taxonID	type	description	language	source
03FCAD28493AFF86FF37FF00A3B2D2A9.taxon	description	urn: lsid: zoobank. org: act: CD 013 F 02 - 6 E 7 B- 4 E 89 - A 47 E- 5 F 307 DD 878 D 3 (Fig. 1)	en	Varga, Oleksandr (2025): First records of the genera Gnathochorisis Förster, 1869 and Symplecis Förster, 1869 (Hymenoptera, Ichneumonidae, Orthocentrinae) from Vietnam. Zootaxa 5647 (6): 571-583, DOI: 10.11646/zootaxa.5647.6.5, URL: https://doi.org/10.11646/zootaxa.5647.6.5
03FCAD28493AFF86FF37FF00A3B2D2A9.taxon	materials_examined	Material examined. Holotype: ♀, VIETNAM: Tonkin, Hoang Lien N. R., 15 km W Sa Pa, ca. 1900 m, Malaise traps, 15 – 21. x. 1999, leg. C. v. Achterberg, RMHN’ 99, RMNH. INS. 1722319 (RMNH). Paratype: 1 ♂, the same locality and date as for holotype, RMNH. INS. 1722320 (RMNH).	en	Varga, Oleksandr (2025): First records of the genera Gnathochorisis Förster, 1869 and Symplecis Förster, 1869 (Hymenoptera, Ichneumonidae, Orthocentrinae) from Vietnam. Zootaxa 5647 (6): 571-583, DOI: 10.11646/zootaxa.5647.6.5, URL: https://doi.org/10.11646/zootaxa.5647.6.5
03FCAD28493AFF86FF37FF00A3B2D2A9.taxon	diagnosis	Diagnosis. Gnathochorisis sapaensis sp. nov. is characterized by the combination of the following characters: face orange, clypeus yellow (Fig. 1 C); antenna with 21 flagellomeres, first flagellomere 3.0 × as long as wide; pronotum partly orange; metapleuron smooth (Fig. 1 G); propodeum with area superomedia elongate, 1.4 × as long as wide (Fig. 1 E); fore wing with vein 3 rs-m present; first metasomal tergite 2.5 × as long as maximum width, with latero-median carinae present; second tergite granulate, 1.2 × as long as apical width (Fig. 1 H); ovipositor 0.9 × as long as hind tibia. Gnathochorisis sapaensis sp. nov. is similar to G. malaisei Humala, 2021 from Myanmar in having the metapleuron smooth, propodeum with area superomedia elongate, and fore wing with vein 3 rs-m present, but differs by the shorter first flagellomere, which is 3.0 × as long as wide (5.3 × in G. malaisei), dark frons (orbits yellow in G. malaisei), and granulate second tergite of metasoma (longitudinally wrinkled anteriorly in G. malaisei).	en	Varga, Oleksandr (2025): First records of the genera Gnathochorisis Förster, 1869 and Symplecis Förster, 1869 (Hymenoptera, Ichneumonidae, Orthocentrinae) from Vietnam. Zootaxa 5647 (6): 571-583, DOI: 10.11646/zootaxa.5647.6.5, URL: https://doi.org/10.11646/zootaxa.5647.6.5
03FCAD28493AFF86FF37FF00A3B2D2A9.taxon	description	Description. Female. Holotype. (Figs 1 A, C, E – H). Body length approximately 3.4 mm. Fore wing 2.9 mm. Head (Figs 1 C, F – G) smooth and densely pubescent. Antenna with 21 flagellomere, first flagellomere 3.0 × as long as wide. Face about 0.7 × as long as wide, sparsely punctate and with traces of granulation; eyes slightly divergent to clypeus, glabrous. Malar space long, 1.7 × the basal width of mandible; subocular sulcus present. Clypeus 0.8 × as long as wide, smooth. Mandible bidentate, not twisted, lower tooth shorter than upper tooth. Temples short and strongly narrowed behind eyes (dorsal view). Frons and vertex smooth and densely pubescent; length of the ocellar-ocular distance 1.3 × maximum diameter of lateral ocellus; occipital carina complete. Mesosoma (Figs 1 E, G) smooth and densely pubescent. Propleuron sparsely pubescent. Pronotum smooth; epomia present, but not reaching mesoscutum. Mesoscutum densely and evenly pubescent; notauli deep anteriorly, with vertical carina. Scutellum densely pubescent. Mesopleuron smooth, densely pubescent anteriorly and ventrally; epicnemial carina present; sternaulus deep on anterior 0.3 of mesopleuron. Metapleuron smooth, with some rugosities on the posterior edge; pleural and submetapleural carinae present. Propodeum smooth, carinated; area superomedia elongate, 1.4 × as long as wide, costulae present; area basalis not delimited by carinae anteriorly; apophyses present, but weak. Legs relatively stout; hind femur 3.2 × as long as wide; third tarsomere of hind tarsus as long as fifth tarsomere; tarsal claws simple. Wings. Fore wing with vein 2 rs-m about 0.8 × the distance between 2 rs-m and 2 m-cu; vein 3 rs-m present; vein 1 cu-a opposite to M & Rs; hind wing with nervellus intercepted below the middle, inclivous; distal abscissa of Cu present. Metasoma (Fig. 1 H) granulate anteriorly and sparsely pubescent. First tergite 2.5 × as long as maximum width, granulate; latero-median carinae present. Second tergite 1.2 × as long as apical width, granulate. The remaining tergites smooth. Ovipositor 0.9 × as long as hind tibia. Colour. Head black except face orange, clypeus, mandible (except apices), scape and pedicel yellow. Mesosoma black except propleuron ventrally, pronotum dorsally and ventrally, subtegular ridge orange, and tegula yellow. Fore and mid legs orange except coxae, trochanters and trochantelli yellow; hind leg orange except coxa largely, trochanter partly, femur except base, tibia except middle, and tarsus entirely fuscous. Metasoma with first tergite black; the remaining tergites brown, except second tergite apically, third tergite basally and apically yellow. Pterostigma and veins brown. Male. Generally resembles female, but differs by the antenna with 22 flagellomeres, first flagellomere 3.8 × as long as wide; more abundant yellow colouration of head; shorter area superomedia, which is about 1.2 × as long as wide; and darker hind legs and third tergite of metasoma (Figs 1 B, D).	en	Varga, Oleksandr (2025): First records of the genera Gnathochorisis Förster, 1869 and Symplecis Förster, 1869 (Hymenoptera, Ichneumonidae, Orthocentrinae) from Vietnam. Zootaxa 5647 (6): 571-583, DOI: 10.11646/zootaxa.5647.6.5, URL: https://doi.org/10.11646/zootaxa.5647.6.5
03FCAD28493AFF86FF37FF00A3B2D2A9.taxon	etymology	Etymology. The specific epithet refers to the type locality closely situated to the city Sa Pa. Adjective.	en	Varga, Oleksandr (2025): First records of the genera Gnathochorisis Förster, 1869 and Symplecis Förster, 1869 (Hymenoptera, Ichneumonidae, Orthocentrinae) from Vietnam. Zootaxa 5647 (6): 571-583, DOI: 10.11646/zootaxa.5647.6.5, URL: https://doi.org/10.11646/zootaxa.5647.6.5
03FCAD28493AFF86FF37FF00A3B2D2A9.taxon	distribution	Distribution. Vietnam.	en	Varga, Oleksandr (2025): First records of the genera Gnathochorisis Förster, 1869 and Symplecis Förster, 1869 (Hymenoptera, Ichneumonidae, Orthocentrinae) from Vietnam. Zootaxa 5647 (6): 571-583, DOI: 10.11646/zootaxa.5647.6.5, URL: https://doi.org/10.11646/zootaxa.5647.6.5
03FCAD284939FF84FF37F948A1B5D480.taxon	description	(Fig. 2)	en	Varga, Oleksandr (2025): First records of the genera Gnathochorisis Förster, 1869 and Symplecis Förster, 1869 (Hymenoptera, Ichneumonidae, Orthocentrinae) from Vietnam. Zootaxa 5647 (6): 571-583, DOI: 10.11646/zootaxa.5647.6.5, URL: https://doi.org/10.11646/zootaxa.5647.6.5
03FCAD284939FF84FF37F948A1B5D480.taxon	materials_examined	Material examined. 1 ♀, VIETNAM: Viet Try, nr Thanh Son, Thuong Cuu, 20 ° 59 ’ N, 105 ° 8 ’ N, 350 – 400 m, Malaise traps, 11 – 16. x. 1999, leg. R. de Vries, RMHN’ 99, RMNH. INS. 1722321 (RMNH); 1 ♀, Dak Lak, Chu Yang Sin N. P., Krong K’Mar, 740 – 900 m, Malaise traps, 02 – 10. vi. 2007, leg. C. v. Achterberg & R. de Vries, RMHN’ 07, RMNH. INS. 1722322 (RMNH).	en	Varga, Oleksandr (2025): First records of the genera Gnathochorisis Förster, 1869 and Symplecis Förster, 1869 (Hymenoptera, Ichneumonidae, Orthocentrinae) from Vietnam. Zootaxa 5647 (6): 571-583, DOI: 10.11646/zootaxa.5647.6.5, URL: https://doi.org/10.11646/zootaxa.5647.6.5
03FCAD284939FF84FF37F948A1B5D480.taxon	diagnosis	Diagnosis (based on Vietnamese specimens). Gnathochorisis leleji is characterized by the combination of the following characters: face brown to black, clypeus yellow; antenna with 22 flagellomere, first flagellomere 4.3 – 4.4 × as long as wide (Fig. 2 B); mesosoma black; metapleuron largely smooth (Fig. 2 A); propodeum with area superomedia transverse, 0.6 × as long as wide; fore wing with vein 3 rs-m absent; second tergite smooth and longitudinally wrinkled, 0.8 – 0.9 × as long as apical width; ovipositor 0.8 × as long as hind tibia and 1.0 × as long as first tergite (Fig. 2 A).	en	Varga, Oleksandr (2025): First records of the genera Gnathochorisis Förster, 1869 and Symplecis Förster, 1869 (Hymenoptera, Ichneumonidae, Orthocentrinae) from Vietnam. Zootaxa 5647 (6): 571-583, DOI: 10.11646/zootaxa.5647.6.5, URL: https://doi.org/10.11646/zootaxa.5647.6.5
03FCAD284939FF84FF37F948A1B5D480.taxon	discussion	Remark. Vietnamese specimens of Gnathochorisis leleji are generally similar to those described from Malaysia, but differ by the slightly wider area superomedia of the propodeum and longer ovipositor, however both characters seem to be variable. Additional female specimen examined (but not listed in the material section) has somewhat shorter first flagellomere (4.0 × as long as wide), more rugulose posteriorly metapleuron (Fig. 2 C) and second tergite of metasoma narrowly banded with yellow apically (Fig. 2 D). The taxonomic status of this specimen remains uncertain until more similar specimens be examined.	en	Varga, Oleksandr (2025): First records of the genera Gnathochorisis Förster, 1869 and Symplecis Förster, 1869 (Hymenoptera, Ichneumonidae, Orthocentrinae) from Vietnam. Zootaxa 5647 (6): 571-583, DOI: 10.11646/zootaxa.5647.6.5, URL: https://doi.org/10.11646/zootaxa.5647.6.5
03FCAD284939FF84FF37F948A1B5D480.taxon	distribution	Distribution. Malaysia (Humala 2021), first record from Vietnam.	en	Varga, Oleksandr (2025): First records of the genera Gnathochorisis Förster, 1869 and Symplecis Förster, 1869 (Hymenoptera, Ichneumonidae, Orthocentrinae) from Vietnam. Zootaxa 5647 (6): 571-583, DOI: 10.11646/zootaxa.5647.6.5, URL: https://doi.org/10.11646/zootaxa.5647.6.5
03FCAD28493FFF82FF37FCE8A3B2D70E.taxon	description	urn: lsid: zoobank. org: act: 942908 DD- 713 A- 405 B- 9 D 30 - 7 E 0478 C 24936 (Fig. 3)	en	Varga, Oleksandr (2025): First records of the genera Gnathochorisis Förster, 1869 and Symplecis Förster, 1869 (Hymenoptera, Ichneumonidae, Orthocentrinae) from Vietnam. Zootaxa 5647 (6): 571-583, DOI: 10.11646/zootaxa.5647.6.5, URL: https://doi.org/10.11646/zootaxa.5647.6.5
03FCAD28493FFF82FF37FCE8A3B2D70E.taxon	materials_examined	Material examined. Holotype: ♀, VIETNAM: Lam Dong, Bidoup Nuiba N. P., near Da Lat, 1650 – 1700 m, Malaise trap 1 – 12, 11 – 19. x. 2005, leg. C. v. Achterberg & R. de Vries, RMHN’ 05, RMNH. INS. 1722323 (RMNH).	en	Varga, Oleksandr (2025): First records of the genera Gnathochorisis Förster, 1869 and Symplecis Förster, 1869 (Hymenoptera, Ichneumonidae, Orthocentrinae) from Vietnam. Zootaxa 5647 (6): 571-583, DOI: 10.11646/zootaxa.5647.6.5, URL: https://doi.org/10.11646/zootaxa.5647.6.5
03FCAD28493FFF82FF37FCE8A3B2D70E.taxon	diagnosis	Diagnosis. Symplecis granulosus sp. nov. is characterized by the combination of the following characters: face dark brown, clypeus yellow (Fig. 3 B); antenna with 18 flagellomeres, first flagellomere 4.3 × as long as wide (Fig. 3 G); pronotum orange (Fig. 3 D); mesoscutum evenly and densely pubescent (Fig. 3 C); propodeum with area superomedia delimited by carinae, 1.6 × as long as wide, costulae present (Fig. 3 E); fore wing with vein 3 rs-m present; hind wing with nervellus not intercepted (Fig. 3 F); first metasomal tergite 2.3 × as long as maximum width; second tergite granulate, 1.2 × as long as apical width (Fig. 3 H); ovipositor 0.6 × as long as first tarsomere of hind tarsus. Symplecis granulosus sp. nov. differs from the other Oriental species by the granulate tergites 2 – 3 (second tergite longitudinally wrinkled and third smooth in the other Vietnamese species). It is similar to S. vietnamensis sp. nov. and S. anaeratabettensis Ranjith, Humala & Priyadarsanan, 2022 in having the completely carinated propodeum, but differs from both species by the more elongate area superomedia, which is 1.6 × as long as wide (1.2 – 1.3 × in S. vietnamensis sp. nov. and 1.0 × in S. anaeratabettensis) and brown-black face (yellow in S. vietnamensis sp. nov. and S. anaeratabettensis).	en	Varga, Oleksandr (2025): First records of the genera Gnathochorisis Förster, 1869 and Symplecis Förster, 1869 (Hymenoptera, Ichneumonidae, Orthocentrinae) from Vietnam. Zootaxa 5647 (6): 571-583, DOI: 10.11646/zootaxa.5647.6.5, URL: https://doi.org/10.11646/zootaxa.5647.6.5
03FCAD28493FFF82FF37FCE8A3B2D70E.taxon	description	Description. Female. Holotype (Fig. 3). Body length approximately 2.9 mm. Fore wing 2.3 mm. Head (Figs 3 B – D, G) smooth and sparsely pubescent. Antenna with 18 flagellomeres, first flagellomere 4.3 × as long as wide. Face about 0.9 × as long as wide, smooth and sparsely pubescent; eyes strongly convergent to clypeus, glabrous. Malar space short, 0.2 × the basal width of mandible; subocular sulcus absent. Clypeus 0.7 × as long as wide, weakly granulate. Mandible bidentate, not twisted, lower tooth shorter than upper tooth. Temples short and strongly narrowed behind eyes (dorsal view). Frons and vertex weakly granulate; length of the ocellar-ocular distance 1.3 × maximum diameter of lateral ocellus; occipital carina complete. Mesosoma (Figs 3 C – E) granulate and densely pubescent. Propleuron densely pubescent. Pronotum smooth and glabrous; epomia present, but not reaching mesoscutum. Mesoscutum densely and evenly pubescent; notauli deep anteriorly, with vertical carina. Scutellum densely pubescent. Mesopleuron granulate, densely pubescent dorsally and ventrally; epicnemial carina present; sternaulus deep on anterior 0.3 of mesopleuron. Metapleuron granulate, with longitudinal carina in lower third; pleural and submetapleural carinae present. Propodeum granulate, carinated; area superomedia elongate, 1.6 × as long as wide, costulae present; apophyses absent. Legs relatively slender; hind femur 5.1 × as long as wide; third tarsomere of hind tarsus 1.2 × as long as fifth tarsomere; tarsal claws simple. Wings (Fig. 3 E). Fore wing with vein 2 rs-m about 0.6 × the distance between 2 rs-m and 2 m-cu; vein 3 rs-m present, but partly unpigmented; vein 1 cu-a opposite to M & Rs; hind wing with nervellus not intercepted, vertical; distal abscissa of Cu absent. Metasoma (Fig. 3 H) longitudinally wrinkled anteriorly and sparsely pubescent. First tergite 2.3 × as long as maximum width, granulate, rugulose laterally; latero-median carinae indistinct. Second tergite 1.2 × as long as apical width, granulate, indistinctly longitudinally wrinkled. Third tergite granulate on anterior 0.8. The remaining tergites smooth. Ovipositor short, about 0.2 × as long as hind tibia and 0.6 × as long as first tarsomere of hind tarsus. Colour. Head brown-black except clypeus, mandible (except apices), scape and pedicel yellow. Mesosoma brown except propleuron, pronotum, tegula and subtegular ridge orange. Legs yellow except hind femur orange, tibia and tarsus brownish. Metasoma brown except second tergite apically yellow. Pterostigma and veins brown. Male. Unknown.	en	Varga, Oleksandr (2025): First records of the genera Gnathochorisis Förster, 1869 and Symplecis Förster, 1869 (Hymenoptera, Ichneumonidae, Orthocentrinae) from Vietnam. Zootaxa 5647 (6): 571-583, DOI: 10.11646/zootaxa.5647.6.5, URL: https://doi.org/10.11646/zootaxa.5647.6.5
03FCAD28493FFF82FF37FCE8A3B2D70E.taxon	etymology	Etymology. The specific epithet is from Latin ‘ grânulôsus’ refers to the granulate body sculpture of this species, especially meso-, metapleuron, propodeum and metasoma. Adjective.	en	Varga, Oleksandr (2025): First records of the genera Gnathochorisis Förster, 1869 and Symplecis Förster, 1869 (Hymenoptera, Ichneumonidae, Orthocentrinae) from Vietnam. Zootaxa 5647 (6): 571-583, DOI: 10.11646/zootaxa.5647.6.5, URL: https://doi.org/10.11646/zootaxa.5647.6.5
03FCAD28493FFF82FF37FCE8A3B2D70E.taxon	distribution	Distribution. Vietnam.	en	Varga, Oleksandr (2025): First records of the genera Gnathochorisis Förster, 1869 and Symplecis Förster, 1869 (Hymenoptera, Ichneumonidae, Orthocentrinae) from Vietnam. Zootaxa 5647 (6): 571-583, DOI: 10.11646/zootaxa.5647.6.5, URL: https://doi.org/10.11646/zootaxa.5647.6.5
03FCAD28493DFF80FF37FF48A3B2D00D.taxon	description	(Fig. 4)	en	Varga, Oleksandr (2025): First records of the genera Gnathochorisis Förster, 1869 and Symplecis Förster, 1869 (Hymenoptera, Ichneumonidae, Orthocentrinae) from Vietnam. Zootaxa 5647 (6): 571-583, DOI: 10.11646/zootaxa.5647.6.5, URL: https://doi.org/10.11646/zootaxa.5647.6.5
03FCAD28493DFF80FF37FF48A3B2D00D.taxon	materials_examined	Material examined. Holotype: ♀, VIETNAM: Tonkin, Hoang Lien N. R., 15 km W Sa Pa, ca. 1900 m, Malaise traps, 15 – 21. x. 1999, leg. C. v. Achterberg, RMHN’ 99, RMNH. INS. 1722324 (RMNH). Paratype: 1 ♀, same locality and date as for holotype ,, RMNH. INS. 1722325 (RMNH).	en	Varga, Oleksandr (2025): First records of the genera Gnathochorisis Förster, 1869 and Symplecis Förster, 1869 (Hymenoptera, Ichneumonidae, Orthocentrinae) from Vietnam. Zootaxa 5647 (6): 571-583, DOI: 10.11646/zootaxa.5647.6.5, URL: https://doi.org/10.11646/zootaxa.5647.6.5
03FCAD28493DFF80FF37FF48A3B2D00D.taxon	diagnosis	Diagnosis. Symplecis hoangliensis sp. nov. is characterized by the combination of the following characters: face and clypeus dark brown (Fig. 4 B); antenna with 18 flagellomeres, first flagellomere 6.1 – 6.3 × as long as wide (Fig. 4 G); pronotum orange (Fig. 4 D); mesoscutum densely pubescent centrally and glabrous laterally (Fig. 4 C); propodeum with area superomedia absent (Fig. 4 E); fore wing with vein 3 rs-m present; hind wing with nervellus not intercepted (Fig. 4 F); first metasomal tergite 2.6 × as long as maximum width; second tergite longitudinally wrinkled, 1.3 × as long as apical width (Fig. 4 G); ovipositor 0.4 – 0.5 × as long as first tarsomere of hind tarsus. Symplecis hoangliensis sp. nov. is similar to S. nudus sp. nov. in having the area superomedia absent, but differs by the densely pubescent centrally mesoscutum (entirely glabrous in S. nudus sp. nov.) and slightly shorter second tergite of metasoma, which is 1.3 × as long as apical width (1.4 – 1.5 × in S. nudus sp. nov.).	en	Varga, Oleksandr (2025): First records of the genera Gnathochorisis Förster, 1869 and Symplecis Förster, 1869 (Hymenoptera, Ichneumonidae, Orthocentrinae) from Vietnam. Zootaxa 5647 (6): 571-583, DOI: 10.11646/zootaxa.5647.6.5, URL: https://doi.org/10.11646/zootaxa.5647.6.5
03FCAD28493DFF80FF37FF48A3B2D00D.taxon	description	Description. Female. (Fig. 4). Body length approximately 3.9 – 4.0 mm. Fore wing 3.0 – 3.1 mm. Head (Figs 4 B – D) smooth and sparsely pubescent. Antenna with 18 flagellomeres, first flagellomere 6.1 – 6.3 × as long as wide. Face about 0.9 × as long as wide, smooth and sparsely pubescent; eyes strongly convergent to clypeus, glabrous. Malar space 0.5 × the basal width of mandible; subocular sulcus absent. Clypeus 0.7 × as long as wide, weakly granulate. Mandible bidentate, not twisted, lower tooth shorter than upper tooth. Temples short and strongly narrowed behind eyes (dorsal view). Frons and vertex smooth; length of the ocellar-ocular distance 1.0 × maximum diameter of lateral ocellus; occipital carina complete. Mesosoma (Figs 4 C – E) smooth and densely pubescent. Propleuron densely pubescent. Pronotum smooth and glabrous; epomia present, but not reaching mesoscutum. Mesoscutum densely and evenly pubescent centrally, glabrous laterally; notauli deep anteriorly, with vertical carina. Scutellum smooth. Mesopleuron smooth, densely pubescent dorsally and ventrally; epicnemial carina present; sternaulus deep on anterior 0.4 of mesopleuron. Metapleuron shiny, with traces of granulation, and with longitudinal carina in lower third; pleural and submetapleural carinae present. Propodeum smooth, with traces of granulation, lateral longitudinal and apical transverse carinae present; apophyses present, but weak. Legs relatively slender; hind femur 5.1 – 5.2 × as long as wide; third tarsomere of hind tarsus 1.3 × as long as fifth tarsomere; tarsal claws simple. Wings (Fig. 4 F). Fore wing with vein 2 rs-m about 1.0 × the distance between 2 rs-m and 2 m-cu; vein 3 rs-m present; vein 1 cu-a weakly basad to M & Rs; hind wing with nervellus not intercepted, slightly inclivous; distal abscissa of Cu absent. Metasoma (Fig. 4 H) longitudinally wrinkled anteriorly and sparsely pubescent. First tergite 2.6 – 2.7 × as long as maximum width, longitudinally wrinkled laterally; latero-median carinae present, but weak. Second tergite 1.3 × as long as apical width, longitudinally wrinkled. The remaining tergites smooth. Ovipositor short, about 0.15 × as long as hind tibia and 0.4 – 0.5 × as long as first tarsomere of hind tarsus. Colour. Head brown-black except mandible (except apices), scape and pedicel orange. Mesosoma black except propleuron, pronotum, tegula and subtegular ridge orange. Fore and mid legs orange except coxae, trochanters and trochantelli yellow; hind leg orange except femur and tibia partly and tarsus entirely brownish. Metasoma with tergites 1 – 2 black; the remaining tergites brown, except second tergite apically, third tergite narrowly basally and apically, and fourth apically yellow. Pterostigma and veins brown. Male. Unknown.	en	Varga, Oleksandr (2025): First records of the genera Gnathochorisis Förster, 1869 and Symplecis Förster, 1869 (Hymenoptera, Ichneumonidae, Orthocentrinae) from Vietnam. Zootaxa 5647 (6): 571-583, DOI: 10.11646/zootaxa.5647.6.5, URL: https://doi.org/10.11646/zootaxa.5647.6.5
03FCAD28493DFF80FF37FF48A3B2D00D.taxon	etymology	Etymology. The specific epithet consists of two words ‘ hoang’ and ‘ lien’ referring to the type locality of the new species situated in the Hoang Lien Nature Reserve. Adjective.	en	Varga, Oleksandr (2025): First records of the genera Gnathochorisis Förster, 1869 and Symplecis Förster, 1869 (Hymenoptera, Ichneumonidae, Orthocentrinae) from Vietnam. Zootaxa 5647 (6): 571-583, DOI: 10.11646/zootaxa.5647.6.5, URL: https://doi.org/10.11646/zootaxa.5647.6.5
03FCAD28493DFF80FF37FF48A3B2D00D.taxon	distribution	Distribution. Vietnam.	en	Varga, Oleksandr (2025): First records of the genera Gnathochorisis Förster, 1869 and Symplecis Förster, 1869 (Hymenoptera, Ichneumonidae, Orthocentrinae) from Vietnam. Zootaxa 5647 (6): 571-583, DOI: 10.11646/zootaxa.5647.6.5, URL: https://doi.org/10.11646/zootaxa.5647.6.5
03FCAD28493CFF8EFF37FA60A39CD3ED.taxon	description	urn: lsid: zoobank. org: act: 64 F 02 AC 7 - 3 E 32 - 4 AB 6 - AC 46 - 3093 D 9 B 0 F 6 D 1 (Fig. 5)	en	Varga, Oleksandr (2025): First records of the genera Gnathochorisis Förster, 1869 and Symplecis Förster, 1869 (Hymenoptera, Ichneumonidae, Orthocentrinae) from Vietnam. Zootaxa 5647 (6): 571-583, DOI: 10.11646/zootaxa.5647.6.5, URL: https://doi.org/10.11646/zootaxa.5647.6.5
03FCAD28493CFF8EFF37FA60A39CD3ED.taxon	materials_examined	Material examined. Holotype: ♀, VIETNAM: Tonkin, Hoang Lien N. R., 10 km SW Sa Pa, ca. 1550 m, Malaise traps, 22 – 29. x. 1999, leg. C. v. Achterberg, RMHN’ 99, RMNH. INS. 1722326 (RMNH). Paratypes: 2 ♀♀, same locality and date as for holotype, RMNH. INS. 1722327, RMNH. INS. 1722328 (RMNH). Other material: 1 ♀, idem, but, 15 km W Sa Pa, ca. 1900 m, Malaise trap, 15 – 21. x. 1999, leg. C. v. Achterberg, RMHN’ 99, RMNH. INS. 1722329 (RMNH).	en	Varga, Oleksandr (2025): First records of the genera Gnathochorisis Förster, 1869 and Symplecis Förster, 1869 (Hymenoptera, Ichneumonidae, Orthocentrinae) from Vietnam. Zootaxa 5647 (6): 571-583, DOI: 10.11646/zootaxa.5647.6.5, URL: https://doi.org/10.11646/zootaxa.5647.6.5
03FCAD28493CFF8EFF37FA60A39CD3ED.taxon	diagnosis	Diagnosis. Symplecis nudus sp. nov. is characterized by the combination of the following characters: face and clypeus dark brown (Fig. 5 B); antenna with 18 – 19 flagellomeres, first flagellomere 5.7 – 5.9 × as long as wide (Fig. 5 H); pronotum orange (Fig. 5 D); mesoscutum largely glabrous except of few setae anteriorly and along notaular lines (Fig. 5 E); propodeum with area superomedia absent (Fig. 5 F); fore wing with vein 3 rs-m present; hind wing with nervellus not intercepted (Fig. 5 G); first metasomal tergite 2.4 – 2.5 × as long as maximum width; second tergite longitudinally wrinkled, 1.4 – 1.5 × as long as apical width (Fig. 5 I); ovipositor 0.4 – 0.5 × as long as first tarsomere of hind tarsus. Symplecis nudus sp. nov. is similar to S. hoangliensis sp. nov. in having the area superomedia absent, but differs by the entirely glabrous meoscutum (densely pubescent centrally in S. hoangliensis sp. nov.) and slightly longer second tergite of metasoma, which is 1.4 – 1.5 × as long as apical width (1.3 × in S. hoangliensis sp. nov.). Symplecis nudus sp. nov. is similar to the Afrotropical S. glabroscutum Varga, 2023 and Holarctic S. glabra Dasch, 1992 in having entirely glabrous mesoscutum, but differs from both species by the not intercepted nervellus of the hind wing (intercepted below the middle in S. glabroscutum and S. glabra). In addition, S. nudus sp. nov. differs from in S. glabroscutum by the slenderer antenna, with first flagellomere 5.7 – 5.9 × as long as wide (4.8 × in S. glabroscutum) and darker second tergite of metasoma (entirely yellow in S. glabroscutum). Symplecis nudus sp. nov. differs from S. glabra also by the reduced carination of the propodeum (area basalis and area superomedia enclosed by the carinae in S. glabra).	en	Varga, Oleksandr (2025): First records of the genera Gnathochorisis Förster, 1869 and Symplecis Förster, 1869 (Hymenoptera, Ichneumonidae, Orthocentrinae) from Vietnam. Zootaxa 5647 (6): 571-583, DOI: 10.11646/zootaxa.5647.6.5, URL: https://doi.org/10.11646/zootaxa.5647.6.5
03FCAD28493CFF8EFF37FA60A39CD3ED.taxon	description	Description. Female. (Fig. 5). Body length approximately 2.9 – 3.0 mm. Fore wing 2.2 – 2.3 mm. Head (Figs 5 B – D, H) smooth and sparsely pubescent. Antenna with 18 – 19 flagellomeres, first flagellomere 5.7 – 5.9 × as long as wide. Face about 0.85 – 0.9 × as long as wide, smooth and sparsely pubescent; eyes strongly convergent to clypeus, glabrous. Malar space 0.3 – 0.4 × the basal width of mandible; subocular sulcus absent. Clypeus 0.7 × as long as wide, weakly granulate. Mandible bidentate, not twisted, lower tooth shorter than upper tooth. Temples short and strongly narrowed behind eyes (dorsal view). Frons and vertex smooth; length of the ocellar-ocular distance 1.1 × maximum diameter of lateral ocellus; occipital carina complete. Mesosoma (Figs 5 D – F) smooth and densely pubescent. Propleuron densely pubescent. Pronotum smooth and glabrous, weakly wrinkled posteriorly; epomia present, but not reaching mesoscutum. Mesoscutum sparsely pubescent anteriorly and along notaular lines; notauli deep anteriorly, with vertical carina. Scutellum smooth. Mesopleuron smooth, densely pubescent dorsally and ventrally; epicnemial carina present; sternaulus deep on anterior 0.4 of mesopleuron. Metapleuron shiny, with traces of granulation, and with longitudinal carina in lower third; pleural and submetapleural carinae present. Propodeum smooth, with traces of granulation, lateral longitudinal and apical transverse carinae present; apophyses present, but weak. Legs relatively slender; hind femur 4.8 – 5.2 × as long as wide; third tarsomere of hind tarsus 1.1 – 1.2 × as long as fifth tarsomere; tarsal claws simple. Wings (Fig. 5 G). Fore wing with vein 2 rs-m about 0.9 – 1.0 × the distance between 2 rs-m and 2 m-cu; vein 3 rs-m present; vein 1 cu-a weakly basad to M & Rs; hind wing with nervellus not intercepted, vertical; distal abscissa of Cu absent. Metasoma (Fig. 5 I) longitudinally wrinkled anteriorly and sparsely pubescent. First tergite 2.4 – 2.5 × as long as maximum width, longitudinally wrinkled laterally; latero-median carinae present, but weak. Second tergite 1.4 – 1.5 × as long as apical width, longitudinally wrinkled. The remaining tergites smooth. Ovipositor short, about 0.15 × as long as hind tibia and 0.4 – 0.5 × as long as first tarsomere of hind tarsus. Colour. Head brown-black except mandible (except apices), scape and pedicel yellow. Mesosoma black except propleuron, pronotum, tegula and subtegular ridge orange. Fore and mid legs orange except coxae, trochanters and trochantelli yellow; hind leg orange except coxa apically, trochanter and trochantellus partly femur and tibia partly and tarsus entirely brownish. Metasoma with tergites 1 – 2 black; the remaining tergites brown, except second tergite apically, third tergite narrowly basally and apically yellow. Pterostigma and veins brown. Male. Unknown.	en	Varga, Oleksandr (2025): First records of the genera Gnathochorisis Förster, 1869 and Symplecis Förster, 1869 (Hymenoptera, Ichneumonidae, Orthocentrinae) from Vietnam. Zootaxa 5647 (6): 571-583, DOI: 10.11646/zootaxa.5647.6.5, URL: https://doi.org/10.11646/zootaxa.5647.6.5
03FCAD28493CFF8EFF37FA60A39CD3ED.taxon	etymology	Etymology. ‘ Nudus’ from Latin means ‘ naked’. The new species is named after the glabrous mesoscutum. Adjective.	en	Varga, Oleksandr (2025): First records of the genera Gnathochorisis Förster, 1869 and Symplecis Förster, 1869 (Hymenoptera, Ichneumonidae, Orthocentrinae) from Vietnam. Zootaxa 5647 (6): 571-583, DOI: 10.11646/zootaxa.5647.6.5, URL: https://doi.org/10.11646/zootaxa.5647.6.5
03FCAD28493CFF8EFF37FA60A39CD3ED.taxon	distribution	Distribution. Vietnam.	en	Varga, Oleksandr (2025): First records of the genera Gnathochorisis Förster, 1869 and Symplecis Förster, 1869 (Hymenoptera, Ichneumonidae, Orthocentrinae) from Vietnam. Zootaxa 5647 (6): 571-583, DOI: 10.11646/zootaxa.5647.6.5, URL: https://doi.org/10.11646/zootaxa.5647.6.5
03FCAD28493CFF8EFF37FA60A39CD3ED.taxon	discussion	Remark. One female has relatively longer first flagellomere, about 7.0 × as long as wide, but otherwise is similar to the holotype. This specimen has both antennae missing except for the first few flagellomeres, thus it was not included in the type seria.	en	Varga, Oleksandr (2025): First records of the genera Gnathochorisis Förster, 1869 and Symplecis Förster, 1869 (Hymenoptera, Ichneumonidae, Orthocentrinae) from Vietnam. Zootaxa 5647 (6): 571-583, DOI: 10.11646/zootaxa.5647.6.5, URL: https://doi.org/10.11646/zootaxa.5647.6.5
03FCAD284932FF8CFF37F900A3B2D271.taxon	description	urn: lsid: zoobank. org: act: 686 D 6 F 52 - 0 E 3 F- 4 AEA- 89 FD-A 20 C 7696 F 2 D 0 (Fig. 6)	en	Varga, Oleksandr (2025): First records of the genera Gnathochorisis Förster, 1869 and Symplecis Förster, 1869 (Hymenoptera, Ichneumonidae, Orthocentrinae) from Vietnam. Zootaxa 5647 (6): 571-583, DOI: 10.11646/zootaxa.5647.6.5, URL: https://doi.org/10.11646/zootaxa.5647.6.5
03FCAD284932FF8CFF37F900A3B2D271.taxon	materials_examined	Material examined. Holotype: ♀, VIETNAM: Tonkin, Hoang Lien N. R., 10 km SW Sa Pa, ca. 1550 m, Malaise traps, 22 – 29. x. 1999, leg. C. v. Achterberg, RMHN’ 99, RMNH. INS. 1722330 (RMNH). Paratypes: 1 ♂, 1 ♀, the same locality as for holotype, but 15 km W Sa Pa, ca. 1900 m, Malaise traps, 15 – 21. x. 1999, leg. C. v. Achterberg, RMHN’ 99, RMNH. INS. 1722331, RMNH. INS. 1722332 (RMNH); 3 ♀♀, Lam Dong, Bidoup Nuiba N. P., near Da Lat, 1650 – 1700 m, Malaise trap 1 – 12, 11 – 19. x. 2005, leg. C. v. Achterberg & R. de Vries, RMHN’ 05, RMNH. INS. 1722333, RMNH. INS. 1722334, RMNH. INS. 1722335 (RMNH).	en	Varga, Oleksandr (2025): First records of the genera Gnathochorisis Förster, 1869 and Symplecis Förster, 1869 (Hymenoptera, Ichneumonidae, Orthocentrinae) from Vietnam. Zootaxa 5647 (6): 571-583, DOI: 10.11646/zootaxa.5647.6.5, URL: https://doi.org/10.11646/zootaxa.5647.6.5
03FCAD284932FF8CFF37F900A3B2D271.taxon	diagnosis	Diagnosis. Symplecis vietnamensis sp. nov. is characterized by the combination of the following characters: face, clypeus and malar space yellow (Fig. 6 B); antenna with 18 – 19 flagellomeres, first flagellomere 4.6 – 4.8 × as long as wide (Fig. 6 J); pronotum yellow (Fig. 6 E); mesoscutum evenly and densely pubescent (Fig. 6 G); propodeum with area superomedia delimited by carinae, 1.2 – 1.3 × as long as wide, costulae present (Fig. 6 H); fore wing with vein 3 rs-m present; hind wing with nervellus intercepted below the middle (Fig. 6 I); first metasomal tergite 2.5 – 2.7 × as long as maximum width; second tergite longitudinally wrinkled, 1.3 – 1.4 × as long as apical width (Fig. 6 K); ovipositor 0.5 – 0.6 × as long as first tarsomere of hind tarsus. Symplecis vietnamensis sp. nov. is similar to Palaearctic S. bicingulata (Gravenhorst, 1829) in having the yellow face, clypeus and pronotum, and evenly pubescent mesoscutum, but differs by the longer first flagellomere, which is 4.6 – 4.8 × as long as wide (3.6 – 3.7 × in S. bicingulata) and more elongate area superomedia, which is 1.2 – 1.3 × as long as wide (about 1.0 × in S. bicingulata). Symplecis vietnamensis sp. nov. resembles the Oriental S. anaeratabettensis in the colouration of face and pronotum, but differs by the black mesopleuron and mesoscutum (largely orange in S. anaeratabettensis) and more elongate area superomedia (about as long as wide in S. anaeratabettensis).	en	Varga, Oleksandr (2025): First records of the genera Gnathochorisis Förster, 1869 and Symplecis Förster, 1869 (Hymenoptera, Ichneumonidae, Orthocentrinae) from Vietnam. Zootaxa 5647 (6): 571-583, DOI: 10.11646/zootaxa.5647.6.5, URL: https://doi.org/10.11646/zootaxa.5647.6.5
03FCAD284932FF8CFF37F900A3B2D271.taxon	description	Description. Female. (Figs 6 A – B, E – K). Body length approximately 3.7 – 4.8 mm. Fore wing 3.0 – 4.0 mm. Head (Figs 6 B, E – F, J) smooth and sparsely pubescent. Antenna with 18 – 19 flagellomeres, first flagellomere 4.6 – 4.8 × as long as wide. Face about 0.8 – 0.9 × as long as wide, smooth and sparsely pubescent; eyes strongly convergent to clypeus, glabrous. Malar space short, 0.2 – 0.3 × the basal width of mandible; subocular sulcus absent. Clypeus 0.9 × as long as wide, weakly granulate. Mandible bidentate, not twisted, lower tooth shorter than upper tooth. Temples short and strongly narrowed behind eyes (dorsal view). Frons and vertex smooth; length of the ocellar-ocular distance 1.3 – 1.4 × maximum diameter of lateral ocellus; occipital carina complete. Mesosoma (Figs 6 E, G – H) smooth and densely pubescent. Propleuron densely pubescent. Pronotum smooth and glabrous; epomia present, but not reaching mesoscutum. Mesoscutum densely and evenly pubescent; notauli deep anteriorly. Scutellum smooth. Mesopleuron smooth, densely pubescent anteriorly and ventrally; epicnemial carina present; sternaulus deep on anterior 0.4 of mesopleuron. Metapleuron shiny, with traces of granulation, with longitudinal carina in lower third; pleural and submetapleural carinae present. Propodeum smooth, carinated; area superomedia elongate, 1.2 – 1.3 × as long as wide, costulae present; area basalis not delimited by carinae anteriorly; apophyses absent. Legs relatively slender; hind femur 5.5 – 5.9 × as long as wide; third tarsomere of hind tarsus 1.4 – 1.5 × as long as fifth tarsomere; tarsal claws simple. Wings (Fig. 6 I). Fore wing with vein 2 rs-m about 0.7 × the distance between 2 rs-m and 2 m-cu; vein 3 rs-m present; vein 1 cu-a weakly basad to M & Rs; hind wing with nervellus weakly intercepted below the middle, vertical; distal abscissa of Cu unpigmented. Metasoma (Fig. 6 K) longitudinally wrinkled anteriorly and sparsely pubescent. First tergite 2.5 – 2.7 × as long as maximum width, weakly granulate, longitudinally wrinkled laterally; latero-median carinae present, but weak. Second tergite 1.3 – 1.4 × as long as apical width, longitudinally wrinkled. The remaining tergites smooth. Ovipositor short, about 0.2 × as long as hind tibia and 0.5 – 0.6 × as long as first tarsomere of hind tarsus. Colour. Headblackexceptface, clypeus, malarspace, mandible (exceptapices), scapeandpedicelyellow. Mesosoma black except propleuron, pronotum, tegula and subtegular ridge yellow. Fore and mid legs orange except coxae, trochanters and trochantelli yellow to ivory; hind leg orange except coxa apically, trochanter and trochantellus partly, femur and tibia partly and tarsus entirely brownish. Metasoma with tergites 1 – 2 black; the remaining tergites brown, except second tergite apically, third tergite basally and apically, and fourth apically yellow. Pterostigma and veins brown. Male. Generally resembles female, but differs by the longer malar space, weakly narrowed eyes, present subocular sulcus, more abundant yellow colouration of head, and shorter first flagellomere, which is 3.7 × as long as wide (Figs 6 C – D).	en	Varga, Oleksandr (2025): First records of the genera Gnathochorisis Förster, 1869 and Symplecis Förster, 1869 (Hymenoptera, Ichneumonidae, Orthocentrinae) from Vietnam. Zootaxa 5647 (6): 571-583, DOI: 10.11646/zootaxa.5647.6.5, URL: https://doi.org/10.11646/zootaxa.5647.6.5
03FCAD284932FF8CFF37F900A3B2D271.taxon	etymology	Etymology. The new species is named after the type locality situated in Vietnam. Adjective.	en	Varga, Oleksandr (2025): First records of the genera Gnathochorisis Förster, 1869 and Symplecis Förster, 1869 (Hymenoptera, Ichneumonidae, Orthocentrinae) from Vietnam. Zootaxa 5647 (6): 571-583, DOI: 10.11646/zootaxa.5647.6.5, URL: https://doi.org/10.11646/zootaxa.5647.6.5
03FCAD284932FF8CFF37F900A3B2D271.taxon	distribution	Distribution. Vietnam.	en	Varga, Oleksandr (2025): First records of the genera Gnathochorisis Förster, 1869 and Symplecis Förster, 1869 (Hymenoptera, Ichneumonidae, Orthocentrinae) from Vietnam. Zootaxa 5647 (6): 571-583, DOI: 10.11646/zootaxa.5647.6.5, URL: https://doi.org/10.11646/zootaxa.5647.6.5
