identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
038787EC0329FFCBCE84E6116ABAFEB4.text	038787EC0329FFCBCE84E6116ABAFEB4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cicindela coerulea Pallas 1773	<div><p>Cicindela (s. str.) coerulea Pallas, 1773</p><p>Redescription. TL = 12.0– 15.2 mm (mean = 13.77 mm, n = 164) in males, 12.2–16.1 mm (mean = 14.4 mm, n = 132) in females.</p><p>Coloration highly variable from unicoloured black, brown, blue, green, olive, cupreous-bronze or purple to bicolored with blue head and pronotum and blue-green or green elytra, or with blue or blue-green head and pronotum and cupreous or cupreous-green elytra.</p><p>Head wider than pronotum, with protruding eyes; genae glabrous, deeply striated; clypeus asetose; frons long, virtually flat, finely striated; vertex with sparse soft hairs, shallowly rugose, with two wide shallow longitudinal depressions in the base of ocular plates; latter shallowly striated, with two long anterior and posterior setae; occiput indistinctly rugose. Labrum pale with dark brown or black narrowly infuscate wave-shaped anterior margin, unidentate, transverse, slightly trapeziform, especially in females, LW/LL = 1.83–2.50 mm (mean = 2.17 mm, n = 164) in males, 1.77–2.44 mm (mean = 2.06 mm, n = 131) in females, with 4–14 (mean 8, n = 299) submarginal setae. Mandibles medium length, in males relatively stout, ML/MW = 5.43–8.0 (mean = 6.58, n = 63), in females thinner, ML/MW = 6.0–8.5 (mean = 7.38, n = 47); black with greenish-blue tinge except long pale lateral margin up to base of apical tooth. Maxillary palpi metallic blue, bluish-green, green or cuprous-green with purple or golden reflection. Labial palpomeres 2 and 3 pale, palpomere 1 brown, palpomere 4 metallic green or bluish green.Antennae relatively long, reach basal third of elytra, antennomeres 1–4 metallic blue, bluish-green or green with purple or golden reflection, antennomeres 5–11 black, densely covered by numerous very short white setae, each with two longer apical setae; scape besides 3–5 apical setae often with 1–3 rarely with 4–5 additional setae or glabrous; antennomere 3 with 4–10 and antennomere 4 with 1–6 short white setae.</p><p>Pronotum transverse, PW/PL = 1.21–1.52 (mean = 1.37, n = 173) in males, 1.28–1.57 (mean = 1.42, n = 138) in females; widest in anterior third, with slightly rounded and convergent to the base lateral margins, slightly wrinkled, with very deep anterior and posterior grooves so that central part of pronotal disc projected as two suboval humps, midline thin but distinct; female coupling sulcus long and shallow, slightly sinuate near the middle. Legs metallic black-violet, blue or bluish-green; fore- and mid-trochanters with single apical seta.</p><p>Elytra relatively long, subparallel, indistinctly dilated near the middle, with angular shoulders; EL/EW = 1.40–1.71 (mean = 2.55, n = 173) in males, 1.40–1.62 (mean = 1.51, n = 138) in females; with smooth punctured microsculpture except more granular humeral area; suture smooth, slightly raised, ended by small apical spine; apical margins finely microserrulated; epipleura some coloured as elytra except narrow brownish outer margin. White elytral pattern extremely variable from completely undeveloped to very wide with humeral lunula, middle band and apical lunula connected by marginal band. Abdominal sternites black, violet, blue or bluish-green with golden-green or greenish-blue tinge, sparsely pubescent in central portions and densely haired by white setae laterally, with submarginal row of long white setae.</p><p>Aedeagus relatively long, distinctly depressed on apical third in right side (see in ventral view), EL/AL = 1.64– 2.0 (mean = 1.80, n = 81), apex lanceolate, clearly curved with sharp hook, short lateral carina and small groove on dorsal side. Internal sac large, distinctly rounded towards to ventral side, fl long, sp long and smoothly curved, mt long, rarely short, as a rule with smoothly curved apex, VA and DA large, B distinctly larger than DLL, VLR prolonged, VB and VLL as flat membranous fields, BLR virtually invisible.</p><p>Posterior margin of female sternum VIII with 2–4 short stout setae on each apex, asetose inner V-shaped margin or with a pair seta on it, and with 6–10 long setae along each margin. Syntergum IX&amp;X with 10–32 long apical setae and 18–38 long setae along each margin. Base of second gonapophyses with 2–6 setae; ventral notch of second gonacoxa with 4–7 long setae; sclerite of bursa copulatrix large, and additional sclerite between gonacoxa small.</p></div>	https://treatment.plazi.org/id/038787EC0329FFCBCE84E6116ABAFEB4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Matalin, Andrey V.	Matalin, Andrey V. (2025): An intraspecific structure of Сicindela (s. str.) coerulea Pallas, 1773 (Coleoptera, Cicindelidae). Zootaxa 5583 (2): 201-246, DOI: 10.11646/zootaxa.5583.2.1, URL: https://doi.org/10.11646/zootaxa.5583.2.1
038787EC032EFFC4CE84E5386D8DFD94.text	038787EC032EFFC4CE84E5386D8DFD94.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cicindela coerulea Pallas 1773	<div><p>Cicindela (s. str.) coerulea coerulea Pallas, 1773</p><p>Figs 1–9, 14–18, 24–44, 65–75</p><p>Cicindela coerulea Pallas, 1773: 724 (Type locality—Irtin [= Irtysh]). =  Cicindela violacea Gebler, 1817: 324 (Type locality—fl. Irtüsch), synonymized by Dejean (1833: 2). =  Cicindela coerulea lucifera Mandl, 1970: 72, Abb. 3 figs 1, 2 (Type locality—Biysk, Altai), syn. nov.  Cicindela coerulea Pallas, 1773: Pallas 1781: Tab. G fig. 12; Fischer von Waldheim 1828: 17; 1829: 5; 1842: 4; Gebler 1830:</p><p>26; 1833: 262; 1847: 268; Dejean 1833: 2; 1836: 3; Dejean &amp; Boisduval 1837: 25 Pl. 3 fig. 4; Sturm 1843: 2; Motschoulski,</p><p>1846: 30; Motschulsky, 1850: 1; Chaudoir 1865: 36; Fleutiaux 1892: 106; Horn 1938: 46, Taf. 63 fig. 22; Rivalier 1950:</p><p>225; Schilder 1953: 564; Werner, 1991: 18, T. 14 figs 100, 101; Dubatolov et al. 1994: 4; Matalin &amp; Zimina 2002: 230.  Cicindela violacea Gebler, 1817: Fischer von Waldheim 1820: 8, Tab. 1 fig. 4; 1821: 102.  Cicindela tricolor coerulea Pallas, 1773: Horn &amp; Roeschke 1891: 50.  Cicindela hybrida nitida Lichtenstein, 1796: Wu 1937: 15.  Cicindela hybrida coerulea Pallas, 1773: Horn 1905a: 158; 1905b: 27; 1915: 337; 1926: 221; Winkler 1924: 4.  Cicindela grumi coerulea Pallas, 1773: Schilder 1953: 564.  Cicindela coerulea tenuifascia Fischer von Waldheim, 1828: Lorenz 1998: 43.  Cicindela coerulea lucifera Mandl, 1970: Cassola &amp; van Nidek 1984: 10; Wiesner 1992: 118; 2020: 184; Kryzhanovskij et al.</p><p>1995: 26; Puchkov &amp; Matalin 2003: 104; Lorenz 2005: 131; Putchkov &amp; Matalin 2017: 227; Makarov et al. 2020.  Cicindela coerulea coerulea Pallas, 1773: Mandl, 1935: 294 figs 9, 10, 296 figs 11, 12; 1936: 9, 14; Cassola &amp; van Nidek 1984:</p><p>10; Wiesner 1992: 117; 2020: 183; Kryzhanovskij et al. 1995: 26; Lorenz 1998: 42; 2005: 131; Puchkov &amp; Matalin 2003:</p><p>104; Shook &amp; Wiesner 2006: 10; Wu &amp; Shook 2010: 77; Wu 2011: 24; Matalin &amp; Dudko 2016a: 140; 2016b: 143; Putchkov &amp; Matalin 2017: 227; Makarov et al. 2020; Štrunc 2020: 142.</p><p>Type material.   Neotype of  Cicindela coerulea Pallas, 1773 (designated here), ♂ (Figs 1, 2, 28, 42)—Irtys r. [handwritten white label],  tricolor coerulea Pall. [handwritten white label], NEOTYPUS,  Cicindela coerulea Pallas, 1773, des. A. V. Matalin, 2024 [printed red label] (ZIN ).</p><p>Paratype of  Cicindela coerulea lucifera Mandl, 1970,   ♂ — Altai, Biysk, bank of Katun’ river. 16.VI.1952, Selivanovskiy [typed white label], Paratypus,  Cicindela coerulea ssp. lucifera m. det. Dr. K. Mandl, 1970 [handwritten and typed red label], coll. K. Mandl [typed white label], 222,  Cicindela coerulea ssp. lucifera Mandl [handwritten white label] (NHMW).</p><p>Additional material.   RUSSIA, Chelyabinsk Oblast: 1♂ —middle stream of  Miass river, coll. I. Moiseenko (cISr) ;   Novosibirsk Oblast: 1♂ —  Karasuk Distr., near Troitskoe, steppe, 28. VI.1967, leg. Zolotarenko ;   1♂ — Karasuk Distr., near Troitskoe,  lake Krotovaya Lyacha, 2.VII.1979, leg. T.D.  Gorelova (both ISEA) ;   8♂♂ 3♀♀ — Inskoy b.,  st. Meret, 21.VI.1898, 18.VIII.1898, 31.VIII.1898 (ZIN, ZMUM, ISEA, NHMW) ;  Tomsk Oblast: 2♀♀ — Tomsk;   1♀ —  Chernyi Yar, 16. VI.1908 (both ZIN) ;   Altai  Krai: 1♂ 1♀ —  Sib. oc., step. Kulunda, 7. VI –19.VII.1922, leg. A. Reichardt ;   1♀ — Slavgorod Kluchi, Omsk reg.,  Slavgorod area, 29. V.1922, leg. A. Reichardt;   3♀♀ — Severnyi, Slavgorod area,  Omsk reg., 24.VII.1922, leg. N.N. Pukhova ;   1♀ — Altai, env.  Kolyvan, VII.1894, leg. F.F. Morawits ;   1♂ — Altai,  Zmeinogorsk reg., env. Novenskoe (al ZIN) ;   1♂ — Lokot,  Rubtsovsk reg., 8.VII.1920, N. Gorbunov ;   1♂ 1♀ — Mikhailovka Distr., env.  Nikolaevka, 10.VII.1980, leg. Yu. Perunov ;   1♀ — Zapadno-Sibirskiy  Krai, near  Biysk, 28. V.1934;   2♂♂ 3♀♀ —near Biysk,  Katun River, 15. VI –18.VIII.1952,  Selivanovsky (all ZMUM) ;  1♂ 1♀ — ibid (cJG);   1♀ —  Barnaul, 26. VI.1909, leg. B. Popov (ISEA);   1♂ 2♀♀ —N  Altai, 20 km S of Biysk, 10–13.VIII.1991, leg. M. Chernyakhovskiy (MSPU) ;   KAZAKHSTAN, North Kazakhstan Region: 2♂♂ 3♀♀ —  Kuybyshev distr., state farm Ruzaevskiy, plough-land, VI –VIII.1982, leg. I.B. Knor (ISEA);   Akmola Region: 1♀ — Selety River, Alchinbay mount., 15 verst from  Gogolevsk, 9.VII.1900, leg. N. Petrovskiy (ISEA) ;   Pavlodar Region: 1♂ —E Kazakhstan, near  Pavlodar, research station of Kaz. IZR, 5. VI.1961, leg. I. Kabak (cEK);   3♂♂ 3♀♀ —  Pavlodar;   3♂♂ — Maraldy-Romanovka,  Pavlodar Area, Semipalatinsk Region, 3.VIII.1922, leg. A. Reichardt ;   2♂♂ 2♀♀ —  Maraldy lake;   1♂ — Aynakol, Pavlodar area,  Akmolinsk region, 8.VIII.[1]922, leg. A. Reichardt (all ZIN) ;   1♂ —state farm  Chernoyarka, steppe, 19. VI.1985, leg. Karabutova;  1♂ — ibid, meadow, 25. VI.1987, leg. Kozhukhanova (ISEA);  12♂♂ 5♀♀ —NE Kazakhstan, right bank of Irtysh Riv., env. of Zhertumsyk (= <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=77.19833&amp;materialsCitation.latitude=52.031113" title="Search Plazi for locations around (long 77.19833/lat 52.031113)">Karla Marksa</a>) vill., 28 km SSE of Pavlodar City, 52°01’52’’N 77°11’54’’E, h=122 m, 23– 24.05.2021, leg. D.A. Zakharov (cDZ) ;   Ulytau Oblast: 2♂♂ 4♀♀ —ur.  Karaagash, 60 km NE Zhana-Arka, 3.VII.1961, leg. G. Medvedev, 19. V.1962, leg. L. Arnoldi (all ZIN);  Abai Region: 1♂ — Talitsa, 1. V.[19]08;  3♂♂ 3♀♀ —  Semipalatinsk (both ZIN) ;   1♀ —  Semipalatinsk, Sibirien (NHMW) ;   2♂♂ 1♀ — Kazakhstan, near  Semipalatinsk, 15–22. V.1991, leg. S. Khvylya (MSPU);   1♂ — Kazakhstan, 60 km NE of  Semipalatinsk, 3.VII.1958 (cEK) ;   1♂ —E Kazakhstan, Semipalatinsk Area,  Novopokrovka, 15.05.1972, leg. A.S. Badenko (cBK) ;   1♂ 1♀ — Semipalatinskiy leskhoz,  Kashtakskoe lesnichestvo, 10.VIII.1955 (ZMUM) ;   1♂ — Dzhertasskiy (= Djartas),  Semipalatinsk oblast, 16. V.1971, leg. Merenkova (ISEA);  East Kazakhstan Region: 2♂♂ 6♀♀ — Ust-Kamenogorsk, 10. VI.30, VI.1908 (ZIN, cPU);  1♀ —E Kazakhstan, Ust-Kamenogorsk City, bank of  Irtysh Riv., 14.06.1979, leg. V.  Ryzhikh (cBK) ;   1♂ — Korosteli, Kaz. №1,  Rubtsovsk Reg., 1.VII.1923, Isaev ;   1♂ 1♀ — Berezovka,  Bukhtarma river, Ust-Kamenogors Reg  .;   2♀♀ —  Kaznakovo, 20. V.1930, F. Zaitsev;   2♂♂ 4♀♀ —  Altai mer., suso Zaysan, 1991 ;  1♂ 1♀ —Irtysch;  2♀♀ —  Belaya river, S Altai, 6.VIII.1930, leg. Lyukyanovich (all ZIN) ;   5♂♂ 3♀♀ — East Kazakhstan reg., <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=83.28732&amp;materialsCitation.latitude=48.72187" title="Search Plazi for locations around (long 83.28732/lat 48.72187)">Kyzylkum</a> sands, 19 km S Kulynzhon, h=223 m, 48.72187°N, 083.28732°E, 09.05.2022, leg. S. V.  Kolov (IZK) ;   3♂♂ 3♀♀ —  Zaysan, 15 km SW of Kaznakovka, sands, 20.05.80, leg. Nikolaev; ibid, 25–31. V.89, M.  Danilevsky (ZIN, MSPU, ISEA, cJG) ;  4♂♂ 2♀♀ —N Zaysan, 300 m, V.1989 (cJW); 1♂ 1♀ — ibid (HMHN);   3♂♂ 2♀♀ — N Zaysan Lake, env. of  Kaznakovka vill., h=398 m, 21–31. V.1989, leg. M. Danilevsky (ZIN, MSPU);  1♂ — ibid, (cBK);  339 ex. — ibid, 29.04.1997, leg. A.A. Klimenko (cAK);  8♂♂ 3♀♀ — ibid, 3.05.1997, leg. S.A. Toropov (cST);  1♂ — ibid, 9.05.2000, leg. S. Nekrashevich (cDF);   10♂♂ 7♀♀ — E Kazakhstan, N  Zaysan Lake, 20.IX.1990, 27.V.1996, 25.V.1997, 25.V.2000, leg. A. Klimenko (cPU, MSPU) ;  1♂ —E Kazakhstan, N Zaysan, N part of Kuludzhanskie sands, 8–10. V.1999, leg. A. Korolyuk, I. Smelyanskiy;  1♀ —cape Krasnoyarsk,  Irtysh River, 22. VI.1925, leg. V.I.  Vereshchagin (both ISEA) ;   1♂ 1♀ —  Aigyrkum desert, 5 km W of Buran vill., 26.05.2003, leg. Zhdanko (cSS); WITHOUT EXACT LOCALITY :   1♂ —  Altai, foothills, VII.1979, ex. koll. G.A. (cISl) ;  2♂♂ —Altai; 1♂ 2♀♀ —Camp. Kirgiss. or.; 2♂♂ 3♀♀ —fl. Irtysh (all ZMUM);   1♀ —  Irtysh (ZIN) ;   1♂ —  Siberie (MNHN)  .</p><p>References. RUSSIA, Tyumen Oblast: Tobolsk (Mandl 1921); Omsk Oblast: N of vill. Malyi Atamas, bank of Irtysh River, 54.014266 N 74.941458 E, 30.V.2021,S.Knyazev(https://www.inaturalist.org/observations/81663853); Tomsk Oblast: Tomsk (Mandl 1921); Altai  Krai: Kolywan (Mandl 1921; Wu 1937); near Mikhaylovskoe; Laptev Log; near Uglovskoe (all Matalin &amp; Dudko 2016a); KAZAKHSTAN, Abai Region: Semipalatinsk (Mandl 1921); Beskaragay District, 51.042429 N 79.465608 E, 18.V.2022, A. Murtazina (https://www.inaturalist.org/ observations/117848591); East Kazakhstan Region: Ust-Bukon; 20 km S Kaznakovka, Bukonskie Peski (both Matalin &amp; Dudko 2016a); Novopokrovka (Matalin &amp; Dudko 2016b).</p><p>Diagnosis. The nominotypical subspecies is easily recognized among other ones by entirely lacking (Figs 1–5, 24–26, 28, 29, 31–35, 38, 39) or strongly reduced (Figs 27, 30, 36, 37, 40, 41) white elytral pattern (HI = 0.0–6.0, mean = 0.4). From other subspecies, except  C. coerulea nitida, the nominotypical subspecies is well distinguished by the long internal sack sharply rounded towards to the ventral side of aedeagus tube (Figs 65–70 vs. Figs 167–170, 190–195, 233–240).</p><p>Redescription. TL = 12.5–15.0 mm (mean = 14.06 mm, n = 41) in males, 13.7–16.0 mm (mean = 14.85 mm, n = 33) in females. Specimens as a rule unicoloured blue, blue-violet or black (Figs 1, 3, 4); sometimes bicolored with blue-violet head and pronotum and blue-green or green elytra, or with blue-green head and pronotum and cupreous or cupreous-green elytra with golden tinge in some specimens (Fig. 5).</p><p>Head often black, violet with bluish tinge or blue with light greenish lustre, rarely blue with bright golden-green sides of clypeus, central portion of frons, vertex and occiput, as well as supraorbital plates. Labrum with 4–14 (mean = 8, n = 78) submarginal setae, LW/LL = 2.0–2.5 (mean = 2.2, n = 41) in males, 1.92–2.2 (mean = 2.07, n = 33) in females (Figs 6–9). Mandibles relatively stout, ML/MW = 5.43–7.78 (mean 6.41, n = 36) in males and 6.0–8.5 (mean 7.3, n = 24) in females. Scape besides 3–5 apical setae with 1–3, rarely with 4–5 additional setae or glabrous.</p><p>Pronotum transverse, PW/PL = 1.28–1.46 (mean 1.35, n = 41) in males and 1.33–1.50 (mean = 1.41, n = 33) in females, black, violet or blue, in some specimens bluish-green with golden tinge on disk and bright blue anterior and posterior grooves and midline (Figs 14–18).</p><p>Elytra with smooth punctured sculpture except slightly granular humeral area, EL/EW = 1.45–1.63 (mean 1.55, n = 41) in males and in females 1.40–1.58 (mean 1.50, n = 33), as a rule without white pattern (Figs 1–5, 24–26, 28, 29, 31–35, 38, 39); sometimes with basal portion of humeral lunula and apical portion of apical lunula (Fig. 30), with narrow apical lunule separated into two small dots (Fig. 37), with apical portion of humeral lunule and subsutural portion of medial band (Fig. 41); or with humeral and apical lunula divided into two small spots and narrow subsutural portion of medial band (Figs 27, 36).</p><p>Aedeagus relatively long (Figs 42–44), EL/AL= 1.7–1.86 (mean 1.78, n = 19) with short, slightly curved basal portion and sharp, medium-sized apical hook. Internal sac long, sharply rounded towards the ventral side so В distinctly or virtually contact with the aedeagus tube, mt large with sharply curved apex, VA and DA the largest, DLL relatively large (Figs 65–70).</p><p>Distribution. RUSSIA: Chelyabinsk, Tyumen, Omsk, Novosibirsk and Tomsk Oblasts, Altai  Krai; KAZAKHSTAN: North Kazakhstan, Akmola, Pavlodar, Ulytau, Abai and East Kazakhstan Regions (Fig. 246).</p><p>Notes.  Cicindela coerulea was described by Pallas (1773) based on collections from his trip to Western Siberia in 1771. In the original description he indicated that the beetles are bright blue, but their colouration is variable from violet-black to completely black. Additionally, Pallas indicated that this species is abundant in different sandy habitats south of Irtin (= Irtysh River). Pallas’  Coleoptera collection is housed partly in ZIN and partly in MFNB. According to Horn and his co-authors (Horn et al. 1990) the portion of the collection of Pallas housed in St.- Petersburg was lost due to a fire. However, the only fire in the Kunstkamera occurred on the night of December 4–5, 1747 (Hartanovich &amp; Hartanovich 2014), and Pallas’s collection had not yet been deposited there by that time. There were no further fires that led to the destruction of collections neither in the Kunstkamera or in the Zoological Museum. Nevertheless, I could not locate a single specimen of  C. coerulea from the collection of Pallas either in ZIN or in MFNB. At the same time in 1781, the illustration of  C. coerulea was published (Pallas 1781: Tab. G fig. 12). According to the Article 73.1.4 of ICZN (1999) the designation of an illustration of a single specimen as a holotype is to be treated as designation of the specimen illustrated; the fact that the specimen no longer exists or cannot be traced does not invalidate the designation. However, Pallas had a series of these specimens and mentioned Article of ICZN that cannot be applied. I believe the types of  C. coerulea are lost and, according to the Article 75.3 of ICZN (1999). a neotype must be designated. Because “Irtin (= Irtysh River)” was indicated by Pallas (1773: 724) as the type area, a male specimen from the collection of ZIN labelled as “Irtys r.” (Figs 1, 2, 28, 42) is designated here as the neotype of  Cicindela coerulea Pallas, 1773 (see chapter Type material).</p><p>The elytral colouration in the nominotypical subspecies is highly variable from black through violet, blue or blue green to greenish-cupreous and bright reddish-cupreous (Figs 24–41). Reddish-cupreous beetles (three males) from the environs of Biysk (Russia, Altai  Krai) were described by Mandl (1972) as  Cicindela coerulea lucifera . However, similarly coloured specimens have been observed in the other parts of the range of the nominotypical subspecies, for example in Zhana-Arka, Kazakhstan (Figs 5, 38, 39). Furthermore, in the same habitat both reddish-cupreous and bluish-green (Figs 30, 35–37, 40, 41) as well as reddish-cupreous and violet-black (Figs 5, 26, 38, 39) specimens are observed. The shape of the aedeagus and the internal sack in these specimens are identical (Figs 65–70). Moreover, the main body proportions in different coloured specimens do not differ significantly, except the differences between males and females (Figs 71–75). Thus, a new synonymy is established:  Cicindela coerulea coerulea Pallas, 1773 =  Cicindela coerulea lucifera Mandl, 1970,  syn. nov.</p><p>During the current study, a male of  C. coerulea coerulea with the label “Issyk-Kul” was found in the collection of ZIN. Considering the great gap between the above-mentioned locality and the main distribution area of the nominotypical subspecies, old age of this specimen as well as the lack of information on the collector, date and exact locality, I consider this record to be mislabelled.</p></div>	https://treatment.plazi.org/id/038787EC032EFFC4CE84E5386D8DFD94	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Matalin, Andrey V.	Matalin, Andrey V. (2025): An intraspecific structure of Сicindela (s. str.) coerulea Pallas, 1773 (Coleoptera, Cicindelidae). Zootaxa 5583 (2): 201-246, DOI: 10.11646/zootaxa.5583.2.1, URL: https://doi.org/10.11646/zootaxa.5583.2.1
038787EC0321FFD9CE84E2B96E04FA68.text	038787EC0321FFD9CE84E2B96E04FA68.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cicindela coerulea Pallas 1773	<div><p>Cicindela (s. str.) coerulea nitida Lichtenstein, 1796</p><p>Figs 10–13, 19–23, 45–54, 76–82, 89–132</p><p>Cicindela nitida Lichtenstein, 1796: 32 (Type locality—Sibiria).</p><p>=  Cicindela tricolor M.F. Adams, 1817: 278 (Type locality—Sibiria ad lacum Baikal et ad ripas fl. Lenae, ultra urbem Jakutzk), synonymized by Horn (1915: 337).</p><p>=  Cicindela tricolor var. obliquefasciata Fischer von Waldheim, 1828: 21, Tab. 1 fig. 3 (Type locality—Sibiriae), synonymized by Roeschke (in Horn &amp; Roeschke 1891: 49).</p><p>=  Cicindela tricolor var. tenuifascia Fischer von Waldheim, 1828: 22, Tab. 1 fig. 6 (Type locality— Altai), synonymized by Roeschke (in Horn &amp; Roeschke 1891: 49).</p><p>=  Cicindela optata Fischer von Waldheim, 1828: 22, Tab. 1 fig. 8 (Type locality—Dauria, ad litora fluvii Amur), synonymized by Motschoulski (1844: 30).</p><p>=  Cicindela tricolor var. saltatoria Gistl, 1837: 38 (Type locality—Sibiria ad lacum Baikal et ad ripas fl. Lenae, ultra urbem Jakutzk, et in montosis Altaï), synonymized by Horn (1915: 337).</p><p>=  Cicindela coerulea selengensis Mandl, 1936: 12 (Type locality—Selengatal, Selengafe), synonymized by Schilder (1953: 564).</p><p>Cicindela tricolor M.F. Adams, 1817: Fischer von Waldheim 1820: 6, Tab. 1 fig. 3, Tab. 17 fig. 3; 1821: 102; 1828: 12, 21; 1829: 5; Dejean 1833: 3; 1836: 3; Dejean &amp; Boisduval 1837: 24 Pl. 3 fig. 3; Sturm 1843: 2; Motschoulski, 1844: 30; Motschulsky, 1850: 1; Chaudoir 1865: 36; Sahlberg 1880: 5; Fleutiaux 1892: 106; Horn &amp; Roeschke 1891: 48, Taf. V fig. 11; Tschitschérine, 1903: 94; Jakobson 1905: Tab. 1 fig. 3; Horn 1905a: 157, 158; 1905b: 27; Poppius 1906: 14; Lafer 1978: 5, figs 5, 14; 1989: 96, fig. 52.4, 97.</p><p>Cicindela hybrida tricolor M.F. Adams, 1817: Jakobson 1905: 190; Horn 1905a: 157, 158.</p><p>Cicindela optata Fischer von Waldheim, 1828: Motschulsky, 1850: 1.</p><p>Cicindela nitida Lichtenstein, 1796: Lutshnik 1916: 95; Mandl 1921: 79; Shilenkov 1994: 8; Kryzhanovskij et al. 1995: 26; Berlov &amp; Berlov 1997: 44; Sundukov 1998: 9; Matalin 2000: 676, fig. 35; Rogatnykh 2006: 30; Rogatnykh &amp; Yakubovich 2009: 108.</p><p>Cicindela hybrida nitida Lichtenstein, 1796: Mandl 1921: 80, 81; Winkler 1924: 4; Horn 1915: 337; 1926: 221; 1930: 402; Wu 1937: 15; Institute of Zoology Academia Sinica 1986: 417, fig. 11-1-3; Hu 1992: 374; Zhu et al. 1999: 26, 27 Tab. 4 fig. 13; Fang &amp; Wu 2001: 69.</p><p>Cicindela grumi nitida Lichtenstein, 1796: Schilder 1953: 564.</p><p>Cicindela nitida tricolor M.F. Adams, 1817: Shilenkov 1987: 7.</p><p>Cicindela nitida optata Fischer von Waldheim, 1828: Berlov &amp; Berlov 1997: 44.</p><p>Cicindela nitida selengensis Mandl, 1936: Kryzhanovskij et al. 1995: 27; Matalin 2000: 676, fig. 34.</p><p>Cicindela coerulea Pallas, 1773: Rogatnykh 2009a: 9; 2009b: 69; Sundukov 2009: 89; Khobrakova 2012: 167; Putchkov 2013: 54 fig. 2.4, 62, 63, 65 fig. 4.1, 68 fig. 7.4, 73 fig. 12.5–6, 80.</p><p>Cicindela coerulea nitida Lichtenstein, 1796: Lutshnik 1924: 21; Okninskyi 1930: 20; Mandl, 1935: 285 fig 1b, 294 figs 1–5, 296 figs 1–5; 1936: 11; 1965: 241; 1974: 137; Horn 1938: 46, Taf. 63 figs 23–25; Nakane 1960: 30, 38 Pl. I fig. d; Mandl 1970: 73 Abb. 3 fig. 6, 74; Alekseeva 1977: 5; Cassola &amp; van Nidek 1984: 10; Werner, 1991: 18, T. 14 figs 102, 104–106, T. 15 figs 107–109; Li 1992: 5; Wiesner 1992: 117; 2020: 183; Putchkov &amp; Shilenkov 1996: 451; Lorenz 1998: 42; 2005: 131; Puchkov &amp; Matalin 2003: 104; Khobrakova &amp; Sharova 2004: 144; Shook &amp; Wiesner 2006: 10; Khobrakova &amp; Dudko 2010: 37; Wu &amp; Shook 2010: 78; Wu 2011: 24; Budilov 2012: 40; Sundukov 2013: 76; Khobrakova et al. 2014: 70; Koshkin et al. 2016: 312; Matalin &amp; Dudko 2016a: 140; 2016b: 143; Tsuji et al. 2016: 3; Putchkov &amp; Matalin 2017: 227; Matalin et al. 2019: 10; National Species List of Korea 2019: 199; Makarov et al. 2020; Putchkov 2020: 169 fig. 45.1, 173 fig. 49.4, 182 fig. 56.5, 199 fig. 61.1; Shilenkov 2021: 41.</p><p>Cicindela coerulea selengensis Mandl, 1936: Mandl, 1935: 294 figs 6–8, 296 figs 6–10, 1936: 12; Horn 1938: 46, Taf. 63 fig. 26; Cassola &amp; van Nidek 1984: 10; Werner, 1991: 18, T. 15 figs 112, 113; Wiesner 1992: 118; 2020: 184; Lorenz 1998: 43; 2005: 131; Puchkov &amp; Matalin 2003: 104; Putchkov &amp; Matalin 2017: 227.</p><p>Type material.   Lectotype of  Cicindela optata Fischer von Waldheim, 1828 (designated here), ♂ (Figs 45, 76, 77)—  C. optata Fisch., Nertshinsk [handwritten white label], Typus [typed pink label], Fischer von Waldheim [typed white label], Staatl. Museum für Tierkunde Dresden [typed white label], Lectotypus,  Cicindela (s. str.) nitida f.  optata F.-W., design. A. V. Matalin, 1997 [handwritten and typed red label]; LECTOTYPUS,  Cicindela optata Fischer von Waldheim, 1828, design. A. V. Matalin, 2024 [printed red label] (MTD).   Paralectotype of  Cicindela optata Fischer von Waldheim, 1828 (designated here), ♂ —Staatl. Museum für Tierkunde Dresden [typed white label], Paralectotypus,  Cicindela (s. str.) nitida f.  optata F.-W., design. A. V. Matalin, 1997 [handwritten and typed red label], PARALECTOTYPUS,  Cicindela optata Fischer von Waldheim, 1828, design. A. V. Matalin, 2024 [printed red label] (MTD).</p><p>Paratypes of  Cicindela coerulea selengensis Mandl: 1♂ (Fig. 113)— Transbaic.,  Selenga fl. [handwritten white label], Сollect.  Hauser [typed white label],  C. tricolor var. [handwritten white label],  v. optata, det. Ganglbauer [typed and handwritten white label],  Cicindela nitida Licht
.
 s. selengensis m., det. Ing. Mandl [typed and handwritten white label], PARATYPUS (selengensis  Mandl) [typed and handwritten red label];  Cicindela
(s. str.) coerulea (Pallas)
 selengensis  Mandl 1936,  Det. J. Probst [printed white label], NHMW-ZOO-COL-0002631 [printed white label];   1♀ —Transbaic., Selenga fl. [handwritten white label], Сollect. Hauser [typed white label],  v. optata, det. Ganglbauer [typed and handwritten white label], PARATYPUS (selengensis Mandl) [typed and handwritten red label], NHMW-ZOO-COL-0002632 [printed white label];  1♀ (Fig. 114)— ibid, NHMW-ZOO-COL-0002633 [printed white label];  1♀ (Fig. 112)—Transbaicalia, Selenga Tal., V – VI.1908 [typed white label], Сollect. Hauser [typed white label],  v. optata, det. Ganglbauer [typed and handwritten white label], PARATYPUS (selengensis Mandl) [typed and handwritten red label], NHMW-ZOO-COL-0002634 [printed white label];   1♀ —Transbaic., Selenga fl. [handwritten white label], Сollect. Hauser” [typed white label],  v. optata, det. Ganglbauer [typed and handwritten white label],  Cicindela nitida Licht. s. selengensis m., det. Ing. Mandl [typed and handwritten white label], PARATYPUS (selengensis Mandl) [typed and handwritten red label], NHMW-ZOO-COL-0002635 [printed white label];   1♂ (Figs 80, 81)— ibid, NHMW-ZOO-COL-0002638 [printed white label]; 1♀ —Cotype [typed red label], Selenga-Tal, Transbaikalien [handwritten white label],  Cicindela nitida Licht. ssp. selengensis m., det. Ing. Mandl [typed and handwritten white label], NHMW-Acquisitions-Nr.: Col-1971-6, coll. E. Grundmann [printed white label], NHMW-ZOO-COL-0002639 [printed white label]  .</p><p>Additional material. RUSSIA, Altai  Republic: 1♂ —Chuysky trakt, Ust-Inya, 15.VII.1907, E.G. Rodd.; 1♀ — Naryn-Gol River, 1956, Pridanieva;  1♀ — Altai, env. Ongudai, leg. A.G. Yakobson (all ZIN); 3♀♀ — ibid, 2– 3. VI.1989, S. Saluk; 1♀ —Chemal, 30.VII.[19]63; 1♂ —mouth of Chulcha River, right tributary of Chulyshman River, 29.VIII.[19]34, V. Geptner; 1♂ —mouth of Malyi Karakem River, tributary of Chulyshman River, 25. VI.1989, leg. V.N. Dremin (all ZMUM); 1♂ — Altai, Seminskiy pass, VII.1979; 2♂♂ 2♀♀ —valley of Chuya River, ZhanaAul, 1850 m, 8.VII.2022, leg. A. et E. Filippov (both MSPU); 3♂♂ 1♀ —Altaigeb., Akash, Mittellauf des Mjonka, Kuraikette, 1600–2300 m, 12.6.1992 (cJW); 2 ex. — Altai, Akash, Mjonka-Fluss, 20.05.1993, leg. A. Budarin; 9 ex. — ibid, 14.06.1991, leg. O. Legezin (both cJG); 1♀ —SE Altai, Kurai, 1.VIII.[1]964, Kerzhner; 1♂ 1♀ —Buguzun River, Chikhacheva Mnt Ridge, ~1900 m, 29.VII.[1]964, Grunin; 1♂ 1♀ —Chagan-Uzun env., bank of Chuya River, 5–6. VI.1989, V. Grichik et S. Saluk;  1♀ — ibid, A.S. Pisanenko (all ZIN);  Republic of Khakassia:  2♂♂ 2♀♀ —Shira Lake, 5.VIII.1916, Khvorov; (ZMUM); 2♂♂ — Tomsk Gouv., Fluss Abakan, Ui-Bat River, 17.VII., A. Jakobson; 1♂ 1♀ — Tomsk Gouv., Tashtyb, 16. VI., A. Jakobson (both ZIN); 1 ex. —Abakan, Krasnoe Lake, 26.07.1993, leg. A. V. Chuvilin (cFB); 2♂♂ 1♀ —Beisk District, near Tabat vill., 19.VIII.[19]63, O. Grigorjev (ISEA); 3♂♂ 3♀♀ —2–3 km S of Abakan, 5.VII.1997, leg. V. Patrikeev; 1♂ —valley of Son River, 2. VI.1969, leg. V. Patrusheva (both MSPU); 3♀♀ —Borets, bank of Son River, 54.46 oN 90.36 oE, h~380 m, 29.06.2011, leg. K. Tomkovich (cOM);  Republic of Tuva: 6♂♂ 9♀♀ —Erzin, sands on the shore of Tere-Khol, 7. VI.1962, leg. D. Berman (ZMUM);  1 ex. — Tuva, Erzin, Erzin-River, 30.05.1990, leg. S. Gundorov (cJG); 2♀♀ —17 km NE of Khadyn Lake, 26. V.1988, leg. V. Zinchenko; 1♀ —Balgazyn, 8. VI.1948, Cherepanov (both ISEA); 1♂ — ibid, 28.VIII.1959, leg. S. V. Sharova (cISl); 1♀ —Balgazyn, 110 km SE of Kyzyl, VII.1956, Galkin; 6♂♂ —Toore-Khem, 11.08.1973, leg. B. Korotyaev; 1♂ —near Erzin, 21.07.1988, leg. S. Byzov (all ZIN); 1♂ 3♀♀ —SE shore of ToreKhol Lake, 31. V –1. VI; 1♂ —Kyzyl District, bank of Biy-Khem River, near Kara-Khaak, 20. V.1997, leg. V. Patrikeev; 2♂♂ 3♀♀ —Shagonar, 26.07.1992, A. Napolov; 2♂♂ —near Kyzyl, bank of Enisey River, 1–6.VII.1997, leg. S. Vastchenko &amp; R. Mishustin; 6♂♂ —Khadyn Lake, 16–19. VI.1997, leg. S. Vastchenko &amp; R. Mishustin; (all MSPU); 1♂ — ibid, 25. V.1998, leg. A. Gorodinski (cXX); 1 ex. — ibid, 20.05.1998, leg. S. Vastchenko (cFB);  1♂ 1♀ — ibid, 25. VI.1998; 2♂♂ —30 km N Balgazyn, 21.6.1994, leg. P. Smrz, Z. Kletecka (both cJW); Krasnoyarsky  Krai: 1♂ —Bazaikha near of Krasnoyarsk, 14. V.[19]01, Krutovsky;  1♂ — ibid, 9. VI.[19]01; 19♂♂ 20♀♀ —Kaptyrevo; 10♂♂ — Krasnoyarsk (all ZIN); 1♂ 1♀ —O Sibiria, Berezovka, leg. Dr. J. Fleischer (cJW); 3♂ 5♀ —Sibiria, Minusinsk, Exp. Zichy, leg. Csiki (HNHM); 1♂ —Minusinsk, Opytnoe Pole, 18. V.1928; 5♂♂ 4♀♀ — ibid, 25– 31.VII.1928 (both ISEA); 1♂ —Sibiria, Krasnoyarsk, A. Zolotarev; 3♂♂ 3♀♀ — ibid, 1899, L.A. Molchanov; 2♀♀ — ibid, VI.1902; 1♂ — ibid, 19. VI.1902; 1♂ 3♀♀ — ibid, VII.1902; 1♀ — ibid, VIII.1902; 1♂ 1♀ —Vostochnoe, 35 km S von Minusinsk, 1928, A. Zhelokhovtsev (all ZMUM); Irkutsk Oblast: 1♀ —Irkutszk, 3.VII.1967, leg. G. Rácz (HNHM); 2 ex. — Irkutsk, 5 km NW of Cap Krestowski, 10 km S of Jelantsy, 22.06.1993, leg. R. Krause, O. Jäger; 4 ex. —Ust Orda, Kuda River, 26.06.2011, leg. W.-H. Liebig; 1 ex. —Nischnij Udinsk, Ude River valley, 28.06.2003, leg. A. Azarov; 1 ex. —Porog, Uda River valley, Privolnoye, 28.06.2003, leg. A. Azarov (all cJG); 2 ex. —Khatar-Khaday, 100 km NE of Irkutsk, 03.08.2000, leg. L. Lehmann (cAP); 1♂ —Baikal Lake, Slyudyanka dist., Kultuk vill., 12.6.2007, leg. S. Rybalkin; 1♀ —Baikal, Olkhon Peninsular, VIII.1973, Medvedev (both cJW); 1♂ — ibid, 15.08.1979, V. Shilenkov (SFU); 1♂ —downstream of Oka River, 23.VII.1999, leg. K. Gongalsky (MSPU); 1♂ —Malyshevka opposite Balagansk, bank of Angara River, 20.VII.[18]98, Ingenitsky; 1♂ —Shivera on Angara, 20.VIII.[18]98, Ingenitsky; 1♂ 2♀♀ —Malta (all ZIN); 2 ex. —Kodat, Mt. Togo, 2500 m, 07.07.2011 (cFB); 86 ex. —Kachug distr., bank of Lena River, h=501 m, 53.99141 N 105.72398 E, 05–15.06.2022, leg. Yu. &amp; N. Seregin (cVS);  Republic of Buryatia: 1♀ —Kyakhta (MNHN);  1♀ — ibid, 05.07.1971; 1♂ 1♀ —near Kyakhta, Deben vill., 11.VIII.1977, leg. V. Shilenkov; 2♂♂ —Tunkinskie Goltsy, near Mondy, h~1599 m, 6–10. VI.1992; 1♂ —Mondy, valley of Irkut River, 20–30.VII.1977, leg. V. Shilenkov (all MSPU); 14 ex. — ibid, 06.2012, leg J. Gebert (cJG); 4 ex. — ibid, 06.06.2006, leg. E. Gavrischuk &amp; O. Pekarsky; 4 ex. — ibid, 01.06.2011, leg. A. &amp; E. Filippov (both cFB); 1♂ —Mondy, Sajan Geb.; 1♂ 2♀♀ — Burjatia, Sajany, Mondy, 1200 m, 15.7.1993; 3♂♂ 1♀ — ibid, 1.8.2002, leg. I. Legezin (all cJW); 1♂ —middle current of Oka River, Arkhabom, 19–21.VIII.1999, leg. K. Gongalsky; 4♂♂ 2♀ ♀ —between Khoronkhoi and Naushki, 50 o 25’38’’N 108 o 08’18’’E, 13–23. V.1999, leg. I. V. Melnik; 2♂♂ 1♀ —2 km E of Beloozersk, 50 o 38’30’’N 105 o 44’18’’E, 1–7.VII.1999, leg. I. V. Melnik; 1♂ 1♀ —5 km NE of Kiran, 50 o 21’43’’N 106 o 46’11’’E, 25–30. VI.1999, leg. I. V. Melnik; 2♂♂ —valley of Dzhida River, near Zakamensk, 19.06.2000, leg. P.Ya. Ustuzhanin; 2♂♂ 1♀ —Tsakir, 7.VII.2004, leg. K. Eskov; 1♂ 4♀♀ —Novaya Komushka, 51 o 47’N 107 o 38’E, 8.08.2015, S. Rudykh; 1♀ —Tunka River, tributary of Irkut River, 22. VI.1914 (all MSPU); 1♂ 2♀♀ —Ust-Kiran, 14.6.1956; 1♀ — ibid, 24.6.1951, N. Filippov; 2♂♂ — ibid; 1♂ —Tory, 17. VI.1914; 1♂ 2♀♀ —Mondy, 13–16.VII.1947, Tsvetaev (all ZMUM); 3♂♂ — ibid, 4–19. VI.1915, S. Rodionoff; 2♂♂ 1♀ — ibid, h=1600–1700 m, 7–8.06.2002, Lvovsky; 1♂ —Ust-Kiran, 20. V.[1]913; 1♂ — ibid, 30. VI.[1]913; 1♂ — ibid, 11.VII.[1]913; 1♂ 2♀♀ —Kyakhta, VII–VIII; 2♀♀ —Olkhon Island, near Ulan-Khushin, 11.VII.1973, N. Ermakova; 1♂ —Verkhneudinsk (= Ulan-Ude), 18.VII.1899, G. Suvorov (all ZIN); 1♂ 1♀ — Transbaikal, Werchne-Udinsk (= Ulan-Ude) (cJW); 3 ex. — ibid, 01.06.2014, leg. A. Filippov; 1 ex. — ibid, 12.06.2014; 1 ex. — ibid, 13.06.2015; 3 ex. —Zaigraevo, Brjanka River valley, 15.06.1997, leg. A. Petrov (all cFB); 1♂ 1♀ —Vjerchne-Ud.; 1♂ —Vjerhnje Ud., u Kulicha Bäd.; 2♀♀ —Kul. Bad. (all ZMUM); 1♂ 1♀ —Transbaikal, Werchne-Udinsk (= Ulan-Ude), leg. Dr. K. Mandl; 1♂ —Mondy, Irkut superior, 900 m, 21.VII.1973, leg. Shilenkov (both HNHM); 2♂♂ 2♀♀ —near Mondy, sandy bank of Irkut River, 09.07.1969; 1♂ —Vitim Plateau, vill. Baisa, 16.VII.1978, leg. T. Zolotarenko; 1♂ 2♀♀ — Dureny, bank of Chikoi River, 1–2.VIII.1986, leg. B. Zakharov et Yu. Chernov (all ISEA);  Zabaykalsky  Krai: 1♀ — Transbaic., Čita, 1910;  6♂♂ 1♀ —Transbaic., Tshita (both ZMUM); 2♂♂ — ibid (MSPU); 1♀ — ibid, Hermann Frieb (cJW); 1♂ — ibid, coll. Jul. Isaac (HNHM); 3♂♂ — ibid, V.1916, Yurkevich; 1♂ — ibid, VII.[1]912, Pisarevsky; 2♂♂ 3♀♀ — ibid, VI –VII.[1]910, I. Zadoplyaev (all ZIN); 1♂ — ibid, 10.07.1929; 2♂♂ —Chita env., 17.07.1959, N. Filippov (both MSPU); 1 ex. —Chita (Tshita), Ingoda-Tal Steppenhang, 14.06.1992, leg. Kostjuk; 1 ex. — ibid, 13.06.1994, leg. R. Krause, O. Jäger (both cJG); 1♀ —Usugli, 10.VII.1958, M. Lurie; 1♂ —Molokovka, P. V. Pisarev; 2♂♂ 1♀ —East Siberia, Sretensk; 1♂ —Sretensk District, Dunaevo env., pad Balagannaya, 17.IX.1957, G. Dlussky; 1♀ —Ust-Dunaevo, 13.IX.1957, G. Dlussky; 2♀♀ —Burgen, bank of Chitinka River, 10.VII.[1]912; 3♂♂ —Amazar, 25. V.1913; 4♂♂ 3♀♀ —near Nerchinsk, V. Podgorbinsky (all ZMUM); 1♂ —Nertschinsk; 1♂ —Bolshie Boty; 1♂ — Ingoda River, pad Argalei, 4. VI.1899, G. Suvorov; 2♂♂ —Aga [= Khyla] (all ZIN); 1♀ —Kalar District, Chara, VI.1975 (ISEA); 2♂♂ —Drovyanaya, 2.VII.1977 (cEK; cAKr); 3♂♂ —Sokhondo Nature Reserve, cordon Agutsa, 16.06.1991, leg. V. Pekin, D. Logunov (ISEA); 1♀ —Chita Region, near Borsya, 08.1995, leg. A. Sedov (cEK); 3♂♂ —Nertschinsk (ZMUM); 1♀ —Transbaikal., Nerchinsk, 16.VII.1915; 1♂ —Abagatui, 17.08.1992, leg. M. Chernyakhovsky; 1♀ —Chita env., 15. VI.1983, leg. G. Nesterov; 1♀ —shore of Tsagan-Nur Lake, 4.VII.2000, leg. I. V. Melnik; 6♂♂ 3♀♀ —5 km W of Nizhny Tsasuchei, floorplan of Onon River, on sand with sparse vegetation, 14–18. VI.1995, leg. D. Fedorenko (all MSPU); 2♀♀ —Chita reg., Onon Riv., Tsasuchei vill., 3. VI.2002, leg. A. Anischenko (cJW);  Republic of  Yakutia: 2♂♂ 2♀♀ —near Yakutsk, 11. VI.–9.VII.1962, Zhelokhovtsev;  1♂ — ibid, 15.06.81, I. Medvedev (both ZMUM); 1♂ — ibid, 28. V.927, Nikiforov (ZIN); 1♂ — ibid, 01. VI.1921, A. Tsvetaev (cEK); 2♂♂ 1♀ —Yakutsk, [1]901, Olenin; 1♂ — ibid, 9. VI.9127, Moskvin; 2♀♀ — ibid, 1883, Herz (all ZIN); 1 ex. — ibid, 01.07.1987, leg Z. Mracek; 2 ex. —Wladimirowka, 18.06.1989, leg. Subota (both cJG); 1♂ —Yakutsk, Botanical Garden, 24. V.1985, leg. V. V. Dubatolov; 4♂♂ — ibid, 18.VIII.1985 (both ISEA); 2♂♂ —40 verst from Yakutsk, Ytyk-Khaya, 9. VI.[19]01, Herz; 2♂♂ 2♀♀ —Ytyk-Khaya, Lena mer; 1♂ —islands on Lena River, near Ytyk-Khaya, 20–21.VII.[19]01, Herz; 1♀ —Dupsya, 8.VII.911, I. Afanasiev; 2♂♂ —Sergeleekh, 3 versty from Yakutsk, 15.VIII.911, I. Afanasiev; 1♂ —between Lena River and Verkhoyansky Mnt Ridge, [18]91, Čerski; 3♂♂ 2♀♀ —Olekminsk, 22. V.–2. VI.[19]08, Buturlin; 1♂ —near Olekminsk, 6. VI.[19]03, Kharitonov; 2♂♂ — ibid, 17. VI.[19]10 (all ZIN); 2♂♂ —right bank of Lena River, Nizhny Bestyakh, 2–17.VII.2001, S. Kurbatov; 3♀♀ — Matta River, ~ 20 km from the confluence with the Sinyaya River, 12.07.2017, leg. A. Ignatenko (both MSPU); 3 ex. —Oimyakon,Tomtor vill., leg. V.Zaritsky (cFB); Amur Oblast: 3♂♂ —30km W of Svobodnyi, near Kostyukovka, 5–9.VII.1995, leg. D. Fedorenko; 1♂ 1♀ —12 km S of Svobodnyi, Malaya Sezemka, bank of Zeya River, on heavily silted sand, 5.VII.1995, leg. D. Fedorenko; 2♂♂ —Innokentievka, 23.VII.1969, leg. Yu. Tretjakov (all MSPU); 1♀ —Kasatkino, 2.IX.1972, V. Zherikhin; 1♂ 1♀ —Arkhara, 4.IX.1972, leg. V. Zherikhin (both ZMUM); 2♂♂ 1♀ —Blagoveshchensk District, cf 15 km W of Natalino, forest, 27–31.VII.2003, leg. B. Kataev (ZIN); 1♂ —near Blagoveshchensk, 14–17.07.1996, leg. V. Bezborodov; 3♂♂ —Zeya District, Gilyui River, 4–6.VIII.1971, L. Morozova (both ISEA); 1♂ —N of railway station Tynda, sand and pebble bank of Lena River, 18.06.1997, leg. R. Dudko, D. Lomakin (MSPU); 1♂ 2♀♀ —Khingan Natural Reserve, cordon Erakta, 9.08.1989, leg. Yu. Tretjakov (cEK);  Khabarovsky  Krai: 1♀ —40 km SE of Mogdy, upstream of Amgun River, 20.07.1997, A. Brinev (MSPU);  2♂♂ — ibid (SFU); 1♀ —near Dzhamku, bank of Amgun River, 20–22.06.1996, leg. A.A. Kuzmin (ISEA); Jewish Autonomous Oblast: 2♂♂ 2♀♀ —near railway station Bira, 14.VII–2.VIII.1994, leg. P. Udovichenko (MSPU); 1♂ 1♀ —below Radde vill., Dichun River, 9.VIII.1980, leg. S. Alekseev (cSA);  Primorsky  Krai: 1♂ —Sidemi (= Narva River), 26. VI.1900, G. Suvorov;  1♀ —lower reaches of Tetyukhe (= Rudnaya) River, 20.VIII.[19]09, Dyukin; 3♂♂ — Vladimir Monomakh (= Monomakhovo), 3.VIII.–2.IX.[19]09, Dyukin; 3♂♂ 1♀ —Sikhote-Alin, 35 km NE of Sergeevka, 1.IX.1986, leg. B. Kataev; 3♂♂ 2♀♀ —Khanka Lake, Troitskoe, 4. V.–27. VI.[19]09, Čersky; 1♀ — Iman (= Dalnerechensk), VI.[18]99, Veselovich (all ZIN); 1♂ 2♀♀ —Khanka Lake, Novokachalinsk, 10. VI.1972, A. Ponomarenko; 1♀ —Ussuri River, Osinovka, 11. VI.[1]917, P. Elsky; 1♀ —fl. Sujfun (= Razdolnaya), Nik.- Ussurijsk, 23. V.; 1♀ —15 km SW Valentin, Glazkovka, 7.VII.1986, A. Antropov; 1♀ —Sudzukhinsky (=Lazovsky) Natural Reserve, Tyapigou (= Pribrezhnaya) River, 24.VII.[19]46, D. Panfilov; 1♂ —cove Kit, 11.VII.[19]46 (all ZMUM); 1♂ 1♀ — SU, Ussuri Region, Archipovka, 17.07.1989, leg. S. Becvar (cJW); 1♂ —Khasan District, Nature Reserve Kedrovaya Pad, 21.VIII.1964, D. Usachev (ZMUM); 1♀ — ibid, 14.06.1968, G. Zolotarenko (ISEA); 1♀ — ibid, 06.06.1962 (MSPU); 1♂ —Gamova Peninsula, pass between cove Vityaz and cove Telyakovskogo, 16.VII.1979, V. V. Dubatolov; 1♂ — Gamova Peninsula, cove Astafieva, 21.VII.1993, V.K. Zinchenko 1♂ —Khasan District, Golubiny Utes, 12. VI.1968, V. Nikolaev (all ISEA); 1♀ — ibid, 22. V.2022, leg. K. Makarov &amp; A. Matalin; 1♂ 2♀♀ —Ussurijsk District, Gornotaezhnaya Stantsiya, 20. VI.1960; 1♂ —Lazovsky Natural Reserve, cordon America, 43 o 16’16’’N 133 o 03’01’’E, 19.VII.2006, leg. K. Makarov; 1♀ —Vladivostok env., VI.1989 (all MSPU); 1♂ — Slavyanka env., VI.1993, E. Zhuravlev (cAKr); 2 ex. —Primorje, 14 km SSW of Krounovka, Krounovka-River, 11.08.1994, leg. P.-H. Schnitter, E. Grill (cJG); 1 ex. —Novonezhino, Venedivnovo, Tschernye Gory, 03.08.1990, leg A. Pütz (cAP); MONGOLIA, Uvs Aimag: 1♂ —Sandgebiet Altanels, 35 km WNW von Somon Tes, 1400 m, 3. VI.1968, leg. Dr. Z. Kaszab (HNHM); 1♀ —Üüreg Lake, 14.08.1970, leg. B. Korotyaev (ZIN); 1 ex. —Burgag Nuur (= Shaazgay Nuur), 03.07.2016, leg. P.-H. Schnitter (cJG); Khovd Aimag: 3 ex. —Baruun Chara, 5 km, Halbwüste, 29.07.1999, leg. A. Stubbe (cPS); Zavkhan Aimag: 1♂ 1♀ —Dzabyn-Gutu-Khudzhirt, 18–21.VII.1877, G. Potanin (ZIN); 1♀ —Choit chunch, 26 km ONO von See Telmen nuur, 2150 m, 13.VII.1968, leg. Dr. Z. Kaszab; 1♀ —Santmargaz sum, Bajan (= Chara) nuur, 18.VIII.1963, leg. A. Bold (both HNHM); 1 ex. —W Somon, Aldarchaan, 26.07.2010, leg. A. &amp; M. Stubbe; 3 ex. —Zawchanmandal, 09.06.2012, leg. J. Müller, A.J. Müller (both cJG); Khövsgöl Aimag: 1♀ —25 km NNE of Ulaan-Uul, 15–16.VII.[1]975, Gurieva; 1♀ —10 km SE of Muren, 23–24.VII.[1]975, Sugonyaev; 1♂ 1♀ —25 km W of Muren, 27. VI.[1]968, Arnoldi (all ZIN); 1 ex. —Khankh, 21.07.1995, leg. E. Novikov (cFB); Bulgan Aimag: 4♂♂ 1♀ —64 km W of Erdenecant, 1260 m, 47°05’N 104°05’E, 3–24. V.1990, leg. Gy. Fábián, M. Hreblay, L. Peregovits &amp; G. Ronkay; 1♀ —11 km W von Somon, Bajannur am see Bajan nuur, 1000 m, 14. VI.1968, leg. Dr. Z. Kaszab; 5♂♂ 5♀♀ — ibid, 1000–1100 m, 24.VII.1968 (all HNHM); 1♂ 2♀♀ — ibid, 24.VIII.1968 (NHMW); 2♀♀ —180 km SW of Ulashivet, Tarn (Lun) River, 47°19.380’N 103°41.751’E, 21.VII.2019, leg. H. Mühle (cJW); 1 ex. —Changai Mts., Orchon Brücke—Ulan Bataar lager steppe, 27.07.1999, leg. A. Stubbe (cPS); Arkhangaj Aimag: 1♀ —30 km E of Tsetserleg, 2–4.VII.[1]975, Narchuk; 1♀ —48 km S of Tevshruulekh, 2–3.VII.[1]975, Gurieva; 1♀ —Tevshruulekh, 13.VIII.1974 (all ZIN); 1♀ — ibid, 24.VIII.1971 (ZMUM); 1♂ — ibid, 16.7.1975 (cJW); 1♀ —63 km O von Urdtamir, Chaalgsin chundi bei Somon Tövschruulech, 1500 m, 28. VI.1966, leg. Zanzan Tombo; 5♂♂ 2♀♀ —NO of Ecke des Sees Ogij Nuur, 1350 m, 2.VII.1964, leg. Dr. Z. Kaszab (both HNHM) 1♂ — ibid, (NHMW); 1♂ — ibid (cJW); 2♂♂ 1♀ — Nord Mongolia, Changai, Leder (ZIN); 1♂ 4♀♀ — ibid (HNHM); 3♂♂ 3♀♀ — ibid (MNHN); Selenge Aimag: 3♂♂ 5♀♀ —Ero-Gol River near Dulaankhaan, 3–4.VIII.[1]975, Gurieva; 1♂ 1♀ — ibid, 30.VIII.[1]975; 1♀ —Shaamar, summer 1984, S. Kuzmin; 2♂♂ —coupling of Suman River and Chuluut River, 29–30. VI.[1]975, Narchuk (all ZIN); 1 ex. —Eröo Somon, Eröö Gol, Hochsteppe, 26.07.1999, leg.A. Stubbe (cPS); 1♀ —Sumber Dist., 5 km S of Baruun Kharaa, 48°51.179’N 106°05.006’E, 792 m, vegetated sand plain, 7. V.2018, leg. J.A. Shetterly; 1♀ —Khongor Dist., 49°22.417’N 105°52.304’E, 668 m, vegetated low sandy dunes, leg. J.A. Shetterly (both cJW); 1 ex. —Ochron-Aue Schotterufer, Suchbaatar, 05.06.2001, leg. Stenzel; 1 ex. —Orchon-Aue, Caragana-Steppe, 06.06.2002, leg. Stenzel (cJG);  4♂♂ 2♀♀ —Transbaic., Selenga fl., Collect. Hauser,  hybrida tricolor det. Ganglbauer;  1♂ —Transbaicalia, Selenga Tal, V – VI, 1, Collect. Hauser (both NHMW); 2♂♂ 2♀♀ —Mongolie sept., Selenga, J. Chaffanjon, 174-96 (MNHN); Darkhan Uul: 1♀ —20 km NNE of Darkhan, 5.VIII.[1]975, Gurieva; 1♂ —25 km E of Darkhan, 30.VII–2. VIII.[1]975, Gurieva; 1♀ —Khara River, tributary of Selenga River, 1. VI.[18]97, Klements; 1♀ —Sharyn River, tributary of Selenga River, 31. V.[18]97, Klements (all ZIN); Ulaanbaatar: 1♂ 2♀♀ —Urga, 13.08.1913, Schultz (HNHM); 9♂♂ —Nalaikh, 40 verst E of Urga, 1–2.VIII.[18]97, Klementz; 1♂ — ibid, 25. VI.[1]967, Zaitsev (both ZIN); Töv (= Central) Aimag: 2♂♂ —Ulan Bator, Nucht im Bogdo uul, 1800–2000 m, 27.VII.1966, leg. Dr. Z. Kaszab; 1♂ —Tel des Tola, zwischen Somon Altanbulag und Somon Tariat, 30 km ONO von Tariat, 1200 m, 25.VII.1966, leg. Dr. Z. Kaszab (both HNHM); 1 ex. —170 km W of Ulaanbataar, 16.08.2007, leg. P. Tyrner; 2 ex. —Terelz, Tuur, 21.07.2003, leg. J. Halada (both cJG); 4♂♂ 2♀♀ —45 km E of Lune, 4.IX.67, Zaitsev; 2♂♂ 4♀♀ —70 km SW of Ulan Bator, sandy steppe, 30.VII.[1]963, Arnoldi, Gurieva; Kerzhner, Zaitsev; 1♀ — ibid, 125 km SW of Ulan Bator, 30–31.VII.[1]963, Kerzhner (ZIN); 3 ex. —SW of Altanbulag, Tola-Aue, 08.08.2009, leg. A. Stubbe (cPS); Khentii Aimag: 1♂ —20–40 km SW von Somon Zargaltchaan, S von Fluss Zencher gol, 1200 m, 23.VIII.1963, leg. Dr. Z. Kaszab (HNHM); 5 ex. —Batshireet 3 km E of Bakh Gol, 03.08.2019, leg. P.-H. Schnitter (cPS); Govisümber Aimag: 2♀ ♀ —25–35 km NNW of Choir, 12.VII.[1]975, Gurieva (ZIN); 1♀ —20 km SO von Čojren, 1200 m, 1.VII.1963, leg. Dr. Z. Kaszab (HNHM); Dornogovi Aimag: 1♂ —12 km NE of Mandakh, 15. VI.[1]971, Kerzhner (ZIN); 1♀ —Erdene-Dalay, 26. V.19[73], Sowjet-Mongol. Biol. Exped. 1973 (ZMUM); 1♂ 2♀♀ —40 km NW von Chara-Eireg, 1150 m, 30. VI.1963, leg. Dr. Z. Kaszab (HNHM); Dornod (= East) Aimag: 1♂ 5♀♀ —Čojbalsan, 1969, leg. Gy. Márton (HNHM); 1♂ —50 km ESE of Choibalsan, 12.VIII.[1]976, Gurieva; 6♂♂ 1♀ — Bui Nuur (= Buir) Lake, VI.[18]99, exp. of Potanin; 1♀ —steppe W of Bui Nuur (= Buir) Lake, 25.VIII.[19]28, A. Ivanov; 1♀ —shore of Bui Nuur (= Buir) Lake, 25 km NE Bayan Nuur Lake, 17. VI.[1]971, Kozlov; 1♂ —10 km W of Dund-Bayan Lake, 27.VIII.[1]975, Gurieva; 1♂ —20 km ENE of Bayan-Uul, 8.VII.[1]976, Gurieva; 1♀ — Khökh Nuur Lake, 24–26. VI.[1]976, Gurieva; 1♂ —Derkhin-Tsagaan-Ovoo, 60 km ENE of Bayan-Burda, 21. VII.[1]971, G. Medvedev; 1♂ —Numragin-Gool River, 32 km SE of Salkhit Mnt, 24.VII.[1]971, G. Medvedev (all ZIN); 6♀♀ —Chamardavas uil, 80 km SO von Somon Chalcingol, 60 m, 13.VIII.1965, leg. Dr. Z. Kaszab (HNHM); 1 ex. —Chalchgol E, Sambariin Tsagaan Nuur, 02.08.2013, leg. P.-H. Schnitter; 1 ex. —Hyanganï Salbar Uuls, Hingan Mts., Nömrög Gol, 02.08.2013, leg. P.-H. Schnitter (both cJG); 4 ex. — ibid; 2 ex. —Tsagaannuur, Khalkhyn Gol, 31.07.2013, leg. P.-H. Schnitter (both cPS); Bayankhongor Aimag: 1♂ —25 km W of Bömbögör, 14. VI.[1]980, G. Medvedev (ZIN); Övörkhangai Aimag: 1♀ —Ongi River, upstream of Khalkha River, 12.VII.[1]926, Kirichenko (ZIN); 1♂ —64 km W of Erdenecant, 1260 m, 47°05’N 104°05’E, 23–24.5.1990, leg. Gy. Fábián, M. Hreblay, L. Peregovits, G. Ronkay (cJW); 1 ex. —Harhorin Karzhorum, Changai Mts., 15.07.2001, leg. K. Gedeon (cPS); Dundgovi Aimag: S of Delgercogt, 46 o 03’01’’N 106 o 21’38’’E, 07.08.2019, leg. A. Rubenyan (MSPU); 1♂ 1♀ — Central Mongolia, Mandal-Gobi, 6.1964, Shuvalov (ZIN); 1 ex. —80 km S of Ulan Bataar, Mandal-Gobi, Halbwüste, 16.07.1999, leg. A. Stubbe (cPS); 2♀♀ —SW of Chuld und Delgerehangaj, Choot bulag, 1480 m, 10. VI.1967, leg. Dr. Z. Kaszab; 2♂♂ — ibid, 12.VII.1967 (both HNHM); Ömnögovi Aimag: 1♂ —60 km W von somon Bulgan, 1120 m, 21. VI.1964, leg. Dr. Z. Kaszab; 1♂ —20 km NO von Somon Bulgan, Bajan zag, 1200 m, 6–7.VII.1967, leg. Dr. Z. Kaszab (both HNHM); 1♀ —Dalansadgat, 29.6.1983, leg. Ulrich Salk Bautzen (cJW); Sükhbaatar Aimag: 3♂♂ 4♀♀ —25 km NW of Bayandelger, 19.VIII.[1]975, Gurieva; 1♂ —sands Ongon-Els, 15 km SSE of Hongor, 5– 6.VII.[1]971, Kerzhner (both ZIN); 1♂ —Ongon-Els, 10 km S von Somon Hongor, 900 m, 3–5.VIII.1965, leg. Dr. Z. Kaszab (NHMW); 1♂ — ibid, 8.VIII.1965 (HNHM); 1♂ —16 km W of Govi-Nuur, 30–31.VIII.1977 (ZMUM); NORTH KOREA: 2♂♂ 1♀ —N Korea, 20–22. V.[18]97, V. Komarow (ZIN); 1♀ — Ryanggang prov., Mupo, Tumangang, 1000 m, 29.IV.1988, leg. O. Merkl &amp; G. Szél; 1♀ —Khamgen prov., Chongjin, 04.06.1991, leg. Ronkay &amp; Vojnits (both HNHM); CHINA, Nei Mongol Autonomus Region: 1♂ —along Kerulen (= Herlen) River, exp. of G. Potanin (ZIN); 1♂ — Inner Mongolia, Chifeng, Keteng, 10.IX.2006, leg. Y. Wang &amp; M.J. Qi (NEFU); Heilongjiang Province: 3♂♂ —Manchuria, railway station Imanpo (= Yimianpozhen), 20–21.VII.[1]914, Emelianov; 7♂♂ 7♀♀ —Manchuria, railway station Hengdaohezizhen (both ZIN); 1♂ –China, Mandzhuria, Gaolindzy st., 17.VII.1973 (cJW); 1♀ —Muleng, Xiaojinshan, 1. VI.1961; 1♂ —Mudanjiang, Ningan, 17–21.VIII.2008 (both NEFU); Liaoning Province: 1♂ 1♀ —PanJinShi, QingyanSi, 300 m, 13.VII.2007, Coll. Liu J.-Z. (cXX); Sichuan Province: 1♂ 1♀ — China, Sichuan, Wenchuan, 23.05.2012, leg. S. Murzin (cJG); Jiangxi Province: 1♂ 1♀ —Jiangxi, Ganzhou, Mount Jiulian, 24.97°N 114.73°E, 620 m, 25.IX.2016, leg. Zhenhua Liu (SYSU); WITHOUT EXACT LOCALITY: 2♂♂ — Dauria; 1♀ —Sibiria; 1♂ —Amur, ex. coll. G. Basilewsky (all ZMUM).</p><p>References. RUSSIA, Altai  Republic: 7 km W of Edigan, right bank of Katun River; Ulus-Cherga; bank of Karakem River, 2 km of mouth; near Chibit; Kuray; Kuray Steppe, floodplain terrace of Kurayka River; Kuray Steppe, NW Kuray; 6–8 km NE of Kokorya, Kyzylshin River, h~1900–2000 м; valley of Kyzyl-Aryk River (all Matalin &amp; Dudko 2016a); Chemal; Chulyshman Upland, upstream of Chulyshman River; bank of Chuya River, near Chagan-Uzun; Chikhacheva Mnt. Ridge, Buguzun River, h~ 1900 m (all Matalin &amp; Dudko 2016b); Krasnoyarsky  Krai: Jenisseisk (Mandl 1921; Wu 1937); near Krasnoyarsk, Mokhovaya, Sobakina, Narmut, Karaulnaya and Bazaikha Rivers (all Okninskyi 1930); Irkutsk Oblast: Katschug (Poppius 1906); Irkutsk (Mandl 1921); Malyshevka opposite Balagansk, bank of Angara River; Shivera, bank of Angara River (both Lafer 1978); Tazheran steppe, salineland on the bank of Holbo-Nur Lake (Shilenkov 2021);  Republic of Buryatia: Vjerhnje Oudinsk (= Ulan-Ude); Kiakhta (both Motchoulski 1844); Botyi (Jakobson 1907); Werchne-Udinsk (= Ulan-Ude); Troizkosawsk (= Kyakhta) (both Mandl 1921; 1931); Kalenovo (Imelhenova 1980); valley of Dzhida River (Shilenkov &amp; Anichtchenko 1998); Ulan-Ude (Khobrakova 2003); Tamgata (Khobrakova &amp; Dudko 2010); Alla River; Angir-Nuur; Bagdarin; Baisa; Barguzin; Beloozersk; Bilutai; Bolshoe Akubycrit Lake; Verkhnyaya Ivolga; Guzhiry; Gunda; Gusinoe Lake; Gusinoozersk; Dalan-Turu; Dzhida; Dzhirga; Dureny; Deben; Zhakhna; Zakamensk; Zun-Murino; Indola; Kabansk; Karasinoe Lake; Karaftit; Kindikan; Kokorino; Kudara-Somon; Kuitun; Kumyn; Kyren; Maisky; Malyi Amalat; Moigoty; Monahovo; Mongolzhon; Mondy; Mostovoi; Murochi; Muya; Nizhnee Beloe Lake; Novaya Komushka; Novaya Kurba; Nukhu-Daban; Orlik; Orkhabom; Perilly; Savva; Sentsa River; Sentsyn-Tala; Seya River; Taezhnyi; Taksimo; Tankhoi; Tapkhar; Temnik (Atsula); Tissa River; Tunka; Turka; Uoyan; Ust’-Kiran; Kyakhta; Ust’- Kyakhta; Ust’-Muya; Ukherik; Khaigas; Khara-Khuzhir; Kharagun; Kharibyaty; Khoito-Gol; Khoyur-Gorkhon River; Khulugaisha; Tscistokhalo Lake; Khulugaisha; Khutel; Shasnur; Shishkovka (all Khobrakova et al. 2014); Kudara, Kentei-Geb. (Mandl 1921); Zabaykalsky  Krai: Tschita (Mandl 1921); Ingoda River, pad Argalei; mouth of Aga River (both Lafer 978); Khilok (Khobrakova et al. 2014);  Republic of  Yakutia: Ytyk-haja (Poppius 1906); Jakutsk (Mandl 1921); Chochur-Muran, bank of Lena River (Lafer 1978); Amur Oblast: Ulunga River; Natalino, bank of Zeya River; Saskal, mouth of Bereya River; Ekimchan, Selemdzha River (all Lafer 1978); 2.5 km E of Blagoveshchensk, floodplane of Zeya River (Rogatnykh 2009a); Jewish Autonomous Oblast: Bastak Nature Reserve (Rogatnykh 2006); Bastak Nature Reserve, kordon Zaimka Ivakina (Budilov 2012); Khabarovsky  Krai: Bureinsky Natural Reserve, near kordon Strela, bank of Bureya River, 51°38’ N 134°15’ E; mouth of Seregekta River, 51°36’ N 134°08’ E (both Koshkin et al. 2016); Primorsky  Krai: Nikolsk-Ussurijsk; Pokrofka; SutschanGebiet (= Partizansky District) (Mandl 1921); Khasan; Gornoe, Ananievka River; Ussury Reserve; Lesnoy Kordon; Benevka River (tributary of Kievka River); Chernigovka (all Lafer 1978); Lazovsky Nature Reserve: Vtoroi Log; Nogeevskaya; America; Petrova; Staraya Kamenka; Pasechnaya; Kievka; Benevskoe; Lazo (all Sundukov 2009); MONGOLIA, Selenge Aimag: Shaamar (Ulykpan 2008); Khövsgöl Aimag: Bayanzürkh (Lafer 1978); Arkhangaj Aimag: Changai-Geb. (Mandl 1921); NO Ecke des Sees Ogij nur (Mandl 1965); Zavkhan Aimag: Santmargaz sum, Bajan (Chara) nur; Songino (both Mandl 1965); Gobi-Altai Aimag: Zavchan-Niederung, ca 20 km N von Zargalan, 47º20’N 95º48’E (Mandl 1974); CHINA, Nei Mongol Autonomus Region: Baotou, 40.747706 N 109.791142 E, 11.06.2017 (https://www.inaturalist.org/observations/196973519); Heilongjiang Province: Suifen (Süd-UssuriGebiet) (Mandl 1921); Liaoning Province: 41.198646 N 123.267497 E, 12.04.2020 (https://www.inaturalist.org/ observations/41983527).</p><p>Diagnosis.  Cicindela coerulea nitida is recognized by complete and isolated white elytral spots (HI = 8.0–16.0, mean = 9.8) with distinctly sinuate medial band ended by round apical extension (typical pattern Figs 89–106). Moreover, from other subspecies except nominotypical one  C. coerulea nitida is clearly distinguished by long and sharply rounded internal sack so В distinctly or virtually contact with the ventral side of aedeagus tube (Figs 117–132 vs. Figs 167–170, 190–195, 233–240).</p><p>Redescription. TL = 12.0– 15.2 mm (mean = 13.76 mm, n = 93) in males, 12.4–15.4 mm (mean = 14.31 mm, n = 67) in females. Colouration of top side variable from unicoloured green, blue, or violet (Figs 76, 78–80) to bicolored with green or bluish-green head and pronotum and cupreous elytra (Fig. 82).</p><p>Head green or bluish-green, rarely bronze-green with golden or golden-cupreous reflection on genae, frons, vertex, occiput and ocular plates as well as with blue or greenish-blue shallow interocular cavities; in some specimens genae bright blue or head completely blue, violet or black. Labrum with 4–14 (mean = 8, n = 160) submarginal setae (Figs 10–13); LW/LL = 1.83–2.44 mm (mean = 2.14 mm, n = 93) in males, 1.77–2.3 mm (mean = 2.05 mm, n = 67) in females. Mandibles relatively stout, ML/MW = 5.71–7.4 (mean = 6.45, n = 20) in males and 7.2–7.75 (mean = 7.48, n = 20) in females. Scape besides 3–5 apical setae with 1–5 additional setae, rarely glabrous.</p><p>Pronotum green or bluish-green with bright blue anterior and posterior grooves as well as midline and golden or cupreous-golden reflection especially on anterior and posterior lobes; in some specimens completely blue or violet, rarely black; PW/PL = 1.27–1.46 (mean = 1.37, n = 93) in males and 1.28–1.57 (mean = 1.41, n = 67) in females (Figs 19–23). Thoracic segments bright green or blue-green with golden or golden-cupreous reflection or violet with blue tinge on mesepimeron, rarely entirely black.</p><p>Elytra generally cupreous with golden or golden-green reflection especially along suture, not infrequently green, blue or violet, rarely black; EL/EW = 1.45–1.71 (mean = 1.55, n = 93) in males and 1.41–1.57 (mean = 1.50, n = 67) in females; white elytral pattern typically complete, narrow or relatively wide, humeral and apical lunulae always isolated from marginal band, middle band distinctly curved with relatively long transverse portion, oblique or virtually vertical branch ended by rounded apical extension (Figs 89–103, 105); in some specimens humeral lunula divided into two separate spots (Figs 104, 106); sometimes middle band coupling via marginal band with apical lunule (Figs 112, 113) or with both humeral and apical lunulae (Figs 114–116); rarely middle band short and lacking rounded apical portion (Figs 110, 111) or white elytral pattern completely undeveloped (Figs 107–109).</p><p>Abdominal sternites green with golden or golden-cupreous tinge, or blue-violet with greenish-gold reflection.</p><p>Aedeagus relatively long (Figs 45–54), EL/AL = 1.64–2.0 (mean = 1.79, n = 30), apex clearly curved with sharp, medium-sized hook. Internal sac long and sharply rounded towards to ventral side so В distinctly or virtually contact with the aedeagus tube, mt large with smoothly curved apex, VA and DA the largest, DLL relatively large (Figs 117–132).</p><p>Distribution. RUSSIA: Republics of Altai, Khakassia, Tuva, Buryatia and  Yakutia; Krasnoyarsky, Zabaykalsky, Khabarovsky and Primorsky  Krai; Irkutsk, Amur and Jewish Autonomous Oblasts; MONGOLIA: Uvs, Khovd, Zavkhan, Khövsgöl, Bulgan, Arkhangaj, Selenge, Töv (= Central), Khentii, Govisümber, Dornogovi, Dornod (= East), Bayankhongor, Gobi-Altai, Övörkhangai, Dundgovi, Ömnögovi and Sükhbaatar Aimags, Darkhan Uul, Ulaanbaatar; NORTH KOREA: Rason City, North Hamgyong and Ryanggang Provinces; CHINA: Nei Mongol Autonomus Region; Heilongjiang, Jilin, Liaoning, Sichuan and Jiangxi Provinces. The eastern part of the Altai Piedmont Plain is the transition zone between  C. coerulea nitida and the nominotypical subspecies; the valley of the Lakes between Gurvansaikhan and Khangai Mountains is the transition zone between  C. coerulea nitida and  C. coerulea kaszabi while the extreme south of Manchurian Plain is the transition zone between  C. coerulea nitida and  C. coerulea shantungensis (Fig. 246).</p><p>Larva. See Putchkov &amp; Shilenkov (1996) and Putchkov (2013; 2020).</p><p>Ecology. This is one of the most common Siberian and Mongolian tiger beetle inhabiting dry steppes, semideserts and open sands both near and far from water.</p><p>Development. The life cycle is biennial with spring–early summer reproduction. In the spring beetles both of new and previous generations copulate, and, after some time, females lay eggs. The larvae develop in soil burrows, and the third-instar larvae overwinter. Teneral beetles emerge in late summer–early autumn of the next year, feed and then overwinter. In spring of third year overwintered immature beetles maturate and start reproducing (Institute of Zoology Academia Sinica 1986).</p><p>Notes. In 1936 Mandl described the green specimens of  Cicindela coerulea from the Selenga River valley as  C. coerulea selengensis . Mainly green colour of the beetles, marginal band connected with apical lunula (HI = 15–16, mean = 15.3) and poorly developed rounded apical extension on medial band (Figs 80, 112–114) were the main criteria for this. Mandl noted that he had only one blue specimen, and he saw one dark brown specimen from the same locality (Mandl 1936: 12). However, in the collection of Mandl (NHMW) five males and two females with the identical locality labels as in the type specimens were found (see chapter Addition material). Among them, one specimen has blue, one specimen has cupreous-green, and five specimens have cupreous elytra. Moreover, the marginal band in these beetles is short or entirely reduced, and all spots are separated from each other. Thus, by the white elytral pattern (HI = 8–10, mean = 9.6) these specimens are indistinguishable from typical  C. coerulea nitida (HI = 8–14, mean = 9.4). Considering the high variability of the colouration as well as of the white elytral pattern in  C. coerulea nitida, and also the location of the Selenga River valley virtually in the centre of the distribution area of the aforementioned subspecies,  C. coerulea selengensis should be considered only as a pattern-coloured morph of  C. coerulea nitida . Thus, I accept the opinion of Schilder (1953: 564), who considered  Cicindela coerulea selengensis Mandl, 1936 as a synonym of  Cicindela coerulea nitida Lichtenstein, 1796 . The absence of significant differences in basic proportions of body between  C. coerulea nitida and  C. coerulea selengensis (Figs 241–245) reinforces this conclusion.</p><p>In the recent Checklist of the tiger beetles of the World (Wiesner 2020), the names cyanescens Poppius, 1905; viridis Poppius, 1908; nigra Mandl, 1921; cobaltina Lutshnik, 1924; poppiusi Lutshnik, 1924; starodubtzevi Lutshnik, 1924 and starovi Lutshnik, 1924 were listed among the synonyms of  C. coerulea nitida . However, according to the Articles 12.1 and 45.6.2 of ICZN (1999) the name is infrasubspecific if its author used one of the terms “ aberration ” and “ morph ” (or any their abbreviations), as well as according to the Articles 12.1 and 45.5 of ICZN (1999) the name is infrasubspecific if it was published as a fourth name as an addition to a trinomen. Thus, the names  Cicindela tricolor ab. cyanescens Poppius, 1905;  Cicindela tricolor ab. viridis Poppius, 1908;  Cicindela hybrida nitida ab. nigra Mandl, 1921;  Cicindela coerulea nitida ab. cobaltina Lutshnik, 1924;  Cicindela coerulea nitida ab. poppiusi Lutshnik, 1924;  Cicindela coerulea nitida ab. starodubtzevi Lutshnik, 1924;  Cicindela coerulea nitida ab. starovi Lutshnik, 1924 are unavailable and should be excluded from the species group.</p><p>Chinese literature sources mention that  C. coerulea nitida is recorded in total in 17 provinces (Institute of Zoology Academia Sinica 1986; Hu 1992; Zhu et al. 1999). It should be noted, that these records refer to at least three subspecies:  C. coerulea nitida,  C. coerulea kaszab i, and  C. coerulea shantungensis . I was unable to examine the specimens from Jilin, Henan, Shaanxi, Jiangsu, Anhui, Zhejiang, Fujian, Guangdong and Yunnan Provinces. However, according to the presently known distribution of different subspecies, I provisionally consider the specimens from Jilin Province to be  C. coerulea nitida and the specimens from Henan and Shaanxi Provinces to be  C. coerulea shantungensis . The localities in Sichuan and especially in Jiangxi provinces are placed far away from the main distribution area of the species. Due to the scant material (male and female in each of them) I cannot reliably establish the subspecific status of these beetles, but preliminarily consider them as  C. coerulea nitida . The records from other Chinese provinces mentioned above are unconfirmed and the status of the specimens from them is still unclear.</p></div>	https://treatment.plazi.org/id/038787EC0321FFD9CE84E2B96E04FA68	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Matalin, Andrey V.	Matalin, Andrey V. (2025): An intraspecific structure of Сicindela (s. str.) coerulea Pallas, 1773 (Coleoptera, Cicindelidae). Zootaxa 5583 (2): 201-246, DOI: 10.11646/zootaxa.5583.2.1, URL: https://doi.org/10.11646/zootaxa.5583.2.1
038787EC033CFFD5CE84E2446D42FD64.text	038787EC033CFFD5CE84E2446D42FD64.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cicindela coerulea Mandl 1965	<div><p>Cicindela (s. str.) coerulea kaszabi Mandl, 1965</p><p>Figs 57–61, 83–88, 133, 134, 145–149, 155–170</p><p>Cicindela coerulea kaszabi 
Mandl, 1965: 241, Abb. 1, 2 (Type locality— Mongolia,  Südgobi Aimak, 60 km W von Somon Bulgan).</p><p>Cicindela hybrida nitida Lichtenstein, 1796: Institute of Zoology Academia Sinica 1986: 417; Hu 1992: 374; Zhu et al. 1999: 26.</p><p>Cicindela coerulea kaszabi Mandl, 1965: Cassola &amp; van Nidek 1984: 10; Wiesner 1992: 118; 2020: 184; Lorenz 1998: 42; 2005: 131; Puchkov &amp; Matalin 2003: 104; Putchkov &amp; Matalin 2017: 227.</p><p>Type material.   Holotype of  C. coerulea kaszabi Mandl, ♂ (Figs 57, 58, 83, 84, 145, 155)— Mongolia,  Südgobi aimak, 60 km W v. Somon Bulgan, 1120 m, Exp. Dr. Z. Kaszab 1964 [typed white label], Nr. 163, 21.VI.1964 [typed white label], Holotypus, 1964,  Cicindela coerulea kaszabi Mandl [handwritten white label], Holotypus,  Cicindela nitida ssp. kaszabi m., det. K. Mandl, 1964 [handwritten and typed red label] (HNHM)  .</p><p>Additional material.   MONGOLIA, Bayan-Ölgii Aimag: 1♂ — Chinese Altai, valley of  Tsagan-Gol River, VII.1899, exp. of Kozlov (ZIN) ;   Ömnögovi Aimag: 1♂ 1♀ — Mongolia, Sud-Gobi aimag, Khongoryn-Els,  60 km WNW of Bayan-Dalay, 30–31.VII.62, Emelyanov (ZIN) ;   Bayankhongor Aimag: 1♂ — Bayan-Khongor env., somon Dalgar,  Gulin-Tal, h = 1760 m, 8.VIII.1991, leg. S. Kasantsev (cSK) ;   6♀♀ —valley of Tuin-Gol River, near  Orog-Nuur Lake, 2–11.VIII.1926, Kirichenko ;   1♂ — Ih Bogdo, valley of  Orog-Nuur River, 25.7.[1]926, P. Kozlov ;   1♀ — Mongolia austr., Bajanchongor aimag,  Jinst Somon, 4.6.1962, leg. Grebentscikov (all ZIN) ;   Övörkhangai Aimag: 1♂ — Mongolia, Ouver-Khangay aimag, near eastern bank of  Taatsiin-Tsagaan Nuur, 2–4.VIII.969, Zaitsev ;  1♀ — Mongolia, Ouver-Khangay aimag, S slope of Ushugin-Obo, 2.VIII.969, Zaitsev (both ZIN);  Govi-Altai Aimag: 1♀ —from Ulan Bator to Beger-Nur, Eglon, 11–16. VI .   [1]949 (ZIN); CHINA, Xinjiang Uyghur  Autonomic Region:   3♂♂ 3♀♀ —  Barkull env., VII.1910, Rukbedl (ZIN) ;   1♂ —  Dsungarei, Karlyk-Tag ;  1♂ — ibid, V – VI.1908 (both NHMW).</p><p>Diagnosis. This subspecies is readily distinguished from others by the complete and wide white elytral pattern (HI = 9.0–16.0, mean = 14.7) with all marks joined by a marginal band as well as by middle band with relatively short transvers portion, oblique branch with poorly developed or virtually undeveloped round apical extension (Figs 83, 85–88, 155–166), as well as by the proportion of pronotum (Figs 145–149, 244). Moreover  C. coerulea kaszabi is clearly distinguished from nominotypical subspecies and  C. coerulea nitida by shorter aedeagus (Figs57, 59–61) and relatively short slightly rounded towards to ventral side internal sack so В does not contact the aedeagus tube (Figs 167–170 vs. Figs 65–70 and Figs 117–132).</p><p>Redescription. TL = 12.1–13.8 mm (mean = 13.06 mm, n = 10) in males, 13.0– 14.4 mm (mean = 13.74 mm, n = 13) in females (Figs 83, 85–88).</p><p>Head often bright green with golden lustre especially on vertex, occiput and orbital plates, with blue basal strips in anterior portion of orbital plates; in some specimens dull green or bluish-green with cupreous tinge on vertex and occiput or purple with bright golden lustre on genae, clypeus, frons, and base of orbital plates; rarely dull bronze with thin green basal strips in anterior portion of orbital plates. Labrum with 5–11 (mean = 8, n = 23) submarginal setae, LW/LL = 2.0–2.38 (mean = 2.19, n = 10) in males, 1.82–2.16 (mean = 2.01, n = 13) in females (Figs 133, 135). Mandibles relatively stout, ML/MW = 6.0–7.33 (mean = 6.7, n = 7) in males, 6.4–8.0 (mean = 7.46, n = 7) in females. Scape besides 3–4 apical setae with 1–4 additional setae, rarely glabrous.</p><p>Pronotum in most specimens green with golden or cupreous-golden lustre and blue anterior and posterior grooves and midline; in some specimens purple with bright golden grooves and midline; rarely dull bronze with green grooves and midline (Figs 145–149); PW/PL = 1.35–1.57 (mean = 1.44, n = 23). Pro- and mesothorax often bright green with golden or cupreous tinge, metathorax blue or bluish-green; epimerum and episternum green or bluish-green with golden or cupreous-golden lustre; in some specimens all thoracic segments purple-bronze except blue metathorax.</p><p>Elytra cupreous with golden lustre or green with cupreous tinge, rarely blue or greenish-blue, purple, purple-bronze or dull bronze; EL/EW = 1.47–1.61 (mean = 1.52, n = 23); white elytral pattern complete and wide with all marks joined by the marginal band, middle band with relatively short transvers portion, smoothly curved arm with poorly developed or virtually undeveloped round apical extension (Figs 83, 155–159, 161, 163); in some specimens humeral lunula separated from other marks (Figs 160, 164–166); rarely both humeral and apical lunulae unconnected with relativity long marginal band (Fig. 162).</p><p>Abdominal sternites blue-violet or bluish-green, in the latest case with golden-cupreous lateral margin.</p><p>Aedeagus relatively short, with small thin apical hook (Figs 57, 59–61), EL/AL= 1.7–1.95 (mean 1.85, n = 9). Internal sac (Fig. 58) relatively short, with slightly rounded towards to ventral side so В does not contact the aedeagus tube (Figs 167–170).</p><p>Distribution. MONGOLIA: Ömnögovi, Bayankhongor, Övörkhangai, Govi-Altai, Bayan-Ölgii Aimags; CHINA: Xinjiang Uyghur Autonomus Region (Fig. 246).</p><p>Notes. Wu (1937: 15) indicated  C. hybrida nitida from Aksu-Musart, East Chinese Turkestan. Considering the current knowledge on the distribution of the subspecies of  C. coerulea, this record could refer to  C. coerulea kaszabi . However, due to lack of any subsequent information on the records of this species from the aforementioned locality I conclude this record is erroneous.</p></div>	https://treatment.plazi.org/id/038787EC033CFFD5CE84E2446D42FD64	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Matalin, Andrey V.	Matalin, Andrey V. (2025): An intraspecific structure of Сicindela (s. str.) coerulea Pallas, 1773 (Coleoptera, Cicindelidae). Zootaxa 5583 (2): 201-246, DOI: 10.11646/zootaxa.5583.2.1, URL: https://doi.org/10.11646/zootaxa.5583.2.1
038787EC0330FFD2CE84E7486EBAF81E.text	038787EC0330FFD2CE84E7486EBAF81E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cicindela coerulea subsp. shantungensis Mandl 1970	<div><p>Cicindela (s. str.) coerulea shantungensis Mandl, 1970</p><p>Figs 55, 56, 134, 136, 150–154, 171–195</p><p>Cicindela coerulea shantungensis Mandl, 1970: 72, Abb. 3 figs 3–5 (Type locality— China, Tsingtau).  Cicindela tricolor M.F. Adams, 1817: Grum-Grzhymailo 1899: 240.  Cicindela hybrida nitida Lichtenstein, 1796: Wu 1937: 15; Institute of Zoology Academia Sinica 1986: 417; Hu 1992: 374; Zhu et al. 1999: 26.  Cicindela coerulea Pallas, 1773: An 2011: 276, fig. 7, 277.  Cicindela coerulea nitida Lichtenstein, 1796: Kim 1998: 354; Werner 1991: 18, T. 14 fig. 103; Kim et al. 2005: 6, fig. 6.  Cicindela coerulea shantungensis Mandl, 1970: Mandl 1981: 28; Cassola &amp; van Nidek 1984: 10; Wiesner 1992: 118; 2020: 184; Lorenz 1998: 43; 2005: 131; Puchkov &amp; Matalin 2003: 105; Shook &amp; Wiesner 2006: 10; Wu &amp; Shook 2010: 78; Wu 2011: 24; Putchkov &amp; Matalin 2017: 227.</p><p>Type material. Paratypes of  C. coerulea shantungensis Mandl, 1972: 1 ♀ (Figs 171, 172)— China, Kiautschou [handwritten white label], Paratypus [typed red label],  Cicindela coerulea ( nitida) shantungensis m., det. K. Mandl 1970 [handwritten and typed white label], coll. K. Mandl [typed light-blue label]; NHMW-ZOO-COLL, 0002628 [printed white label]; 1♀ (Figs 152, 173, 174, 178)— China, Kiautschou [handwritten white label], Paratypus [typed red label],  Cicindela coerulea ( nitida) shantungensis m., det. K. Mandl 1970 [handwritten and typed white label], coll. K. Mandl [typed light-blue label], 222,  Cicindela coerulea ( nitida) shantungensis Mandl [handwritten white label]; NHMW-ZOO-COLL, 0002627 [printed white label] (both NHMW).</p><p>Additional material.   CHINA: Liaoning Province: 4♂♂ 3♀♀ — Huludao City, Jianchang County, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=119.78307&amp;materialsCitation.latitude=40.53912" title="Search Plazi for locations around (long 119.78307/lat 40.53912)">Yangmadianzi Township</a>, 40.53912°N 119.78307°E, 15.VII.2022, leg. Xuekui Wang (cXX) ;   Hebei Province: 1♂ — China,  Can zhou (= Cangzhou), 1500 m, VI.1991, leg. J.M. Bousquet (cJW);   3♂♂ 2♀♀ — Qinhuangdao City, Haigang District, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=119.55833&amp;materialsCitation.latitude=40.01954" title="Search Plazi for locations around (long 119.55833/lat 40.01954)">Duzhuang Town</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=119.55833&amp;materialsCitation.latitude=40.01954" title="Search Plazi for locations around (long 119.55833/lat 40.01954)">Xiaojiagang Village</a>, 40.019537°N 119.558324°E, 19. V.2023, leg. Zhiyong Bai (cXX) ;   Beijing  Municipality:   3♂♂ 1♀ —  Nord Pekin, Jehol, A. David, 1865 (MNHN) ;  2♂♂ — Peking (ZIN);  Shanxi Province: 1♀ — Shan-si, 18. VI.1884, G. Potanin (ZIN);  Shandong Province: 1♂ —1898, China, Kiautschou (NHMW);   1♀ —  Tsingtau (HNHM) ;  2 ex. — ibid, S. Glaue;  2 ex. — ibid, 31.VIII.1919, E.S. Märker (both MNHUB);  Gansu Province: 1♂ —sanads Pin-Faun-Cha, 8.IV.[18]90, Grum [- Grzhymailo] (ZIN);   1♂ —  Kansou or., K’ing-iang-fou (ZMUM) ;   Qinghai Province: 1♂ 3♀♀ —canyon of Moudjik (= Si-He)  River, 3–7.VII.[18]90, Grum [- Grzhymailo] (ZIN); WITHOUT EXACT LOCALTY :   1♀ —  Collect. Konschegg (ZMUM)  .</p><p>References. CHINA, Beijing Municipality: Peking (Mandl 1951); Peking, 39 o 55’N 116 o 25’E (Mandl 1981); Beijing, Yanqing District, 40.38684 N 115.86325 E, 09.X.2011, Terry Townshend (https://www.inaturalist. org/observations/50020234); Beijing, 40.64235 N 117.46375 E, 22.V.2019, Shan Gui, (https://www.inaturalist. org/observations/27913762); ibid, 40.02167 N 116.14653 E, 40.06845 N 116.1495 E, 40.14761 N 116.0647 E, 40.14709 N 116.08837 E, 40.10333 N 116.0267 E, 40.12321 N 116.13432 E, VI.2023, Zhongqi Wang (https:// www.inaturalist.org/observations/188438646); Beijing, Mentougou District, 40.06896 N 116.02095 E, 02.VI.2023, Yihang Li (https://www.inaturalist.org/observations/167686381); Shanxi Province: Mien Shan, 3700, 37 oN 112 o 30’E (Mandl 1981); Shandong Province: Tsingtau, Kiautschou (Mandl 1970); Qinghai Province: Hainan Tibetan Autonomous Prefecture, Guinan County, 35.68101 N 101.060027 E, 08.07.2016 (https://www.inaturalist. org/observations/69748970); Hainan Tibetan Autonomous Prefecture, Xinghai County, 36.04694 N 100.11991 E, 13.IX.2020 (https://www.inaturalist.org/observations/62014051); ibid, 36.223347 N 100.11401 E, 22.08.2021 (https://www.inaturalist.org/observations/92649596); Gansu Province: Jingyuan County, Baiyin City, 36.94422 N 105.15422 E, 05.VII.2024 (https://www.gbif.org/occurrence/4908088997); SOUTH KOREA: Seoraksan Mountain Natural Reserve; Jeju-do, Hallasan Mountain Natural Reserve (both An 2011); Gyehwado, Gimje-gun (Kim et al. 2005); 36.7 N 128.1 E, 16.07.2013 (https://www.gbif.org/occurrence/3444542042); 36.7 N 126.6 E, 16.05.2019 (https://www.gbif.org/occurrence/2825849468); Henan Province: Jiyuan City, Sanmenxia City (Lingbao), Luoyang City (Yanshi District and Luoning County), Zhengzhou City, Kaifeng City, Zhumadian City (Xiping County), Nanyang City, Xinyang City (Gushi County) (all Zhu et al. 1999).</p><p>Diagnosis.  Cicindela coerulea shantungensis differs from other subspecies by a reduced and relatively narrow white elytral pattern (HI = 5.0–9.0, mean = 7.3) with humeral lunula divided on two isolated spots and oblique branch of middle band without rounded apical extension (Figs 171, 173, 175–189), by relatively long and thin basal portion of aedeagus (Figs 55, 56), and by proportion of pronotum (Figs 150–154, 244).</p><p>Redescription. TL = 13.5–15.2 mm (mean = 14.69 mm, n = 7) in males, 15.2–16.1 mm (mean = 15.71 mm, n = 7) in females (Figs 171, 173, 175–177).</p><p>Head blue or bluish-green, genae bright blue or green, clypeus, supraorbital plates, central portion of frons, vertex and occiput cupreous with golden tinge. Labrum with 6–13 (mean = 8, n = 15) submarginal setae, LW/LL = 2.0–2.3 (mean = 2.13, n = 7) in males, 1.77–2.23 (mean = 1.97. n = 7) in females (Figs 134, 136). Mandibles relatively thin, ML/MW = 6.0–6.33 (mean = 6.17, n = 3) in males, 6.0–8.0 (mean = 7.07, n = 3) in females. Scape besides 3–4 apical setae with a single, rarely with 2–4 additional setae, in some specimens glabrous.</p><p>Pronotum green or bluish-green with golden-cupreous reflection especially in central portion of anterior and posterior lobes, deep anterior and posterior grooves as well as thin midline bright blue; PW/PL = 1.21–1.37 (mean = 1.32, n = 15) in males, 1.32–1.45 (mean = 1.38, n = 12) in females (Figs 150–154). Thoracic segments blue or bluish-green, in some specimens all epimerae and episternae green or blue-green with light golden lustre.</p><p>Elytra cupreous often with golden lustre or purple-cupreous, with distinct granulation most developed on humeri especially in females; EL/EW = 1.4–1.7 (mean = 1.57, n = 27); white elytral pattern as a rule relatively narrow, humeral lunula divided on two isolated spots, rarely with virtually invisible medial portion, middle band in most specimens with short or long thin oblique branch without or rarely with slightly rounded apical extension; marginal band undeveloped, rarely only marked; apical lunule complete with thin medial portion (Figs 171, 173, 175–189).</p><p>Abdominal sternites blue or greenish-blue with light golden lustre.</p><p>Aedeagus with relatively long and thin basal portion and small stout apical hook (Figs 55, 56), EL/AL = 1.65–2.0 (mean = 1.8, n = 10). Internal sac relatively short but rounded towards to ventral side so В contact with the aedeagus tube (Figs 190–195).</p><p>Distribution. CHINA: Beijing Municipality, Liaoning, Hebei, Shandong, Shanxi, Henan, Shaanxi, Gansu, Qinghai Provinces; SOUTH KOREA: Gangwon and Jeonbuk States, South Chungcheong, North Gyeongsang and Jeju Provinces (Fig. 246).</p></div>	https://treatment.plazi.org/id/038787EC0330FFD2CE84E7486EBAF81E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Matalin, Andrey V.	Matalin, Andrey V. (2025): An intraspecific structure of Сicindela (s. str.) coerulea Pallas, 1773 (Coleoptera, Cicindelidae). Zootaxa 5583 (2): 201-246, DOI: 10.11646/zootaxa.5583.2.1, URL: https://doi.org/10.11646/zootaxa.5583.2.1
038787EC0334FFEFCE84E4C16EB6F9F6.text	038787EC0334FFEFCE84E4C16EB6F9F6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cicindela coerulea Pallas 1773	<div><p>Cicindela (s. str.) coerulea przewalskii Dokhtouroff, 1887</p><p>Figs 62, 137, 139, 196, 197, 203–206, 215–220, 233, 237</p><p>Cicindela przewalskii Dokhtouroff, 1887: 439 (Type locality—rives du fleuve Yang tzékiang [= ЯН-ЦЗЫ-ЦЗЯН] et de la rivière</p><p>Kon-tschun-tschu [= КОН-чюН-чу], Tschun-Tschu-Oumà [= Чум-ча-ума] et Bydjun [= БЫ-джуН], 12 000–14 000 pds.</p><p>d’altitude). = przevalskii [auct.] = przewalskyi [auct.]  Cicindela przewalskii Dokhtouroff, 1887: Horn &amp; Roeschke 1891: 50; Fleutiaux 1892: 106; Tschitschérine, 1903: 96.  Cicindela hybrida przewalskii Dokhtouroff, 1887: Jakobson 1905: 190; Horn 1905a: 157, 158; 1905b: 26; 1915: 336; 1926:</p><p>221; 1930: 402; 1938: 45, Taf. 63 figs 6, 7; Winkler 1924: 4; Mandl 1935: 292 figs 72, 73, 296 fig. 62, 299, 300 fig. 16; Wu</p><p>1937: 14; Cassola &amp; van Nidek 1984: 10; Werner, 1991: 16, T. 12 fig. 79; Wiesner 1992: 118.  Cicindela sahlbergi przewalskii Dokhtouroff, 1887: Schilder 1953: 564.  Cicindela coerulea przewalskii Dokhtouroff, 1887: Gebert 1995: 7, 25 Abb. 9; Lorenz 1998: 43; 2005: 131; Puchkov &amp; Matalin</p><p>2003: 104; Shook &amp; Wiesner 2006: 10; Wu &amp; Shook 2010: 78; Wu 2011: 24; Putchkov &amp; Matalin 2017: 227; Wiesner</p><p>2020: 184.</p><p>Type material.   Lectotype of  Cicindela przewalskii Dokhtouroff (designated here), ♂ (Figs62, 137, 196,197, 203, 215, 233, 237)—Amdo, 10–18. VI.1884, Przevalsky [typed white label], silver circle,  C. Przewalskii Dokht. [handwritten white label], Type Dokhtourow [typed white label], LECTOTYPUS,  Cicindela przewalskii Dokhtouroff, 1887, des. A. V. Matalin, 2024 [printed red label] (ZIN). Paralectotypes of  Cicindela przewalskii Dokhtouroff (designated here):   1♀ (Fig. 139, 217)—Amdo, 10–18. VI.1884, Przevalsky [typed white label], silver circle,  Przewalskii Dokht., Tschitscherin det. [handwritten and typed white label] ,   PARALECTOTYPUS,  Cicindela przewalskii Dokhtouroff, 1887, design. A. V. Matalin, 2024 [printed red label]; 3♀♀ (Figs 205, 206, 218–220)—Amdo, 18–24. VI.1884, Przevalsky [typed white label], silver circle,  Przewalskii Dokht., Tschitscherin det. [handwritten and typed white label], PARALECTOTYPUS,  Cicindela przewalskii Dokhtouroff, 1887, des. A. V. Matalin, 2024 [printed red label];   1♀ —Amdo, 24–25. VI.1884, Przevalsky [typed white label], silver circle,  Przewalskii Dokht., Tschitscherin det. [handwritten and typed white label], PARALECTOTYPUS,  Cicindela przewalskii Dokhtouroff, 1887, design. A. V. Matalin, 2024 [printed red label];   1♂ (Fig. 204, 216)— Przewals. [handwritten white label], golden circle,  Przewalskii Dokht.,   ♂, 14.96, A. Semenow det. [handwritten and typed white label], PARALECTOTYPUS,  Cicindela przewalskii Dokhtouroff, 1887, des. A. V. Matalin, 2024 [printed red label] (all ZIN);   2♂ m 1♀ —Amdo, 1884, Przevalsky [typed white label], PARALECTOTYPUS,  Cicindela przewalskii Dokhtouroff, 1887, des. A. V. Matalin, 2024 [printed red label] (SDEI).</p><p>Diagnosis. This subspecies is easily distinguished from others by purple-bronze or purple-cupreous elytra with complete and relatively wide white pattern with virtually transverse, slightly sinuate middle band ending in hook-shaped apical portion and narrowly connecting with wide apical lunula (Figs 196, 215–220), as well as by the proportion of labrum (Figs 137, 139, 245). From nominotypical subspecies and  C. coerulea nitida C. coerulea przewalskii is clearly distinguished by short, slightly rounded towards to ventral side internal sack so В with no contact with the aedeagus tube (Figs 233, 237 vs. Figs 65–70 and Figs 117–132). Moreover  C. coerulea przewalskii significantly differs from  C. coerulea grumi by the proportions of pronotum and labrum (Figs 137, 139 vs. Figs 138, 140–144; Figs 203–206 vs. Figs 207–214; Figs 244, 245).</p><p>Redescription. TL = 12.3 mm (n = 2) in males, 12.2–13.6 mm (mean = 12.72 mm, n = 5) in females (Fig. 196).</p><p>Head bicolour, genae, clypeus, antennal plates, frontal margins and two lateral striae on vertex bright green with golden-cupreous lustre; central portion of frons and vertex, supraorbital plates and occiput bright purple-cupreous. Labrum with 6–10 (mean = 8, n = 7) submarginal setae (Figs 137, 139), LW/LL = 2.30–2.44 (mean = 2.36, n = 7). Mandibles relatively stout, ML/MW = 7.75 (n = 2) in males and 7.5–7.75 (mean 7.58, n = 5) in females. Scape with 3–4 apical setae, glabrous, or with only a single additional seta.</p><p>Pronotum purple or purple-cupreous with golden reflection, anterior and posterior grooves as well as midline bright green and blue (Figs 203–206); PW/PL = 1.36–1.40 (mean = 1.37, n = 7). Prothorax bright green with golden tinge, mesothorax blue-violet, metathorax bluish-green with bright golden-cupreous reflection laterally; pro- and mesepimerum as well as metepisternum bright purple-cupreous with golden-green lustre; mesepisternum in males cupreous-purple with light golden lustre, in females bluish-green forward of coupling sulcus and purple-cupreous behind it.</p><p>Elytra purple-bronze or purple-cupreous with sparse bright green punctures in the base near suture, in some specimens with thin green lateral margin; EL/EW = 1.47–1.55 (mean = 1.51, n = 7); white elytral pattern complete, relatively wide; humeral lunula always separate, middle band virtually transverse, slightly sinuate, with hook-shaped apical portion, narrowly connecting with wide apical lunula by a short marginal band (Figs 196, 215–220).</p><p>Abdominal sternites bluish-green or blue-violet, lateral margin sternites 1 and 2 with golden-green or greenblue tinge.</p><p>Aedeagus relatively short (Fig. 62), EL/AL= 1.91–1.93 (mean 1.92, n = 2). Internal sac with short slightly rounded towards to ventral side internal sack so В does not contact the aedeagus tube (Figs 233, 237).</p><p>Distribution. CHINA: Qinghai Province (Fig. 246).</p><p>Notes. The coordinates of the environs of one of the typical localities of  C. coerulea przewalskii, namely Kon-tschun-tschu [= КОН-чюН-чу] River, are indicated in the book of Przewalskyi (1888: 176) and correspond to 33 o 47.1’N 95 o 54.5’E.</p></div>	https://treatment.plazi.org/id/038787EC0334FFEFCE84E4C16EB6F9F6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Matalin, Andrey V.	Matalin, Andrey V. (2025): An intraspecific structure of Сicindela (s. str.) coerulea Pallas, 1773 (Coleoptera, Cicindelidae). Zootaxa 5583 (2): 201-246, DOI: 10.11646/zootaxa.5583.2.1, URL: https://doi.org/10.11646/zootaxa.5583.2.1
038787EC030AFFEACE84E2E66CCEFBF0.text	038787EC030AFFEACE84E2E66CCEFBF0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cicindela coerulea Pallas 1773	<div><p>Cicindela (s. str.) coerulea grumi Tschitschérine, 1903</p><p>Figs. 63, 64, 138, 140–144, 198–202, 207–214, 221–232, 234–236, 238–240</p><p>Cicindela przewalskii var. grumi Tschitschérine, 1903: 98 (Type locality—Mons Nan-Shan: region do la riv. Babo-che (= EdzinGol), 11 000’–12 700’ d’altitude). =  Cicindela przewalskii var. kozlovi Tschitschérine, 1903: 99(Type locality—Thibet or.: region Dabasun-gobi (au SE du lac Dalaï-dabasu et au N de la Tamyrtyn-ula), synonymized by Schilder (1953: 564). = przevalskii var. grumi [auct.] = przevalskii  var. kozlovi [auct.] = kozlowi [auct.]</p><p>Cicindela hybrida grumi Tschitschérine, 1903: Jakobson 1905: 190; Horn 1905a: 157, 158; 1905b: 27; 1915: 336; 1926: 221; 1930: 402; Winkler 1924: 4; Wu 1937: 14.</p><p>Cicindela grumi grumi Tschitschérine, 1903: Schilder 1953: 564.</p><p>Cicindela coerulea grumi Tschitschérine, 1903: Mandl, 1935: 294 fig. 11, 296 fig. 13; 1936: 13; Horn 1938: 46, Taf. 63 figs 27, 28, Taf. 64 fig. 1; Cassola &amp; van Nidek 1984: 10; Werner, 1991: 18, T. 15 fig. 110; Wiesner 1992: 118; 2020: 184; Lorenz 1998: 42; 2005: 131; Puchkov &amp; Matalin 2003: 104; Shook &amp; Wiesner 2006: 10; Wu &amp; Shook 2010: 78; Wu 2011: 24; Putchkov &amp; Matalin 2017: 227.</p><p>Cicindela hybrida kozlovi Tschitschérine, 1903: Jakobson 1905: 190; Horn 1905a: 158; 1905b: 26; 1915: 336; 1926: 221; 1930: 401; Winkler 1924: 3; Wu 1937: 14.</p><p>Cicindela coerulea kozlovi Tschitschérine, 1903: Mandl, 1935: 294 figs 12, 13, 296 fig. 14, 300 fig. 18; 1936: 14; Horn 1938: 46, Taf. 64 fig. 2; Cassola &amp; van Nidek 1984: 10; Werner, 1991: 18, T. 15 fig. 111; Wiesner 1992: 118; 2020: 184; Lorenz 1998: 42; 2005: 131; Puchkov &amp; Matalin 2003: 104; Shook &amp; Wiesner 2006: 10; Wu &amp; Shook 2010: 78; Wu 2011: 24; Putchkov &amp; Matalin 2017: 227.</p><p>Type material.   Lectotype of  Cicindela przewalskii var. grumi Tschitschérine, 1903 (designated here), ♂ (Figs 138, 198, 199, 234, 238)—р. БадО-хО и ХЫ-хО-дО, 13.VIII.90, Грум. [handwritten white label], silver circle, Type,  v. grumi m., Tschitschérine det. [handwritten and typed white label], LECTOTYPUS,  Cicindela przewalskii var. grumi Tschitschérine, 1903, des. A. V. Matalin, 2024 [printed red label] (ZIN). Paralectotypes of  Cicindela przewalskii var. grumi Tschitschérine, 1903 (designated here):   2♂♂ 4♀♀ (Figs 63, 140, 207–210, 221–226)—same labelled as Lectotype, but PARALECTOTYPUS,  Cicindela przewalskii var. grumi Tschitschérine, 1903, des. A. V. Matalin, 2024 [printed red label];   3♂♂ —р. БадО-хО и ХЫ-хО-дО, 13.VIII.90, Гр. ГржимайлО” [handwritten white label], PARALECTOTYPUS,  Cicindela przewalskii var. grumi Tschitschérine, 1903, des. A. V. Matalin, 2024 [printed red label];   1♀ —р. БадО-хО и ХЫ-хО-дО, 13.VIII.90, Гр. ГржимайлО [handwritten white label], Cic. gemmata potanini Dokht., A. Semenov-Tian-Shansky det., VI.30 [handwritten and typed label], PARALECTOTYPUS,  Cicindela przewalskii var. grumi Tschitschérine, 1903, des. A. V. Matalin, 2024 [printed red label] (all ZIN);   3 ex. —р. БадО-хО и ХЫ-хО-дО, 13.VIII.90, Гр. ГржимайлО [handwritten white label], PARALECTOTYPUS,  Cicindela przewalskii var. grumi Tschitschérine, 1903, des. A. V. Matalin, 2024 [printed red label] (SDEI).</p><p>Lectotype of  Cicindela przewalskii var. kozlovi Tschitschérine, 1903 (designated here), ♂ (Figs 141, 200, 201, 211, 227, 236, 240)—ДабасуН ГОби, ЭКсп. КОЗлОва, 1/ 2.VIII.1901 [handwritten white label], golden circle, Type, Przewalskii  v. kozlovi m., Tschitschérin det. [handwritten and typed white label], LECTOTIPUS,  Cicindela przewalskii var. kozlovi Tschitschérine, 1903, des. A. V. Matalin, 2024 [printed red label] (ZIN). Paralectotypes of  Cicindela przewalskii var. kozlovi Tschitschérine, 1903 (designated here):   2♂♂ 1♀ (Figs 64, 143, 213, 228– 230)—ДабасуН ГОби, ЭКсп. КОЗлОва, 1/ 2.VIII.1901 [handwritten white label], golden circle, Type,  v. Kozlovi m., Tschitschérine det. [handwritten and typed white label], PARALECTOTIPUS,  Cicindela przewalskii var. kozlovi Tschitschérine, 1903, des. A. V. Matalin, 2024 [printed red label] (ZIN);   3 ex. — ДабасуН ГОби, ЭКсп. КОЗлОва, 2.VIII.1901 [handwritten white label], PARALECTOTIPUS,  Cicindela przewalskii var. kozlovi Tschitschérine, 1903, des. A. V. Matalin, 2024 [printed red label] (SDEI).</p><p>Additional material.   CHINA, Qinghai Province: 1♀ —  Quinghai-Lake, eastern edge, 07.07.2011, leg. M. Schülke (cDW) ;   4♂♂ 4♀♀ — Qinghai Prov., Qinghai  Lake nature reserves, east of lake, 36.83 oN 100.75 oE, 3202 m, 18.IV.2015, leg. Zhenhua Liu, Menjie Jin (SYSU) ;   3♂♂ 1♀ — China NW, Qinghai, Wulan (=  Ulan), h ~ 3700–4000 m, 16– 21.07.1997, leg. A. Wrzecionko (SIZ) ;  1♂ — ibid, 1–6.VII.1995;  2♀♀ — ibid, 3500 m, 24– 25.7. 1992, leg. S. Kocman (both cJW);   2 ex. — China, Qinghai-Haixi,  Wulan (=  Ulan), 40 km N Chakayan Nuur ( Lake), 25.07.1992, leg. S. Kocman ;  2 ex. — ibid, 01.07.1995, leg. A. Wrzecionko (both cJG);  2 ex. — ibid, 09.07.2001 (cFB).</p><p>References.   CHINA, Qinghai Province: Haibei Tibetan Autonomous Prefecture, Haiyan County, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.54047&amp;materialsCitation.latitude=37.15033" title="Search Plazi for locations around (long 100.54047/lat 37.15033)">Ganzihe Township</a>, 37.150331 N 100.540469 E, 15.07.2021, 23.07.2022, Zhang Qianyi (https://www.inaturalist.org/ observations/193423392#data_quality_assessment)</p><p>Diagnosis. This subspecies is easily distinguished from other subspecies by transverse or virtually transverse, slightly sinuate middle band often with rounded apical portion (Figs 198, 200, 202, 221–232), as well as by the proportion of labrum and pronotum (Figs 244, 245). From nominotypical subspecies and  C. coerulea nitida C. coerulea grumi is clearly distinguished by short slightly rounded towards to ventral side internal sack so В uncontact with the aedeagus tube (Figs 234–236, 238–240 vs. Figs 65–70 and Figs 117–132). Moreover  C. coerulea grumi significantly differs from  C. coerulea przewalskii by the proportions of pronotum and labrum (Figs 207–214 vs. Figs 203–206; Figs 138, 140–144 vs. Figs 137, 139; Figs 244, 245).</p><p>Redescription. TL = 12.1–14.0 mm (mean = 13.14 mm, n = 12) in males, 12.8–15.0 mm (mean = 14.11, n = 8) in females. Most specimens unicolor brown or cupreous-brown, sometimes black, blue or green, rarely purple-cupreous (Figs 198, 200, 202).</p><p>In brown specimens genae, clypeus, antennal plates, lateral portions of frons and shallow intraocular cavities bright green with cupreous or golden-cupreous reflection, base of genae bright blue; central portion of frons and vertex as well as supraorbital plates bright cupreous, sometimes with golden-purple lustre; occiput cupreous-bronze; shallow interocular cavities bright blue or greenish-blue; in green specimens genae with light golden-cupreous tinge while clypeus and intraocular cavities with bright blue lustre; in some specimens head completely blue or black. Labrum with 6–13 (mean = 8, n = 12) submarginal setae, LW/LL = 2.09–2.33 (mean = 2.18, n = 12) in males, 2.0– 2.27 (mean = 2.16, n = 86) in females (Figs 138, 140–144). Mandibles relatively stout, ML/MW = 6.0–8.0 (mean = 7.08, n = 9) in males and 7.2–8.25 (mean = 7.63, n = 6) in female. Scape besides 3–4 apical setae glabrous, rarely with no more than two additional setae.</p><p>Pronotum purple-cupreous or cupreous-green with golden reflection, anterior and posterior grooves as well as midline bright blue, bluish-green or golden-green; in some specimens completely green, blue or black (Figs 207–214); PW/PL = 1.41–1.52 (mean 1.45, n = 13) in males and 1.43–1.57 (mean = 1.49, n = 10) in females. Proand mesothorax bluish-green with light golden tinge; metathorax blue-violet, cupreous-green laterally with distinct golden reflection; pro- and mesepimeron as well as metepisternum bright cupreous with golden-green reflection; mesepisternum green with golden-cupreous or cupreous-bronze lustre; in some specimens thoracic segments entirely black or black-brown.</p><p>Elytra brown, cupreous-bronze or purple-cupreous with dense golden, blue or greenish-blue punctures sometimes black, blue or green; EL/EW = 1.49–1.63 (mean = 1.56, n = 13) in males and 1.49–1.60 (mean 1.54, n = 10) in females; white elytral pattern complete, relatively wide; humeral and apical lunulae always isolated from slightly curved middle band with short marginal portion, in some specimens middle band lacking rounded apical extension (Figs 198, 200, 202, 221–232).</p><p>Abdominal sternites black, blue or bluish-green with light golden or golden-green tinge.</p><p>Aedeagus relatively short with relatively short and wide basal portion (Figs 63, 34), EL/AL= 1.74–1.96 (mean 1.83, n = 11). Internal sac with short slightly rounded towards to ventral side internal sack so В uncontact with the aedeagus tube (Figs 234–236, 238–240).</p><p>Distribution. CHINA: Qinghai Province (Fig. 246).</p><p>Notes.  Cicindela przewalskii var. grumi and  Cicindela przewalskii var. kozlovi were described from northern and southern environs of Kuko Nor (= Qinghai) Lake, respectively (Tschitschérine 1903). Re-examination of the types of both these taxa leads me to consider them conspecific. The similarity of the structure of the labrum (compare Fig. 138 with Figs 141, 142 and Fig. 140 with Figs 143, 144), pronotum (compare 207–210 with Figs 211–214) and internal sac of the aedeagus (compare Fig. 234 with Figs 235, 236 and Fig. 238 with Figs 239, 240), as well as the complete coincidence of the main proportions of the body (Figs 243–245) help justify this opinion. At the same time, both taxa clearly differ from the most related to them,  C. coerulea przewalskii, by the form of pronotum (Figs 207–214 vs. Figs 203–206), white elytral pattern (Figs 221–232 vs. Figs 215–220) and the main proportions of the body (Figs 243–245). Thus, I accept the opinion of Schilder (1953: 564), who considered  Cicindela kozlovi Tschitschérine, 1903 as a synonym of  Cicindela grumi Tschitschérine, 1903 .</p></div>	https://treatment.plazi.org/id/038787EC030AFFEACE84E2E66CCEFBF0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Matalin, Andrey V.	Matalin, Andrey V. (2025): An intraspecific structure of Сicindela (s. str.) coerulea Pallas, 1773 (Coleoptera, Cicindelidae). Zootaxa 5583 (2): 201-246, DOI: 10.11646/zootaxa.5583.2.1, URL: https://doi.org/10.11646/zootaxa.5583.2.1
038787EC030FFFEACE84E0FC6AA2F80E.text	038787EC030FFFEACE84E0FC6AA2F80E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cicindela coerulea Pallas 1773	<div><p>Identification key to the subspecies of  Cicindela (s. str.) coerulea</p><p>1(2) White elytral pattern undeveloped or strongly reduced (Figs 24–41). 12.5–16.0 mm......  C. coerulea coerulea Pallas, 1773</p><p>2(1) White elytral pattern well developed (Figs 89–106, 110–116, 155–166, 178–189, 215–232).......................... 3</p><p>3(4) White elytral pattern narrow with unconnected marks, humeral lunula divided on two isolated spots, rarely with virtually invisible medial portion, middle band in most specimens with short or long thin oblique branch without rounded apical extension, rarely with slightly developed it, marginal band undeveloped, rarely only marked (Figs 178–189). Aedeagus with relatively long basal portion (Figs 55, 56). 13.5–16.1 mm ......................  C. coerulea shantungensis Mandl, 1970</p><p>4(3) White elytral pattern medium-sized or wide with isolated marks, rarely with connected of some marks or with coupling all of them. Aedeagus with relatively short basal portion (Figs 42–54, 57, 59–64)....................................... 5</p><p>5(6) Medial band virtually transverse or slightly curved with very short oblique branch or without it (Figs 215–232).......... 9</p><p>6(5) Medial band distinctly sinuate with well-developed oblique branch (Figs 89–106, 112–116, 155–166).................. 7</p><p>7(8) White elytral pattern medium-sized, in most specimens with isolated marks (Figs 89–106), rarely with coupling middle band and apical lunula (Figs 112–114) or all marks connected by marginal band (Figs 115, 116), extremely rarely with undeveloped (Figs 110, 111) or clearly reduced (Figs 107–109) marks. Internal sac long and sharply rounded towards to ventral side so В distinctly or virtually contact with the aedeagus tube (Figs 117–132). 12.0– 15.4 mm ...  C. coerulea nitida Lichtenstein, 1796</p><p>8(7) White elytral pattern often wide and all marks connected by marginal band (Figs 83, 155–159, 161, 163); in some specimens humeral lunula virtually separated from other marks (Figs 160, 164–166); rarely both humeral and apical lunulae unconnected with relatively long marginal band (Fig. 162). Internal sac short so В does not contact the aedeagus tube (Figs 167–170). 12.1–14.5 mm ...............................................................  C. coerulea kaszabi Mandl, 1965</p><p>9(10) Labrum shorter, LW/LL = 2.3–2.44. Pronotum narrower, PW / PL = 1.36–1.4, with slightly rounded subparallel lateral sides (Figs 203–206). Middle band narrowly connected with apical lunula (Figs 215–220). 12.2–13.6 mm ......................................................................................  C. coerulea przewalskii Dokhtouroff, 1887</p><p>10(9) Labrum longer, LW/LL = 2.0–2.33. Pronotum wider, PW / PL = 1.41–1.57, with virtually straight, convergent at its base with the sides (Figs 207–214). All elytral spots separated from each other (Figs 221–232). 12.1–15.0 mm.........................................................................................  C. coerulea grumi Tschitschérine, 1903</p></div>	https://treatment.plazi.org/id/038787EC030FFFEACE84E0FC6AA2F80E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Matalin, Andrey V.	Matalin, Andrey V. (2025): An intraspecific structure of Сicindela (s. str.) coerulea Pallas, 1773 (Coleoptera, Cicindelidae). Zootaxa 5583 (2): 201-246, DOI: 10.11646/zootaxa.5583.2.1, URL: https://doi.org/10.11646/zootaxa.5583.2.1
