identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03F1311657606B49BAC6FD3EFECE687F.text	03F1311657606B49BAC6FD3EFECE687F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hexarhopalus (Hexarhopalus) kaminskii Purchart 2025	<div><p>Hexarhopalus (Hexarhopalus) kaminskii sp. nov.</p><p>(Figs 2a–l)</p><p>Type material.   Holotype ♂ (NMPC): {white, printed} VIETNAM north / Hà Giang prov., vii.2024 /  Thành Ph ố  Hà Giang / local collector leg  .  Paratypes. (4 ♂ + 4 ♀ LPCB, 1 ♂ MKCP): same data as the holotype .</p><p>Differential diagnosis. Due to its black body and large confluent tubercles in elytral interstriae the new species resembles  H. higurashii Masumoto &amp; Akita, 2022 (Laos) and  H. julioi Jang &amp; Chen, 2024 (China).  Hexarhopalus higurashii differs from the new species by tubercles in elytral interstriae arranged regularly in rows, different pronotal surface being more rugose and by having more or less shiny head (compare Figs 1a, d with Fig. 3 in Masumoto &amp; Akita 2022).  Hexarhopalus julioi can be distinguished by differently arranged rugosity on the pronotal surface, by distinct oblique indentation in posterior pronotal angles (indistinct in the new species) and mainly by a roughly punctured vertex (compare Figs 1d, e with Figs 2a, b in Jang et al. 2022). It is also similar to  H. ruzzieri sp. nov.,  H. tuberculatus (Pic, 1928) (Figs 4f, g),  H. tuberculipennis Kaszab, 1960 (Figs 6h, p; for habitus see Fig. 14 in Kaszab 1960) and  H. loebli Bečvář &amp; Purchart, 2008 (Figs 6e, i; for habitus see Fig. 4 in Bečvář &amp; Purchart 2008). All these species, however, differ from the new species by having their tubercles in elytral rows isolated and well separated, whereas they are confluent in the new species.</p><p>Description. Size of the holotype 11.6 × 6.5 mm. Body black, elongate oval, strongly convex (Figs 1a–c).</p><p>Head narrower than pronotum in ratio 1:1.5 (Figs 1a, d). Whole surface shagreened, strongly matt; with fine very short (as long as one eye facet) yellow hairs in sparse and extremely shallow punctures. Frontoclypeal suture weakly marked, complete. Ocular sulcus moderately deep. Distance between eyes 2× wider than their transverse diameter (DV). Labrum shiny, strongly transverse, flat, apically with long yellow setae. Antenna shiny, 1.28× longer than pronotum, covered with yellow adherent hairs. Palpomeres shiny, apical maxillary palpomere securiform. Mentum with well visible lateral wings, middle part of mentum with two deep longitudinal grooves.</p><p>Pronotum coarsely sculptured, longer little than broad, widest before middle, sides rounded in apical third quarter, straight in basal fourth (Figs 1a, d), PW:PL ratio is 1:1.05. Pronotal margins completely bordered but obliterated in middle of anterior border. The latter is straight, basal margin rounded in middle. Oblique indentation in posterior angles shallow but well visible (Fig. 1e). Dorsal surface shagreened, strongly matt, covered with irregular shiny sculptures, with exception of midlongitudinal groove which is lacking any sculptures or rugosities. Both sides of pronotum with large swelling situated antero-laterally. Prothoracic hypomeron shagreened, smooth, glabrous.</p><p>Elytra ovate, robustly convex, 1.48× longer than its width and 2.23× longer and 1.44× wider than pronotum, broadest behind midlength (DV), highest at or promptly behind midlength (LV). Basal margin slightly raised but not carinated. Elytral surface shagreened with exception of tubercles and large protuberances situated densely in slightly convex elytral interstriae. Small isolated tubercles situated mostly along elytral suture and at apical declivity of elytra; larger tubercles situated on steep sides of elytra. Large longitudinal or transversal protuberances situated mainly on disc of elytra (Fig. 1a). Apterous. Scutellum narrowly triangular (Fig. 1d).</p><p>Prosternum strongly shagreened, matt. Prosternal process steeply inclined anteriorly, with shallow grove between coxae, almost vertically declivous behind coxae, apical part horizontal, flat, apex rounded. Mesoventrite coarse in apical half, shagreened, with shiny midlongitudinal carina in middle, posterior part steeply inclined towards mesocoxae, shagreened, matt.Metaventrite narrow (narrower than mesocoxa), shagreened, matt.Abdominal ventrites glabrous, covered densely with very shallow punctures; abdominal ventrites I–III shagreened, matt; abdominal ventrites IV–V shiny (Fig. 1b).</p><p>Legs shiny, densely punctate; protibiae distinctly curved in both sexes.</p><p>Aedeagus, spiculum gastrale, male inner sternite VIII, and male inner tergites VII and VIII are shown in Figs 1f–l.</p><p>Variability of size: 14.9–17.3 × 6.2–7.2 mm.</p><p>Etymology. Named in honour of my colleague and friend Marcin Jan Kamiński (Poland), a prominent expert on  Tenebrionidae .</p></div>	https://treatment.plazi.org/id/03F1311657606B49BAC6FD3EFECE687F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Purchart, Luboš	Purchart, Luboš (2025): Four new species of the genus Hexarhopalus Fairmaire, 1891 (Coleoptera, Tenebrionidae, Stenochiinae) from Vietnam and India. Zootaxa 5631 (1): 121-136, DOI: 10.11646/zootaxa.5631.1.5, URL: https://doi.org/10.11646/zootaxa.5631.1.5
03F1311657626B4FBAC6FD4AFE5168B3.text	03F1311657626B4FBAC6FD4AFE5168B3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hexarhopalus (Hexarhopalus) ruzzieri Purchart 2025	<div><p>Hexarhopalus (Hexarhopalus) ruzzieri sp. nov.</p><p>(Figs 2a–l)</p><p>Type material.   Holotype ♂ (NMPC): {white, printed} VIETNAM, Quảng Bình prov. / Lệ Thủy distr., iv.2023 /  Kim Th ủy env., 700 m / local collector leg  .  Paratypes. (1 ♀ NMPC, 1 ♂ BMNH, 1 ♂ ERPC, 2 ♂ + 4 LPCB): same data as holotype; (3 ♂ + 2 ♀ + 1 LPCB): {white, printed} VIETNAM, Yên Bái prov. / Yên Bái env., ix.2020 / local collector leg.;  (4 ♂ + 4 ♀ LPCB, 1 ♀ NHMB): {white, printed} VIETNAM, Yên Bái prov. /  Mù Cang Ch ải env. / ix.2023, 1600 m / local collector leg  .</p><p>Differential diagnosis. Owing to its black body colour and presence of large and well separated tubercles in elytral interstriae, it can only be confused with  H. tuberculatus (Pic, 1928) (Figs 4f, g),  H. tuberculipennis Kaszab, 1960 (see Fig. 14 in Kaszab 1960) and  H. loebli Bečvář &amp; Purchart, 2007 (see Fig. 4 in Bečvář &amp; Purchart 2008).  Hexarhopalus ruzzieri sp. nov., however, differs at first glance from  H. tuberculipennis and  H. loebli by its distinct and shiny rugosity, which covers the entire pronotal surface (Figs 2a, d), whereas it is completely smooth in  H. loebli (Figs 6e, m) or with several tubercles and three shallow depressions in  H. tuberculipennis (Figs 6h, p).  Hexarhopalus tuberculatus differs from the new species by the pronotal surface covered by large shiny granules, by large, deep and broad midlongitudinal groove on pronotal disc and by deep well developed oblique indentations in posterior angles of the pronotum (see Fig. 1A in Bečvář &amp; Purchart 2008), whereas in the new species, the pronotal surface is covered by irregular rugosities, pronotal disc with shallow midlongitudinal impression and oblique shallow indentations (Figs 2d, e). The new species also resembles  H. kaminskii sp. nov.,  H. higurashii Masumoto &amp; Akita, 2022 and  H. julioi Jang &amp; Chen, 2024 but differs from them by isolated tubercles in elytral interstriae becoming confluent in the latter three species.</p><p>Description. Size of holotype 14.6 × 6.3 mm. Body black, elongate oval, strongly convex (Figs 2a–c).</p><p>Head narrower than pronotum in ratio 1:1.46 (Figs 2a, d). Whole surface densely and coarsely punctate, in places with several shiny rugosities. Frontoclypeal suture weakly marked, complete, broadly U-shaped. Ocular sulcus moderately deep. Distance between eyes 2× wider than their transverse diameter (DV). Labrum brown, shiny, transverse, flat, apically with long yellow setae. Antenna dark brown, shiny, 1.23× longer than pronotum, covered with yellow adherent hairs. Palpomeres shiny, apical maxillary palpomere securiform. Mentum with well visible lateral wings, middle part of mentum with two shallow longitudinal grooves.</p><p>Pronotum slightly but evenly convex (LV), coarsely sculptured, nearly as long as broad, widest at middle, sides rounded in anterior three fourth, depressed posterolaterally in the posterior fourth, straight in basal fourth (Figs 2a, d). PW:PL ratio is 1:1.05. Pronotal margins completely bordered, except for anterior margin which is completely unbordered. The latter concave (in DV), anterior angles obtuse. Basal pronotal margin rounded in middle. Oblique indentation in posterior angles shallow but well obvious (Fig. 2e). Dorsal surface densely covered with shiny rugosities, space between them shagreened, thus strongly matt. Dorsum with shallow midlongitudinal impression. Prothoracic hypomeron shagreened, glabrous, mostly smooth but transversally wrinkled along posterior pronotal border.</p><p>Elytra ovate, evenly convex with slightly flattened disc (LV), 1.4× longer than its width and 2.26× longer and 1.54× wider than pronotum, broadest at midlength (DV), highest at midlength (LV). Basal margin not bordered or carinated. Whole elytral surface shagreened with exception of large and small tubercles situated in flat elytral interstriae. Striae indicated by small punctures of same size as eye facet arranged regularly in rows. Large tubercles in elytral interstriae arranged more or less regularly in rows, small tubercles (approx. as large as eye facet) arranged irregularly (Figs 1a, c). Scutellum broadly triangular (Fig. 1d). Apterous.</p><p>Prosternum shagreened, matt, longitudinally wrinkled. Prosternal process steeply inclined anteriorly, with distinct grove between coxae, almost vertically declivous behind coxae, apical part horizontal, flat, apex rounded. Mesoventrite coarse in apical half, shagreened, with shiny midlongitudinal carina in middle, posterior part steeply inclined towards mesocoxae, shegreend, matt. Metaventrite narrow (narrower than mesocoxa), shagreened, matt, sparsely and shallowly punctate. Abdominal ventrites glabrous, covered densely with very shallow punctures, shagreened, matt (Fig. 2b).</p><p>Legs shiny, densely punctate; protibiae curved in both sexes.</p><p>Aedeagus, spiculum gastrale, male inner sternite VIII, male inner tergite VII and VIII see Figs 2f–l.</p><p>Variability of size: 13.6–15.7 × 5.9–6.5 mm.</p><p>Etymology. Named in honour of my colleague and friend Enrico Ruzzier (Italy), a prominent expert on  Tenebrionidae and  Mordellidae .</p></div>	https://treatment.plazi.org/id/03F1311657626B4FBAC6FD4AFE5168B3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Purchart, Luboš	Purchart, Luboš (2025): Four new species of the genus Hexarhopalus Fairmaire, 1891 (Coleoptera, Tenebrionidae, Stenochiinae) from Vietnam and India. Zootaxa 5631 (1): 121-136, DOI: 10.11646/zootaxa.5631.1.5, URL: https://doi.org/10.11646/zootaxa.5631.1.5
03F1311657646B4DBAC6FC86FE3869AF.text	03F1311657646B4DBAC6FC86FE3869AF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hexarhopalus (Leprocaulus) arunachalpradeshensis Purchart 2025	<div><p>Hexarhopalus (Leprocaulus) arunachalpradeshensis sp. nov.</p><p>(Figs 3a–k)</p><p>Type material.   Holotype, sex not examined (BMNH): {white, printed} NE INDIA; ARUNACHAL PR. / betw. Dirang &amp; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=92.36667&amp;materialsCitation.latitude=27.316668" title="Search Plazi for locations around (long 92.36667/lat 27.316668)">Bomdila Pass</a> / 27°19´N 92°22´E; 2200m / L. Dembický leg.; 15.vi2004  .  Paratypes. (2 ♂ BMNH, 1 ♂ LPCB): same data as holotype; (1 BMNH): NE INDIA; ARUNACHAL PR. / W of BOMBDILA; 2600m; / 27°16´N 92°24´E; / L. Dembický leg.; 17.v.2004 .</p><p>Condition of holotype: Last two protarsomeres on right front leg missing.</p><p>Differential diagnosis. In general appearance, due to its rugose or roughly punctured pronotal disc and convex to carinate elytral interstriae with rather large tubercles, the new species resembles  H. eva Bečvář &amp; Purchart, 2008 (Thailand),  H. jendeki Bečvář &amp; Purchart, 2008 (India),  H. difformis (Pic, 1922) (Laos, Thailand, Vietnam, China),  H. xui Ren &amp; Xu, 2011 (China, Yunnan),  H. sinjaevi Bečvář &amp; Purchart, 2008 (Vietnam),  H. qiului Jiang, Bai, Ren &amp; Wang, 2020 (China, Yunann) and  H. xuhaoi Jiang, Bai, Ren &amp; Wang, 2020 (China, Xizang). Compared with  H. eva and  H. jendeki, the new species can be reliably distinguished mainly by smooth prothoracic hypomeron, which is densely and coarsely wrinkled in the two latter species, and by the different surface of the pronotal disc, which is densely and coarsely rugose in  H. eva (Figs 6b, j) and  H. jendeki, whereas it is densely and roughly punctured in  H. arunachalpradeshensis sp. nov. Compared with  H. difformis,  H. sinjaevi and  H. xui, it differs mainly in the absence of midlongitudinal depression ( H. difformis, see Fig. 17 in Jiang et al. (2020)) or groove ( H. sinjaevi — Figs 6g, o,  H . xui, see Fig. 22 in Ren &amp; Xu 2011).  Hexarhopalus qiului (see Figs 12, 13 in Jiang et al. (2020)) can be distinguished from the new species mainly by its roughly and irregularly wrinkled pronotal surface covered by short hairs (roughly punctured and glabrous in the new species), by head covered by short hairs (completely glabrous in the new species) and large shiny tubercles in elytral interstriae, especially in the first and third inner interstriae (tubercles are much smaller in the new species). From  H. xuhaoi (see Fig 15 in Jiang et al. 2020) it differs mainly by completely glabrous head and pronotal surface and a roughly punctured surface of the pronotum (head and pronotum covered by short hairs, pronotum sparsely and finely punctured in  H. xuhaoi).</p><p>Description. Size of holotype 9.4 × 3.5 mm. Brown, dull-shiny, with antennae and mouthparts at least partly reddish brown. Body elongate subcylindrical, glabrous (Figs 3a–b).</p><p>Head dark brown, glabrous, shagreened and therefore dull, shallowly but densely punctured, punctures´ diameter same as diameter of eye facet, narrower than pronotum in ratio 1:1.22 (Figs 3a, c). Frontoclypeal suture well developed, moderately deep, reaching clypeal margin. Ocular sulcus moderately deep, distance between eyes 2.32× wider than their transverse diameter (DV). Antenna pale brown, pubescent, 1.5× longer than pronotum.</p><p>Pronotum dark brown, shagreened, dull, subquadrate, broadest at middle, PW:PL ratio is 1: 1.15 (Figs 3a, c). Entire surface of pronotum irregularly and coarsely punctate, with punctures´ diameter slightly larger than eye facet. Oblique indentation in posterior angles relatively long, deep and smooth (Fig. 3d). Anterior margin of pronotum straight, not rimmed. Base of pronotum completely rimmed. Prothoracic hypomeron smooth, shagreened, dull, impunctate, separated from pronotal disc by lateral groove.</p><p>Elytra brown, shagreened, 1.86× longer than its width, 3.25× longer and 1.52× wider than pronotum, dorsally flattened (LV), very slightly dilated posteriad, broadest at posterior third (Fig. 3a). Declivity of elytra before apex moderate (LV). Elytral humeri well developed. Elytral striae formed of dense row of separated deep and large rounded to longitudinal punctures. First two inner elytral interstriae flat to convex with irregularly situated granules of the same size as punctures in elytral striae. Remaining interstriae convex to slightly carinate (especially in anterior part of elytra) with more or less regularly situated shiny granules on their top. Base of elytra rimmed with moderately raised carina. Winged. Scutellum triangular (Fig. 3c).</p><p>Prosternum brown, shagreened, glabrous, dull. Prosternal process with distinct and rather deep midlongitudinal groove between procoxae, apically broadly rounded, as wide as procoxa. Mesosternum pale brown, shagreened, dull, wrinkled, glabrous; in middle with shiny midlongitudinal carina. Metasternum pale brown, shagreened, dull, very finely and shallowly but densely punctate, finely pubescent. Approximately 2.5× longer than mesocoxa. Abdominal ventrites pale brown, shagreened, dull, very finely and shallowly but densely punctate, finely pubescent, abdominal ventrite I–III rimmed laterally (Fig. 3b).</p><p>Legs. Brown, simply, finely and densely punctate. Femora claviform, glabrous. Tibiae straight, finely pubescent. Tarsi dorsally pubescent.</p><p>Aedeagus, spiculum gastrale, male inner sternite VIII, male inner tergite VII and VIII are shown in Figs 3e–k.</p><p>Variability of size: 8.6–10.0 × 3.0– 3.5 mm.</p><p>Etymology. The specific epithet refers to the area of the species occurrence, Arunachal Pradesh; the name is treated as an adjective.</p></div>	https://treatment.plazi.org/id/03F1311657646B4DBAC6FC86FE3869AF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Purchart, Luboš	Purchart, Luboš (2025): Four new species of the genus Hexarhopalus Fairmaire, 1891 (Coleoptera, Tenebrionidae, Stenochiinae) from Vietnam and India. Zootaxa 5631 (1): 121-136, DOI: 10.11646/zootaxa.5631.1.5, URL: https://doi.org/10.11646/zootaxa.5631.1.5
03F1311657666B43BAC6FBFAFCC66FDF.text	03F1311657666B43BAC6FBFAFCC66FDF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hexarhopalus (Leprocaulus) binhanus (Pic 1922) Purchart 2025	<div><p>Hexarhopalus (Leprocaulus) binhanus (Pic, 1922) comb. n.</p><p>(Figs 4a–d, 6a, i)</p><p>Derosphaerus binhanus Pic, 1922: 26 .— Gebien (1941): 353 (648).</p><p>Type material.   Lectotype ♂ (MNHN, designated here): TYPE {red, printed} // Hoa—Binh [province] / (Tonkin) {white, handwritten} // binhanus  n. sp. {white, handwritten} // Museum Paris / Coll. M. Pic {white, printed} // LECTOTYPE /  Derosphaerus /  binhanus Pic, 1922 / des. L. Purchart, 2023 {red, handwritten}  .</p><p>Additional material.   Vietnam: (1 BMNH): LACTHO / Tonkin /  de Cooman // 385 // Necrobioid. / gibbicollis / Fleut. // Necrobioides / gibbicollis / Fleut.;   (4 ♂ + 4 ♀ MNHN):  LACTHO / Tonkin /  de Cooman // MUSEUM PARIS / COL. P. ARDOIN / 1978;   (1 BMNH): Tonkin / Laos / [the other side of the label] 386 // Necrobioid. / gibbicollis / Fleut. //  Derosphaerus ! / det. Schawaller 2010 ;   Thailand: (1 BMNH): THAI, 9.–13.IV.1991 / <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=35.0&amp;materialsCitation.latitude=2.0" title="Search Plazi for locations around (long 35.0/lat 2.0)">THIMONGHTA</a>, 350 m / 15 02 °N 98 35 °E / P. Pacholatko leg ;   Laos: (1 ♀ MNHN): JAN 1963 / Vientiane / LAOS—  A. Baudon // cum typo / comparat /  Derosphaerus / binhanus / P. ARDOIN det. 1963 // MUSEUM PARIS / COL. P. ARDOIN / 1978;  (1 ♀ MNHN):   17.VII.1975 /  Khongsédone / LAOS / J. Rondon leg. // MUSEUM PARIS / COL. P. ARDOIN / 1978;   (1 ♂ MNHN): 5.VII.1965 /  Paksong / Sud LAOS / J. Rondon leg. // MUSEUM PARIS / COL. P. ARDOIN / 1978;  (1 ♂ MNHN): 8.VI.1965 / Phou Khao Khouai / Vientiane — LAOS / J. Rondon leg. // MUSEUM PARIS / COL. P. ARDOIN / 1978; (1 ♂ MNHN): 30.X.1966 / Ban Van Eua / Vientiane — LAOS / J. Rondon leg. // MUSEUM PARIS / COL. P. ARDOIN / 1978; (1 ♀ MNHN): 1.V.1964 / Phontiou / Thakek — LAOS / J. Rondon leg. // MUSEUM PARIS / COL. P. ARDOIN / 1978;  (1 ♀ MNHN): VII.1964 /  Plaine des Jarres / LAOS — A. Baudon;   (2 MNHN): Boa Binh / Tonkin //  Derosphaerus / binhanus // Museum Paris / Coll. M. Pic.</p><p>Remark. I had the opportunity to study the type of  Derosphaerus binhanus Pic, 1922 deposited in MNHN together with a smaller series of non-type specimens belonging to this species. I found out that it actually belongs to the genus  Hexarhopalus due to constantly present oblique indentation in the posterior corners of the pronotum, which can be quite inconspicuous but obvious. Therefore, I transfer this species from  Derosphaerus to  Hexarhopalus as  H. binhanus (Pic, 1922) comb. n.</p><p>In additional material the locality “ Tonkin Laos ” is stated in one specimen (BMNH), but Tonkin in Laos does not exist. I assume that the collector of this specimen did not record accurate locality data while travelling in these two areas which were parts of the former French Indochina. Therefore, he probably just labelled this specimen with the locality label Tonkin Laos.</p><p>Differential diagnosis. In general appearance  H. binhanus (Pic, 1922) comb. n. can, owing to the convex elytral interstriae without tubercles and smooth, glabrous pronotum, only be confused with  H. kaszabi Bečvář &amp; Purchart, 2008 (Borneo) and  H. lilligi Bečvář &amp; Purchart, 2008 (Thailand, Laos). From both species, however, it differs by shallow and extremely weakly developed lateral oblique indentation in posterior pronotal angles (Fig. 6i), which is well developed and deep in the latter species (Figs 6k, l). In addition, it differs from  H. lilligi by brown colour and finer punctation of the pronotal surface (in  H. lilligi the colour is black and pronotal surface coarsely punctate Fig. 6d). Compared with  H. kaszabi, it further differs by completely obliterated lateral edge of pronotum whereas it is very shortly developed in posterior pronotal angles in  H. kaszabi (Fig. 6k).</p><p>Redescription. Size of lectotype 10.8 × 4.1 mm. Winged, brown, dull shiny. Body elongate subcylindrical, glabrous (Figs 4a–c).</p><p>Head glabrous, narrower than pronotum in ratio 1:1.48 (Fig. 6a). Vertex shagreened and therefore dull. Frons shagreened and finely punctate, punctures´ diameter smaller than diameter of eye facet. Clypeus shiny and finely but densely punctate with punctures´ diameter same as of eye facet, anterior margin emarginated. Labrum transverse, reddish-brown, keeled transversally, with long yellow setae anteriorly. Frontoclypeal suture well developed, rather shallow, reaching clypeal margin. Ocular sulcus moderately deep, distance between eyes 1.09× wider than their transverse diameter (DV). Antenna pale brown, pubescent, 1.05× longer than pronotum.</p><p>Pronotum shagreened, dull-shiny, slightly transverse, PW:PL ratio is 1:1.13, broadest at middle (Fig. 6a). Entire surface of pronotum irregularly and densely punctate, with punctures´ diameter approx. as large as eye facet and intervals between punctures same or 2× larger than their diameter. Oblique indentation in posterior angles extremely slightly developed, almost obliterated, very shallow (Fig. 6i). Anterior margin of pronotum straight, only very slightly rimmed at anterior corners. Sides of pronotum rounded. Base of pronotum completely and deeply rimmed. Prothoracic hypomeron smooth, shagreened, dull, impunctate, not separated from pronotal disc by lateral groove which is completely absent.</p><p>Elytra shagreened, 1.85× longer than its width, 3.4× longer and 1.54× wider than pronotum, dorsally flattened (LV), slightly dilated posteriad, broadest at posterior third (Fig. 1a–b). Declivity of elytra before apex moderate (LV). Elytral humeri well developed. Elytral striae deep with row of separated deep rounded punctures. All elytral interstriae convex. Base of elytra not rimmed. Scutellum triangular (Fig. 6a).</p><p>Prosternum brown, shagreened, glabrous, dull. Prosternal process with distinct and rather deep midlongitudinal groove between procoxae, apically broadly rounded, slightly narrower than procoxa. Mesosternum brown, shiny, glabrous; in middle with shiny midlongitudinal carina. Metasternum brown, shagreened, dull-shiny, very finely, shallowly and sparsely punctate.Approximately 1.7× longer than mesocoxa.Abdominal ventrites brown, shagreened, dull-shiny, finely, shallowly and rather densely punctate, glabrous.</p><p>Legs brown, simply, finely and densely punctate. Femora claviform, glabrous. Tibiae straight, finely pubescent. Tarsi dorsally pubescent.</p><p>Variability of size: 9.5–11.9 × 3.6–4.4 mm.</p><p>Distribution. Vietnam. New records for Thailand and Laos.</p></div>	https://treatment.plazi.org/id/03F1311657666B43BAC6FBFAFCC66FDF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Purchart, Luboš	Purchart, Luboš (2025): Four new species of the genus Hexarhopalus Fairmaire, 1891 (Coleoptera, Tenebrionidae, Stenochiinae) from Vietnam and India. Zootaxa 5631 (1): 121-136, DOI: 10.11646/zootaxa.5631.1.5, URL: https://doi.org/10.11646/zootaxa.5631.1.5
03F1311657686B41BAC6FA2AFEEB6EE3.text	03F1311657686B41BAC6FA2AFEEB6EE3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hexarhopalus (Leprocaulus) kontumensis Purchart 2025	<div><p>Hexarhopalus (Leprocaulus) kontumensis sp. nov.</p><p>(Figs 5a–e)</p><p>Type material.  Holotype, sex not examined (NMPC): {white, printed} VIETNAM centr. / Kon Tum prov., v. 2023 / local collector leg .   Paratypes. (2 LPCB): {white, printed} VIETNAM centr. / Kon Tum prov., iv.2024 /  Ngọc Linh vill. env. / local collector leg  .</p><p>Condition of the holotype. Right hind leg is missing.</p><p>Differential diagnosis. Owing to its dark brown colour, presence of tubercles in elytral interstriae and smooth pronotal disc, it can only be confused with  H. particularis (Pic, 1922) from Vietnam (Fig. 4e) and  H. schawalleri Bečvář &amp; Purchart, 2008 from Laos (for figure of habitus see Bečvář &amp; Purchart (2008)). Both mentioned species, however, differ from  H. kontumensis sp. nov. by the presence of two large and smooth, dorso-laterally situated longitudinal swellings ranging from anterior angles posteriad laterally toward posterior third of pronotum (Figs 6f, n).</p><p>Description. Size of holotype 11.3 × 4.2 mm. Brown, with femorae and labrum pale brown, shiny. Body elongate subcylindrical, glabrous (Figs 5a–c).</p><p>Head glabrous, finely shagreened, very sparsely and very shallowly micro-punctured, punctures´ diameter much smaller than eye facet; narrower than pronotum in ratio 1:1.44 (Figs 5a, d). Frontoclypeal suture well developed, moderately deep, reaching clypeal margin. Ocular sulcus moderately deep, distance between eyes 1.25× wider than their transverse diameter (DV). Antenna, pubescent, 1.14× longer than pronotum.</p><p>Pronotum dark brown, finely shagreened, slightly shiny, subquadrate, little wider than long, broadest at anterior third, PW:PL ratio 1: 1.09 (Figs 5a, d), flattened (LV). Sides of pronotum slightly rounded in anterior third, then gradually narrowing towards pronotal base, slightly depressed posterolaterally. Entire surface of pronotum very shallowly and sparsely punctate, diameter of punctures little smaller than eye facet. Disc of pronotum with fine, shallow but obvious midlongitudinal grove. Oblique indentation in posterior angles relatively long, deep and smooth (Fig. 5e). Anterior margin of pronotum concave (DV), rimmed but obliterated in middle. Base of pronotum completely rimmed, rounded in middle (DV). Prothoracic hypomeron smooth, finely shagreened, glabrous, separated from pronotal disc by fine, in places obliterated border.</p><p>Elytra finely shagreened, glabrous, 1.86× longer than its width, 2.91× longer and 1.46× wider than pronotum, dorsally flattened (LV), subcylindrical, very slightly dilated posteriad, broadest at posterior third (Fig. 5a). Declivity of elytra before apex moderate (LV). Elytral humeri well developed. Elytral striae formed of dense row of separated moderately deep rounded punctures, twice as large as eye facet. Elytral interstriae slightly convex bearing row of low but distinct tubercles, space between tubercles approx. of same size as the width of respective elytral interstria. Base of elytra slightly raised. Scutellum triangular (Figs 5d). Winged.</p><p>Prosternum shagreened, glabrous, dull. Prosternal process with distinct and rather deep midlongitudinal groove between procoxae, apically broadly rounded, as wide as procoxa. Mesosternum shagreened, dull, glabrous; in middle with shiny midlongitudinal carina. Metasternum shagreened, dull, very finely and shallowly punctate, finely pubescent. Approximately 2.5× longer than mesocoxa. Abdominal ventrites shagreened, dull, very finely and shallowly but densely punctate, finely pubescent (Fig. 5b).</p><p>Legs finely and sparsely punctate with very shallow punctures, sparsely covered with very fine adherent setae.</p><p>Variability of size: 11.3–12.0 × 4.2–4.4 mm.</p><p>Etymology. The specific epithet refers to the area of species occurrence, Kon Tum province; the name is treated as an adjective.</p></div>	https://treatment.plazi.org/id/03F1311657686B41BAC6FA2AFEEB6EE3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Purchart, Luboš	Purchart, Luboš (2025): Four new species of the genus Hexarhopalus Fairmaire, 1891 (Coleoptera, Tenebrionidae, Stenochiinae) from Vietnam and India. Zootaxa 5631 (1): 121-136, DOI: 10.11646/zootaxa.5631.1.5, URL: https://doi.org/10.11646/zootaxa.5631.1.5
03F13116576A6B41BAC6FAFEFBE06FA7.text	03F13116576A6B41BAC6FAFEFBE06FA7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hexarhopalus lilligi Becvar & Purchart 2008	<div><p>Hexarhopalus lilligi Bečvář &amp; Purchart, 2008</p><p>(1 NMPC, 1 LPCB): Laos, Sekong prov., ca. 12 km S Sekong, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=106.75166&amp;materialsCitation.latitude=15.245" title="Search Plazi for locations around (long 106.75166/lat 15.245)">Tad Faek</a> waterfalls (at light), 15°14.7´N, 106°45.1´E, 118 m, Jiří Hájek leg., 8.+ 12.v.2010  .</p><p>Remark. First two specimens since holotype description. New record for Laos.</p></div>	https://treatment.plazi.org/id/03F13116576A6B41BAC6FAFEFBE06FA7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Purchart, Luboš	Purchart, Luboš (2025): Four new species of the genus Hexarhopalus Fairmaire, 1891 (Coleoptera, Tenebrionidae, Stenochiinae) from Vietnam and India. Zootaxa 5631 (1): 121-136, DOI: 10.11646/zootaxa.5631.1.5, URL: https://doi.org/10.11646/zootaxa.5631.1.5
03F13116576A6B41BAC6F9F2FBD16D47.text	03F13116576A6B41BAC6F9F2FBD16D47.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hexarhopalus masumotoi Becvar & Purchart 2008	<div><p>Hexarhopalus masumotoi Bečvář &amp; Purchart, 2008</p><p>(1 LPCB): Malaysia / Borneo, Sabah, / Keningau distr.,  Trus Madi / Mt., h.— 1160 m / 20–21. iii. 2012  .</p><p>Remark. Second known specimen of this species so far. The specimen was collected in type locality (Keningau env.) and its habitus perfectly agrees with all the characteristics mentioned (e.g. colour, pronotal and elytral surface, etc.) in the holotype description except for the size, which is 9.3 mm (7 mm in holotype).</p></div>	https://treatment.plazi.org/id/03F13116576A6B41BAC6F9F2FBD16D47	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Purchart, Luboš	Purchart, Luboš (2025): Four new species of the genus Hexarhopalus Fairmaire, 1891 (Coleoptera, Tenebrionidae, Stenochiinae) from Vietnam and India. Zootaxa 5631 (1): 121-136, DOI: 10.11646/zootaxa.5631.1.5, URL: https://doi.org/10.11646/zootaxa.5631.1.5
03F13116576A6B41BAC6F892FC276D9F.text	03F13116576A6B41BAC6F892FC276D9F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hexarhopalus sinjaevi Becvar & Purchart 2008	<div><p>Hexarhopalus sinjaevi Bečvář &amp; Purchart, 2008</p><p>(1 LPCB): Vietnam N, Yên Bái prov. /  Yên Bái env., vi.2023 / local collector leg.</p><p>Remark. Third known specimen of this rarely collected taxon.</p></div>	https://treatment.plazi.org/id/03F13116576A6B41BAC6F892FC276D9F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Purchart, Luboš	Purchart, Luboš (2025): Four new species of the genus Hexarhopalus Fairmaire, 1891 (Coleoptera, Tenebrionidae, Stenochiinae) from Vietnam and India. Zootaxa 5631 (1): 121-136, DOI: 10.11646/zootaxa.5631.1.5, URL: https://doi.org/10.11646/zootaxa.5631.1.5
03F13116576D6B44BAC6F8A0FB0669C3.text	03F13116576D6B44BAC6F8A0FB0669C3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hexarhopalus Fairmaire 1891	<div><p>List of the species of the genus  Hexarhopalus Fairmaire, 1891</p><p>Remark. Masumoto &amp; Akita (2022) described  Hexarhopalus maoi Masumoto &amp; Akita, 2022 from Laos. According to the description and the photo of habitus provided in their paper I have strong doubts that this species belongs to genus  Hexarhopalus due to lacking claviform femorae (present in all known species) and indentation in posterior angles of pronotum which can be quite weakly developed and thus is inconspicuous but always obvious.</p><p>Subgenus  Hexarhopalus Fairmaire, 1891</p><p>1.  H. attenuatus (Pic, 1922) China (Yunnan)</p><p>2.  H. becvari Purchart, 2010 Borneo (Sabah)</p><p>3.  H. birmanicus Kaszab, 1960 Myanmar</p><p>4.  H. borneensis (Kaszab, 1982) Borneo (Sabah)</p><p>5.  H. bouchardi Bečvář &amp; Purchart, 2008 Borneo (Sabah)</p><p>6.  H. bremeri Bečvář &amp; Purchart, 2008 Borneo (Sabah)</p><p>7.  H. ferreri Purchart, 2010 Borneo (Sarawak)</p><p>8.  H. higurashii Masumoto &amp; Akita, 2022 Laos</p><p>9.  H. iwani Purchart, 2010 Borneo</p><p>10.  H. julioi Jiang &amp; Chen, 2024 China (Yunnan)</p><p>=  H. ferreri Jiang, Zhou, Liu, Huang &amp; Chen, 2022: 539 (nec Purchart, 2010: 5)</p><p>11.  H. kaminskii sp. nov. Vietnam</p><p>12.  H. liuyixiaoi Jiang, Bai, Ren &amp; Wang, 2020 China (Hunan, Hubei) 13.  H. loebli Bečvář &amp; Purchart, 2008 West Malaysia</p><p>14.  H. mangshanicus Ren &amp; Xu, 2011 China (Hunan, Guangdong) 15.  H. maoi Masumoto &amp; Akita, 2022 Laos</p><p>16.  H. montanus (Kaszab, 1982) Borneo (Sabah)</p><p>17.  H. murudensis Purchart, 2010 Borneo (Sarawak)</p><p>18.  H. problematicus Bečvář &amp; Purchart, 2008 Thailand</p><p>19.  H. rolandi Purchart, 2010 Borneo (Sabah)</p><p>20.  H. ruzzieri sp. nov. Vietnam</p><p>21.  H. sculpticollis Fairmaire, 1891 China (Zhejiang, Anhui) 22.  H. sculptilis Kaszab, 1960 Myanmar, China (Yunnan) 23.  H. sculptithorax Kaszab, 1960 Thailand</p><p>24.  H. tianbaoyanensis Jiang, Zhou, Liu, Huang &amp; Chen, 2022 China (Fujian)</p><p>25.  H. tuberculatus (Pic, 1928) Laos</p><p>26.  H. tuberculipennis Kaszab, 1960 West Malaysia</p><p>27.  H. yeziyangi Jiang, Zhou, Liu, Huang &amp; Chen, 2022 China (Jiangxi)</p><p>28.  H. yunnanensis Jiang, Li, Ji, Engel &amp; Wang, 2021 China (Yunnan)</p><p>29.  H. zhuxiangi Jiang, Li, Ji, Engel &amp; Wang, 2021 China (Guangxi)</p><p>Subgenus  Leprocaulus Fairmaire, 1896</p><p>30.  H. andoi Bečvář &amp; Purchart, 2008 Borneo (Sabah)</p><p>31.  H. arunachalpradeshensis sp. nov. India</p><p>32.  H. barclayi Purchart, 2010 Borneo (Sarawak)</p><p>33.  H. binhanus (Pic, 1922) comb. n. Vietnam, Thailand, Laos</p><p>34.  H. bisinuatus Ren &amp; Xu, 2011 China (Guizhou, Hunan, Sichuan, Fujian)</p><p>35.  H. cameroni Bečvář &amp; Purchart, 2008 West Malaysia, (Thailand?)</p><p>36.  H. clavipes (Fairmaire, 1896) Sumatra, Java</p><p>37.  H. crockeri Bečvář &amp; Purchart, 2008 Borneo (Sabah)</p><p>38.  H. difformis (Pic, 1922) Laos, Thailand, Vietnam, China (Yunnan)</p><p>39.  H. eva Bečvář &amp; Purchart, 2008 Thailand</p><p>40.  H. granulipennis (Blair, 1919) Sumatra</p><p>41.  H. grimmi Bečvář &amp; Purchart, 2008 Sumatra</p><p>42.  H. jendeki Bečvář &amp; Purchart, 2008 India</p><p>43.  H. kaszabi Bečvář &amp; Purchart, 2008 Borneo (Sabah)</p><p>44.  H. kontumensis sp. nov. Vietnam</p><p>45.  H. kubani Bečvář &amp; Purchart, 2008 Thailand</p><p>46.  H. lilligi Bečvář &amp; Purchart, 2008 Thailand, Laos</p><p>47.  H. loeffleri (Kaszab, 1982) Borneo (Sabah)</p><p>48.  H. masumotoi Bečvář &amp; Purchart, 2008 Borneo (Sabah)</p><p>49.  H. merkli Bečvář &amp; Purchart, 2008 West Malaysia, Thailand</p><p>50.  H. nanlingensis Jiang, Li, Ji, Engel &amp; Wang, 2021 China (Guangdong)</p><p>51.  H. pacholatkoi Bečvář &amp; Purchart, 2008 Thailand, Myanmar, Laos</p><p>52.  H. particularis (Pic, 1922) Vietnam</p><p>53.  H. punctithorax (Kaszab, 1982) Vietnam</p><p>54.  H. qiujianyueae Jiang, Bai, Ren &amp; Wang, 2020 China (Yunnan)</p><p>55.  H. qiului Jiang, Bai, Ren &amp; Wang, 2020 China (Yunnan)</p><p>56.  H. rotundicollis (Pic, 1922) Borneo (Sabah)</p><p>57.  H. schawalleri Bečvář &amp; Purchart, 2008 Laos</p><p>58.  H. seniori Bečvář &amp; Purchart, 2008 Thailand, Laos</p><p>59.  H. sinjaevi Bečvář &amp; Purchart, 2008 Vietnam</p><p>60.  H. sumatranus (Kaszab, 1982) Sumatra</p><p>61.  H. tibangi Bečvář &amp; Purchart, 2008 Borneo (border of Sarawak and East Kalimantan)</p><p>62.  H. xuhaoi Jiang, Bai, Ren &amp; Wang, 2020 China (Xizang)</p><p>63.  H. xui Ren &amp; Xu, 2011 China (Yunnan)</p></div>	https://treatment.plazi.org/id/03F13116576D6B44BAC6F8A0FB0669C3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Purchart, Luboš	Purchart, Luboš (2025): Four new species of the genus Hexarhopalus Fairmaire, 1891 (Coleoptera, Tenebrionidae, Stenochiinae) from Vietnam and India. Zootaxa 5631 (1): 121-136, DOI: 10.11646/zootaxa.5631.1.5, URL: https://doi.org/10.11646/zootaxa.5631.1.5
