identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03FB65461A1E7A18FF1ADCF8CDB47685.text	03FB65461A1E7A18FF1ADCF8CDB47685.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Actinopus chilikuti Ríos-Tamayo & Goloboff 2025	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Actinopus chilikuti sp. nov.</p>
            <p>(Figs 1–5)</p>
            <p> Actinopus septemtrionalis Ríos-Tamayo &amp; Goloboff, 2018: 26 (only female from Formosa province). Misidentification. </p>
            <p>
                  Holotype. ARGENTINA: Santiago del Estero Prov: 1 ♂ (FML-ARAN 02273) from  
                <a title="Search Plazi for locations around (long -61.958523/lat -25.972542)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-61.958523&amp;materialsCitation.latitude=-25.972542">Parque Nacional “Copo”</a>
                 , seccional  
                <a title="Search Plazi for locations around (long -61.958523/lat -25.972542)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-61.958523&amp;materialsCitation.latitude=-25.972542">La Portada</a>
                 (25°58’21.15”S 61°57’30.68”W), 23 November 2023, Ríos-Tamayo. D &amp; P. Goloboff leg.  Paratypes. Same data as holotype, 1 ♀ (FML-ARAN 02274), 24 November 2023, Ríos-Tamayo. D &amp; J.C Stazonelli leg. ;  3 ♂♂ (FML-ARAN 02275), 23 November 2023, Ríos-Tamayo. D, J.C Stazonelli &amp; P. Goloboff leg.
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                 Other material examined.   ARGENTINA: Formosa Prov: 1 ♀ (MACN-Ar 27307), tissue sample (MACN-Ar 27307-t1) from  
                <a title="Search Plazi for locations around (long -61.801502/lat -24.300068)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-61.801502&amp;materialsCitation.latitude=-24.300068">Reserva Natural Formosa</a>
                 : Seccional Rio Teuco, (24°18’0.25”S 61°48’5.41”W), 27 June–5 July 2010, Izquierdo. M, Kopuchian. C and Calderón. L. leg. 
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            <p>Etymology. The specific epithet works as a noun in apposition referring to the small size of the specimens of this species. In Quechua Aymara (very commonly spoken in Santiago del Estero), ch’ilikuti means “little, small”.</p>
            <p> Diagnosis. Members of  Actinopus chilikuti sp. nov. are small spiders. Males can be distinguished from all other species with Type I morphology in Argentina by having a copulatory bulb with a more curved embolus (Figs 2C–F, 4C), with keels well developed (similar to  A. reycali ) and PA developed (smaller in  A. reycali , and  A. clavero but bigger in  A. septemtrionalis , Fig 4). Females can be differentiated from  A. reycali and  A. septemtrionalis by the shorter and more slender spermathecae (Fig 3D, similar to  A. clavero ). It is distinguished from  A. clavero by a larger number of retrolateral thorns on tibia II (30 vs 20, Fig 3G) and fewer on tibia I (0 vs 5). </p>
            <p>Description. Male holotype (FML-ARAN 02273). Total length: 6.61. Carapace (Fig 1A): length 3.30, width 2.93; cephalic region: length 1.89, width 2.01; clypeus small, 7 bristles between PME, 4 behind each PME–PLE. Fovea: length 0.44, width 0.80. Eyes, diameters and interdistances (Fig 1C): AME 0.19 (large), ALE 0.27, PME 0.14, PLE 0.16, ocular group length 0.54, anterior width 1.32, 1.23 posterior width; AME–ALE 0.20, PME–PLE 0.06, AME–PME 0.19, ALE–PLE 0.19, AME–AME 0.09, PME-PME 0,74. Chelicerae: apex with 7 recumbent and elongated bristles over fang base. Rastellum with 4 marginal blunt spines and 2 small dorsal ones. Cheliceral furrow with 4 large promarginal and 5 retromarginal teeth and 4 denticles, near promarginal teeth. Labium (Fig 1B): length 0.52, width 0.55. Maxillae (Fig 1B): anterior length 0.78, posterior length 1.18, width 0.78. Sternum (Fig 1B): Center flat, length 2.15, maximum width 1.66, minimum width 0.87. Postlabial sigilla square, posterior sigilla small and well defined. Abdomen (Fig 1E, F): length 2.99, covered by small bristles. PMS: length 0.32; PLS: length of basal:medial:apical segments 0.38:0.17:0.13. Lengths of legs and palp: I: 3.92, 1.46, 2.52, 2.83, 1.60, 12.33. II: 3.70, 1.40, 2.40, 2.76, 1.54, 11.80. III: 2.92, 1.42, 1.77, 2.74, 1,59, 10.44. IV: 3.67, 1.51, 2.87, 3.04, 1.85, 12.94. Palp: 4.15, 1.98, 3.50,–, 0.81, 10.44. Chaetotaxy: Trochanter: III, p 1; I-II-IV, 0. Femora: I-II, 0; III, d 1-1-2; IV, 0. Patellae: I- II 0; III, d 1-1-1-1, p 16, 14 on the margin (crown of thorns); IV, p 21.Tibiae: I, r 1-1-2, v 1-2-1-2-1-3; II, r 13, v 1-1-2-3; III, v 2-2-2, r 2-2, d 1, with apical crown of 14 thorns; IV, v 1-1-1-1-1-3 (spiniform). Metatarsi: I, r 1-1-1-1-1-1-2, v 2-2-2, p 1-1-2-1-1; II, r 1-1-1-2-2, v 2-1-1-1-4 (apical), p 1-1; III, p 1-1-2-1-1-2, v 1-1-1-1, r 1-1-2-1-2-3; IV, v 2-2, p 1-2-3-2-1-3, r 1-0-2-1-2. Tarsi: I, r 2-1-2-2-1-4, v 2-2-1-1-1, p 1-1-1-1; II, r 18, v 1-1-1-1- 1-1-1, p 1-1-1-1-1; III, r 1-2-1-1, p 1-1-1; IV, r 1, p 1-2-2-2-1. Total number of retrolateral spines on tibiae I, II (Fig 1D): 4 and 13, respectively. Palp: tibia long, thin (Fig 2A, B). Bulb (Fig 2C–F) slim and slender. TA developed, PA developed. Embolus without denticles and with PS, PI, and Pac well developed. Pseudoscopula: Tarsi: I–II, sparse; III-IV, dense, uniform. Metatarsi: I-II absent, III apical and sparse, IV 1:3 apical. Trichobothria: Tibiae: I 3-3; II-II 4-3; IV 5-5. Metatarsi: I-II 8; III 6; IV 7. Tarsi: I -II 5; III 4; IV 5. Colour in alcohol: cephalic region and chelicerae reddish brown, carapace lighter. Abdomen brown; book lungs with light markings. Sternum reddish brown with a lighter center mark. Legs like cephalic region, legs III-IV with lighter tarsi–metatarsi; palp with tibia lighter. Spinnerets yellowish.</p>
            <p>Female paratype (FML-ARAN 02274). Total length: 9.57. Carapace (Fig 3A): length 4.58, width 4.44; cephalic region: length 3.20, width 4.40; clypeus small with 12 bristles; 10 bristles between PME, 4 behind each PME – PLE, 5 bristles and one larger on both sides of ocular group. Fovea: length 0.47, width 1.08. Eyes, diameters and interdistances (Fig 3F): AME 0.11, ALE 0.27, PME 0.21, PLE 0.17, ocular group 0.80, anterior width 2.39, 2.09 posterior width; AME – ALE 0.73, PME – PLE 0.11, AME – PME 0.47, ALE – PLE 0.53, AME – AME 0.54, PME – PME 1.32. Chelicerae: cheliceral apex with 9 recumbent and elongated bristles covering base of fang. Rastellum with 10 marginal, blunt spines and 5 small dorsal. Cheliceral furrow with 6 promarginal and 5 retromarginal teeth and 8 denticles in furrow, near promargin. Labium (Fig 3B): length 1.13, width 0.97, with 12 anterior cuspules. Maxillae (Fig 3B): anterior length 1.24, posterior length 1.67, width 1.35, with ~95/100 cuspules 1:3 ventral anterior, expanding to anterior face. Sternum (Fig 3B): Center flat, length: 3.38, maximum wide 2.67, minimum width 1.63. Postlabial sigilla square, posterior ones shallow, well defined. Abdomen (Fig 3C, E): length 5.02, covered by brown bristles. PMS: length 0.56; PLS with basal: medial: apical articles 0.62:0.23:0.20. Lengths of legs and palp: I: 2.76, 1.83, 1.46, 1.64, 0.91, 8.60. II: 2.79, 1.86, 1.44, 1.71, 0.85, 8.65. III: 2.67, 2.09, 1.11, 1.73, 0.78, 8.38. IV: 3.33, 1.95, 2.05, 2.04, 0.79, 10.16. Palp: 2.46, 1.66, 1.65, –, 1.81, 7.58.</p>
            <p>Chaetotaxy: All femora: 0. Patellae: I–II, 0; III, p 4 + 9 on margin, r 7 on margin; IV, p-d 38; palp, p 1-1. Tibiae: I, v 1-1-1; II, r 30, v 1-1 spiniform setae; III, d 1, with apical crown of 15 thorns, r 12; IV, 0; palp, p 1-1-1-1-1-1, r 1-1-2-1-1. Metatarsi: I, r 7, v 1-1-2, p 14; II, r 1-2-1-1-1-1, p 1-1-1-1-2, v 1-1-2 near to the r ones; III, r 17 on row, v 2, p 10 on row; IV, p 2-1-2, d 1 anterior. Tarsi: I, r 10, v2, p 6; II, p 11, v 2, r 8; III, r 11, v-p 16; IV, v-p 24; palp, r 16, p 10, v 2 apical. Total number of retrolateral spines on tibiae I and II (Fig 3G): 0 and 30, respectively. Trichobothria: Tibiae: I 3-3; II-III 3-4; IV 5-6. Metatarsi: I-II 9; III-IV 7. Tarsi: I 6; II 5; III-IV 4. Colour: cephalic region and chelicerae greenish brown, carapace lighter. Abdomen brown; book lungs with light markings. Sternum greenish brown with a lighter center mark (Fig 3B), labium and maxillae darker. Legs like cephalic region. Spinnerets yellowish.</p>
            <p>Distribution. Provinces of Santiago del Estero and Formosa (Fig 5).</p>
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	https://treatment.plazi.org/id/03FB65461A1E7A18FF1ADCF8CDB47685	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Ríos-Tamayo, Duniesky;Goloboff, Pablo A.	Ríos-Tamayo, Duniesky, Goloboff, Pablo A. (2025): A new trapdoor spider of the genus Actinopus (Araneae: Mygalomorphae: Actinopodidae) from Argentina. Zootaxa 5563 (1): 255-263, DOI: 10.11646/zootaxa.5563.1.17, URL: https://doi.org/10.11646/zootaxa.5563.1.17
03FB65461A197A1BFF1AD9A7CB97749A.text	03FB65461A197A1BFF1AD9A7CB97749A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Actinopus Perty 1833	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Key to known species of  Actinopus with Type I Morphology </p>
            <p> 1. Males.............................................................................................. 2 - Females............................................................................................. 6 2. Small size (body length &lt;11.00 mm); color light brown or beige; spiniform bristles on dorsum of femora III and P on trochanter III................................................................................................. 3 - Larger size (body length&gt;12.00), color dark reddish brown, without spiniform bristles on dorsum of femora III or P on trochanter III........................................................  A. reycali Ríos-Tamayo &amp; Goloboff, 2018 . 3. Bulb keels and ATA very small, scarcely visible...........................  A. clavero Ríos-Tamayo &amp; Goloboff, 2018 . - Bulb keels and ATA developed, remarkably visible........................................................... 4 4. Chelicerae with small anterior retrolateral keels; PA very developed (Fig 4D....................................... ...........................................................  A. septemtrionalis Ríos-Tamayo &amp; Goloboff, 2018 . - Chelicerae without keels; PA developed (Fig 4C, 4E .......................................................... 5 5. Two bulb keels (Fig 4E); fovea wide (&gt;1.80)....................................  A. cochabamba Ríos-Tamayo, 2016 . - Three bulb keels (Fig 4C); fovea narrow (&lt;1.00)...............................................  A. chilikuti sp. nov. 6. Spermathecae elongated (fig 7D, E in Ríos-Tamayo &amp; Goloboff, 2018, p. 19)..............................  A. reycali - Spermathecae short (Fig 3D)............................................................................ 6 6. Apical portion of the spermathecae broad (fig 9D, E in Ríos-Tamayo &amp; Goloboff, 2018, p. 24, fig. 2E in Ríos-Tamayo, 2016, p. 187).............................................................................................. 7 - Apical portion of the spermathecae slender................................................................. 8 7. Few retrolateral spines on tibia II (&lt;20).......................................................  A. septemtrionalis - Many retrolateral spines on tibia II (&gt;30)........................................................  A. cochabamba 8. Few retrolateral spines on tibia I..................................................................  A. clavero - Absence of retrolateral spines on tibia I.....................................................  A. chilikuti sp. nov.</p>
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	https://treatment.plazi.org/id/03FB65461A197A1BFF1AD9A7CB97749A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Ríos-Tamayo, Duniesky;Goloboff, Pablo A.	Ríos-Tamayo, Duniesky, Goloboff, Pablo A. (2025): A new trapdoor spider of the genus Actinopus (Araneae: Mygalomorphae: Actinopodidae) from Argentina. Zootaxa 5563 (1): 255-263, DOI: 10.11646/zootaxa.5563.1.17, URL: https://doi.org/10.11646/zootaxa.5563.1.17
03FB65461A197A1AFF1ADD01CAEC72DD.text	03FB65461A197A1AFF1ADD01CAEC72DD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Actinopus argenteus Rios-Tamayo & Goloboff 2018	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Actinopus argenteus Ríos-Tamayo &amp; Goloboff 2018: 26 . </p>
            <p>
                 Material examined.   ARGENTINA: Santiago del Estero Prov:  
                <a title="Search Plazi for locations around (long -61.958523/lat -25.972542)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-61.958523&amp;materialsCitation.latitude=-25.972542">Parque Nacional “Copo”</a>
                 , seccional  
                <a title="Search Plazi for locations around (long -61.958523/lat -25.972542)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-61.958523&amp;materialsCitation.latitude=-25.972542">La Portada</a>
                 (25°58’21.15”S 61°57’30.68”W), 23 November 2023, Ríos-Tamayo. D., P. Goloboff, &amp; J.C Stazonelli leg, 7 ♂ (FML-ARAN 02294);  same data, 2 ♀ (FML-ARAN 02298) . 
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            <p>Diagnosis: See Ríos-Tamayo &amp; Goloboff, 2018: 27.</p>
            <p>Distribution: ARGENTINA: Center of Santiago del Estero, Córdoba and Catamarca provinces.</p>
            <p>New record: ARGENTINA, North of Santiago del Estero province, Parque Nacional “Copo”.</p>
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	https://treatment.plazi.org/id/03FB65461A197A1AFF1ADD01CAEC72DD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Ríos-Tamayo, Duniesky;Goloboff, Pablo A.	Ríos-Tamayo, Duniesky, Goloboff, Pablo A. (2025): A new trapdoor spider of the genus Actinopus (Araneae: Mygalomorphae: Actinopodidae) from Argentina. Zootaxa 5563 (1): 255-263, DOI: 10.11646/zootaxa.5563.1.17, URL: https://doi.org/10.11646/zootaxa.5563.1.17
03FB65461A187A1AFF1ADB7CCED9711D.text	03FB65461A187A1AFF1ADB7CCED9711D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Actinopus coylei Rios-Tamayo & Goloboff 2018	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Actinopus coylei Ríos-Tamayo &amp; Goloboff 2018: 36 . </p>
            <p>
                 Material examined.   ARGENTINA: Santiago del Estero Prov:  
                <a title="Search Plazi for locations around (long -61.958523/lat -25.972542)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-61.958523&amp;materialsCitation.latitude=-25.972542">Parque Nacional “Copo”</a>
                 , seccional  
                <a title="Search Plazi for locations around (long -61.958523/lat -25.972542)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-61.958523&amp;materialsCitation.latitude=-25.972542">La Portada</a>
                 (25°58’21.15”S 61°57’30.68”W), 23 November 2023, Ríos-Tamayo. D., P. Goloboff, &amp; J.C Stazonelli leg, 1 ♂ (FML-ARAN 02293);  same data, 1 ♂ (FML-ARAN 02297);  6 km de Secc. “  Salum ”, camino a Secc “  Guayacan ”, Ríos-Tamayo. D., P. Goloboff, &amp; J.C Stazonelli, 2 ♀ (FML-ARAN 02295)  . 
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            <p>Diagnosis: See Ríos-Tamayo &amp; Goloboff, 2018: 36.</p>
            <p>Distribution: ARGENTINA: Córdoba and San Luis provinces.</p>
            <p>New record: ARGENTINA, North of Santiago del Estero province, Parque Nacional “Copo”.</p>
            <p>Natural history</p>
            <p> Adult males of  A. argenteus ,  A. coylei and the new species here described were found walking during the night, in seccional La Portada after a big storm. It is known, females of the genus prefer to build their burrows in sloping surfaces (Ríos-Tamayo &amp; Goloboff, 2018). In this trip many females of the species mentioned, were observed living sympatrically in the flat areas around and inside of the Seccional La Portada as well as in other areas of the National Park. </p>
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	https://treatment.plazi.org/id/03FB65461A187A1AFF1ADB7CCED9711D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Ríos-Tamayo, Duniesky;Goloboff, Pablo A.	Ríos-Tamayo, Duniesky, Goloboff, Pablo A. (2025): A new trapdoor spider of the genus Actinopus (Araneae: Mygalomorphae: Actinopodidae) from Argentina. Zootaxa 5563 (1): 255-263, DOI: 10.11646/zootaxa.5563.1.17, URL: https://doi.org/10.11646/zootaxa.5563.1.17
