taxonID	type	description	language	source
03FA570AFC0CFFB77BF6F94FFD296C26.taxon	description	Besides the genera and subgenera considered in the present paper, the generic complex also includes the Afrotropical genus Kruegerilema Volynkin & László, 2021 endemic to Sao Tome Island (see: Volynkin & László 2021), which is morphologically very similar to Brunia.	en	Volynkin, Anton V., Černý, Karel (2025): On the taxonomy of the Brunia Moore generic complex with the description of a new genus, a new subgenus and four new species (Lepidoptera: Erebidae: Arctiinae: Lithosiini: Lithosiina). Ecologica Montenegrina 87: 46-76, DOI: 10.37828/em.2025.87.2, URL: https://doi.org/10.37828/em.2025.87.2
03FA570AFC0FFFB77BF6FE60FC54693F.taxon	diagnosis	Diagnosis. Many species of the genus display a moderate sexual dimorphism: males have intense dark suffusion on the forewing distally whereas females have a more unicolorous forewing, in many species with pale suffusion along the costal margin. However, in certain Asian species sexual dimorphism is limited. The male genitalia of the genus are characterised by the combination of the following features. (1) The scaphium area is setose. (2) The subscaphium is broad and moderately or weakly sclerotised. (3) The processus momenti and tendon form a conjuga. (4) The lamella centralis is present, fold-like and setose. (5) The vesica with several diverticula bearing large cornuti of various shapes. In the female genitalia, the postvaginal region of the 8 th sternite is heavily sclerotised and may bear various swollen and setose sculptures; and the posterior sclerotised section of the corpus bursae is broad and bears various protrusions.	en	Volynkin, Anton V., Černý, Karel (2025): On the taxonomy of the Brunia Moore generic complex with the description of a new genus, a new subgenus and four new species (Lepidoptera: Erebidae: Arctiinae: Lithosiini: Lithosiina). Ecologica Montenegrina 87: 46-76, DOI: 10.37828/em.2025.87.2, URL: https://doi.org/10.37828/em.2025.87.2
03FA570AFC0FFFB77BF6FE60FC54693F.taxon	distribution	Distribution. The genus is widely distributed in the Afrotropical, Indomalayan and northern Australasian realms. Note. The taxonomy of the genus worldwide is under revision by Volynkin (in prep.).	en	Volynkin, Anton V., Černý, Karel (2025): On the taxonomy of the Brunia Moore generic complex with the description of a new genus, a new subgenus and four new species (Lepidoptera: Erebidae: Arctiinae: Lithosiini: Lithosiina). Ecologica Montenegrina 87: 46-76, DOI: 10.37828/em.2025.87.2, URL: https://doi.org/10.37828/em.2025.87.2
03FA570AFC0FFFB77BF6FB09FA686824.taxon	diagnosis	Diagnosis. Male adults have rather unicolorous body. In the male genitalia, species of the nominate subgenus are characterised by the condition of the distal section of the phallus, which has longitudinal wrinkles making it flexible and expandable. In the female genitalia, the ductus bursae is broad, heavily gelatinous or sclerotised, very short in certain species.	en	Volynkin, Anton V., Černý, Karel (2025): On the taxonomy of the Brunia Moore generic complex with the description of a new genus, a new subgenus and four new species (Lepidoptera: Erebidae: Arctiinae: Lithosiini: Lithosiina). Ecologica Montenegrina 87: 46-76, DOI: 10.37828/em.2025.87.2, URL: https://doi.org/10.37828/em.2025.87.2
03FA570AFC0EFFB67BF6FA0AFBB56A95.taxon	materials_examined	Type species: Katha montana Bucsek, 2012, by present designation.	en	Volynkin, Anton V., Černý, Karel (2025): On the taxonomy of the Brunia Moore generic complex with the description of a new genus, a new subgenus and four new species (Lepidoptera: Erebidae: Arctiinae: Lithosiini: Lithosiina). Ecologica Montenegrina 87: 46-76, DOI: 10.37828/em.2025.87.2, URL: https://doi.org/10.37828/em.2025.87.2
03FA570AFC0EFFB67BF6FA0AFBB56A95.taxon	diagnosis	Diagnosis. Species of Brunia (Brunikatha) subgen. n. (Figs 5 – 21) are externally similar to Bucsekatha gen. n. (Figs 22 – 35) and Sany (Figs 36 – 38), and differ from the nominate subgenus of Brunia (Figs 1 – 4) in the blackish-brown head and thorax in males. In the male genitalia, the new subgenus (Figs 56 – 65) is distinguished from Brunia s. str. (Figs 54, 55) by two features. (1) The processus momenti is reduced, which makes the conjuga very short and consisting of the tendon and the thin sclerotised plate connecting the tendon and the processus momenti in the nominate subgenus. (2) The phallus condition: in Brunia s. str., its distal section has longitudinal wrinkles making it expandable whereas it is evenly sclerotised and lacks the apical wrinkles in Brunia (Brunikatha) subgen. n. The female genitalia of the two subgenera (Figs 89 – 97) display no fundamental differences.	en	Volynkin, Anton V., Černý, Karel (2025): On the taxonomy of the Brunia Moore generic complex with the description of a new genus, a new subgenus and four new species (Lepidoptera: Erebidae: Arctiinae: Lithosiini: Lithosiina). Ecologica Montenegrina 87: 46-76, DOI: 10.37828/em.2025.87.2, URL: https://doi.org/10.37828/em.2025.87.2
03FA570AFC0EFFB67BF6FA0AFBB56A95.taxon	etymology	Etymology. The subgenus name is an aggregate of the genus-group names Brunia and Katha. Note. Species of the subgenus are herein subdivided into two species-groups clearly different in their morphology of the genitalia of both sexes.	en	Volynkin, Anton V., Černý, Karel (2025): On the taxonomy of the Brunia Moore generic complex with the description of a new genus, a new subgenus and four new species (Lepidoptera: Erebidae: Arctiinae: Lithosiini: Lithosiina). Ecologica Montenegrina 87: 46-76, DOI: 10.37828/em.2025.87.2, URL: https://doi.org/10.37828/em.2025.87.2
03FA570AFC09FFB17BF6FF0BFDAC6C26.taxon	description	Note. The taxonomy of the group is under revision by Volynkin & Černý (in prep.).	en	Volynkin, Anton V., Černý, Karel (2025): On the taxonomy of the Brunia Moore generic complex with the description of a new genus, a new subgenus and four new species (Lepidoptera: Erebidae: Arctiinae: Lithosiini: Lithosiina). Ecologica Montenegrina 87: 46-76, DOI: 10.37828/em.2025.87.2, URL: https://doi.org/10.37828/em.2025.87.2
03FA570AFC09FFB17BF6FD72FD156EC3.taxon	description	Note. Two species included in the species-group are reviewed below.	en	Volynkin, Anton V., Černý, Karel (2025): On the taxonomy of the Brunia Moore generic complex with the description of a new genus, a new subgenus and four new species (Lepidoptera: Erebidae: Arctiinae: Lithosiini: Lithosiina). Ecologica Montenegrina 87: 46-76, DOI: 10.37828/em.2025.87.2, URL: https://doi.org/10.37828/em.2025.87.2
03FA570AFC09FFB07BF6FB9DFDD16FD7.taxon	description	(Figs 9 – 15, 58 – 61, 93, 94)	en	Volynkin, Anton V., Černý, Karel (2025): On the taxonomy of the Brunia Moore generic complex with the description of a new genus, a new subgenus and four new species (Lepidoptera: Erebidae: Arctiinae: Lithosiini: Lithosiina). Ecologica Montenegrina 87: 46-76, DOI: 10.37828/em.2025.87.2, URL: https://doi.org/10.37828/em.2025.87.2
03FA570AFC09FFB07BF6FB9DFDD16FD7.taxon	materials_examined	Type material examined. Photograph of the holotype (by monotypy) (Fig. 9): male, “ Horisha | Formosa | - 1917 ” / “ L. | usuguronis ” / red label “ Type | Matsumura ” (HUFA). Additional material examined: series of specimens of both sexes from Taiwan, Northern Vietnam (Nghe An, Hoa Binh and Ninh Binh Provinces), Northern Thailand (Chiang Mai Province) and Northeast India (Meghalaya) (MWM / ZSM). Note. Wu et al. (2020: fig. 9 k) illustrated the male genitalia belonging to the genus Cernyia Bucsek, 2012 as those of the holotype of Lithosia usuguronis. However, the only species of the genus Cernyia occurring in Taiwan, Cernyia ranrunensis (Matsumura, 1927), a male specimen of which was also illustrated by Wu et al. (2020: fig. 5 c) as “ Cer. [nyia] usuguronis ”, is externally fundamentally different from the holotype of usuguronis and is very similar to other species in the genus Cernyia. On the other hand, all examined Taiwanese moths externally identical to the holotype of Lithosia usuguronis belong to a single species of the genus Brunia, which is widespread from Northeast India to Taiwan. Thus, it is clear that the Cernyia male genitalia illustrated by Wu et al. (2020: fig. 9 k) do not belong to the holotype specimen of Lithosia usuguronis. This unfortunate mistake is probably due to a Cernyia abdomen being misglued to the holotype of Lithosia usuguronis in the past, or was caused by the accidental confusion of the genitalia of different specimens happened during a series of dissections (for the similar case see Volynkin (2020 )).	en	Volynkin, Anton V., Černý, Karel (2025): On the taxonomy of the Brunia Moore generic complex with the description of a new genus, a new subgenus and four new species (Lepidoptera: Erebidae: Arctiinae: Lithosiini: Lithosiina). Ecologica Montenegrina 87: 46-76, DOI: 10.37828/em.2025.87.2, URL: https://doi.org/10.37828/em.2025.87.2
03FA570AFC09FFB07BF6FB9DFDD16FD7.taxon	diagnosis	Diagnosis. The forewing length is 13.0 – 14.5 mm in males and 15.0 – 16.0 mm in females. Brunia usuguronis is externally similar to B. montana but is somewhat larger. The male genital capsule of B. usuguronis is distinguished from B. montana by the shorter vinculum, the narrower and more apically rounded tip of the valva, and the shorter sacculus with a distal saccular process not reaching the valva apex. Compared to B. montana, the vesica of B. usuguronis has a markedly broader and somewhat longer main chamber, its proximal dorsal and medial lateral diverticula are considerably longer and broader, the proximal ventral diverticulum is longer, distally tapered and bears a serrulate plate-shaped cornutus (in B. montana it is distally dilated and bears a robust thorn-shaped cornutus), and the distal diverticulum is longer, broader, hook-like curved and bears a cornutus with a long plate-like base laterally whereas in B. montana the distal diverticulum is shortly conical and bears a massive horn-shaped cornutus apically. In the female genitalia, B. usuguronis differs from B. montana in the somewhat longer ductus bursae, the larger and sclerotised, semiglobular lateral protrusion bursae on the right side, the lack of the heavily sclerotised ventro-lateral pocket on the left side of the corpus bursae anteriorly of the appendix bursae, which is typical of B. montana, and the proximally broader and more weakly sclerotised appendix bursae.	en	Volynkin, Anton V., Černý, Karel (2025): On the taxonomy of the Brunia Moore generic complex with the description of a new genus, a new subgenus and four new species (Lepidoptera: Erebidae: Arctiinae: Lithosiini: Lithosiina). Ecologica Montenegrina 87: 46-76, DOI: 10.37828/em.2025.87.2, URL: https://doi.org/10.37828/em.2025.87.2
03FA570AFC09FFB07BF6FB9DFDD16FD7.taxon	distribution	Distribution. The species is found in Northeast India (Meghalaya), Northern Thailand (Chiang Mai and Nan Provinces), Northern Laos (Huaphanh Province) (Bucsek 2020, as montana), Northern Vietnam (Nghe An, Hoa Binh and Ninh Binh Provinces), and the island of Taiwan.	en	Volynkin, Anton V., Černý, Karel (2025): On the taxonomy of the Brunia Moore generic complex with the description of a new genus, a new subgenus and four new species (Lepidoptera: Erebidae: Arctiinae: Lithosiini: Lithosiina). Ecologica Montenegrina 87: 46-76, DOI: 10.37828/em.2025.87.2, URL: https://doi.org/10.37828/em.2025.87.2
03FA570AFC08FFB07BF6FD10FA6C68D1.taxon	description	(Figs 16 – 21, 62 – 65, 95 – 97)	en	Volynkin, Anton V., Černý, Karel (2025): On the taxonomy of the Brunia Moore generic complex with the description of a new genus, a new subgenus and four new species (Lepidoptera: Erebidae: Arctiinae: Lithosiini: Lithosiina). Ecologica Montenegrina 87: 46-76, DOI: 10.37828/em.2025.87.2, URL: https://doi.org/10.37828/em.2025.87.2
03FA570AFC08FFB07BF6FD10FA6C68D1.taxon	materials_examined	Type material examined. Holotype (Figs 16, 62): male, “ Malaysia, Pahang distr. | Cameron Highlands | Tanah Rata env., 1,450 – 1,500 m | 4 ° 28 ' 25,3 '' N, 101 ° 22 ' 43,7 '' E | 10.4.2010 – 24.4.2010 | leg. Karol Bucsek ” / red label “ Holotypus | Katha montana ♂ | Karol Bucsek 2012 ” / “ Museum Witt | München ” / “ Genitalpräparat | Heterocera | Nr. 19.826 | Museum Witt München ” [ex slide Mal 166 by K. Bucsek, remounted by A. Volynkin] (MWM / ZSM). Additional material examined. Series of specimens of both sexes from Thailand (Phetchabun Province), West Malaysia (Pahang) (the paratypes listed by Bucsek (2012), and Indonesia (Sumatra Island) (MWM / ZSM, CKC).	en	Volynkin, Anton V., Černý, Karel (2025): On the taxonomy of the Brunia Moore generic complex with the description of a new genus, a new subgenus and four new species (Lepidoptera: Erebidae: Arctiinae: Lithosiini: Lithosiina). Ecologica Montenegrina 87: 46-76, DOI: 10.37828/em.2025.87.2, URL: https://doi.org/10.37828/em.2025.87.2
03FA570AFC08FFB07BF6FD10FA6C68D1.taxon	diagnosis	Diagnosis. The forewing length is 11.0 – 12.5 mm in males and 13.0 – 13.5 mm in females. The species is morphologically similar to B. usuguronis and the detailed comparison is provided above in the diagnosis of the latter species.	en	Volynkin, Anton V., Černý, Karel (2025): On the taxonomy of the Brunia Moore generic complex with the description of a new genus, a new subgenus and four new species (Lepidoptera: Erebidae: Arctiinae: Lithosiini: Lithosiina). Ecologica Montenegrina 87: 46-76, DOI: 10.37828/em.2025.87.2, URL: https://doi.org/10.37828/em.2025.87.2
03FA570AFC08FFB07BF6FD10FA6C68D1.taxon	distribution	Distribution. The species is found in Northeast India (Arunachal Pradesh) (Kirti & Singh 2015), Central Thailand (Phetchabun Province) (present study), Western Malaysia (Pahang) (Bucsek 2012), and the island of Sumatra (Indonesia) (present study).	en	Volynkin, Anton V., Černý, Karel (2025): On the taxonomy of the Brunia Moore generic complex with the description of a new genus, a new subgenus and four new species (Lepidoptera: Erebidae: Arctiinae: Lithosiini: Lithosiina). Ecologica Montenegrina 87: 46-76, DOI: 10.37828/em.2025.87.2, URL: https://doi.org/10.37828/em.2025.87.2
03FA570AFC08FFB37BF6FA12FA166F13.taxon	type_taxon	Type species: Bucsekatha sunda sp. n., by present designation.	en	Volynkin, Anton V., Černý, Karel (2025): On the taxonomy of the Brunia Moore generic complex with the description of a new genus, a new subgenus and four new species (Lepidoptera: Erebidae: Arctiinae: Lithosiini: Lithosiina). Ecologica Montenegrina 87: 46-76, DOI: 10.37828/em.2025.87.2, URL: https://doi.org/10.37828/em.2025.87.2
03FA570AFC08FFB37BF6FA12FA166F13.taxon	diagnosis	Diagnosis. Species of the new genus (Figs 22 – 35) are externally very similar to Brunia (Brunikatha) subgen. n. (Figs 5 – 21) and Sany (Figs 36 – 38), and identification requires the examination of the genitalia structures. In the male genitalia, Bucsekatha gen. n. (Figs 66 – 78) is morphologically similar to Sany (Figs 79, 80) sharing the combination of the following features. (1) The scaphium area is setose. (2) The subscaphium area is membranous. (3) The ventral wall of the anellus bears a caulis subdivided into two lobes. (4) The phallus is cylindrical with a ductus ejaculatorius originating laterally on the left side (vs. Brunia and a number of other Lithosiini having a ductus ejaculatorius originating dorsally). (5) The dorsal carina bears a thorn-shaped subapical process directed dorsad. (6) The vesica has a short main chamber, which bears a short but basally broad medial cornutus and has one to three short diverticula proximally and medially, and an elongate, more or less tubular distal diverticulum bearing a cornutus apically or subapically. However, the male genitalia of Bucsekatha gen. n. differ from Sany in the lobes of the caulis being completely fused with anellus and separated from each other (whereas they are ventrally fused with each other and dorsally extended into loose processes in Sany), the broader lobular valva with a convex ventral margin (it is very narrow with almost parallel margins in Sany), the distal section of the editum expanded into a broad setose soft lobe (autapomorphic feature), the reduced, tubercle-like lamella centralis (absent in Sany), the weakly setose apex of the valva (it is covered with robust spine-like setae in Sany), and the broader sacculus with a well-developed distal process (the latter is reduced or completely absent in Sany). The female genitalia ground plan of Bucsekatha gen. n. (Figs 98 – 105) is very similar to Sany (Fig. 106) but the latter has an appendix bursae situated rather laterally whereas it is situated ventrally or ventro-laterally in Bucsekatha gen. n.	en	Volynkin, Anton V., Černý, Karel (2025): On the taxonomy of the Brunia Moore generic complex with the description of a new genus, a new subgenus and four new species (Lepidoptera: Erebidae: Arctiinae: Lithosiini: Lithosiina). Ecologica Montenegrina 87: 46-76, DOI: 10.37828/em.2025.87.2, URL: https://doi.org/10.37828/em.2025.87.2
03FA570AFC08FFB37BF6FA12FA166F13.taxon	etymology	Etymology. The genus name is an aggregate of the surname Bucsek and the genus-group name Katha. The new genus is named after Karol Bucsek, renowned expert in the Asian Arctiinae, who greatly contributed to the knowledge of the Lithosiini of the Malay Peninsula, Philippine Archipelago and Laos including the taxonomy of the Brunia generic complex.	en	Volynkin, Anton V., Černý, Karel (2025): On the taxonomy of the Brunia Moore generic complex with the description of a new genus, a new subgenus and four new species (Lepidoptera: Erebidae: Arctiinae: Lithosiini: Lithosiina). Ecologica Montenegrina 87: 46-76, DOI: 10.37828/em.2025.87.2, URL: https://doi.org/10.37828/em.2025.87.2
03FA570AFC0BFFB37BF6FC66FD086B7D.taxon	description	(Figs 22, 23, 66, 98)	en	Volynkin, Anton V., Černý, Karel (2025): On the taxonomy of the Brunia Moore generic complex with the description of a new genus, a new subgenus and four new species (Lepidoptera: Erebidae: Arctiinae: Lithosiini: Lithosiina). Ecologica Montenegrina 87: 46-76, DOI: 10.37828/em.2025.87.2, URL: https://doi.org/10.37828/em.2025.87.2
03FA570AFC0BFFB37BF6FC66FD086B7D.taxon	materials_examined	Type material examined. Holotype: male, “ Sarawak: Gunong | Mulu Nat. Pak | R. G. S. Exped. 1977 – 8 | J. D. Holloway et al. | B. M. 1978 – 206 ” / “ Site 15. February | Camp 2.5, Mulu | 1000 m. 413461 | Lower l. montane f. | MV – understorey. ” / “ + 3 ” / “ Holotype | “ Eilema ” | brevivalva | Holloway ” / round label with red margin “ Type ” / blue label “ Arctiidae | genitalia slide | No. 2639 ♂ ” / “ Figured specimen | Moths of Borneo | Part 7 ” / QR-code label with unique ID “ NHMUK 010401787 ” (NHMUK). Additional material examined. INDONESIA: 4 males, 1 female, Borneo, Kalimantan Selatan [Prov.], 1100 m, 30 km E of Kandangan, rainforest, 15 km NE of Loksado, 2 ° 52 ' S 115 ° 38 ' E, I. 1998, Jakl leg., gen. slide Nos: ZSM Arct. 2021 - 541 ♂, 2021 - 542 ♀ (MWM / ZSM).	en	Volynkin, Anton V., Černý, Karel (2025): On the taxonomy of the Brunia Moore generic complex with the description of a new genus, a new subgenus and four new species (Lepidoptera: Erebidae: Arctiinae: Lithosiini: Lithosiina). Ecologica Montenegrina 87: 46-76, DOI: 10.37828/em.2025.87.2, URL: https://doi.org/10.37828/em.2025.87.2
03FA570AFC0BFFB37BF6FC66FD086B7D.taxon	diagnosis	Diagnosis. The forewing length is 12.0 mm in males and 13.5 mm in females. The male genital capsule of B. brevivalva differs clearly from other congeners in the reduced, very short dorsal (valvular) section of the valva and the markedly longer distal saccular process. In the female genitalia, the species is distinct by the narrow and anteriorly tapered posterior sclerotised section of the corpus bursae.	en	Volynkin, Anton V., Černý, Karel (2025): On the taxonomy of the Brunia Moore generic complex with the description of a new genus, a new subgenus and four new species (Lepidoptera: Erebidae: Arctiinae: Lithosiini: Lithosiina). Ecologica Montenegrina 87: 46-76, DOI: 10.37828/em.2025.87.2, URL: https://doi.org/10.37828/em.2025.87.2
03FA570AFC0BFFB37BF6FC66FD086B7D.taxon	distribution	Distribution. Endemic to the island of Borneo (Holloway 2001). The present report of the species from Kalimantan Selatan Province of Indonesia is first for the country.	en	Volynkin, Anton V., Černý, Karel (2025): On the taxonomy of the Brunia Moore generic complex with the description of a new genus, a new subgenus and four new species (Lepidoptera: Erebidae: Arctiinae: Lithosiini: Lithosiina). Ecologica Montenegrina 87: 46-76, DOI: 10.37828/em.2025.87.2, URL: https://doi.org/10.37828/em.2025.87.2
03FA570AFC0BFFB27BF6F946FAB96857.taxon	description	(Figs 24 – 27, 67 – 69, 99, 100)	en	Volynkin, Anton V., Černý, Karel (2025): On the taxonomy of the Brunia Moore generic complex with the description of a new genus, a new subgenus and four new species (Lepidoptera: Erebidae: Arctiinae: Lithosiini: Lithosiina). Ecologica Montenegrina 87: 46-76, DOI: 10.37828/em.2025.87.2, URL: https://doi.org/10.37828/em.2025.87.2
03FA570AFC0BFFB27BF6F946FAB96857.taxon	materials_examined	Type material. Holotype (Figs 24, 67): male, “ SW Thailand | Ranong-prov. Ranong, 380 m | 10 ° 01 ' 32 '' N, 98 ° 40 ' 13 '' E | 17, 20. XI. 2006 leg. K. Cerný ” / “ Slide | AV 9177 ♂ | A. Volynkin ” (MWM / ZSM, ex CKC). Paratypes. THAILAND: 6 males, 1 female, Ranong Prov., Ranong, 380 m, 10 ° 01 ' 32 '' N 98 ° 40 ' 13 '' E, 3 – 4. XII. 2005, K. Černý leg., gen. slide Nos: AV 8286 ♂, AV 9150 ♀ (CKC); 1 female, same data as previous but ex ovo 2006, gen. slide No.: AV 9172 ♀ (CKC); 1 male, Ranong, 543 m, 10 ° 02 ' 01 ' N 98 ° 40 ' 17.66 '' E, 7. IV. 2013, K. Černý leg., gen. slide No.: AV 8281 ♂ (CKC); 1 female, Surat Thani Prov., Khao Sok NP, 199 m, 8 ° 53 ' 23 '' N 98 ° 30 ' 25 '' E, 6. XII. 2005, K. Černý leg., gen. slide No.: AV 9153 ♀ (CKC); 3 males, Nakhon Si Tammarat [Prov.], Amphoe Ron Pibun, | Ang Ka Tai Hill, 900 – 950 m, 8 ° 14.16 ' N 99 ° 48.12 ' E, 9 – 10. IV. 2013, K. Černý leg., gen. slide No.: AV 8280 ♂ (CKC); 1 male, Chumphon Prov., Pa Toh Ditr., Ban Lang Tang, 162 m, 7. XII. 2005, 9 ° 46 ' 5 '' N 98 ° 46 ' 59 '' E, K. Černý leg., gen. slide No.: AV 8282 ♂ (CKC); MALAYSIA: 2 males, 1 female, Perak, 50 km NE of Gerik (aerial path), Belum-Temenggor: Titiwangsa, 5 ° 36 ' 17.4 '' N 101 ° 32 ' 34.0 '' E, 30. iii – 13. iv. 2015, 1100 m, E. Jendek & O. Šauša leg., gen. slide Nos: ZSM Arct. 2021 - 518 ♂, 2021 - 595 ♀, 2021 - 599 ♂ (prepared by A. Volynkin) (MWM / ZSM); INDONESIA: 1 male, 1 female, N Sumatra, 15 km NE of Prapat, HW 4, 1150 m, 3 – 6. I. 1994, Karel Černý leg., gen. slide Nos: AV 8274 ♂, AV 9185 ♀ (CKC); 1 female, N Sumatra, Simalungun, Prapat, HW 3, 9. II. 1984, Dr Diehl leg., gen. slide No.: AV 9484 ♀ (CKC); 1 male, Sumatra, Aek Tarum, Gunung Malayu, 8. V. 1983, Dr Diehl leg., gen. slide No.: AV 9491 ♂ (CKC); 1 male, Sumatra, Bandar Lampung, Bukit Rigis Mts, Gunung Terang, 1022 m, 5 ° 3 ' 43 '' S 104 ° 27 ' 6 '' E, 13 – 15. XI. 2003, [K.] Černý leg., gen. slide No.: AV 9176 ♂ (CKC); 1 female, W Sumatra, 22 km E of Padang, Bukit Subang, microwave stat. [ion], 0 ° 40 ' S 100 ° 32 ' E, montane forest, 1200 m, 19 – 20. X. 1981, A. Schintlmeister et al. leg., gen. slide No.: AV 9184 ♀ (CKC); 1 male, South Sumatra, Lake Ranau env., 800 m, 4 ° 507 ' 38 '' S 104 ° 3 ' [00 ''] E, 17. XI. 2003, K. Černý leg. (CKC). Note. The population from the Malay Peninsula lacks the small medial diverticulum, which is present in the specimens from Sumatra. However, the taxonomic value of this feature is unclear and, as no other morphological differences were found between the specimens from the two populations, they are herein considered as belonging to the same biological species.	en	Volynkin, Anton V., Černý, Karel (2025): On the taxonomy of the Brunia Moore generic complex with the description of a new genus, a new subgenus and four new species (Lepidoptera: Erebidae: Arctiinae: Lithosiini: Lithosiina). Ecologica Montenegrina 87: 46-76, DOI: 10.37828/em.2025.87.2, URL: https://doi.org/10.37828/em.2025.87.2
03FA570AFC0BFFB27BF6F946FAB96857.taxon	diagnosis	Diagnosis. The forewing length is 11.5 – 13.0 mm in males and 14.0 mm in females. The male genital capsule of B. sunda sp. n. differs from B. chonga sp. n. and B. pseudobrevivalva in the longer distal saccular process and the shorter, narrower and more weakly sclerotised lobes of the caulis. The phallus of B. sunda sp. n. differs from the similar species in the markedly smaller carinal process. The vesica structure of B. sunda sp. n. is most similar to B. chonga sp. n., from which the former differs in the larger medial cornutus. In the female genitalia, B. sunda sp. n. differs from other congeners in the markedly narrower ductus bursae.	en	Volynkin, Anton V., Černý, Karel (2025): On the taxonomy of the Brunia Moore generic complex with the description of a new genus, a new subgenus and four new species (Lepidoptera: Erebidae: Arctiinae: Lithosiini: Lithosiina). Ecologica Montenegrina 87: 46-76, DOI: 10.37828/em.2025.87.2, URL: https://doi.org/10.37828/em.2025.87.2
03FA570AFC0BFFB27BF6F946FAB96857.taxon	distribution	Distribution. The new species is found on the Malay Peninsula (South Thailand and West Malaysia) and the island of Sumatra (Indonesia).	en	Volynkin, Anton V., Černý, Karel (2025): On the taxonomy of the Brunia Moore generic complex with the description of a new genus, a new subgenus and four new species (Lepidoptera: Erebidae: Arctiinae: Lithosiini: Lithosiina). Ecologica Montenegrina 87: 46-76, DOI: 10.37828/em.2025.87.2, URL: https://doi.org/10.37828/em.2025.87.2
03FA570AFC0BFFB27BF6F946FAB96857.taxon	etymology	Etymology. The specific epithet is derived from Sundaland, in the western part of which the new species is found. The name is a noun in the nominative singular.	en	Volynkin, Anton V., Černý, Karel (2025): On the taxonomy of the Brunia Moore generic complex with the description of a new genus, a new subgenus and four new species (Lepidoptera: Erebidae: Arctiinae: Lithosiini: Lithosiina). Ecologica Montenegrina 87: 46-76, DOI: 10.37828/em.2025.87.2, URL: https://doi.org/10.37828/em.2025.87.2
03FA570AFC0AFFBD7BF6FA90FC1B6CEB.taxon	description	(Figs 28 – 32, 70 – 75, 101 – 103)	en	Volynkin, Anton V., Černý, Karel (2025): On the taxonomy of the Brunia Moore generic complex with the description of a new genus, a new subgenus and four new species (Lepidoptera: Erebidae: Arctiinae: Lithosiini: Lithosiina). Ecologica Montenegrina 87: 46-76, DOI: 10.37828/em.2025.87.2, URL: https://doi.org/10.37828/em.2025.87.2
03FA570AFC0AFFBD7BF6FA90FC1B6CEB.taxon	materials_examined	Type material. Holotype (Figs 28, 70): male, “ Cambodia, Kampot Prov., | Bokor N. P., Hill Station, 1025 m, | 19 – 21. I. 2006 | leg. G. Csorba & G. Ronkay ” / “ Slide | ZSM Arct. | 2021 - 520 ♂ | A. Volynkin ” (MWM / ZSM). Paratypes. CAMBODIA: 1 male, 2 females, same data as in holotype, gen. slide No.: ZSM Arct. 2021 - 521 ♀ (prepared by A. Volynkin) (MWM / ZSM); 1 male, 1 female, Kirirom Nat. Res., 650 m, 7 – 9. XI. 2007, S. Murzin leg., gen. slide Nos: ZSM Arct. 2021 - 597 ♂, 2021 - 598 ♀ (prepared by A. Volynkin) (MWM / ZSM); THAILAND: 2 males, 2 females, Saraburi [Nakhon Nayok Prov.], Khao Yai NP, Mt. Khao Kaew [Khao Khiao], 1244 m, 14 ° 21 ' 56 '' N 101 ° 24 ' E, 31. V. 2005, K. Černý leg., gen. slide Nos: AV 8278 ♂, AV 8284 ♂, AV 9155 ♀, AV 9173 ♀ (CKC); 4 males, 1 female, Saraburi [Nakhon Nayok Prov.], Khao Yai NP, Khao Khieo [foot of Khao Khiao Mt.], 752 m, 14 ° 24 ' 20 '' N 101 ° 22 ' 14 '' E, 1. VI. 2005, K. Černý leg., gen. slide Nos: AV 8279 ♂, AV 8283 ♂, AV 9154 ♀ (CKC); 2 males, Chanthaburi Prov., 150 m, 33 km N of Chanthaburi, 12 ° 48.17 ' N 102 ° 17.53 ' E, 30. X. 2010, K. Černý leg., gen. slide Nos: AV 8285 ♂, AV 9179 ♂ (CKC).	en	Volynkin, Anton V., Černý, Karel (2025): On the taxonomy of the Brunia Moore generic complex with the description of a new genus, a new subgenus and four new species (Lepidoptera: Erebidae: Arctiinae: Lithosiini: Lithosiina). Ecologica Montenegrina 87: 46-76, DOI: 10.37828/em.2025.87.2, URL: https://doi.org/10.37828/em.2025.87.2
03FA570AFC0AFFBD7BF6FA90FC1B6CEB.taxon	diagnosis	Diagnosis. The forewing length is 11.0 – 13.0 mm in males and 14.0 – 15.0 mm in females. The male genitalia of B. chonga sp. n. are similar to B. sunda sp. n. but distinguished by the shorter and thinner distal saccular process, the broader and more heavily sclerotised lobes of the caulis, the markedly larger carinal process, and the smaller medial cornutus. The female genitalia of B. chonga sp. n. differ from B. sunda sp. n. in the considerably broader ductus bursae and the somewhat more anteriorly curved sclerotised anterior section of the corpus bursae.	en	Volynkin, Anton V., Černý, Karel (2025): On the taxonomy of the Brunia Moore generic complex with the description of a new genus, a new subgenus and four new species (Lepidoptera: Erebidae: Arctiinae: Lithosiini: Lithosiina). Ecologica Montenegrina 87: 46-76, DOI: 10.37828/em.2025.87.2, URL: https://doi.org/10.37828/em.2025.87.2
03FA570AFC0AFFBD7BF6FA90FC1B6CEB.taxon	distribution	Distribution. The new species is found in Central and Southeast Thailand (Nakhon Nayok and Chanthaburi Provinces) and Southwest Cambodia (Kampot and Kampong Speu Provinces).	en	Volynkin, Anton V., Černý, Karel (2025): On the taxonomy of the Brunia Moore generic complex with the description of a new genus, a new subgenus and four new species (Lepidoptera: Erebidae: Arctiinae: Lithosiini: Lithosiina). Ecologica Montenegrina 87: 46-76, DOI: 10.37828/em.2025.87.2, URL: https://doi.org/10.37828/em.2025.87.2
03FA570AFC0AFFBD7BF6FA90FC1B6CEB.taxon	etymology	Etymology. The specific epithet is derived from the Chong people, who inhabit western Cambodia, and south-eastern and eastern Thailand. The name is a noun in the nominative singular.	en	Volynkin, Anton V., Černý, Karel (2025): On the taxonomy of the Brunia Moore generic complex with the description of a new genus, a new subgenus and four new species (Lepidoptera: Erebidae: Arctiinae: Lithosiini: Lithosiina). Ecologica Montenegrina 87: 46-76, DOI: 10.37828/em.2025.87.2, URL: https://doi.org/10.37828/em.2025.87.2
03FA570AFC05FFBD7BF6FAFBFAF96B3C.taxon	distribution	Distribution. South China: Hainan Island (Fang 2000).	en	Volynkin, Anton V., Černý, Karel (2025): On the taxonomy of the Brunia Moore generic complex with the description of a new genus, a new subgenus and four new species (Lepidoptera: Erebidae: Arctiinae: Lithosiini: Lithosiina). Ecologica Montenegrina 87: 46-76, DOI: 10.37828/em.2025.87.2, URL: https://doi.org/10.37828/em.2025.87.2
03FA570AFC05FFBD7BF6FE3DF85E69A9.taxon	description	(Figs 33 – 35, 76 – 78, 104, 105)	en	Volynkin, Anton V., Černý, Karel (2025): On the taxonomy of the Brunia Moore generic complex with the description of a new genus, a new subgenus and four new species (Lepidoptera: Erebidae: Arctiinae: Lithosiini: Lithosiina). Ecologica Montenegrina 87: 46-76, DOI: 10.37828/em.2025.87.2, URL: https://doi.org/10.37828/em.2025.87.2
03FA570AFC05FFBD7BF6FE3DF85E69A9.taxon	materials_examined	Material examined. Series of specimens of both sexes from Northeast India (Assam), Laos (Luang Prabang and Champasak Provinces), and Vietnam (Nghe An, Lam Dong and Da Nang Provinces) (MWM / ZSM, CKC and CAV).	en	Volynkin, Anton V., Černý, Karel (2025): On the taxonomy of the Brunia Moore generic complex with the description of a new genus, a new subgenus and four new species (Lepidoptera: Erebidae: Arctiinae: Lithosiini: Lithosiina). Ecologica Montenegrina 87: 46-76, DOI: 10.37828/em.2025.87.2, URL: https://doi.org/10.37828/em.2025.87.2
03FA570AFC05FFBD7BF6FE3DF85E69A9.taxon	diagnosis	Diagnosis. The forewing length is 12.0 – 13.5 mm in males and 12.5 – 13.5 mm in females. The male genital capsule of B. pseudobrevivalva is very similar to B. chonga sp. n., from which B. pseudobrevivalva differs clearly in the vesica having a markedly shorter distal diverticulum bearing a larger terminal cornutus. Additionally, compared to B. chonga sp. n., B. pseudobrevivalva has a shorter proximal ventral diverticulum displaced more laterally, a markedly shorter medial ventral diverticulum, and a considerably larger lateral diverticulum displaced more dorsally. In the female genitalia of B. pseudobrevivalva, the posterior sclerotised section of the corpus bursae is markedly broader and less curved than in the similar congener, and the appendix bursae is situated ventrally whereas it is situated ventro-laterally on the left side in B. chonga sp. n.	en	Volynkin, Anton V., Černý, Karel (2025): On the taxonomy of the Brunia Moore generic complex with the description of a new genus, a new subgenus and four new species (Lepidoptera: Erebidae: Arctiinae: Lithosiini: Lithosiina). Ecologica Montenegrina 87: 46-76, DOI: 10.37828/em.2025.87.2, URL: https://doi.org/10.37828/em.2025.87.2
03FA570AFC05FFBD7BF6FE3DF85E69A9.taxon	distribution	Distribution. The species is widespread and found in Northeast India (Assam and Mizoram) (Singh et al. 2024), Laos (Huaphanh, Luang Prabang and Champasak Provinces) (Bucsek 2020), and Vietnam (Thanh Hoa, Nghe An, Gia Lai, Bac Giang, and Da Nang Provinces) (Dubatolov & Bucsek 2016)	en	Volynkin, Anton V., Černý, Karel (2025): On the taxonomy of the Brunia Moore generic complex with the description of a new genus, a new subgenus and four new species (Lepidoptera: Erebidae: Arctiinae: Lithosiini: Lithosiina). Ecologica Montenegrina 87: 46-76, DOI: 10.37828/em.2025.87.2, URL: https://doi.org/10.37828/em.2025.87.2
03FA570AFC05FFBD7BF6F906FAEB6AB2.taxon	materials_examined	Type species: Lithosia brevipennis sandakana Draudt, 1914, by original designation.	en	Volynkin, Anton V., Černý, Karel (2025): On the taxonomy of the Brunia Moore generic complex with the description of a new genus, a new subgenus and four new species (Lepidoptera: Erebidae: Arctiinae: Lithosiini: Lithosiina). Ecologica Montenegrina 87: 46-76, DOI: 10.37828/em.2025.87.2, URL: https://doi.org/10.37828/em.2025.87.2
03FA570AFC05FFBD7BF6F906FAEB6AB2.taxon	diagnosis	Diagnosis. Species of the genus (Figs 36 – 38) are externally very similar to Brunia (Brunikatha) subgen. n. (Figs 5 – 21) and Bucsekatha gen. n. (Figs 22 – 35), and identification requires the examination of the genitalia structures. Sany is morphologically similar to Bucsekatha gen. n. and the detailed comparison is provided above in the diagnosis of the latter genus. Note. The genus comprises two species reviewed below.	en	Volynkin, Anton V., Černý, Karel (2025): On the taxonomy of the Brunia Moore generic complex with the description of a new genus, a new subgenus and four new species (Lepidoptera: Erebidae: Arctiinae: Lithosiini: Lithosiina). Ecologica Montenegrina 87: 46-76, DOI: 10.37828/em.2025.87.2, URL: https://doi.org/10.37828/em.2025.87.2
03FA570AFC04FFBF7BF6F90FF84D6F6C.taxon	materials_examined	Type species: “ Eilema ” trimacula Holloway, 2001, by the original designation. Note. The genus was characterised and reviewed by Volynkin (2021).	en	Volynkin, Anton V., Černý, Karel (2025): On the taxonomy of the Brunia Moore generic complex with the description of a new genus, a new subgenus and four new species (Lepidoptera: Erebidae: Arctiinae: Lithosiini: Lithosiina). Ecologica Montenegrina 87: 46-76, DOI: 10.37828/em.2025.87.2, URL: https://doi.org/10.37828/em.2025.87.2
03FA570AFC04FFBF7BF6F90FF84D6F6C.taxon	diagnosis	Diagnosis. Species of the genus (Figs 39 – 52) display a substantial sexual dimorphism with the females having a fuscous forewing with yellowish costal and outer areas whereas males of Dubatoloviana have two cellular and one subapical spots, which are absent in other genera in the Brunia generic complex. Additionally, unlike in other related genera, the male forewing of Dubatoloviana has a triangular patch of darker, presumably androconial scales antemedially between the vein A 1 and the anal wing margin but not reaching the latter. The male genitalia ground plan of Dubatoloviana (Figs 81 – 88) is similar to Bucsekatha gen. n. (Figs 66 – 78) and Sany (Figs 79, 80) but, unlike in those genera, Dubatoloviana is characterised by the broader and strongly swollen uncus, the lack of the caulis, the heavily sclerotised distal section of the costa forming a “ cucullus ” with a spur-shaped tip, the small and membranous valvula, and the phallus with a ductus ejaculatorius situated dorsally and a dorsal carina lacking the subapical process. The vesica configuration of Dubatoloviana is similar to Bucsekatha gen. n. and Sany but the main chamber lacks the medial cornutus, and the distal diverticulum is short and directed dorsally. The female genitalia of Dubatoloviana (Figs 107 – 112) are very similar to Bucsekatha gen. n. (Figs 98 – 105) and Sany (Fig. 106) but have an appendix bursae situated laterally and curved dorsad.	en	Volynkin, Anton V., Černý, Karel (2025): On the taxonomy of the Brunia Moore generic complex with the description of a new genus, a new subgenus and four new species (Lepidoptera: Erebidae: Arctiinae: Lithosiini: Lithosiina). Ecologica Montenegrina 87: 46-76, DOI: 10.37828/em.2025.87.2, URL: https://doi.org/10.37828/em.2025.87.2
03FA570AFC04FFBC7BF6FF0CFA206E9F.taxon	description	(Figs 36, 37, 79, 106)	en	Volynkin, Anton V., Černý, Karel (2025): On the taxonomy of the Brunia Moore generic complex with the description of a new genus, a new subgenus and four new species (Lepidoptera: Erebidae: Arctiinae: Lithosiini: Lithosiina). Ecologica Montenegrina 87: 46-76, DOI: 10.37828/em.2025.87.2, URL: https://doi.org/10.37828/em.2025.87.2
03FA570AFC04FFBC7BF6FF0CFA206E9F.taxon	materials_examined	Type material examined. Holotype (Figs 36, 79): male, “ Malaysia, Pahang distr. | Endau Rompin State park | Camping by The Kincin River, 50 m | 2 ° 37 ' 10,4 '' N, 103 ° 20 ' 10,9 '' E | 1.3.2011 – 13.3.2011 | leg. Karol Bucsek ” / red label “ Holotypus | Sany pseudosandakana ♂ | Karol Bucsek 2012 ” / “ Museum Wit | München ” / “ Genitalpräparat | Heterocera | Nr. 37.368 | Museum Witt München ” [prepared by A. Volynkin] (MWM / ZSM). Additional material examined. MALAYSIA: series of specimens of both sexes (paratypes) listed by Bucsek (2012) (MWM / ZSM); INDONESIA: 1 male, N Sumatra, Prapat, HW 3, 4. X. 1983, Dr Diehl leg., gen. slide No.: AV 9490 ♂ (CKC).	en	Volynkin, Anton V., Černý, Karel (2025): On the taxonomy of the Brunia Moore generic complex with the description of a new genus, a new subgenus and four new species (Lepidoptera: Erebidae: Arctiinae: Lithosiini: Lithosiina). Ecologica Montenegrina 87: 46-76, DOI: 10.37828/em.2025.87.2, URL: https://doi.org/10.37828/em.2025.87.2
03FA570AFC04FFBC7BF6FF0CFA206E9F.taxon	diagnosis	Diagnosis. The forewing length is 11.0 – 11.5 mm in males and 11.5 – 12.0 mm in females. The male genital capsule of S. pseudosandakana is distinguished from S. sandakana by the narrower vinculum, the somewhat shorter but broader valva, the longer cucullus covered with larger setae, and the lack of the distal saccular process, which is tiny thorn-shaped in S. sandakana. The vesica of S. pseudosandakana differs from S. sandakana in the narrower main chamber, and the markedly shorter, proximally broader and distally curved distal diverticulum bearing a larger cornutus postmedially whereas in S. sandakana, the tiny cornutus is situated apically.	en	Volynkin, Anton V., Černý, Karel (2025): On the taxonomy of the Brunia Moore generic complex with the description of a new genus, a new subgenus and four new species (Lepidoptera: Erebidae: Arctiinae: Lithosiini: Lithosiina). Ecologica Montenegrina 87: 46-76, DOI: 10.37828/em.2025.87.2, URL: https://doi.org/10.37828/em.2025.87.2
03FA570AFC04FFBC7BF6FF0CFA206E9F.taxon	distribution	Distribution. The species is known from the Malay Peninsula (West Malaysia: Pahang) (Bucsek 2012) and the island of Sumatra (Indonesia) (present study).	en	Volynkin, Anton V., Černý, Karel (2025): On the taxonomy of the Brunia Moore generic complex with the description of a new genus, a new subgenus and four new species (Lepidoptera: Erebidae: Arctiinae: Lithosiini: Lithosiina). Ecologica Montenegrina 87: 46-76, DOI: 10.37828/em.2025.87.2, URL: https://doi.org/10.37828/em.2025.87.2
03FA570AFC04FFBC7BF6FBE8FA8F6B35.taxon	description	(Figs 38, 80) Lithosia brevipennis subsp. sandakana Draudt in Seitz, 1914, Die Gross-Schmetterlinge der Erde, 10: 210 (TL: “ Borneo (Sandakan) ”).	en	Volynkin, Anton V., Černý, Karel (2025): On the taxonomy of the Brunia Moore generic complex with the description of a new genus, a new subgenus and four new species (Lepidoptera: Erebidae: Arctiinae: Lithosiini: Lithosiina). Ecologica Montenegrina 87: 46-76, DOI: 10.37828/em.2025.87.2, URL: https://doi.org/10.37828/em.2025.87.2
03FA570AFC04FFBC7BF6FBE8FA8F6B35.taxon	materials_examined	Type material examined. Holotype (by monotypy) (Figs 38, 80): male, “ Sandakan. | 91 – 115. ” / “ Eilema | brevipennis | ab. sandakana | Draudt | Holotype. Strand | Hampson Ab. 1. ” / round label with red margins “ Type ” / blue label “ Arctiidae | genitalia slide | No. 4893 ♂ ” / “ Figured specimen | Moths of Borneo | part 7 ” / QR-code label with unique ID “ NHMUK 010401830 ” (NHMUK). Note. The name sandakana was introduced by Draudt (1914) for Hampson’s ‘ Ab. [erration] 1 ’ of Ilema brevipennis. In his book, Hampson (1900) cited a single male associated by him to that aberration, which should be considered as the holotype of sandakana by monotypy.	en	Volynkin, Anton V., Černý, Karel (2025): On the taxonomy of the Brunia Moore generic complex with the description of a new genus, a new subgenus and four new species (Lepidoptera: Erebidae: Arctiinae: Lithosiini: Lithosiina). Ecologica Montenegrina 87: 46-76, DOI: 10.37828/em.2025.87.2, URL: https://doi.org/10.37828/em.2025.87.2
03FA570AFC04FFBC7BF6FBE8FA8F6B35.taxon	diagnosis	Diagnosis. The forewing length is 11.0 mm in the male holotype. The detailed comparison with S. pseudosandakana is provided above in the diagnosis of the latter species. The female is unknown.	en	Volynkin, Anton V., Černý, Karel (2025): On the taxonomy of the Brunia Moore generic complex with the description of a new genus, a new subgenus and four new species (Lepidoptera: Erebidae: Arctiinae: Lithosiini: Lithosiina). Ecologica Montenegrina 87: 46-76, DOI: 10.37828/em.2025.87.2, URL: https://doi.org/10.37828/em.2025.87.2
03FA570AFC04FFBC7BF6FBE8FA8F6B35.taxon	distribution	Distribution. The species is known only from its holotype collected in Sabah (Malaysia) (Holloway 2001), and is probably endemic to the island of Borneo.	en	Volynkin, Anton V., Černý, Karel (2025): On the taxonomy of the Brunia Moore generic complex with the description of a new genus, a new subgenus and four new species (Lepidoptera: Erebidae: Arctiinae: Lithosiini: Lithosiina). Ecologica Montenegrina 87: 46-76, DOI: 10.37828/em.2025.87.2, URL: https://doi.org/10.37828/em.2025.87.2
03FA570AFC07FFBF7BF6FC48F8B76BE4.taxon	description	(Figs 45, 46, 84, 110)	en	Volynkin, Anton V., Černý, Karel (2025): On the taxonomy of the Brunia Moore generic complex with the description of a new genus, a new subgenus and four new species (Lepidoptera: Erebidae: Arctiinae: Lithosiini: Lithosiina). Ecologica Montenegrina 87: 46-76, DOI: 10.37828/em.2025.87.2, URL: https://doi.org/10.37828/em.2025.87.2
03FA570AFC07FFBF7BF6FC48F8B76BE4.taxon	materials_examined	Type material. Holotype (Figs 45, 84): male, “ Sumatra | Prapat HW 3 | 26. – 30. XI. 1982 | Dr. Diehl leg. ” / “ Slide | AV 8372 ♂ | A. Volynkin ” (MWM / ZSM, ex CKC). Paratype. INDONESIA: 1 female, W Sumatra, Mt. Talang, 0 ° 57 ' S 100 ° 24 ' E, 18. IX. 1991, secondary forest, 1600 m, Graul & Schintlmeister leg., gen. slide No.: AV 8373 ♀ (CKC).	en	Volynkin, Anton V., Černý, Karel (2025): On the taxonomy of the Brunia Moore generic complex with the description of a new genus, a new subgenus and four new species (Lepidoptera: Erebidae: Arctiinae: Lithosiini: Lithosiina). Ecologica Montenegrina 87: 46-76, DOI: 10.37828/em.2025.87.2, URL: https://doi.org/10.37828/em.2025.87.2
03FA570AFC07FFBF7BF6FC48F8B76BE4.taxon	diagnosis	Diagnosis. The forewing length is 12.0 mm in the male holotype and 13.5 mm in the female paratype. Dubatoloviana diehli sp. n. is externally very similar to D. pahanga but has a somewhat narrower forewing. Additionally, compared to D. pahanga, the male of the new species has more distinct forewing markings, and the female has a straighter costal margin of the forewing with a narrower outer ochreous yellow area at the apex. The male genital capsule of D. diehli sp. n. differs clearly from D. pahanga in the longer and less downcurved tip of the uncus, the thicker “ cucullus ”, and the narrower unilobate distal saccular process, which is bifurcate in D. pahanga. The phallus of the new species is somewhat longer and distally broader than in D. pahanga. The vesica configurations of the two species are similar but in D. diehli sp. n., the main chamber is somewhat longer than in the congener and the distal diverticulum bears a cornutus, which is absent in D. pahanga. In the female genitalia, D. diehli sp. n. is distinguished from D. pahanga by the wider gelatinous antevaginal plate and the shorter posterior sclerotised section of the corpus bursae.	en	Volynkin, Anton V., Černý, Karel (2025): On the taxonomy of the Brunia Moore generic complex with the description of a new genus, a new subgenus and four new species (Lepidoptera: Erebidae: Arctiinae: Lithosiini: Lithosiina). Ecologica Montenegrina 87: 46-76, DOI: 10.37828/em.2025.87.2, URL: https://doi.org/10.37828/em.2025.87.2
03FA570AFC07FFBF7BF6FC48F8B76BE4.taxon	distribution	Distribution. Indonesia (Sumatra Island).	en	Volynkin, Anton V., Černý, Karel (2025): On the taxonomy of the Brunia Moore generic complex with the description of a new genus, a new subgenus and four new species (Lepidoptera: Erebidae: Arctiinae: Lithosiini: Lithosiina). Ecologica Montenegrina 87: 46-76, DOI: 10.37828/em.2025.87.2, URL: https://doi.org/10.37828/em.2025.87.2
03FA570AFC07FFBF7BF6FC48F8B76BE4.taxon	etymology	Etymology. The new species is dedicated to the memory of the late Dr. med. Eduard Wilhelm Diehl (1917 – 2003), famous researcher of Sumatran insects and collector of the holotype. The name is a noun in the genitive case.	en	Volynkin, Anton V., Černý, Karel (2025): On the taxonomy of the Brunia Moore generic complex with the description of a new genus, a new subgenus and four new species (Lepidoptera: Erebidae: Arctiinae: Lithosiini: Lithosiina). Ecologica Montenegrina 87: 46-76, DOI: 10.37828/em.2025.87.2, URL: https://doi.org/10.37828/em.2025.87.2
03FA570AFC07FFBE7BF6F93EFD3F6E1D.taxon	description	(Figs 47 – 50, 85, 86, 111, 112)	en	Volynkin, Anton V., Černý, Karel (2025): On the taxonomy of the Brunia Moore generic complex with the description of a new genus, a new subgenus and four new species (Lepidoptera: Erebidae: Arctiinae: Lithosiini: Lithosiina). Ecologica Montenegrina 87: 46-76, DOI: 10.37828/em.2025.87.2, URL: https://doi.org/10.37828/em.2025.87.2
03FA570AFC07FFBE7BF6F93EFD3F6E1D.taxon	materials_examined	Type material examined. Lectotype (designated by Hampson (1900) as “ Type ”) (Figs 47, 111): female, “ Tenasserim Valley, | E. of Tovoy [Dawei], | Burmah [Myanmar] | Doherty. ” / “ Lithosia | fumidisca | Hmpn. type. ♀ ” / “ Presented by | J. J. Joicey Esq. | Brit. Mus. 1934 - 444. ” / round label with red margin “ Type | H. T ” / blue label “ Arctiidae | genitalia slide No. 2854 ♀ ” (NHMUK). Additional material examined. THAILAND: 1 male, Chumphon Prov., 162 m, Pa Toh, Ban Lang Tang, 9 ° 46 ' 5 '' N, 98 ° 46 ' 59 '' E, 3. V. 2008, K. Černý leg., gen. slide No.: AV 8368 ♂ (CKC); 1 male, same data as previous but 2 – 3. XII. 2007, gen. slide No.: AV 8370 ♂ (CKC); 1 male, 1 female, same data as previous but 7. XII. 2005, gen. slide No.: AV 8457 ♀ (CKC). Notes. (1) Lithosia fumidisca was originally described from an unspecified number of female specimens originating from two localities (“ Sikhim; Tenasserim Valley ”) (Hampson 1894). Subsequently, Hampson (1900) cited only one female from Tenasserim Valley as “ type ”, which, according to the Article 74.5 of ICZN (1999), should be considered as lectotype designation. (2) The species was placed in the genus Brunia by Dubatolov & Zolotuhin (2011).	en	Volynkin, Anton V., Černý, Karel (2025): On the taxonomy of the Brunia Moore generic complex with the description of a new genus, a new subgenus and four new species (Lepidoptera: Erebidae: Arctiinae: Lithosiini: Lithosiina). Ecologica Montenegrina 87: 46-76, DOI: 10.37828/em.2025.87.2, URL: https://doi.org/10.37828/em.2025.87.2
03FA570AFC07FFBE7BF6F93EFD3F6E1D.taxon	diagnosis	Diagnosis. The forewing length is 10.0 – 11.5 mm in males and 11.0 – 12.0 mm in females. The male genitalia of D. fumidisca are similar to D. navneetsinghi but differ in the narrower arms of the tegumen, the longer valva, and the larger proximal diverticulum. As the female of D. navneetsinghi remains unknown (see below), the female genitalia of D. fumidisca were compared to the morphologically closest D. trimacula instead, from which D. fumidisca can be distinguished by the larger setose postvaginal lobes of the 8 th sternite, the presence of a sclerotised swollen fascia on the posterior margin of the 7 th sternite, and the broader posterior sclerotised section of the corpus bursae.	en	Volynkin, Anton V., Černý, Karel (2025): On the taxonomy of the Brunia Moore generic complex with the description of a new genus, a new subgenus and four new species (Lepidoptera: Erebidae: Arctiinae: Lithosiini: Lithosiina). Ecologica Montenegrina 87: 46-76, DOI: 10.37828/em.2025.87.2, URL: https://doi.org/10.37828/em.2025.87.2
03FA570AFC07FFBE7BF6F93EFD3F6E1D.taxon	distribution	Distribution. The species is known from south-eastern Myanmar (Tanintharyi Region) (Hampson 1894) and Southern Thailand (Chumphon Province) (present study).	en	Volynkin, Anton V., Černý, Karel (2025): On the taxonomy of the Brunia Moore generic complex with the description of a new genus, a new subgenus and four new species (Lepidoptera: Erebidae: Arctiinae: Lithosiini: Lithosiina). Ecologica Montenegrina 87: 46-76, DOI: 10.37828/em.2025.87.2, URL: https://doi.org/10.37828/em.2025.87.2
03FA570AFC06FFBE7BF6FC66FD1768BD.taxon	description	(Figs 51, 87)	en	Volynkin, Anton V., Černý, Karel (2025): On the taxonomy of the Brunia Moore generic complex with the description of a new genus, a new subgenus and four new species (Lepidoptera: Erebidae: Arctiinae: Lithosiini: Lithosiina). Ecologica Montenegrina 87: 46-76, DOI: 10.37828/em.2025.87.2, URL: https://doi.org/10.37828/em.2025.87.2
03FA570AFC06FFBE7BF6FC66FD1768BD.taxon	materials_examined	Type material. Holotype (Figs 51, 87): male, “ Vietnam, Prov. Tuyen Quang | Na Hang Nature Reserve, 300 m | 105 ° 5 ' E, 22 ° 3 ' N, 22. II. – 5. III. 1997 | leg. G. Csorba ” / “ Slide | CW 00099 ♂ [prepared by A. Volynkin] ” (MWM / ZSM).	en	Volynkin, Anton V., Černý, Karel (2025): On the taxonomy of the Brunia Moore generic complex with the description of a new genus, a new subgenus and four new species (Lepidoptera: Erebidae: Arctiinae: Lithosiini: Lithosiina). Ecologica Montenegrina 87: 46-76, DOI: 10.37828/em.2025.87.2, URL: https://doi.org/10.37828/em.2025.87.2
03FA570AFC06FFBE7BF6FC66FD1768BD.taxon	diagnosis	Diagnosis. The forewing length is 13.0 mm in the male holotype. Dubatoloviana vieta sp. n. differs externally from the closely related D. navneetsinghi and D. fumidisca in the longer and narrower forewing with a more intense greyish-brown subterminal suffusion at the anal corner. The male genitalia of D. vieta sp. n. are most similar to D. fumidisca but can be distinguished by the longer, narrower and straighter uncus, the slightly shorter valva with a narrower cucullus and a proximally narrower distal saccular process. Additionally, the vinculum of the new species is longer than in the similar congener.	en	Volynkin, Anton V., Černý, Karel (2025): On the taxonomy of the Brunia Moore generic complex with the description of a new genus, a new subgenus and four new species (Lepidoptera: Erebidae: Arctiinae: Lithosiini: Lithosiina). Ecologica Montenegrina 87: 46-76, DOI: 10.37828/em.2025.87.2, URL: https://doi.org/10.37828/em.2025.87.2
03FA570AFC06FFBE7BF6FC66FD1768BD.taxon	distribution	Distribution. The new species is currently known only from its type locality in Northern Vietnam (Tuyen Quang Province). The record of ‘ Eilema trimacula Holloway’ from Laos (Kishida 2020) probably also belongs to D. vieta sp. n. but requires confirmation.	en	Volynkin, Anton V., Černý, Karel (2025): On the taxonomy of the Brunia Moore generic complex with the description of a new genus, a new subgenus and four new species (Lepidoptera: Erebidae: Arctiinae: Lithosiini: Lithosiina). Ecologica Montenegrina 87: 46-76, DOI: 10.37828/em.2025.87.2, URL: https://doi.org/10.37828/em.2025.87.2
03FA570AFC06FFBE7BF6FC66FD1768BD.taxon	etymology	Etymology. The specific epithet is derived from the Viet people native to modern-day Northern Vietnam and Southern China. The name is a noun in the nominative singular.	en	Volynkin, Anton V., Černý, Karel (2025): On the taxonomy of the Brunia Moore generic complex with the description of a new genus, a new subgenus and four new species (Lepidoptera: Erebidae: Arctiinae: Lithosiini: Lithosiina). Ecologica Montenegrina 87: 46-76, DOI: 10.37828/em.2025.87.2, URL: https://doi.org/10.37828/em.2025.87.2
03FA570AFC06FFB87BF6F987FC49692F.taxon	description	(Figs 52, 88) Dubatoloviana navneetsinghi Volynkin, 2021, Ecologica Montenegrina, 47: 45, figs 1, 2, 7, 10 (Type locality: “ NE India, Assam, Nambor Reserv [e] Forest, Garampani, 100 m, 26 ° 30 ' N, 93 ° 55 ' E ”).	en	Volynkin, Anton V., Černý, Karel (2025): On the taxonomy of the Brunia Moore generic complex with the description of a new genus, a new subgenus and four new species (Lepidoptera: Erebidae: Arctiinae: Lithosiini: Lithosiina). Ecologica Montenegrina 87: 46-76, DOI: 10.37828/em.2025.87.2, URL: https://doi.org/10.37828/em.2025.87.2
03FA570AFC06FFB87BF6F987FC49692F.taxon	materials_examined	Type material examined. Holotype (Figs 52, 88): male, “ N-E. India | Assam, Nambor Reserv [e] | Forest, Garampani, H = 100 m | 26 ° 30 ' N, 93 ° 55 ' E, | 21 – 29. Nov. 1997. | lg. V. Siniaev [recte: Sinyaev] & M. Murzin | Museum Witt ” / “ Slide | ZSM Arct. 2021 - 097 ♂ | A. Volynkin ” (MWM / ZSM). Paratype: male, same data as in holotype (MWM / ZSM). Note. The twenty-two female specimens originally included into the type series of D. navneetsinghi (Volynkin 2021) in fact represent a dark form of Brunia gibonica (Fig. 53) while the female of D. navneetsinghi remains unknown.	en	Volynkin, Anton V., Černý, Karel (2025): On the taxonomy of the Brunia Moore generic complex with the description of a new genus, a new subgenus and four new species (Lepidoptera: Erebidae: Arctiinae: Lithosiini: Lithosiina). Ecologica Montenegrina 87: 46-76, DOI: 10.37828/em.2025.87.2, URL: https://doi.org/10.37828/em.2025.87.2
03FA570AFC06FFB87BF6F987FC49692F.taxon	distribution	Distribution. The species is known only from its type locality in Assam, Northeast India (Volynkin 2021). Acknowledgements The senior author expresses his sincere thanks to the following colleagues for their kind assistance during visits to the collections under their care: Axel Hausmann, Mei-Yu Chen and Ulf Buchsbaum (ZSM); Alberto Zilli and Geoff Martin (NHMUK); and Wolfram Mey (MfN). Vladimir Dubatolov (Novosibirsk, Russia) and the late Vadim Zolotuhin (Ulyanovsk, Russia) are acknowledged for the photo of the holotype of Lithosia usuguronis Matsumura kindly provided. Images of specimens deposited in NHMUK are used with permission and are copyright of the Trustees of the Natural History Museum, London and made available under Creative Commons License 4.0, CC-BY (https: // creativecommons. org / licenses / by / 4.0 /).	en	Volynkin, Anton V., Černý, Karel (2025): On the taxonomy of the Brunia Moore generic complex with the description of a new genus, a new subgenus and four new species (Lepidoptera: Erebidae: Arctiinae: Lithosiini: Lithosiina). Ecologica Montenegrina 87: 46-76, DOI: 10.37828/em.2025.87.2, URL: https://doi.org/10.37828/em.2025.87.2
03FA570AFC06FFB87BF6F987FC49692F.taxon	description	Draudt, M. (1914) Arctiidae. – Tiger moths (Miltochrista to Stenosa). In: Seitz, A. (Ed.), Die Gross-Schmetterlinge der Erde. 10. Die Indo-Australischen Spinner und Schwärmer. Alfred Kernen, Stuttgart, pp. 135 – 223. [In German] Dubatolov, V. V. & Bucsek, K. (2016) New lichen-moth taxa (Lepidoptera, Arctiidae, Lithosiinae) from Vietnam. Euroasian entomological journal, 15 (3), 228 – 238. Dubatolov, V. V. & Zolotuhin, V. V. (2011) Does Eilema Hübner, [1819] (Lepidoptera, Arctiidae, Lithosiinae) present one or several genera? Euroasian entomological journal, 10 (3), 367 – 379 + 380, pl. VII. Fang, C. L. (2000) Lepidoptera. Arctiidae. Fauna Sinica (Insecta). 19. Science Press, Beijing, 590 pp. [in Chinese with English summary] Hampson, G. F. (1894) The Fauna of British India, including Ceylon and Burma Fauna Br. India (Moths). Vol. 2. Taylor and Francis, London, pp. v-xxii, 1 – 609. Hampson, G. F. (1900) Catalogue of the Arctiadae (Nolinae, Lithosiinae) in the collection of the British Museum. In: Catalogue of the Lepidoptera Phalaenae in the collection of the British Museum, 2. British Museum (Natural History), London, xx + 589 pp., pl. XVIII – XXXV. Holloway, J. D. (2001) The Moths of Borneo, part 7. Family Arctiidae, subfamily Lithosiinae. Malayan Nature Journal, 55, 279 – 486. ICZN (1999) International Code of Zoological Nomenclature. 4 th Edition. International Trust for Zoological Nomenclature, The Natural History Museum, London, 306 pp. Kirti, J. S. & Singh, N. (2015) Arctiid Moths of India. Vol. 1. Nature Books India, New Delhi, 205 pp. Kononenko, V. S. (2010) Noctuidae Sibiricae. Vol. 2. Micronoctuidae, Noctuidae: Rivulinae – Agaristinae (Lepidoptera). Entomological Press, Sorø, 475 pp. Krüger, M. (2015) Generic classification of Afrotropical footman moths sensu stricto (Lepidoptera: Erebidae: Arctiinae: Lithosiini (partim )). Transvaal Museum Monograph, 15, 1 – 176. Lafontaine, J. D. & Mikkola, K. (1987) Las-och-nyckel systemen i de inre genitalierna av Noctuidae (Lepidoptera) som taksonomiska kaennetecken. [Lock-and-key systems in the inner genitalia of Noctuidae (Lepidoptera) as a taxonomic character.] Entomologiske Meddelelser, 55, 161 – 167. [In Swedish with English summary] Matsumura, S. (1927) New species and subspecies of moths from the Japanese Empire. Journal of the College of Agriculture, Hokkaido Imperial University, Sapporo, Japan, 19 (1), 1 – 91, pls. 1 – 5. [http: // hdl. handle. net / 2115 / 12601] Moore, F. (1878) A revision of certain genera of European and Asiatic Lithosiinae, with characters of new genera and species. Proceedings of the general meetings for scientific business of the Zoological Society of London, 1878, 3 – 37, pls. 1 – 3. https: // doi. org / 10.1111 / j. 1469 - 7998.1878. tb 07927. x Singh, N., Singh, S., Datta, H. S. & Kirti, J. S. (2024) Taxonomic review of the genera Cernyia Bucsek and Denteilema Dubatolov: resolving a group of cryptic species (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Annales Zoologici (Warszawa), 74 (1), 129 – 141. https: // dx. doi. org / 10.3161 / 00034541 ANZ 2024.74.1.007 Volynkin, A. V. (2020) Re-characterization of two Lithosiini species recently described from the island of Taiwan, with establishing of a new combination and the corrected checklist of the genus Aberrasine Volynkin & Huang, 2019 (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Ecologica Montenegrina, 38, 119 – 126. http: // dx. doi. org / 10.37828 / em. 2020.38.15 Volynkin, A. V. (2021) Taxonomic review of the genus Dubatoloviana Bucsek with description of a new species from Northeast India (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Ecologica Montenegrina, 47, 43 – 49. http: // dx. doi. org / 10.37828 / em. 2021.47.7 Volynkin, A. V. (2024) On the terminology of the genitalia structures of lichen moths (Lepidoptera: Erebidae: Arctiinae: Lithosiini) with some references to Noctuidae. Ecologica Montenegrina, 73, 176 – 207. https: // dx. doi. org / 10.37828 / em. 2024.73.18 Volynkin, A. V. & László, G. M. (2021) Kruegerilema, a new genus for a new species endemic to São Tomé Island (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Bonn zoological Bulletin, 70 (2), 333 – 338. https: // doi. org / 10.20363 / BZB- 2021.70.2.333 Watson, A., Fletcher, D. S. & Nye, I. W. B. (1980) The generic names of moths of the world. Vol. 2. Noctuoidea (part): Arctiidae, Cocytiidae, Ctenuchidae, Dilobidae, Dioptidae, Lymantriidae, Notodontidae, Strepsimanidae, Thaumetopoeidae, Thyretidae. Trustees of the British Museum (Natural History), London, 228 pp. https: // doi. org / 10.5962 / bhl. title. 119421 Wu, S., Fu, C. - M., Tzuoo, H. - R., Shih. L. - C., Chang, W. - C., Lin, H. - H. (2020) An annotated checklist of macro moths in mid- to high-mountain ranges of Taiwan (Lepidoptera: Macroheterocera). Formosan Entomologist, 40, 10 – 83. https: // doi. org / 10.6662 / TESFE. 202002 _ 40 (1). 002	en	Volynkin, Anton V., Černý, Karel (2025): On the taxonomy of the Brunia Moore generic complex with the description of a new genus, a new subgenus and four new species (Lepidoptera: Erebidae: Arctiinae: Lithosiini: Lithosiina). Ecologica Montenegrina 87: 46-76, DOI: 10.37828/em.2025.87.2, URL: https://doi.org/10.37828/em.2025.87.2
