distinctly arcuate, hyaline in apical third, each with short small pilus at apex. Female gonocoxites and epipleurite (Figs 105, 106). Gonocoxites scapiform, elongated, fragile, less sclerotized, not fused, completely movable; basal gonocoxite 1 widened, with tree long and one minute setae, apical gonocoxite 2 slender, conspicuously elongated, cylindrical in apical half, with two long setae at apex, with three small and two medium sized setae at lateral margin. Epipleurite formed by hyaline velum, furnished with a single seta and a pair of setae. Variation. The lateral margin of the pronotum varies from slightly convex to straight. There is also one specimen with indistinctly concave part at the lateral margin. The sharp tooth at the posterior angle projects laterally in most specimens not as far as the maximum width of the margin. In two cases, it is indistinctly wider than the widest part of the lateral margin. At the intermediate tibia, the number of tubercles above the spur vary from seven to nine, whereas the setae are always present.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F9731FFFA453D4A53AFC3EF9F4.taxon	distribution	Distribution. Known from Sabah and Brunei on the island of Borneo.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F9731EFFA253D4A594FC03FF4A.taxon	description	(Figs 4, 39, 74)	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F9731EFFA253D4A594FC03FF4A.taxon	materials_examined	Type material. Holotype: ♂, with labels and data: white, black printed “ S-VIETNAM Nam Cat Tien Nat. Park 1. – 15. 5. 1994 Pacholatko & Dembicky ” (CADW). Paratype: 1 ♂, same data as holotype (CADW).	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F9731EFFA253D4A594FC03FF4A.taxon	description	Description. Measurements. Body length 6.05 / 6.14 mm, width 1.91 / 1.96 mm, ratio L / W of pronotum 0.84 / 0.86, ratio L / W of elytra 1.74 / 1.76. Colour glossy, piceous. Supraantennal plate indistinctly brightened at margin, pronotum anteriorly to anterior transverse line fuscous. Antennae, mouthparts, intermediate and hind legs fuscous. Head about a quarter narrower than pronotum. Clypeus nearly straight anteriorly, with its lateral teeth slightly projecting; wing of moderate size, slightly less projecting anteriorly than clypeus, distinctly separated from clypeus by notch; supraantennal plate convex laterally; clypeus, wing, and supraantennal plate reflexed margined. Supraantennal plate extended up to mid-eye level, ending at mid-eye level as moderately blunt knob, separated from wing by distinct notch, separated from supraorbital carina by sharp furrow. Clypeus transverse, slightly convex, separated from frons by flat furrow. Frons moderately convex, with central pore at middle, with scattered small punctures. Supraantennal plate separated from clypeus by sharp and from frons by moderately wide furrow continuing posteriorly up to mid-eye level. With blunt distinct supraorbital carina at posterior half of eye. With two supraorbital setigerous punctures situated at mid-eye level and at gena-level. Neck constriction developed by punctures and distinct step. Eye convex, projecting laterally but not hemispherical. Gena distinct, enclosing eye posteriorly like a band. Antenna moderately long, just not reaching basal channel of pronotum, antennomeres four to ten subelongate (L / W around 1.18). Labrum slightly undulate, nearly straight, isodiametrically reticulated, seven-setose. Mandible of moderate size. Mentum with lateral lobe flattened, rectangular, reflexed margined, isodiametrically reticulated, median tooth hollowed out, as high as lobes. Pronotum (Fig. 39) with disk slightly convex in lateral view, in frontal view flattened at median line, moderately convex laterally. Quadrate, distinctly wider than long. Anterior margin nearly straight. Reflexed lateral margin conspicuously crenate, regularly moderately convex, widest at middle; anterior angle rounded-off, slightly retracted; posterior angle rounded, marked by anteriorly pointed tooth. Lateral channel wide, foveolate, reticulated, basal part of lateral margin from posterior angle to flange of base slanted with angle of less than 20 °, convex, flange short but distinct, basal channel moderately developed, basal carina sharp. Median line crenulated, joining anterior transverse line and base; anterior transverse line narrow, complete. Surface with microscopic punctures, with scattered medium sized punctures, with rough transverse wrinkles laterally and basally, base isodiametrically reticulated. Elytron (Fig. 4) with disk slightly flattened in anterior half in lateral view, regularly convex in frontal view. Outline elongated, distinctly less than twice as long as wide, with maximum width slightly behind middle. Humerus distinct, formed by lateral margin which joins stria five laterally. Reflexed lateral margin indistinctly crenulated posterior humerus. Lateral channel moderately wide. Scutellar striole of moderate length, distinct; with distinct setigerous puncture at base of first stria, with tubercle at base of third interval. Striae moderately deep, with indistinct punctures, one to four free at base, five and six joining at humerus, seven shortened behind humerus, striae one and two running up to apex, three and four, and five and six joining apically. Intervals moderately convex, eight carinate apically, seven at apex with very short step-like carina. Third interval with four setigerous punctures adjoining third stria. Surface of intervals smooth and glossy on disk, with distinct isodiametric reticulation at base, at apical fifth, and on interval seven and eight. Hind wing fully developed. Lower surface. Proepisternum with distinct isodiametric reticulation, with sharp transverse wrinkles. Sternites of abdomen with isodiametric reticulation, slightly shingle-like at middle. Abdominal sternum VII with the two anal setigerous punctures widely separated, at middle slightly projecting posteriorly where it is flattened, with narrow marginal furrow at middle. Legs. Protibia with three sharp spines of moderate length and a small spine basally, distinctly sulcate dorsally, movable spur moderately arcuate, surface with longitudinal reticulation. Mesotibia with distinct and moderately long protuberance preapically with acuminate apex and with its seta inserted laterally, with eight setae furnished tubercles above protuberance, the apical four ones sharp, with few fine setae at the inner side. Front leg with tarsomeres slightly widened, first tarsomere elongated, with carina baso-laterally, laterally with acute tubercle furnished with three short setae. Male genitalia (Fig. 74). Median lobe moderately and regularly convex at middle, indistinctly curved at apical lamella, apical lamella with somewhat flattened regularly acuminated lamella. Parameres moderately slender, acuminated and slightly distorted in apical third, apex with three medium sized setae. Female gonocoxites and epipleurite unknown. Variation. Among the two specimens, the conspicuous crenation at the lateral margin of the pronotum varies in intensity. The labrum is more or less straight.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F9731EFFA253D4A594FC03FF4A.taxon	distribution	Distribution. Known from the Nam Cat Tien National Park in Vietnam.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F9731EFFA253D4A594FC03FF4A.taxon	diagnosis	Diagnosis. A medium sized piceous species with square pronotum and interval seven and eight of the elytron reticulated. It differs mainly from all the other species with a square pronotum by the unique extraordinarily crenated lateral margin of the pronotum.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F9731EFFA253D4A594FC03FF4A.taxon	etymology	Etymology. The name refers to the pronotum (in Latin collaris) which lateral margin is conspicuously crenated (in Latin crenatus). The combination is used in the feminine form (crenata).	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97318FFA153D4A20BFAECFA5E.taxon	description	(Figs 5, 40, 75, 107)	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97318FFA153D4A20BFAECFA5E.taxon	materials_examined	Type material. Holotype: ♂, with labels and data: white, black printed “ BORNEO, Sabah, Danum Valley, Raiforest Lodge, 19. - 20. X. 2009. leg. U. + H. J. BREMER ” (SMNS). Paratypes: 1 ♀, 1 spec., same data as holotype (CMBB); 1 spec., “ Borneo – MALAYSIA: Sabah: Mt. Kinabalu Park vic. Serinsim sub-station ” / “ 6 ° 12 ’ N 116 ° 33 ’ E L. F. 180 - 250 m 30. VIII. 1998 D. Bartsch & C. Häuser leg. “ / „ Clivina helferi group det Balkenohl 2000 ” (SMNS); 1 ♂, “ BORNEO, SABAH, NW., Kota Marundu, Taritipan env., 13. v. 1999, M. Sniźek leg. ” / “ helferi Gr. ” (CADW); 1 spec., “ Malaysia, Sabah, Batu Punggul Resort env. 24. VI. - 1. VII. 1996, 11 f, intercept trap ” (NHMW); 1 ♂, “ Sarawak, Kapit dist. Rumah Ugap vill., Sut riv., 3. - 9. 3. 1994 J. Horák leg. ” (CADW); 1 spec., “ BORNEO, Brunei 2010 Temburong Prov. 14. - 15. ii. Kuala Belalong F. stn., at light, BMNH (E) 2010 - 15 ” (NHMUK); 1 ♀, 3 specs, “ SARAWAK (Borneo), ca 25 km E KAPIT, III. 1994, Kodada leg. ” (NHMW); 1 spec., same data but “ ca 40 km SE KAPIT ” (NHMW); 1 spec., “ MALAYSIA – Sabah prov. Banjaran Crocker Mts. GUNUNG ALAB peak 30. IV. - 27. V. 1996, 1650 - 1800 m M. Štrba & R. Hergovitis leg. ” / “ VIII. helferi Gr. det. Dr. A. Dostal 2000 ”. This specimen was received with an empty abdomen. (CADW); 1 ♂, “ BORNEO: SABAH Bingkor N Keningau 400 - 500 m, 19. - 20. XI. 1996 leg. W. SCHAWALLER ” / “ Clivina group helferi, nr. wallacei PUTZ. det. Balkenohl, 1998 ” (SMNS).	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97318FFA153D4A20BFAECFA5E.taxon	description	Description. Measurements. Body length 6.31 – 7.19 mm (ẍ = 6.59 mm *), width 1.91 – 2.28 mm (ẍ = 2.05 mm *), ratio L / W of pronotum 0.82 – 0.85 (ẍ = 0.84 *), ratio L / W of elytra 1.77 – 1.88 (ẍ = 1.81 *); (* n = 10). Colour glossy, fuscous. Supraantennal plate with margin and clypeal-wing translucent. Antennae, mouthparts, intermediate and hind legs yellowish. Elytron with tip of apex brightened. Head a quarter narrower than pronotum. Clypeus straight anteriorly, with its lateral teeth slightly projecting; wing of moderate size, not more protruding anteriorly than clypeus, distinctly separated from clypeus by notch; supraantennal plate convex laterally, with straight part in anterior half; clypeus, wing, and supraantennal plate finely reflexed margined. Supraantennal plate extended up to mid-eye level, separated from wing by distinct notch, separated from supraorbital carina by a step. Clypeus transverse, with transverse flat V-like elevation, separated from frons by flat furrow. Frons moderately convex, with central pore at middle, with scattered small punctures. Supraantennal plate separated from clypeus and frons by moderately wide furrow continuing posteriorly up to mid-eye level as moderately wide supraorbital furrow. With blunt distinct supraorbital carina at posterior half of eye. Posterior keel of supraorbital plate and supraorbital carina separated by distinct step. With two supraorbital setigerous punctures situated at mid-eye level and at the posterior end of eye. Neck constriction developed by punctures and distinct step, step pointed posteriorly at middle. Eye distinctly hemispherical projecting laterally. Gena small. Antenna moderately long, not reaching base of pronotum, antennomeres four to ten elongate (L / W around 1.55). Labrum nearly straight, isodiametrically reticulated, seven-setose. Mandible of moderate size. Mentum with lateral lobe rectangular anteriorly, flat, isodiametrically reticulated, median tooth as high as lateral lobes, hollowed out, slightly rounded apically. Pronotum (Fig. 40) with disk somewhat flattened but slightly convex in lateral view, in frontal view flattened at middle and moderately convex laterally. Quadrate, wider than long. Anterior margin slightly excised. Reflexed lateral margin smooth, slightly convex at middle, attenuating in anterior half, widest in posterior third; anterior angle distinct but rounded; posterior angle marked by laterally distinctly projecting tooth. Lateral channel wide (lateral view), reticulated, basal part of lateral margin from posterior angle to flange of base slanted with angle of slightly more than 20 °, indistinctly convex, flange short but distinct, basal channel moderately developed, basal carina sharp. Median line moderately narrow, joining anterior transverse line and base; anterior transverse line narrow, complete. Surface with microscopic punctures, bilaterally with three group of punctures in basal half, arrangement of puncture-groups resembles to an indistinct widened letter Y, base isodiametrically reticulated. Elytron (Fig. 5) with disk flattened in anterior half in lateral view, regularly convex in frontal view. Outline long oval, distinctly less than twice as long as wide, with maximum width behind middle. Humerus rounded, formed by lateral margin which joins stria five. Reflexed lateral margin subcrenulated posterior humerus. Lateral channel moderately wide. Scutellar striole of moderate length, not deep; with distinct setigerous puncture at base of first stria, with small tubercle at base of third interval. Striae moderately deep, with indistinct punctures, one to four free at base, five and six joining at humerus, seven shortened behind humerus, striae one and two running up to apex, three and four, and five and six just not joining apically. Intervals moderately convex, eight carinate apically. Third interval with four setigerous punctures adjoining third stria. Surface of intervals smooth and glossy on disk, with distinct isodiametric reticulation at base, at apical tip, and on interval eight, with indistinct reticulation at interval seven. Hind wing fully developed. Lower surface with proepisternum completely covered with distinct isodiametric reticulation, with some transverse wrinkles. Sternites of abdomen isodiametrically reticulated, transverse at middle, the three middle segments with flat impression laterally. Abdominal sternum VII with the two anal setigerous punctures widely separated, at middle slightly projecting posteriorly where it is flattened, with narrow marginal furrow at middle (in both sexes). Legs. Protibia with three spines of moderate length and a small but distinct one basally, sulcate dorsally, movable spur nearly straight, surface with nearly isodiametric reticulation. Mesotibia with distinct and moderately long protuberance preapically with acuminate apex and with its seta inserted laterally, with nine setae furnished tubercles above protuberance, with few fine setae at the inner side. Front leg with tarsomeres slightly widened, first tarsomere elongated, slender and with carina basally, laterally with three setae furnished tubercles. Male genitalia (Fig. 75). Median lobe with nearly straight part at middle, curved basally and slightly at root of apical lamella, apex with flattened duck bill-like elongated lamella. Parameres moderately wide, acuminated and slightly distorted in apical third, apex asetose. Female gonocoxites and epipleurite (Fig. 107). Gonocoxites scapiform, extraordinary slender, less sclerotized, not fused, completely movable; basal gonocoxite 1 elongated, with three setae, apical gonocoxite 2 slender, distinctly elongated, indistinctly curved towards apex, cylindrical in apical half, with two long setae at apex, with two moderately long setae at lateral margin at middle. Epipleurite with two setae distant from each other. Variation. The reticulation on interval seven is more or less expressed. Specimens from Sarawak exhibit a strip of reticulation, only. The flat clypeal-furrow on the head is slightly rugose in some specimens, and the clypeus is more or less elevated. The specimen from Bingkor is rather small.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97318FFA153D4A20BFAECFA5E.taxon	distribution	Distribution. Known from the island of Borneo: Sabah, Brunei, Sarawak.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97318FFA153D4A20BFAECFA5E.taxon	diagnosis	Diagnosis. A larger sized dark fuscous species with square pronotum and interval seven and eight of the elytron reticulated. It differs mainly from the most similar species C. quadrataconvexa Balkenohl by the anteriorly distinctly attenuated pronotum with laterally projecting sharp teeth of the posterior angle, the wider basal channel, and with intervals seven and eight of the elytron reticulated. It differs from C. tuberculifer sp. nov. mainly by the elytron with small tubercle at the base of interval three, the more intense reticulation on intervals seven and eight, the smaller eyes, and the smaller size.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97318FFA153D4A20BFAECFA5E.taxon	etymology	Etymology. The name refers to the square pronotum which is in frontal view laterally distinctly convex, and is expressed by the Latin adjective convexus (= arched) and quadratus (= square) in the feminine genus.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F9731BFFA053D4A53FFAE5F842.taxon	description	(Figs 6, 41, 76)	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F9731BFFA053D4A53FFAE5F842.taxon	materials_examined	Type material. Holotype: ♂, with labels and data: white, black printed “ 1 Jan 16 1 Wao, Lanao del Sur, Mindanao JANUARY 2004 ” / backside “ resid. col. leg. ” / yellow, handwritten in blue ink “ Coll. Balkenohl “ (CMBB). Remark. The left front leg is missing in the holotype.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F9731BFFA053D4A53FFAE5F842.taxon	description	Description. Measurements. Body length 6.21 mm, width 1.74 mm, ratio L / W of pronotum 0.83, ratio L / W of elytra 1.93. Colour glossy, fuscous. Supraantennal plate slightly brightened. Antennae, mouthparts, intermediate and hind legs yellowish, trochanters yellow. Head a quarter narrower than pronotum. Clypeus straight anteriorly, with its lateral teeth somewhat projecting; wing of moderate size, not more protruding anteriorly than clypeus, separated from clypeus by notch; supraantennal plate convex laterally; clypeus, wing, and supraantennal plate finely reflexed margined. Supraantennal plate extended nearly up to mid-eye level, separated from wing by distinct obtuse notch. Clypeus transverse, separated from frons by flat furrow, the two clypeal setae with wide pore. Frons moderately convex, with central pore at middle, clypeus and frons with scattered small punctures. Supraantennal plate separated from clypeus and frons by moderately wide reticulated furrow, continuing posteriorly up to mid-eye level as moderately wide supraorbital furrow. With blunt distinct supraorbital carina at posterior half of eye. Posterior keel of supraorbital plate and supraorbital carina separated by distinct narrow furrow. With two supraorbital setigerous punctures situated at mid-eye level and at end of gena-level. Neck constriction developed by distinct step and band of punctures. Eye distinctly hemispherical projecting laterally. Gena small, with convex ridge posteriorly. Antenna moderately long, just reaching posterior angles of pronotum, antennomeres four to ten sub-elongate (L / W around 1.33). Labrum straight, isodiametrically reticulated, seven-setose. Mandible of moderate size. Mentum with lateral lobe distinctly convex laterally, concave internally, tip flannel-like, surface with fine askew wrinkles, cleft small, prolonged posteriorly as blunt carina, median tooth higher than lateral lobes, rounded and hollowed out at apex. Pronotum (Fig. 41) with disk slightly convex in lateral view, in frontal view slightly flattened at middle and moderately convex laterally. Trapezoidal, wider than long. Anterior margin slightly excised. Reflexed lateral margin smooth, nearly straight in anterior half, convex in posterior half, distinctly attenuating in anterior half, widest at posterior third; anterior angle rounded-off; posterior angle marked by rounded tooth. Lateral channel moderately narrow, reticulated, basal part of lateral margin from posterior angle to flange of base slanted with angle of around 20 °, convex, flange short but distinct, basal channel moderately developed, basal carina sharp. Median line moderately narrow, joining anterior transverse line and base; anterior transverse line narrow, complete. Surface with scattered microscopic punctures, bilaterally with short, slight line-like impression, base isodiametrically reticulated. Elytron (Fig. 6) with disk flattened in anterior half in lateral view, regularly convex in frontal view. Outline long oval, nearly two times longer than wide, maximum width slightly behind middle. Humerus rounded, formed by lateral margin which joins interval five. Reflexed lateral margin subcrenulated posterior humerus. Lateral channel moderately wide. Scutellar striole of moderate length, not deep; with longitudinal small tubercle at base, with distinct setigerous puncture at base of first stria, with distinct tubercle at base of third interval. Striae moderately deep, nearly impunctate, one to four free at base, five and six joining at humerus, seven shortened behind humerus, striae one and two running up to apex, three and four, and five and six joining apically. Intervals moderately convex, eight with relatively long narrow carina apically, seventh with step-like ridge at apex. Third interval with four setigerous punctures adjoining third stria. Surface of intervals smooth and glossy on disk, with distinct isodiametric reticulation at base, apical third, and on interval six to eight. Hind wing fully developed. Lower surface with proepisternum covered laterally with dense small punctures, punctures in median quarter less dense and larger, with some fine transverse wrinkles. Sternites of abdomen isodiametrically reticulated. Abdominal sternum VII with the two anal setigerous punctures widely separated, at middle slightly projecting posteriorly where it is indistinctly hollowed-out, with narrow indistinct marginal furrow at middle. Legs. Protibia with three spines of moderate length and a small but distinct one basally, the apical one somewhat widened. Tibia finely sulcate dorsally, movable spur sharp, moderately curved, surface with longitudinal reticulation. Mesotibia with distinct and moderately long protuberance preapically with acuminate apex and with its seta inserted laterally, with ten setae furnished tubercles above protuberance, with few fine setae at the inner side. Front leg with tarsomeres slightly widened, first tarsomere elongated, laterally with two setae furnished tubercles. Male genitalia (Fig. 76). Median lobe distinctly and regularly curved, apical lamella stretched out into asymmetric flattened slightly distorted spatula. Parameres moderately wide, acuminated and slightly distorted in apical third, apophyses small, apex asetose. Female gonocoxites and epipleurite unknown. Variation. The carina of interval eight at apex is interrupted at the left side. The left elytron is slightly longer than the right one.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F9731BFFA053D4A53FFAE5F842.taxon	distribution	Distribution. Known from the type locality Wao on the island of Mindanao, Philippines.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F9731BFFA053D4A53FFAE5F842.taxon	diagnosis	Diagnosis. A medium sized slender fuscous species with trapezoidal pronotum and intervals six to eight of the elytron reticulated. It differs mainly from the most similar species C. convexaquadrata sp. nov. by the more slender habitus, the trapezoid shape of the pronotum, and by the reticulation on intervals six to eight of the elytron. In addition, it is distinguished by its long and narrow carina in the apical third of interval eight.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F9731BFFA053D4A53FFAE5F842.taxon	etymology	Etymology. The name is derived from the trapeziform shape of the pronotum, and is expressed by the Greek adjective trapӗziŏn (= trapezoid) and the Latin substantive collum (pronotum) in the feminine genus.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97305FFBE53D4A3BAFD10FBAA.taxon	description	(Figs 7, 42, 77, 108)	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97305FFBE53D4A3BAFD10FBAA.taxon	materials_examined	Type material. Holotype: ♂, with labels and data: white, handwritten in black ink and black framed, " N-India, UP Rishikesh, 450 m leg. Werner 8 / 88 ” (CMBB). Paratypes: 1 ♂, 2 ♀♀, same data as holotype (CMBB, OLML); 1 ♀, same data as holotype but with printed label; 1 ♂, white, black printed “ KISTNA DT. Uppaluru V- 51 Biffi ” (CMBB). The species is described by Balkenohl (2023). New records. China: 1 ♂, CHINA: B. M. 1980 - 491 P. M. Hammond / PEKING: N. suburbs 30. viii. 80 / manure and straw heap. (NHMUK); 1 ♂, 1 ♀, 4 specs., CHINA (Yunnan) Baoshan Pref., Longchuan river, 24 km SE Tengchong 24 ° 53 ’ 32 ” N / 98 ° 40 ’ 34 ” E 1210 m (sandy river bank at mouth of a brook. Under stones) 26. VIII. 2009 D. W. Wrase (07) / sp. 2 / CLIVINA cfr. westwoodi Ptz. P. Bulirsch det. 2010 / COLL. WRASE BERLIN / COLL. WRASE GUSOW-PLATKOW GERM. (CDWGP); Vietnam: 1 ♀, Tonkin Tujen-Quang. July 1924. H. Stevens. / Sladen-Godman Trust Exped. B. M. 1924 - 460. (NHMUK); 1 ♂, Tonkin. Le Kep. 1.11.1915. R. V. de Salvaza. / Brit. Mus. 1921 - 89 / Clivina sp. (NHMUK); 1 ♀, VIETNAM N Hanoi VI- 1991 Strnad Jan leg. (CADW); 1 spec., Indo China. Annam Kathai 18.10.1915 R. V. de Salvaza / Brit. Mus. 1921 - 89 / Clivina sp. / CLIVINA helferi group P. Bulirsch det. 2011 (NHMUK); Thailand: 2 ♂♂, 1 ♀, 4 specs. THAILAND Nan Province Lianan cabbage farm, 18.576 N 100.759 E; LN ” Light Trap. 11 - 12. x. 2012, leg. D. L. J. Quicke BMNH (E) 2015 - 75 (NHMUK); 1 ♂, THAILAND PROV. NAN, BO KHUA 19. 4. - 4. 7. 2004 MORAVEC PETR (CMBB); 4 ♀♀, 2 specs., N. W. THAILAND Mae Hong Son Prov. Nam Tok Mae Surin NP N 19 21.494 E 97 59.018 plastic FIT with light river beech 12. vii. 2006 / BMNH (E) 2006.128 H. Mendel & MVL Barkley / one of them with add. label CLIVINA (s. str.) helferi group P. Bulirsch det. 2015 (NHMUK); 4 specs., same data but NP N 19 21.31 E 97 59.02, one with barcode NHMUK 010369304 (NHMUK); 2 ♂♂, Thailand, Chiang Dao Hill Resort, 30. V. - 2. VI. 2017, K. Takahashi (CMBB); 2 ♂♂, 1 ♀, 2 specs, THAILAND Mae Ping, Licht, 8. - 6.9.1991 leg. Malicky (NHMW); Laos: 1 ♀, Laos. Nape. Oct. 1915. R. V. de Salvaza / Clivina sp. / Brit. Mus. 1921 - 89 (NHMUK); Cambodia: 1 ♂; CAMBODIA Siem Reap 25. V. 2005 Light trap Leg. J. Constant & K. Smets, Coll. I. R. Sc. N. B. (RBINS); Malaysia: 1 spec., MALAYA Kuala Lumpur. / Sel: Museum Nr: L. Gardens. Coll. H. M. Pendleburry 28. I. 1936 / Ex F. M. S. Museum. B. M. 1955 - 354. / CLIVINA (s. str.) helferi group P. Bulirsch det. 2015 (NHMUK); Nepal: 1 ♀, NEPAL, P: Narayani Sauhara, Rapti Ufer, LF, 150 m NN, 16. - 18. VI. 2007, leg. M. Hartmann 27 ° 34 ’ 29 ” N, 84 ° 29 ’ 55 ” E / CLIVINA helferi gr. Ptz. P. Bulirsch det. 2008 (NKME); 1 spec., NEPAL (Chitwan) Gunganagar, 5 km W Rampur (light trap) IV 2004 Y. Gc leg. IAAS / SSMP-WG / Clivina helferi group det. Balkenohl / COLL. WRASE GUSOW-PLATKOW GERM. (CDWGP); 1 spec., NEPAL (Nawalparasi) Gaindakot, 3 km N Bharatpur (light trap) VI 2004 Y. Gc leg. IAAS / SSMP-WG / CLIVINA helferi group P. Bulirsch det. 2006 / COLL. WRASE GUSOW-PLATKOW GERM. (CDWGP); Myanmar: 1 ♂, MYANMAR: Kachin State Putao, at light ca. 550 m, 28.5. - 3.6.1999 leg. Schillhammer & Schuh (56) (NHMW); 1 ♂, MYANMAR: Kachin State Myitkyina, Sumpra Hotel 26. - 27.5.1999 at light, leg. Schuh & Schillhammer (55) (NHMW); India: 1 ♂, INDIA bor. UTTAR PRADESH bor. KOTDWARA env. 1. - 3. 8. 1994 lgt. M. SNIŽEK / VIII helferi (CADW); 1 ♂, INDIA bor. UTTAR PRADESH HARIDWAR, CHILA, 330 m, 09. - 14.08.1994 lgt. M. SNIŽEK (CADW); 1 ♂, same data but 300 m, (CADW); 1 ♂, 1 ♀, NE INDIA, ARUNACHAL PR. 8 km S Jamiri SESSA vicinity 27 ° 07 ’ - 09 ’ N 92 ° 34 ’ E, 350 m L. Dembický leg., 26. v. - 4. vi. 2005 / CLIVINA westwoodi Putzeys, 1866 P. Bulirsch det. 2012. Compendious description. A larger sized piceous species with the following measurements: Body length 6.07 – 6.94 mm (ẍ = 6.54 mm *), width 2.03 – 2.61 mm (ẍ = 2.27 mm *), ratio L / W of pronotum 0.89 – 0.95 (ẍ = 0.92 *), ratio L / W of elytra 1.89 – 2.02 (ẍ = 1.95 *); (* n = 10). Head with labrum seven-setose, nearly straight. Median part of clypeus separated from clypeal-wing by distinct notch, clypeal-wings less projecting as lateral teeth of clypeus. Eye semicircular protruding laterally, gena indistinct. Clypeo-frontal furrow distinct, rugose. Neck constriction developed as distinct step. Frons with central impression at middle, with scattered small punctures, facial carina distinct. Pronotum (Fig. 42) quadrate, wider than long, distinctly convex in frontal view, moderately convex in lateral view, with lateral margin slightly convex at middle, widest at middle, anterior angles rounded off, posterior angles marked by distinct tooth, lateral margin subcrenulated (in lateral view), surface bilaterally with line-like punctures laterally in basal half, with two clouds of punctures anterior to the line-like punctures, glossy on disc, with finely scattered punctures; basal part of lateral margin from posterior angle to flange of base slanted with angle of less than 20 °, basal channel and carina of flange narrow. Proepisternum with distinct isodiametric reticulation, laterally with fine transverse wrinkles. Elytron (Fig. 7) subcylindrical in frontal view, lateral margin smooth, striae one to four ending free at base, interval three with four setigerous punctures; Intervals moderately convex, intervals six, seven and eight as well as base and quarter of apex isodiametrically reticulated, interval eight carinate in apical quarter, seventh with very short carina at apex, third at base with small but distinct tubercle. Lower surface with abdomen with isodiametric reticulation, two apical setigerous punctures widely separated. Hind wing fully developed. Intermediate tibia with eight tubercles with setae above the spur at the lateral side. Male genitalia (Fig. 77). Median lobe regularly curved, with straight apex, apex with flattened and regularly acuminated lamella. Endophallus with group of bristles. Parameres moderately wide, acuminated and distorted in apical third, at apex with one short seta. Female gonocoxites and epipleurite (Fig. 108). Gonocoxites scapiform, elongated, moderately sclerotized, not fused, completely movable; basal gonocoxite 1 elongated, with two long setae, apical gonocoxite 2 slender, distinctly elongated, cylindrical in apical half, with two long setae at apex, with one long seta at middle, with two small setae at lateral margin in basal half. Epipleurite with two groups of three setae, each.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97305FFBE53D4A3BAFD10FBAA.taxon	description	Variation. The clypeal-furrow on the frons varies slightly in its rugosity. In few specimens, there are slight impressions on the frons laterally and posteriorly of the clypeal-furrow. The supraantennal plates vary slightly in its lateral convexity. In one specimen from Thailand (Prov. Nan, Bo Khua) the tubercle at base of the third interval is missing.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97305FFBE53D4A3BAFD10FBAA.taxon	distribution	Distribution. The species seems to be widely distributed from Nepal, northern India, and south-western China, to Vietnam, Thailand, Laos, Cambodia, and Malaysia.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97304FFBD53D4A7ABFB04F92E.taxon	description	(Figs 8, 43, 78, 109)	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97304FFBD53D4A7ABFB04F92E.taxon	materials_examined	Type material. Holotype: ♂, with labels and data: white, black printed “ NEPAL East, Kosi Zone, Dhankutta distr., Arun valley, Dharan env., 500 m, 22 - 23. v. 1996, P. Čechovský lgt. ” / “ Clivina sp. ad. wallacei DET. DR. A. DOSTAL 2011 ” (CADW). Paratype: 1 ♀, same data as holotype (CADW).	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97304FFBD53D4A7ABFB04F92E.taxon	description	Description. Measurements. Body length 4.85 / 4.96 mm, width 1.45 / 1.53 mm, ratio L / W of pronotum 0.90 / 0.91, ratio L / W of elytra 1.78 / 1.79. Colour glossy, fuscous. Supraantennal plate slightly brightened. Antennae, mouthparts, intermediate and hind legs yellowish. Head less than a quarter narrower than pronotum. Clypeus straight anteriorly, with its lateral teeth slightly projecting; wing of moderate size, not more protruding anteriorly than clypeus, distinctly separated from clypeus by notch; supraantennal plate convex laterally, slightly convex in anterior half; clypeus, wing, and supraantennal plate finely reflexed margined. Supraantennal plate extended up to mid-eye level, ending at mid-eye level as blunt knob, separated from wing by distinct notch, separated from supraorbital carina by deep moderately wide furrow. Clypeus transverse, slightly convex, separated from frons by flat furrow. Frons moderately convex, with central longitudinal pore at middle, with scattered small punctures. Supraantennal plate separated from clypeus and frons by moderately wide furrow continuing posteriorly up to mid-eye level as moderately wide supraorbital furrow. With blunt distinct supraorbital carina at posterior half of eye. With two supraorbital setigerous punctures situated at mid-eye level and posterior gena-level. Neck constriction developed by punctures and distinct step. Eye convex, projecting laterally but not hemispherical. Gena distinct, enclosing eye posteriorly. Antenna moderately long, not reaching basal carina of pronotum, antennomeres four to ten moniliform (L / W around 1.06). Labrum nearly straight, isodiametrically reticulated, seven-setose. Mandible of moderate size. Mentum with lateral lobe pentagonal with rounded angles, with irregular and oblique wrinkles, median tooth spatulate, slightly higher as lateral lobes. Pronotum (43) with disk somewhat flattened but indistinctly convex in lateral view, in frontal view flattened at middle and moderately convex laterally. Quadrate, wider than long. Anterior margin nearly straight. Reflexed lateral margin indistinctly sub-crenulate, regularly moderately convex, widest at middle; anterior angle rounded-off; posterior angle rounded, marked by obtuse tooth. Width of lateral channel medium sized, reticulated, basal part of lateral margin from posterior angle to flange of base slanted with angle of around 20 °, convex, flange short but distinct, basal channel moderately developed, basal carina fine but sharp. Median line narrow, crenulated, joining anterior transverse line and base; anterior transverse line narrow, complete. Surface with microscopic punctures, with scattered medium sized punctures, base isodiametrically reticulated. Elytron (Fig. 8) with disk flattened in anterior half in lateral view, regularly convex in frontal view. Outline regularly long oval, distinctly less than twice as long as wide, with maximum width about at middle. Humerus rounded, formed by lateral margin which joins stria five laterally by a short carina. Reflexed lateral margin nearly smooth posterior humerus. Lateral channel moderately wide. Scutellar striole of moderate length, not deep; with distinct setigerous puncture at base of first stria, with conspicuous tubercle at base of third interval. Striae moderately deep, punctate-striate, one to four free at base, five and six joining at humerus, seven shortened behind humerus, striae one and two running up to apex, three and four, and five and six just not joining apically. Intervals slightly convex, seven and eight carinate apically. Third interval with four setigerous punctures adjoining third stria. Surface of intervals smooth and glossy on disk, with distinct isodiametric reticulation at base, at apical tip, and on interval seven and eight. Hind wing fully developed. Lower surface. Proepisternum with distinct isodiametric reticulation, with some fine transverse wrinkles laterally. Sternites of abdomen with shingle-like reticulation, transverse at middle. Abdominal sternum VII with the two anal setigerous punctures widely separated, at middle slightly projecting posteriorly where it is flattened, with narrow marginal furrow at middle (in both sexes). Legs. Protibia with three spines of moderate length and a small knob basally, distinctly sulcate dorsally, movable spur moderately arcuate, surface with intricate reticulation. Mesotibia with distinct and moderately long protuberance preapically with acuminate apex and with its seta inserted laterally, with eight setae furnished tubercles above protuberance, with few fine setae at the inner side. Front leg with tarsomeres slightly widened, first tarsomere elongated, slender and with carina baso-laterally, laterally with acute tubercle furnished with four short setae. Male genitalia (Fig. 78). Median lobe slender, moderately convex, more curved at root of apical lamella, apex with flattened wide lamella. Parameres moderately slender, with three setae at apex. Female gonocoxites and epipleurite (Fig. 109). Gonocoxites scapiform, extraordinary slender, less sclerotized, not fused, completely movable; basal gonocoxite 1 elongated, with four setae, apical gonocoxite 2 slender, distinctly elongated, indistinctly curved in its whole length, cylindrical in apical half, with two long setae at apex, with one long setae at lateral margin at middle, with one minute seta. Epipleurite with three setae, two close together and one isolated. Variation. The separation by a notch between the clypeus and the clypeal-wings varies between the two specimens, as well as the rugosity of the clypeal-furrow.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97304FFBD53D4A7ABFB04F92E.taxon	distribution	Distribution. Known from the East of Nepal.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97304FFBD53D4A7ABFB04F92E.taxon	diagnosis	Diagnosis. A small sized fuscous species with square pronotum and interval seven and eight of the elytron reticulated. It differs mainly from the most similar species C. quadrataconvexa Balkenohl and convexaquadrata sp. nov. by the conspicuous tubercle at the base of interval three of the elytron, the different reticulation pattern of the lateral intervals, the smaller eyes, and its small size.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97304FFBD53D4A7ABFB04F92E.taxon	etymology	Etymology. The name refers to the base of the elytron which carries a large tubercle, and is expressed by a combination of the Latin substantive tuberculum (= tubercle) and the Latin verb ferre (= to carry).	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97307FFBB53D4A42FFBEEFEFA.taxon	description	(Figs 9, 44, 79)	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97307FFBB53D4A42FFBEEFEFA.taxon	materials_examined	Type material. Holotype: ♂, with labels and data: white, black printed “ Kankau (Koshun) Formosa H. Sauter 1912 ” / red, black framed and black printed “ TYPE ” / white, handwritten in black ink and black printed “ C. depressa Kt. det. K. Kult, 1948 ” / white, black printed “ COLLECTIO KAREL KULT COLL. A. DOSTAL, 1999 ” (CADW). Remarks. Because the specimen is the only available type material, it is the holotype by default. Missing parts are: left front leg, right terminal tarsomere, right hind tibia and tarsomeres. The head and anterior half of the pronotum shows micro-scratches giving the surface a dull appearance. There is a short description in Kult (1951).	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97307FFBB53D4A42FFBEEFEFA.taxon	description	Redescription. Measurements. Holotype: Body length 6.08 mm, width 1.67 mm; ratio L / W of pronotum 1.01; ratio L / W of elytra 1.99. Colour glossy, almost uniformly fuscous to ferruginous. Supraantennal plate and wing with margin more pale, mandibles piceous. Head a sixth narrower than pronotum. Clypeus with central part nearly straight, laterally with rounded teeth, separated from clypeal-wing by small but distinct notch; wing not as advanced as clypeal-teeth, obtuse; separated from supraantennal plate by obtuse notch, supraantennal plate laterally nearly straight in anterior half; clypeus, wing, and supraantennal plate reflexed margined. Supraantennal plate extended posteriorly into an obtuse keel, separated from wing by a slightly widened furrow. Clypeus with obtuse transverse carina, clypeal-furrow indicated laterally by flat impressions. Frons slightly convex, with pore-like impressions at middle, separated from supraantennal plate and supraorbital carina by furrow, with clypeal seta in front; supraorbital carinae of moderate size, fused with frons in posterior half of the carina, with two supraorbital setigerous punctures at middle of eye and posterior end of the gena. Neck constriction closed, formed by band of medium sized punctures and step from frons. Eye regularly sized, gena small but distinct. Antenna long, reaching up to flange of pronotum. Labrum nearly straight, irregularly reticulated, six-setose. Mentum laterally isodiametrically reticulated, transverse at middle, lobe oval, flattened, with fine transverse obtuse carinae, median tooth wide, rounded anteriorly, indistinctly higher than lobes. Pronotum (Fig. 44) with disk distinctly flattened in frontal and lateral view. Sub-quadrate, as long as wide. Reflexed lateral margin smooth, slightly convex laterally, with short straight but anteriorly attenuating part at level of anterior setigerous puncture, widest at middle; anterior angle distinct, rounded, posterior angle indicated by somewhat rounded tooth; lateral channel moderately wide, irregularly reticulated, continuing up to base as narrow, convex margin. Median line moderately narrow, deeply engraved, complete, with short transverse wrinkles basally. Anterior transverse line fine, joining median line, just reaching anterior margin. Disk with small sized punctures resembling very indistinctly to a letter Y, with few flattened transverse wrinkles laterally, with isodiametric reticulation at base, with posterior setigerous puncture distant from margin by two diameters of puncture. Lateral margin from posterior angle to flange of base slanted with angle of around 45 °. Basal channel on flange wider than lateral channel, distinctly carinate and posteriorly with transverse reticulation. Elytron (Fig. 9) with disk in frontal and lateral view distinctly flattened. Shape of elytra elongate, about two times longer than wide, with subparallel margin anterior middle, maximum width at middle. Humerus distinct, rounded, slightly prominent. Reflexed lateral margin subcrenulated posterior humerus. Row of setigerous punctures in the lateral channel interrupted anterior middle by a short distance. Scutellar striole distinct; setigerous tubercle at base of first stria distinct, with distinct tubercle at base of third interval, basal declivity with isodiametric reticulation. Striae not deep, nearly impunctate, one to four free at base, five and six joining at humerus, one and two, three and four, and five and six joining at apex. Intervals slightly convex, interval three with five setigerous punctures joining third stria, interval eight carinate at apex. Surface of interval one to six glossy on disk, interval seven and eight with isodiametric reticulation, apical quarter of elytron with isodiametric reticulation. Hind wing fully developed. Lower surface with proepisternum isodiametrically reticulated, transverse wrinkles not visible. Sternite of abdomen laterally with isodiametric reticulation, with transverse reticulation at middle. The two marginal setigerous punctures are widely separated. Legs with profemur with indistinct reticulation. Protibia covered with irregular reticulation, sulcate dorsally, with three spines, apical spine indistinctly widened apically, movable spur distinctly shorter than apical spine, arcuate. First tarsomere elongated, distinctly flattened at base, with acute tubercle at middle of tarsomere. Intermediate tibia with seven tubercles at the lateral side, with slender spur near apex, all setae furnished. Tarsomeres of front and intermediate leg slightly widened. Male genitalia (Fig. 79). Median lobe regularly arcuate at middle, apically continuously attenuated and flattened, tip of apex slightly curved laterally. Parameres moderately slender, with small apophysis, hyaline apically, with one seta at apex, each. Female gonocoxites and epipleurite unknown. Variation unknown.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97307FFBB53D4A42FFBEEFEFA.taxon	distribution	Distribution. Known from the type locality Kankau in the South of Taiwan.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97307FFBB53D4A42FFBEEFEFA.taxon	diagnosis	Diagnosis. A medium sized fuscous to ferruginous species with square pronotum and interval seven and eight of the elytron reticulated. It differs mainly from the most similar species C. subdepressa Kult by the pronotum which appears longer than wide and by the much shorter flange at base. In addition, the labrum has six setigerous punctures (five in C. subdepressa), and on the elytron there are five setigerous punctures.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97301FFBB53D4A29BFC3FFA92.taxon	description	(Figs 10, 45, 80)	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97301FFBB53D4A29BFC3FFA92.taxon	materials_examined	Type material. Holotype: ♂, with labels and data: white, black printed “ Lope de Vega, Northern Samar, Eastern Visayas FEBRUARY 2002 ” / backside “ loc. coll. lgt. ” / “ Coll. M. Balkenohl ” (CMBB). Remark. The left front leg is missing. The subspecies differs from the nominotypical subspecies mainly in the following characters.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97301FFBB53D4A29BFC3FFA92.taxon	description	Description. Measurements. Body length 5.48 mm, width 1.48 mm, ratio L / W of pronotum 0.98, ratio L / W of elytra 2.03. Head with labrum distinctly convex anteriorly, six-setose. Frons more intensively covered with small sized punctures. Pronotum (Fig. 45) with lateral margin slightly to moderately convex; basal channel of flange somewhat widened. Elytron (Fig. 10) double as long as wide. Interval three with four setigerous punctures; interval seven with short step-like carina at apex. Male genitalia (Fig. 80). Median lobe apically more abruptly attenuated to tip of apex as in nominotypical subspecies. Female gonocoxites and epipleurite unknown. Variation unknown.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97301FFBB53D4A29BFC3FFA92.taxon	distribution	Distribution. Philippines, Samar Island.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97301FFBB53D4A29BFC3FFA92.taxon	diagnosis	Diagnosis. The subspecies differs from the nominotypical subspecies mainly by the elytron with interval three with five setigerous punctures, interval seven with a short step-like carina at apex, the anteriorly convex margin of the labrum, the smaller size.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97301FFBB53D4A29BFC3FFA92.taxon	etymology	Etymology. The name refers to the island of Samar in the Philippines.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97301FFBA53D4A673FB7EF906.taxon	description	(Figs 11, 46, 81)	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97301FFBA53D4A673FB7EF906.taxon	materials_examined	Type material. Holotype: ♂, with labels and data: white, black printed “ Malaysia, Sabah, Batu Punggul Resort env., 24. VI. - 1. VII. 1996, 11 c. vegetation debris and forest floor litter accumulated around large trees near river ” (NHMW).	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97301FFBA53D4A673FB7EF906.taxon	description	Description. Measurements. Body length 5.77 mm, width 1.59 mm, ratio L / W of pronotum 0.89, ratio L / W of elytra 1.99. Colour glossy, piceous. Supraantennal plate indistinctly translucent. Antennae, mouthparts, intermediate and hind legs rufous-fuscous. Head a sixth narrower than pronotum. Clypeus with central part straight, with slightly projecting teeth laterally, separated from clypeal-wing by distinct notch; wing distinct, rounded, wider but not as projecting as clypeal-teeth, separated from supraantennal plate by distinct notch, supraantennal plate with margin laterally obtuse-angular, clypeus, wing, and supraantennal plate reflexed margined. Supraantennal plate extended posteriorly into a convex keel, separated from wing by flat furrow. Clypeus continuing into frons without furrow, slightly convex, with scattered small punctures. Frons with pore at middle, separated from supraantennal plate and supraorbital carina by sharp furrow, with clypeal seta in front; supraorbital carinae moderately long, narrow, widened posteriorly, with two supraorbital setigerous punctures at middle of eye and at end of the gena-level. Neck constriction closed, formed by a step from frons. Eye relatively large, gena small. Antenna elongate, reaching up middle of pedunculus. Labrum excised anteriorly, conspicuously bilobate, isodiametrically reticulated, seven-setose. Mentum isodiametrically reticulated, with oblique wrinkles, lobe with lateral margin latero-basally straight, with obtuse angle, latero-apically straight, with apex rounded, medially straight to cleft, median tooth wide, much lower than lobes, at apex bent dorsally. Pronotum (Fig. 46) with disk flattened in lateral view, moderately convex in frontal view. Shape cordate, wider than long. Reflexed lateral margin smooth, slightly convex, widest at middle; anterior angle distinctly angled, posterior angle nearly completely rounded off, visible by knob-like tooth; lateral channel moderately narrow, isodiametrically reticulated, continuing up to base as narrow, convex margin. Median line engraved, complete, joining anterior transverse line. Anterior transverse line distinct, complete. Disk nearly completely covered with small sized punctures, with some transverse wrinkles, with isodiametric reticulation at lateral margin and at base, with posterior setigerous puncture removed from margin by diameter of pore. Lateral margin from posterior angle to flange of base slanted with angle of around 45 °. Basal channel on flange moderately narrow, rugose, with sharp carina. Elytron (Fig. 11) with disk indistinctly convex in lateral view, distinctly convex in frontal view. Shape of elytra elongate, about two-fifth longer than wide, with lateral margin nearly parallel, maximum width at middle. Humerus distinct, rounded. Reflexed lateral margin slightly crenulate behind humerus. Row of setigerous punctures in the lateral channel regular in size, slightly more distant at middle. Scutellar striole distinct; setigerous tubercle at base of first stria distinct, with distinct tubercle at base of third interval, basal declivity perpendicularly falling to pedunculus, with isodiametric reticulation. Striae moderately deep, impunctate, one to four free at base, five and six joining at humerus, one and two ending free at apex, three and four and five and six joining at apex. Intervals moderately convex, interval three with four setigerous punctures joining third stria, interval five joining lateral margin at humerus as very short carina, interval seven with rim at apex, eight carinate at apex. Surface of interval one to six glossy, interval seven and eight with distinct isodiametric reticulation, apical quarter of elytron isodiametrically reticulated at all intervals. Hind wing fully developed. Lower surface. Proepisternum isodiametrically reticulated, with fine wrinkles laterally. Sternite of abdomen with isodiametric reticulation, more scaly medially. Last visible sternite with the two marginal setigerous punctures widely separated, with slight projection posteriorly at middle where it is flattened, with narrow marginal furrow at middle. Legs. Protibia slender, with longitudinal to irregular reticulation, with narrow sulcus dorsally, with three spines, movable spur nearly straight, basal tarsomere slender at base, carinate, with acute tubercle and two wide setae at middle of tarsomere. Intermediate tibia with nine tubercles at the lateral side, with slender spur near apex, all setae furnished. Tarsomeres of front leg widened. Male genitalia (Fig. 81). Median lobe distinctly arcuate in basal part, straight in apical half, apically regularly tapered, somewhat flattened. Parameres moderately slender, with small apophysis, slightly distorted, hyaline apically, with one minute seta at apex. Female gonocoxites and epipleurite unknown. Variation not observed.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97301FFBA53D4A673FB7EF906.taxon	distribution	Distribution. Known from Sabah on the island of Borneo.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97301FFBA53D4A673FB7EF906.taxon	diagnosis	Diagnosis. A medium sized piceous species with peltate pronotum and intervals seven and eight of the elytron reticulated. It is the only species with peltate pronotum and having the labrum seven-setose. In addition, the frons is covered by medium sized punctures. Moreover, it is the only species of the group with a distinctly bilobed labrum.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97301FFBA53D4A673FB7EF906.taxon	etymology	Etymology. The name refers to the labrum which is distinctly bilobed (Latin: bilobata).	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97300FFB853D4A5C7FEBDFDFE.taxon	description	(Figs 12, 47, 82, 110)	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97300FFB853D4A5C7FEBDFDFE.taxon	materials_examined	Type material. Holotype: ♂, with labels and data: yellow, black printed “ PHILIPPINES: LEYTE SW Abuyog, 28. II. 1991 river bank leg. SCHAWALLER & al. ” / white, printed “ Clivina nr. wallacei PUTZ. det. Balkenohl II. 93 ” (SMNS). Paratypes: 1 ♂, 2 ♀♀, 12 specs, same data as holotype (SMNS, CMBB). Remark. In one of the paratypes, an intermediate leg is missing.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97300FFB853D4A5C7FEBDFDFE.taxon	description	Description. Measurements. Body length 4.59 – 5.19 mm (ẍ = 5.12 mm *), width 1.24 – 1.48 mm (ẍ = 1.39 mm *), ratio L / W of pronotum 0.97 – 0.99 (ẍ = 0.98 *), ratio L / W of elytra 1.95 – 1.99 (ẍ = 1.98 *); (* n = 4). Colour glossy, fuscous, with rufous tinge. Supraantennal plate, wing of clypeus, and apex of elytron slightly translucent. Antennae, mouthparts, intermediate and hind legs nearly of same colour as body. Head a quarter narrower than pronotum. Clypeus with central part indistinctly excised, with indistinctly projecting rounded teeth laterally, separated from clypeal-wing by slight notch; wing wider than teeth of clypeus, obtuse-angled, not as projecting as clypeal-teeth, separated from supraantennal plate by obtuse-angled notch, supraantennal plate with margin laterally convex, clypeus reflexed margined. Supraantennal plate extended posteriorly into keel, separated from wing by narrow furrow. Clypeus transverse, moderately convex, smooth, clypeal-furrow flattened, more distinct laterally. Frons depressed, slightly hollowed out in posterior part, separated from supraantennal plate and supraorbital carina by distinct furrow, with clypeal seta in front; supraorbital carinae sharp anteriorly, obtuse posteriorly, with two supraorbital setigerous punctures at middle and end of gena-level. Neck constriction open at middle. Eye relatively small, somewhat flattened, gena distinct, enclosing eye posteriorly by one third. Antenna elongate, reaching distinctly over base of elytron. Labrum indistinctly excised anteriorly, finely isodiametrically reticulated, six-setose. Mentum with lobes pentagonal, finely margined, with irregular to isodiametric reticulation, median tooth nearly as high as lobe, hollowed out at tip, rectangular. Pronotum (Fig. 47) with disk flattened in lateral view, flattened at middle and moderately convex laterally in frontal view. Shape cordate to square, as long as wide, seemingly longer than wide. Reflexed lateral margin sub-crenulate, slightly convex at middle, widest at middle; anterior angle distinct, posterior angle distinct, with distinct anteriorly projecting somewhat rounded tooth; lateral channel narrow, distinctly reticulated, continuing up to base as sharp, convex margin. Median line complete, engraved, joining anterior transverse line. Anterior transverse line distinct, complete. Disk in basal half with puncture pattern resembling to the Geek letter gamma, with transverse wrinkles, with isodiametric reticulation at base and laterally, with anterior setigerous puncture slightly removed from lateral channel, with posterior setigerous puncture removed from margin by diameter of pore. Lateral margin from posterior angle to flange of base slanted with angle of around 45 °. Basal channel on flange regularly developed, somewhat rugose, with sharp carina. Elytron (Fig. 12) with disk flattened in anterior two thirds lateral view, flattened at middle and moderately convex in frontal view. Shape of elytra elongate, nearly two times longer than wide, with lateral margin straight anterior middle but slightly diverging, maximum width posterior middle. Humerus distinct, rounded. Reflexed lateral margin finely crenulate in anterior half. Lateral channel slightly narrower at middle. Row of setigerous punctures in the lateral channel somewhat distant at middle, scutellar striole distinct; setigerous tubercle at base of first stria distinct, with distinct tubercle at base of third interval, basal declivity falling perpendicularly to pedunculus, with isodiametric reticulation. Striae moderately deep, impunctate, one to four free at base, five and six joining at humerus, one and two, three and four, and five and six joining apically. Intervals slightly convex, interval three with four setigerous punctures joining third stria, interval five joining lateral margin at humerus as short carina, interval seven convex at apex, eight carinate at apex. Surface of interval one to six smooth, glossy, seven and eight as well as apex with isodiametric reticulation. Hind wing fully developed. Lower surface. Proepisternum isodiametrically reticulated, with fine wrinkles. Sternite of abdomen with isodiametric reticulation laterally, shingle-like at middle. Last visible sternite with the two marginal setigerous punctures widely separated, with moderately wide projection posteriorly at middle where it is slightly hollowed out, with narrow marginal furrow at middle (in both sexes). Legs slender, protibia with fine longitudinal reticulation, with slight sulcus dorsally, with three acute spines and obtuse tooth, the frontal spine somewhat widened, movable spur moderately arcuate, basal tarsomere slender at base, carinate, with acute tubercle with two setae and small tubercle at middle of tarsomere. Intermediate tibia with nine tubercles at the lateral side, with slender spur near apex, all setae furnished. Tarsomeres of front leg slightly widened. Male genitalia (Fig. 82). Median lobe moderately and regularly arcuate, in lateral view nearly parallel at middle, apex moderately elongated into a slightly arcuate spatula. Parameres moderately slender, with moderately developed apophyses, slightly distorted towards apices, hyaline apically, asetose. Female gonocoxites and epipleurite (Fig. 110). Basal gonocoxite somewhat widened, with two long setae apically, gonocoxite II elongated basally, slightly curved, cylindrical in apical half in cross section, with three medium sized and one small seta, with two long setae apically positioned close together. Epipleurite wide-oval, with one medium sized seta. Variation. Among the four specimens, the depression on the frons varies from completely explanated to slightly transversally hollowed out. The borders around the flattened area are more or less obtuse carinate.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97300FFB853D4A5C7FEBDFDFE.taxon	distribution	Distribution. Known from the type locality Abuyog, province Leyte in the Philippines.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97300FFB853D4A5C7FEBDFDFE.taxon	diagnosis	Diagnosis. A medium sized slender fuscous to rufous species with square to cordate pronotum and intervals seven and eight with isodiametric reticulation. A species sharing most of the basic characters (with exception of the colour) is C. gamma Andrewes. However, in C. gamma the female gonocoxites are of the very different falcate type, the pronotum with its margin is straight and more convex, on the elytron there is an apical setigerous punctures at stria two (beside the regular four setigerous punctures), and interval eight is completely carinate.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97300FFB853D4A5C7FEBDFDFE.taxon	etymology	Etymology. The name refers to the head with its slightly hollowed out frons (Latin: caput = head; depressus, a, um = depressed).	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97302FFB753D4A19FFF1BF9B9.taxon	description	(Figs 13, 48, 111)	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97302FFB753D4A19FFF1BF9B9.taxon	materials_examined	Type material. Holotype: ♀, with labels and data: white, black printed “ INDONESIEN Samabusa 25 km östl. Nabire COROVA - RIVER leg. Dr. D. DASCHE 14. - 18.02.95 “ / “ Clivina group helferi det. Balkenohl, 1999 ”, “ COLL. WRASE BERLIN ” / green, black printed “ COLL. WRASE GUSOW-PLATKOW GERM. ” (CDWGP). Paratype: 1 ♀, with labels and data: all white, black printed “ Palmer R. at Black R. Papua July 22 - 31, 1936 ArchboldExped. ” / “ Borrowed fr AMNH 1961 ” / “ Clivina wallacei Pz. det. Darl. 68 ” / barcode “ MCZ-ENT 00806446 ” (MCZ). Remarks. The paratype shows several damages: the two right terminal antennomeres are missing. In both of the front legs, the tarsomeres are missing. The right intermediate leg is missing as well as three tarsomeres of the left hind leg, and the left tibia with tarsomeres is dislocated. These damages gave reason to select the complete specimen as holotype. The paratype designated is mentioned in Darlington (1971) under the name Clivina wallacei Putzeys as the “ first surely recorded ” C. wallacei from the island. The whereabouts of the other two specimens mentioned in Darlington (1971) remain unclear but should be investigated in order to clear occurrence of C. wallacei wallacei in Papua / New Guinea.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97302FFB753D4A19FFF1BF9B9.taxon	description	Description. Measurements. Holotype: Body length 5.47 mm, width 1.52, ratio L / W of pronotum 0.99, ratio L / W of elytra 1.95. Paratype: Body length 5.31 mm, width 1.46, ratio L / W of pronotum 0.99, ratio L / W of elytra 2.0. Colour fulvous. Head and pronotum glossy, elytra opaque. Lateral margins of head fuscous. Antennae, mouthparts, intermediate and hind legs yellowish. Head about a fifth narrower than pronotum. Clypeus with central part straight, projection of lateral teeth distinct, separated from clypeal-wing by distinct notch; wing distinct, less high as lateral teeth but as high as central part of clypeus, separated from supraantennal plate by obtuse-angled notch, supraantennal plate with margin regularly convex, clypeus, wing and supraantennal plate reflexed margined. Supraantennal plate extended posteriorly into a blunt keel, separated from wing by narrow furrow. Clypeus transverse, convex, smooth, clypeal-furrow wide, flat, with indistinct reticulation, frons moderately convex, with scattered small punctures, with pore at middle, separated from supraantennal plate and supraorbital carina by moderately wide furrow, with clypeal seta in front; supraorbital carinae narrow, with two supraorbital setigerous punctures at middle of eye and at posterior gena-level. Neck constriction closed, developed as step from frons, with punctures posteriorly. Eye prominent, gena distinct, convex posteriorly. Antenna long, reaching over base of elytron, antennomeres four to ten elongate (L / W 2.0). Labrum pentagonal, anteriorly with flat angle and pointed at middle, irregularly to isodiametrically reticulated, six-setose. Lobe of mentum flat, regularly oval at tip, median tooth about as high as lobe, bent dorsally, rounded anteriorly. Pronotum (Fig. 48) with disk flat in lateral view, in frontal view moderately convex laterally, flattened at middle. Shape peltate, seemingly longer than wide but measured as long as wide. Reflexed lateral margin smooth, slightly convex, widest at middle; anterior angle rounded but distinct, posterior one widely rounded, with tooth indistinct; lateral channel narrow, reticulated; margin continuing up to base as sharp, convex margin. Median line complete, moderately wide, engraved, joining anterior transverse line. Anterior transverse line narrow, complete. Disk with few scattered punctures at middle, with transverse wrinkles, with isodiametric reticulation at base and laterally, with setigerous punctures situated beside channel. Lateral margin from posterior angle to flange of base slanted with angle of around 40 °. Basal channel on flange slightly widened, rugose, basal carina as wide as channel, sharp anteriorly. Elytron (Fig. 13) with disk in lateral view flattened in anterior two thirds, in frontal view slightly convex at middle and moderately convex laterally. Shape of elytra elongate, around two times longer than wide, with lateral margin nearly straight but diverging in anterior third, maximum width posterior middle. Humerus distinct, rounded. Reflexed lateral margin slightly crenulate behind humerus (50 - fold). Row of setigerous punctures in lateral channel regular in size, slightly more distant at middle, scutellar striole distinct; setigerous tubercle at base of first stria distinct, with distinct tubercle at base of third interval, basal declivity perpendicularly falling to pedunculus. Striae moderately deep, nearly impunctate, one to four free at base, five and six joining humerus, seventh shortened at humerus, one and two ending free at apex, three and four and five and six joining apically. Intervals slightly convex on disk, convexity more distinct laterally, interval three with four setigerous punctures joining third stria, interval five joining lateral margin at humerus as short carina, interval seven convex up to apex, eight carinate at apex. Surface of all intervals with distinct and regular isodiametric reticulation already traceable at magnification of 40 - fold. Hind wing fully developed. Lower surface. Proepisternum with isodiametric reticulation. Sternite of abdomen with isodiametric reticulation. Last visible sternite with the two marginal setigerous punctures widely separated, with slight flattened projection posteriorly at middle, with narrow marginal furrow at middle. Legs. Protibia with longitudinal reticulation, with wide sulcus dorsally, with three apically slender spines and distinct tubercle, movable spur moderately arcuate, basal tarsomere slender at base, carinate, with one acute and one blunt tubercle with three setae. Intermediate tibia with eight tubercles at the lateral side, with slender spur near apex, all setae furnished. Tarsomeres of front legs slightly widened. Male genitalia unknown. Female gonocoxites and epipleurite (Fig. 111). Basal gonocoxite inconspicuous, apically with two small setae, apical gonocoxite of moderate size, medially with one very long seta, with three moderately long setae, with two long setae apically. Epipleurite triangle-like, with two setae at middle. Variation. In the two specimens, the neck constriction is more or less punctured. The anterior transverse line of the pronotum is more or less engraved.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97302FFB753D4A19FFF1BF9B9.taxon	distribution	Distribution. Known from two localities in Indonesia, Papua at the Palmer River and from Samabusa near Nabire.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97302FFB753D4A19FFF1BF9B9.taxon	diagnosis	Diagnosis. A slender medium sized fulvous species with peltate pronotum, all intervals of the elytron reticulated, and with six-setose labrum. The most similar species are Clivina dealata brachyptera Darlington, C. hartmanni sp. nov., and C. spatulifigura sp. nov., all of them with isodiametric reticulation on all intervals of the elytron. Out of these, C. opacielytra sp. nov. is the only species with a six-setose and anteriorly pointed labrum. Mainly due to its falciform gonocoxites, C. dealata brachyptera belongs to a different species group, and the reticulation pattern on the elytron is different. In C. hartmanni sp. nov. and in C. spatulifigura sp. nov., the labrum is distinctly excised anteriorly. Moreover, both of them have interval seven of the elytron step-like carinate at apex.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97302FFB753D4A19FFF1BF9B9.taxon	etymology	Etymology. The name refers to the elytra (Greek: elytra = ceilings of alae) which are distinctly and completely isodiametrically reticulated giving the surface a dull opaque appearance (Latin: opacus, a, um = dull, like with shadow).	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F9730DFFB553D4A55AFC81FF4A.taxon	description	(Figs 14, 49, 83)	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F9730DFFB553D4A55AFC81FF4A.taxon	materials_examined	Type material. Holotype: ♂, with labels and data: grey, handwritten in blue ink “ PHILIPPINES ” / “ ROXAS Palawan ” / backside “ locCOLL ” / “ white, black printed “ MAY 2001 ” / white, black printed “ Coll. Balkenohl ” (CMBB).	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F9730DFFB553D4A55AFC81FF4A.taxon	description	Description. Measurements. Body length 4.35 mm, width 1.21 mm, ratio L / W of pronotum 0.97, ratio L / W of elytra 1.89. Colour glossy, fuscous. Supraantennal plate at the margin slightly translucent. Antennae, mouthparts, intermediate and hind legs yellowish. Head about a sixth narrower than pronotum. Clypeus with central part straight, with distinctly projecting rounded teeth laterally, separated from clypeal-wing by distinct obtuse-angles notch; wing wider than teeth of clypeus, rounded, not as projecting as clypeal-teeth, separated from supraantennal plate by obtuse-angled notch, supraantennal plate with margin laterally convex, clypeus distinctly reflexed margined. Supraantennal plate extended posteriorly into blunt keel, separated from wing by flat furrow. Clypeus transverse, slightly convex, smooth, clypeal-furrow flattened. Frons completely flat, with flat pore-like depression at middle, separated from supraantennal plate by relatively narrow furrow, with clypeal seta in front; supraorbital carinae blunt, nearly continuing to flat frons, with two supraorbital setigerous punctures at middle of eye level and posterior to gena-level. Neck constriction open at middle, with distinct step laterally. Eye relatively small, somewhat flattened, more convex anteriorly, gena distinct, enclosing eye posteriorly by nearly one quarter. Antenna elongate, reaching up to base of pronotum. Labrum slightly excised anteriorly, laterally rounded, finely isodiametrically reticulated, six-setose. Mentum with lobes flat, distinctly convex laterally, slightly concave medially, indistinctly margined, with few askew fine carinae, median tooth wide, higher as lobe, hollowed out at tip, nearly rectangular. Pronotum (Fig. 49) with disk flattened in lateral view, flattened at middle and slightly to moderately convex laterally in frontal view. Shape sub-cordate, nearly as long as wide. Reflexed lateral margin sub-crenulate (100 - fold), slightly convex at middle, widest at middle; anterior angle distinct, posterior angle indicated by distinct anteriorly projecting tooth; lateral channel moderately narrow, reticulated, continuing up to base as sharp, slightly convex margin. Median line complete, engraved. Anterior transverse line distinct laterally, fading at middle, indistinctly joining median line. Disk with finely scattered punctures, in basal half with group of median sized punctures, with fine transverse wrinkles laterally, with isodiametric reticulation at base and laterally, with posterior setigerous puncture somewhat removed from margin. Lateral margin from posterior angle to flange of base slanted with angle of around 45 °. Basal channel on flange widened, rugose, with sharp carina. Elytron (Fig. 14) with disk flattened in anterior two thirds in lateral view, flattened at middle and slightly to moderately convex in frontal view. Shape of elytra elongate, nearly two times longer than wide, with lateral margin straight anterior middle but slightly diverging, maximum width posterior middle. Humerus distinct, rounded. Reflexed lateral margin finely crenulate posterior humerus. Row of setigerous punctures in the lateral channel somewhat more distant at middle, scutellar striole distinct; setigerous tubercle at base of first stria distinct, with small tubercle at base of third interval, basal declivity falling perpendicularly to pedunculus, with isodiametric reticulation. Striae moderately deep, impunctate, one to four free at base, five and six joining at humerus, one and two ending free at apex, three and four, and five and six joining apically. Intervals slightly convex, interval three with four setigerous punctures joining third stria, interval five joining lateral margin at humerus as short carina, interval seven convex at apex, eight carinate at apex. Surface of intervals two to eight with isodiametric reticulation, reticulation tapering in intensity towards middle, first interval nearly smooth. Hind wing fully developed. Lower surface. Proepisternum isodiametrically reticulated, completely covered with dense transverse wrinkles. Sternite of abdomen with isodiametric reticulation laterally, transverse at middle. Last visible sternite with the two marginal setigerous punctures widely separated, with moderately wide projection posteriorly at middle where it is slightly hollowed out, with narrow marginal furrow at middle. Legs slender, protibia with fine longitudinal reticulation, with widened sulcus dorsally, with three spines, spatulate-like, with obtuse tooth, movable spur moderately arcuate, basal tarsomere slender at base, carinate, with acute tubercle with two setae and a small tubercle at middle of tarsomere. Intermediate tibia with nine tubercles at the lateral side, with slender spur near apex, all setae furnished. Tarsomeres of same width. Male genitalia (Fig. 83). Median lobe relatively small, widened basally in dorsal and lateral view, moderately and regularly arcuate in basal half, indistinctly arcuate in apical half, apex arcuate at tip, in lateral view apically regularly and widely acuminate, somewhat flattened. Parameres slender, with moderately developed apophyses, slightly distorted towards apices, with small seta at apex, each. Female gonocoxites and epipleurite unknown. Variation unknown.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F9730DFFB553D4A55AFC81FF4A.taxon	distribution	Distribution. Known from Roxas on the island of Palawan, Philippines.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F9730DFFB553D4A55AFC81FF4A.taxon	diagnosis	Diagnosis. A small sized slender fuscous species with sub-cordate pronotum and intervals two to eight with isodiametric reticulation. A species sharing most of the basic characters is C. dilutureticulata sp. nov. However, and beside other characters, in C. dilutureticulata sp. nov. the frons of the head is moderately convex and covered with scattered fine punctures, the neck constriction is completely closed by a distinct sharp step, and the antennae are reaching over the base of the elytron.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F9730DFFB553D4A55AFC81FF4A.taxon	etymology	Etymology. The name is a patronym in honour of Hannes Baur (NHMB) who provided me with full logistical support during the last years and helped me to localize material.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F9730FFFB453D4A20BFC69FCC2.taxon	description	(Figs 15, 50, 84)	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F9730FFFB453D4A20BFC69FCC2.taxon	materials_examined	Type material. Holotype: ♂, with labels and data: white, black printed “ PHILIPPINEN - Mindoro 28 km S Calapan Balete 100 - 700 m leg. Jäch 27. - 29.11. 1992 ” (NHMW). Paratypes: 1 ♂, same data as holotype (CMBB); 1 ♂, “ PHILIPPINEN: Mindoro or. Baco, Hidden Paradise 19. - 20.11.1993 leg. H. Zettel (27) ” / “ Clivina VII. subdepressa Kult det. Alex. Dostal 1994 ” (CADW).	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F9730FFFB453D4A20BFC69FCC2.taxon	description	Description. Measurements. Body length 5.72 – 6.04 mm (ẍ = 5.91 mm *), width 1.67 – 1.74 mm (ẍ = 1.71 mm *), ratio L / W of pronotum 0.97 – 0.99 (ẍ = 0.98 *), ratio L / W of elytra 1.88 – 1.93 (ẍ = 1.90 *); (* n = 3). Colour glossy, fuscous. Supraantennal plate and clypeal-wing at the margin slightly translucent. Intermediate and hind legs somewhat paler, mouthparts and first four antennomeres yellowish. Head about a quarter narrower than pronotum. Clypeus with central part indistinctly excised, with distinctly projecting somewhat acute teeth laterally, separated from clypeal-wing by distinct obtuse-angles notch; wing wider than teeth of clypeus, rounded, not as projecting as clypeal-teeth, separated from supraantennal plate by obtuse-angled notch, supraantennal plate with margin laterally convex, clypeus, clypeal-wing, and anterior part of supraantennal plate distinctly reflexed margined. Supraantennal plate extended posteriorly into blunt keel, separated from wing by narrow furrow. Clypeus transverse, slightly convex, smooth, clypeal-furrow flattened. Frons moderately convex, with flat pore-like depression at middle, with scattered fine punctures, separated from supraantennal plate by moderately wide furrow, with clypeal seta in front; supraorbital carinae blunt, separating frons from supraorbital furrow, with two supraorbital setigerous punctures at middle of eye level and posterior to gena-level. Neck constriction closed, with distinct sharp step, with second narrow carina. Eye relatively small, somewhat flattened, gena distinct, enclosing eye posteriorly by nearly one quarter. Antenna elongate, reaching over base of elytron. Labrum straight anteriorly, laterally rounded, finely isodiametrically reticulated, six-setose. Mentum with lobes oval, angled at tip, obtusely margined, with isodiametric reticulation, median tooth wide, higher than lobe, hollowed out at tip. Pronotum (Fig. 50) with disk distinctly flattened in lateral view, flattened at middle and slightly convex laterally in frontal view. Shape sub-cordate, nearly as long as wide. Reflexed lateral margin sub-crenulate (80 - fold), moderately convex at middle, widest at middle; anterior angle distinct, posterior angle indicated by distinct anteriorly projecting tooth; lateral channel moderately narrow, reticulated, continuing up to base as sharp, distinctly convex margin. Median line complete, engraved. Anterior transverse line distinct, joining median line. Disk with finely scattered punctures, baso-medially with group of few median sized punctures, with few fine transverse wrinkles laterally, with isodiametric reticulation at base and laterally, with posterior setigerous puncture somewhat removed from margin. Lateral margin at flange slanted with angle of around 30 °. Basal channel on flange of regular width, rugose, with sharp carina. Elytron (Fig. 15) with disk flattened in anterior half in lateral view, flattened at middle and moderately convex in frontal view. Shape of elytra elongate, nearly two times longer than wide, with lateral margin straight anterior middle but slightly diverging, maximum width posterior middle. Humerus distinct, rounded. Reflexed lateral margin sub-crenulate (80 - fold) posterior humerus. Lateral channel slightly narrower at middle, row of setigerous punctures more distant at middle, scutellar striole distinct; setigerous tubercle at base of first stria distinct, with small tubercle at base of third interval, basal declivity falling perpendicularly to pedunculus, with isodiametric reticulation. Striae moderately deep, impunctate, one to four free at base, five and six joining at humerus, one and two ending free at apex, three and four, and five and six joining apically. Intervals slightly convex, interval three with four setigerous punctures joining third stria, interval five joining lateral margin at humerus as short carina, interval seven convex at apex, eight carinate at apex. Surface of intervals two to eight with isodiametric reticulation, reticulation tapering in intensity towards middle, first interval nearly smooth. Hind wing fully developed. Lower surface. Proepisternum isodiametrically reticulated, with fine flat transverse wrinkles laterally. Sternite of abdomen with isodiametric reticulation laterally, transverse at middle. Last visible sternite with the two marginal setigerous punctures widely separated, with moderately wide projection posteriorly at middle where it is flattened, with narrow marginal furrow at middle. Legs slender, protibia with fine longitudinal reticulation, with indistinct sulcus dorsally, with three spines, spatulate-like, with obtuse tooth, movable spur moderately arcuate, basal tarsomere slender at base, carinate, with acute tubercle with two setae and a small tubercle with one seta at middle of tarsomere. Intermediate tibia with nine tubercles at the lateral side, with slender spur near apex, all setae furnished. Tarsomeres of front leg somewhat widened. Male genitalia (Fig. 84). Median lobe slender, distinctly curved in basal half in dorsal view, nearly straight in apical half, slightly cracked at beginning of apical spatula, apex acute, in lateral view apically regularly acuminate, somewhat flattened. Parameres robust, with distinct apophyses, distorted, with small seta at apex, each. Female gonocoxites and epipleurite unknown. Variation. In one of the paratypes, the neck constriction is not doubled.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F9730FFFB453D4A20BFC69FCC2.taxon	distribution	Distribution. Known from the island of Mindoro, Philippines.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F9730FFFB453D4A20BFC69FCC2.taxon	diagnosis	Diagnosis. A medium sized slender fuscous species with sub-cordate pronotum and intervals two to eight with isodiametric reticulation. A species sharing most of the basic characters is C. bauri sp. nov. However, and beside other characters, in C. bauri sp. nov. the frons of the head is completely flat with the supraorbital carinae nearly continuing to the flat frons, the neck constriction is open at middle and the frons continues to the neck, and the antennae are much shorter, i. e. reaching up to base of the pronotum.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F9730FFFB453D4A20BFC69FCC2.taxon	etymology	Etymology. The name refers to the elytra with its reticulation on the intervals which is more and more fading to the middle (Latin: dilutus = diluted; reticulum = small sized net).	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F9730EFFB353D4A083FB05F832.taxon	description	(Figs 16, 51, 85)	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F9730EFFB353D4A083FB05F832.taxon	discussion	Remark. The short description of Kult (1951, p. 26) specifies two individuals, a “ TYPE ” from the Philippine Islands and a “ Paratype ” from Sarawak, both of them indicated as females. Investigation revealed, that both are males, and display distinct differences in the outer integument and of the male genitalia. So, they are not conspecific. The “ TYPE ” from the Philippine Islands is designated here as lectotype, the “ Paratype ” is described as a new species (see C. geiseri sp. nov.).	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F9730EFFB353D4A083FB05F832.taxon	materials_examined	Type material. Lectotype by present designation: ♂, with labels and data: white, handwritten in black ink “ Philippines Isl. Pulobatu ” / red, black framed and black printed “ TYPE ” / white, handwritten in black ink and black printed “ C. subdepressa Kt. det. K. Kult, 1948 ” / white, black printed “ COLLECTIO KAREL KULT COLL. A. DOSTAL, 1999 ” (CADW). Remark. The median lobe is slightly damaged at the oroficium and the parameres are severely damaged, looking like a dissection attempt. Missing parts are: left front leg, right intermediate tarsomeres, left hind tibia and tarsomeres. At the right front leg, the apical spine is broken by half. Additional material. 1 ♂, MINDANAO, MISAMIS OCC. DON VICTORIANO, 1700 m, 1. - 3. MAY 1996 BOLM lgt. (SMNS).	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F9730EFFB353D4A083FB05F832.taxon	description	Redescription. Measurements. Lectotype: Body length 4.78 mm, width 1.45 mm; ratio L / W of pronotum 0.95; ratio L / W of elytra 1.78. Second specimen (non-type): Body length 5.15 mm, width 1.52 mm; ratio L / W of pronotum 0.97; ratio L / W of elytra 1.87. Colour glossy, ferruginous. Supraantennal plate and clypeal-wing translucent. Intermediate and hind legs yellowish. Head a fifth narrower than pronotum. Clypeus with central part indistinctly convex, laterally with distinctly projecting teeth, separated from clypeal-wing by distinct obtuse notch; wing slightly toothed anteriorly, not as projecting as clypeal-teeth, separated from supraantennal plate by obtuse-angled notch, supraantennal plate with margin laterally rectangular, rounded at angle; clypeus, wing, and supraantennal plate in posterior part reflexed margined. Supraantennal plate extended posteriorly into an obtuse keel, separated from wing by a moderate furrow. Clypeus with transverse slightly convex field, clypeal-furrow indistinct. Frons moderately convex, with longitudinal pore at middle, with indistinct small punctures, separated from supraantennal plate and supraorbital carina by furrow, with clypeal seta in front; supraorbital carinae of moderate size, fused with frons in posterior half of the carina, with two supraorbital setigerous punctures at middle of eye and posterior end of the gena. Neck constriction closed, formed by a step from frons, neck completely covered with rough punctures. Eye regularly sized, gena small but distinct. Antenna long, reaching up to base of elytron. Labrum nearly straight, trapezoid, distinctly attenuated anteriorly, isodiametrically reticulated basally, five-setose. Mentum isodiametrically reticulated, without keel at middle, lobe wide-oval, flattened, cleft for palpomeres small, short, median tooth wide, distinctly higher than lobes. Pronotum (Fig. 51) with disk distinctly flattened in frontal and lateral view. Distinctly cordate, slightly wider than long. Reflexed lateral margin smooth, convex laterally in anterior half, widest slightly anterior middle; anterior angle widely rounded off, posterior angle distinct, not sharp, with concavity anterior to posterior angle; lateral channel moderately wide, somewhat flattened, irregularly to isodiametrically reticulated, continuing up to base as narrow, nearly straight margin. Median line wide, engraved, complete, with short transverse wrinkles basally. Anterior transverse line formed by connected punctures, not joining median line, not reaching anterior margin. Disk with few scattered punctures in basal half, with few transverse wrinkles laterally and basally, with isodiametric reticulation laterally at base, with impression of posterior setigerous puncture with large diameter, conspicuously distant from margin. Lateral margin from posterior angle to flange of base slanted with angle of around 45 °. Basal channel on flange conspicuously widened, wider than lateral channel, with narrow, obtuse carina. Elytron (Fig. 16) with disk in frontal and lateral view distinctly flattened. Shape of elytra elongate, about two-fifth longer than wide, with lateral margin indistinctly convex at middle and diverging, maximum width posterior middle. Humerus rounded off and prominent anteriorly. Reflexed lateral margin subcrenulated in anterior half. Row of setigerous punctures in the lateral channel more distant at middle. Scutellar striole distinct; setigerous tubercle at base of first stria distinct, with small tubercle at base of third interval, basal declivity overhanging for a short distance, with isodiametric reticulation. Striae moderately deep, nearly impunctate, one to four free at base, five and six joining at humerus, one and two ending free at apex, three and four, and five and six joining at apex. Intervals slightly convex, interval three with four setigerous punctures joining third stria, interval seven and eight not carinate, seventh running to apex as obtuse convexity, convexity of eighth ending before apex. Surface of interval one to seven glossy on disk, interval eight with isodiametric reticulation. Hind wing fully developed. Lower surface with proepisternum isodiametrically reticulated, with fine transverse wrinkles. Sternite of abdomen laterally with isodiametric reticulation, with transverse shingle-like reticulation at middle. Last visible sternite with the two marginal setigerous punctures are widely separated, with two small lappets. Legs. Profemur with transverse rugae. Protibia covered with longitudinal reticulation, with indistinct sulcus dorsally, with three spines; first tarsomere elongated, distinctly flattened at base, with acute tubercle at middle of tarsomere. Intermediate tibia with six tubercles at the lateral side, with slender spur near apex, all setae furnished. Tarsomeres of front and intermediate leg widened. Addition from the non-type specimen. The acute tubercle at middle of the front tarsomere is furnished with a short but robust seta. The apical spine is indistinctly widened apically. Male genitalia (Fig. 85) with median lobe moderately and regularly arcuate at middle, apically slightly widened and somewhat flattened. Parameres moderately slender, with small apophysis, nearly not distorted, hyaline apically, asetose. Female gonocoxites and epipleurite unknown. Variation. In the non-type specimen, the reticulation of interval eight is more distinct as well as the row of setigerous punctures in the lateral channel. In addition, the specimen shows a very small tubercle at the base of interval four.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F9730EFFB353D4A083FB05F832.taxon	distribution	Distribution. Known from the Philippine Islands, Pulobatu and Mindanao.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F9730EFFB353D4A083FB05F832.taxon	diagnosis	Diagnosis. A small sized ferruginous species with distinctly cordate pronotum and interval eight of the elytron reticulated. Three of the species with distinctly cordate pronotum, C. geiseri sp. nov., C. subdepressa Kult, and C. depressiforma sp. nov. show in addition a conspicuously elongated flange of the pronotum with flat basal carina. C. subdepressa Kult differs from all of them mainly by the five-setose labrum, the nearly smooth surface of the pronotum, and the smooth lateral margin of the pronotum. In addition, C. depressiforma sp. nov. differs by the pronotum with the laterally and acutely projecting teeth of the posterior angles and the coreaceate-like opaque surface, and the conspicuously rounded off anterior angles. In C. geiseri sp. nov., the surface of the pronotum is glossy and shows some medium-sized punctures. Moreover, the male aedeagi of the three species are very different.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97308FFB153D4A3BAFACAFD6E.taxon	description	(Figs 17, 52, 86, 112)	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97308FFB153D4A3BAFACAFD6E.taxon	materials_examined	Type material. Holotype: ♂, with labels and data: white, black printed “ BRUNEI-Temburong Belalong Field Res. C. Mix. Dipterocarp For. ~ 100 m 2 - 8.5.95 Heiss (CADW). Paratypes: 1 ♂, 1 ♀, same data as holotype, the male with additional handwritten labels “ helferi Gr. ” and “ nov. Gen ?? ” (CADW / CMBB).	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97308FFB153D4A3BAFACAFD6E.taxon	description	Description. Measurements. Body length 4.91 – 5.75 mm (ẍ = 5.13 mm *), width 1.28 – 1.56 mm (ẍ = 1.39 mm *), ratio L / W of pronotum 1.06 – 1.09 (ẍ = 1.08), ratio L / W of elytra 1.89 – 1.94 (ẍ = 1.92 *); (* n = 3). Colour glossy, fuscous with ferruginous tinge. Supraantennal plate at margin slightly translucent. Antennae, mouthparts, and tarsomeres yellowish. Antennomeres darker from joint four onwards. Head a fifth narrower than pronotum. Clypeus with central part straight, with slightly projecting sharp teeth laterally, separated from clypeal-wing by distinct notch; wing distinct, much less projecting as clypeal-teeth, somewhat rounded, separated from supraantennal plate by acute-angled notch, supraantennal plate with margin laterally convex, clypeus, wing, and supraantennal plate slightly reflexed margined. Supraantennal plate extended posteriorly into a widened blunt keel, separated from wing by small sharp furrow. Clypeus and frons developed as continuous moderately convex field, with few scattered micro-punctures, with indistinct flat impression centrally, separated from supraantennal plate and supraorbital carina by wide roughly reticulated furrow, with clypeal seta in front; supraorbital carinae short, blunt, with two supraorbital setigerous punctures at middle of eye level and at level of neck constriction. Neck constriction closed, formed by distinct step from frons, neck covered with irregular coreaceate-like structure. Eye small and flattened in dorsal view, main part situated ventrally, gena and supraorbital carina somewhat enlarged. Antenna elongate, reaching distinctly up to base of elytron. Labrum slightly convex, isodiametrically reticulated, seven-setose. Mentum isodiametrically reticulated, lobe with askew wrinkles, with increasing convexity anteriorly, cleft for palpomeres small, short, median tooth acuminated to tip, as high as lobes. Pronotum (Fig. 52) conspicuously flattened frontal view, slightly convex in lateral view. Shape elongate, sub-cordate, longer than wide. Reflexed lateral margin slightly crenulate, slightly convex, widest at middle, teeth of posterior angles seemingly wider than margin; anterior angle widely rounded off, posterior angle marked by distinct anteriorly projecting tooth, with short concavity anterior to posterior angle; lateral channel narrow, isodiametrically reticulated, continuing up to base as narrow, straight margin. Median line engraved, complete, joining and surpassing anterior transverse line. Anterior transverse line distinct, complete. Disk covered with medium sized punctures, with fine transverse wrinkles, with isodiametric reticulation basally and laterally, with impression of posterior setigerous puncture distinct, conspicuously distant from margin. Lateral margin from posterior angle to flange of base slanted with angle of around 45 °. Basal channel on flange conspicuously widened, coreaceate, with narrow carina. Elytron (Fig. 17) with disk in frontal view sub-moderately convex, in lateral view flattened in anterior half. Shape of elytra distinctly elongated, about two times longer than wide, with lateral margin indistinctly convex in anterior two thirds, indistinctly diverging, maximum width slightly posterior middle. Humerus rounded off. Base distinctly retracted. Reflexed lateral margin slightly crenulate in anterior half. Row of setigerous punctures in the lateral channel of equal distance. Scutellar striole distinct; setigerous tubercle at base of first stria distinct, with tubercle at base of third interval extraordinary small, nearly invisible, basal declivity perpendicularly falling to pedunculus, with isodiametric reticulation. Striae moderately deep, nearly impunctate, one to four free at base, five and six joining at humerus, one and two, three and four, and five and six joining at apex. Intervals somewhat flattened but still slightly convex, interval three with four setigerous punctures joining third stria, interval five joining lateral margin at humerus as short carina, interval seven and eight not carinate, seventh running to apex as obtuse convexity, convexity of eighth ending before apex. Surface of interval one to seven smooth, glossy, interval eight widened, with distinct and clear isodiametric reticulation. Hind wing fully developed. Lower surface. Proepisternum distinctly isodiametrically reticulated. Sternite of abdomen with isodiametric reticulation. Last visible sternite with the two marginal setigerous punctures widely separated, with somewhat widened projection at margin (in both sexes). Legs. Protibia slender, with longitudinal reticulation, with moderately wide sulcus dorsally, with three slender spines and obtuse tooth basally, basal tarsomere slender, slightly convex, and flattened at base, carinate laterally, with acute tubercle and two setae at middle of tarsomere. Profemur with distinct transverse nearly parallel wrinkles dorsally and ventrally. Intermediate tibia with eight tubercles at the lateral side, with slender spur near apex, all setae furnished. Tarsomeres of front and intermediate leg widened. Male genitalia (Fig. 86). Median lobe relatively short, moderately arcuate at middle, straight to apex. In lateral view moderately robust, widely acuminated. Parameres regularly convex at middle, with small and elongated apophyses, distorted at base, hyaline apically, asetose. Female gonocoxites and epipleurite (Fig. 112). Basal gonocoxite crescent-shaped, with two long setae, apical gonocoxite relatively stout, nearly straight, with long seta at basal third, with two very long setae apically. Epipleurite lappet-like, basally with two setae close together. Variation. On the frons of the head, the numbers of punctures postero-laterally vary. On the disk of the pronotum, the punctures vary in number.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97308FFB153D4A3BAFACAFD6E.taxon	distribution	Distribution. Known from Brunei on the island of Borneo.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97308FFB153D4A3BAFACAFD6E.taxon	diagnosis	Diagnosis. A medium sized fuscous to ferruginous species with cordate pronotum and interval eight of the elytron distinctly reticulated. The other three species with conspicuously elongated flange of the pronotum, and with missing carinae on the intervals, C. geiseri sp. nov., C. subdepressa Kult, and C. depressiforma sp. nov. show a different shape of the pronotum which is in all three cases distinctly wider than long and conspicuously cordate whereas in C. gracilicollis sp. nov. it is distinctly longer than wide and more elongate than cordate. In addition, in C. gracilicollis sp. nov. the base of the pronotum is conspicuously retracted.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97308FFB153D4A3BAFACAFD6E.taxon	etymology	Etymology. The name refers to the slender pronotum (Latin: gracilis = slender; collum = pronotum).	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F9730BFFB053D4A06FFE36F80A.taxon	description	(Figs 18, 53, 87, 113)	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F9730BFFB053D4A06FFE36F80A.taxon	materials_examined	Type material. Holotype: ♂, with labels and data: yellow, black printed “ BORNEO: SABAH Bingkor N Keningau 400 - 500 m, 19. - 20. XI. 1996 leg. W. SCHAWALLER ” / white, black printed “ Clivina nr. vigil DARL. det. Balkenohl, 1998 ” / “ Coll. M. Balkenohl ” (SMNS). Paratypes: 8 specs, same data as holotype (SMNS, CMBB); 2 specs, with labels and data: yellow, black printed “ BORNEO: SABAH, Bingkor N Keningau 400 - 500 m, at light; 20. XI. 1996, leg. D. Grimm ” / “ Clivina nr. vigil DARL. det. Balkenohl, 1998 ” / “ comp. ≠ vigil aus AUS / Baehr ” (SMNS); 2 specs, with labels and data: white, black printed “ BORNEO, Sabah, Danum Valley, Rainforest Lodge, 19. - 20. X. 2009, leg. U. + H. J. BREMER ” (SMNS); 1 ♂, 1 ♀, with labels and data: white, black printed “ SARAWAK (Borneo) ca 40 km SE KAPIT 03.1994, J. Kodada leg. ” (NHMW); 2 ♀♀, with labels and data: white, black printed “ Malaysia, Sabah, Crocker Range National park, Long-kogungan env., ca 750 - 850 m a. s. l., 19. - 21. VI. 1996, 7 c (NHMW); 1 ♀, with labels and data: white, black printed “ BRITISH N. BORNEO Tawau Residency Kalabakan R., Tawau 48 km W, 9 - 18. XI. ’ 58 ” / “ Primary Forest ” / “ T. C. Maa Collector BISHOP ” (NHMUK); 2 ♀♀, with labels and data: white, black printed “ MALAYSIA – Sabah prov. Banjaran Crocker Mts. GUNUNG ALAB peak 30. IV. - 27. V. 1996. 1650 - 1800 m M. Strba & R. Hergovits leg. ” / “ VII. helferi Gr. det. Dr. A. DOSTAL 2000 ” / red, handwritten “ sp. nov. ad. depressa ” (CADW).	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F9730BFFB053D4A06FFE36F80A.taxon	description	Description. Measurements. Body length 5.23 – 6.02 mm (ẍ = 5.47 mm *), width 1.55 – 1.77 mm (ẍ = 1.66 mm *), ratio L / W of pronotum 0.89 – 0.93 (ẍ = 0.90), ratio L / W of elytra 1.74 – 1.81 (ẍ = 1.77 *); (* n = 10). Colour glossy, fuscous to ferruginous. Supraantennal plate at margin and clypeal-wing translucent. Antennae, mouthparts, intermediate and hind legs yellowish. Antennomeres darker from joint four onwards. Head a quarter narrower than pronotum. Clypeus with central part straight, with slightly projecting sharp teeth laterally, separated from clypeal-wing by distinct notch; wing distinct, much less projecting as clypeal-teeth, rounded, separated from supraantennal plate by obtuse-angled notch, supraantennal plate with margin laterally obtuse-angularly rounded, clypeus, wing, and supraantennal plate slightly reflexed margined. Supraantennal plate extended posteriorly into a somewhat widened keel, separated from wing by furrow. Clypeus and frons developed as continuous moderately convex field, with scattered punctures, separated from supraantennal plate and supraorbital carina by wide roughly reticulated furrow, with clypeal seta in front; supraorbital carinae narrow, with two supraorbital setigerous punctures at middle of eye and distinctly posterior end of the gena. Neck constriction closed, formed by a step from frons, neck covered with irregular partly connected punctures. Eye prominent, regularly sized, gena distinct. Antenna elongate, reaching distinctly over base of elytron. Labrum nearly sub-circular, irregularly reticulated, seven-setose. Mentum isodiametrically reticulated, lobe with askew wrinkles, convex anteriorly, median tooth nearly rectangular at tip, not as high as lobes. Pronotum (Fig. 53). Conspicuously flattened but still slightly convex in frontal and lateral view. Distinctly cordate, wider than long. Reflexed lateral margin slightly crenulate, convex, widest at middle and at teeth of posterior angles; anterior angle conspicuously and widely rounded off, bent ventrally, posterior angle marked by distinct laterally projecting tooth, with short concavity anterior to posterior angle; lateral channel moderately wide, somewhat flattened, irregularly to isodiametrically reticulated, continuing up to base as narrow, straight margin. Median line engraved, complete, joining anterior transverse line. Anterior transverse line distinct, complete. Disk covered with partly connected punctures, with numerous transverse wrinkles, with isodiametric rough reticulation basally and laterally, with impression of posterior setigerous puncture distinct, conspicuously distant from margin. Lateral margin from posterior angle to flange of base slanted with angle of around 45 °. Basal channel on flange conspicuously widened, coreaceate, with narrow, obtuse carina. Elytron (Fig. 18) with disk in frontal view moderately convex, in lateral view flattened in anterior half. Shape of elytra elongate, about two-fifth longer than wide, with lateral margin nearly straight at middle but slightly diverging, maximum width slightly posterior middle. Humerus rounded off. Reflexed lateral margin slightly crenulate in anterior half. Row of setigerous punctures in the lateral channel wider separated at middle. Scutellar striole distinct, impressed anteriorly; setigerous tubercle at base of first stria distinct, with tubercle at base of third interval, basal declivity perpendicularly falling to pedunculus, with isodiametric reticulation. Striae moderately deep, indistinctly punctate-striate, one to four free at base, five and six joining at humerus, one and two ending free at apex, three and four, and five and six joining at apex. Intervals somewhat flattened but still slightly convex, interval three with four setigerous punctures joining third stria, interval five joining lateral margin at humerus as short carina, interval seven and eight not carinate, seventh running to apex as obtuse convexity, convexity of eighth ending before apex. Surface of interval one to seven glossy but covered with indistinctly and evanescent reticulation, interval eight with distinct and clear isodiametric reticulation. Hind wing fully developed. Lower surface. Proepisternum isodiametrically reticulated, with indistinct transverse wrinkles laterally. Sternite of abdomen with isodiametric reticulation, weaker medially. Last visible sternite with the two marginal setigerous punctures widely separated, with two small but distinct lappets at margin (in both sexes). Legs. Protibia slender, with longitudinal reticulation, with moderately deep sulcus dorsally, with three slender spines, basal tarsomere fractuate and flattened at base, carinate laterally, with acute tubercle and two setae at middle of tarsomere. Intermediate tibia with eight tubercles at the lateral side, with slender spur near apex, all setae furnished. Tarsomeres of front leg widened. Male genitalia (Fig. 87). Median lobe relatively short, moderately arcuate at middle, apically regularly rounded, slightly flattened. In lateral view moderately acuminated. Parameres moderately slender, with small apophysis, slightly distorted, slightly more arcuate and hyaline apically, asetose. Female gonocoxites and epipleurite (Fig. 113). Basal gonocoxite crescent-shaped, with two long setae, apical gonocoxite elongated, slightly convex, with long seta at middle and shorter one sub-medially, with two very long setae apically. Epipleurite lappet-like, with two setae close together and one isolated towards middle. Variation. The clypeus with central part is straight in the holotype but in some of the paratypes slightly undulate. On the elytron, the reticulation on intervals one to seven varies in distinctness from clearer to seemingly washed out.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F9730BFFB053D4A06FFE36F80A.taxon	distribution	Distribution. Known from Sabah and Sarawak on the island of Borneo	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F9730BFFB053D4A06FFE36F80A.taxon	diagnosis	Diagnosis. A medium sized fuscous to ferruginous species with distinctly cordate pronotum and intervals of the elytron more or less distinctly reticulated. Three of the species with distinctly cordate pronotum, C. geiseri sp. nov., C. subdepressa Kult, and C. depressiforma sp. nov. show in addition a conspicuously elongated flange of the pronotum with flat basal carina. C. depressiforma sp. nov. differs distinctly as only species by the pronotum with the laterally and acutely projecting teeth of the posterior angles, the coreaceate-like surface, and the conspicuously rounded off anterior angles. In addition, in C. depressiforma interval eight of the elytron is clearly and distinctly reticulated isodiametrically and all of the other intervals are indistinctly and evanescent reticulated. Moreover, the male aedeagi of the three species are very different.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F9730BFFB053D4A06FFE36F80A.taxon	etymology	Etymology. The name refers to the pronotum with its extraordinary depressed dorsal surface. (Latin: depressus = depressed; forma = form).	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97335FF8E53D4A3BAFE3DFC56.taxon	description	(Figs 19, 54, 88, 114)	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97335FF8E53D4A3BAFE3DFC56.taxon	materials_examined	Type material. Holotype: ♂, with labels and data: white, black printed “ SARAWAK: [underlined in yellow] foot of Mt. Dulit, junction of rivers Tinjar & Lejok. 29. viii. 1932. ” / white, black printed “ Recent Clearing ” / white, black printed “ Old secondary forest ” / white, black printed “ Oxford Univ. Exp. B. M. Hobby & A. W. Moore. B. M. 1933 - 254 ” white, black printed and handwritten in grey ink “ Clivina sp. H. E. Andrewes det. ” / white, black printed and handwritten in black ink “ C. subdepressa Kt. det. K. Kult 1950 ” / red, black framed and black printed “ PARATYPE ” / circular, yellow framed, black printed “ Para-type ” (NHMUK). Paratypes: 1 ♀, with labels and data: white, black printed SARAWAK (Borneo), ca 25 km E KAPIT III. 1994, Kodada leg. (NHMW); 2 ♂♂, same data as HT (NHMW); 2 ♀♀, same data but ca 40 km E KAPIT (NHMW). Remark. The holotype represents the paratype designated by Kult (1951) as C. subdepressa which is not conspecific with the holotype of C. subdepressa (see remarks under C. subdepressa).	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97335FF8E53D4A3BAFE3DFC56.taxon	description	Description. Measurements. Body length 4.49 – 4.87 mm (ẍ = 4.72 mm *), width 1.31 – 1.41 mm (ẍ = 1.36 mm *), ratio L / W of pronotum 0.93 – 0.97 (ẍ = 0.95 *), ratio L / W of elytra 1.82 – 1.88 (ẍ = 1.85 *); (* n = 6). Colour glossy, fulvous. Supraantennal plate and clypeal-wing translucent. Antennae, mouthparts, intermediate and hind legs yellowish. Head a quarter narrower than pronotum. Clypeus with central part straight, with distinctly projecting teeth laterally, separated from clypeal-wing by distinct notch; wing distinct, not as projecting as clypeal-teeth, separated from supraantennal plate by obtuse-angled notch, supraantennal plate with margin laterally obtuse-angular, clypeus, wing, and supraantennal plate slightly reflexed margined. Supraantennal plate extended posteriorly into a somewhat widened keel, separated from wing by furrow. Clypeus with transverse slightly convex field, clypeal-furrow flattened. Frons moderately convex, with indistinct pore at middle, with scattered small punctures, separated from supraantennal plate and supraorbital carina by wide furrow, with clypeal seta in front; supraorbital carinae narrow, with two supraorbital setigerous punctures at middle of eye and distinctly posterior end of the gena. Neck constriction closed, formed by a step from frons, neck covered with punctures. Eye regularly sized, gena small but somewhat bulgy. Antenna elongate, reaching over base of elytron. Labrum straight anteriorly, laterally rounded, irregularly reticulated, six-setose. Mentum isodiametrically reticulated, lobe laterally convex, tip convex, medially straight, flattened, median tooth wide, slightly higher than lobes. Pronotum (Fig. 54) with disk distinctly flattened in frontal and lateral view. Distinctly cordate, wider than long. Reflexed lateral margin slightly crenulate, convex, widest at middle; anterior angle widely rounded off, posterior angle distinctly marked by tooth, with short concavity anterior to posterior angle; lateral channel moderately wide, somewhat flattened, irregularly to isodiametrically reticulated, continuing up to base as narrow, indistinctly convex margin. Median line engraved, complete, joining and surpassing anterior transverse line. Anterior transverse line distinct, complete. Disk bilaterally with cloud of medium sized punctures in basal half, with numerous transverse wrinkles, with isodiametric reticulation laterally at base, with impression of posterior setigerous puncture with large diameter, conspicuously distant from margin. Lateral margin from posterior angle to flange of base slanted with angle of around 45 °. Basal channel on flange conspicuously widened, rugose, with narrow, obtuse carina. Elytron (Fig. 19) with disk in frontal and lateral view distinctly flattened. Shape of elytra elongate, about two-fifth longer than wide, with lateral margin nearly straight at middle but slightly diverging, maximum width posterior middle. Humerus rounded off. Reflexed lateral margin slightly crenulate in anterior half. Row of setigerous punctures in the lateral channel regular in size and distance. Scutellar striole distinct; setigerous tubercle at base of first stria distinct, with small tubercle at base of third interval, basal declivity perpendicularly falling to pedunculus, with isodiametric reticulation. Striae moderately deep, indistinctly punctate-striate, one to four free at base, five and six joining at humerus, one and two ending free at apex, three and four, and five and six joining at apex. Intervals slightly convex, interval three with four setigerous punctures joining third stria, interval five joining lateral margin at humerus as short carina, interval seven and eight not carinate, seventh running to apex as obtuse convexity, convexity of eighth ending before apex. Surface of interval one to seven glossy, interval eight with distinct isodiametric reticulation. Hind wing fully developed. Lower surface. Proepisternum isodiametrically reticulated, with indistinct transverse wrinkles. Sternite of abdomen with isodiametric reticulation. Last visible sternite with the two marginal setigerous punctures widely separated, with two slight embayments at margin. Legs. Protibia slender, with longitudinal reticulation, with sulcus dorsally, with three spines; basal tarsomere distinctly flattened at base, carinate, with acute tubercle and seta at middle of tarsomere. Intermediate tibia with eight tubercles at the lateral side, with slender spur near apex, all setae furnished. Tarsomeres of front and intermediate leg widened. Male genitalia (Fig. 88). Median lobe moderately and regularly arcuate at middle, apically duckbill-shaped, somewhat flattened. Parameres moderately slender, with small apophysis, slightly distorted, hyaline apically, asetose. Female gonocoxites and epipleurite (Fig. 114). Basal gonocoxite with two long setae, apical gonocoxite elongated, slightly convex, with short and long seta basally, with two very long setae apically. Epipleurite with two setae. Variation. In the holotype, the clypeal-furrow is indistinct at middle. In the small series of six specimens, the wing of the clypeus is more or less toothed anteriorly or rounded.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97335FF8E53D4A3BAFE3DFC56.taxon	distribution	Distribution. Known from Sarawak on the island of Borneo.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97335FF8E53D4A3BAFE3DFC56.taxon	diagnosis	Diagnosis. A small sized fulvous species with distinctly cordate pronotum and interval eight of the elytron reticulated. Three of the species with distinctly cordate pronotum, C. geiseri sp. nov., C. subdepressa Kult, and C. depressiforma sp. nov. show in addition a conspicuously elongated flange of the pronotum with flat basal carina. Clivina depressiforma sp. nov. differs mainly by the pronotum with the laterally and acutely projecting teeth of the posterior angles and the coreaceate-like opaque surface, and the conspicuously rounded off anterior angles. Clivina subdepressa Kult differs mainly by the five-setose labrum, the nearly smooth surface of the pronotum, and the smooth lateral margin of the pronotum. Moreover, the male aedeagi of the three species are very different.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97335FF8E53D4A3BAFE3DFC56.taxon	etymology	Etymology. The name is a patronym in honour of Dr. Michael Geiser (NHMUK) who supported me admirably with full access to the Scaritinae Collection during my stay at the NHMUK, and who helped me to localize a lot of material of this contribution.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97334FF8D53D4A737FD50FA02.taxon	description	(Figs 20, 55, 89)	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97334FF8D53D4A737FD50FA02.taxon	materials_examined	Type material. Holotype: ♂, with labels and data: white, black printed “ Lope de Vega, Samar, Central Visayas JANUARY 2020 ” / backside handwritten in black ink “ reg. coll. leg. ” / white, black printed “ Coll. Balkenohl ” (CMBB). Remark. The right front leg is missing as well as the left tibia onwards.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97334FF8D53D4A737FD50FA02.taxon	description	Description. Measurements. Body length 5.35 mm, width 1.51 mm, ratio L / W of pronotum 0.94, ratio L / W of elytra 1.91. Colour glossy, piceous. Supraantennal plate at margin and legs fuscous. Antennae and mouthparts somewhat more pale. Head about a sixth narrower than pronotum. Clypeus with central part straight, with indistinctly projecting rounded teeth laterally, separated from clypeal-wing by obtuse-angled notch; wing slightly wider than teeth of clypeus, rounded, not as projecting as clypeal-teeth, separated from supraantennal plate by nearly rectangular notch, supraantennal plate with margin laterally convex, clypeus, wing, and anterior part of supraantennal plate distinctly reflexed margined. Supraantennal plate extended posteriorly into blunt keel, separated from wing by narrow furrow. Clypeus continuing into frons without clypeal-furrow. Clypeus slightly convex, frons moderately convex, with small indistinct flat pore-like depression at middle, separated from supraantennal plate by moderately wide furrow, with clypeal seta in front; supraorbital carinae blunt, with second smaller supraantennal carina, with two supraorbital setigerous punctures at middle of eye level and at posterior gena-level. Supraantennal plates, clypeus, and frons densely covered with scattered small punctures. Neck constriction closed at middle, developed as band of medium sized punctures and slight step. Eye prominent, gena small but distinct, enclosing eye posteriorly by about one quarter. Antenna elongate, reaching up to base of elytron. Labrum excised anteriorly, seemingly bilobed, finely isodiametrically reticulated, seven-setose. Mentum with lobes flat, convex laterally, with apex acute, isodiametrically reticulated, distinctly margined, median tooth flat, as high as lobe. Pronotum (Fig. 55) with disk flat in lateral view, moderately and regularly convex in frontal view. Shape sub-cordate to square, indistinctly wider than long. Reflexed lateral margin sub-crenulate (120 - fold), slightly convex at middle, widest at middle; anterior angle distinct, posterior angle distinct, indicated by distinct anteriorly projecting tooth; lateral channel moderately narrow, becoming wider posteriorly, reticulated, continuing up to base as sharp, convex margin. Median line complete, engraved. Anterior transverse line narrow, joining median line. Disk densely covered with small scattered punctures, with flat transverse wrinkles laterally, with isodiametric reticulation at base, with posterior setigerous puncture removed from margin by two diameters of pore. Lateral margin from posterior angle to flange of base slanted with angle of around 30 °. Basal channel on flange distinctly widened, rugose, with sharp carina. Elytron (Fig. 20) with disk flattened in anterior half in lateral view, distinctly convex in frontal view. Shape of elytra elongate, nearly two times longer than wide, with lateral margin almost straight in anterior two thirds but slightly diverging, maximum width posterior middle. Humerus distinct, rounded, base somewhat retracted at middle. Reflexed lateral margin crenulate in anterior quarter. Row of setigerous punctures in the lateral channel of equally distance, scutellar striole distinct; setigerous tubercle at base of first stria distinct, with distinct tubercle at base of third interval, basal declivity falling perpendicularly to pedunculus, with isodiametric reticulation. Striae not very deep, impunctate, irregularly reticulated, one to four free at base, five and six joining at humerus, one to four ending free at apex, five and six joining apically. Intervals slightly convex, interval three with four setigerous punctures joining third stria, interval five joining lateral margin at humerus as short carina, interval seven and eight carinate at apex. Surface of intervals six to eight with isodiametric reticulation, interval one to five densely covered with small scattered punctures. Hind wing fully developed. Lower surface. Proepisternum isodiametrically reticulated, with small scattered punctures, with few transverse wrinkles laterally. Sternite of abdomen with isodiametric reticulation. Last visible sternite with the two marginal setigerous punctures widely separated, with moderately projection posteriorly at middle, with narrow marginal furrow at middle. Legs. Front tibia and front tarsomeres unknown. Intermediate tibia with nine tubercles at the lateral side, with slender spur near apex, all setae furnished. Tarsomeres of intermediate and hind legs of same width. Male genitalia (Fig. 89). Median lobe, distinctly arcuate in basal half, indistinctly arcuate in apical half, apex regularly arcuate to tip, in lateral view apically with duckbill-like spatula. Parameres distinctly arcuate, with distinct apophyses, with small seta at apex, each. Female gonocoxites and epipleurite unknown. Variation unknown.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97334FF8D53D4A737FD50FA02.taxon	distribution	Distribution. Known from Lope de Vega on the island of Samar, Philippines.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97334FF8D53D4A737FD50FA02.taxon	diagnosis	Diagnosis. A medium sized piceous species with sub-cordate to square pronotum, intervals six to eight with isodiametric reticulation. In addition, interval one to five, head, pronotum, and proepisternum is densely covered with scattered small punctures. It is the only species of the group with this puncture pattern on the whole dorsal surface of the body.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97334FF8D53D4A737FD50FA02.taxon	etymology	Etymology. The name refers to the intense scattered punctures on the head, pronotum and elytron (Latin: passim = scattered; punctatus = with punctures).	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97337FF8C53D4A6C3FE43F80A.taxon	description	(Figs 21, 56, 115)	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97337FF8C53D4A6C3FE43F80A.taxon	materials_examined	Type material. Holotype: ♀, with labels and data: white, black printed “ SARAWAK (Borneo), ca 40 km SE KAPIT 03.1994, J. Kodada leg. ” (NHMW). Paratype: 1 ♀, same data as holotype but “ ca 25 km E KAPIT ” (CMBB). Remark. In the paratype, the right hind leg and some tarsomeres of the left hind leg are missing. In addition, the elytra seem to be somewhat more slender, possibly due to a damage.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97337FF8C53D4A6C3FE43F80A.taxon	description	Description. Measurements. Holotype / paratype: Body length 6.16 mm / 5.92 mm, width 1.86 mm / 1.63 mm, ratio L / W of pronotum 0.90 / 0.92, ratio L / W of elytra 1.76 / 1.94. Colour glossy, piceous. Supraantennal plate indistinctly translucent. Antennae, mouthparts, intermediate and hind legs rufous-fuscous. Head a quarter narrower than pronotum. Clypeus with central part straight, with slightly but distinctly projecting teeth laterally, separated from clypeal-wing by distinct notch; wing distinct, rounded, wider but not as projecting as clypeal-teeth, separated from supraantennal plate by distinct notch, supraantennal plate with margin laterally obtuse-angular, clypeus, wing, and supraantennal plate reflexed margined. Supraantennal plate extended posteriorly into a blunt keel, separated from wing by furrow. Clypeus continuing into frons without furrow, slightly convex, with scattered small punctures laterally. Frons with indistinct indentation at middle, separated from supraantennal plate and supraorbital carina by furrow, with clypeal seta in front; supraorbital carinae blunt, with second shorter carina laterally, with two supraorbital setigerous punctures at middle of eye and posterior to gena-level. Neck constriction closed, formed by a straight step from frons. Eye moderately large, gena distinct but small. Antenna elongate, reaching up to base of elytron. Labrum slightly excised anteriorly, isodiametrically reticulated, seven-setose. Mentum isodiametrically reticulated, with oblique wrinkles, lobe with lateral margin convex, with acute apical tip, medially concave to cleft, median tooth much lower than lobes, acute, not carinate. Pronotum (Fig. 56) with disk slightly convex in lateral view, in frontal view flattened at middle and moderately convex laterally. Shape square to cordate, wider than long. Reflexed lateral margin distinctly serrate, nearly parallel, widest at middle; anterior angle somewhat rounded but well-marked, posterior angle distinct, with robust tooth; lateral channel distinctly widened, rugose and reticulated, continuing up to base as narrow, straight margin. Median line engraved, complete, joining and surpassing anterior transverse line. Anterior transverse line distinct, complete. Disk with pattern of smaller punctures resembling with some phantasy to the letter Y, with some transverse wrinkles, with isodiametric reticulation at lateral margin and at base, with posterior setigerous puncture removed from margin by diameter of pore. Lateral margin from posterior angle to flange of base slanted with angle of around 30 °. Basal channel on flange wide, rugose, with sharp carina. Elytron (Fig. 21) with disk flattened in anterior half in lateral view, distinctly convex in frontal view. Shape of elytra elongate, about two-fifth longer than wide, with lateral margin indistinctly convex and diverging, maximum width behind middle. Humerus distinct, rounded. Reflexed lateral margin slightly crenulate behind humerus. Row of setigerous punctures in the lateral channel regular in size and distance. Scutellar striole distinct; setigerous tubercle at base of first stria distinct, with small tubercle at base of third interval, basal declivity perpendicularly falling to pedunculus, with isodiametric reticulation. Striae moderately deep, indistinctly punctate, one to four free at base, five and six joining at humerus, one and two ending free at apex, three and four and five and six indistinctly joining at apex. Intervals slightly convex at middle, moderately convex laterally, interval three with four setigerous punctures joining third stria, interval five joining lateral margin at humerus as very short carina, interval seven with distinct carina at apex, eight more sharp carinate at apex. Surface of all intervals isodiametrically reticulated, reticulation more distinct laterally. Hind wing fully developed. Lower surface. Proepisternum with fine isodiametrical reticulation, with extremely fine wrinkles laterally. Sternite of abdomen with isodiametric reticulation, weaker medially. Last visible sternite with the two marginal setigerous punctures widely separated, with slight projection posteriorly at middle where it is slightly hollowed out and smooth, with narrow marginal furrow at middle. Legs. Protibia slender, with longitudinal reticulation, with narrow sulcus dorsally, with three spines, movable spur moderately arcuate, basal tarsomere slender at base, carinate, with acute tubercle and two wide setae at middle of tarsomere. Intermediate tibia with nine tubercles at the lateral side, with slender spur near apex, all setae furnished. Tarsomeres of front leg widened. Male genitalia unknown. Female gonocoxites and epipleurite (Fig. 115). Basal gonocoxite with two long and two minute setae, apical gonocoxite elongated, with one long seta at base and one long seta at beginning of apical third, with two very long setae apically. Epipleurite trapezoid-shaped, with one long seta arising from tubercle-like lappet. Variation. In the paratype, the elytra are slenderer, and the lateral margin of the pronotum is not as conspicuously serrate as in the holotype.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97337FF8C53D4A6C3FE43F80A.taxon	distribution	Distribution. Known from Sarawak on the island of Borneo.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97337FF8C53D4A6C3FE43F80A.taxon	diagnosis	Diagnosis. A medium sized piceous species with square to cordate shaped pronotum and all intervals of the elytron reticulated. The most similar species C. punctaticollis sp. nov., differs mainly by the elytra with intervals one to six glossy, the pronotum with the lateral margin sub-serrate, narrow lateral channel, much smaller tooth at the posterior angle, and the different puncture pattern on the disk. In addition, it is much smaller.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97337FF8C53D4A6C3FE43F80A.taxon	etymology	Etymology. The name refers to the pronotum with its lateral border (Latin: margo, marginis) which is distinctly serrated (Latin: serrate).	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97331FF8A53D4A3BAFC21FD52.taxon	description	(Figs 22, 57, 90, 116)	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97331FF8A53D4A3BAFC21FD52.taxon	materials_examined	Type material. Holotype: ♂, with labels and data: white, black printed “ INDONESIA, Irian Jaya Jayapura district Genyem, 50 m NN, IV. - V. 1999, leg. M. Balke ” / “ Sammlung NATURKUNDE- MUSEUM ERFURT ” (NKME). Paratype: 1 ♀, same data as holotype (CMBB).	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97331FF8A53D4A3BAFC21FD52.taxon	description	Description. Measurements. Body length 5.86 / 5.91 mm, width 1.79 / 1.77 mm, ratio L / W of pronotum 0.88 / 0.88, ratio L / W of elytra 1.77 / 1.78. Colour glossy, fuscous. Supraantennal plate at margin slightly translucent. Antennae, mouthparts, intermediate and hind legs yellowish. Head about a fifth narrower than pronotum. Clypeus with central part straight, projection of lateral teeth just traceable, separated from clypeal-wing by small notch; wing rounded, nearly as high as central part of clypeus, separated from supraantennal plate by obtuse-angled notch, supraantennal plate with margin convex at middle, clypeus reflexed margined. Supraantennal plate extended posteriorly into a blunt decreasing convexity, separated from wing by small furrow. Clypeus and frons forming continuous moderate convexity, with scattered small punctures, clypeal-furrow not traceable. Frons with pore at middle, separated from supraantennal plate and supraorbital carina by flat furrow, with clypeal seta in front; supraorbital carinae moderately developed, with two supraorbital setigerous punctures at middle of eye and at middle of gena-level. Neck constriction widely open, indicated laterally. Eye prominent but somewhat flattened, gena enclosing eye posteriorly by nearly a third. Antenna elongate, reaching over base of elytron by one to two antennomeres. Labrum excised anteriorly, finely irregularly reticulated, seven-setose. Lobe of mentum oval acuminated, flat, median tooth flat, distinctly lower than lobe, rectangular at apex. Pronotum (Fig. 57) with disk flattened in lateral view, in frontal view laterally moderately convex, flattened at middle. Shape square to sub-cordate, wider than long. Reflexed lateral margin smooth, widest at middle; anterior and posterior angles distinct, posterior one with distinct projecting sharp tooth; lateral channel moderately narrow, continuing up to base as sharp, sigmoid margin. Median line complete, engraved at middle, joining anterior transverse line. Anterior transverse line distinct, complete. Disk covered with scattered micro-punctures, with transverse wrinkles, with isodiametric reticulation at base, with indistinct impression in basal third, with posterior setigerous puncture situated directly at tooth of posterior angle. Lateral margin from posterior angle to flange of base slanted with angle of around 30 °. Basal channel on flange somewhat widened, rugose, with sharp carina. Elytron (Fig. 22) with disk flattened in anterior half in lateral view, moderately convex in frontal view. Shape of elytra elongate, slightly less than two times longer than wide, with lateral margin indistinctly convex at middle, diverging posteriorly, maximum width posterior middle. Humerus rounded off, base retracted. Reflexed lateral margin slightly crenulate behind humerus. Row of setigerous punctures in lateral channel regular in size, more distant at middle, with about every fourth seta extraordinarily elongated, scutellar striole distinct; setigerous tubercle at base of first stria distinct, with small but distinct tubercle at base of third interval, basal declivity perpendicularly falling to pedunculus, with isodiametric reticulation. Striae moderately deep, striate-punctate, one to four free at base, five joining humerus, six ending free at humerus, one and two ending free at apex, three and four and five and six joining apically. Intervals moderately convex, interval three with four setigerous punctures joining third stria, interval five joining lateral margin at humerus as short carina, interval seven with step-like carina at apex, eight carinate at apex. Surface of all intervals with distinct isodiametric reticulation already well visible at 25 - fold, slightly less reticulated at disk of first interval. Hind wing fully developed. Lower surface. Proepisternum regularly isodiametrically reticulated, with very fine wrinkles laterally. Sternite of abdomen with regular isodiametric reticulation. Last visible sternite with the two marginal setigerous punctures widely separated, with indistinct projection posteriorly at middle, with narrow marginal furrow at middle (in both sexes). Legs. Protibia moderately slender, with longitudinal reticulation, with sulcus dorsally, with three slender acute spines and indistinct tubercle, movable spur moderately arcuate, basal tarsomere convex, slender at base, carinate, with acute tubercle with three setae. Intermediate tibia with ten tubercles at the lateral side, with slender spur near apex, all setae furnished. Tarsomeres of intermediate and hind leg laterally widened, looking serrate. Male genitalia (Fig. 90). Median lobe moderately arcuate at basal half, more straight in apical half, indistinctly bent to apex. Apex like an elongated cone. Parameres slender, with small apophyses, slightly distorted, with flat hyaline elongation at apex. Female gonocoxites and epipleurite (Fig. 116). Basal gonocoxite with one long, three medium long, and two small setae, apical gonocoxite elongated, slightly curved, oval in cross section, with one long seta at end of basal third, with one small seta basally, with two long setae apically. Epipleurite triangle-like, with two setae arising from flat tubercles. Variation. With exception of measurements, not observed.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97331FF8A53D4A3BAFC21FD52.taxon	distribution	Distribution. Known from Irian Jaya, Indonesia.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97331FF8A53D4A3BAFC21FD52.taxon	diagnosis	Diagnosis. A medium sized fuscous species with sub-cordate to square pronotum, all intervals of the elytron reticulated, and with seven-setose labrum. The most similar species with completely reticulated elytra are C. spatulifigura sp. nov., C. opacielytra sp. nov., and C. dealata antecessor Darlington. The last mentioned one is a member of the vulgivaga species-group due to its falcate gonocoxites. It also differs mainly by the smaller eyes and enlarged genae, coreaceate episternum and the acute tubercle at the front trochanter. Differences in C. opacielytra are mainly the six-setose and pentagonal labrum, and the distinctly peltate shaped pronotum. C. spatulifigura differs by its distinctly closed neck constriction, the slightly excised clypeus, and the smaller body size. Moreover, all of the genitalia are different.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97331FF8A53D4A3BAFC21FD52.taxon	etymology	Etymology. The name is a patronym in honour of Matthias Hartmann (NKME) who provided me with full support during many years and loaned material of this and other species.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97330FF8953D4A03BFD48F996.taxon	description	(Figs 23, 58, 91, 117)	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97330FF8953D4A03BFD48F996.taxon	materials_examined	Type material. Holotype: ♂, with labels and data: yellow, black printed “ BORNEO: SABAH Bingkor N Keningau 400 - 500 m, 19. - 20. XI. 1996 leg. W. SCHAWALLER “ / “ Clivina group helferi nr. wallacei PUTZ. det. Balkenohl, 1998 ” (SMNS). Paratypes: 1 ♀, 1 ♂, same data as holotype; one of the paratypes with additional information “ at light ” (SMNS, CMBB); 1 ♂, “ BORNEO, SABAH, C., Telupid Miruru, 25 km 8. v. 1999. Z. Smrź leg. ” / “ helferi-Gr. ” (CADW); 1 ♂, “ BRUNNEI-Temburong Belalong Field Res. C. Mix. Dipterocarp. For. ~ 100 m, 2 - 8.5.95 Heiss ” (CADW). Remark. In the female paratype, the right front and intermediate leg is missing.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97330FF8953D4A03BFD48F996.taxon	description	Description. Measurements. Body length 4.91 – 5.05 mm (ẍ = 4.99 mm *), width 1.38 – 1.39 mm (ẍ = 1.39 mm *), ratio L / W of pronotum 0.98 – 0.99 (ẍ = 0.98), ratio L / W of elytra 1.91 – 1.95 (ẍ = 1.94 *); (* n = 5). Colour glossy, fuscous. Supraantennal plate and clypeal-wing at margin slightly translucent. Antennae, mouthparts, intermediate and hind legs yellowish. Head about a fifth narrower than pronotum. Clypeus with central part straight, projection of lateral teeth distinct, separated from clypeal-wing by distinct notch; wing rounded, distinctly less high as central part of clypeus, separated from supraantennal plate by obtuse-angled notch, supraantennal plate slightly elongated, with margin convex at middle, clypeus, wing and supraantennal plate reflexed margined. Supraantennal plate extended posteriorly into a blunt keel, separated from wing by small furrow. Clypeus transverse, convex, smooth, clypeal-furrow wide, flat, with small punctures, frons moderately convex, with pore at middle, separated from supraantennal plate and supraorbital carina by flat furrow, with clypeal seta in front; supraorbital carinae moderately wide, blunt, with two supraorbital setigerous punctures at middle of eye and at middle of gena-level. Neck constriction closed, developed as step from frons, with indistinct punctures posteriorly. Eye prominent, gena small. Antenna elongate, reaching just over base of elytron. Labrum excised anteriorly, irregularly reticulated, seven-setose. Lobe of mentum slightly convex laterally, rectangular acuminated at tip, medially straight, with fine isodiametric reticulated, median tooth as high as lobe, wide, hollowed out anteriorly. Pronotum (Fig. 58) with disk indistinctly convex in lateral view, in frontal view moderately convex, flattened at median line. Shape square to sub-cordate, as long as wide. Reflexed lateral margin smooth, widest at beginning of posterior third, nearly straight in anterior half but slightly converging; anterior and posterior angles distinct, posterior one with distinct projecting somewhat rounded tooth; lateral channel narrow, reticulated, continuing up to base as sharp, convex margin. Median line complete, engraved at middle, joining and surpassing anterior transverse line. Anterior transverse line distinct, complete. Disk with few scattered punctures, with transverse wrinkles, with isodiametric reticulation at base, with indistinct impression in basal third, with posterior setigerous puncture situated beside tooth of posterior angle. Lateral margin from posterior angle to flange of base slanted with angle of around 40 °. Basal channel on flange regularly developed, rugose, with sharp carina. Elytron (Fig. 23) with disk flattened in anterior half in lateral view, convex in frontal view. Shape of elytra elongate, around two times longer than wide, with lateral margin nearly straight at middle, slightly diverging posteriorly, maximum width posterior middle. Humerus distinct, rounded. Reflexed lateral margin slightly crenulate behind humerus (50 - fold). Row of setigerous punctures in lateral channel regular in size, slightly more distant at middle, scutellar striole distinct; setigerous tubercle at base of first stria distinct, with distinct tubercle at base of third interval, basal declivity perpendicularly falling to pedunculus, with isodiametric reticulation. Striae moderately deep, nearly impunctate, one to four free at base, five and six joining humerus, one and two ending free at apex, three and four and five and six joining apically. Intervals moderately convex, convexity more distinct laterally, interval three with four setigerous punctures joining third stria, interval five joining lateral margin at humerus as short carina, interval seven with step-like carina at apex, eight carinate at apex. Surface of all intervals with distinct isodiametric reticulation already well visible at 40 - fold. Hind wing fully developed. Lower surface. Proepisternum with fine isodiametric reticulation, with fine transverse wrinkles. Sternite of abdomen with regular isodiametric reticulation. Last visible sternite with the two marginal setigerous punctures widely separated, with slight flattened projection posteriorly at middle, with narrow marginal furrow at middle (in both sexes). Legs. Protibia moderately developed, with longitudinal reticulation, with wide sulcus dorsally, with three apically rounded spines and indistinct tubercle, movable spur moderately arcuate, basal tarsomere slender at base, carinate, with acute tubercle with two setae. Intermediate tibia with eight tubercles at the lateral side, with slender spur near apex, all setae furnished. Tarsomeres of front legs slightly widened. Male genitalia (Fig. 91). Median lobe regularly arcuate in basal half, nearly straight in apical half, slightly bent to apex. Apex formed as elongate spatula. Parameres moderately slender, with small apophyses, slightly distorted, with flat hyaline elongation at apex, with two setae apically. Female gonocoxites and epipleurite (Fig. 117). Basal gonocoxite with apex curved laterally into a tubercle with long seta, medially with two long and two medium sized setae, apical gonocoxite elongated, curved in basal third, oval in cross section, with one small seta in basal third and one long seta at end of apical third, with two long setae apically. Epipleurite trapezoid, with one seta. Variation. On the head and pronotum the density of punctures varies slightly. In one of the paratypes, the labrum is five-setose due to a gap of punctures on the left side. In the specimen from Temburong the reticulation at middle of the elytron is slightly less expressed.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97330FF8953D4A03BFD48F996.taxon	distribution	Distribution. Known from Sabah and Brunei on the island of Borneo.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97330FF8953D4A03BFD48F996.taxon	diagnosis	Diagnosis. A small to medium sized fuscous species with sub-cordate pronotum, all intervals of the elytron reticulated, and with seven-setose labrum. The most similar species is C. hartmanni sp. nov. from Irian Jaya also with isodiametric reticulation on all intervals of the elytron. In C. spatulifigura sp. nov. however, the main differences are as follows: the pronotum is as long as wide, the clypeus with its teeth, wings and notches are distinctly developed, there is a distinct clypeal-furrow between the clypeus and the frons, the neck constriction is closed, and the eyes are more prominent. In addition, the male and female genitalia are different.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97330FF8953D4A03BFD48F996.taxon	etymology	Etymology. The name refers to the form (Latin: forma = form) of the male aedeagus with its distinctly elongated spatulate apex (Latin: spatula = spatula, scoop).	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97333FF8753D4A577FB6DFE4E.taxon	description	(Figs 24, 59, 92, 118)	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97333FF8753D4A577FB6DFE4E.taxon	materials_examined	Type material. Holotype: ♂, with labels and data: white, black printed “ MALAYSIA: Sabah Lahad Datu, Ulu Segama For. Res. Danum Valley Forest Centre 01 ° 57 ’ N 117 ° 48 ’ E, 450 m alt xi. 2005, 1 ° Forest coll. Mann, Slade & Villanueva ” / “ Lowland mixed Dipterocarp frst around buildings at artificial light OUMNH- 2005 - 062 ” / “ Clivina group helferi PUTZ. det. Balkenohl, 2020 ” / “ desiderata ex Oxford ” (OUMNH). Paratypes: 1 ♀, same data as holotype (CMBB); 1 ♂, 3 ♀♀, with labels and data: white, black printed “ BORNEO, Sabah Danum Valley, Rainforest Lodge, 19. - 10. X. 2009, leg. U. + H. J. BREMER ” (SMNS).	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97333FF8753D4A577FB6DFE4E.taxon	description	Description. Measurements. Body length 3.84 – 4.53 mm (ẍ = 4.24 mm *), width 1.11 – 1.28 mm (ẍ = 1.22 mm *), ratio L / W of pronotum 0.86 – 0.88 (ẍ = 0.87), ratio L / W of elytra 1.91 – 1.93 (ẍ = 1.92 *); (* n = 6). Colour glossy, fuscous to piceous. Supraantennal plate, lateral margin of pronotum and elytra slightly translucent. Antennae, mouthparts, intermediate and hind legs yellowish. Head a seventh narrower than pronotum. Clypeus with central part straight, with indistinctly projecting rounded teeth laterally, separated from clypeal-wing by indistinct obtuse notch; wing slightly wider than teeth of clypeus, flatly rounded, not as projecting as clypeal-teeth, separated from supraantennal plate by obtuse-angled slight notch, supraantennal plate with margin convex, clypeus indistinctly reflexed margined. Supraantennal plate extended posteriorly into an obtuse keel, separated from wing by indistinct furrow. Clypeus and frons forming continuous slight convexity, with confluent densely scattered small punctures and isodiametric reticulation, flat clypeal-furrow nearly not traceable. Frons with flat pore at middle, separated from supraantennal plate and supraorbital carina by flat reticulated furrow, with clypeal seta in front; supraorbital carinae moderately developed, with two supraorbital setigerous punctures at middle of eye and posterior gena-level. Neck constriction nearly closed, formed by a slight step from frons and reticulation posteriorly. Eye prominent, gena small but distinct. Antenna elongate, reaching over base of elytron. Labrum slightly excised anteriorly, finely isodiametrically reticulated, seven-setose. Mentum oval acuminated, with isodiametric reticulation, median tooth distinctly lower than lobe, acute at apex. Pronotum (Fig. 59) with disk extraordinary flattened in lateral view, in frontal view laterally moderately convex, depressed at median line. Shape sub-cordate, wider than long. Reflexed lateral margin nearly smooth, with fine crenulation visible at 100 - fold, convex at middle, widest at middle; anterior and posterior angles distinct, posterior one with distinct projecting tooth; lateral channel narrow, continuing up to base as sharp, straight margin. Median line complete, depressed, engraved at middle, joining anterior transverse line. Anterior transverse line distinct, complete. Disk covered with micro and medium sized punctures, with rough transverse wrinkles laterally, with isodiametric reticulation at base, with posterior setigerous puncture removed from margin by diameter of pore. Lateral margin from posterior angle to flange of base slanted with angle of around 45 °. Basal channel on flange somewhat widened, rugose, with sharp carina. Elytron (Fig. 24) with disk distinctly flattened in anterior half in lateral view, moderately convex in frontal view. Shape of elytra elongate, about two times longer than wide, with lateral margin nearly straight at middle, indistinctly diverging, maximum width posterior middle. Humerus rounded off. Base distinctly retracted. Reflexed lateral margin indistinctly crenulate behind humerus (120 - fold). Lateral channel slightly narrower at middle. Row of setigerous punctures in the lateral channel nearly regular in size and distance, scutellar striole distinct; setigerous tubercle at base of first stria distinct, with distinct tubercle at base of third interval, basal declivity perpendicularly falling to pedunculus, with isodiametric reticulation. Striae moderately deep, impunctate, one to four free at base, five and six joining at humerus, one to four ending free at apex, five and six joining weakly apically. Intervals slightly to moderately convex, interval three with four setigerous punctures joining third stria, interval five joining lateral margin at humerus as short carina, interval seven completely flat at apex, eight step-like carinate at apex. Surface of interval one to five at middle of disk smooth and glossy, all other intervals with distinct isodiametric reticulation, apically third of elytron reticulated as well as base. Hind wing fully developed. Lower surface. Proepisternum isodiametrically reticulated, with fine wrinkles. Sternite of abdomen with isodiametric reticulation. Last visible sternite with the two marginal setigerous punctures widely separated, with moderately narrow projection posteriorly at middle where surface is smooth, with narrow marginal furrow at middle (in both sexes). Legs. Protibia moderately slender, with indistinct longitudinal reticulation, with sulcus dorsally, with three slender acute spines and small but distinct tooth, movable spur moderately arcuate, basal tarsomere slender at base, carinate, with acute tubercle with two setae. Intermediate tibia with nine tubercles at the lateral side, with slender spur near apex, all setae furnished. Tarsomeres of front leg somewhat widened. Male genitalia (Fig. 92). Median lobe distinctly arcuate at basal part, slightly arcuate up to apex, in lateral view widened at middle, short acuminated to apex. Basal oroficium not distorted. Parameres slender, with small apophyses, slightly distorted, hyaline at apex, each with one minute setae apically. Female gonocoxites and epipleurite (Fig. 118). Basal gonocoxite with three long setae, apical gonocoxite elongated, oval in cross section, flattened in basal part, with one long seta at base, one long seta at middle, with two very long setae apically. Epipleurite relatively wide, asetose. Variation. In two of the paratypes, the reticulation on the head is not as distinct developed on the top of the supraantennal plates and in the posterior part of the frons. Among the six specimens, the lateral margin of the pronotum varies in its convexity, and the micro and medium sized punctures on the disk vary in number.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97333FF8753D4A577FB6DFE4E.taxon	distribution	Distribution. Known from Sabah on the island of Borneo.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97333FF8753D4A577FB6DFE4E.taxon	diagnosis	Diagnosis. A small sized fuscous to piceous species with sub-cordate pronotum, intervals one to five at middle of the disk smooth and glossy, and with seven-setose labrum. The most similar species are C. circumopaca sp. nov. from Sulawesi and C. circumreticulata sp. nov. from Borneo, both having as well the middle of the elytra smooth and glossy. C. caputopaca sp. nov. differs from both of them mostly by the pronotum with its depressed median line, the elytron with interval seven completely flat and interval eight developed as step-like carina, the slightly developed clypeal-wings and clypeal-notches, the reticulated head, and the smaller size.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97333FF8753D4A577FB6DFE4E.taxon	etymology	Etymology. The name refers to the head (Latin: caput = head) with its intense cover of reticulation giving the surface a dull opaque appearance (Latin: opacus, a, um = darken, dull, like with shadow).	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F9733DFF8653D4A10FFCA3FAB5.taxon	description	(Figs 25, 60, 93, 119)	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F9733DFF8653D4A10FFCA3FAB5.taxon	materials_examined	Type material. Holotype: ♂, with labels and data: white, black printed “ BORNEO – SABAH 1995 Ban jaran Mattland Balu Pungul 25. - 27.5. Ivo Jenis leg. ” / “ Clivina dealata DARL. det. Balkenohl X. 97 ” (CMBB). Paratypes: 5 ♂♂, 3 ♀♀, “ BRUNEI-Temburong Belalong Field Res. C. Mix. Diperocarp For. ~ 100 m, 2 - 8.5.95 Heiss ” (CADW); 1 ♂, 1 ♀, “ BORNEO BRUNEI Temburong Kuala Belalong R. Borcherding leg. ” / “ VI / VII 1995 KBFSC ” / “ Clivina wallacei PUTZ. det. M. Baehr’ 12 ” / “ COLL. WRASE BERLIN ” / “ COLL. WRASE GUSOW-PLATKOW GERM. ” (CDWGP); 1 ♀, “ SARAWAK Kapit dist. Rumah Ugap vill., Sut riv., 3. - 9.3.1994 J. Horák leg. ” (CADW); 1 ♂, “ BORNEO: SARAWAK Nanga Pelagus nr. Kapit 180 - 585 m, 7 - 14. VIII. ‘ 58 ” / “ T. C. Maa Collector ” / “ Clivina (s. str.) helferi group P. Bulirsch det. 2015 ” (NHMUK). Remark. In one of the paratypes (from NHMUK), the right front and hind leg, the tarsomeres of the right intermediate leg, two apical tarsomeres of the left front leg, and two right apical antennomeres are missing. The specimen was pinned and shows considerable damages on the ventral side.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F9733DFF8653D4A10FFCA3FAB5.taxon	description	Description. Measurements. Body length 5.17 – 6.38 mm (ẍ = 5.76 mm *), width 1.5 – 1.82 mm (ẍ = 1.61 mm *), ratio L / W of pronotum 0.86 – 0.91 (ẍ = 0.88 *), ratio L / W of elytra 1.84 – 1.95 (ẍ = 1.91 *); (* n = 10). Colour glossy, piceous. Supraantennal plate slightly translucent. Antennae, mouthparts, intermediate and hind legs yellowish. Head a fifth narrower than pronotum. Clypeus with central part slightly concave, with slightly projecting rounded teeth laterally, separated from clypeal-wing by small notch; wing wider than teeth of clypeus, rounded, not as projecting as clypeal-teeth, separated from supraantennal plate by obtuse-angled notch, supraantennal plate with margin laterally convex, more distinct at middle, clypeus and wing indistinctly reflexed margined. Supraantennal plate extended posteriorly into an obtuse keel, separated from wing by flat furrow. Clypeus moderately convex, with scattered small punctures, clypeal-furrow indistinct, flattened. Frons slightly to moderately convex, with indistinct flat pore at middle, with scattered small punctures, separated from supraantennal plate and supraorbital carina by distinct furrow, with clypeal seta in front; supraorbital carinae moderately developed, with two supraorbital setigerous punctures at middle and end of eye level. Neck constriction nearly closed, formed by a step from frons and punctures posteriorly. Eye well developed, gena small but distinct. Antenna elongate, just reaching up to middle of pedunculus. Labrum excised anteriorly, finely isodiametrically reticulated, seven-setose. Mentum with irregularly askew reticulated surface, lobe slightly hollowed out, laterally reflexed margined, laterally convex, medially straight, apex acute, median tooth acute, hollowed out apically, distinctly lower than lobes, clefts moderately developed. Pronotum (Fig. 60) with disk slightly convex in lateral view, moderately convex in frontal view. Shape square to cordate, wider than long. Reflexed lateral margin smooth, slightly convex, widest at middle; anterior angle distinctly marked, posterior angle indistinct, marked by tooth; lateral channel narrow, indistinctly reticulated, continuing up to base as narrow, indistinctly convex margin. Median line complete, engraved at middle, narrow at extremities, joining anterior transverse line. Anterior transverse line distinct, complete. Disk completely covered with medium sized punctures, with some transverse wrinkles, with isodiametric reticulation at base, with posterior setigerous puncture removed from margin by diameter of pore. Lateral margin from posterior angle to flange of base slanted with angle of around 35 °. Basal channel on flange regularly developed, somewhat rugose, with sharp carina. Elytron (Fig. 25) with disk distinctly flattened in lateral view, moderately convex in frontal view with laterally increasing convexity. Shape of elytra elongate, slender, around two-fifth longer than wide, with lateral margin indistinctly convex at middle but slightly diverging, maximum width posterior middle. Humerus rounded but distinct. Reflexed lateral margin indistinctly crenulate behind humerus (120 - fold). Row of setigerous punctures in the lateral channel regular in size and distance, scutellar striole distinct; setigerous tubercle at base of first stria distinct, with distinct small tubercle at base of third interval, basal declivity perpendicularly falling to pedunculus, with isodiametric reticulation. Striae moderately deep, impunctate, one to four free at base, five and six joining at humerus, one to four ending free at apex, five and six joining apically. Intervals slightly convex, interval three with four setigerous punctures joining third stria, interval five joining lateral margin at humerus as short carina, interval seven with obtuse step at apex, eight carinate at apex. Surface of interval one to four glossy at middle, all other intervals with distinct isodiametric reticulation, apically nearly half of elytron reticulated as well as base. Hind wing fully developed. Lower surface. Proepisternum isodiametrically reticulated, with few fine wrinkles laterally. Sternite of abdomen with isodiametric reticulation. Last visible sternite with the two marginal setigerous punctures widely separated, with moderately wide projection posteriorly at middle where it is transversely flattened, with narrow marginal furrow at middle (in both sexes). Legs. Protibia slender, with fine longitudinal reticulation, with slight sulcus dorsally, with three slender acute spines and obtuse tooth, movable spur acute, distinctly curved, basal tarsomere slender at base, carinate, with acute tubercle and three setae at middle of tarsomere. Intermediate tibia with nine tubercles at the lateral side, with slender spur near apex, all setae furnished. Tarsomeres of front leg widened. Male genitalia (Fig. 93). Median lobe slender, distinctly arcuate at basal part and at middle, nearly straight to apex, apically flattened-oval in dorsal view, duck-bill like in lateral view. Parameres moderately slender, with moderate apophyses, slightly distorted, hyaline apically, each with one apical seta. Female gonocoxites and epipleurite (Fig. 119). Basal gonocoxite with two long and three medium sized setae, apical gonocoxite elongated, nearly cylindrical in cross section, with one long and one short seta at base and one long seta at beginning of apical third, with two very long setae apically. Epipleurite sub-trapezoid shaped, with one long seta laterally. Variation. In one of the paratypes, the lateral margin of the pronotum is more distinctly convex and the tooth of the posterior angle is more distinctly developed.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F9733DFF8653D4A10FFCA3FAB5.taxon	distribution	Distribution. Known from the island of Borneo (Sabah, Sarawak, Brunei).	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F9733DFF8653D4A10FFCA3FAB5.taxon	diagnosis	Diagnosis. A small to medium sized piceous species with square to cordate pronotum and intervals one to four smooth and glossy on the disk of the elytron. The most similar species with smooth and glossy intervals one to four of the elytron and a similar body size is C. circumopaca sp. nov. from Sulawesi. It differs by the straight labrum which is in addition six-setose. Moreover, the pronotum is distinctly trapezoid-shaped and the elytra are more convex.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F9733DFF8653D4A10FFCA3FAB5.taxon	etymology	Etymology. The name refers to the elytra with its reticulation on the intervals basally, apically, and laterally (Latin: circum = surrounded by; reticulum = small sized net).	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F9733CFF8553D4A656FA64F834.taxon	description	(Figs 26, 61, 94, 120)	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F9733CFF8553D4A656FA64F834.taxon	materials_examined	Type material. Holotype: ♂, with labels and data: white, black printed “ C Sulawesi, 50 m, 1999 6 km E Tambarana, 120.28,06 E 1.11,15 S 9. - 11. July Bolm lgt. ” / “ Clivina spec. helferi-group det. Balkenohl 2001 ” (SMNS). Paratypes: 2 ♂♂, 2 ♀♀, same data as holotype (SMNS, CMBB).	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F9733CFF8553D4A656FA64F834.taxon	description	Description. Measurements. Body length 5.21 – 5.38 mm (ẍ = 5.29 mm *), width 1.54 – 1.58 mm (ẍ = 1.56 mm *), ratio L / W of pronotum 0.89 – 0.90 (ẍ = 0.89 *), ratio L / W of elytra 1.79 – 1.89 (ẍ = 1.85 *); (* n = 5). Colour glossy, fuscous. Supraantennal plate and wing of clypeus slightly translucent. Antennae, mouthparts, intermediate and hind legs yellowish. Head about a quarter narrower than pronotum. Clypeus with central part straight, with indistinctly projecting rounded teeth laterally, separated from clypeal-wing by slight notch; wing wider than teeth of clypeus, rounded, not as projecting as clypeal-teeth, separated from supraantennal plate by obtuse-angled notch, supraantennal plate with margin laterally convex, more distinct at middle, clypeus, wing, and supraantennal plate indistinctly reflexed margined. Supraantennal plate extended posteriorly into an obtuse keel, separated from wing by narrow furrow. Clypeus transverse, moderately convex, with scattered small punctures, clypeal-furrow distinct. Frons slightly to moderately convex, with indistinct flat pore at middle, with scattered small punctures, separated from supraantennal plate and supraorbital carina by distinct furrow, with clypeal seta in front; supraorbital carinae moderately developed, with two supraorbital setigerous punctures at middle and end of eye level. Neck constriction closed, formed by a slight step from frons and punctures posteriorly. Eye prominent, gena small but distinct. Antenna elongate, reaching over base of pronotum. Labrum straight anteriorly, finely isodiametrically reticulated, six-setose. Mentum slightly translucent, slightly reflexed margined, with isodiametric reticulation, lobe flat, oval, with apex rectangular, median tooth higher than lobe, hollowed out at tip, clefts relatively small. Pronotum (Fig. 61) with disk slightly convex in lateral view, moderately convex in frontal view. Shape trapezoid to cordate, wider than long. Reflexed lateral margin smooth, nearly straight at middle, widest at posterior third; anterior angle distinct, posterior angle distinct, with distinct laterally projecting tooth; lateral channel moderately narrow, indistinctly reticulated, continuing up to base as sharp, slightly convex margin. Median line complete, engraved at middle, joining anterior transverse line. Anterior transverse line distinct, complete. Disk completely covered with micro-punctures, with few slight transverse wrinkles laterally, with isodiametric reticulation at base, bilaterally with two groups of punctures and impression in basal third, with posterior setigerous puncture removed from margin by diameter of pore. Lateral margin from posterior angle to flange of base slanted with angle of around 30 °. Basal channel on flange regularly developed, somewhat rugose, with sharp carina. Elytron (Fig. 26) with disk flattened in anterior half in lateral view, moderately and regularly convex in frontal view. Shape of elytra elongate, somewhat widened, less than about two-fifth longer than wide, with lateral margin distinctly convex at middle and slightly diverging, maximum width posterior middle. Humerus rounded but seemingly angled. Reflexed lateral margin indistinctly crenulate behind humerus (100 - fold). Lateral channel slightly narrower at middle. Row of setigerous punctures in the lateral channel nearly regular in size and distance, scutellar striole distinct; setigerous tubercle at base of first stria distinct, with distinct tubercle at base of third interval, basal declivity falling convex to pedunculus, with isodiametric reticulation. Striae moderately deep, striate-punctate at middle, punctures less distinct laterally, striae one to four free at base, five and six joining at humerus, one to four ending free at apex, five and six joining apically. Intervals moderately convex, interval three with four setigerous punctures joining third stria, interval five joining lateral margin at humerus as short carina, interval seven with very short obtuse step at apex, eight carinate at apex. Surface of interval one to five at middle of disk smooth and glossy, all other parts with distinct isodiametric reticulation, apically third of elytron reticulated as well as base. Hind wing fully developed. Lower surface. Proepisternum isodiametrically reticulated, with few fine wrinkles laterally, proepisternum with distinct narrow furrow. Sternite of abdomen with shingle-like reticulation. Last visible sternite with the two marginal setigerous punctures widely separated, with moderately wide projection posteriorly at middle where it is slightly hollowed out, with narrow marginal furrow at middle (in both sexes). Legs. Protibia moderately slender, with fine longitudinal reticulation, with slight sulcus dorsally, with three slender acute spines and obtuse tooth, movable spur slightly arcuate, basal tarsomere slender at base, carinate, with acute tubercle with two setae and a small tubercle at middle of tarsomere. Intermediate tibia with nine tubercles at the lateral side, with slender spur near apex, all setae furnished. Tarsomeres of front leg somewhat widened. Male genitalia (Fig. 94). Median lobe moderately arcuate at basal part, straight to apex, in lateral view widened at middle, extended acuminated to apex, base with basal oroficium in lateral view moderately distorted. Parameres robust, with well-developed apophyses, slightly distorted, hyaline at apex, each with two apical setae. Female gonocoxites and epipleurite (Fig. 120). Basal gonocoxite with two long and two short setae, apical gonocoxite elongated, oval in cross section, with one long seta at basal third, with two very long setae apically. Epipleurite shaped like oval lappet, with one long seta. Variation. In two of the paratypes, the supraorbital carinae are somewhat confluent with the frons, and the lateral margin of the pronotum is more or less trapezoid-shaped.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F9733CFF8553D4A656FA64F834.taxon	distribution	Distribution. Known from the type locality Tambarana on the island of Sulawesi.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F9733CFF8553D4A656FA64F834.taxon	diagnosis	Diagnosis. A small to medium sized fuscous species with trapezoid to cordate pronotum and intervals one to five at middle of the disk smooth and glossy. The similar species with smooth and glossy inner intervals of the elytron and a similar body size is C. circumreticulata sp. nov. from Borneo. It differs by the excised labrum which is in addition seven-setose. Moreover, the pronotum is distinctly square to cordate-shaped and the elytra are more slender and subparallel.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F9733CFF8553D4A656FA64F834.taxon	etymology	Etymology. The name refers to the elytra with its opaque appearance due to the reticulation on the lateral intervals, basally, and apically (Latin: circum = surrounded by; opacus, a, um = dull, darken, like with shadow).	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F9733EFF8353D4A3BAFBAAFDA2.taxon	description	(Figs 27, 62, 95)	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F9733EFF8353D4A3BAFBAAFDA2.taxon	materials_examined	Type material. Holotype: ♂, with labels and data: white, black framed, black printed and handwritten in black ink “ Philippinen Ost Luzon Kasiguran ”, backside, handwritten in black ink “ J. Boetcher 9. 1915 ” / white, black printed and handwritten in black ink “ Clivina attenuata- helferi-group 2023 det. M. Balkenohl 2017 ” (MFNB). Remark. The following parts are missing in the holotype: Left terminal antennomere, left hind tibia with tarsomeres, right intermediate tarsomeres two to four.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F9733EFF8353D4A3BAFBAAFDA2.taxon	description	Description. Measurements. Body length 6.91 mm, width 1.88 mm, ratio L / W of pronotum 0.92, ratio L / W of elytra 1.97. Colour glossy, fuscous. Supraantennal plate slightly brightened. Antennae, mouthparts, intermediate and hind legs yellowish. Head a fifth narrower than pronotum. Clypeus gently excised anteriorly, with lateral teeth distinctly projecting; wing of moderate size, not more protruding anteriorly than clypeus, distinctly separated from clypeus by notch; supraantennal plate nearly angled laterally; clypeus distinctly reflexed margined, wing and supraantennal plate finely reflexed margined. Supraantennal plate extended up to mid-eye level, separated from wing by distinct notch, separated from supraorbital furrow by distinct groove. Clypeus transverse, moderately convex, separated from frons by wide flat furrow. Frons moderately convex, smooth, with central pore at middle. Supraantennal plate separated from clypeus and frons by moderately wide furrow continuing posteriorly up to mid-eye level as moderately wide supraorbital furrow. With blunt distinct supraorbital carina at posterior half of eye. With two supraorbital setigerous punctures situated at mid-eye level and posterior gena-level. Neck constriction closed, developed by punctures and distinct step. Eye convex, projecting laterally but not hemispherical. Gena distinct, enclosing eye posteriorly by nearly a quarter. Antenna somewhat elongated, reaching basal carina of pronotum, antennomeres four to ten elongate (L / W around 1.6). Labrum nearly straight, isodiametrically reticulated, seven-setose. Mandible of moderate size. Mentum with lateral lobe oval with acute angles, flattened, isodiametrically reticulated, distinctly convex laterally, straight medially, median tooth spatulate, hollowed out, as high as lateral lobes. Pronotum (Fig. 62) with disk flattened in lateral view, in frontal view moderately convex. Distinctly quadrate, wider than long. Anterior margin slightly undulate. Reflexed lateral margin smooth, straight, widest at middle; anterior and posterior angles distinct, posterior one with sharp tooth. Lateral channel narrow to medium sized, reticulated, basal part of lateral margin from posterior angle to flange of base slightly sinuate, slanted with angle of around 30 °, flange distinct, with deep basal channel and sharp basal carina. Median line moderately narrow, slightly crenulated, joining anterior transverse line and base; anterior transverse line narrow, complete. Surface nearly smooth, with indistinct flat impression basally at each side, base isodiametrically reticulated. Elytron (Fig. 27) with disk flattened to indistinctly concave in anterior two thirds in lateral view, regularly and moderately convex in frontal view. Outline regularly long oval, nearly two times as long as wide, with maximum width behind middle. Humerus rounded, formed by lateral margin which joins middle of stria five laterally. Reflexed lateral margin indistinctly crenulated posterior humerus. Lateral channel moderately wide. Scutellar striole of moderate length, not deep; with distinct setigerous puncture at base of first stria, with distinct tubercle at base of third interval. Base straight. Striae moderately deep, punctate-striate, one to four free at base, five and six joining posterior humerus, seven shortened behind humerus, striae one and two running up to apex, three and four, and five and six joining apically. Intervals moderately convex, eights apically with long narrow carina, seventh with short step-like carina apically. Third interval with four setigerous punctures adjoining third stria. Surface of intervals smooth and glossy on disk, with distinct isodiametric reticulation at base, at apical tip, and on interval seven and eight. Hind wing fully developed. Lower surface. Proepisternum with distinct isodiametric reticulation, with dense transverse wrinkles laterally. Sternites of abdomen with isodiametric reticulation, slightly transverse at middle, with few fine punctures laterally. Abdominal sternum VII with the two anal setigerous punctures widely separated, at middle widely flattened and slightly projecting posteriorly, with narrow marginal furrow at middle. Legs. Protibia with three spines of moderate length and a small one basally, with flat sulcus basally, movable spur nearly straight, surface with longitudinal reticulation. Mesotibia with distinct and moderately long protuberance preapically with acuminate apex and with its seta inserted laterally, with nine setae furnished tubercles above protuberance, with few fine setae at the inner side. Front leg with tarsomeres slightly widened, first tarsomere elongated, slender and with carina baso-laterally, laterally with two tubercles furnished with short setae. Male genitalia (Fig. 95). Median lobe moderately robust, distinctly curved in basal three quarters, straight apically, apex with flattened wide acute-angled lamella. Parameres relatively wide, distorted apically, asetose. Female gonocoxites and epipleurite unknown. Variation. At one side, striae three and four are not joining at apex.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F9733EFF8353D4A3BAFBAAFDA2.taxon	distribution	Distribution. Known from the East of the island of Luzon, Philippines.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F9733EFF8353D4A3BAFBAAFDA2.taxon	diagnosis	Diagnosis. A larger sized fuscous species with square to subcordate pronotum, interval seven and eight of the elytron reticulated, and seven-setose labrum. Beside the unique apical lamella of the aedeagus, it differs mainly from the most and very similar species C. westwoodi Putzeys, C. wallacei Putzeys and C. paris sp. nov. by the seven-setose labrum, the parallel and nearly straight lateral margin of the pronotum, the elongated and narrow carina at the apex of the interval eight, and the distinctly reflexed margined clypeus.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F9733EFF8353D4A3BAFBAAFDA2.taxon	etymology	Etymology. The name is a patronym in honour of Bernd Jäger (MFNB) who supported me over many years by loaning type and other material as well as answering questions to historic specimens.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97339FF8253D4A1A3FCA8FC1E.taxon	description	(Figs 28, 63, 96, 121)	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97339FF8253D4A1A3FCA8FC1E.taxon	materials_examined	Type material. Holotype: ♀, with labels and data: white, black printed “ Dobodura Papua, N. G. Mar-July, 1944 Darlington ” / white, black printed “ Darlington at B. M. 1947 - 48 Notes p. ” / white, handwritten in black ink “ Clivina vigil Darl. ” / red, black printed and handwritten in black ink “ M. C. Z. Type 30 / 59 ” / white, black printed, with black / blue image “ Jan. - Jul. 2005 MCZ Image Database ” / white, black printed, with barcode “ MCZ-ENT 00030159 ” (MCZ). Paratype: 1 ♂, with labels and data: white, black printed “ Dobodura Papua, N. G. Mar-July, 1944 Darlington ” / red, black printed and handwritten in black ink “ paratype Clivina vigil Darl. ” / white, black printed, with barcode “ MCZ-ENT 00535205 ” (MCZ). Additional material. 1 ♀, INDONESIA, Irian Jaya Jayapura district Genyem, 50 m NN, IV. - V. 1999, leg. M. Balke / Sammlung NATURKUNDE-MUSEUM ERFURT / Clivina group helferi det. Balkenohl, 2001 (NKME); 1 ♀, INDONESIEN: 14. - 18.2.95 Samabusa 25 km östl. Nabire COROVA – RIVER leg. Dr. D. DASCHE / COLL. WRASE BERLIN / Clivina group helferi det. Balkenohl, 1999 / COLL. WRASE GUSOW-PLATKOW GERM. (CDWGP). Remark. In the holotype, the three terminal tarsomeres are missing at the intermediate tibia. In this dried specimen, the reproductive tract was partly everted with missing of the two terminal gonocoxites. In the paratype, the left antennae is shortened to six antennomeres due to an anomaly. The species is described and keyed out by Darlington (1962). Compendious description. A medium sized fuscous species with the following measurements: Body length 5.83 – 6.46 mm (ẍ = 6.16 mm *), width 1.65 – 1.83 mm (ẍ = 1.77 mm *), ratio L / W of pronotum 0.91 – 0.98 (ẍ = 0.94 *), ratio L / W of elytra 1.82 – 1.99 (ẍ = 1.94 *); (* n = 4). Head with labrum seven-setose, straight anteriorly. Median part of clypeus separated from clypeal-wing by small obtuse notch, lateral teeth of clypeus indistinctly projecting, clypeal-wing not as far projecting as clypeal-teeth, with convex margin, separated from supraantennal plates by distinct obtuse notch. Eye prominent, gena small but distinct. Clypeo-frontal furrow flat, indistinct; clypeus transverse; neck constriction open, laterally indicated by punctures and reticulation. Frons moderately convex, with pore at middle, with scattered small punctures. Antennae elongate, just reaching over base of pronotum. Pronotum (Fig. 63) cordate, wider than long, slightly convex in frontal and moderately convex in lateral view, with lateral margin convex, widest at middle, anterior angle distinct, posterior angle distinct, with distinct tooth projecting anteriorly, lateral margin microscopically crenate (100 - fold), disk basally with small group of punctures, with isodiametric reticulation laterally and at base, with posterior setigerous puncture moved away from posterior angle by diameter of pore; lateral margin from posterior angle to flange of base convex, slanted with angle of around 30 °, basal channel conspicuously widened, rugose, carina of flange sharp. Proepisternum with isodiametric reticulation, with fine transverse wrinkles, medially with punctures. Elytron (Fig. 28) elongate, flattened in basal half in lateral view, lateral margin slightly convex, subcrenulated posterior humerus, striae one to four ending free at base, interval three with four setigerous punctures; striae nearly impunctate; intervals slightly to moderately convex, intervals seven and eight distinctly isodiametrically reticulated, with weaker reticulation on interval six (see variation), base and apical third isodiametrically reticulated, interval eight carinate in apical third, seven convex apically, striole distinct, setigerous tubercle at base distinct, with distinct tubercle at base of third interval. Abdomen with isodiametric reticulation, transverse at middle, two apical setigerous punctures widely separated. Hind wing fully developed. Legs. Intermediate tibia with ten tubercles with setae above the spur at the lateral side. Male genitalia (Fig. 96). Median lobe moderately arcuate in basal half, nearly straight in apical half, with indistinct angle to apical spatula; in lateral view with widened spatula. Parameres moderately slender, distorted and hyaline in apical quarter, with short seta each. Female gonocoxites and epipleurite (Fig. 121). Basal gonocoxite fragile, hyaline, with two long and three medium sized setae, apical gonocoxite elongated, straight, cylindrical in cross section, with five setae in basal half, with two long setae at apex. Epipleurite trapezoid, with two long and one small setae.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97339FF8253D4A1A3FCA8FC1E.taxon	description	Variation. In the holotype, there are transverse wrinkles on the frons of the head near the pore. In the paratype, the anterior setigerous punctures of the pronotum at the left side is doubled. Among the four specimens, the group of punctures on the pronotum varies distinctly. The reticulation on intervals six and five is more or less fading to a different degree.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97339FF8253D4A1A3FCA8FC1E.taxon	distribution	Distribution. The species occurs in Papua / New Guinea.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97338FF8153D4A0FFFD76F9CE.taxon	description	(Figs 29, 64, 122)	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97338FF8153D4A0FFFD76F9CE.taxon	materials_examined	Type material. Lectotype by present designation: ♀, with labels and data: yellow, handwritten in black ink “ Castanea P. N. Guin. (Stev.) ” / grey, handwritten in black ink and black printed “ Clivina castanea Putz. dét. J. Putzeys ” / white, dark blue framed and printed “ Soc. Ent. Belg. Coll. PUTZEYS ” / white, handwritten in black ink and printed “ Clivina Westwoodi Putz. (N. Guinea) = C. Wallacei Putz. H. E. Andrewes det 1928 ” (RBINS). Remarks. The lectotype is glued at the pedunculus. Putzeys (1863) described a species under the name of Clivina castanea based on two specimens from New Guinea and Sri Lanka. Later, when he found the name castanea was already occupied by Westwood (1837), he renamed it in Clivina westwoodi (Putzeys 1867, p. 109). Andrewes (1926, 1929 a) mentioned, the specimens are deposited in RBINS. They belong to different species, and the specimen from New Guinea belongs to C. wallacei (Putzeys 1867, described by Putzeys p. 127 / see also label quotation of the LT above as well as remark in Andrewes 1926, p. 373). The specimen from New Guinea is available in RBINS. It is not identical with C. wallacei Putzeys, as asserted by Andrewes (1926, p. 373). It remains to be a miracle where the specimen from Sri Lanka is deposited. It is not in RBINS or NHMUK as I could check during personal visits, and could not be located in any of the other institutions mentioned. At this point in time, the specimen from New Guinea seems to be the only type specimen available. Therefore, it has been selected as lectotype and the extended diagnosis is based on this lectotype. In case the material from Sri Lanka will show up by any chance, that specimen has to be reassessed. This designation was done in order to maintain stability of the nomenclature. There are several specimens listed under the redescription of Andrewes (1929 a) as representatives of C. westwoodi Putzeys. These specimens are available in NHMUK. They were investigated and listed here for completeness but not used for the compendious description. They are similar to C. westwoodi but consist of more than one different form. This is true for external characters, the ratios of measurements, and for characters of the genitalia. In the absence of Putzeys’ material from Sri Lanka, I refrain from further taxonomic action. It is the following material: 1 spec., Dehra Dun, U. P., S. Bahadur III. 1909 / For. Res. Inst. Dehra Dun / 204 / H. E. Andrewes Coll. B. M. 1945 - 97. This specimen is a member of the different Clivina lobata - species group. It was also quoted as C. westwoodi by Saha & Biswas (1985). 1 ♂, Belgaum. Kh. / H. E. Andrewes Coll. B. M. 1945 - 97. / Clivina Westwoodi Putz. Compared with type H. E. A.; backside: ‘ Figure another prothorax strongly developed’; 1 ♂, Belgaum, Bombay H. E. Andrewes / H. E. Andrewes Coll. B. M. 1945 - 97. / Clivina Westwoodi Putz. Compared with type H. E. A.; backside: ‘ Only stalk of figure on the prothorax visible’; 1 ♀, Gerrappa L 29 6 07 / T. R. Bell collection B. M. 1934 - 394 / Clivina Westwoodi Putz. H. E. Andrewes det. / CLIVINA westwoodi Putzeys, 1866 P. Bulirsch det. 2012; The quoted specimen from Sri Lanka, Colombo (G. M. Henry) (Andrewes 1929 a) is not available. 1 ♀, JAPAN Kobe Harada 25. VI. ’ 19 / J. E. A. Lewis. 1920 — 105 / 806 / Clivina Westwoodi Putz. H. E. Andrewes det. The specimen from Japan fits with the description of Nakane (1978) of C. westwoodi Putzeys. Compendious description. A medium sized fuscous species with the following measurements: Lectotype: Body length 5.72 mm, width 1.61 mm, ratio L / W of pronotum 1.91, ratio L / W of elytra 1.93. Head with labrum with three setiferous punctures at each side, with a setigerous puncture at middle situated slightly backwards, resulting in seven-setose, straight anteriorly. Median part of clypeus slightly excised, separated from clypeal-wing by distinct notch, lateral teeth of clypeus moderately projecting, rounded, clypeal-wing not as far projecting as clypeal-teeth, rounded, separated from supraantennal plates by distinct notch. Eye prominent, gena small. Clypeo-frontal furrow somewhat rugose, distinct; clypeus transverse; neck constriction open at middle, with indistinct punctures posteriorly. Frons with flat pore at middle, smooth. Antennae subelongate, not reaching base of pronotum. Pronotum (Fig. 64) square to sub-cordate, wider than long, slightly convex in frontal and moderately convex in lateral view, with lateral margin slightly convex, widest at middle, angles distinct, posterior angle blunt, with small rounded tooth, lateral margin nearly smooth, disk with some scattered punctures in basal half, with isodiametric reticulation at base; lateral margin from posterior angle to flange of base slanted with angle of around 35 °, basal channel of flange of regular size, basal carina not sharp. Proepisternum with distinct isodiametric reticulation, with few extraordinary fine transverse wrinkles. Elytron (Fig. 29) elongate, flattened in basal half in lateral view, distinctly convex in frontal view, base straight, lateral margin nearly straight in basal half but slightly diverging, subcrenulated posterior humerus, striae one to four ending free at base, interval three with four setigerous punctures; striae impunctate on disk; intervals moderately convex, interval seven and eight, as well as base and apical third of elytron isodiametrically reticulated, interval eight carinate in apical fifth, seven with shortened step-like carina at apex, striole not deep but well visible, setigerous tubercle at base distinct, with distinct tubercle at base of third interval. Lower surface with abdomen with isodiametric reticulation, more confluent at middle, two apical setigerous punctures widely separated. Hind wing fully developed. Legs with intermediate tibia with nine tubercles with setae above the spur at the lateral side. Male genitalia unknown. Female gonocoxites and epipleurite (Fig. 122). Basal gonocoxite with apex acuminated conically with long seta at apex and second long seta subapically, apical gonocoxite elongated, slightly curved, cylindrical in cross section, with two setae basally, with one long seta at middle, with two long setae apically. Epipleurite sub-trapezoid, laterally with two setae close together.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97338FF8153D4A0FFFD76F9CE.taxon	distribution	Distribution. Known from New Guinea.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F9733BFF9F53D4A58FFE65FE32.taxon	description	(Figs 30, 65, 97, 123)	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F9733BFF9F53D4A58FFE65FE32.taxon	materials_examined	Type material. Lectotype by present designation: ♂, with labels and data: yellow, handwritten in black ink “ C. Wallacei Ptz. Mak. (Wall. Schm.) ” / grey, handwritten in black ink and printed “ Clivina Wallacei Ptz. dét. J. Putzeys ” / white, black framed, red printed “ Syntype ” / white, dark blue framed and printed “ Soc. Ent. Belg. Coll. PUTZEYS ” / white, handwritten in black ink and printed “ Clivina Wallacei Putz. cotype H. E. Andrewes 1928 ” (RBINS). Additional material. 1 ♀, Dorey / Bowring 63.47 * / Clivina Wallacei Putz. Compared with type H. E. A. (NHMUK); 1 ♂, 59.58. Dory, New Guinea. / Clivina Wallacei Putz det. K. Kult 47 (NHMUK); 1 ♂, Bantimoeroeng Haros 4. 30 W. Celebes / Clivina wallacei Putz. H. E. Andrewes det. (CADW); 1 ♂, Celebes Dorey. / Compared with TYPE K. Kult 1946 / C. Wallacei Putz. det. K. Kult 47 / COLLECTIO KAREL KULT COLL. A. DOSTAL, 1999 (CADW); 2 ♂♂, 2 ♀♀, N – SULAWESI Dumoga Bone NP / Doluduo, Licht (16) 22. IV. 1992 leg. Jäch (NHMW); 2 specs, S-Sulawesi Prov., Wasuponda env., 600 m, 2.33.13 S 121.13.05 E, 8. - 9. Jul 2001, Bolm lgt. (SMNS). Remarks. In the lectotype, the right front leg is missing. Putzeys (1867, p. 127) writes at the end of his description [translated from French]: “ M. Schaum found two exemplars in the collection Wallace which he acquired. They came from Makassar (Célèbes). He gave me one. M. de Chaudoir also has two individuals which came from Dorey (Célèbes). ” The specimen from M. Schaum is available in RBINS and served for the designation of the lectotype. The whereabouts of the second specimen is unclear. Inquiry at the MFNB revealed the second specimen is not in MFNB in the Schaum collection (written confirmation by the curator Bernd Jäger). The two specimens Putzeys quoted to be in the collection Chaudoir could not be located while checking the Chaudoir collection during my personal visit at the MNHN in November 2021. Andrewes (1926, p. 373 and 1929 b, p. 355) obviously misinterpreted ‘ Dorey’ as a location in New Guinea. He quotes two specimens to be located in RBINS, including the ‘ type’. The type was located in RBINS and is now designated as lectotype. The second specimen was dirty and after cleaning identified as member of the Clivina lobata - species group. The two other specimens from ‘ Dorey’ Andrewes quoted to be deposited in NHMUK were localized there. Darlington (1962, p. 330, 331, 366) cleared the misinterpretations regarding the geographic position of ‘ Dorey’ also spelled as ‘ Dory’. He thinks, this location is located in Celebes (today Sulawesi) and not in New Guinea. Future collections will confirm if the species occurs outside Sulawesi. The species is described by Putzeys (1867), redescribed by Andrewes (1926), and keyed out with descriptive information by Darlington (1962). Compendious description. A medium sized fuscous species with the following measurements: Lectotype: Body length 5.39 mm, width 1.52 mm, ratio L / W of pronotum 1.93, ratio L / W of elytra 1.94. All material: Body length 5.39 – 5.74 mm (ẍ = 5.54 mm *), width 1.52 – 1.79 mm (ẍ = 1.61 mm *), ratio L / W of pronotum 0.89 – 0.94 (ẍ = 0.92 *), ratio L / W of elytra 1.79 – 1.96 (ẍ = 1.91 *); (* n = 5). Head with labrum seven-setose, straight anteriorly. Median part of clypeus separated from clypeal-wing by distinct notch, lateral teeth of clypeus moderately projecting, rounded, clypeal-wing not as far projecting as clypeal-teeth, separated from supraantennal plate by distinct notch. Eye prominent, gena small but distinct. Clypeo-frontal furrow flattened but distinct; clypeus transverse; neck constriction closed, developed as step with punctures posteriorly. Frons with flat pore at middle, with scattered small punctures. Antennae subelongate, not reaching base of pronotum. Pronotum (Fig. 65) square to sub-cordate, wider than long, slightly convex in frontal and moderately convex in lateral view, with lateral margin slightly convex, widest at middle, angles distinct, posterior angle with tooth somewhat rounded and projecting anteriorly, lateral margin sub-crenate, disk with short impression bilaterally in basal half, with isodiametric reticulation laterally and at base; lateral margin from posterior angle to flange of base slanted with angle of around 30 °, basal channel and carina of flange of regular size. Proepisternum with isodiametric reticulation, laterally with some transverse wrinkles. Elytron (Fig. 30) elongate, flattened in basal half in lateral view, lateral margin slightly convex, subcrenulated in anterior quarter, striae one to four ending free at base, interval three with four setigerous punctures; striae nearly impunctate; intervals slightly to moderately convex, interval seven and eight, and half of interval six as well as base and apical third of elytron isodiametrically reticulated, interval eight carinate in apical fifth, interval seven with shortened step-like carina at apex, striole distinct, setigerous tubercle at base distinct, with distinct tubercle at base of third interval. Abdomen with isodiametric reticulation, transverse at middle, two apical setigerous punctures widely separated. Hind wing fully developed. Legs. Intermediate tibia with nine tubercles with setae above the spur at the lateral side. Male genitalia (Fig. 97). Median lobe relatively short, regularly arcuate in basal half, nearly straight in apical half, apex parable-like acuminated. Parameres moderately slender, distorted and hyaline in apical quarter, with medium long seta each. Female gonocoxites and epipleurite (Fig. 123). Basal gonocoxite with apex acuminated into a tubercle with long seta, with second long seta in apical third, apical gonocoxite elongated, slightly curved, cylindrical in cross section, with two setae basally, with one medium sized seta in basal third and one at middle, with long seta at end of apical third, with two long setae apically. Epipleurite trapezoid, with small seta at each edge.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F9733BFF9F53D4A58FFE65FE32.taxon	description	Variation. In one of the male specimens, the lateral margin of the pronotum is slightly more convex. On the frons of the head the number of scattered punctures vary.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F9733BFF9F53D4A58FFE65FE32.taxon	distribution	Distribution. The species occurs on the island of Sulawesi. The occurrence in New Guinea needs confirmation (see remarks above).	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97325FF9E53D4A2D3FAB3FDA4.taxon	description	(Figs 31, 66, 98, 124)	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97325FF9E53D4A2D3FAB3FDA4.taxon	materials_examined	Type material. Holotype: ♀, with labels and data: white, black printed “ Philippine Is. Coll. Bottcher, B. M. 1929 - 201. ” / “ Philippine Islands Manila 8: VII: 1914 Coll. Bottcher ” / white, handwritten in black ink and printed “ Wallacei ssp. philippinensis Kult det. K. Kult 47 ” / red, black framed, black printed “ TYPE ” (NHMUK). Paratypes: 1 ♂, with labels and data: white, black printed “ Philippine Is. Coll. Bottcher, B. M. 1929 - 201. ” / “ Philippine Islands Manila 8: X: 1914 Coll. Bottcher ” / white, handwritten in black ink and printed “ Wallacei ssp. philippinensis Kult det. K. Kult 47 ” / “ red, black framed, black printed “ PARATYPE ” / circle, yellow framed, black printed “ Paratype ” (NHMUK); 1 spec., same data as holotype but with additional label “ Clivina sp.? det. Ing. Jedlička ” [damaged, glued spec.] (NHMUK); 1 ♂, same data as holotype but “ 2: XI: 1914 ” (NHMUK); 1 spec., white, handwritten in back ink “ Philipp Islands ” / “ Semper ” / black printed “ Fry. Coll. 1905.100. ” / white, circle, yellow framed “ Paratype ” / red, black printed and black framed “ PARATYPE ” / white, handwritten in black ink and black printed “ C. Wallacei s. philippinensis Kt. Det. K. Kult 1948 (NHMUK). Additional material. Luzon: 1 ♀, 1 ♂, Los Baños P. I. Baker / 2569 / Ex Mus. Coll. Agric. Phil Is. / F. van Emden Bequest. B. M. 1960 - 129. / wallacei (or subsp.) / wallacei Putz. / Gesch. 8. 1938 von Andrews / Clivina vulgivaga Boh. H. E. Andrewes det.; the male with additional label: Wallacei ssp. philippinensis Kult det. K. Kult 47 (NHMUK); 1 ♀, Philippine Is. Coll. Bottcher. B. M. 1929 - 201. / Philippine Islands Manila 30. X. 1914 Coll. Bottcher (NHMUK); 3 specs, same data but 8. VII. 1914 (NHMUK); 1 spec., same data but 7. I. 1916 (NHMUK); 2 specs, same data but 20. X. 1914 (NHMUK); 1 spec., same data but with additional label C. Wallacei s. philippinensis Kt. det. K. Kult 1948 (NHMUK); 1 spec., Phil. Isl. / C. Wallacei s. philippinensis Kt. det. K. Kult 1948 (NHMUK); 1 spec., Manila Philipp. / H. E. Andrewes Coll. B. M. 1945 - 97. / Wallacei ssp. Philippinensis Kult det. K. Kult 47 (NHMUK); 1 spec., Philippine Islands Manila 6. II. 1914 Coll. Bottcher / Philippine Is. Coll. Bottcher. B. M. 1929 - 201. / From British Museum Collection / C. Wallacei s. philippinensis Kt. det. K. Kult 1948 / COLLECTIO KAREL KULT COLL. A. DOSTAL, 1999 (CADW); 1 ♀, PHILIP.: Luzon Mountain Prov., Sagada Echo Valley, 23. / 24.2.1999 leg. Schödl (21) (NHMW); 1 ♀, Norzagaray Bulacan, Central Luzon, January 2023, loc. Coll. leg. (CMBB); 1 ♂, Mt. Makiling Luzon Baker / Ex Mus. Coll. Agric. Phil Is. / H. E. Andrewes Coll. B. M. 1945 - 97. (NHMUK). Marinduque: 1 ♂, Philippines Marinduque Dec. 2013 / Isoclivina helferi-gr. (CADW). Mindanao: 1 spec., Davao Mindanao Baker / H. E. Andrewes Coll. B. M. 1945 - 97. (NHMUK); 1 ♀, MINDANAO, 30 km E of MALYBALAY, BUSDI 5. - 9. MAY 1996 BOLM lgt. 1000 m / Clivina group helferi det. Balkenohl, 1999 (SMNS). Leyte: 1 ♂, 4 ♀, PHILIPPINES: LEYTE VISCA N BayBay cultiv. land, 1991 leg. SCHAWALLER & al. / 1.3.1991 / Clivina helferi-group det. Balkenohl VI. 93 (SMNS); 1 spec., same data but sec. forest 100 - 200 m (SMNS); 1 ♀, 1 spec., PHILIPPINES: LEYTE SW Abuyog, 28. II. 1991 river bank leg. SCHAWALLER & al. (SMNS). The subspecies is described by Kult (1951). Additional description. Measurements. Body length 5.29 – 6.05 mm (ẍ = 5.73 mm *), width 1.49 – 1.82 mm (ẍ = 1.66 mm *), ratio L / W of pronotum 0.89 – 0.95 (ẍ = 0.93 *), ratio L / W of elytra 1.84 – 1.95 (ẍ = 1.89 *); (* n = 10). The subspecies seems to be slightly larger than the nominotypical subspecies. It differs mainly in the following characters: colour piceous. Head with neck constriction more distinctly closed by band of punctures, with slight step. Pronotum (Fig. 66) with lateral margin slightly contracted anteriorly, channel wider. Elytron (Fig. 31) with intervals more convex, interval seven at apex almost convex, with very short rim at apex. Male genitalia (Fig. 98). Median lobe of moderate length, arcuate basally, somewhat bulgy in apical half, apex like elongated slight parable. Parameres relatively stable, moderately slender, distorted in apical quarter, with medium long seta each. Female gonocoxites and epipleurite (Fig. 124). Basal gonocoxite with apex regularly acuminated, with two long setae, apical gonocoxite elongated, slightly curved, cylindrical in cross section in apical two thirds, with two medium sized and one small setae basally, with two long setae apically. Epipleurite trapezoid, with two setae.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97325FF9E53D4A2D3FAB3FDA4.taxon	description	Variation. The lateral margin of the pronotum is more or less convex. In some of the specimens including paratypes, one side is straighter whereas the other side is more convex.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97325FF9E53D4A2D3FAB3FDA4.taxon	distribution	Distribution. Known from the following Philippine Islands: Luzon, Marinduque, Mindanao, and Leyte.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97324FF9E53D4A1A6FB72F941.taxon	description	(Figs 32, 67, 99, 125)	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97324FF9E53D4A1A6FB72F941.taxon	materials_examined	Type material. Holotype: ♂, with labels and data: white, handwritten in black ink “ SOLOMON ISLANDS ” [yellow underlined] “ San Cristoval 24: VII: 65 Warahito R. 175 ’ P. J. M. Greenslade. Roy. Soc. Exped. B. M. 1966 - 1 ” / black printed “ at light ” (NHMUK). Paratypes: 1 ♂, same data as holotype (NHMUK); 1 ♀, with labels and data: white, handwritten in black ink “ SOLOMON ISLANDS ” [yellow underlined] “ Guadalcanal Umasani R. 18702 5 - VII- 65 P. Greenslade Roy. Soc. Exped. B. M. 1966 - 1 ” / “ River bank under stones ” (NHMUK); 1 ♂ with labels and data: white, black printed “ SOLOMON ISLANDS: ” [yellow underlined] San Cristoval, S. Wainoni. 25. vii. 65 P. N. Lawrence. B. M. 1966 - 1. ” / “ at light ” / handwritten in black ink and printed “ Clivina wallacei Putz. det. C. Louwerens ” (NHMUK). The subspecies differs from the nominotypical subspecies mainly in the following characters.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97324FF9E53D4A1A6FB72F941.taxon	description	Description. Measurements. Holotype: Body length 5.18 mm, width 1.43 mm; ratio length / width of pronotum 0.96; ratio length / width of elytra 1.92. All material: Body length 4.71 - 5.21 mm (ẍ = 5.02 mm *), width 1.35 – 1.46 mm (ẍ = 1.41 mm *), ratio L / W of pronotum 0.93 – 0.96 (ẍ = 0.94 *), ratio L / W of elytra 1.92 – 1.98 (ẍ = 1.96 *); (* n = 4). Head with clypeo-frontal furrow flat, indistinct, neck constriction more closed by band of irregular punctures. Pronotum (Fig. 67) with lateral margin indistinctly convex, nearly straight, with slight impression basally at level of posterior setigerous puncture. Elytron (Fig. 32) with shape seemingly more slender with lateral margin not as much convex as in the nominative subspecies. Male genitalia (Fig. 99). Median lobe narrowed towards apex, somewhat thickened apically, distorted and fractuate at apex. Female gonocoxites and epipleurite (Fig. 125). Gonocoxite slightly wider, epipleurite with one medium sized setae. Variation not observed.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97324FF9E53D4A1A6FB72F941.taxon	distribution	Distribution. Solomon Islands.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97324FF9E53D4A1A6FB72F941.taxon	diagnosis	Diagnosis. The subspecies differs from the nominotypical subspecies mainly by the more prominent eye, the pronotum which is more slender and shows slight depressions at level of posterior setigerous punctures, and the male aedeagus with narrowed distorted and fractuate apex.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97324FF9E53D4A1A6FB72F941.taxon	etymology	Etymology. The name refers to the Solomon Islands, where the specimens were found.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97324FF9C53D4A402FD3AFD36.taxon	description	(Figs 33, 68, 100, 126)	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97324FF9C53D4A402FD3AFD36.taxon	materials_examined	Type material. Holotype: ♂, with labels and data: white, black printed “ SARAWAK (Borneo), ca 40 km SE KAPIT 03.1994, J. Kodada leg. ” (NHMW). Paratypes: 1 ♂, same data as holotype (CMBB); 1 ♀, with labels and data: white, black printed “ BRUNEI: Temburong Kuala Belalong FSC, 60 - 300 m ” / “ Clivina helferi group P. Bulirsch det. 2013 ” (NHMW). 1 ♀, with labels and data: white, black printed “ BORNEO BRUNEI Temburong Kuala Belalong R. Borcherding leg. ” / “ VI / VII 1995 KBFSC ” / “ Clivina nr. wallacei PUTZ. det. Balkenohl IX. 96 ” / “ COLL. WRASE BERLIN ” / “ COLL. WRASE GUSOW-PLATKOW GERM. ” (CDWGP). Remark. In the female paratype (CDWGP), the right elytron is missing.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97324FF9C53D4A402FD3AFD36.taxon	description	Description. Measurements. Body length 4.03 – 4.54 mm (ẍ = 4.28 mm *), width 1.21 – 1.34 mm (ẍ = 1.27 mm *), ratio L / W of pronotum 0.83 – 0.87 (ẍ = 0.85 *), ratio L / W of elytra 1.85 – 1.89 (ẍ = 1.87 *); (* n = 4). Colour glossy, piceous. Supraantennal plate and clypeal-wing translucent. Antennae, mouthparts, intermediate and hind legs yellowish. Head a sixth narrower than pronotum. Clypeus with central part straight, with slightly projecting teeth laterally, separated from clypeal-wing by obtuse notch; wing well visible, rounded, not as projecting as clypeal-teeth, separated from supraantennal plate by obtuse notch, supraantennal plate with margin laterally obtuse-angular, clypeus, wing, and supraantennal plate indistinctly reflexed margined. Supraantennal plate extended posteriorly into a convex keel, separated from wing by furrow. Clypeus slightly convex, with scattered small punctures, clypeal-furrow indistinct, flattened. Frons relatively wide, slightly convex, with pore at middle, with scattered small punctures, separated from supraantennal plate and supraorbital carina by wide furrow, with clypeal seta in front; supraorbital carinae moderately long, narrow, with two supraorbital setigerous punctures at middle of eye and distinctly posterior end of the gena-level. Neck constriction closed, formed by a step from frons. Eye well developed but somewhat flattened, gena small. Antenna elongate, just reaching up to base of elytron. Labrum excised anteriorly, isodiametrically reticulated, seven-setose. Mentum isodiametrically reticulated, lobe slightly hollowed out, laterally reflexed margined, bilaterally tapered to acute apex, median tooth acute, distinctly lower than lobes, clefts small. Pronotum (Fig. 68) with disk distinctly flattened in lateral view, slightly convex in frontal view. Shape square to cordate, wider than long. Reflexed lateral margin smooth, nearly parallel, indistinctly convex, widest at middle; anterior angle distinctly marked, posterior angle distinctly marked by tooth; lateral channel moderately narrow, isodiametrically reticulated, continuing up to base as narrow, indistinctly convex margin. Median line engraved, complete, joining anterior transverse line. Anterior transverse line distinct, complete. Disk nearly completely covered with medium sized partly connected punctures, with some transverse wrinkles laterally, with isodiametric reticulation laterally and at base, with posterior setigerous puncture removed from margin by diameter of pore. Lateral margin from posterior angle to flange of base slanted with angle of around 40 °. Basal channel on flange regularly developed, rugose, with sharp carina. Elytron (Fig. 33) with disk distinctly flattened in lateral view, slightly to moderately convex in frontal view. Shape of elytra elongate, about two-fifth longer than wide, with lateral margin nearly straight at middle but slightly diverging, maximum width posterior middle. Humerus rounded but distinct. Reflexed lateral margin slightly crenulate behind humerus. Row of setigerous punctures in the lateral channel regular in size, slightly more distant at middle. Scutellar striole distinct; setigerous tubercle at base of first stria distinct, with distinct tubercle at base of third interval, basal declivity perpendicularly falling to pedunculus, with isodiametric reticulation. Striae moderately deep, nearly impunctate, one to four free at base, five and six joining at humerus, one to six ending free at apex. Intervals slightly convex, interval three with four setigerous punctures joining third stria, interval five joining lateral margin at humerus as short carina, interval seven flattened at apex, eight carinate at apex. Surface of interval one to six glossy, interval eight and half of width of interval seven with distinct isodiametric reticulation, apical fifth of elytron isodiametrically reticulated on all intervals. Hind wing fully developed. Lower surface. Proepisternum isodiametrically reticulated, without wrinkles. Sternite of abdomen with isodiametric reticulation, scalier medially. Last visible sternite with the two marginal setigerous punctures widely separated, with slight projection posteriorly at middle where it is flattened, with narrow marginal furrow at middle (in both sexes). Legs. Protibia slender, with longitudinal reticulation, with sulcus dorsally, with three spines, movable spur distinctly curved, basal tarsomere slender at base, carinate, with acute tubercle and two setae at middle of tarsomere. Intermediate tibia with eight tubercles at the lateral side, with slender spur near apex, all setae furnished. Tarsomeres of front leg widened. Male genitalia (Fig. 100). Median lobe distinctly arcuate basal part and at middle, slightly arcuate in apical half, apically regularly acuminate, somewhat flattened. Parameres moderately slender, with small apophysis, slightly distorted, hyaline apically, asetose. Female gonocoxites and epipleurite (Fig. 126). Basal gonocoxite with two long setae, apical gonocoxite elongated, oval in cross section, with one long seta at base and one long seta above middle, with two very long setae apically. Epipleurite triangle-shaped, with one long seta arising from tubercle, with two medium sized setae. Variation. In one of the paratypes, the neck constriction is indistinctly closed at middle. In the paratypes, the striae three and four and five and six of the elytron are more or less indistinctly joining at apex, and the extend of the reticulation on interval seven varies.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97324FF9C53D4A402FD3AFD36.taxon	distribution	Distribution. Known from Sarawak and Brunei on the island of Borneo.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97324FF9C53D4A402FD3AFD36.taxon	diagnosis	Diagnosis. A small sized piceous species with cordate to square pronotum and intervals seven and eight of the elytron reticulated. The most similar species C. bilobata sp. nov. differs by the labrum which is distinctly bilobate and excised, the pronotum with the indistinct posterior angle and lateral teeth, its more convex lateral margin, and the nearly smooth disk. In addition, C. bilobata sp. nov. is a larger species. Certain similarities can also be observed in C. margineserrata sp. nov., but that species shows a straight labrum, a pronotum with lateral margin distinctly crenulated, the lateral channel wide, and a widened basal channel. In addition, all intervals are reticulated.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97324FF9C53D4A402FD3AFD36.taxon	etymology	Etymology. The name refers to the pronotum (Latin: collum = pronotum) with its intense cover of medium sized punctures (Latin: punctatus = with punctures).	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97326FF9B53D4A1D7FED6FBEA.taxon	description	(Figs 34, 69, 101)	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97326FF9B53D4A1D7FED6FBEA.taxon	materials_examined	Type material. Holotype: ♂, with labels and data: beige, handwritten in black ink “ Filipinas Sanchez. ” / white, handwritten in black ink “ no 23 Manila ” / “ white, black framed, black printed “ MNCN _ Ent 392232 ” (MNCN).	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97326FF9B53D4A1D7FED6FBEA.taxon	description	Description. Measurements. Body length 5.67 mm, width 1.65, ratio L / W of pronotum 0.89, ratio L / W of elytra 1.88. Colour glossy, piceous. Antennae, mouthparts, intermediate and hind legs fuscous. Head about a quarter narrower than pronotum. Clypeus with central part straight, with rounded slightly projecting teeth laterally, separated from clypeal-wing by indistinct notch; wing not as projecting as clypeal-teeth, separated from supraantennal plate by obtuse-angled notch, supraantennal plate with margin distinctly convex at middle, slightly convex anteriorly, clypeus, wing, and supraantennal plate slightly reflexed margined. Supraantennal plate extended posteriorly into a widened keel, separated from wing by narrow furrow. Clypeus transverse, convex, clypeal-furrow flattened, wider and more distinct laterally. Frons distinctly convex, with pore at middle, nearly smooth, separated from supraantennal plate and supraorbital carina by distinct furrow, with clypeal seta in front; supraorbital carinae distinct, with two supraorbital setigerous punctures at middle of eye and at posterior end of gena. Neck constriction closed, formed by a conspicuous step from frons, with row of medium sized punctures posteriorly. Eye prominent, gena small. Antenna elongate, reaching up to basal channel of pronotum. Labrum trapezoid, straight anteriorly, isodiametrically reticulated, six-setose. Mentum irregularly reticulated, lobe flat, laterally convex, medially slightly concave, tip angular, median tooth wide, flat, oval at tip, distinctly higher than lobes. Pronotum (Fig. 69) with disk distinctly convex in frontal and moderately convex in lateral view. Shape square to sub-cordate, distinctly contracted anteriorly, looking somewhat trapezoid, distinctly wider than long. Reflexed lateral margin microscopically crenulate (80 - fold), convex, widest at basal third; anterior angle rounded off, posterior angle distinctly marked by tooth, lateral channel moderately wide, isodiametrically reticulated, continuing up to base as narrow, convex margin. Median line engraved, complete, joining anterior transverse line. Anterior transverse line distinct, complete. Disk bilaterally with smaller sized punctures in basal half, medially as circular group, laterally as band continuing anteriorly, with few slight transverse wrinkles laterally, with isodiametric reticulation at base, with impression of posterior setigerous puncture with large diameter, distant from margin by diameter of pore. Lateral margin from posterior angle to flange of base slanted with angle of around 25 °. Basal channel on flange wide, rugose, with narrow, obtuse carina. Elytron Fig. 34) with disk in frontal view moderately and in laterally view slightly convex. Shape of elytra elongate, slightly less than double as long as wide, with lateral margin slightly convex at middle, maximum width at middle. Humerus distinct but rounded. Reflexed lateral margin slightly crenulate in anterior quarter (40 - fold). Row of setigerous punctures in the lateral channel regular in size and distance. Scutellar striole distinct; setigerous tubercle at base of first stria distinct, with tubercle at base of third interval, basal declivity perpendicularly falling to pedunculus, with isodiametric reticulation. Striae moderately deep, impunctate, one to four free at base, five and six joining at humerus, one to four ending free at apex, five and six joining at apex. Intervals moderately convex, interval three with four setigerous punctures joining third stria, interval five joining lateral margin at humerus as short carina, interval carinate apically, seventh at apex with step-like carina. Surface of interval one to seven glossy, interval eight with distinct isodiametric reticulation, apex reticulated by a quarter. Hind wing fully developed. Lower surface. Proepisternum isodiametrically reticulated, with fine transverse wrinkles laterally. Sternite of abdomen laterally with isodiametric reticulation, shingle-like at middle. Last visible sternite with the two marginal setigerous punctures widely separated, with slight projection posteriorly at middle where it is flattened and smooth, with narrow marginal furrow at middle. Legs. Protibia of moderate size, with longitudinal reticulation, with sulcus dorsally, with three spines and obtuse tubercle, basal tarsomere distinctly flattened at base, carinate, laterally with two tubercles and three seta at middle of tarsomere. Intermediate tibia with nine tubercles at the lateral side, with slender spur near apex, all setae furnished. Tarsomeres of front and intermediate leg indistinctly widened. Male genitalia (Fig. 101). Median lobe extraordinary slender, nearly straight, slightly arcuate at apical tip, in lateral view shaped like elongated duckbill. Parameres robust, with distinct apophyses, slightly distorted, each with small seta at apex. Female gonocoxites and epipleurite unknown. Variation. The clypeus shows asymmetric wings. On the pronotum, the puncture pattern is slightly different on the two sides. On the right elytron, striae between intervals six to eight are somewhat confluent due to individual wear.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97326FF9B53D4A1D7FED6FBEA.taxon	distribution	Distribution. Known from the type locality Manila in the Philippines.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97326FF9B53D4A1D7FED6FBEA.taxon	diagnosis	Diagnosis. A medium sized piceous species with square to sub-cordate pronotum and interval eight of the elytron reticulated. It differs from the most similar taxa C. wallacei wallacei Putzeys and C. wallacei philippinensis Kult mainly by the head with the convex frons with distinct step-like vaulted neck constriction, and by the nearly straight and conspicuously slender male aedeagus.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97326FF9B53D4A1D7FED6FBEA.taxon	etymology	Etymology. The name is a patronym in honour of Mercedes Paris (noun in apposition), (MNCN), who made me aware of this material during my visit at the MNCN in spring 2024 and who supported me with this and other contributions.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97321FF9A53D4A7EBFEE9F996.taxon	description	(Figs 35, 70, 102, 127)	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97321FF9A53D4A7EBFEE9F996.taxon	materials_examined	Type material. Holotype: ♂, with labels and data: white, black printed “ Sumatra Palembang M. KNAPPERT. ” / white, black printed “ Ex coll. B. H. Klynstra ” / red, black printed “ Type ” / white, black printed and handwritten in black ink “ Clivina integra Andr. Type H. E. Andrewes det. ” / white, black printed “ H. E. Andrewes Coll. B. M. 1945 - 97. ” (NHMUK). Paratype: 1 ♀, with labels and data: white, black printed “ Sumatra Palembang M. KNAPPERT. ” / white, handwritten in black ink “ integra Andr. ” / red, black printed, black framed “ PARATYPUS ” / white, handwritten in black ink and black printed “ Clivina integra Andr. det. C. J. Louwerens ” / white, black printed “ F. van Emden Bequest. B. M. 1960 - 129. ” / white, handwritten in black ink and black printed “ Gesch. 3. 1954 von Louwerens ” (NHMUK). Remark. The historically designated and available ‘ type material’ of C. integra consists of the ‘ type’ and four ‘ co-types’. The ‘ type’ and one ‘ co-type’ are conspecific, fit with the description, and represent C. integra whereas the other three ‘ co-types’ represent another species (see also the remarks under C. similata sp. nov.). Some of the historic specimens investigated are damaged, mostly in the legs, including the type material. The species is described by Andrewes (1929 b). Additional material. Sumatra: 1 ♂, J. B. CORPORAAL. Sumatra’s O. K. Bandar, Baroe. 850 M 7. 2. 21 / Ex coll. Klynstra / H. E. Andrewes Coll. B. M. 1945 - 97. (NHMUK); 1 ♀, Sumatra: Mjöberg / Tjinta Radja / Ex Mus. Stockholm / coll. Bänninger / Clivina integra Andr. H. E. Andrewes det. (ETHZ); 1 ♂, SUMATRA, ALCEH KETAMBE, KUTACANE 499 m, 7. - 19. 10. 91 leg. W. BARRIOS / Helferi-Gruppe (CADW); 3 ♀♀, Coll. I. R. Sc. N. B. SUMATRA Indrapoura Côte Ouest / Collection E. Rousseau. (RBINS); Nias: 1 ♀, INDONESIA, Nias Island, Südteil LF, 03. III. 1995 leg. U. Buchsbaum / Clivina helferi-group det. Balkenohl IX. 96 (NKME); Malaysia: 1 ♂, MALAYSIA – Pahang Taman Negara NP 12. - 14.7.1993 leg. H. Forster (NHMW); Thailand: S-Thailand, Betong Gunung Cang dun vill. Yala dist., 25.3. - 22.4.1993 J. Horák leg. / VIII helferi Gr. (CADW); Compendious description. A small sized piceous species with the following measurements: Body length 4.10 – 5.32 mm (ẍ = 4.95 mm *), width 1.27 – 1.63 mm (ẍ = 1.48 mm *), ratio L / W of pronotum 0.87 – 0.93 (ẍ = 0.91 *), ratio L / W of elytra 1.77 – 1.87 (ẍ = 1.82 *); (* n = 10). Head with labrum seven-setose, straight anteriorly. Median part of clypeus separated from clypeal-wing by distinct notch, lateral teeth of clypeus moderately projecting, angulate, clypeal-wing not as far projecting as clypeal-teeth. Eye prominent, gena small but distinct. Clypeo-frontal furrow flattened, more distinct laterally. Neck constriction developed as line of impressed punctures, deeper laterally, open at middle. Frons with indistinct impression at middle, covered with small punctures. Pronotum (Fig. 70) glossy on disc, square, seemingly slightly wider than long, flattened at middle and moderately convex in frontal view, flattened in lateral view, with lateral margin slightly and regularly convex, widest at middle, anterior angles somewhat rounded but clearly marked, posterior angles somewhat rounded, clearly less projecting laterally than lateral margin, lateral margin slightly crenate, surface bilaterally with short impression, continued anteriorly as group of punctures, with isodiametric reticulation at base; lateral margin from posterior angle to flange of base slanted with angle of around 35 °, basal channel and carina of flange of regular size. Proepisternum with isodiametric reticulation, latero-anteriorly with some transverse wrinkles. Elytron (Fig. 35) elongate, flattened in lateral view, lateral margin slightly convex, subcrenulated in anterior quarter, striae one to four ending free at base, interval three with four setigerous punctures; striae impunctate; intervals moderately convex, interval eight, half of interval seven as well as base of interval six isodiametrically reticulated, intervals seven and eight carinate in apical fifth, interval three at base with minute tubercle. Abdomen with isodiametric reticulation, transverse at middle, two apical setigerous punctures widely separated. Hind wing fully developed. Intermediate tibia with nine tubercles with setae above the spur at the lateral side. Male genitalia (Fig. 102). Median lobe regularly arcuate in basal half, straight in apical half, with apical lamella slightly bent at apex, base with basal oroficium in lateral view moderately distorted. In lateral view duck-bill like. Parameres slender, spatulate, and hyaline in apical third, asetose (at 500 - fold). Female gonocoxites and epipleurite (Fig. 127). Gonocoxites scapiform, elongated, fragile, less sclerotized, not fused, completely movable; basal gonocoxite 1 widened, with two medium-sized setae, apical gonocoxite 2 slender, conspicuously elongated, cylindrical in apical half, with one fine seta basally and one long setae a third before apex, with two long setae at apex. Epipleurite formed by hyaline velum, furnished with single seta basally.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97321FF9A53D4A7EBFEE9F996.taxon	description	Variation. On the frons of the head, the flattened clypeo-frontal furrow is more or less distinct, and the neck constriction is slightly more complete in the holotype. On the pronotum, the number of punctures vary. On the elytron, the minute tubercle at the base of interval three varies in size. The reticulation on interval seven varies from a small stripe to half of the interval.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97321FF9A53D4A7EBFEE9F996.taxon	distribution	Distribution. Known from the islands of Sumatra and Nias, and from the southern part of the Malay Peninsula.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97320FF9853D4A577FF0BFC5C.taxon	description	(Figs 36, 71, 103, 128)	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97320FF9853D4A577FF0BFC5C.taxon	materials_examined	Type material. Holotype: ♂, with labels and data: white, black printed “ Medan Mjöb. ” / “ Ex Mus. Stockholm ” / “ H. E. Andrewes Coll. B. M. 1945 - 97. ” (NHMUK); Paratypes: 1 ♂, with labels and data: white, black printed “ Dohrn Sumatra Liangagas ” / “ Ex Mus. Stettin ” / “ H. E. Andrewes Coll. B. M. 1945 - 97. ” / “ 370 ” (NHMUK); 1 ♀, with labels and data: white, black printed “ Medan Mjöb. ” / “ Ex Mus. Stockholm ” / “ coll. Bänninger ” / handwritten in black ink and black printed “ Clivina integra And. Cotype H. E. Andrewes 28.10.1929 Co-Type ” (ETHZ).; 1 ♀, “ Sumatra ” / “ Coll. Kraatz ” / “ T. M. ” / “ Ex Deutsch Ent Mus ” / “ H. E. Andrewes Coll. B. M. 1945 - 97. ” / circular, green framed, black printed “ Co-type ” (NHMUK); 1 spec. [received with empty abdomen], with white labels and data written in black ink: “ Perak L. C. ” / “ Doherty ” / “ fossor-group ” / black printed “ Fry Coll. 1905.100. ” / handwritten in black ink and black printed “ C. integra ssp. Andr. det. K. Kult, 1948 ” (NHMUK); 1 ♀, white labels, black printed “ Sumatra ” / “ Coll. Kraatz ” / “ Ex Deutsch Ent Mus ” / circular, green framed, black printed “ Co-type ” / handwritten in grey ink “ Clivina integra Andr. Cotype ” / white, black printed “ COLLECTIO KAREL KULT COLL. A. DOSTAL, 1999 ” / red, black printed “ PARATYPUS Coll. K. KULT Dostal fecit 1999 ” (CADW); 1 ♀, white labels, black printed “ Sumatra Medan Mjöberg ” / “ Ex Mus. Stockholm ” / handwritten in black ink and black printed “ C. integra Andr. det. K. Kult, 1948 ” / white, black printed “ COLLECTIO KAREL KULT COLL. A. DOSTAL, 1999 ” (CADW); 1 ♀, with white labels and data written in black ink: “ Perak L. C. ” / “ Doherty ” / black printed “ Fry Coll. 1905.100. ” / handwritten in black ink and black printed “ C. integra ssp. Andr. det. K. Kult, 1948 ” / white, black printed “ COLLECTIO KAREL KULT COLL. A. DOSTAL, 1999 ” (CADW); Remark. The holotype is a ‘ co-type’ of C. integra designated by Andrewes (1929 b). Two of the other available ‘ co-types’ of C. integra are now representing paratypes of C. neglecta sp. nov. (see also remark under C. integra).	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97320FF9853D4A577FF0BFC5C.taxon	description	Description. Measurements. Body length 4.58 – 5.42 mm (ẍ = 4.96 mm *), width 1.43 – 1.63 mm (ẍ = 1.53 mm *), ratio L / W of pronotum 0.81 – 0.87 (ẍ = 0.86 *), ratio L / W of elytra 1.77 – 1.86 (ẍ = 1.81 *); (* n = 8). Colour glossy, piceous. Supraantennal plate slightly brightened. Antennae, mouthparts, intermediate and hind legs yellowish. Head a quarter narrower than pronotum. Clypeus straight to indistinctly excised anteriorly, with its lateral teeth rounded and slightly projecting; wing of moderate size, rounded, less protruding than clypeus, distinctly separated from clypeus by notch; supraantennal plate convex laterally, slightly convex in anterior half; clypeus, wing, and supraantennal plate reflexed margined. Supraantennal plate extended up to mid-eye level, ending at mid-eye level as blunt knob, separated from wing by distinct notch, separated from supraorbital carina by deep moderately wide furrow. Clypeus with transverse blunt elevation, separated from frons by flat but distinct furrow. Frons moderately convex, with indistinct pore at middle, with scattered small punctures. Supraantennal plate separated from clypeus and frons by moderately wide furrow continuing posteriorly up to mid-eye level as moderately wide supraorbital furrow. With blunt distinct supraorbital carina at posterior half of eye. With two supraorbital setigerous punctures situated at mid-eye level and posterior gena-level. Neck constriction developed by distinct step, punctured posteriorly. Eye prominent, projecting laterally, sub-hemispherical. Gena distinct, enclosing eye posteriorly as small stripe. Antenna moderately long, just reaching base of elytron, antennomeres four to ten elongate (L / W 1.43). Labrum nearly straight, isodiametrically reticulated, seven-setose. Mandible of moderate size. Mentum completely covered with isodiametric reticulation, lateral lobe distinctly convex laterally, angulate anteriorly, straight medially, median tooth wide, angulate, as high as lateral lobes. Pronotum (Fig. 71) with disk indistinctly convex in lateral view, in frontal view flattened at middle and moderately convex laterally. Trapezoid, wider than long. Anterior margin indistinctly excised. Reflexed lateral margin smooth, convex and slightly widening posteriorly, widest at posterior third; anterior angle rounded-off; posterior angle distinctly marked by moderately sharp tooth. Lateral channel relatively narrow, reticulated, margin from posterior angle to base slanted by angle of about 40 °, indistinctly convex, flange short but distinct, basal channel and carina of flange of regular size. Median line distinct, crenulated, joining and surpassing anterior transverse line, joining base; anterior transverse line narrow, complete. Surface with microscopic punctures, basal declivity isodiametrically reticulated. Elytron (Fig. 36) with disk indistinctly convex to flatten in anterior half in lateral view, regularly convex in frontal view. Outline regularly long oval, distinctly less than twice as long as wide, with maximum width behind middle. Humerus distinct, rounded, formed by lateral margin which joins stria five laterally by a short carina. Reflexed lateral margin subcrenulated posterior humerus. Lateral channel moderately wide. Scutellar striole of moderate length, not deep; with distinct setigerous puncture at base of first stria, with small but distinct tubercle at base of third interval. Striae moderately deep, indistinctly punctate, one to four free at base, five and six joining at humerus, striae one and two running up to apex, three and four, and five and six joining apically. Intervals convex, seven and eight carinate apically. Third interval with four setigerous punctures adjoining third stria. Surface of intervals smooth and glossy on disk, with distinct isodiametric reticulation at basal declivity, at apical fifth, and on interval seven and eight. Hind wing fully developed. Lower surface. Proepisternum with distinct isodiametric reticulation, with transverse wrinkles laterally. Sternites of abdomen with isodiametric reticulation laterally, shingle-like and transverse at middle. Abdominal sternum VII with the two anal setigerous punctures widely separated, at middle slightly projecting posteriorly where it is flattened, with narrow marginal furrow at middle (in both sexes). Legs. Protibia with three spines of moderate length and a small knob basally, widely sulcate dorsally, movable spur slightly arcuate, surface with longitudinal reticulation. Mesotibia with distinct and moderately long protuberance preapically with acuminate apex and with its seta inserted laterally, with seven setae furnished tubercles above protuberance, with few fine setae at the inner side. Front leg with tarsomeres of about same width as other tarsomeres, first tarsomere elongated, with carina baso-laterally, laterally with acute tubercle furnished with two short setae. Male genitalia (Fig. 103). Median lobe regularly arcuate in basal half, in apical half nearly straight for a short distance, with apical lamella straight to apex, base with basal oroficium in lateral view at axis of median lobe. In lateral view spoon-like widened. Parameres slender, spatulate, and hyaline in apical third, asetose (at 500 - fold). Female gonocoxites and epipleurite (Fig. 128). Gonocoxites scapiform, extraordinary slender, less sclerotized, not fused, completely movable; basal gonocoxite 1 elongated, with four setae, apical gonocoxite 2 slender, distinctly elongated, straight, narrowed in basal half, cylindrical in apical half, with two long setae at apex, with one long setae at lateral margin in apical third, with one minute seta. Epipleurite formed by hyaline velum, furnished with two setae close together basally. Variation. On the frons of the head, the central impression varies in size and depth, and the small punctuation varies from nearly smooth to slightly punctured. On the elytron, the punctuation of the striae is more or less distinct.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97320FF9853D4A577FF0BFC5C.taxon	distribution	Distribution. Known from the island of Sumatra, and from the southern part of the Malay Peninsula.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97320FF9853D4A577FF0BFC5C.taxon	diagnosis	Diagnosis. A small sized piceous species with interval eight of the elytron reticulated. It differs mainly from the most similar species C. integra Andrewes and C. neglecta sp. nov. by its trapezoid shape of the pronotum, the distinct neck constriction, and the median lobe of the male aedeagus in dorsal view with apical lamella straight to apex and in lateral view spoon-like widened.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97320FF9853D4A577FF0BFC5C.taxon	etymology	Etymology. The name refers to the close similarity of the species to that of C. integra Andrewes and C. neglecta sp. nov.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97322FF9153D4A739FE48F86D.taxon	description	(Figs 37, 72, 104)	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97322FF9153D4A739FE48F86D.taxon	materials_examined	Type material. Holotype: ♂, with labels and data: white, black printed “ Sumatra Mjöberg ” / “ Kota Tjane ” / “ F. van Emden Bequest. B. M. 1960 - 129. ” / handwritten in black ink and black printed “ Gesch. 8. 1938 von Andrewes ” / “ Clivina integra Andr. H. E. Andrewes det. (NHMUK). Remark. The specimen is slightly abraded. Five terminal antennomeres at the left side and some frontal tarsomeres are missing, and the right front femur is broken. At the left hind leg, the three terminal tarsomeres are missing.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97322FF9153D4A739FE48F86D.taxon	description	Description. Measurements. Body length 4.44 mm, width 1.37 mm, ratio L / W of pronotum 0.91, ratio L / W of elytra 1.82. Colour glossy, ferruginous. Supraantennal plate slightly brightened. Antennae, mouthparts and legs yellowish. Head a quarter narrower than pronotum. Clypeus straight anteriorly, with its lateral teeth rounded and indistinctly projecting; wing not prominent, rounded, slightly less protruding than clypeus, separated from clypeus by small notch; supraantennal plate convex laterally, straight in anterior half; clypeus reflexed margined. Supraantennal plate extended up to mid-eye level, ending at mid-eye level as blunt knob, separated from wing by indistinct flat notch, separated from supraorbital carina by deep moderately wide furrow. Clypeus with transverse blunt horseshoe-like elevation, separated from frons by flat but distinct furrow. Frons moderately convex, smooth. Supraantennal plate separated from clypeus and frons by moderately wide furrow continuing posteriorly up to mid-eye level as moderately wide supraorbital furrow. With blunt distinct supraorbital carina at posterior half of eye. With two supraorbital setigerous punctures situated at mid-eye level and posterior gena-level. Neck constriction developed as step, distinct laterally, indistinct and flat at middle, laterally punctured posteriorly. Eye convex but somewhat flattened. Gena distinct, enclosing eye posteriorly as small stripe. Antenna somewhat shortened in comparison to body size, just reaching over posterior setigerous punctures of pronotum, antennomeres four to ten elongate (L / W around 1.88). Labrum slightly excised, isodiametrically reticulated, seven-setose. Mandible of moderate size. Mentum at middle with isodiametric reticulation, lateral lobe with fine coreaceate-like structure, with oblique carinae, lateral margin nearly straight anteriorly, obtuse angulated anteriorly, convex medially, median tooth widened apically, slightly higher as lateral lobes. Pronotum (Fig. 72) with disk slightly convex in lateral view, in frontal view moderately convex at middle and distinctly laterally. Subsquare, wider than long, resembling to sub-globosity in dorsal view. Anterior margin excised. Reflexed lateral margin nearly smooth, with indistinct microscopic notches, distinctly convex, widest at middle; anterior and posterior angles distinctly rounded-off; tooth at posterior angle nearly integrated in rounding of posterior angle. Lateral channel relatively narrow, reticulated, margin from posterior angle to base slanted by angle of about 45 °, indistinctly convex, flange short but distinct, basal channel and carina of flange of regular size. Median line distinct, crenulated, joining anterior transverse line, joining base; anterior transverse line complete. Surface with short impression bilaterally, with few punctures in basal half, basal declivity isodiametrically reticulated. Elytron (Fig. 37) with disk flattened in anterior half in lateral view, regularly convex in frontal view. Outline long oval, distinctly less than twice as long as wide, with maximum width behind middle. Humerus distinctly marked, formed by lateral margin which joins stria five laterally by a very short carina. Reflexed lateral margin nearly smooth posterior humerus. Lateral channel moderately wide. Scutellar striole of moderate length, not deep; with distinct setigerous puncture at base of first stria, with minute tubercle at base of third interval. Striae moderately deep, indistinctly punctate, one to four free at base, five and six joining at humerus, striae one and two running up to apex, three and four, and five and six joining apically. Intervals moderately convex, eights flattened, seven and eight carinate apically. Third interval with four setigerous punctures adjoining third stria. Surface of intervals smooth and glossy on disk, with distinct isodiametric reticulation at basal declivity, at apical fifth, and on interval eight. Hind wing fully developed. Lower surface. Proepisternum with distinct isodiametric reticulation, with transverse wrinkles laterally in posterior part. Sternites of abdomen with isodiametric reticulation laterally, shingle-like and transverse at middle. Abdominal sternum VII with the two anal setigerous punctures widely separated, at middle slightly projecting posteriorly where it is flattened, with narrow marginal furrow at middle. Legs. Protibia with three spines of moderate length and a small knob basally, slightly sulcate dorsally, movable spur wide, slightly arcuate, surface with longitudinal reticulation. Mesotibia with distinct and moderately long protuberance preapically with acuminate apex and with its seta inserted laterally, with seven setae furnished tubercles above protuberance, with few fine setae at the inner side. Front leg with tarsomeres of about same width as other tarsomeres, first tarsomere elongated, with carina baso-laterally, laterally with acute tubercle furnished with two short setae. Male genitalia (Fig. 104). Median lobe arcuate in basal third, straight at middle, conspicuously arcuate in apical fifth including lamella, base with basal oroficium in lateral view at axis of median lobe. Lamella in lateral view duck-bill shaped. Parameres slender apically, spatulate, and hyaline in apical third, asetose (at 500 - fold). Female gonocoxites and epipleurite unknown. Variation unknown.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97322FF9153D4A739FE48F86D.taxon	distribution	Distribution. Known from the type locality Kota Tjane in the North of Sumatra.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97322FF9153D4A739FE48F86D.taxon	diagnosis	Diagnosis. A small sized ferruginous species with interval eight of the elytron flattened and reticulated. It differs mainly from the most similar species C. integra Andrewes and C. similata sp. nov. by its subsquare shape of the pronotum with all angles distinctly rounded, the flattened eyes, the indistinct flattened neck constriction, and the median lobe of the male aedeagus in dorsal view which is conspicuously arcuate in apical fifth including the lamella, and the duck-bill shaped lamella in lateral view.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
03F487F97322FF9153D4A739FE48F86D.taxon	etymology	Etymology. The name is derived from the Latin expression ‘ neglectus est’ which means ‘ the overlooked one’, because the specimen of this species was sorted under type and other material of C. integra Andrewes. Specimen incertae sedis The following specimen could not be interpreted by its unclear character states, and severe damages: 1 spec., Sumatra / Coll. Kraatz / Andrewes det. / Clivina integra Andr. redet. K. Kult 44 / COLLECTIO KAREL KULT COLL. A. DOSTAL, 1999 (CADW). The specimen is glued at several parts, the right front leg is missing, and it was received with an empty hind body. Therefore, information on the genitalia is completely missing. It is not conspecific with the type and other material of C. integra. The data of this specimen have not been included under the determined material. Concluding remarks: On the basis of the limited material investigated, taxa of the Clivina westwoodi - species group seem to be mainly distributed in the eastern part of the Oriental region including New Guinea / Papua. It looks like that in the many islands a rich Clivinini fauna has been evolved with a high degree of endemism on some of the islands like for example in Borneo, Sulawesi, and New Guinea / Papua. Such high degree of endemism in Carabidae was also reported by Liebherr (2017) for the genus Mecyclothorax Sharp in New Guinea / Papua. This hypothesis is supported by the ascertainment that the species are not reported for Australia in the recently published profound revisions of the genus Clivina and allied genera by Baehr (2008, 2015, 2017) and are obviously not occurring there according to current knowledge. At the first glance, the number of 637 specimens totally investigated is seemingly high. In reality, it is a small cut-out only. Many of the islands are only explored by random sampling, if at all, e. g. the Philippine Island with its about 7000 islands, Sulawesi, or New Guinea / Papua. Often, the samplings exist of few specimens, sometimes only one specimen of a species was found. Moreover, the morphological differences are in some cases not large respectively the characters and combination of characters were not known in this specificity from other regions. So, differential diagnoses based on the habitus only reach limits. Fortunately, characters of the male and female genitalia are distinct in all the cases evaluated. A good example is the pair C. westwoodi and C. wallacei wallacei which are similar but show a different external female reproductive tract. Taking the explanations regarding the collection locality and the assumption of Darlington (1962) into account, the hypothesis that C. wallacei wallacei occurs in Sulawesi and not in New Guinea / Papua might be probably true. The new finds on Sulawesi support this. The lectotype of C. westwoodi from New Guinea is the only specimen actually available and needs such confirmation as well. Therefore, the revision is seen as an initial stage, and exact conclusions on the distribution cannot be drawn at the current point in time.	en	Balkenohl, Michael (2025): On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini). Zootaxa 5592 (1): 1-70, DOI: 10.11646/zootaxa.5592.1.1, URL: https://doi.org/10.11646/zootaxa.5592.1.1
