identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03FA87F7FFCAFF0478EF9D33FEB5FD83.text	03FA87F7FFCAFF0478EF9D33FEB5FD83.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Spilogona hennigi Sorokina 2025	<div><p>Spilogona hennigi sp. nov.</p><p>Figs 1 A–E</p><p>Limnaricia pauxilla (Holmgren, 1869); Frey, 1915: 28 (West Taymyr, New-Siberian Islands). Misidentification.</p><p>Spilogona hirticauda (Malloch, 1921); Hennig, 1959b: 302 (West Taymyr, New-Siberian Islands). Misidentification.</p><p>Notes. Over the past 100 years, questions have been raised around three species: Spilogona hirticauda (Malloch, 1921), Spilogona obsoleta (Malloch, 1920), and Spilogona novaesibiriae (Frey, 1915) . Malloch (1921) described Melanochelia hirticauda from St. Paul Island, Alaska. Later, Huckett (1932) compared paratypes of Melanochelia hirticauda with the male holotype of Limnophora obsoleta and considered them identical species. However, Hennig (1959b) resurrected Spilogona hirticauda (Malloch) as a possibly valid species occurring in Alaska and the Far East of Russia, but this was not followed by Huckett (1965) and Pont (1986). Hennig’s doubts were caused by Frey’s paper (1915), where an unknown species was described and illustrated with the genitalia under the name “ Limnaricia pauxilla (Holmgren, 1869) ” (Fig. 1C, D). Collin (1930) illustrated a similar form of surstylus in an Alaskan specimen (Savonoki, Naknek Lake) under the name “ Limnophora obsoleta Malloch ”, and synonymized Spilogona pauxilla (Holmgren) with Spilogona megastoma (Boheman, 1866) . Hennig studied Frey’s specimens from West Taymyr and the New-Siberian Islands, as well as Collin’s drawings of the Alaskan specimen (Collin, 1930: Plate XIX, 16b1), and concluded that these specimens belong to Spilogona hirticauda . Hennig indicated that these specimens are very close to Spilogona novaesibiriae and Spilogona obsoleta but differ significantly from both in the male genitalia (Fig. 1E).</p><p>Relatively recently, Sorokina and Michelsen (2014) supported the view that Spilogona hirticauda (Malloch) is a synonym of Spilogona obsoleta and Spilogona novaesibiriae . This opinion, like the opinion of probably many other researchers, was based on the fact that Frey’s illustrations (1915: Taf. II, Figs 23–24) and Hennig’s illustrations (1959a: Taf. XV, Fig. 312) show Spilogona obsoleta from different perspectives.</p><p>However, I recently received muscid material for identification from Franz Josef Land. The specimens are very similar to Spilogona novaesibiriae . I examined the male genitalia and they are absolutely identical with Frey’s illustrations (1915: Taf. II, Figs 23-24) and Hennig’s illustrations (1959a: Taf. XV, Fig. 312), and they are completely different from the genitalia of Spilogona obsoleta from the same perspective. My first thought was that Hennig was right and Spilogona hirticauda is a valid species. Unfortunately, the type specimens of Spilogona hirticauda were not dissected and it is not possible at this time to study the type to confirm this statement. Huckett (1932) studied a paratype and noted that the description of Melanochelia hirticauda is entirely consistent with Spilogona obsoleta .</p><p>After studying Malloch’s description of Melanochelia hirticauda (1921), I found several distinctive differences from the specimens from Franz Josef Land and, therefore, from Hennig’s description. In addition to the structure of the male genitalia, these differences include the width of frons, the face in profile, the chaetotaxy of legs, and colour of female scutum. In addition, the locality of Melanochelia hirticauda is significantly further south than the locality of specimens from Franz Josef Land and of Frey’s specimens from the New Siberian Islands. I have thus concluded that the species discussed by Hennig is not Spilogona hirticauda but is a species new to science.</p><p>The new species appears therefore to be an exclusively arctic species that also inhabits the high Arctic of the Nearctic region, and so it may be found among series of Spilogona obsoleta in museum collections. As regards of the Alaskan specimen (Savonoski, Naknek Lake) described by Collin (1930: 272, Plate XIX, 16b 1), the surstylus is similar to the new species in its bend, but the shape is different: the surstylus does not protrude much above the epandrium; the cerci are also wide, as in Spilogona obsoleta . In view of these characters and the more southern locality of the Alaskan specimen, it is most unlikely that it belongs to the new species.</p><p>Type material: Holotype ♂, RUSSIA, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=52.8&amp;materialsCitation.latitude=80.333336" title="Search Plazi for locations around (long 52.8/lat 80.333336)">Franz Josef-Land</a>: Hooker Island, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=52.8&amp;materialsCitation.latitude=80.333336" title="Search Plazi for locations around (long 52.8/lat 80.333336)">stream</a>, 37 m, 80°20’N 52°48’E, 10.viii.2016, leg. A. Krasheninnikov (SZMN) . Paratypes: RUSSIA, Franz Josef Land: 1♂ 2♀♀, Hayes Island, Cape Observatorskii, tundra, 3.viii.2012, leg. I. Mizin ; 1♀, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=58.5&amp;materialsCitation.latitude=81.083336" title="Search Plazi for locations around (long 58.5/lat 81.083336)">Kane Island</a>, Cape Gagara (Loon), coastal tundra, 81°05’N 58°30’E, 9.viii.2013, leg. M. Gavrilo ; 3♀♀, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=52.8&amp;materialsCitation.latitude=80.333336" title="Search Plazi for locations around (long 52.8/lat 80.333336)">Hooker Island</a>, Tikhaya Bay, tundra, 80°20’N 52°48’E, 4, 5, 17.viii.2014, leg. M. Gavrilo ; 2♀♀, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=52.8&amp;materialsCitation.latitude=80.333336" title="Search Plazi for locations around (long 52.8/lat 80.333336)">Hooker Island</a>, stream, 80°20’N 52°48’E, 10.viii.2016, leg. A. Krasheninnikov ; 1♀, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=52.766666&amp;materialsCitation.latitude=80.333336" title="Search Plazi for locations around (long 52.766666/lat 80.333336)">Hooker Island</a>, Tikhaya Bay, polar station, 80°20’N 52°46’E, 41 m, 10.viii.2016, leg. A. Krasheninnikov ; 7♀♀, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=55.833332&amp;materialsCitation.latitude=80.36667" title="Search Plazi for locations around (long 55.833332/lat 80.36667)">Aldzhera Island</a>, stream, 80°22’N 55°50’E, 13 m, 17.viii.2016, leg. A. Krasheninnikov (all in SZMN).</p><p>Etymology. The species name is a patronym in honour of the famous dipterologist W. Hennig, who was the first to describe and illustrate it.</p><p>Diagnosis. The new species is very similar to Spilogona novaesibiriae (Frey, 1915) and can be separated from it as follows:</p><p>- Mid femur with 5–9 strong pv setae in basal half which are as long as diameter of femur or a little longer, without av setae; hind femur with short strong pv setae which are usually shorter than diameter of femur or sometimes a little longer; scutellum with downwardly-directed preapical setulae on upper border of declivities. Male: frontal vitta wide, distance between eye margins on upper part of frons as wide as frontal triangle or greater; surstylus semi-oval in shape, rounded at apex, without a platform with setae projecting above epandrium; cerci in lateral view broader, with convex inner margin. Female: scutum light, with three brown longitudinal stripes and sometimes with light brown dust between them.................... S. novaesibiriae</p><p>- Mid femur with a numerous long hair-like pv setae which are longer than diameter of femur and often with short hair-like setae on av-v surfaces; hind femur with elongate hair-like pv setae in basal half; scutellum without downwardly-directed preapical setulae on upper border of declivities. Male: frontal vitta narrower, distance between eye margins on upper part of frons narrower than frontal triangle; surstylus triangular in shape, pointed at apex, with a platform with setae projecting above epandrium; cerci in lateral view narrow, more or less straight. Female: scutum darker, with four black longitudinal stripes and brown dust between them................................................................. S. hennigi sp. nov.</p><p>Description. Hennig gave a good description of this species under the name “ Spilogona hirticauda ” and pointed out its great similarity to Spilogona novaesibiriae (Hennig, 1959b: 302) . All the differences found between these two species are indicated in the diagnosis above.</p><p>As further differences, Hennig in his key (1959a: 260) also described a white wing and a completely yellow haltere for Spilogona novaesibiriae but a yellowish wing and a brown-black haltere for “ Spilogona hirticauda ”. However, these characters are variable as the series of Spilogona novaesibiriae as well as in the series of Spilogona hennigi sp. nov.</p><p>Comment. Because the type series has been extracted and pinned from alcohol, the flies are not in very good condition, which makes description difficult.</p><p>Distribution. PALAEARCTIC: Russia (Arctic Siberia: Franz Josef Land, West Taymyr, New-Siberian Islands).?NEARCTIC.</p></div>	https://treatment.plazi.org/id/03FA87F7FFCAFF0478EF9D33FEB5FD83	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sorokina, Vera S.	Sorokina, Vera S. (2025): New taxonomic notes on the genus Spilogona Schnabl, 1911 (Diptera: Muscidae) in Arctic Russia, with the description of three new species. Zootaxa 5584 (1): 1-25, DOI: 10.11646/zootaxa.5584.1.1, URL: https://doi.org/10.11646/zootaxa.5584.1.1
03FA87F7FFC8FF0A78EF9E7EFC15FDA7.text	03FA87F7FFC8FF0A78EF9E7EFC15FDA7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Spilogona gorodkovi Sorokina 2025	<div><p>Spilogona gorodkovi sp. nov.</p><p>Figs 2 A–I</p><p>Type material: Holotype ♂, RUSSIA, Wrangel Island: 4 km N Somnitil’naya Bay, valley of the River Somnitil’naya, on flowers of Potentilla emarginata, 28.vii.1972, leg. K. Gorodkov (ZISP) . Paratypes: RUSSIA, Wrangel Island: 2♂♂ 2♀♀, same data as holotype (SZMN) ; 3♀♀, 5 km N Somnitil’naya Bay, valley of the River Somnitil’naya, 150 m, 25.vii.1972, leg. K. Gorodkov (SZMN) ; 1♂, Somnitil’naya Bay, south of island, bank of the River Somnitil’naya, on flowers of Potentilla emarginata, 26.vii.1966, leg. K. Gorodkov (ZISP) ; 1♂ 1♀, Somnitil’naya Bay, south of island, tundra, on flowers of Dryas integrifolia, 19.vii.1966, leg. K. Gorodkov (ZISP) ; 5♀♀, 5 km N Somnitil’naya Bay, south of island, middle part of the River Somnitil’naya, Mineeva Mts, 23.vii.1971, leg. K. Gorodkov (ZISP) ; 1♂, 15 km W Rodzhers Bay, south of island, 18.vii.1971, leg. K. Gorodkov (ZISP) .</p><p>Etymology. The species name is a patronym in honour of the celebrated Soviet dipterist Kirill B. Gorodkov (Russia), who collected the type series of this species as well many other new Muscidae .</p><p>Diagnosis. The new species is similar to Nearctic Spilogona rufitarsis (Stein, 1920) because of 4 pairs of postsutural dorsocentral setae, facial edge not projecting beyond level of profrons, broad frons, anepisternum without interspatial setae, mid tibia without v setae, hind femur with pv setae on proximal half, scutellum without downwardly-directed preapical setulae on upper border of declivities, grey abdomen with black spots and a narrow brown median stripe between them. But the new species differs from Spilogona rufitarsis by the wider frons, dusted prementum, black fore tibia and tarsi, darker colour of thorax, blackish haltere, wide parafacial which is 1.2–2.0 times width of postpedicel at level of base of antenna, and postgenital plate with spinules.</p><p>Description: Male: body 5.6–5.8 mm, wing 5.0– 5.2 mm.</p><p>Head: Ground-colour black. Frons, fronto-orbital plate, gena and parafacial shining silvery-grey, with dark reflections when viewed in profile. In lateral view, facial edge not projecting beyond level of profrons (Fig. 2A). Eye large, bare or with very sparse and inconspicuous hairs. Fronto-orbital plates separated by a wide black frontal vitta. Distance between eye margins on upper part of frons about as wide as distance between the first pair of dc setae, sometimes a little narrower. Frontal triangle small, grey dusted, not extending to middle of frons. Frons with 8–9 pairs of strong frontal setae and hair-like setae between them. Parafacial at level of base of antenna 1.2–1.3 times width of postpedicel, very weakly narrowing below. Height of gena about 1/4 of height of eye and 1.8–2 times width of postpedicel.Antenna black, postpedicel approximately 1.5 times as long as wide. Arista black, thickened at base, short pubescent, with the longest individual hairs shorter than basal diameter of arista. Palpus black. Proboscis short, thin, prementum broadly dusted in middle. Labella large.</p><p>Thorax: Ground-colour black, densely grey dusted, with three distinct brown longitudinal stripes widening diffusely to scutellum (Fig. 2B). Presutural acr setae in 2 even rows, 2+4 dc setae. Notopleuron bare, or with individual hairs in upper part. Anepisternum without interspatial setae. Katepisternal setae 1+1. Scutellum brown dusted except grey at apex, without downwardly-directed preapical setulae on upper border of declivities.</p><p>Wing: Light brownish tinged. Costa without spinules or costal spine. Calypters whitish, lower one with broadly yellow margins. Haltere brown with blackish knob.</p><p>Legs: Ground colour black, grey dusted. Fore tibia with 1 p seta. Mid femur with a row of hair-like pv and avv setae, which become more numerous and elongate towards base of pv surface, with 2 pd–p preapical setae. Mid tibia with 0–1 ad and 3 pd setae, without any v setae. Hind femur with 6–7 strong av setae in distal third and with hair-like setae at base on av-v-pv surfaces which are longer on pv surface. Hind tibia with 1–3 short av, 4–5 ad, 3–4 pd, without pv setae.</p><p>Abdomen: Black with dense grey dust, conical. Tergites 1+2–4 with black triangular spots separated by a narrow brown median stripe, tergite 5 with a diffuse brown median stripe. Sternite 1 with hairs, sternite 5 somewhat protruding, a little longer then sternite 4, with a deep median notch and a pointed caudal margin, with shining apex (Figs 2E, I).</p><p>Terminalia: In lateral view cerci wide, curved, sharply tapered towards the apex; surstylus rounded at apex, with hairs inside, not longer then cerci; epandrium large, hemispherical; hypandrium 2 times shorter then epandrium (Figs 2D, G); cercal plate triangular (Fig. 2H).</p><p>Female: body 5.8–6.5 mm, wing 5.5–5.8 mm.</p><p>Similar to male but differing as follows:</p><p>Head: Frons, fronto-orbital plate, gena and parafacial grey or brown dusted (Fig. 2C). Frontal triangle brownish dusted, extending to middle of frons. Width of frontal vitta 3–4 times width of a fronto-orbital plate. Fronto-orbital plate with 2 pairs of orbital setae. Parafacial at level of base of antenna 2 times width of postpedicel. Height of gena about 1/3 of height of eye. Proboscis thickened.</p><p>Thorax: Scutum more matt. Scutellum with brown spots on lateral angles.</p><p>Legs: Mid femur with a row of hair-like pv which become more elongate towards base; only several short av-v hairs at base. Mid tibia with 0 ad and 2–3 pd. Hind femur with 4–5 strong av setae in distal third; hair-like setae at base short and sparse on av-v surface, longer on pv surface. Hind tibia with 3–4 ad, 2–3 pd pv setae.</p><p>Abdomen: Postgenital plate with spinules and hairs (Fig. 2F).</p><p>Distribution. PALAEARCTIC: Arctic Russia (Wrangel Island).</p></div>	https://treatment.plazi.org/id/03FA87F7FFC8FF0A78EF9E7EFC15FDA7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sorokina, Vera S.	Sorokina, Vera S. (2025): New taxonomic notes on the genus Spilogona Schnabl, 1911 (Diptera: Muscidae) in Arctic Russia, with the description of three new species. Zootaxa 5584 (1): 1-25, DOI: 10.11646/zootaxa.5584.1.1, URL: https://doi.org/10.11646/zootaxa.5584.1.1
03FA87F7FFC6FF0878EF9E9AFCC3FB5F.text	03FA87F7FFC6FF0878EF9E9AFCC3FB5F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Spilogona latizygoma Sorokina 2025	<div><p>Spilogona latizygoma sp. nov.</p><p>Figs 3 A–I</p><p>Spilogona tendipes (Malloch, 1920); Sorokina &amp; Michelsen 2014: 516. Wrangel Island. Misidentification.</p><p>Type material: Holotype ♂, RUSSIA, Wrangel Island: Somnitil’naya Bay, valley of the River Somnitil’naya, Mineeva Mts, 150 м, on flowers of Potentilla emarginata, 19.vii.1966, leg. K. Gorodkov (ZISP).</p><p>Paratypes: RUSSIA, Wrangel Island: 1♀, Somnitil’naya Bay, valley of the River Somnitil’naya, south of island, Mineeva Mts, 250 м, 22.vii.1966, leg. K. Gorodkov (ZISP) ; 2♀♀, Somnitil’naya Bay, south of island, 22, 26.vii.1966, leg. K. Gorodkov (SZMN) ; 2♂♂ 4♀♀, middle part of the <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-179.15&amp;materialsCitation.latitude=71.01667" title="Search Plazi for locations around (long -179.15/lat 71.01667)">River Neozhidannaya</a>, 71°01’N 179°09’W, yellow pan traps, 12–16.vii.2016, leg. L. Volkova (SZMN, 1♂ — BUIC-DIP1761, 1♀ — BUIC-DIP1762) ; 1♀, middle part of the <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-179.13333&amp;materialsCitation.latitude=71.01667" title="Search Plazi for locations around (long -179.13333/lat 71.01667)">River Neozhidannaya</a>, valley of river, 71°01’N 179°08’W, yellow pan traps, 10.vi.–27.vii.2016, leg. L. Khruleva (SZMN) ; 1♀, middle part of the <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-179.15&amp;materialsCitation.latitude=71.01667" title="Search Plazi for locations around (long -179.15/lat 71.01667)">River Neozhidannaya</a>, 71°01’N 179°09’W, yellow pan traps, 8, 9.vii.2019, leg. L. Khruleva (SZMN) ; 2♂♂ 1♀, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-179.88333&amp;materialsCitation.latitude=71.26667" title="Search Plazi for locations around (long -179.88333/lat 71.26667)">Tundrovii Mt.</a>, 71°16’N 179°53’W, yellow pan traps, 8–10.vii.2018, leg. U. Babii (SZMN) ; 2♀♀, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-179.8&amp;materialsCitation.latitude=71.3" title="Search Plazi for locations around (long -179.8/lat 71.3)">Tundrovii Mt.</a>, 71°18’N 179°48’W, yellow pan traps, 7–11.vii.2019, leg. U. Babii &amp; O. Khruleva (SZMN) ; 1♀, middle part of the <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-179.75&amp;materialsCitation.latitude=71.166664" title="Search Plazi for locations around (long -179.75/lat 71.166664)">River Mamontovaya</a>, 71°10’N 179°45’W, yellow pan traps, 2–7.vii.2015, leg. O. Khruleva (SZMN, BUIC-DIP1763) ; 1♀, upper reaches of the <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-170.31667&amp;materialsCitation.latitude=71.21667" title="Search Plazi for locations around (long -170.31667/lat 71.21667)">River Neizvestnaya</a>, 71º13’N 170º19’W, yellow pan traps, 4–11.vii.2015, leg. O. Khruleva (SZMN) .</p><p>Etymology. The species name is based on the Latin words “latus” and “zygoma” meaning “wide cheekbones”, because the parafacial is very broad.</p><p>Diagnosis. The new species is very similar to Spilogona tendipes (Malloch, 1920) because of its large size, covering of dense grey dusting with a bluish or greenish tinge and having paired dark marks visible on abdomen, a projecting lower face, small eyes, very broad parafacial, long and slender haustellum with a shiny prementum and slender and elongate labella, 4 pairs of postsutural dorsocentral setae, hind tarsus with a v seta near base, and hypoproct with recurrent spines. But the new species differs substantially from Spilogona tendipes by the absence of ventral seta on mid tibia, distinct long pv in basal half of hind femur, numerous hair-like av setae in basal half of all femora, long ground-setulae of scutum, long setae on sternites 3–5, more greenish tinge of body, almost inconspicuous hairs on eye, and the male terminalia.</p><p>Description: Male: body 5.3–5.7 mm, wing 4.8–5.0 mm.</p><p>Head: Ground-colour black. Frons, fronto-orbital plate, gena and parafacial shining silvery-grey or silverybrown, with dark reflections when viewed in profile. In lateral view, facial edge projecting beyond level of profrons (Fig. 3A). Eye small, bare or with very sparse and inconspicuous hairs. Fronto-orbital plates separated by a wide frontal vitta. Distance between eye margins on upper part of frons about as wide as ocellar triangle (Fig. 3B). Frontal triangle extending to middle of frons. Frons with 12–14 pairs of strong frontal setae. Parafacial at level of base of antenna 1.5–2 times width of postpedicel, not narrowing below. Height of gena about 1/3–1/4 of height of eye and 1.5–2 times width of postpedicel. Antenna black, postpedicel approximately 1.5 times as long as wide. Arista black, bare, thickened in basal half. Palpus black. Proboscis long, slender, prementum shining. Labella slender and elongate, crocheted relative to the proboscis.</p><p>Thorax: Ground-colour black, densely grey dusted with a bluish or greenish tinge, with two brown indistinct presutural longitudinal stripes when viewed from in front. Ground-setulae of scutum relatively long, hair-like. Presutural acr setae in 3–4 uneven rows, 2+4 dc setae. Notopleuron bare, or with individual hairs. Anepisternum with 2–3 interspatial setae. Katepisternal setae 1+1. Scutellum grey dusted, with downwardly-directed preapical setulae on upper border of declivities.</p><p>Wing: Light brownish tinged, with dark base. Costa without spinules or costal spine. Calypters whitish, with light yellow margins. Haltere brown with blackish knob.</p><p>Legs: Ground colour black, grey dusted. Fore tibia with 2–4 p setae. Fore femur with numerous long hair-like setae on posterior surface, with a row of short hair-like av setae. Mid femur with a row of long pv setae and with a row of short hair-like av-v setae, with 2 pd–p preapical setae. Mid tibia with 2–3 ad and 3 pd setae, without any v setae. Hind femur with a row of long av setae becoming shorter on basal third, with a row of long pv setae in basal half, with numerous long hair like setae on basal part of posterior surface. Hind tibia with 3–5 av, 4–6 ad, with an uneven row of long pd-p setae in middle part, without pv setae. First tarsomere of hind tarsus with strong v seta near base.</p><p>Abdomen: Black with dense bluish-grey dust, conical. Tergites 1+2–4 with large brown triangular marks separated by a brown median stripe, sometimes fused, tergite 5 with a brown median stripe. Sternite 1 bare or with individual hairs, sternites 2–5 with numerous long setae, especially sternite 5, sternite 5 large, longer then sternite 4, with a deep median notch, and a pointed caudal margin (Figs 3E, I).</p><p>Terminalia: In lateral view cerci wide in proximal half and narrow, rod-shaped in distal half, curved; surstylus rounded at apex, with a notch in ventral margin, shorter than cerci; epandrium large, hemispherical; hypandrium 2 times shorter then epandrium, with a wavy ventral margin (Figs 3D, G). Cercal plate oval with two apical prolongations (Fig. 3H).</p><p>Female: body 6.2–7.3 mm, wing 5.7–6.7 mm.</p><p>Similar to male but differing as follows:</p><p>Head: Frons, fronto-orbital plate, gena and parafacial densely brown dusted, contrasting with the greenish-bluish-grey occiput (Fig. 3C). Frontal vitta matt grey, contrasting with brown fronto-orbital plate and frontal triangle when viewed from above. Width of frontal vitta 2–2.3 times width of a fronto-orbital plate. Frontal triangle brown dusted, extending to lunule. Fronto-orbital plate with an uneven row of frontal and orbital setae. Parafacial at level of base of antenna 2–2.3 times width of postpedicel. Height of gena about 1/2–1/3 of height of eye.</p><p>Thorax: Colour lighter, with greenish-bluish-grey dust. Scutum with 3 brown longitudinal stripes, sometimes indistinct, not extending to scutellum.</p><p>Abdomen: Postgenital plate with weak individual spinules.</p><p>Distribution. PALAEARCTIC: Arctic Russia (Wrangel Island).</p><p>DNA barcode. BOLD BIN: BOLD:ADZ7218. See Table 1 for GenBank accession numbers.</p><p>Remarks. Spilogona latizygoma morphological is very similar to Spilogona tendipes . But based on DNA barcodes, these species are genetically different; the p-distance between them is 3.36%. The intraspecific distance of Spilogona latizygoma specimens ranges from 0.0% to 0.16%.</p></div>	https://treatment.plazi.org/id/03FA87F7FFC6FF0878EF9E9AFCC3FB5F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sorokina, Vera S.	Sorokina, Vera S. (2025): New taxonomic notes on the genus Spilogona Schnabl, 1911 (Diptera: Muscidae) in Arctic Russia, with the description of three new species. Zootaxa 5584 (1): 1-25, DOI: 10.11646/zootaxa.5584.1.1, URL: https://doi.org/10.11646/zootaxa.5584.1.1
03FA87F7FFC4FF0978EF994AFF76FB5B.text	03FA87F7FFC4FF0978EF994AFF76FB5B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Spilogona ciliatocosta (Schnabl 1915)	<div><p>Spilogona ciliatocosta (Schnabl, 1915)</p><p>Limnophora (Spilogona) ciliatocosta Schnabl in Becker et al., 1915: 31. Type-locality: “in der Karskaja Tundra” [RUSSIA, West Siberia, Yugorskiy peninsula, Kara district; vii.1909; collector F. Zaitsev] [seen].</p><p>Spilogona contigua Huckett, 1965: 18, 190, 201 (key), and 216. Type-locality: “ Spence Bay, Northwest Territories, 7.14.51 [i.e. 14.vii.1951], J.G. Chillcott ” [CANADA, NWT] [seen]. Syn. nov.</p><p>Spilogona ciliatocosta (Schnabl, 1915); Sorokina, 2012a: 331 (Taymyr Peninsula); Sorokina &amp; Thidrikh, 2021: 219 (Chukotka AO, incl. Wrangel Island) .</p><p>Notes. The type material of Limnophora ciliatocosta (in ZISP) and Spilogona contigua (in CNC) was examined. The descriptions of both species and the male terminalia are the same. However, the illustrations of the male terminalia of Spilogona ciliatocosta by Hennig (1959a: Taf. XV, Fig. 299) and of Spilogona contigua by Huckett (1965: 352, Fig. 165) are slightly different in the width of the surstylus. I examined many specimens of Palaearctic Spilogona ciliatocosta and Nearctic Spilogona contigua and I have concluded that the width of the surstylus depends on the angle of viewing the terminalia and is variable. The cercal plate of these species is very distinctive because of the different length of right and left cerci (Huckett, 1965: 346, Fig. 96). Hennig in his monograph illustrated the male terminalia (Hennig, 1959a: Taf. XIII, Fig. 276; 1959b: Taf. XVIII, Fig. 361) of a specimen from Wrangel Island named as “ Limnophora macropyga Frey ” (det. A. Stackelberg). These figures correspond well with both ciliatocosta and contigua . I examined this specimen and found that the terminalia is mounted on a permanent slide preparation, which is why it is slightly compressed, and the surstylus thus appears wider than that of the Hennig’s figure of the type of Limnophora ciliatocosta . Unfortunately, the terminalia of the syntype of Limnophora ciliatocosta is also mounted on a poorly prepared permanent mount, and it was impossible to describe the form and exact width of the surstylus.</p><p>Specimens of both sexes of these species are distinctive, with a completely black body, facial edge projecting beyond level of profrons, mid tibia with v setae and hind femur without pv setae, scutellum with downwardly-directed preapical setulae on upper border of declivities, shining prementum, and costa with costal spinules. I therefore conclude that Spilogona contigua Huckett, 1965 is a new junior synonym of Limnophora ciliatocosta Schnabl, 1915 .</p><p>In the ZISP, apart from the male syntype of Limnophora ciliatocosta there is a female named as Limnophora ciliatocosta . The locality is the same as the male (Karskaja Tundra). However, this specimen does not agree with the male syntype of Limnophora ciliatocosta . Hennig (1959a) also had doubts about the identity of this female. The female syntype of Limnophora ciliatocosta does not have the facial edge projecting beyond level of profrons, costa with very strong costal spines, mid tibia without v setae and hind femur with several pv setae, proboscis short and thickened, prementum dusted, body with grey dust, not black, thorax grey with three distinct brown longitudinal stripes and with 2+3 dc setae, abdomen wet, but with traces of paired brown spots, postgenital plate with hairs. This female is definitely a different species and the specimen runs to Spilogona tornensis (Ringdahl, 1926) .</p><p>Spilogona ciliatocosta is very similar to Spilogona tundrae (Schnabl, 1915) . These species can be separated as following couple:</p><p>1 Larger species (5–6 mm), costal setulae weak. Male: fore tibia with p seta in middle; fronto-orbital plates separated by a wide frontal vitta, which is as wide as width of outer margins of ocelli or wider; abdomen shining; anal segment swollen, when viewed in profile bulging towards the dorsal region. Female: abdomen blackish............................. S. tundrae</p><p>- Smaller species (4–4.5 mm), costal setulae coarsened proximad of costal spine. Male: fore tibia without p seta in middle; fronto-orbital plates touching or separated by a narrow frontal vitta; abdomen subshining; anal segment flattish or appressed dorsad, not bulging toward the dorsal region. Female: abdomen bronzy....................................... S. ciliatocosta</p><p>All of the characters given above for Spilogona ciliatocosta and Spilogona tundrae are variable, but are more commonly as stated in the couplet. The male terminalia provides the most reliable identification character.</p><p>Distribution. PALAEARCTIC: Russia (Arctic Siberia and Far East). NEARCTIC: Alaska, Yukon, NWT, Quebec.</p></div>	https://treatment.plazi.org/id/03FA87F7FFC4FF0978EF994AFF76FB5B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sorokina, Vera S.	Sorokina, Vera S. (2025): New taxonomic notes on the genus Spilogona Schnabl, 1911 (Diptera: Muscidae) in Arctic Russia, with the description of three new species. Zootaxa 5584 (1): 1-25, DOI: 10.11646/zootaxa.5584.1.1, URL: https://doi.org/10.11646/zootaxa.5584.1.1
03FA87F7FFC5FF0E78EF9906FB58FE4B.text	03FA87F7FFC5FF0E78EF9906FB58FE4B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Spilogona churchillensis Huchett 1965	<div><p>Spilogona churchillensis Huchett, 1965</p><p>Figs 4 A–C</p><p>Spilogona churchillensis Huckett, 1965: 213 . Type-locality: “Churchill, Manitoba, 7.22.47 [i.e. 22.vii.1947], T. N. Freeman ” [CANADA, Manitoba] [seen].</p><p>Spilogona quinquesetosa (Schnabl, 1915); Sorokina, 2012b: 212 (Altai Mts); Sorokina, 2018: 244 (Altai Mts). Misidentification.</p><p>Notes. Spilogona churchillensis was synonymized with Spilogona quinquesetosa in 2018 (Sorokina, 2018). However, the re-examination of the types of both species and DNA analyses shows that Spilogona churchillensis differs from Spilogona quinquesetosa . The holotype of Spilogona quinquesetosa is in bad condition: without abdomen and mid and hind legs. According to the original descriptions and examination of the types some morphological differences between these two species have been found:</p><p>1 Scutellum without downwardly-directed preapical setulae on upper border of declivities; posthumeral and notopleural regions pale grey, in contract to darker adjoining surface; mid tibia usually with one v seta; hind femur with short pv setae in basal 1/3, sometimes these setae elongated; sternite 5 with a wide rectangular median notch (Fig. 4C); male cerci narrower (Fig. 4A, B)....................................................................................... S. churchillensis</p><p>- Scutellum with downwardly-directed preapical setulae on upper border of declivities; posthumeral and notopleural regions dark brown; mid tibia usually with two or more v setae; hind femur without pv setae; sternite 5 with a deep triangular median notch (Fig. 4D); male cerci broader................................................................ S. quinquesetosa</p><p>The DNA barcoding confirmed the differences between Spilogona churchillensis and Spilogona quinquesetosa . The DNA barcodes for specimens under the name “ quinquesetosa ” from the Chukotka population and the Altai population were very different, with an intraspecific p-distance of from 4.26% to 4.59%. Altai specimens of “ quinquesetosa ” (Sorokina, 2018) proved to be genetically closest to Spilogona churchillensis from Manitoba, Canada (BOLD).</p><p>Thus, Spilogona churchillensis Huckett, 1965 is valid species and is not a junior synonym of Spilogona quinquesetosa (Schnabl, 1915) . Spilogona churchillensis has an arcto-alpine distribution, whereas Spilogona quinquesetosa has an arctic distribution.</p><p>Distribution. PALAEARCTIC: Siberia (Altai Mts). NEARCTIC: Manitoba.</p><p>DNA barcode. BOLD BIN: BOLD:AAB7605. See Table 1 for GenBank accession numbers.</p></div>	https://treatment.plazi.org/id/03FA87F7FFC5FF0E78EF9906FB58FE4B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sorokina, Vera S.	Sorokina, Vera S. (2025): New taxonomic notes on the genus Spilogona Schnabl, 1911 (Diptera: Muscidae) in Arctic Russia, with the description of three new species. Zootaxa 5584 (1): 1-25, DOI: 10.11646/zootaxa.5584.1.1, URL: https://doi.org/10.11646/zootaxa.5584.1.1
03FA87F7FFC3FF0F78EF9C83FE7DF9FE.text	03FA87F7FFC3FF0F78EF9C83FE7DF9FE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Spilogona cordyluraeformis (Schnabl 1915)	<div><p>Spilogona cordyluraeformis (Schnabl, 1915)</p><p>Figs 4 E–K</p><p>Scatocoenosia cordyluraeformis Schnabl in Becker et al., 1915: 3. Type-locality: “in der Karskaja Tundra” [RUSSIA, West Siberia, Yugorskiy Peninsula, Kara district; actually Yamalo-Nenetskiy AO, east slope of Polar Urals, Mt Minisey, 180 m, 21.vii.1909, F. Zaitsev] [seen].</p><p>Spilogona albinepennis Huckett, 1965: 210 . Type-locality: “ Spence Bay, Northwest Territories, 7.30 ... 51 [i.e. 30.vii.1951] (A.E.R. Downe)” [CANADA, NWT] [seen]. Syn. nov.</p><p>Spilogona dorsostriata Huckett, 1965: 217 . Type-locality: “Point Barrow, Alaska, 7.18.52 [i.e. 18.vii.1952]” [USA, Alaska; collector P.D. Hurd] [seen]. Syn. nov.</p><p>Spilogona cordyluraeformis (Schnabl, 1915); Sorokina &amp; Pont, 2013: 581 (Polar Urals, Cape Otto Schmidt, Wrangel Island); Sorokina, 2017: 53 (Yamalo-Nenez AO, Taymyr, Wrangel Island).</p><p>Spilogona sp. 1; Sorokina &amp; Khruleva, 2012: 557 (Wrangel Island); Sorokina &amp; Tridrikh, 2021: 221 (Cape Otto Schmidt).</p><p>Notes. Huckett (1965) described two very similar species from different populations of the arctic coastal regions: Spilogona albinepennis (Nunavut /NWT, Quebec) and Spilogona dorsostriata (Alaska). Both species are very distinctive, with a very broad frons in male which is about as broad as in female, very small size, with small yellow subequal calypters, black unmarked abdomen with subshining margins, often with subshining/shining stripes on scutum, and whitish wings in male and brownish tinged along veins in female.</p><p>According to Huckett (1965: 217), Spilogona dorsostriata differs from Spilogona albinepennis by “four black glossy stripes on mesonotum, which often fuse or merge together, thus partly blurring their identity as separate stripes”. In my opinion, the pattern of scutal stripes cannot be a reliable character for describing a new species, given that reliable characters such as the width of the frons, small subequal calypters etc. are the same in both species. The types of both species were examined by the author in Canadian National Collection. Additionally, 77 females and 74 males with the same characters were found in unsorted material from different populations in Canada (Yukon, NWT, Quebec). It was impossible to separate these specimens into two species, as the dusting varied greatly, from completely matt, to glossy streaks, to narrow glossy stripes and to broad glossy marks (Figs 4E–K). I also dissected the male terminalia in specimens from different populations with different scutal markings, and they were the same .</p><p>The descriptions and the male terminalia of Spilogona dorsostriata and Spilogona albinepennis are identical with those in the recently discussed Spilogona cordyluraeformis . The differences between all these species consist only in the different degrees of brown pollination on the scutum. Scatocoenosia cordyluraeformis was described only from the female and for a long time raised questions as to its correct family assignment. A large series of males and females of this species from Cape Otto Schmidt, Taymyr and Wrangel Island enabled us to resolve these issues (Sorokina &amp; Pont, 2013; Sorokina &amp; Tridrikh, 2021). The female holotype of Scatocoenosia cordyluraeformis has a shining scutum with a median dusted vitta. Specimens from Taymyr have a shining, or subshining, or dusted scutum; specimens from Wrangel Island mostly have a completely dusted scutum; specimens from Cape Otto Schmidt have a completely shining scutum, or with a narrow median dusted vitta. In my opinion, the dusting of the scutum is a variable character.</p><p>I can therefore conclude that Spilogona albinepennis Huckett, 1965 and Spilogona dorsostriata Huckett, 1965 are new junior synonyms of Scatocoenosia cordyluraeformis Schnabl, 1915 .</p><p>Distribution. PALAEARCTIC: Russia (Yamalo-Nenez AO, Taymyr, Wrangel Island). NEARCTIC: Alaska, Yukon, Nunavut/NWT, Quebec.</p></div>	https://treatment.plazi.org/id/03FA87F7FFC3FF0F78EF9C83FE7DF9FE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sorokina, Vera S.	Sorokina, Vera S. (2025): New taxonomic notes on the genus Spilogona Schnabl, 1911 (Diptera: Muscidae) in Arctic Russia, with the description of three new species. Zootaxa 5584 (1): 1-25, DOI: 10.11646/zootaxa.5584.1.1, URL: https://doi.org/10.11646/zootaxa.5584.1.1
03FA87F7FFC3FF0C78EF9AA1FB5BFD57.text	03FA87F7FFC3FF0C78EF9AA1FB5BFD57.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Spilogona depressula (Zetterstedt 1845)	<div><p>Spilogona depressula (Zetterstedt, 1845)</p><p>Aricia depressula Zetterstedt, 1845: 1460 . Type-locality: “ Hab. in Svecia in primis boreali rarissime; in Helsingia ad Arbrå 24 Aug. [lectotype]; in alpe Mullfjellet Jemtlandiae 27 Jul. 1840 specimina duo legi. In Smolandia etiam mense Jul. ad Skärsjö a D. Ljungh bis captam vidi. In Scania ad Lund quoque semel deprehensa.” [SWEDEN].</p><p>Limnophora (Spilogona) reflecta Huckett, 1932: 316 . Type-locality: “ 1♂, Bozeman, Montana, June 14, 1916; 1♂, June 15, 1916; 1♀, June 30, 1916. ( Mont. Exp. Station)” [USA, Montana]. Syn. nov.</p><p>Spilogona depressula (Zetterstedt, 1845); Sorokina, 2017: 53 (Yamalo-Nenez AO); Sorokina, 2018: 237 (Altai Krai, Altai Mts); Sorokina et al., 2018: 327 (Magadan region); Tridrikh &amp; Sorokina, 2021: 827 (Magadan region).</p><p>Notes. The series of Nearctic specimens of Spilogona reflecta (in Canadian National Collection) and the series of Palaearctic specimens of Spilogona depressula were examined. The descriptions of both species and the male terminalia are the same. I can therefore conclude that Limnophora (Spilogona) reflecta Huckett, 1932 is a new junior synonym of Aricia depressula Zetterstedt, 1845 .</p><p>Additional material examined. RUSSIA, Republic of Mordovia : 1♂ 1♀, Temnikov, bank of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=43.19939&amp;materialsCitation.latitude=54.62589" title="Search Plazi for locations around (long 43.19939/lat 54.62589)">Moksha River</a>, 54°37’33.2”N 43°11’57.8”E, 3.viii.2020, leg. K. Tomkovich (ZMUM) . Novosibirsk area: 1 ♂, Novosibirsk, the arboretum, 30.v.2005, leg. V. Sorokina; 1♀, the same data, but 12.vii.2008 . Magadan area: 1♀, ~ 100 km NNE Seimchan, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=153.26666&amp;materialsCitation.latitude=63.65" title="Search Plazi for locations around (long 153.26666/lat 63.65)">Nature Reserve</a> “Magadansky”, 63°39’N 153°16’E, larch forest, sweeping in grass, 4.vii.2019, leg. N. Tridrikh ; 1♀, the same data, but in yellow pan trap, 10.vii.2019; 1♀, ~ 100 km NNE Seimchan, Nature Reserve “Magadansky”, 63°39’N 153°17’E, valley of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=153.28334&amp;materialsCitation.latitude=63.65" title="Search Plazi for locations around (long 153.28334/lat 63.65)">Suksukan River</a>, thickets of young willow, sweeping in grass, 13.vii.2019, leg. N. Tridrikh ; 8♀♀, the same data, but 17.vii.2019; 1♀, the same data, but sweeping over river pebble, 24.vii.2019; 2♀♀, the same data, but on house window, 8.vii.2019 (SZMN, except where stated otherwise).</p><p>Distribution. PALAEARCTIC: Austria, Czech Republic, Denmark, Estonia, Finland, Germany, Latvia, Norway, Poland, Russia (Europe, Siberia, Far East), Sweden. NEARCTIC: Alaska to NWT &amp; Manitoba, Montana, Washington, California.</p><p>DNA barcode. BOLD BIN: BOLD:AAL6514. See Table 1 for GenBank accession numbers.</p></div>	https://treatment.plazi.org/id/03FA87F7FFC3FF0C78EF9AA1FB5BFD57	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sorokina, Vera S.	Sorokina, Vera S. (2025): New taxonomic notes on the genus Spilogona Schnabl, 1911 (Diptera: Muscidae) in Arctic Russia, with the description of three new species. Zootaxa 5584 (1): 1-25, DOI: 10.11646/zootaxa.5584.1.1, URL: https://doi.org/10.11646/zootaxa.5584.1.1
03FA87F7FFC0FF0C78EF9F0AFC70F987.text	03FA87F7FFC0FF0C78EF9F0AFC70F987.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Spilogona fulvibasis Huckett 1965	<div><p>Spilogona fulvibasis Huckett, 1965</p><p>Fig. 5 A</p><p>Spilogona fulvibasis Huckett, 1965: 220 . Type locality: “ Mount McKinley National Park, Alaska, 1932, F.W. Morand ” [USA, Alaska] [seen].</p><p>Spilogona fulvibasis Huckett, 1965; Sorokina et al., 2018: 327 (Magadan region).</p><p>Notes. Huckett (1965:220) described Spilogona fulvibasis from Alaska and indicated in his monograph that the typedepository was the USNM, no. 67311. However, the holotype and the allotype of a “ Spilogona fulvisquama Huckett ” are located in the Main Collection precisely where Spilogona fulvibasis should be located (Garry Ouellette, pers. comm.). Dr Garry Ouellette (USNM) kindly sent me the holotype and allotype of Spilogona fulvisquama for study. These specimens have the same locality and collection date as the original description of Spilogona fulvibasis (Fig. 5A). I examined the types and compared them with the original description of Spilogona fulvibasis, and found that the specimens named as Spilogona fulvisquama completely fit the original description of Spilogona fulvibasis . We can assume that Huckett mislabelled these type-specimens, or changed his mind about the name when completing his manuscript.</p><p>This species is distinctive with an intensive yellow base of the wings, including veins, and densely yellowish calypters (Fig. 5A); also it is large in size (8–9 mm) with haired notopleuron, anepisternum without interspatial setae, scutellum without preapical setulae on upper border of declivities, mid tibia without v setae, hind femur without pv setae, and proboscis short with dusted prementum.</p><p>In the holotype the left fore leg is missing, whilst the allotype is without mid and hind legs.</p><p>Distribution. PALAEARCTIC: Russia (Far East). NEARCTIC: Alaska.</p></div>	https://treatment.plazi.org/id/03FA87F7FFC0FF0C78EF9F0AFC70F987	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sorokina, Vera S.	Sorokina, Vera S. (2025): New taxonomic notes on the genus Spilogona Schnabl, 1911 (Diptera: Muscidae) in Arctic Russia, with the description of three new species. Zootaxa 5584 (1): 1-25, DOI: 10.11646/zootaxa.5584.1.1, URL: https://doi.org/10.11646/zootaxa.5584.1.1
03FA87F7FFC0FF0D78EF9A7AFA03FE4B.text	03FA87F7FFC0FF0D78EF9A7AFA03FE4B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Spilogona lapponica (Ringdahl 1932)	<div><p>Spilogona lapponica (Ringdahl, 1932)</p><p>Figs 5 B–G</p><p>Limnophora (Spilogonoides) lapponica Ringdahl, 1932: 158 . Type-locality: “Ich fing die Art in zwei männlichen Stücken am Strande vom Torneträsk (Regio subarkt.) in dem nördlichsten Lappland, Juli” [SWEDEN, Torne Lappmark].</p><p>Spilogona compacta Huckett, 1965: 198, 208 (key), and 214. Type locality: “ Indian House Lake, Quebec, 7.30 ... 54 [i.e. 30.vii.1954], W.R. Richards ” [CANADA, Quebec] [seen]. Syn. nov.</p><p>Spilogona lapponica (Ringdahl, 1932); Tridrikh &amp; Sorokina, 2021: 827, 830 (Magadan region).</p><p>Notes. The type material of Spilogona compacta (in Canadian National Collection) and European and Asian specimens of Spilogona lapponica were examined. The descriptions of both species and the male terminalia are the same. Males of this species are very distinctive with a very broad frons; a stout thick abdomen with an unusually large sternite 5 (Fig. 5B, F); costa with a strong costal spine in both sexes (Figs 5D, E); mid tibia usually with ad setae and hind femur usually with several short pv setae. I conclude that Spilogona compacta Huckett, 1965 is a new junior synonym of Spilogona lapponica (Ringdahl, 1932) .</p><p>There is no description of the female of Spilogona lapponica in Hennig (1959b), but there is the description of the female of Spilogona compacta in Huckett (1965: 215).</p><p>Additional material examined. RUSSIA, Magadan area: 3♂♂ 2♀♀, ~ 100 km NNE Seimchan, Nature Reserve “Magadansky”, valley of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=153.28334&amp;materialsCitation.latitude=63.65" title="Search Plazi for locations around (long 153.28334/lat 63.65)">Suksukan River</a>, 63°39’N 153°17’E, thickets of young willow, sweeping in grass, 17.vii.2019, leg. N. Tridrikh (SZMN) .</p><p>Distribution. PALAEARCTIC: Norway, Russia (Europe, Far East), Sweden. NEARCTIC: Alaska, Quebec.</p></div>	https://treatment.plazi.org/id/03FA87F7FFC0FF0D78EF9A7AFA03FE4B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sorokina, Vera S.	Sorokina, Vera S. (2025): New taxonomic notes on the genus Spilogona Schnabl, 1911 (Diptera: Muscidae) in Arctic Russia, with the description of three new species. Zootaxa 5584 (1): 1-25, DOI: 10.11646/zootaxa.5584.1.1, URL: https://doi.org/10.11646/zootaxa.5584.1.1
03FA87F7FFC1FF0D78EF9E36FF76F87C.text	03FA87F7FFC1FF0D78EF9E36FF76F87C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Spilogona padlei Huckett 1965	<div><p>Spilogona padlei Huckett, 1965</p><p>Figs 5 H–J</p><p>Spilogona padlei Huckett, 1965: 228 . Type-locality: “Padlei, Northwest Territories, 7.25.50 [i.e. 25.vii.1950], R. E. Duckworth ” [CANADA, NWT] [seen].</p><p>Spilogona perambulata Huckett, 1965: 228 . Type-locality: “Richardson Highway mi. 236, Alaska, 6.26.51 [i.e. 26.vi.1951], J.R. McGillis ” [USA, Alaska] [seen]. Syn. nov.</p><p>Spilogona padlei Huckett, 1965; Sorokina &amp; Thidrikh, 2021: 225 (Chukotka AO).</p><p>Notes. Huckett described Spilogona padlei and Spilogona perambulata in the same monograph (Huckett, 1965). Huckett indicated in the description (1965: 228): “The species padlei and perambulata have much in common. The former however may be distinguished from the latter and related forms by the presence of a fine ventral bristle on mid tibia.”</p><p>The type material of Spilogona padlei and Spilogona perambulata (in Canadian National Collection) has been examined. The descriptions of both species and the male terminalia are the same. Both species have sternite 1 with fine hairs, anepisternum without interspatial seta, scutellum with 2 long downwardly-directed preapical setulae on upper border of declivities, and male mid femur with 1–2 stout pv setae at base. However, Huckett noted in the description of Spilogona padlei “… mid femur with 2 or 3 fine pv,..”, and “… mid femur with a few weak pv …” for Spilogona perambulata . The mid femur of Spilogona padlei really does have a row of weak short pv setae in addition to stout pv seta. Huckett probably did not notice the stout setae because the mid legs are tucked close to the thorax. The mid tibia of Spilogona padlei is actually unlike that of Spilogona perambulata by possessing very short v setae. Since only 6 specimens (5 males and 1 female) of Spilogona padlei have been described from one locality (NWT, Padlei), it can be assumed that the short v setae are a variation of this character in one generation and not enough to describe a separate species.</p><p>The male terminalia of Spilogona padlei and Spilogona perambulata from different generations is the same (Fig. 5H). The sternite 5 is especially distinctive because the two small tubercles at middle are visible in dry specimens (Fig. 5I). The females of both species are also identical; the ovipositor is especially distinctive, with large coarse hook-shaped spinules on the postgenital plate (Fig. 5J).</p><p>I therefore conclude that Spilogona perambulata Huckett, 1965 and Spilogona padlei Huckett, 1965 are both new synonyms, and as they were both published in the same work and on the same page I select as first reviser Spilogona padlei as the valid name for this species.</p><p>Additional material examined. RUSSIA, Taymyr Peninsula: 1♂, Verkhnyaya Taimyra River, mouth of Logata River, willow forest in floodplain, 30.vii.1967, leg. K. Gorodkov (ZISP). Wrangel Island: 1♂ 7♀, lower reaches of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-179.61667&amp;materialsCitation.latitude=70.933334" title="Search Plazi for locations around (long -179.61667/lat 70.933334)">Somnitel′naya River</a>, 70º56’N 179º37’W, yellow pan trap, 7–10.vii.2019, leg. P. Kulemeev &amp; O. Khruleva. Magadan area: 2♀, 120 km NNE Seimchan, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=153.41667&amp;materialsCitation.latitude=63.733334" title="Search Plazi for locations around (long 153.41667/lat 63.733334)">Lyadinoi</a> stream, 63°44’N 153°25’E, yellow pan trap, 10.vii.2019, leg. N. Tridrikh ; 2♀, ~ 100 km NNE Seimchan, Nature Reserve “Magadansky”, 63°39’N 153°17’E, valley of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=153.28334&amp;materialsCitation.latitude=63.65" title="Search Plazi for locations around (long 153.28334/lat 63.65)">Suksukan River</a>, thickets of young willow, white pan trap, 13.vii.2019, leg. N. Tridrikh ; 2♀, the same data, but yellow pan trap, 21–24.vii.2019; 1♀, the same data, but pebble beach, pan trap, 24.vii.2019; 1♀, 10 km N Evensk, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=159.26666&amp;materialsCitation.latitude=61.916668" title="Search Plazi for locations around (long 159.26666/lat 61.916668)">Tainyi</a> stream, 61°55’N 159°16’E, 25.vii.2020 leg. N. Tridrikh ; 1♀, 10 km W Evensk, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=159.38333&amp;materialsCitation.latitude=61.866665" title="Search Plazi for locations around (long 159.38333/lat 61.866665)">Malaya Garmanda River</a>, 61°52’N 159°23’E, sweeping in grass, 01.vii.2020, leg. N. Tridrikh (SZMN except where stated otherwise) .</p><p>Distribution. PALAEARCTIC: Russia (Arctic Siberia, NE of Far East). NEARCTIC: NWT, Alaska, Yukon, Quebec.</p></div>	https://treatment.plazi.org/id/03FA87F7FFC1FF0D78EF9E36FF76F87C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sorokina, Vera S.	Sorokina, Vera S. (2025): New taxonomic notes on the genus Spilogona Schnabl, 1911 (Diptera: Muscidae) in Arctic Russia, with the description of three new species. Zootaxa 5584 (1): 1-25, DOI: 10.11646/zootaxa.5584.1.1, URL: https://doi.org/10.11646/zootaxa.5584.1.1
03FA87F7FFDFFF1378EF9C83FB5BF9AB.text	03FA87F7FFDFFF1378EF9C83FB5BF9AB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Spilogona pusilla Huckett 1932	<div><p>Spilogona pusilla Huckett (1932)</p><p>Fig. 6 B</p><p>Limnophra (Spilogona) pusilla Huckett, 1932b: 134, March. Type locality: “ Tennessee Pass, Colorado (10240 feet), J.M. Aldrich [USA] [seen].</p><p>Limnophra (Spilogona) albisquama Ringdahl, 1932: 156, October. Type-locality: “Mehrere Männchen bei Abisko und Kiruna in der subarktischen Region von mir gefangen, Juli 1918 ” [SWEDEN]. Syn. nov.</p><p>Spilogona albisquama Ringdahl (1932); Sorokina, 2017: 52 (Nenetz AO); Sorokina, 2018: 234 (Altai Mts); Tridrikh &amp; Sorokina, 2021: 826, 832 (Magadan region).</p><p>Notes. Huckett (1932a: 62, key, March; 1932b: 134, June) described Limnophra (Spilogona) pusilla from specimens from Colorado (USA) and Nettilling Lake, Baffin Land (Canada). In the same year Ringdahl (1932, October) described two species Spilogona albisquama and Spilogona subfuscisquama from specimens from Abisko (Sweden) and pointed out their minor differences, mainly in the colour of calypters and male terminalia. Later Huckett (1965) synonymized Spilogona subfuscisquama Ringdahl with Spilogona pusilla because both species have calypters tinged, the lower one with margin markedly dark brownish infuscated. However, Huckett’s figures of the male terminalia of Spilogona pusilla (Huckett, 1965: 353, Plate XVI, fig. 173) agreed well Hennig’s figure of the male terminalia of Spilogona albisquama (Hennig, 1959a: Taf. XIV, fig. 288). Hennig examined Ringdahl’s specimens of Spilogona albisquama and Spilogona subfuscisquama (Hennig, 1959b: Taf. XVIII, fig. 354), and gave good illustrations of the male terminalia of both species which are very different. Ringdahl’s figures of the male terminalia of Spilogona albisquama and Spilogona subfuscisquama are also different (Ringdahl, 1956: 180, figs 76, 78). Moreover, Hennig pointed to weak elongate pv setae on the hind femur of Spilogona albisquama and their absence in Spilogona subfuscisquama (Hennig, 1959a: 265, key).</p><p>Specimens of Spilogona pusilla in the CNC were examined. The paratype of Spilogona pusilla [Nettilling Lake, Baffin Land, July 9, 1925, J.D.D. Soper] and a long series of specimens from Clyde, Baffin Island, belong to Spilogona humeralis Huckett, 1965 . Among specimens of Spilogona pusilla I also found female specimens of Spilogona atrisquamula Hennig, 1959 and male specimens of Spilogona subfuscisquama from Salmita Mines (NWT). All correctly identified specimens of Spilogona pusilla in this collection have hind femur on basal half with short, weak but distinct pv setae. The terminalia are the same in specimens from different localities which fit the terminalia of Spilogona albisquama, and in these specimens the colour of calypters varies from light yellow to brownish.</p><p>I examined and dissected the holotype of Spilogona pusilla . The type has hind femur with short weak pv setae, lower calypter a little darkened but not much, and the terminalia fits Spilogona albisquama . I therefore conclude that Limnophora (Spilogona) albisquama Ringdahl, 1932 is a new junior synonym of Limnophora (Spilogona) pusilla Huckett, 1932, but Spilogona subfuscisquama Ringdahl (1932) is valid species (see below).</p><p>Additional material examined. RUSSIA, Magadan area: 1♀, 100 km NNE Seimchan, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=153.26666&amp;materialsCitation.latitude=63.633335" title="Search Plazi for locations around (long 153.26666/lat 63.633335)">Nature Reserve</a> “Magadansky”, 63°38’N 153°16’E, shrub swamp, sweeping in grass, 15.vii.2019, leg. N. Tridrikh ; 1♀, the same data, but larch woodland, 24.vii.2019; 1♀, the same data, but 63°39’N 153°17’, cotton grass swamp, 25.vii.2019; 1♀, the same data, but 63°41’N 153°22E’, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=153.36667&amp;materialsCitation.latitude=63.683334" title="Search Plazi for locations around (long 153.36667/lat 63.683334)">Karasevoe lake</a>, 25.vii.2019 (all SZMN).</p><p>Distribution. PALAEARCTIC: Finland, Norway, Russia (Siberia and Far East), Sweden. NEARCTIC:Alaska, Yukon, NWT, Manitoba, Quebec, Labrador.</p><p>DNA barcode. BOLD BIN: BOLD:ACR4125. See Table 1 for GenBank accession numbers.</p></div>	https://treatment.plazi.org/id/03FA87F7FFDFFF1378EF9C83FB5BF9AB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sorokina, Vera S.	Sorokina, Vera S. (2025): New taxonomic notes on the genus Spilogona Schnabl, 1911 (Diptera: Muscidae) in Arctic Russia, with the description of three new species. Zootaxa 5584 (1): 1-25, DOI: 10.11646/zootaxa.5584.1.1, URL: https://doi.org/10.11646/zootaxa.5584.1.1
03FA87F7FFDFFF1078EF9A97FB63FDD8.text	03FA87F7FFDFFF1078EF9A97FB63FDD8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Spilogona quinquesetosa (Schnabl 1915)	<div><p>Spilogona quinquesetosa (Schnabl, 1915)</p><p>Fig. 4D</p><p>Limnophora (Spilogona) quinquesetosa Schnabl in Becker et al., 1915: 33. Type-locality: Russia, Yugorskiy peninsula, Kara district [seen].</p><p>Spilogona barrowensis Huckett, 1965: 212 . Type locality: “ Point Barrow, Alaska, 7.20.52 [i.e. 20.vii.1952]” [USA, Alaska; collector P.D. Hurd] [seen]. Syn. nov.</p><p>Spilogona quinquesetosa (Schnabl, 1915); Sorokina &amp; Thidrikh, 2021: 226 (Chukotka AO, including Wrangel Island).</p><p>Notes. The type material of Limnophora quinquesetosa Schnabl (in ZISP) and Spilogona barrowensis Huckett (in Canadian National Collection) was examined. The descriptions of both species and the male terminalia are the same. This is black species with brown dust, without marks on scutum and abdomen, a shining prementum, with preapical setulae on upper border of declivities of scutellum, facial edge not projecting beyond level of profrons, strong av, v, pv setae on mid tibia, and hind femur without pv setae. According to Huckett’s description (1965), the posthumeral and notopleural regions are pale in Spilogona barrowensis, but in the holotype and the paratypes of this species these regions are brown dusted as in Limnophora quinquesetosa .</p><p>I therefore conclude that Spilogona barrowensis Huckett, 1965 is a new junior synonym of Limnophora (Spilogona) quinquesetosa Schnabl, 1915 .</p><p>Additional material examined. RUSSIA, Yamalo-Nenets AO: 3♀, Gydanskiy Proliv, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=72.45&amp;materialsCitation.latitude=72.96667" title="Search Plazi for locations around (long 72.45/lat 72.96667)">Shokalsky Island</a>, 72°58’N 72°27’E, swamp by a stream (1♀), arctic fox hole (1♀), tundra (1♀), 4–25.viii.2016, leg. A. Nikhaeva &amp; M. Bizin (BUIC-DIP2175, BUIC-DIP2176) (all SZMN).</p><p>DNA barcode. BOLD BIN: BOLD:AEK7965. See Table 1 for GenBank accession numbers.</p><p>Distribution. PALAEARCTIC: Russia (Arctic Siberia). NEARCTIC: Alaska (Point Barrow).</p></div>	https://treatment.plazi.org/id/03FA87F7FFDFFF1078EF9A97FB63FDD8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sorokina, Vera S.	Sorokina, Vera S. (2025): New taxonomic notes on the genus Spilogona Schnabl, 1911 (Diptera: Muscidae) in Arctic Russia, with the description of three new species. Zootaxa 5584 (1): 1-25, DOI: 10.11646/zootaxa.5584.1.1, URL: https://doi.org/10.11646/zootaxa.5584.1.1
03FA87F7FFDCFF1078EF9E87FC25FAD1.text	03FA87F7FFDCFF1078EF9E87FC25FAD1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Spilogona scutulata (Schnabl 1911)	<div><p>Spilogona scutulata (Schnabl, 1911)</p><p>Figs 5 K–M</p><p>Limnophora (Spilogona) scutulata Schnabl in Schnabl &amp; Dziedzicki, 1911: 284. Type-locality: “Vier ♂ Exemplare in Warschau, Vorstadt Park Praga. Juli” [POLAND, Warsaw].</p><p>Tetramerinx brevicornis Malloch, 1917: 226 . Type-locality: “Waukegan, Ill., August 23, 1906. Taken on shore of Lake Michigan. Two specimens ” [USA, Illinois]. Syn. nov.</p><p>Notes. The series of Nearctic specimens of Spilogona brevicornis and European specimens of Spilogona scutulata were examined. The descriptions of both species and the male terminalia are the same. These species are very distinctive in both sexes, with strong presutural acr setae in 2 even rows which are strikingly stronger than the ground hairs (Figs 5K, L), and hind femur without pv setae; males with rather broad fronto-orbital plates, which touch and are very silvery, shining; very light grey scutum with a brown spot between post dc; light abdomen with small spots, and a brown dusted scutellum; sternite 5 with with a broad median notch and a pointed caudal margins (Fig. 5M).</p><p>I therefore conclude that Tetramerinx brevicornis Malloch, 1917 is a new junior synonym of Limnophora (Spilogona) scutulata Schnabl, 1911 .</p><p>Distribution. PALAEARCTIC: Austria, Belgium, China (Shanxi, Gansu), Czech Republic, Denmark, Finland, Germany, Great Britain, Hungary, Norway, Poland, Russia (Europe), Slovakia, Sweden. NEARCTIC: Oregon, NWT to Colorado &amp; S Dakota, New York, Manitoba, Quebec, Maine.</p></div>	https://treatment.plazi.org/id/03FA87F7FFDCFF1078EF9E87FC25FAD1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sorokina, Vera S.	Sorokina, Vera S. (2025): New taxonomic notes on the genus Spilogona Schnabl, 1911 (Diptera: Muscidae) in Arctic Russia, with the description of three new species. Zootaxa 5584 (1): 1-25, DOI: 10.11646/zootaxa.5584.1.1, URL: https://doi.org/10.11646/zootaxa.5584.1.1
03FA87F7FFDCFF1678EF998AFC70FC00.text	03FA87F7FFDCFF1678EF998AFC70FC00.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Spilogona subfuscisquama (Ringdahl 1932)	<div><p>Spilogona subfuscisquama (Ringdahl, 1932)</p><p>Fig. 6 C</p><p>Limnophora (Spilogona) subfuscisquama Ringdahl, 1932: 156 . Type-locality: “ 2♂ in der subarktischen Region bei Abisko von mir gefangen, Juli 1918 ” [SWEDEN, Torne Lappmark].</p><p>Notes. Huckett (1965) synonymized Spilogona subfuscisquama with Spilogona pusilla . However, the holotype of Spilogona pusilla agrees well with Spilogona albisquama but not with Spilogona subfuscisquama (see above). The male terminalia of Spilogona subfuscisquama is very different from the figures of Spilogona albisquama given by Ringdahl (1956) and by Hennig (1959a), and from the figures of the male terminalia of Spilogona pusilla given by Huckett (1965).</p><p>Among specimens of Spilogona pusilla in the CNC I found specimens of Spilogona subfuscisquama from Salmita Mines (N.W.T). In this collection, some specimens of Spilogona subfuscisquama from Fort Churchill (Manitoba) were also found among Spilogona aenea Huckett, 1965 . All these specimens had hind femur without any pv setae and the male terminalia as in Hennig’s figures of Spilogona subfuscisquama (Hennig, 1959b: Taf. XVIII, fig. 354). The colour of the calypters of these specimens is variable from light yellow to brownish.</p><p>Spilogona subfuscisquama (Ringdahl, 1932) is thus a valid species with an arctic-subarctic distribution, which is different from the similar Spilogona pusilla with a wider subarctic-subalpine distribution. The colour of the calypters is variable in both species and cannot be used for the identification of these species.</p><p>Material examined. CANADA, NWT: 4♂♂, Salmita Mines, 64°05’N 111°15’W, N.W. T ., 4.vii.1953, leg. J.G. Chillcott (CNC) . CANADA, Manitoba: 3♂♂, Fort Churchill, 23.vi.1952, leg. C.D. Bird (CNC) .</p><p>The very similar Spilogona subfuscisquama and Spilogona pusilla are closely allied to species such as Spilogona humeralis, Spilogona atrisquamula, Spilogona malaisei Ringdahl, 1920 and Spilogona wrangeli Hennig, 1959 by their small size (3–4 mm), subovate abdomen, brown or blackish thorax without stripes, anepisternum without interspatial setae, scutellum with downwardly-directed preapical setulae on upper border of declivities, and mid tibia without v setae.</p><p>Males of these species can be separated by the following couplets:</p><p>1 Both calypters dark brown................................................................... S. atrisquamula</p><p>- Calypters yellow, or lower calypter light brownish........................................................... 2</p><p>2 Scutum seen from in front with anterior border of presutural region more or less greyish; sternite V with a blunt rounded caudal margin (Fig. 6A)............................................................................. S. humeralis</p><p>- Scutum concolorous on presutural region, deep brown; sternite V with a pointed caudal margin (Figs 6B, C)............. 3</p><p>3 Abdomen dark grey with indistinct and ill-defined black marks; facial edge a little projecting beyond level of profrons, or not ................................................................................................... 4</p><p>- Abdomen pale or whitish-grey, with distinct and well-defined black marks; facial edge not projecting beyond level of profrons............................................................................................. 5</p><p>4 Hind femur without pv setae; costa without spinules, with short setae; surstylus curved and shorter than cerci (Fig. 6D)................................................................................................ S. malaisei</p><p>- Hind femur with weak elongated pv setae in basal half; costa with spinules; surstylus not curved and as long as cerci (Fig. 6E)........................................................................................... S. wrangeli</p><p>5 Hind femur with weak, but distinct elongated pv setae in basal half; sternite V with a deep trapezoidal median notch (Fig. 6B); prementum completely shining................................................................... S. pusilla</p><p>- Hind femur without pv setae; sternite V with a deep triangular median notch (Fig. 6C); prementum often with light dust in middle................................................................................ S. subfuscisquama</p><p>Females of all these species are very similar and difficult to distinguish. More specimens from different localities need to be studied to identify possible differences. Spilogona malaisei and Spilogona wrangeli are the most distinct from the others, as in the following couplets:</p><p>1 Abdomen seen from behind entirely black and mostly glossy, without dorsal marks................................. 2</p><p>- Abdomen grey and with paired brownish marks on dorsum.......................................................3</p><p>2 Costa without spinules, with short setae........................................................... S. malaisei</p><p>- Costa with spinules........................................................................... S. wrangeli</p><p>3 Facial edge not projecting; scutum pale grey, with or without clear brown marks; scutellum wholly pale grey............................................................................. S. pusilla &amp;? S. atrisquamula &amp;? S. subfuscisquama</p><p>- Facial edge a little projecting; scutum and scutellum with deep brownish dust, at least at basal angles of scutellum................................................................................................... S. humeralis</p><p>Spilogona pusilla is distributed not only on the plains but also in mountainous areas, where specimens are characterized by a darker colour not only on the lower calypter but also on the scutum, as in Spilogona humeralis . The latter species can be separated from Spilogona pusilla by the slightly projecting facial edge. Females of Spilogona subfuscisquama, like males, probably have the prementum with light dust in the middle, and can be separated from the other species by this character.</p><p>Distribution. PALAEARCTIC: Sweden. NEARCTIC: NWT, Manitoba.</p></div>	https://treatment.plazi.org/id/03FA87F7FFDCFF1678EF998AFC70FC00	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sorokina, Vera S.	Sorokina, Vera S. (2025): New taxonomic notes on the genus Spilogona Schnabl, 1911 (Diptera: Muscidae) in Arctic Russia, with the description of three new species. Zootaxa 5584 (1): 1-25, DOI: 10.11646/zootaxa.5584.1.1, URL: https://doi.org/10.11646/zootaxa.5584.1.1
03FA87F7FFDAFF1778EF9FFFFBD5FE4B.text	03FA87F7FFDAFF1778EF9FFFFBD5FE4B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Spilogona tendipes (Malloch 1920) F	<div><p>Spilogona tendipes (Malloch, 1920)</p><p>Figs 7 A–H</p><p>Notes. In Sorokina &amp; Michelsen (2014) Spilogona tendipes was recorded for the first time from Wrangel Island and from NE Greenland from Zackenberg, all based on females. However, a few doubts were expressed about the assignment of these females to Spilogona tendipes: (1) the type locality lies in the warmer maritime climatic zone on the south coast of Alaska; (2) Huckett (1932, 1965) emphasized the presence of abundant setulae on the declivities of the scutellum in the male holotype, but these declivities are practically bare in these females.</p><p>Later, male and female specimens similar to Spilogona tendipes from Wrangel Island were found in Gorodkov’s collections in the ZISP, and additional material of the same species has been collected on Wrangel Island. Further material similar to Spilogona tendipes from the Magadan region and Altai Mts of Russia was also collected recently. The good condition of these flies and the presence of males made it possible to study more carefully all the characters of the collected specimens.</p><p>Specimens from the Magadan region and Altai Mts agreed fully with the description of Spilogona tendipes; these localities are not further north than the type locality of Spilogona tendipes . Specimens from Wrangel Island are very similar to Spilogona tendipes, but there are not only some morphological differences between these species, including the genitalia, but they are also genetically distinct. The specimens from Wrangel Island, including the females listed in Sorokina &amp; Michelsen (2014), thus proved to be a species new for science, Spilogona latizygoma sp. nov., described above.</p><p>Spilogona tendipes and Spilogona latizygoma sp. nov. can be separated as in following couple:</p><p>- Mid tibia without v setae; hind femur with distinct long pv in basal half; all femora with numerous hair-like av setae in basal half; scutum with long ground-setulae; sternites 3–5 with long setae (Fig. 3A); eye with inconspicuous hairs; male terminalia as in Fig. 3G, H; female hypoproct with weak individual spinules (Fig. 3F)...................... S. latizygoma sp. nov.</p><p>- Mid tibia with a short v seta; hind femur without distinct long pv in basal half; all femora without numerous hair-like av setae in basal half; scutum with short ground-setulae; sternites 3–5 with short setae (Fig. 7B); eye with distinct hairs; male terminalia as in Fig. 7F, G; female hypoproct with numerous spinules (Fig. 7E)..................................... S. tendipes</p><p>Material examined. RUSSIA, Magadan region: 1♂ 2♀, Yablonevyi Pass, 120 km NNE Magadan, 900 m,</p><p>60°59’N 151°53 Е, 17.vii.2014, leg. N. Vikhrev (1♀ (BUIC-DIP1759) in SZMN, 1♀ 1♂ in ZMUM); 1♀ 1♂, Yablonevyi Pass, 120 km NNE Magadan, 783 м , 60°35’N 151°32 Е, yellow pan trap, 11.vii.2017, leg. V. Sorokina (BUIC-DIP1758, SZMN). Altai Republic: 1♂, Kosh-Agach district, Akbul River, on stones near the water, 2075 m , 49º39’N 88º1’Е, 12.vii.2009, leg. V. Sorokina (SZMN). CANADA, Yukon: 1♀, Richardson Mts, 67º58’N 136º29’W, 1–5.vii.1987, leg. M. Polak &amp; M. Wood (SZMN) .</p><p>Distribution. PALAEARCTIC:? Greenland (NE), Russia (Magadan region, Altay). NEARCTIC: Alaska, Yukon.</p><p>DNA barcode. BOLD BIN: BOLD:ADZ7217. See Table 1 for GenBank accession numbers.</p><p>Remarks. Spilogona tendipes is morphologically very similar to Spilogona latizygoma . But based on DNA barcodes, these species are genetically different; the p-distance between them is 3.36%.</p></div>	https://treatment.plazi.org/id/03FA87F7FFDAFF1778EF9FFFFBD5FE4B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sorokina, Vera S.	Sorokina, Vera S. (2025): New taxonomic notes on the genus Spilogona Schnabl, 1911 (Diptera: Muscidae) in Arctic Russia, with the description of three new species. Zootaxa 5584 (1): 1-25, DOI: 10.11646/zootaxa.5584.1.1, URL: https://doi.org/10.11646/zootaxa.5584.1.1
03FA87F7FFD8FF1478EF9C83FABCFAB9.text	03FA87F7FFD8FF1478EF9C83FABCFAB9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Spilogona wrangeli Hennig 1959	<div><p>Spilogona wrangeli Hennig, 1959</p><p>Figs 6 E, F</p><p>Spilogona wrangeli Hennig, 1959: 360 . Type-locality: “O-w Wrangelja, werch. r. Klen k SSW ot pika Berri, 19.–21.VIII. 38 g. Gekker” [RUSSIA, Far East, Wrangelya I., upper Klen river SSW of Mt. Berry] [seen].</p><p>Spilogona coactilis Huckett, 1965: 214 . Type-locality: “ Herschel Island, Yukon Territory, 7.11.53 [i.e. 11.vii.1953], J.S. Waterhouse)” [CANADA, Yukon] [seen]. Syn. nov.</p><p>Spilogona malaisei (Ringdahl, 1920); Sorokina &amp; Khruleva 2012: 556 (Wrangel Island). Misidentification.</p><p>Spilogona wrangeli Hennig, 1959; Sorokina &amp; Thidrikh, 2021: 229 (Wrangel Island).</p><p>Notes. The holotypes of Spilogona coactilis (in Canadian National Collection) and Spilogona wrangeli (in ZISP) were examined. Both species had been described from arctic Islands, Wrangel Island and Herschel Island, and had not since been found in other localities. Spilogona coactilis is known only from the holotype and one additional male. A series of Spilogona wrangeli has been found recently on Wrangel Island (Sorokina &amp; Thidrikh, 2021). The descriptions and the male terminalia are the same for both species. The exception was a v seta on the mid tibia, which Huckett described for Spilogona coactilis; Spilogona wrangeli is without a v seta on the mid tibia. I examined the holotype of Spilogona coactilis . The specimen is a little crumpled, legs pressed against the thorax, one mid tibia in glue; with difficulty I was able to see that the mid tibia is in fact without v setae. Males of these species are small in size (3.5–4 mm), facial edge often a little projecting beyond level of profrons, but sometimes not, subovate abdomen, blackish thorax without stripes, anepisternum without interspatial setae, scutellum with downwardly-directed preapical setulae on upper border of declivities, costa with spinules, mid tibia without v setae, hind femur with weak pv setae, and abdomen dark with indistinct black expansive marks.</p><p>I therefore conclude that Spilogona coactilis Huckett, 1965 is a new junior synonym of Spilogona wrangeli Hennig, 1959 .</p><p>The female of both Spilogona coactilis and Spilogona wrangeli has been not described. It is very similar to Spilogona malaisei, like the male, and both can be separated as in following couplet:</p><p>1 Costa without spinules, with short setae............................................................ S. malaisei</p><p>- Costa with spinules........................................................................... S. wrangeli</p><p>Distribution. PALAEARCTIC: Russia (Wrangel Island). NEARCTIC: Yukon (Herschel Island).</p><p>DNA barcode. BOLD BIN: BOLD:AED9512. See Table 1 for GenBank accession numbers.</p><p>Remarks. Spilogona wrangeli is morphologically very similar to Spilogona malaisei, especially females. But based on DNA barcodes, these species are genetically different; the p-distance between them is 2.04–2.15%.</p></div>	https://treatment.plazi.org/id/03FA87F7FFD8FF1478EF9C83FABCFAB9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sorokina, Vera S.	Sorokina, Vera S. (2025): New taxonomic notes on the genus Spilogona Schnabl, 1911 (Diptera: Muscidae) in Arctic Russia, with the description of three new species. Zootaxa 5584 (1): 1-25, DOI: 10.11646/zootaxa.5584.1.1, URL: https://doi.org/10.11646/zootaxa.5584.1.1
