taxonID	type	description	language	source
0380C53FFF92FF8D6394FA00FD1B1E58.taxon	description	(Figure 3)	en	Szederjesi, Tímea, Pop, Antonia Adriana, Horváth, Edit, Krízsik, Virág, Csuzdi, Csaba (2025): Remarks on the Dendrobaena alpina (Rosa, 1884) species complex in the Carpathian Basin (Oligochaeta, Lumbricidae). Zootaxa 5589 (1): 201-209, DOI: 10.11646/zootaxa.5589.1.16, URL: https://doi.org/10.11646/zootaxa.5589.1.16
0380C53FFF92FF8D6394FA00FD1B1E58.taxon	materials_examined	Holotype. ICB- 437 Romania, Retezat Mts, Stanuleti, N 45.29752 ° E 22.83293 °, 30.07.2005, leg. Cs. Csuzdi, M. Csuzdi, V. V. Pop. Paratypes. ICB- 436 1 ex., HNHM / 15027 15 ex. Romania, Retezat Mts, Rotunda, N 45.3150 ° E 22.77716 °, 30.07.2005, leg. Cs. Csuzdi, M. Csuzdi, V. V. Pop. HNHM / 15600 12 ex., Romania, Retezat Mts, Valea Lapusnicului, 6 km after Rotunda, spruce forest, 30.06.2005, leg. Cs. Csuzdi, M. Csuzdi, V. V. Pop. HNHM / 15030 10 ex., HNHM / 15601 7 ex., Romania, Retezat Mts, Val. Lapusnicului, Stănuleţi, 30.06.2005, leg. Cs. Csuzdi, M. Csuzdi, V. V. Pop.	en	Szederjesi, Tímea, Pop, Antonia Adriana, Horváth, Edit, Krízsik, Virág, Csuzdi, Csaba (2025): Remarks on the Dendrobaena alpina (Rosa, 1884) species complex in the Carpathian Basin (Oligochaeta, Lumbricidae). Zootaxa 5589 (1): 201-209, DOI: 10.11646/zootaxa.5589.1.16, URL: https://doi.org/10.11646/zootaxa.5589.1.16
0380C53FFF92FF8D6394FA00FD1B1E58.taxon	diagnosis	Diagnosis. Length 56 – 77 mm, diameter 2 – 3 mm, setae widely paired. Colour slight pinkish anterio-dorsally. First dorsal pore in 5 / 6. Clitellum on 27, 28 – 33, tubercles on 30 – 32. Male pore great slit on 15, surrounded by a pair of small glandular crescents. Nephridial pores irregularly alternate between b and above d. Three pairs of vesicles in 9,11,12; spermathecae two pairs in 9 / 10 – 10 / 11 open in M. Well-developed calciferous diverticula in 11 – 12. Hearts in segments 6 – 10, nephridial bladders simple, sausage-shaped. External characters. Holotype, tail cut-off specimen. Paratypes 56 – 77 mm long and 2 – 3 mm wide. Number of segments 123 – 140. Colour slight pinkish anteriorly. Prostomium epilobous ½ closed. First dorsal pore at intersegmental furrow 5 / 6. Setae widely paired. Setal arrangement behind clitellum: aa: ab: bc: cd: dd = 1.88: 1: 1.19: 1: 2.81. Male pores great slits on segment 15, surrounded by glandular crescents, protruding slightly into the neighbouring segments. Nephridial pores irregularly alternate between setal line b – d. Clitellum on segments 27, 28 – 33. Tubercula pubertatis slightly glandular on 30 – 32, sometimes just a small notch. Glandular tumescences usually lacking, rarely on 33 ab. Internal characters. All septa membranous. Testes and funnels paired in segments 10 – 11, covered with perioesophageal testis sac. Three pairs of seminal vesicles in 9, 11, and 12. Spermathecae two pairs in 9 / 10, 10 / 11 with external openings near the mid-dorsal line (M). Calciferous glands well-developed in segments 11 – 12. Paired hearts appear in segments 6 – 10. Nephridial bladders simple, sausage-shaped. Crop in segments 15 – 16, and gizzard in segments 17 – 18. Typhlosolis bilobous. Longitudinal musculature of pinnate type.	en	Szederjesi, Tímea, Pop, Antonia Adriana, Horváth, Edit, Krízsik, Virág, Csuzdi, Csaba (2025): Remarks on the Dendrobaena alpina (Rosa, 1884) species complex in the Carpathian Basin (Oligochaeta, Lumbricidae). Zootaxa 5589 (1): 201-209, DOI: 10.11646/zootaxa.5589.1.16, URL: https://doi.org/10.11646/zootaxa.5589.1.16
0380C53FFF92FF8D6394FA00FD1B1E58.taxon	etymology	Etymology. The new species is named in honour of Victor and Victor V. Pop in recognition of their tremendous work in earthworm research.	en	Szederjesi, Tímea, Pop, Antonia Adriana, Horváth, Edit, Krízsik, Virág, Csuzdi, Csaba (2025): Remarks on the Dendrobaena alpina (Rosa, 1884) species complex in the Carpathian Basin (Oligochaeta, Lumbricidae). Zootaxa 5589 (1): 201-209, DOI: 10.11646/zootaxa.5589.1.16, URL: https://doi.org/10.11646/zootaxa.5589.1.16
0380C53FFF92FF8D6394FA00FD1B1E58.taxon	discussion	Remarks. The new species is very similar to certain other Carpathian, Balkanic and Anatolian species (Table 3). However, it differs from D. armeniaca by the position of the spermatheal openings (M vs. d – M); from the pale D. misirlioglui by its slight pinkish pigmentation; from D. orientaloides by its shorter clitellum and tubercles (27,28 – 33 vs. 25 – 33; 30 – 32 vs. 30 – 32, 1 / n 33), its lighter pigmentation and the position of the last hearts (10 vs. 11); and from the pale D. popi by its slight pinkish pigmentation, the position of the spermathecal openings (M vs. M> d) and the position of the last hearts (10 vs. 11).	en	Szederjesi, Tímea, Pop, Antonia Adriana, Horváth, Edit, Krízsik, Virág, Csuzdi, Csaba (2025): Remarks on the Dendrobaena alpina (Rosa, 1884) species complex in the Carpathian Basin (Oligochaeta, Lumbricidae). Zootaxa 5589 (1): 201-209, DOI: 10.11646/zootaxa.5589.1.16, URL: https://doi.org/10.11646/zootaxa.5589.1.16
0380C53FFF93FF8E6394F897FEF51A7C.taxon	materials_examined	Material examined. HNHM / 17203 10 ex. Romania, Munţii Parâng, between Novaci and Rânca, 1132 m, N 45.23141 ° E 23.68690 °, alpine pasture with scattered beech trees, 12.11.2016, leg. Cs. Csuzdi, L. Dányi, V. V. Pop, T. Szederjesi. HNHM / 17219 3 ex. Romania, Munţii Vâlcan, N of Vaidei, bank of ŞuŞita Verde, 425 m, N 45.19456 ° E 23.26883 °, beech forest, 11.11.2016., leg. Cs. Csuzdi, L. Dányi, V. V. Pop, T. Szederjesi. HNHM / 17222 15 ex. Munţii Parâng, stream valley NW of Novaci, 505 m, N 45.19327 ° E 23.66583 °, chestnut forest patch, 13.11.2016., leg. Cs. Csuzdi, L. Dányi, V. V. Pop, T. Szederjesi.	en	Szederjesi, Tímea, Pop, Antonia Adriana, Horváth, Edit, Krízsik, Virág, Csuzdi, Csaba (2025): Remarks on the Dendrobaena alpina (Rosa, 1884) species complex in the Carpathian Basin (Oligochaeta, Lumbricidae). Zootaxa 5589 (1): 201-209, DOI: 10.11646/zootaxa.5589.1.16, URL: https://doi.org/10.11646/zootaxa.5589.1.16
0380C53FFF93FF8E6394F897FEF51A7C.taxon	diagnosis	Diagnosis. Length 90 – 130 mm, diameter 3 – 4 mm, setae widely paired. Colour greiysh, pigmentation lacking. First dorsal pore in 5 / 6. Clitellum on ½ 26, 26 – 33, ½ 34, tubercles on 30 – 32. Male pore large on 15. Nephridial pores irregularly alternate between b and above d. Two pairs of seminal vesicles in 11,12; spermathecae two pairs in 9 / 10 – 10 / 11 open in M. Well-developed calciferous diverticula in 11 – 12. Hearts in segments 6 – 11, nephridial bladders sausage-shaped with a small ectal loop.	en	Szederjesi, Tímea, Pop, Antonia Adriana, Horváth, Edit, Krízsik, Virág, Csuzdi, Csaba (2025): Remarks on the Dendrobaena alpina (Rosa, 1884) species complex in the Carpathian Basin (Oligochaeta, Lumbricidae). Zootaxa 5589 (1): 201-209, DOI: 10.11646/zootaxa.5589.1.16, URL: https://doi.org/10.11646/zootaxa.5589.1.16
0380C53FFF93FF8E6394F897FEF51A7C.taxon	discussion	Remarks. Dendrobaena popi is close to the unpigmented D. misirlioglui but differs clearly by the number of vesicles (Table 3).	en	Szederjesi, Tímea, Pop, Antonia Adriana, Horváth, Edit, Krízsik, Virág, Csuzdi, Csaba (2025): Remarks on the Dendrobaena alpina (Rosa, 1884) species complex in the Carpathian Basin (Oligochaeta, Lumbricidae). Zootaxa 5589 (1): 201-209, DOI: 10.11646/zootaxa.5589.1.16, URL: https://doi.org/10.11646/zootaxa.5589.1.16
0380C53FFF90FF8F6394F9D2FCB6181A.taxon	materials_examined	Material examined. HNHM / 17091 2 ex. Turkey, ŞavŞat, Artvin, SSE of the village, Abies forest mixed with deciduous trees, N 41.23276 ° E 42.37156 °, 1311 m, 25.04.2015, leg. L. Dányi, M. B. Kaydan. HNHM / 17552 3 ex. Azerbaijan, Qax district, Lǝkit, large stream and its littoral, 530 m, N 41.48803 ° E 46.85581 °, 07.05.2019, leg. D. Murányi.	en	Szederjesi, Tímea, Pop, Antonia Adriana, Horváth, Edit, Krízsik, Virág, Csuzdi, Csaba (2025): Remarks on the Dendrobaena alpina (Rosa, 1884) species complex in the Carpathian Basin (Oligochaeta, Lumbricidae). Zootaxa 5589 (1): 201-209, DOI: 10.11646/zootaxa.5589.1.16, URL: https://doi.org/10.11646/zootaxa.5589.1.16
0380C53FFF90FF8F6394F9D2FCB6181A.taxon	diagnosis	Diagnosis. Length 40 – 60 mm, diameter 2 – 3 mm, setae widely paired. Colour slight reddish. First dorsal pore in 5 / 6. Clitellum on ½ 27, 28 – 33, ½ 34 tubercles on 30 – 32. Male pore moderate on 15. Nephridial pores irregularly alternate between b and above d. Three pairs of seminal vesicles in 9, 11, 12, those in 9 usually smaller than the others; spermathecae two pairs in 9 / 10 – 10 / 11 open between d <M. Calciferous diverticula in 11 – 12. Hearts in segments 6 – 10, nephridial bladders sausage-shaped (Table 3).	en	Szederjesi, Tímea, Pop, Antonia Adriana, Horváth, Edit, Krízsik, Virág, Csuzdi, Csaba (2025): Remarks on the Dendrobaena alpina (Rosa, 1884) species complex in the Carpathian Basin (Oligochaeta, Lumbricidae). Zootaxa 5589 (1): 201-209, DOI: 10.11646/zootaxa.5589.1.16, URL: https://doi.org/10.11646/zootaxa.5589.1.16
0380C53FFF90FF8F6394F9D2FCB6181A.taxon	discussion	Remarks. Dendrobaena armeniaca is most close to D. orientaloides, but differs from it by its shorter clitellum and the opening of the spermathecae (between d <M vs. d)	en	Szederjesi, Tímea, Pop, Antonia Adriana, Horváth, Edit, Krízsik, Virág, Csuzdi, Csaba (2025): Remarks on the Dendrobaena alpina (Rosa, 1884) species complex in the Carpathian Basin (Oligochaeta, Lumbricidae). Zootaxa 5589 (1): 201-209, DOI: 10.11646/zootaxa.5589.1.16, URL: https://doi.org/10.11646/zootaxa.5589.1.16
