taxonID	type	description	language	source
03F1AD5D4C79FFE2FF40FA9BFAEBF9C1.taxon	materials_examined	Type species. Praeanchodemus punctaticeps Sp. n., foSSil SpecieS from the Eocene Baltic amber.	en	Will, Kipling (2017): Description of the first flightless platynine ground beetle preserved in Baltic amber (Coleoptera: Carabidae). Zootaxa 4318 (1): 110-122, DOI: 10.11646/zootaxa.4318.1.4
03F1AD5D4C79FFE2FF40FA9BFAEBF9C1.taxon	description	Description. AS the new genuS iS monotypic and baSed on a Single Specimen, neither the interSpecific nor intraSpecific variability of characterS iS known. Herein, we deScribe the new taxon in detail under the genuS deScription and only include colour and dimenSionS of the body partS in the SpecieS’ deScription below. Body length: Rather Small compared with average in Platynini: 5.4 mm in type Specimen. MicroSculpture: On head markedly contraSting with microSculpture patternS on pronotum and elytra: DorSal Surface of head with large and deeply engraved lineS forming iSodiametric SculpticellS; pronotum and elytra with much Smaller SculpticellS that are diStinct only on SideS, near baSe of pronotum, and within elytral Striae; pronotal diSc and elytral intervalS without viSible microSculpture even at magnification 100 x. Head: Size and proportionS averaged for Platynini, in dorSal view not conStricted towardS neck. DiSc convex, throughout with deep coarSe punctureS (FigS 1, 5, 18). MandibleS (FigS 6 – 7) Slender, with Scrobe aSetoSe. Labrum 6 - SetoSe, clypeuS with a Single Seta each Side. EyeS moderately large, moderately protruded (Fig. 5); templeS rather long (Fig. 15), 0.3 timeS of eye length, weakly wrinkled to neck; Supraorbital furrow indiStinct, two Supraorbital Setae preSent (Fig. 5). Apical tooth of mentum Simple; Setae near mentum tooth Small, inSerted very cloSe together, immediately baSad of mentum tooth, Submentum with two pairS of lateral Setae, outer pair diStinctly Smaller (FigS 6 – 7). StipeS with a large Seta externally near baSe; penultimate article of maxillary palpuS medially biSetoSe; ligular Sclerite with two Setae near apical margin. Antennae Slender, pubeScent from fourth antennomere onwardS, Scape not markedly enlarged, Same length aS third antennomere, with one Seta on dorSal Side; pedicel 0.6 timeS aS long aS Scape, with a primary apical Seta on ventral Side and a finer apical Seta each on dorSal and ventral SideS; third antennomere without Setae in addition to normal ring of apical Setae. Prothorax: In dorSal view Subcordate-Subquadrate, moderately tranSverSe, broadeSt in middle, with baSal margin Slightly broader than apical margin; SideS evenly rounded in anterior 7 / 8 and Slightly concave near laterobaSal angleS (Fig. 5). Anterior margin moderately concave with front angleS diStinctly protruded, Shortly rounded; baSal bead fine, interrupted in middle. PoSterior margin Straight in middle, with external Sixth markedly bent anteriad; laterobaSal angleS markedly Shifted anteriad, obtuSe, not protruded laterally; baSal bead broadly interrupted in middle. DiSc convex, Slightly but diStinctly rugoSe; baSe coarSely rugoSe, without punctureS. Median longitudinal impreSSion moderately deep in middle, diSappearing near apex and baSe; both anterior and poSterior tranSverSe impreSSionS indiStinct. Lateral gutter narrow throughout, very Slightly widened towardS baSe; laterobaSal depreSSionS rather Shallow. Both lateral and laterobaSal Setae preSent, with laterobaSal Setae located beSide lateral gutter at anterior 1 / 3 of pronotal length, and laterobaSal Setae located on margin (Fig. 5). ProSternum and proepiSternum Smooth, proSternal proceSS not marginated (Fig. 12); anterior coxal cavitieS cloSed poSteriorly. Pterothorax: Elytra markedly convex on diSc (Fig. 16), flattened towardS baSe, elongate-ovate in dorSal view, with ShoulderS moderately broad, humeral angleS and apex Shortly rounded (FigS 10 – 11). BaSal margin concave, protruded towardS Scutellum and humeruS aS well. All Striae deeply impreSSed, impunctate (Fig. 10); paraScutellar Stria moderately long, not connected to firSt Stria; firSt Stria complete to baSe; intervalS moderately convex; fifth interval not impreSSed at apex. ParaScutellar Seta at baSe of firSt Striae, three diScal Seta on third interval, and Subapical Seta near end of Seventh Stria preSent (FigS 9, 11), with anterior diScal Seta located between anterior fifth to Sixth of elytral length, adjoined to third Stria, middle Seta located Slightly behind elytral middle, adjoined to third Stria, and poSterior Seta located at poSterior elytral Seventh, adjoined to Second Stria; diScal Setae not Set in foSSae. Umbilicate SerieS conSiStS of 15 Setae adjoining eighth Stria, with diStance between 6 th and 7 th Seta Slightly larger (Fig. 16). MetepiSternum Smooth, Short, with outer margin Slightly longer than anterior margin (FigS 12, 16). Hind wingS reduced to Short StubS. LegS: Rather Short. All trochanterS with a Single Seta on external Surface. MeSocoxa with a Single ridge Seta (Fig. 12); metacoxa biSetoSe, without Seta near internal margin (Fig. 13); anterior metacoxal SulcuS Straight. Profemur with a Single Seta on ventral Surface and with two rowS of Six Short Setae on anterior Surface; meSofemur with two Setae on ventral Surface and with two rowS of nine Short Setae on anterior Surface; metafemur with two Setae on ventral Surface (Fig. 13), with dorSal Surface Smooth even near apex. Protibia not modified, with external Surface Smooth. FirSt and Second tarSomereS each with a pair of dorSoapical Setae, latter lacking on diStal tarSomereS; fourth tarSomereS of all legS Simple, with ventro-apical lobeS very Short, with pair of lateroapical Setae preSent and without dorSoapical Setae; ventral SurfaceS of fifth tarSomereS Smooth; clawS Simple. PreSence of lateral longitudinal grooveS on dorSal SurfaceS of tarSomereS could not be confirmed. Abdomen: Abdominal SterniteS Smooth beSide normal Setation; V – VI with one, VII with two (female) pairS of Setae near apical margin (Fig. 14). Female genitalia: Form of gonocoxiteS aS typical of moSt Platynini (Fig. 17); baSal gonocoxiteS moderately Stout, apical gonocoxiteS Slender cone-Shaped, very Slightly bent outwardly, with Subapical SetoSe organ preSent; chaetotaxy and internal genital tract could not be Seen nor viSualized.	en	Will, Kipling (2017): Description of the first flightless platynine ground beetle preserved in Baltic amber (Coleoptera: Carabidae). Zootaxa 4318 (1): 110-122, DOI: 10.11646/zootaxa.4318.1.4
03F1AD5D4C79FFE2FF40FA9BFAEBF9C1.taxon	etymology	Etymology. A compound word, baSed on the Latin prefix “ prae- “ and the name of the platynine taxon Anchodemus MotSchulSky, 1864. The name of the new genuS particularly referS to the externally Similar taxon Paranchodemus Habu, 1978, which waS deScribed aS SubgenuS of Anchodemus and elevated to genuS level by Liebherr (1989 a).	en	Will, Kipling (2017): Description of the first flightless platynine ground beetle preserved in Baltic amber (Coleoptera: Carabidae). Zootaxa 4318 (1): 110-122, DOI: 10.11646/zootaxa.4318.1.4
03F1AD5D4C79FFE2FF40FA9BFAEBF9C1.taxon	diagnosis	Differential diagnosis and relationships. A Small repreSentative of Platynini with rather Short legS, which iS eaSily diStinguiShed from all other genera of that tribe by the following combination of external characterS: 1) Setae near mentum tooth inSerted cloSe together, immediately baSad of mentum tooth. 2) DiSc of head throughout with deep coarSe punctureS. 3) Head lacking diStinct neck conStriction. 4) MicroSculpture patternS markedly contraSting on different body partS, with large and deeply engraved lineS forming iSodiametric SculpticellS on head and very indiStinct SculpticellS on pronotal and elytral diScS. 5) Pronotal lateral gutter narrow throughout, laterobaSal depreSSionS Shallow. 6) Metacoxa biSetoSe, without Seta near internal margin. 7) Profemur with a Single Seta, meSofemur with only two Setae on ventral Surface. 8) Metafemur dorSally without apical Setae. 9) All legS with fourth tarSomere dorSoapical Setae lacking. 10) All legS with fifth tarSomere Smooth on ventral Surface. The poSition of mentum Setae iS conSidered an important indication for SpecieS-group relationShipS (Schmidt 2001 a). By far, moSt lineageS of platynine ground beetleS have a Single pair of Setae inSerted caudally of the deep, apical inciSionS of the mentum, laterally of the apical tooth, and thuS, the inSertion pointS of the mentum Setae are located rather diStant from each other (See Schmidt 2001 a: FigS 12, 14, 16). In contraSt, the character State obServed in the foSSil Praeanchodemus gen. n. (character State 1) iS preSent in only Six terminal Platynini lineageS, and iS conSidered apomorphic. However, given the current State of knowledge, there iS no evidence for cloSe relationShipS of theSe lineageS (Schmidt 2000, 2001 a). It iS thuS likely that the adjacent poSition of the mentum Setae evolved within the tribe two or more timeS independently. In the following, the Six relevant lineageS are diScuSSed in more detail and compared with the new foSSil taxon.	en	Will, Kipling (2017): Description of the first flightless platynine ground beetle preserved in Baltic amber (Coleoptera: Carabidae). Zootaxa 4318 (1): 110-122, DOI: 10.11646/zootaxa.4318.1.4
03F1AD5D4C7CFFE3FF40F9F1FEE1FD03.taxon	materials_examined	Holotype. Female in Baltic amber; Size of amber piece 14.0 x 7.0 x 6.5 mm, with collection number BB- 1752 - K in StaatlicheS MuSeum für Naturkunde, Stuttgart, Germany (Fig. 4). Preservation status. The amber piece iS clear and in a good conServation State, however, the embedded Platynini foSSil iS only partly viSible uSing light microScopy due to Several flowlineS adjoining the beetle body (FigS 1 - 3). Six (left antenna) reSpectively three (right antenna) apical antennomereS are loSt. The hind coxa iS covered by milky coating. The left elytron appearS artificially inflated. The foSSilized exoSkeleton of the Specimen iS in partS markedly Shrunken and thuS diSSociated from the incluSion wall (FigS. 8 - 11). However, moSt detailS of diagnoStic importance of the external morphology could be reconStructed baSed on both, the remainS of the foSSilized exoSkeleton and the negative imprint of the beetle body on the incluSion wall, uSing micro-CT (FigS. 5 - 17).	en	Will, Kipling (2017): Description of the first flightless platynine ground beetle preserved in Baltic amber (Coleoptera: Carabidae). Zootaxa 4318 (1): 110-122, DOI: 10.11646/zootaxa.4318.1.4
03F1AD5D4C7CFFE3FF40F9F1FEE1FD03.taxon	description	Syninclusions. No SynincluSionS except for a few dirt particleS. Description. See diagnoSiS and deScription chapterS of the monotypic genuS above. MeaSurementS: Body length 5.4 mm; length of head (from tip of half-opened mandible to cervical collar) 1.1 mm; width of head between eyeS 0.9 mm; length of pronotum along midline 1.05 mm; width of pronotum 1.3 mm; length of right elytron (from baSe of Scutellum to apex) 3.3 mm; maximum width of right elytron 0.9 mm. ConSequently, the maximum width of elytra Should be approximately 1.5 mm; the inflated left elytron waS not meaSured. ProportionS: Width of pronotum / width of head = 1.43; width / length of pronotum = 1.23; width of pronotum / width of pronotal baSe between laterobaSal angleS = 1.22; length / width of elytra uSing two timeS width of right elytron = 1.64; width of pronotum / width of elytra uSing two timeS width of right elytron = 1.40. Colour: Unicoloured dark and Shiny, probably Slightly metallic, however, not clearly diStinguiShable in the foSSil Specimen.	en	Will, Kipling (2017): Description of the first flightless platynine ground beetle preserved in Baltic amber (Coleoptera: Carabidae). Zootaxa 4318 (1): 110-122, DOI: 10.11646/zootaxa.4318.1.4
03F1AD5D4C7CFFE3FF40F9F1FEE1FD03.taxon	etymology	Etymology. The SpecieS epithet referS to the markedly punctate head which iS a conSpicuouS character of the new taxon.	en	Will, Kipling (2017): Description of the first flightless platynine ground beetle preserved in Baltic amber (Coleoptera: Carabidae). Zootaxa 4318 (1): 110-122, DOI: 10.11646/zootaxa.4318.1.4
