taxonID	type	description	language	source
038A879EFFAE8C7CFF67FB1E500DF967.taxon	description	In his paper on the Meganola of Côte d’Ivoire, László (2020) addressed the apparent surplus of names in the species complex and synonymised several taxa, including M. septima Hacker, 2012, M. octava Hacker, 2012, M. heteromorpha Hacker, 2012, M. dissoluta Hacker, 2012, M. brachyvalva Hacker, 2012, and M. pseudofuscata Hacker, 2012, with M. monofascia, treating it as a widely distributed pan-African taxon. However, the present study has shown that the western and central African M. monofascia populations differ externally from the southern-eastern African nominotypical population in a more creamy and often darker brownish tinge. Furthermore, although both populations feature identical genitalia characters, the DNA barcode analysis supports their distinctness. Thus, M. dissoluta Hacker, 2012 stat. rev., stat. n. is revived here from synonymy as a western-central African subspecies of M. monofascia. Taxon content	en	László, Gyula M. (2025): Integrative taxonomic review of the Meganola monofascia van Son species group sensu Hacker et al. (Lepidoptera, Noctuoidea, Nolidae, Nolinae, Nolini). Ecologica Montenegrina 87: 135-191, DOI: 10.37828/em.2025.87.7, URL: https://doi.org/10.37828/em.2025.87.7
038A879EFFAD8C7DFF67FE335056FD17.taxon	description	(Figs 1 – 5, 93 – 94, 144)	en	László, Gyula M. (2025): Integrative taxonomic review of the Meganola monofascia van Son species group sensu Hacker et al. (Lepidoptera, Noctuoidea, Nolidae, Nolinae, Nolini). Ecologica Montenegrina 87: 135-191, DOI: 10.37828/em.2025.87.7, URL: https://doi.org/10.37828/em.2025.87.7
038A879EFFAD8C7DFF67FE335056FD17.taxon	materials_examined	Material examined (all in coll. ANHRT). Mozambique. Maputo Special Reserve, see László & Vetina (2019); 1 male, 2 females, Manica Province, Chimanimani National Reserve, Moribane Forest, Ndzou Camp, (Moist Forest), 630 m, 19 ° 44 ' 01.4 " S, 33 ° 20 ' 15.1 " E, 3 – 5. viii. 2018, actinic and MV light trap, leg. László, G., Miles, W., Vetina, A., gen. slide Nos: LGNA 638 (male), LGNA 641 (female). RSA. 1 male, Krüger N. P., Krüger Gate, Garden of Protea Hotel, 300 m, 22 – 23. iii. 2014., leg. M. Hluchy & G. László, gen. slide No.: LGNA 208; 1 male, Limpopo Province, 50 km NW of Mokopane, Waterberg Wilderness Reserve, 1420 m, 24 ° 12 ’ 26 ” S, 28 ° 52 ’ 17 ” E, 8 – 9. iv. 2011, leg. R. Yakovlev. Zambia. 1 male, Hillwood, Ikelenge, 1400 m, 11 ° 16 ’ 02 ” S, 24 ° 18 ’ 59 ” E, 30. iv – 11. v. 2014, light trap, leg. Smith, R., Takano, H., Chmurova, L., Smith, L., gen. slide No.: LGNA 103; 1 male, same site, 25 – 27. xi. 2014, leg. Smith, R., Takano, H., gen. slide No.: LGNA 393; 1 male, same site, 28. v – 4. vi. 2020, leg. Chizuwa, D., Choongo, W.; 1 male, same site and collectors, 5 – 11. vi. 2021; 2 males, 1 female, Kapishya Hot Springs, Shiwa N'gandu Estate, 1437 m, 11 ° 10 ’ 13 ” S, 31 ° 36 ’ 00 ” E, i – iii. 2016, M. T. Harvey coll., leg. Smith, R., Takano, H., gen. slide No.: LGNA 1554 (female); 1 female, Ngonye Falls, Zambezi River, 980 m, 16 ° 39 ' 39 " S, 23 ° 34 ' 24 " E, 17 – 18. iv. 2014, light trap, leg. Smith, R., Takano, H., Chmurova, L., Smith, L., gen. slide No.: LGNA 98; 1 female, Kasanka River Pontoon, Kasanka N. P., 1191 m, 12 ° 34 ' 23 " S, 30 ° 14 ' 05 " E, 2 – 4. xii. 2012, light trap, leg. Smith, R. & Takano, H., gen. slide No.: LGNA 116; 1 male, Bruce-Miller Farm, Choma, 1191 m, 16 ° 37 ' 58.6 " S, 27 ° 02 ' 45.8 " E, 28. ii – 8. iii. 2019, MV light trap, leg. Dérozier, V., Imakando, M., Miles, W., Mulvaney, L.; long series of both sexes, Bruce-Miller Farm Nansai (Southern miombo woodland), 1179 m, 16 ° 38 ' 32 " S, 27 ° 01 ' 47 " E, 25. iv – 1. v. 2022, MV light trap, leg. Miles, W., Mulvaney, L., gen. slide No.: LGNA 1553 (male); 1 male, 1 female, Ijobwe, Sioma Ngwezi NP, 1020 m, 16 ° 53 ' 55 " S, 23 ° 35 ' 54 " E, 18 – 20. iv. 2014, light trap, leg. Smith, R., Takano, H., Chmurova, L., Smith, L., gen. slide No.: LGNA 368 (female); 1 female, Kambishi, Jiwundu, 1340 m, 11 ° 54 ' 39 " S, 25 ° 29 ' 05 " E, 18 – 19. x. 2014, light trap, leg. Smith, R., Takano, H., Oram, D., gen. slide No.: LGNA 85; 3 males, 1 female, Jiwundu Swamp (Miombo / Riverine forest mosaic), 1340 m, 11 ° 51 ' 54 " S, 25 ° 33 ' 20 " E, 29. x – 4. xi. 2018, actinic light trap, leg. Aristophanous, M., Dérozier, V., László, G., Oram, D., gen. slide Nos: LGNA 1008, LGNA 999, LGNA 896 (males), LGNA 1006 (female); 1 male, 3 males, Muchinga Prov., Benyanga village, 1250 m, 10 ° 40 ' 41 " S, 33 ° 27 ' 45 " E, 07 – 12. xii. 2023, MV and actinic light trap, leg. László, G., Morgan, L., Volynkin, A.; gen. slide No.: LGNA 1705; 4 males, 2 females, Mutinondo Wilderness Area, Northern Province, 1460 m, 12 ° 27 ' 06 " S, 31 ° 17 ' 30 " E, 17 – 20. v. 2019, actinic light trap, leg. Dérozier, V., Imakando, M., László, G., Miles, W., gen. slide Nos: LGNA 1028, LGNA 1898 (females), LGNA 1025 (male); 1 female, Muchinga Prov., Muyombe, Mama Muwowo's Lodge, 1230 m, 10 ° 32 ' 40 " S, 33 ° 26 ' 05 " E, 29 – 30. xi. 2023, actinic light trap, leg. Bashford, M., Collins, A., László, G., Morgan, L., Volynkin, A.; 2 males, Eastern Province, 48 km North of Lundazi, Lake Beu, 1140 m, 11 ° 53 ' 56 " S, 33 ° 08 ' 51 " E, 26 – 27. xi. 2023, MV light trap, leg. Bashford, M., Collins, A., László, G., Morgan, L., Volynkin, A.; 1 male, 3 females, Danger Hill, 30 km N of Mpika, Muchinga Province, 1684 m, 11 ° 37 ' 38 " S, 31 ° 33 ' 56 " E, 27 – 30. iv. 2019 Actinic Light Trap, leg. Dérozier, V., László, G., Miles, W., gen. slide Nos: LGNA 1027, LGNA 1029 (females); 1 male, Muchinga Province, Jombo village, 1400 m, 10 ° 27 ' 01 " S, 33 ° 14 ' 30 " E, 30. xi – 05. xii. 2023, MV light trap, leg. Bashford, M., Collins, A., László, G., Morgan, L., Volynkin, A.; 1 female, 1396 m, Chitunta Plain (Miombo / Dambo mosaic), 11 ° 29 ' 12 " S, 24 ° 24 ' 18 " E, 11 – 17. vi. 2021, Fish Bait, Chizuwa, D., Choongo, W. leg., gen. slide No.: LGNA 1712; 5 males, 1 female, Senka Hill, Mukulizi Forest Reserve, Muchinga Province, 1566 m, 09 ° 05 ' 43 " S, 32 ° 05 ' 06 " E, 1 – 6. v. 2019, MV, LepiLED and UV LED Light Trap, Dérozier, V., László, G., Miles, W., gen. slide No.: LGNA 1032 (male), LGNA 1134 (female); 7 males, Zambezi Rapids (Miombo / Riverine forest mosaic), 1205 m, S 11 ° 7 ' 30 ", E 24 ° 11 ' 6 ", 4 – 9. xi. 2018, MV, actinic and LepiLED light trap, leg. Aristophanous, M., Dérozier, V., László, G., Oram, D.; 4 males, Lunzua Falls, 20 km S of Mpulungu, Northern Province, 1416 m, 08 ° 55 ' 38 " S, 31 ° 09 ' 31 " E, 11 – 16. v. 2019, MV light trap, leg. Dérozier, V., László, G., Miles, W., gen. slide No.: LGNA 1026; 2 males, 1 female, Kalungu, North of Isoka, S 9 ° 40 ' 52 ", E 32 ° 42 ' 50 ", 1280 m, 30. iv – 1. v. 2019, actinic light trap, leg. Dérozier, V., László, G., Miles, W.; 1 male, Kalambo Falls, 30 km N of Mbala, Northern Province, 1189 m, 08 ° 35 ' 50 " S, 31 ° 14 ' 26 " E, 9 – 11. v. 2019, actinic light trap, leg. Dérozier, V., László, G., Miles, W.; 1 male, Source of the Zambezi, 1494 m, 11 ° 22 ' 09 " S, 24 ° 18 ' 30 " E, 12 – 13. v. 2015, light trap, leg. Smith, R., Takano, H. & Aristophanous, M.; 1 female, Chilambwe Falls, Kafubu River, 1420 m, S 09 ° 50 ' 13 "; E 30 ° 43 ' 35 ", 8 – 12. ii. 2019, MV Light Trap, Dérozier, V., Mulvaney, L., Smith, R., Takano, H., gen. slide No.: LGNA 1030. Genetic information. This species has been assigned the BIN URI: BOLD ABZ 2648. The intraspecific divergence calculated from 13 samples is in the range of 0.00 – 0.77 %. The nominotypical subspecies of M. monofascia diverges from M. monofascia dissoluta by 1.87 – 2.84 %. Another morphologically similar taxon is M. perfusca, diverging at 1.71 – 2.04 %.	en	László, Gyula M. (2025): Integrative taxonomic review of the Meganola monofascia van Son species group sensu Hacker et al. (Lepidoptera, Noctuoidea, Nolidae, Nolinae, Nolini). Ecologica Montenegrina 87: 135-191, DOI: 10.37828/em.2025.87.7, URL: https://doi.org/10.37828/em.2025.87.7
038A879EFFAD8C7DFF67FE335056FD17.taxon	distribution	Distribution. Confirmed records of nominotypical M. monofascia are known from the RSA, Uganda, Ethiopia, Tanzania, Kenya, DRC, Burundi (as M. septima, M. octava, M. heteromorpha, M. pseudofuscata and M. brachyvalva (Hacker et al. 2012, Hacker 2014 )), Mozambique (László & Vetina 2019) and Zambia (László 2020).	en	László, Gyula M. (2025): Integrative taxonomic review of the Meganola monofascia van Son species group sensu Hacker et al. (Lepidoptera, Noctuoidea, Nolidae, Nolinae, Nolini). Ecologica Montenegrina 87: 135-191, DOI: 10.37828/em.2025.87.7, URL: https://doi.org/10.37828/em.2025.87.7
038A879EFFAC8C63FF67FAC35728FEC9.taxon	description	(Figs 6 – 13, 95 – 97, 148)	en	László, Gyula M. (2025): Integrative taxonomic review of the Meganola monofascia van Son species group sensu Hacker et al. (Lepidoptera, Noctuoidea, Nolidae, Nolinae, Nolini). Ecologica Montenegrina 87: 135-191, DOI: 10.37828/em.2025.87.7, URL: https://doi.org/10.37828/em.2025.87.7
038A879EFFAC8C63FF67FAC35728FEC9.taxon	materials_examined	Material examined (all in coll. ANHRT). Cameroon. 4 males, 3 females, Adamawa Region, Adamawa Plateau, 7.3 km West of Bazanga, Chute de Tello, 1246 m, 07 ° 13 ' 50.6 " N, 13 ° 56 ' 29.2 " E, 24 – 30. ix. 2018, Cold Cathode UV and MV light trap, leg. Sáfián, Sz., Simonics, G., gen. slide Nos.: LGNA 568, LGNA 570 (males), LGNA 866, LGNA 1787 (females). Gabon. 5 males, 1 female, Mikongo (Rougier), Monts de Cristal (Secondary Forest), 430 m, 0 ° 29 ' 47 " N, 11 ° 10 ' 42 " E, 28. vii – 12. viii. 2019, MV, actinic and LepiLED light trap, leg. Albert, J-L., Aristophanous, M., Bie Mba, J., Dérozier, V., Moretto, P., gen. slide Nos: LGNA 1150, LGNA 1270, LGNA 1271 (males), LGNA 1544 (female). Guinea. 2 males, 1 female, Seredou camp, Foret Classee de Ziama (upland forest), 872 m, 8 ° 21 ' 14.53 " N, 9 ° 19 ' 31.93 " W, 24. ii – 06. iii. 2019, general coll., leg. Sáfián, Sz., Simonics, G., Florczyk, K., Koivogui, S.; 3 males, Guinee Forestiere Monts Nimba UNESCO World Heritage Site, Mont Richard Molard camp and ridge (High-Altitude Grassland & Forest), 1400 - 1752 m, 07 ° 36 ' 19 " N, 08 ° 25 ' 30 " W, 1 – 7. vii. 2019, 250 W Blended, UV Cold Cathode and LepiLED light trap, leg. Dérozier, V., Miles, W., Sáfián, Sz.; 9 males, 1 female, Guinee Forestiere, Bossou Forest and Institut de Recherche Environmentale de Bossou (Lowland Forest-Farmland), 690 m, 07 ° 38 ' 32 " N, 08 ° 30 ' 30 " W, 24 – 30. vi. 2019, 8 W UV Cold Cathode, LepiLED and MV light trap, leg. Dérozier, V., Suah Dore, J., Koivagui, S., Miles, W., Sáfián, S., Warner, R., gen. slide No.: LGNA 1555 (male); 1 female, Dalaba, Foret de Tinka, 1289 m, 10 ° 43 ' 14 " N, 12 ° 15 ' 22 " W, 25 – 28. ix. 2019, MV light trap, leg. Geiser, M., Leno, M., Koivagui, S., Miles, W., Mulvaney, L., Sáfián, Sz., gen. slide No.: LGNA 1761; 1 male, Nimba Mts, SMFG concession area, (Societe des Mines de Fer de Guinee), Cite 1, 700 m, 7 ° 42 ' 2.83 " N, 8 ° 23 ' 58.60 " W, 16 – 25. vii. 2017, General coll. at light, leg. Sáfián, Sz.; 1 male, 1 female, Ditinn, Chute de Ditinn, 771 m, 10 ° 49 ' 08 " N, 12 ° 11 ' 30 " W, 18 – 25. ix. 2019, MV light trap, leg. Geiser, M., Leno, M., Koivagui, S., Miles, W., Mulvaney, L., Sáfián, Sz., gen. slide No.: LGNA 1610 (male); 3 females, Geipa Camp, Foret de Diecke, 435 m, 7 ° 26 ' 7.06 " N, 8 ° 50 ' 47.87 " W, 05 – 14. iv. 2019, light trap, blended bulb (250 W), leg. Sáfián, Sz., Koivogui, S., gen. slide No.: LGNA 1148; long series of both sexes, 619 km ESE of Conakry, Nzerekore Region, Prefecture de Lola, Ziela env., 540 – 600 m, x. 2017, 7 ° 42 ' N, 8 ° 21 ' W, leg. local collectors, gen. slide Nos: LGNA 1540, LGNA 1541, LGNA 1542, LGNA 1752. Côte d’Ivoire. 3 males, Mt. Tonkoui Peak, 07 ° 27 ' 15.2 " N; 07 ° 38 ' 12.5 " W, 1171 m, 1 – 8. xi. 2015, light trap, leg. Aristophanous, M., Moretto, P., Ruzzier, E., gen. slide Nos: LGNA 237, LGNA 245, LGNA 246; 1 male, 1 female, same site and collectors, 12 – 18. vii. 2015, gen. slide Nos: LGNA 26 (male), LGNA 27 (female); long series of both sexes, same site, 19 – 27. xi. 2019, leg. Aristophanous, M., Dérozier, V., Moretto, P., Ouattara, S.; 1 male, Tai NP, Tai Research Station (SRET), 174 m, 05 ° 50 ' 00 " N, 07 ° 20 ' 32.0 " W, 25. iii – 17. iv. 2017, MV light trap, leg. Aristophanous, A., Aristophanous, M., Geiser, M., Moretto, P.; 1 female, Park National d'Azagny, entrée Sonaye (Secondary forest), 60 m, 05 ° 14 ' 32 " N, 04 ° 48 ' 05 " W, 25 – 28. xi. 2021, MV light trap, leg. Moretto, P., Mulvaney, L., Takano, H., gen. slide No.: LGNA 1543; 1 female, Azagny NP, 59 m, 05 ° 14 ' 33.7 " N, 04 ° 48 ' 06.2 " W, 26. xi – 1. xii. 2015, light trap, leg. Aristophanous, M., Moretto, P., Ruzzier, E., gen. slide No.: LGNA 256; 1 male, Park National d'Azagny, entrée Sonaye (Secondary forest), 60 m, 05 ° 14 ' 32 " N, 04 ° 48 ' 05 " W, 25 – 28. xi. 2021, MV light trap, leg. Moretto, P., Mulvaney, L., Takano, H., gen. slide No.: LGNA 1865; 1 male, 40 m, Banco National park, 05 ° 23 ' 3.8 " N, 04 ° 03 ' 11.2 " W, 29. xi – 5. xii. 2019, LepiLED light trap, leg. Aristophanous, M., Dérozier, V., Moretto, P., Ouattara, S., gen. slide No.: LGNA 1539; 2 males, Park National du Mont Sângbé (forest / savannah mosaic), 422 m, 08 ° 07 ' 05 " N, 07 ° 19 ' 09 " W, 14 – 20. xi. 2021, MV light trap, leg. Moretto, P., Mulvaney, L., Takano, H.; 2 females, Station d'Ecologie de Lamto (Riverine Forest), 95 m, 06 ° 13 ' 2 " N, 05 ° 1 ' 32 " W, 21 – 25. xi. 2021, MV light trap, leg. Moretto, P., Mulvaney, L., Takano, H. Liberia. 1 male, 1 female, Nimba county, ENNR, Nimba Mts, main ridge (montane forest), 1327 m, 7 ° 31 ' 1.3 " N, 8 ° 31 ' 1.0 " W, 9. xii. 2017, light trap (blended bulb 250 W), leg. Aristophanous, M., Sáfián, Sz., Simonics, G., Smith, L., gen. slide Nos: LGNA 583, LGNA 584; 1 male, Nimba County, Nimba Mts, ENNR, Cellcom road, 750 m, 7 ° 33 ' 3.78 " N, 8 ° 31 ' 46.49 " W, 16 – 28. xii. 2018, Cold Cathode UV light trap (8 W), leg. Sáfián, Sz., Simonics, G., gen. slide No.: LGNA 1146; 1 male, Nimba county, Nimba Mts, ENNR, Grassfield FDA Field Station, 494 m, 7 ° 29 ' 32.45 " N, 8 ° 34 ' 54.05 " W, 22. xi. 2018, light trap, blended bulb (250 W), leg. Sáfián, Sz., Simonics, G.; 1 male, Mount Nimba, Cellcom Road, ENNR, Nimba County, 750 - 1100 m, 7 ° 32 ' 35.89 " N, 8 ° 31 ' 20.63 " W, 14 – 28. xii. 2017, Cold Cathode light bucket trap, leg. Sáfián, Sz., Simonics, G.; 1 male, Nimba Mts, East Nimba Nature Reserve, Little stream, 1250 - 1300 m, 7 ° 31 ' 27.99 " N, 8 ° 30 ' 39.89 " W, 14 – 15. iii. 2017, 8 W Cold Cathode light bucket trap, leg. Sáfián, Sz., Simonics, G.; 4 males, 1 female, Nimba Mts, ENNR, Nimba county (Cellcom road), 1000 - 1100 m, 7 ° 32 ' 45.88 " N, 8 ° 31 ' 21.04 " W, 16 – 28. xii. 2018, Cold Cathode UV light trap (8 W), leg. Sáfián, Sz., Simonics, G., gen. slide Nos: LGNA 1551 (male), LGNA 974 (female); 2 males, 1 female, Lofa County, Wologizi Mts, Ridge Camp 2, 883 m, 8 ° 07 ' 20.79 " N, 9 ° 56 ' 50.75 " W, 22 – 30. xi. 2018, Cold Cathode UV light trap (8 W), leg. Sáfián, Sz., Simonics, G., gen. slide No.: LGNA 1147 (male). Republic of Congo. 1 male, 1 female, Nouabale-Ndoki National Park, Bomassa camp, 341 m, 02 ° 12 ' 36.9 " N, 16 ° 11 ' 30.2 " E, 06 – 14. v. 2023 actinic and LepiLED light trap, leg. Dérozier, V., Kirk-Spriggs, A., László, G., Mvouende, S., gen. slide No.: ANHRT 02482; 1 male, same site, 16 – 23. ix. 2022, MV ligh trap, leg. Dérozier, V., Fouka, B., Kirk-Spriggs, A., Takano, H., gen. slide No.: ANHRT 02719, 1 male, same site and collectors, 10 – 16. x. 2022; 1 female, Nouabale-Ndoki National Park, Bomassa camp, 353 m, 02 ° 12 ' 36.9 " N, 16 ° 11 ' 30.2 " E, 18. vi – 3. vii. 2022, MV light trap, leg. Dérozier, V., gen. slide No.: ANHRT 02541; 2 males, 2 female, 377 m, Likouala Prov., Nouabale-Ndoki NP, Makao camp (Secondary forest), 2 ° 35 ' 42.2 " N, 17 ° 10 ' 08.3 " E, 15 – 21. v. 2023, MV light trap, leg. Dérozier, V., Kirk-Spriggs, A., László, G., Mvouende, S., gen. slide No.: ANHRT 02704; 1 female, Nouabale-Ndoki National Park, Mondika camp, 365 m, 02 ° 21 ' 50.63 " N, 16 ° 16 ' 25.82 " E, 07 – 14. ii. 2023, MV light trap, leg. Bakala N., M., Dérozier, V., Kirk-Spriggs, A., László, G.; 1 male, 2 females, Sangha Prov., Nouabale-Ndoki National Park, Mbeli camp, (Gilbertiodendron forest), 372 m, 02 ° 14 ' 23.8 " N, 16 ° 23 ' 52.1 " E, 1 – 10. x. 2022, LepiLED light trap, leg. Dérozier, V., Fouka, B., Kirk-Spriggs, A., Takano, H., gen. slide Nos: ANHRT 02212 (male), ANHRT 02211 (female); 1 male, same site, 14 – 20. ii. 2023, MV light trap, leg. Bakala N., M., Dérozier, V., Kirk-Spriggs, A., László, G.; 1 male, 1 female, Sangha Prov., Nouabale-Ndoki National Park, Bomassa (Secondary forest), 02 ° 12 ' 25.7 " N, 16 ° 11 ' 39.1 " E, 380 m, 18 – 22. vi. 2022, actinic light trap, leg. Dérozier, V., gen. slide Nos: LGNA 1693 (male), LGNA 1694 (female); 3 males, 1 female, Odzala-Kokoua National Park, Imbalanga Camp, 480 m, 00 ° 45 ' 47 " N, 15 ° 15 ' 39 " E, 05 – 11. iv. 2024, MV and actinic light trap, leg. Bashford, M., László, G., Talani, M., Yaba Ngouma, S., gen. slide No.: LGNA 1401 (female); 4 males, 2 females, Odzala-Kokoua National Park, Camp Imbalanga, 490 m, 00 ° 45 ' 59 '' N, 15 ° 15 ' 20 '' E, 13 – 20. ix. 2024, MV, actinic and LepiLED light trap, leg. Bashford, M., László, G., Talani, M., Volynkin, A.; 1 male, 1 female, same site, 14 – 18. xi. 2024, MV and LepiLED light trap, leg. Bashford, M., Elliott, I., Kirk-Spriggs, A.; 1 male, Odzala-Kokoua National Park, Bangassou Forest near Lobo, 400 m, 00 ° 32 ' 50 " N, 14 ° 51 ' 47 " E, 23 – 26. ix. 2024, actinic light trap, leg. Bashford, M., László, G., Talani, M., Volynkin, A, Yaba Ngouma, S.; 1 male, Odzala-Kokoua National Park, Moba, 380 m, 00 ° 48 ' 52 " N, 15 ° 05 ' 15 " E, 19 – 20. ix. 2024, actinic light trap, leg. Bashford, M., László, G., Talani, M., Volynkin, A., Yaba Ngouma, S.; 2 females, Odzala-Kokoua National Park, Lobo Research Camp, 390 m, 00 ° 35 ' 04 " N, 14 ° 53 ' 12 " E, 20 – 27. ix. 2024, MV light trap, Bashford, M., László, G., Volynkin, A., gen. slide No.: LGNA 1832; 8 males, 1 female, Odzala-Kokoua National Park, Mbomo HQ, 570 m, 00 ° 26 ' 13 '' N, 14 ° 42 ' 01 '' E, 28. ix – 01. x. 2024, MV, actinic and LepiLED light trap, leg. Bashford, M., László, G., Talani, M., Volynkin, A., Yaba Ngouma, S., gen. slide Nos: LGNA 1435, LGNA 1833 (males). Senegal. Long series of both sexes, Simenti, Niokolo-Koba NP, 30 m, 13 ° 1 ' 33 " N, 13 ° 17 ' 4 " W, 3 – 16. vi. 2019, actinic light trap, leg. Aristophanous, M., Moretto, P., Mulvaney, L., gen. slide Nos: LGNA 1557, LGNA 1558, LGNA 1559 (males), LGNA 1561, LGNA 1562, LGNA 1563 (females). Sierra Leone. 1 female, Kono Province, Gori Hills, Near Giehum, 373 m, 08 ° 27 ' 47 " N, 10 ° 46 ' 17 " W, 22 – 29. ii. 2020, leg. Kalnoi, G., Sinyaev, V.; 1 male, Kalainkay nr. Kamabai, Northern Prov., 80 m, 3 – 6. xi. 2015, 09 ° 10 ' 52 " N, 11 ° 56 ' 44 " W, light trap, R. Goff coll., leg. Smith, R., Takano, H., gen. slide No.: LGNA 289; 1 male, Mansonia village at the foothills of Loma Mts, 420 m, 09 ° 07 ' 47 " N, 11 ° 05 ' 06 " W, 6. vi. 2016, light trap, leg. Takano, Miles & Goff, gen. slide No.: LGNA 471. Taxonomic note. László (2020) synonymised M. dissoluta with M. monofascia based on their identical genital morphology. However, this study demonstrated that the western and central African populations of M. monofascia can be distinguished from the nominotypical taxon found in southeastern Africa by the more brownish tint of their forewing ground colour. In addition to the phenotypical differences, the two populations also exhibit significant genetic divergence. Since the name dissoluta applies to the western African M. monofascia, it has been reinstated from synonymy and classified as a subspecies of the latter taxon: M. monofascia dissoluta stat. rev., stat. n. Genetic information. Meganola monofascia dissoluta has been assigned the BIN URI: BOLD ABA 7515. The intraspecific divergence of COI- 5 P sequences calculated from 23 samples is in the range of 0.00 – 1.71 % where the specimens from Senegal diverge at the highest rate. Meganola monofascia dissoluta diverges from the nominotypical subspecies by 1.87 – 2.84 %. The PWD between this taxon and the BIN sharing M. perfusca (Hampson, 1911) is in the range of 0.77 – 1.39 %.	en	László, Gyula M. (2025): Integrative taxonomic review of the Meganola monofascia van Son species group sensu Hacker et al. (Lepidoptera, Noctuoidea, Nolidae, Nolinae, Nolini). Ecologica Montenegrina 87: 135-191, DOI: 10.37828/em.2025.87.7, URL: https://doi.org/10.37828/em.2025.87.7
038A879EFFAC8C63FF67FAC35728FEC9.taxon	distribution	Distribution. Confirmed records of Meganola monofascia dissoluta are known from Nigeria (type locality), DRC and Republic of Congo (as M. septima), Côte d’Ivoire, Senegal, and Ghana (as M. furvitincta) (Hacker et al. 2012, Hacker 2014), Cameroon, and Sierra Leone (as M. monofascia) (László 2020). In this present study, the taxon has been newly recorded from Gabon, Guinea, and Liberia.	en	László, Gyula M. (2025): Integrative taxonomic review of the Meganola monofascia van Son species group sensu Hacker et al. (Lepidoptera, Noctuoidea, Nolidae, Nolinae, Nolini). Ecologica Montenegrina 87: 135-191, DOI: 10.37828/em.2025.87.7, URL: https://doi.org/10.37828/em.2025.87.7
038A879EFFB18C61FF67FEB45065FC6E.taxon	description	(Figs 14 – 16, 98 – 99, 146)	en	László, Gyula M. (2025): Integrative taxonomic review of the Meganola monofascia van Son species group sensu Hacker et al. (Lepidoptera, Noctuoidea, Nolidae, Nolinae, Nolini). Ecologica Montenegrina 87: 135-191, DOI: 10.37828/em.2025.87.7, URL: https://doi.org/10.37828/em.2025.87.7
038A879EFFB18C61FF67FEB45065FC6E.taxon	materials_examined	Material examined (all in coll. ANHRT). Côte d’Ivoire. 1 male, Mt. Tonkoui Peak, 07 ° 27 ' 15.2 " N; 07 ° 38 ' 12.5 " W, 1171 m, 1 – 8. xi. 2015, light trap, leg. Aristophanous, M., Moretto, P., Ruzzier, E., gen. slide No.: LGNA 1734; 2 females, same site, 19 – 27. xi. 2019, leg. Aristophanous, M., Dérozier, V., Moretto, P., Ouattara, S. Gabon. 1 female, Mikongo (Rougier), Monts de Cristal (Secondary Forest), 430 m, 0 ° 29 ' 47 " N, 11 ° 10 ' 42 " E, 28. vii – 12. viii. 2019, actinic light trap, leg. Albert, J-L., Aristophanous, M., Bie Mba, J., Dérozier, V., Moretto, P. Guinea. 2 females, Guinee Forestiere, Bossou Forest and Institut de Recherche Environmentale de Bossou (Lowland Forest-Farmland), 690 m, 07 ° 38 ' 32 " N, 08 ° 30 ' 30 " W, 24 – 30. vi. 2019, MV light trap, leg. Dérozier, V., Suah Dore, J., Koivagui, S., Miles, W., Sáfián, S., Warner, R.; 3 females, 619 km ESE of Conakry, Nzerekore Region, Prefecture de Lola, Ziela env., 540 - 600 m, x. 2017, 7 ° 42 ' N, 8 ° 21 ' W, leg. local collectors. Liberia. 3 females, Lofa County, Wologizi Mts, Ridge Camp, 865 m, 8 ° 07 ' 10 " N, 9 ° 57 ' 11 " W, 24 – 29. xi. 2017, Cold Cathode light bucket and blended bulb (250 W) light trap, leg. Aristophanous, M., Sáfián, Sz., Simonics, G. & Smith, L., gen. slide No.: LGNA 1259; 1 female, Lofa county, Wologizi Mts, Rosewood Camp, 585 m, 8 ° 06 ' 14.9 " N, 9 ° 58 ' 27.3 " W, 18. xi – 1. xii. 2018 MV light trap (8 W), leg. Sáfián, Sz., Simonics, G.; 1 female, Lofa County, Wologizi Mts, base camp forest, 611 m, 8 ° 07 ' 17 " N, 9 ° 57 ' 42 " W, 20. xi – 01. xii. 2017, MV light trap (125 W), leg. Aristophanous, M., Sáfián, Sz., Simonics, G. & Smith, L.; 1 female, Lofa County, Wologizi Mts, Elephant Ridge, 1002 m, 8 ° 7 ' 1.46 " N, 9 ° 55 ' 24.18 " W, 23 – 26. xi. 2018, LepiLED light trap, leg. Sáfián, Sz., Simonics, G.; 1 female, Lofa County, Zuwulor Village School, 230 m, 7 ° 54 ' 52 " N, 9 ° 31 ' 08 " W, 8. xi. 2017, light trap (blended bulb 250 W), leg. Aristophanous, M., Sáfián, Sz., Simonics, G. & Smith, L.; 1 female, Lofa County, Foya Proposed Protected Area, 530 m, 7 ° 56 ' 36 " N, 10 ° 16 ' 36 " W, 10 – 19. xi. 2017, 8 W actinic light bucket trap, leg. Aristophanous, M., Sáfián, Sz., Simonics, G. & Smith, L.; 1 male, 750 m, Nimba County, Nimba Mts, ENNR, Cellcom road, 7 ° 33 ' 3.78 " N, 8 ° 31 ' 46.49 " W, 16 – 28. xii. 2018, Cold Cathode UV Light Trap (8 W), leg. Sáfián, Sz., Simonics, G.; 1 female, Nimba County, Nimba Mts, ENNR, Blue lake road, 583 m, 7 ° 33 ' 38 " N, 8 ° 30 ' 28.5 " W, 7. xii. 2017, general collection, leg. Aristophanous, M., Sáfián, Sz., Simonics, G., Smith, L.; 3 females, Nimba Mts, ENNR, Nimba county (Cellcom road), 1000 – 1100 m, 7 ° 32 ' 45.88 " N, 8 ° 31 ' 21.04 " W, 16 – 28. xii. 2018, Cold Cathode UV Light Trap (8 W), leg. Sáfián, Sz., Simonics, G.; 1 female, Nimba County, Nimba Mts, ENNR, Cellcom Road, 1300 m, 7 ° 31 ' 2.18 " N, 8 ° 31 ' 1.90 " W, 24 – 28. xii. 2018, light trap blended bulb (250 W) & LepiLED, leg. Sáfián, Sz., Simonics, G.; 1 female, Nimba county, ENNR, Nimba Mts, main ridge (montane forest), 1327 m, 7 ° 31 ' 1.3 " N, 8 ° 31 ' 1.0 " W, 9. xii. 2017, light trap (blended bulb 250 W), leg. Aristophanous, M., Sáfián, Sz., Simonics, G., Smith, L. Republic of Congo. 2 males, Likouala Prov., Nouabale-Ndoki National Park, Makao forest, 349 m, 02 ° 36 ' 42.5 " N, 17 ° 09 ' 23.8 " E, 15 – 21. v. 2023, actinic light trap, leg. Dérozier, V., Kirk-Spriggs, A., László, G., Mvouende, S., gen. slide Nos: ANHRT 02502, ANHRT 02505; 1 male, Sangha Prov., Nouabale-Ndoki National Park, Bomassa camp (Secondary forest), 341 m, 02 ° 12 ' 36.9 " N, 16 ° 11 ' 30.2 " E, 16 – 23. ix. 2022, MV light trap, Dérozier, V., Fouka, B., Kirk-Spriggs, A., Takano, H.; 1 male, Nouabale-Ndoki National Park, Mombongo camp, 352 m, 02 ° 10 ' 30.7 " N, 16 ° 8 ' 37.7 " E, 02 – 07. ii. 2023, MV light trap, leg. Bakala N., M., Dérozier, V., Kirk-Spriggs, A., László, G.; 1 male, Nouabale-Ndoki NP, Makao camp (Secondary forest), 377 m, 2 ° 35 ' 42.2 " N, 17 ° 10 ' 08.3 " E, 15 – 21. v. 2023, MV Light Trap, leg. Dérozier, V., Kirk-Spriggs, A., László, G., Mvouende, S.; 1 male, Sangha Prov., Nouabale-Ndoki National Park, Bomassa forest, (Secondary forest), 358 m, 02 ° 11 ' 58.1 " N, 16 ° 11 ' 16.9 " E, 17 – 22. ix. 2022, Actinic Light Trap, leg. Dérozier, V., Fouka, B., Kirk-Spriggs, A., Takano, H., gen. slide No.: LGNA 1374; 1 male, 1 female, Nouabale-Ndoki National Park, Mondika camp, 365 m, 02 ° 21 ' 50.63 " N, 16 ° 16 ' 25.82 " E, 07 – 14. ii. 2023, actinic light trap, leg. Bakala N., M., Dérozier, V., Kirk-Spriggs, A., László, G., gen. slide Nos: ANHRT 03281 (male), LGNA 1375 (female); 1 male, 2 females, Odzala-Kokoua National Park, Mbomo HQ, 570 m, 00 ° 26 ' 13 '' N, 14 ° 42 ' 01 '' E, 28. ix – 01. x. 2024, MV and LepiLED light trap, leg. Bashford, M., László, G., Volynkin, A., gen. slide No.: LGNA 1517 (female); 1 male, Odzala-Kokoua National Park, Lobo Research Camp, 390 m, 00 ° 35 ' 04 " N, 14 ° 53 ' 12 " E, 20 – 27. ix. 2024, MV light trap, leg. Bashford, M., László, G., Volynkin, A., gen. slide No.: LGNA 1468; 1 female, Odzala-Kokoua National Park, Bangassou Forest near Lobo, 400 m, 00 ° 32 ' 50 " N, 14 ° 51 ' 47 " E, 25 – 26. ix. 2024, LepiLED light trap, Bashford, M., László, G., Talani, M., Volynkin, A, Yaba Ngouma, S., gen. slide No.: LGNA 1480. Genetic information. This highly distinctive species was found to be BIN-sharing with M. monofascia dissoluta (BIN URI: BOLD ABA 7515). The intraspecific divergence calculated from three samples is 0.00 %. Meganola perfusca diverges from M. monofascia dissoluta by 0.77 – 1.39 %, from M. monofascia monofascia by 1.71 – 2.04 %.	en	László, Gyula M. (2025): Integrative taxonomic review of the Meganola monofascia van Son species group sensu Hacker et al. (Lepidoptera, Noctuoidea, Nolidae, Nolinae, Nolini). Ecologica Montenegrina 87: 135-191, DOI: 10.37828/em.2025.87.7, URL: https://doi.org/10.37828/em.2025.87.7
038A879EFFB18C61FF67FEB45065FC6E.taxon	distribution	Distribution. Meganola perfusca was reported from Nigeria (type locality), Burkina Faso, Côte d’Ivoire, and Ghana (as M. melanosomalis) (Hacker et al. 2012). This present study confirmed the occurrence of the species in Gabon, Guinea, Liberia and the Republic of Congo. Taxonomic note. Hacker et al. (2012) treated this species under a distinct eponymous species group comprising similarly dark species. However, as the genitalia of M. perfusca are hardly distinguishable in both sexes from those of the M. monofascia complex, furthermore, M. perfusca shows very close genetic affinities to M. monofascia dissoluta (the two species are BIN sharing), the correct assignment of this peculiar species in the M. monofascia complex is beyond any doubt.	en	László, Gyula M. (2025): Integrative taxonomic review of the Meganola monofascia van Son species group sensu Hacker et al. (Lepidoptera, Noctuoidea, Nolidae, Nolinae, Nolini). Ecologica Montenegrina 87: 135-191, DOI: 10.37828/em.2025.87.7, URL: https://doi.org/10.37828/em.2025.87.7
038A879EFFB08C61FF67FB2B5010FF97.taxon	description	The facies of the taxa of this species group are confusingly similar to those of the M. monofascia complex, featuring a well-developed, blackish, broad-based triangular medio-costal patch and a variably developed medial fascia of the forewing. The shared diagnostic male genitalia characters of the species group are the relatively narrow, elongate, apically evenly rounded valva, the variably long, apically pointed harpe and the variably long aedeagus possessing a characteristic rounded carina with a small subapical ventral tooth. In the female genitalia, the species of this complex display a very short sclerotised sinus vaginalis, a heavily sclerotised distal section of the ductus bursae and a single, short and rounded signum bursae accompanied distally by a longitudinal scobinated band. Taxon content	en	László, Gyula M. (2025): Integrative taxonomic review of the Meganola monofascia van Son species group sensu Hacker et al. (Lepidoptera, Noctuoidea, Nolidae, Nolinae, Nolini). Ecologica Montenegrina 87: 135-191, DOI: 10.37828/em.2025.87.7, URL: https://doi.org/10.37828/em.2025.87.7
038A879EFFB78C66FF67FF5F53D6FE80.taxon	description	(Figs 18 – 21, 100 – 101, 147)	en	László, Gyula M. (2025): Integrative taxonomic review of the Meganola monofascia van Son species group sensu Hacker et al. (Lepidoptera, Noctuoidea, Nolidae, Nolinae, Nolini). Ecologica Montenegrina 87: 135-191, DOI: 10.37828/em.2025.87.7, URL: https://doi.org/10.37828/em.2025.87.7
038A879EFFB78C66FF67FF5F53D6FE80.taxon	materials_examined	Material examined (all in coll. ANHRT). Malawi. 1 male, Mzuzu Wildlife Sanctuary, (Afromontane forest), 1294 m, 11 ° 27 ' 30.2 " S, 34 ° 03 ' 11.3 " E, 22 – 27. vi. 2022, MV light trap, leg. Kirk-Spriggs, A., Miles, W., gen. slide No.: LGNA 1583. Mozambique. Maputo Special Reserve, see László & Vetina (2019); 9 males, 1 female, Manica Province, Chimanimani National Reserve, Moribane Forest, Ndzou Camp, (Moist Forest), 630 m, 19 ° 44 ' 01.4 " S, 33 ° 20 ' 15.1 " E, 3 – 5. viii. 2018, actinic and LepiLED light trap, László, G., Miles, W., Vetina, A., gen. slide Nos: LGNA 637, LGNA 639, LGNA 640 (males), LGNA 644 (female). RSA. 1 male, Krüger NP, P. Krüger Gate, Garden of Protea Hotel, 300 m, 22 – 23. iii. 2014, leg. M. Hluchy & G. László, gen. slide No.: LGNA 207. Uganda. 1 female, Bwindi Impenetrable N. P., Cuckooland Lodge, 1700 m, 01 ° 00 ' 17 " S, 29 ° 43 ' 06 " E, 28 – 29. vi. 2022, leg. Naumann, S., Ott, E., Schintlmeister, A., Sulak, H., gen. slide No.: LGNA 1716. Genetic information. Both subspecies of this species belong to the BIN URI: BOLD ABU 7264. The intraspecific divergence of the nominotypical taxon calculated from four samples is 0.00 – 0.82 %. The PWD between the two subspecies is in the range of 0.61 – 1.31 %. The morphologically closest relatives of M. bispermutata bispermutata are M. diagrapha and M. saotomeana sp. n., diverging by 2.81 – 3.34 % and 2.66 – 3.35 %, respectively. Taxonomic note. Hacker (Hacker et al. 2012) described two additional species that show no noticeable morphological divergence from M. bispermutata: M. derrae from southern Kenya and M. stigmatolalis from the Southern Province of Ethiopia. Since they lack distinctive morphological characters, these two species are considered northern populations of M. bispermutata bispermutata, and consequently, they are synonymised here: M. derrae syn. n. and M. stigmatolalis syn. n.	en	László, Gyula M. (2025): Integrative taxonomic review of the Meganola monofascia van Son species group sensu Hacker et al. (Lepidoptera, Noctuoidea, Nolidae, Nolinae, Nolini). Ecologica Montenegrina 87: 135-191, DOI: 10.37828/em.2025.87.7, URL: https://doi.org/10.37828/em.2025.87.7
038A879EFFB78C66FF67FF5F53D6FE80.taxon	distribution	Distribution. Meganola bispermutata occurs sympatrically with M. monofascia in the RSA and Mozambique, where the identification of the two species can only be confirmed by genitalia dissection. The species has also been recorded from southern Kenya under the name M. derrae and southern Ethiopia as M. stigmatolalis (Hacker et al. 2012). In this study, M. bispermutata has been newly recorded from Malawi and Uganda. Interestingly, despite extensive sampling efforts by the ANHRT, it has not been found in Zambia.	en	László, Gyula M. (2025): Integrative taxonomic review of the Meganola monofascia van Son species group sensu Hacker et al. (Lepidoptera, Noctuoidea, Nolidae, Nolinae, Nolini). Ecologica Montenegrina 87: 135-191, DOI: 10.37828/em.2025.87.7, URL: https://doi.org/10.37828/em.2025.87.7
038A879EFFB78C67FF67F950515CFFB5.taxon	description	(Figs 22 – 25, 102 – 103, 148)	en	László, Gyula M. (2025): Integrative taxonomic review of the Meganola monofascia van Son species group sensu Hacker et al. (Lepidoptera, Noctuoidea, Nolidae, Nolinae, Nolini). Ecologica Montenegrina 87: 135-191, DOI: 10.37828/em.2025.87.7, URL: https://doi.org/10.37828/em.2025.87.7
038A879EFFB78C67FF67F950515CFFB5.taxon	materials_examined	Material examined (all in coll. ANHRT). Côte d’Ivoire. 1 male, Mt. Tonkoui Peak, 1171 m, 07 ° 27 ' 15.2 " N, 07 ° 38 ' 12.5 " W, 19 – 27. xi. 2019, LepiLED light trap, leg. Aristophanous, M., Dérozier, V., Moretto, P., Ouattara, S.; 1 male, Abidjan, Banco Forest (Parc National du Banco), 39 – 48 m, 05 ° 23 ' 03.8 " N, 04 ° 03 ' 11.2 W, 21 – 30. iv. 2017, MV light trap, leg. Aristophanous, A., Aristophanous, M., Geiser, M., Moretto, P. Guinea. 1 male, Guinee Forestiere, Bossou Forest and Institut de Recherche Environmentale de Bossou (Lowland Forest-Farmland), 690 m, 07 ° 38 ' 32 " N, 08 ° 30 ' 30 " W, 24 – 30. vi. 2019, MV light trap, leg. Dérozier, V., Suah Dore, J., Koivagui, S., Miles, W., Sáfián, S., Warner, R., gen. slide No.: LGNA 1163; 1 female, Nimba Mts, SMFG concession area (Societe des Mines de Fer de Guinee) 600 Forest, 1500 m, 7 ° 39 ' 49.93 " N, 8 ° 22 ' 22.19 " W, 21 – 30. viii. 2017, Cold Cathode UV light trap (8 W), leg. Sáfián, Sz., Simonics, G., gen. slide No.: LGNA 1247; 9 males, 3 females, Guinee Forestiere Monts Nimba UNESCO World Heritage Site, Mont Richard Molard camp and ridge (High-Altitude Grassland & Forest), 1400 – 1752 m, 07 ° 36 ' 19 " N, 08 ° 25 ' 30 " W, 1 – 7. vii. 2019, UV Cold Cathode, LepiLED and 250 W Blended light trap, leg. Dérozier, V., Miles, W., Sáfián, Sz., gen. slide Nos: LGNA 1545 (male), LGNA 1585, LGNA 1552 (females); 4 males, 1 female, 619 km ESE of Conakry, Nzerekore Region, Prefecture de Lola, Ziela env., 540 – 600 m, x. 2017, 7 ° 42 ' N, 8 ° 21 ' W, leg. local collectors, gen. slide Nos: LGNA 1571, LGNA 1760, LGNA 1793 (males). Republic of Congo. Sangha Prov., Nouabale-Ndoki National Park, Bomassa camp (Secondary forest), 341 m, 02 ° 12 ' 36.9 " N, 16 ° 11 ' 30.2 " E, 16 – 23. ix. 2022, MV light trap, leg. Dérozier, V., Fouka, B., Kirk-Spriggs, A., Takano, H., gen. slide No.: LGNA 1388; 1 male, Nouabale-Ndoki National Park, Mombongo camp, 352 m, 02 ° 10 ' 30.7 " N, 16 ° 8 ' 37.7 " E, 02 – 07. ii. 2023, actinic light trap, leg. Bakala N., M., Dérozier, V., Kirk-Spriggs, A., László, G., gen. slide No.: LGNA 1392; 1 female, Nouabale-Ndoki National Park, Mbeli camp, 372 m, 02 ° 14 ' 23.8 " N, 16 ° 23 ' 52.1 " E, 14 – 20. ii. 2023, MV light trap, Bakala N., M., Dérozier, V., Kirk-Spriggs, A., László, G., gen. slide No.: ANHRT 02311; 1 female, same site, 1 – 10. x. 2022, leg. Dérozier, V., Fouka, B., Kirk-Spriggs, A., Takano, H., gen. slide No.: ANHRT 02221; 1 female, Nouabale-Ndoki NP, Makao camp (Secondary forest), 377 m, 2 ° 35 ' 42.2 " N, 17 ° 10 ' 08.3 " E, 15 – 21. v. 2023, MV light trap, leg. Dérozier, V., Kirk-Spriggs, A., László, G., Mvouende, S., gen. slide No.: LGNA 1410; 1 female, Nouabale-Ndoki National Park, Mondika camp, 365 m, 02 ° 21 ' 50.63 " N, 16 ° 16 ' 25.82 " E, 07 – 14. ii. 2023, MV light trap, leg. Bakala N., M., Dérozier, V., Kirk-Spriggs, A., László, G., gen. slide No.: LGNA 1399; 1 male, 1 female, Odzala-Kokoua National Park, Lobo Research Camp, 390 m, 00 ° 35 ' 04 " N, 14 ° 53 ' 12 " E, 20 – 27. ix. 2024, MV and LepiLED light trap, leg. Bashford, M., László, G., Volynkin, A., Yaba Ngouma, S., gen. slide No.: LGNA 1491; 1 male, Odzala-Kokoua National Park, Mbomo HQ, 570 m, 00 ° 26 ' 13 '' N, 14 ° 42 ' 01 '' E, 19 – 22. iv. 2024, MV light trap, leg. Bashford, M., László, G., Talani, M., Yaba Ngouma S., gen. slide No.: LGNA 1353. Sierra Leone. 1 male, Mansonia village at the foothills of Loma Mts, 420 m, 09 ° 07 ' 47 " N, 11 ° 05 ' 06 " W, 6. vi. 2016, light trap, leg. Takano, Miles & Goff, gen. slide No.: LGNA 461; 1 male, 120 m, Tiwai Island, Moa River, 07 ° 33 ' 00 " N, 11 ° 21 ' 09 " W, 17 – 22. vi. 2016, light trap, leg. Takano, Miles & Goff, gen. slide No.: LGNA 469. Genetic information. This taxon has been assigned the BIN URI: BOLD ABU 7264; BIN-sharing with M. bispermutata bispermutata. A single female specimen from Guinea (gen. slide No.: LGNA 1552), showing no morphological differences to other West African specimens, belongs to a different BIN URI: BOLD AFG 3904. The intraspecific divergence calculated from six samples (excluding the diverging specimen) is 0.00 – 0.31 %. This subspecies diverges from the nominotypical M. bispermutata by 0.61 – 1.31 %. The morphologically closest taxa are M. diagrapha and M. saotomeana sp. n., diverging by 2.82 – 2.99 % and 2.03 – 2.35 %, respectively. Taxonomic note. The western-central African populations of M. bispermutata exhibit identical genital morphology to those found in southern-eastern Africa. However, they can be distinguished by their somewhat smaller size and paler forewing markings observed in West African individuals. Due to these minor but consistent differences in external appearance, a West African subspecies, separate from the nominotypical M. bispermutata can be recognised. The name M. cryptochra Hacker, introduced for a species from Cameroon, can be applied to the western M. bispermutata populations: M. bispermutata cryptochra stat. n.	en	László, Gyula M. (2025): Integrative taxonomic review of the Meganola monofascia van Son species group sensu Hacker et al. (Lepidoptera, Noctuoidea, Nolidae, Nolinae, Nolini). Ecologica Montenegrina 87: 135-191, DOI: 10.37828/em.2025.87.7, URL: https://doi.org/10.37828/em.2025.87.7
038A879EFFB78C67FF67F950515CFFB5.taxon	distribution	Distribution. The western-central African M. cryptochra was earlier known from Cameroon (type locality) (Hacker et al. 2012). The specimens from Côte d’Ivoire, Guinea, Liberia, the Republic of Congo and Sierra Leone represent new country records.	en	László, Gyula M. (2025): Integrative taxonomic review of the Meganola monofascia van Son species group sensu Hacker et al. (Lepidoptera, Noctuoidea, Nolidae, Nolinae, Nolini). Ecologica Montenegrina 87: 135-191, DOI: 10.37828/em.2025.87.7, URL: https://doi.org/10.37828/em.2025.87.7
038A879EFFB58C64FF67FF5F5092FF4D.taxon	description	(Figs 26 – 29, 104 – 105, 149)	en	László, Gyula M. (2025): Integrative taxonomic review of the Meganola monofascia van Son species group sensu Hacker et al. (Lepidoptera, Noctuoidea, Nolidae, Nolinae, Nolini). Ecologica Montenegrina 87: 135-191, DOI: 10.37828/em.2025.87.7, URL: https://doi.org/10.37828/em.2025.87.7
038A879EFFB58C64FF67FF5F5092FF4D.taxon	materials_examined	Material examined (all in coll. ANHRT). Côte d’Ivoire. 2 males, 1 female, Mt. Tonkoui Peak, 1171 m, 07 ° 27 ' 15.2 " N, 07 ° 38 ' 12.5 " W, 19 – 27. xi. 2019, actinic, LepiLED and MV light trap, leg. Aristophanous, M., Dérozier, V., Moretto, P., Ouattara, S., gen. slide Nos: LGNA 1149, LGNA 1592 (males), LGNA 1566 (female). Guinea. 6 males, 4 females, Guinee Forestiere Monts Nimba UNESCO World Heritage Site, Mont Richard Molard camp and ridge (High-Altitude Grassland & Forest), 1400 – 1752 m, 07 ° 36 ' 19 " N, 08 ° 25 ' 30 " W, 1 – 7. vii. 2019, 250 W blended bulb light trap, leg. Dérozier, V., Miles, W., Sáfián, Sz., gen. slide Nos: LGNA 1289, LGNA 1547, LGNA 1565 (males), LGNA 1283, LGNA 1164, LGNA 1165, LGNA 1550 (females); 3 males, 1 female, Guinee Forestiere, Bossou Forest and Institut de Recherche Environmentale de Bossou (Lowland Forest-Farmland), 690 m, 07 ° 38 ' 32 " N, 08 ° 30 ' 30 " W, 24 – 30. vi. 2019, MV and 8 W UV Cold Cathode light trap, leg. Dérozier, V., Suah Dore, J., Koivagui, S., Miles, W., Sáfián, S., Warner, R.; 2 males, 1 female, 619 km ESE of Conakry, Nzerekore Region, Prefecture de Lola, Ziela env., 540 – 600 m, x. 2017, 7 ° 42 ' N, 8 ° 21 ' W, leg. local collectors, gen. slide No.: LGNA 1757 (female). Liberia. 883 m, Lofa County, Wologizi Mts, Ridge Camp 2, 8 ° 07 ' 20.79 " N, 9 ° 56 ' 50.75 " W, 22 – 30. xi. 2018, Cold Cathode UV (8 W) and blended bulb (250 W) light trap, leg. Sáfián, Sz., Simonics, G., gen. slide Nos: LGNA 1201 (male), LGNA 1282, LGNA 978 (females); 1 female, Nimba County, Nimba Mts, ENNR, Cellcom road, 750 m, 7 ° 33 ' 3.78 " N, 8 ° 31 ' 46.49 " W, 16 – 28. xii. 2018, Cold Cathode UV light trap (8 W), leg. Sáfián, Sz., Simonics, G. Genetic information. This species has been assigned the BIN URI: BOLD AAP 8748. The intraspecific divergence calculated from seven samples is 0.00 %. The morphologically closest taxa are M. saotomeana sp. n., M. bispermutata cryptochra, and M. bispermutata bispermutata, diverging by 2.02 – 2.19 %, 2.82 – 2.99 % and 2.81 – 3.34 %, respectively. The externally highly similar M. pachygrapha diverges by 4.42 – 4.77 %. Taxonomic notes. Meganola kakamega has been synonymized with M. bispermutata by László & Vetina (2019). However, the examination of extensive material from Central Africa suggests that the Kakamega population, which resembles M. bispermutata, is more likely to belong to the western-central African species M. diagrapha.	en	László, Gyula M. (2025): Integrative taxonomic review of the Meganola monofascia van Son species group sensu Hacker et al. (Lepidoptera, Noctuoidea, Nolidae, Nolinae, Nolini). Ecologica Montenegrina 87: 135-191, DOI: 10.37828/em.2025.87.7, URL: https://doi.org/10.37828/em.2025.87.7
038A879EFFB58C64FF67FF5F5092FF4D.taxon	distribution	Distribution. This species was earlier published from Burkina Faso (type locality) and Kenya- Kakamega (as M. kakamega) (Hacker et al. 2012). The specimens from Côte d’Ivoire, Guinea and Liberia represent new country records.	en	László, Gyula M. (2025): Integrative taxonomic review of the Meganola monofascia van Son species group sensu Hacker et al. (Lepidoptera, Noctuoidea, Nolidae, Nolinae, Nolini). Ecologica Montenegrina 87: 135-191, DOI: 10.37828/em.2025.87.7, URL: https://doi.org/10.37828/em.2025.87.7
038A879EFFB48C65FF67FF5F5060FDFC.taxon	description	(Figs 30 – 33, 106 – 107, 150)	en	László, Gyula M. (2025): Integrative taxonomic review of the Meganola monofascia van Son species group sensu Hacker et al. (Lepidoptera, Noctuoidea, Nolidae, Nolinae, Nolini). Ecologica Montenegrina 87: 135-191, DOI: 10.37828/em.2025.87.7, URL: https://doi.org/10.37828/em.2025.87.7
038A879EFFB48C65FF67FF5F5060FDFC.taxon	materials_examined	Holotype. Male, “ SAO TOME 1321 m / Bom Successo, Antenna / (plantation-dry scrub forest) / 0 ° 16 ' 31 " N, 6 ° 36 ' 14 " E / 22 - 25. i. 2018 MV Light Trap / Aristophanous, M., Lima, D., / Miles, W., Turner, Clive R. leg. / ANHRT: 2018.1 ” // “ ANHRTUK / 00043828 ” / “ SAMPLED FOR: / DNA (2022) ” // “ Gen. slide No. / LGN A 1605 ♂ / prep. by Gy. M. Laszlo ” (ANHRT). Paratypes (all in coll. ANHRT). São Tomé. 7 males, 7 females, with the same data as the holotype, gen. slide Nos: LGNA 1288, LGNA 1606 (females); 1 male, 1 female, Antenna, Bom Successo, 0 ° 16 ’ 33.7 ” N, 6 ° 36 ’ 19.7 ” E, 19. x. 2016, MV light trap, leg. Turner, C. R., Tasane, T., gen. slide No.: LGNA 405 (male); 1 male, same site and collectors, 20. x. 2016, gen. slide No.: LGNA 388; 2 males, same site and collectors, 29. x. 2016.	en	László, Gyula M. (2025): Integrative taxonomic review of the Meganola monofascia van Son species group sensu Hacker et al. (Lepidoptera, Noctuoidea, Nolidae, Nolinae, Nolini). Ecologica Montenegrina 87: 135-191, DOI: 10.37828/em.2025.87.7, URL: https://doi.org/10.37828/em.2025.87.7
038A879EFFB48C65FF67FF5F5060FDFC.taxon	diagnosis	Diagnosis. Forewing length 11 – 13 mm in males, 13 – 15 mm in females. Meganola saotomeana sp. n. is an island vicariant of M. diagrapha and easily distinguished by its larger size, notably darker and more extensive forewing markings, especially in terms of the more distinct postmedial line and the considerably broader and more contrasting subterminal and terminal lines. In the male genitalia, the clasping apparatus of both species has a nearly identical configuration, possessing a similarly short and pointed harpe; however, the valva of the new species is slightly more dilated postmedially compared to M. diagrapha. Similar to the other species of the M. bispermutata complex, the shape of the aedeagus bears the most distinctive diagnostic characters. In M. saotomeana sp. n., the aedeagus is 25 – 30 % longer with a more elongate and narrower distal section than in M. diagrapha. Compared to M. bispermutata, the clasping apparatuses are also nearly identical, being slightly broader in M. saotomeana sp. n., and the aedeagus of the new species is also longer (by ca. 20 %) with a noticeably longer coecum. In the female genitalia, the new species has a considerably smaller ostium bursae continuing in a notably narrower and longer distal sclerotised section of the ductus bursae compared to M. diagrapha. Furthermore, the narrow longitudinal scobinated field of the signum bursae is narrower and less sclerotised in the new species than in M. diagrapha. Compared to M. bispermutata, the new species is readily distinguished by the more rounded and markedly smaller ostium bursae, the longer distal sclerotisation of the ductus bursae, the considerably narrower and longer ductus bursae, the smaller corpus bursae and the much longer longitudinal scobinated field of the signum bursae.	en	László, Gyula M. (2025): Integrative taxonomic review of the Meganola monofascia van Son species group sensu Hacker et al. (Lepidoptera, Noctuoidea, Nolidae, Nolinae, Nolini). Ecologica Montenegrina 87: 135-191, DOI: 10.37828/em.2025.87.7, URL: https://doi.org/10.37828/em.2025.87.7
038A879EFFB48C65FF67FF5F5060FDFC.taxon	etymology	Etymology. This new species is named after its type locality, the island of São Tomé. Genetic information. Meganola saotomeana sp. n. has been assigned the BIN URI: BOLD AEG 4865. The intraspecific divergence calculated from four samples is 0.00 – 0.15 %. The closest relative is M. diagrapha diverging by 2.02 – 2.19 %.	en	László, Gyula M. (2025): Integrative taxonomic review of the Meganola monofascia van Son species group sensu Hacker et al. (Lepidoptera, Noctuoidea, Nolidae, Nolinae, Nolini). Ecologica Montenegrina 87: 135-191, DOI: 10.37828/em.2025.87.7, URL: https://doi.org/10.37828/em.2025.87.7
038A879EFFB48C6BFF67F910572DFA2D.taxon	description	(Figs 34 – 39, 108 – 110, 151)	en	László, Gyula M. (2025): Integrative taxonomic review of the Meganola monofascia van Son species group sensu Hacker et al. (Lepidoptera, Noctuoidea, Nolidae, Nolinae, Nolini). Ecologica Montenegrina 87: 135-191, DOI: 10.37828/em.2025.87.7, URL: https://doi.org/10.37828/em.2025.87.7
038A879EFFB48C6BFF67F910572DFA2D.taxon	materials_examined	Material examined (all in coll. ANHRT). Cameroon. 1 female, SW slope of Mount Cameroon, Elephant camp, 1091 m, 4 ° 07 ' 01.20 " N, 9 ° 04 ' 22.51 " E, 18 – 23. ii. 2017, light trap (105 W energy saving bulb), leg. Janecek, S., Mertens, J., Potocky. P., Sáfián, Sz., Tropek, R., gen. slide No.: LGNA 808. Côte d’Ivoire. Mt. Tonkoui Peak, 07 ° 27 ' 15.2 " N, 07 ° 38 ' 12.5 " W, 1171 m, 19 – 27. xi. 2019, actinic, LepiLED and MV light trap, leg. Aristophanous, M., Dérozier, V., Moretto, P., Ouattara, S., gen. slide Nos: LGNA 1782, LGNA 1549, LGNA 1290 (males); LGNA 1280, LGNA 1781 (females); 2 females, same site, 1 – 8. xi. 2015, leg. Aristophanous, M., Moretto, P., Ruzzier, E., gen. slide Nos: LGNA 86, LGNA 253. Guinea. Long series of both sexes, Guinee Forestiere Monts Nimba UNESCO World Heritage Site, Mont Richard Molard camp and ridge (High-Altitude Grassland & Forest), 1400 - 1752 m, 07 ° 36 ' 19 " N, 08 ° 25 ' 30 " W, 1 – 7. vii. 2019, 250 W Blended and LepiLED light trap, leg. Dérozier, V., Miles, W., Sáfián, Sz., gen. slide Nos: LGNA 1144, LGNA 1291, LGNA 1161 (males), LGNA 1276 (female); 4 females, Guinee Forestiere, Bossou Forest and Institut de Recherche Environmentale de Bossou (Lowland Forest-Farmland), 690 m, 07 ° 38 ' 32 " N, 08 ° 30 ' 30 " W, 24 – 30. vi. 2019, 8 W UV Cold Cathode, LepiLED and MV light trap, leg. Dérozier, V., Suah Dore, J., Koivagui, S., Miles, W., Sáfián, S., Warner, R., gen. slide Nos: LGNA 1286, LGNA 1287, LGNA 1548; 1 male, Geipa Camp, Foret de Diecke, 435 m, 7 ° 26 ' 7.06 " N, 8 ° 50 ' 47.87 " W, 05 – 14. iv. 2019, Cold Cathode UV Light Trap (8 W), leg. Sáfián, Sz., Koivogui, S., gen. slide No.: LGNA 975; 5 males, 10 females, 619 km ESE of Conakry, Nzerekore Region, Prefecture de Lola, Ziela env., 540 – 600 m, x. 2017, 7 ° 42 ' N, 8 ° 21 ' W, leg. local collectors. Kenya. 1 male, Gatamaiyo Forest, Aberdare Mountains, 2260 m, 0 ° 58 ' 26.2 " S, 36 ° 41 ' 49.03 " E, 22 – 23. vii. 2017, light trap (blended bulb 250 W), leg. Sáfián Sz., gen. slide No.: LGNA 806; 1 male, Western, Kakamega F. R. Prim. For., 0 ° 21.34 ' N, 34 ° 51.39 ' E, 1600 m, 15 Watt Black Light, 9 - I- 2002, F. N. Namu, gen. slide No.: LGNA 1762. Liberia. 1 male, Lofa County, Wologizi Mts, Ridge Camp, 865 m, 8 ° 07 ' 10 " N, 9 ° 57 ' 11 " W, 24 – 29. xi. 2017, light trap (blended bulb 250 W), leg. Aristophanous, M., Sáfián, Sz., Simonics, G. & Smith, L., gen. slide No.: LGNA 1168; 2 females, same site, 22 – 30. xi. 2018, Cold Cathode UV light trap (8 W), leg. Sáfián, Sz., Simonics, G., gen. slide No.: LGNA 977; 2 males, Mount Nimba, Cellcom Road, ENNR, Nimba County, 750 – 1100 m, 7 ° 32 ' 35.89 " N, 8 ° 31 ' 20.63 " W, 14 – 28. xii. 2017, Cold Cathode light bucket trap, leg. Sáfián, Sz., Simonics, G.; 1 female, Nimba County, Nimba Mts, ENNR, Cellcom road, 750 m, 7 ° 33 ' 3.78 " N, 8 ° 31 ' 46.49 " W, 16 – 28. xii. 2018, Cold Cathode UV light trap (8 W), leg. Sáfián, Sz., Simonics, G., gen. slide No.: LGNA 1281. Republic of Congo. 2 males, Nouabale-Ndoki NP, Makao camp (Secondary forest), 377 m, 2 ° 35 ' 42.2 " N, 17 ° 10 ' 08.3 " E, 15 – 21. v. 2023, MV light trap, leg. Dérozier, V., Kirk-Spriggs, A., László, G., Mvouende, S., gen. slide Nos: LGNA 1387, LGNA 1390; 1 male, Odzala-Kokoua National Park, Imbalanga Camp, 480 m, 00 ° 45 ' 47 " N, 15 ° 15 ' 39 " E, 05 – 09. iv. 2024, MV light trap, Bashford, M., László, G., Talani, M., Yaba Ngouma, S., gen. slide No.: LGNA 1396; 2 males, Odzala-Kokoua National Park, Kokoua base, 540 m, 00 ° 28 ' 39 " N, 15 ° 16 ' 42 " E, 05 – 13. ix. 2024, MV light trap, leg. Bashford, M., László, G., Talani, M., Volynkin, A, Yaba Ngouma, S., gen. slide No: LGNA 1777; 1 male, Odzala-Kokoua National Park, Lobo Research Camp, 390 m, 00 ° 35 ' 04 " N, 14 ° 53 ' 12 " E, 20 – 27. ix. 2024, MV light trap, leg. Bashford, M., László, G., Volynkin, A., gen. slide No.: LGNA 1776. Sierra Leone. 1 male, Mansonia village at the foothills of Loma Mts, 420 m, N 09 ° 07 ' 47 ", W 11 ° 05 ' 06 ", 6. vi. 2016, light trap, leg. Takano, Miles & Goff, gen. slide No.: LGNA 1251; 1 female, Kono Province, Gori Hills, Near Giehum, 373 m, 08 ° 27 ' 47 " N, 10 ° 46 ' 17 " W, 22 – 29. ii. 2020, leg. Kalnoi, G., Sinyaev, V. Uganda. 1 male, Bwindi Impenetrable N. P., Cuckooland Lodge, 1700 m, 01 ° 00 ' 17 " S, 29 ° 43 ' 06 " E, 28 – 29. vi. 2022, leg. Naumann, S., Ott, E., Schintlmeister, A., Sulak, H., gen. slide No.: LGNA 1718. Genetic information. This species belongs to the BIN URI: BOLD AAP 8746. The intraspecific divergence calculated from eight samples is 0.00 - 0.46 %. The morphologically closest taxa are M. bispermutata cryptochra, M. bispermutata bispermutata and M. saotomeana sp. n. diverging by 3.44 – 3.94 %, 3.76 – 4.10 % and 3.78 – 4.27 %, respectively. Taxonomic notes. Meganola pachygrapha is readily distinguished from the other members of the M. bispermutata complex by the conspicuously short and stout aedeagus of the male genitalia. This character is identical in M. murzinialis as illustrated in Hacker et al. (2012) and confirmed by dissections of topotypical specimens from the Odzala-Kokoua National Park. Since M. pachygrapha and M. murzinialis display no distinctive morphological characters, the latter is considered a junior synonym: M. murzinialis syn. n. There is multiple confusion regarding M. xantholeuca in Hacker et al. (2012: 498): The females illustrated as M. xantholeuca belong to M. pachygrapha. Additionally, the male holotype of M. xantholeuca is misclassified and actually belongs to the genus Ezishnola László, Ronkay & Witt, 2010. Furthermore, the holotype genitalia illustrated in the original description closely resemble those of M. nyungwe Hacker, 2012. These taxonomic issues will be addressed in detail in an upcoming paper on the genus Ezishnola (László, in prep.).	en	László, Gyula M. (2025): Integrative taxonomic review of the Meganola monofascia van Son species group sensu Hacker et al. (Lepidoptera, Noctuoidea, Nolidae, Nolinae, Nolini). Ecologica Montenegrina 87: 135-191, DOI: 10.37828/em.2025.87.7, URL: https://doi.org/10.37828/em.2025.87.7
038A879EFFB48C6BFF67F910572DFA2D.taxon	distribution	Distribution. Meganola pachygrapha is a widespread species with an equatorial distribution. It has been recorded from Ethiopia (type locality), Uganda, Kenya, South Sudan, Bioko Island (as M. pachygrapha), the Republic of Congo (as M. murzinialis), and Côte d’Ivoire (as M. xantholeuca) (Hacker et al. 2012). The specimens from Cameroon, Guinea, Liberia, and Sierra Leone represent new country records.	en	László, Gyula M. (2025): Integrative taxonomic review of the Meganola monofascia van Son species group sensu Hacker et al. (Lepidoptera, Noctuoidea, Nolidae, Nolinae, Nolini). Ecologica Montenegrina 87: 135-191, DOI: 10.37828/em.2025.87.7, URL: https://doi.org/10.37828/em.2025.87.7
038A879EFFBA8C6BFF67FDE550E5FE3C.taxon	description	(Figs 40 – 42, 111 – 112, 152)	en	László, Gyula M. (2025): Integrative taxonomic review of the Meganola monofascia van Son species group sensu Hacker et al. (Lepidoptera, Noctuoidea, Nolidae, Nolinae, Nolini). Ecologica Montenegrina 87: 135-191, DOI: 10.37828/em.2025.87.7, URL: https://doi.org/10.37828/em.2025.87.7
038A879EFFBA8C6BFF67FDE550E5FE3C.taxon	materials_examined	Material examined (all in coll. ANHRT). Côte d’Ivoire. 2 males, Denguele Classified Forest, 479 m, 09 ° 30 ' 0.6 " N, 07 ° 40 ' 51.1 " W, 11 – 18. xi. 2019, actinic light trap, leg. Aristophanous, M., Dérozier, V., Moretto, P., Outtara, S., gen. slide Nos: LGNA 1962, LGNA 1963. Guinea. 1 male, Nimba Mts, SMFG concession area, (Societe des Mines de Fer de Guinee), Cite 1, 700 m, 7 ° 42 ' 2.83 " N, 8 ° 23 ' 58.60 " W, 25. viii – 05. ix. 2017, general coll., leg. Sáfián Sz., Simonics, G., gen. slide No.: LGNA 1151; 1 female, Geipa Camp, Foret de Diecke, 435 m, 7 ° 26 ' 7.06 " N, 8 ° 50 ' 47.87 " W, 05 – 14. iv. 2019, light trap, blended bulb (250 W), leg. Sáfián, Sz., Koivogui, S., gen. slide No.: LGNA 988; 1 male, 2 females, 619 km ESE of Conakry, Nzerekore Region, Prefecture de Lola, Ziela env., 540 – 600 m, x. 2017, 7 ° 42 ' N, 8 ° 21 ' W, leg. local collectors, gen. slide No.: LGNA 1538 (male), LGNA 1696, LGNA 1754 (females). Genetic information. This species has been assigned the BIN URI: BOLD AEG 5922. The intraspecific divergence calculated from three samples is 0.15 – 0.31 %. The closest relative of M. dananae is M. baaka sp. n. with a PWD of 3.15 – 3.31 %. Meganola bispermutata bispermutata and M. bispermutata cryptochra diverge from M. dananae by 3.80 – 4.19 % and 3.80 – 4.28 %, respectively. Taxonomic note. Owing to the then-lack of DNA barcodes, László (2020) erroneously illustrated the male of M. pachygrapha as the male of M. dananae. The correct matching of the males and females of these species was clarified by the DNA barcode analyses during this study, and the confirmed male and female genitalia are illustrated under Figs 111 – 112, 152.	en	László, Gyula M. (2025): Integrative taxonomic review of the Meganola monofascia van Son species group sensu Hacker et al. (Lepidoptera, Noctuoidea, Nolidae, Nolinae, Nolini). Ecologica Montenegrina 87: 135-191, DOI: 10.37828/em.2025.87.7, URL: https://doi.org/10.37828/em.2025.87.7
038A879EFFBA8C6BFF67FDE550E5FE3C.taxon	distribution	Distribution. Meganola dananae is a west African species with confirmed records from Côte d’Ivoire (Hacker et al. 2012) and Guinea (present study).	en	László, Gyula M. (2025): Integrative taxonomic review of the Meganola monofascia van Son species group sensu Hacker et al. (Lepidoptera, Noctuoidea, Nolidae, Nolinae, Nolini). Ecologica Montenegrina 87: 135-191, DOI: 10.37828/em.2025.87.7, URL: https://doi.org/10.37828/em.2025.87.7
038A879EFFBA8C68FF67F91551BFFDBF.taxon	description	(Figs 43 – 46, 113 – 114, 153)	en	László, Gyula M. (2025): Integrative taxonomic review of the Meganola monofascia van Son species group sensu Hacker et al. (Lepidoptera, Noctuoidea, Nolidae, Nolinae, Nolini). Ecologica Montenegrina 87: 135-191, DOI: 10.37828/em.2025.87.7, URL: https://doi.org/10.37828/em.2025.87.7
038A879EFFBA8C68FF67F91551BFFDBF.taxon	materials_examined	Holotype. Male, “ GABON 430 m / Mikongo (Rougier), Monts / de Cristal (Secondary Forest) / 0 ° 29 ' 47 " N, 11 ° 10 ' 42 " E / 28. vii – 12. viii. 2019 MV LightTrap / Albert, J-L., Aristophanous, M., / Bie Mba, J., Dérozier, V., / Moretto, P. Leg. / ANHRT: 2019.17 ” // “ ANHRTUK / 00192260 ” / “ SAMPLED FOR: / DNA (2022) ” // “ Gen. slide No. / LGN A 1582 ♂ / prep. by Gy. M. Laszlo ” (ANHRT). Paratypes (all in coll. ANHRT). Republic of Congo. 1 male, Sangha Prov., Noubale-Ndoki National Park, Ndoki formation (Secondary forest), 352 m, 02 ° 12 ' 47.7 " N, 16 ° 23 ' 45.8 " E, 29. ix – 1. x. 2022, actinic light trap, leg. Dérozier, V., Fouka, B., Kirk-Spriggs, A., Takano, H., gen. slide No.: ANHRT 02216; 2 males, 3 females, Likouala Prov., Nouabale-Ndoki National Park, Makao forest (Secondary forest), 349 m, 02 ° 36 ' 42.5 " N, 17 ° 09 ' 523.8 " E, 24 – 29. ix. 2022, actinic and LepiLED light trap, leg. Dérozier, V., Fouka, B., Kirk-Spriggs, A., Takano, H., gen. slide Nos: LGNA 1318, ANHRT 02224 (males), LGNA 1384, LGNA 1389, ANHRT 2223 (females); 2 males, Nouabale-Ndoki National Park, Mbeli camp, 372 m, 02 ° 14 ' 23.8 " N, 16 ° 23 ' 52.1 " E, 14 – 20. ii. 2023, actinic light trap, leg. Bakala N., M., Dérozier, V., Kirk-Spriggs, A., László, G., gen. slide Nos: ANHRT 02239, ANHRT 02267; 7 males, 1 female, Odzala-Kokoua National Park, Kokoua base, 540 m, 00 ° 28 ' 39 " N, 15 ° 16 ' 42 " E, 05 – 13. ix. 2024, MV light trap, Bashford, M., László, G., Talani, M., Volynkin, A, Yaba Ngouma, S., gen. slide Nos: LGNA 1423, LGNA 1487, LGNA 1831 (males); 3 males, Odzala-Kokoua National Park, Mbomo HQ, 570 m, 00 ° 26 ' 13 '' N, 14 ° 42 ' 01 '' E, 28. ix – 01. x. 2024, MV light trap, leg. Bashford, M., László, G., Volynkin, A., gen. slide Nos: LGNA 1433, LGNA 1465, LGNA 1485; 1 female, Odzala-Kokoua National Park, Lobo Research Camp, 390 m, 00 ° 35 ' 04 " N, 14 ° 53 ' 12 " E, 25 – 26. ix. 2024, LepiLED light trap, leg. Bashford, M., László, G., Volynkin, A., Yaba Ngouma, S., gen. slide No.: LGNA 1490.	en	László, Gyula M. (2025): Integrative taxonomic review of the Meganola monofascia van Son species group sensu Hacker et al. (Lepidoptera, Noctuoidea, Nolidae, Nolinae, Nolini). Ecologica Montenegrina 87: 135-191, DOI: 10.37828/em.2025.87.7, URL: https://doi.org/10.37828/em.2025.87.7
038A879EFFBA8C68FF67F91551BFFDBF.taxon	diagnosis	Diagnosis. Forewing length 7 – 9 mm in males, 9 – 10 mm in females. Meganola baaka sp. n. is a central African allopatric sibling of the West African M. dananae, distinguished in the following external features: the new species has a notably more extensive and contrasting elongate-triangular costal patch and considerably darker terminal area of the forewing, which has a whitish grey sheen in contrast to the pale ochreous-grey ground colour of M. dananae. In the male genitalia, both species feature a narrow, apically rounded valva and a remarkably long, pointed harpe; however, the new species has a noticeably longer tegumen without an angled medial dilation as observed in M. dananae. The aedeagi of both species are similarly long, with an elongate distal section distinguishing them from the other taxa of the M. bispermutata complex, having markedly shorter aedeagi. In the female genitalia, M. baaka sp. n. has a somewhat shorter and broader anterior apophysis, a noticeably narrower, elongate-quadrangular ostium bursae (it is rounded, U-shaped in M. dananae), a narrower distal sclerotised section of the ductus bursae, and a shorter (by ca. 30 %) ductus bursae.	en	László, Gyula M. (2025): Integrative taxonomic review of the Meganola monofascia van Son species group sensu Hacker et al. (Lepidoptera, Noctuoidea, Nolidae, Nolinae, Nolini). Ecologica Montenegrina 87: 135-191, DOI: 10.37828/em.2025.87.7, URL: https://doi.org/10.37828/em.2025.87.7
038A879EFFBA8C68FF67F91551BFFDBF.taxon	etymology	Etymology. This new species is dedicated to the Baaka people, a major ethnic group in central Africa. Genetic information. Meganola baaka sp. n. belongs to the BIN URI: BOLD AAF 7471. As only one specimen was sequenced in this study, intraspecific divergence of the COI 5 P sequences could not be calculated. The species’ closest relative is M. dananae, diverging by 3.15 – 3.31 %.	en	László, Gyula M. (2025): Integrative taxonomic review of the Meganola monofascia van Son species group sensu Hacker et al. (Lepidoptera, Noctuoidea, Nolidae, Nolinae, Nolini). Ecologica Montenegrina 87: 135-191, DOI: 10.37828/em.2025.87.7, URL: https://doi.org/10.37828/em.2025.87.7
038A879EFFBA8C68FF67F91551BFFDBF.taxon	distribution	Distribution. Meganola baaka sp. n. is a central African vicariant of the West African M. dananae with confirmed records only from Gabon and the Republic of Congo.	en	László, Gyula M. (2025): Integrative taxonomic review of the Meganola monofascia van Son species group sensu Hacker et al. (Lepidoptera, Noctuoidea, Nolidae, Nolinae, Nolini). Ecologica Montenegrina 87: 135-191, DOI: 10.37828/em.2025.87.7, URL: https://doi.org/10.37828/em.2025.87.7
038A879EFFB98C68FF67F9945646FF76.taxon	description	Taxonomic note. The female paratype specimen illustrated by Hacker et al. (2012) is not conspecific with the male holotype of M. subderrae but related to M. microfascia Hacker, 2012.	en	László, Gyula M. (2025): Integrative taxonomic review of the Meganola monofascia van Son species group sensu Hacker et al. (Lepidoptera, Noctuoidea, Nolidae, Nolinae, Nolini). Ecologica Montenegrina 87: 135-191, DOI: 10.37828/em.2025.87.7, URL: https://doi.org/10.37828/em.2025.87.7
038A879EFFB98C68FF67F9945646FF76.taxon	distribution	Distribution. This species is known only from the Kakamega forest in Kenya.	en	László, Gyula M. (2025): Integrative taxonomic review of the Meganola monofascia van Son species group sensu Hacker et al. (Lepidoptera, Noctuoidea, Nolidae, Nolinae, Nolini). Ecologica Montenegrina 87: 135-191, DOI: 10.37828/em.2025.87.7, URL: https://doi.org/10.37828/em.2025.87.7
038A879EFFB88C69FF67FF5F5036FC8C.taxon	description	Several species in this complex are highly reminiscent of members of the M. monofascia-bispermutata species-groups, although often displaying more contrasting transverse lines. In the male genitalia, the valva is shorter and broader than in the M. bispermutata complex and apically evenly rounded, unlike in the M. monofascia species group. The harpe is considerably shorter and more triangular in the M. furvitincta species group than in the other complexes. The most characteristic feature of this species group is the well-developed, often excessively long, flat and straight laminar, or thorn-shaped and curved carina process, the length and shape of which is an important specific character. The shared female genitalia characters of the species group are the very short, rounded anterior apophysis, the well-developed, heavily sclerotised somewhat infundibular antrum and the small transverse signum bursae surrounded by an extensive ovoid field of fine longitudinal scobination continued in a narrow scobinate area towards the base of the ductus bursae. Taxon content	en	László, Gyula M. (2025): Integrative taxonomic review of the Meganola monofascia van Son species group sensu Hacker et al. (Lepidoptera, Noctuoidea, Nolidae, Nolinae, Nolini). Ecologica Montenegrina 87: 135-191, DOI: 10.37828/em.2025.87.7, URL: https://doi.org/10.37828/em.2025.87.7
038A879EFFB88C6EFF67FB44565AFEC9.taxon	description	(Figs 47 – 51, 115 – 117, 154)	en	László, Gyula M. (2025): Integrative taxonomic review of the Meganola monofascia van Son species group sensu Hacker et al. (Lepidoptera, Noctuoidea, Nolidae, Nolinae, Nolini). Ecologica Montenegrina 87: 135-191, DOI: 10.37828/em.2025.87.7, URL: https://doi.org/10.37828/em.2025.87.7
038A879EFFB88C6EFF67FB44565AFEC9.taxon	materials_examined	Material examined (all in coll. ANHRT). Côte d’Ivoire. 7 males, 4 females, Mt. Tonkoui Peak, 1171 m, 07 ° 27 ' 15.2 " N, 07 ° 38 ' 12.5 " W, 19 – 27. xi. 2019, actinic, LepiLED and MV light trap, leg. Aristophanous, M., Dérozier, V., Moretto, P., Ouattara, S., gen. slide Nos: LGNA 1586, LGNA 1587, LGNA 1590, LGNA 1703, LGNA 1964 (males), LGNA 1588, LGNA 1589 (females). Guinea. 2 males, Dalaba, Foret de Tinka, 1289 m, 10 ° 43 ' 14 " N, 12 ° 15 ' 22 " W, 25 – 28. ix. 2019, LepiLED light trap, leg. Geiser, M., Leno, M., Koivagui, S., Miles, W., Mulvaney, L., Sáfián, Sz., gen. slide Nos: LGNA 1243, LGNA 1250; 1 male, 619 km ESE of Conakry, Nzerekore Region, Prefecture de Lola, Ziela env., 540 – 600 m, x. 2017, 7 ° 42 ' N, 8 ° 21 ' W, leg. local collectors, gen. slide No.: LGNA 1794; 1 female, Seredou camp, Foret Classee de Ziama (upland forest), 872 m, 8 ° 21 ' 14.53 " N, 9 ° 19 ' 31.93 " W, 24. ii – 06. iii. 2019, general coll., leg. Sáfián, Sz., Simonics, G., Florczyk, K., Koivogui, S., gen. slide No.: LGNA 1604. Liberia. 1 male, Nimba Mts, ENNR, Nimba county (Cellcom road), 1000 – 1100 m, 7 ° 32 ' 45.88 " N, 8 ° 31 ' 21.04 " W, 02 – 14. xii. 2017, Cold Cathode light bucket, leg. Aristophanous, M., Sáfián, Sz., Simonics, G., Smith, L., gen. slide No.: LGNA 491; 2 males, Nimba Mts camp, ENNR, Nimba county, 1165 m, 7 ° 31 ' 45 " N, 8 ° 31 ' 37 " W, 03 – 13. xii. 2017, light trap (blended bulb 250 W), leg. Aristophanous, M., Sáfián, Sz., Simonics, G., Smith, L., gen. slide Nos: LGNA 560, LGNA 616; 1 male, 1 female, Nimba County, Nimba Mts, ENNR, Cellcom road, 750 m, 7 ° 33 ' 3.78 " N, 8 ° 31 ' 46.49 " W, 16 – 28. xii. 2018, Cold Cathode UV light trap (8 W), leg. Sáfián, Sz., Simonics, G., gen. slide Nos: LGNA 1244 (male), LGNA 973 (female); 1 male, Mount Nimba Main Ridge, ENNR, Nimba County, 1300 m, 7 ° 31 ' 1.53 " N, 8 ° 31 ' 1.24 " W, 14 – 28. xii. 2017, light trap blended bulb (250 W), leg. Sáfián, Sz., Simonics, G., gen. slide No.: LGNA 1591; 1 female, Nimba Mts, Mt. Gangra Summit, 978 m, 7 ° 32 ' 45.82 " N, 8 ° 38 ' 9.36 " W, 17 – 25. iii. 2017, Light Trap (250 w blended bulb) & cold cathode UV light bucket trap (8 w), leg. Sáfián, Sz., Simonics, G., gen. slide No.: LGNA 1274; 1 male, Nimba county, ENNR, Nimba Mts, main ridge (montane forest), 1327 m, 7 ° 31 ' 1.3 " N, 8 ° 31 ' 1.0 " W, 9. xii. 2017, light trap (blended bulb 250 W), leg. Aristophanous, M., Sáfián, Sz., Simonics, G., Smith, L.; 1 male, Lofa County, Wologizi Mts, Ridge camp, 865 m, 8 ° 07 ' 10 " N, 9 ° 57 ' 11 " W, 24 – 29. xi. 2017, light trap (blended bulb 250 W), leg. Aristophanous, M., Sáfián, Sz., Simonics, G., Smith, L.; 1 male, Lofa County, Foya Proposed Protected Area, 530 m, 7 ° 56 ' 36 " N, 10 ° 16 ' 36 " W, 10 – 19. xi. 2017, MV light trap (125 W), leg. Aristophanous, M., Sáfián, Sz., Simonics, G., Smith, L., gen. slide No.: LGNA 863. Republic of Congo. 1 male, Odzala-Kokoua National Park, Kokoua base, 540 m, 00 ° 28 ' 39 " N, 15 ° 16 ' 42 " E, 05 – 13. ix. 2024, MV light trap, leg. Bashford, M., László, G., Talani, M., Volynkin, A, Yaba Ngouma, S., gen. slide No.: LGNA 1774. Taxonomic notes. The genitalia illustrations of M. furvitincta presented in Hacker et al. (2012) are incorrect, as they were not based on examination of primary types. This discrepancy was addressed in László (2020), who designated and dissected the lectotype of this species. The male genitalia shown in Figures E and F in Hacker et al. (2012) actually belong to M. monofascia, while the female genitalia depicted in Figure G represent an undescribed species, which will be discussed below. Additionally, Figure H illustrates the copulatory organ of M. monofascia, although it is incorrectly referred to as M. bispermutata by László (2020). Following the clarification of the male genital morphology, genetic analyses conclusively identified the female of M. furvitincta (see Text fig. 2 and Table 1). The genitalia illustration of the female holotype of the externally similar M. spherographa Hacker, 2012 displays the same features as those observed in M. furvitincta. Therefore, M. spherographa syn. n. is considered conspecific with M. furvitincta. However, there is, indeed, a second, closely related species that occurs sympatrically with M. furvitincta in West Africa. The male genitalia described under M. spherographa (Hacker et al. 2012: 449, figs D, E) actually belong to this other species, which is described below as new to science. Genetic information. Despite their distinctive morphology, M. furvitincta furvitincta, M. furvitincta poliographa, M. spherographata sp. n. and M. aarviki all belong to the same BIN URI: BOLD AAP 3225. The intraspecific divergence of M. furvitincta furvitincta (n = 13) ranges from 0.00 % to 0.46 %. The PWD between M. furvitincta furvitincta and M. furvitincta poliographa, M. spherographata sp. n. and M. aarviki are 0.11 – 1.24 %, 0.46 – 2.19 % and 1.86 – 2.02 %, respectively. Additionally, close relatives such as M. fontainei and M. trigonographa diverge by 2.24 – 2.84 % and 2.51 – 2.82 %, respectively.	en	László, Gyula M. (2025): Integrative taxonomic review of the Meganola monofascia van Son species group sensu Hacker et al. (Lepidoptera, Noctuoidea, Nolidae, Nolinae, Nolini). Ecologica Montenegrina 87: 135-191, DOI: 10.37828/em.2025.87.7, URL: https://doi.org/10.37828/em.2025.87.7
038A879EFFB88C6EFF67FB44565AFEC9.taxon	distribution	Distribution. The nominotypical subspecies of M. furvitincta is a western-central African taxon with confirmed records from Guinea, Côte d’Ivoire, Liberia, the Republic of Congo (present research), Sierra Leone (type locality) and Nigeria (as M. spherographa syn. n. (Hacker et al. 2012). The specimens identified as M. furvitincta in Hacker et al. (2012) and Hacker (2014) have been misidentified; apart from the syntype, the illustrated specimens actually belong to M. monofascia.	en	László, Gyula M. (2025): Integrative taxonomic review of the Meganola monofascia van Son species group sensu Hacker et al. (Lepidoptera, Noctuoidea, Nolidae, Nolinae, Nolini). Ecologica Montenegrina 87: 135-191, DOI: 10.37828/em.2025.87.7, URL: https://doi.org/10.37828/em.2025.87.7
038A879EFFBF8C6CFF67F88956B4F923.taxon	description	(Figs 52 – 55, 118 – 119, 155)	en	László, Gyula M. (2025): Integrative taxonomic review of the Meganola monofascia van Son species group sensu Hacker et al. (Lepidoptera, Noctuoidea, Nolidae, Nolinae, Nolini). Ecologica Montenegrina 87: 135-191, DOI: 10.37828/em.2025.87.7, URL: https://doi.org/10.37828/em.2025.87.7
038A879EFFBF8C6CFF67F88956B4F923.taxon	materials_examined	Material examined (all in coll. ANHRT). Malawi. 2 males, Mzuzu Wildlife Sanctuary, (Afromontane forest), 1294 m, 11 ° 27 ' 30.2 " S, 34 ° 03 ' 11.3 " E, 22 – 27. vi. 2022, LepiLED and MV light trap, leg. Kirk-Spriggs, A., Miles, W., gen. slide Nos: LGNA 1788, LGNA 1902. Uganda. 2 females, Bwindi Impenetrable N. P., Cuckooland Lodge, 1700 m, 01 ° 00 ' 17 " S, 29 ° 43 ' 06 " E, 28 – 29. vi. 2022, leg. Naumann, S., Ott, E., Schintlmeister, A., Sulak, H., gen. slide Nos: LGNA 1717, LGNA 1720. Zambia. 2 females, Zambezi Rapids, (Miombo / Riverine forest mosaic), 1205 m, S 11 ° 7 ' 30 ", E 24 ° 11 ' 6 ", 4 – 9. xi. 2018, MV light trap, leg. Aristophanous, M., Dérozier, V., László, G., Oram, D., gen. slide Nos: LGNA 758, LGNA 1009; 1 male, Kalene Hill (Miombo woodland), 1440 m, S 11 ° 11 ' 11 ", E 24 ° 12 ' 5 ", 6. xi. 2018, actinic light trap, leg. Aristophanous, M., Dérozier, V., László, G., Oram, D., gen. slide No.: LGNA 744; 3 males, Hillwood, Ikelenge (Miombo / Riverine forest mosaic), 1400 m, S 11 ° 16 ' 02 ", E 24 ° 18 ' 59 ', 23 – 30. xi. 2019, actinic and MV light trap, leg. Bashford, M., Miles, W., Mulvaney, L., Smith, R.; 1 male, same site, 28. v – 4. vi. 2020, leg. Chizuwa, D., Choongo, W.; 2 males, Nyangombe Falls (Miombo / Riverine forest mosaic), 1300 m, 11 ° 48 ' 25 " S, 24 ° 32 ' 12 " E, 17 – 23. xi. 2019, actinic and MV light trap, leg. Bashford, M., Miles, W., Mulvaney, L., Smith, R.; 1 male, Chitunta Plain (Miombo / Dambo mosaic), 1396 m, 11 ° 29 ' 12 " S, 24 ° 24 ' 18 " E, 29. xi – 4. xii. 2019, actinic light trap, leg. Bashford, M., Miles, W., Mulvaney, L.; 1 female, Jiwundu Swamp (Miombo / Riverine forest mosaic), 1340 m, S 11 ° 51 ' 54 ", E 25 ° 33 ' 20 ", 29. x – 4. xi. 2018, MV light trap, leg. Aristophanous, M., Dérozier, V., László, G., Oram, D.; 1 male, Lyangu, Liuwa Plain NP, 1035 m, 14 ° 46 ' 51 " S, 22 ° 34 ' 44 ", 12 – 15. xi. 2013, light trap, leg. Smith, Takano & Oram, gen. slide No.: LGNA 22; 1 male, Lunzua Falls, 20 km S of Mpulungu, Northern Province, 1416 m, 08 ° 55 ' 38 " S, 31 ° 09 ' 31 " E, 11 – 16. v. 2019, MV light trap, Dérozier, V., László, G., Miles, W.; 1 female, Kundalila Falls, 12 km SE of Kanona, Central Province, 1500 m, 13 ° 09 ' 17 " S, 30 ° 42 ' 07 " E, 26 – 27. iv. 2019, actinic light trap, Dérozier, V., László, G., Miles, W., gen. slide No.: LGNA 1011; 1 male, Kapishya Hot Springs, Shiwa N'gandu Estate, 1437 m, S 11 ° 10 ' 13 ", E 31 ° 36 ' 00 ", i – iii. 2017, M. T. Harvey coll., leg. Oram, D., Miles, W., Smith, L., gen. slide No.: LGNA 743; 1 female, Chilambwe Falls, Kafubu River, 1420 m, 09 ° 50 ' 13 " S, 30 ° 43 ' 35 " E, 8 – 12. ii. 2019, MV light trap, leg. Dérozier, V., Mulvaney, L., Smith, R., Takano, H., gen. slide No.: LGNA 934; 1 female, Mutinondo Wilderness Area, Mpika, Northern Prov., 1460 m, 12 ° 27 ' 06 " S, 31 ° 17 ' 30 " E, 16 – 20. iii. 2017, MV, leg. Oram, D., Miles, W., Smith, L.; 9 males, 1 female, Choma, Bruce-Miller Farm Nansai (Southern Miombo woodland), 1179 m, 16 ° 38 ' 32 " S, 27 ° 01 ' 47 " E, 25. iv – 1. v. 2022, MV light trap, leg. Miles, W., Mulvaney, L.; 1 male, 2 females, Danger Hill, 30 km N of Mpika, Muchinga Province, 1684 m, 11 ° 37 ' 38 " S, 31 ° 33 ' 56 " E, 27 – 30. iv. 2019, actinic and MV light trap, Dérozier, V., László, G., Miles, W., gen. slide Nos: LGNA 1010 (male), LGNA 1034 (female); 7 males, 3 females, same site, 13 – 15. xii. 2023, leg. László, G., Morgan, L., Volynkin, A., gen. slide Nos: LGNA 1972 (male), LGNA 1707, LGNA 1970, LGNA 1971 (female). Genetic information. This species has been assigned the BIN URI: BOLD AAP 3225; BIN-sharing with M. furvitincta furvitincta, M. spherographata sp. n. and M. aarviki. The intraspecific divergence of M. furvitincta poliographa (n = 5) ranges from 0.00 % to 1.52 %. It diverges from M. furvitincta furvitincta, M. spherographata sp. n. and M. aarviki by 0.11 – 1.24 %, 0.00 – 2.51 % and 2.17 %, respectively. Additionally, M. furvitincta poliographa shows a divergence from M. trigonographa and M. fontainei of 2.35 – 3.15 % and 2.82 – 3.53 %, respectively. Taxonomic notes. László (2020) synonymised this taxon with M. furvitincta. However, an examination of several specimens resembling M. furvitincta from southern-central Africa revealed subtle but consistent differences in external and genital morphology between the two species. Externally, M. poliographa has much darker and more contrasting forewing markings than M. furvitincta. In the male genitalia, the former taxon has a slightly curved and ca. 20 – 30 % shorter flap-like laminar projection of the carina than that of M. furvitincta; in addition, the dorsal margin of the distal section of the aedeagus is smooth in M. furvitincta poliographa, whilst it is slightly serrated in the nominotypical taxon. Based on these diagnostic morphological characters, the name poliographa has been reinstated as a subspecies to represent the southern-central African populations of M. furvitincta: M. furvitincta poliographa stat. rev., stat. n.	en	László, Gyula M. (2025): Integrative taxonomic review of the Meganola monofascia van Son species group sensu Hacker et al. (Lepidoptera, Noctuoidea, Nolidae, Nolinae, Nolini). Ecologica Montenegrina 87: 135-191, DOI: 10.37828/em.2025.87.7, URL: https://doi.org/10.37828/em.2025.87.7
038A879EFFBF8C6CFF67F88956B4F923.taxon	distribution	Distribution. Meganola furvitincta poliographa is a southern-central African taxon expanding northward along the Rift valley, up to Uganda (type locality). The subspecies has further confirmed records from Zambia (as M. mbala (Hacker et al. 2014 )) and Malawi (present study).	en	László, Gyula M. (2025): Integrative taxonomic review of the Meganola monofascia van Son species group sensu Hacker et al. (Lepidoptera, Noctuoidea, Nolidae, Nolinae, Nolini). Ecologica Montenegrina 87: 135-191, DOI: 10.37828/em.2025.87.7, URL: https://doi.org/10.37828/em.2025.87.7
038A879EFFBD8C6CFF67FEF751EFFBBC.taxon	description	Taxonomic note. The validity of this taxon was examined by László (2020), who suggested that it may only be a pale form of M. furvitincta. However, a thorough examination of the M. furvitincta complex indicates that this taxon is valid and probably confined to the Usambara Mountains. This mountain range is part of the Eastern Arc Mountains, which are known for their exceptional diversity and endemism (Rodgers & Homewood 1982). Although the clasping apparatus of both subspecies of M. furvitincta and M. usambarana is identical, the latter species features a distinctly narrower and approximately 20 % longer aedeagus in the male genitalia. Additionally, the female genitalia of M. usambarana are notably different from those of M. furvitincta, characterized by a funnel-shaped antrum that is about 1.5 times longer and slightly curved. Based on these unique morphological traits, M. usambarana is upheld as a valid species. However, specimens identified under this taxon in Hacker et al. (2012) from Rwanda, Uganda, and DRC (Lubumbashi) belong to M. furvitincta poliographa.	en	László, Gyula M. (2025): Integrative taxonomic review of the Meganola monofascia van Son species group sensu Hacker et al. (Lepidoptera, Noctuoidea, Nolidae, Nolinae, Nolini). Ecologica Montenegrina 87: 135-191, DOI: 10.37828/em.2025.87.7, URL: https://doi.org/10.37828/em.2025.87.7
038A879EFFBD8C6DFF67FBD756A2FFDB.taxon	description	(Figs 56 – 60, 120 – 122, 156)	en	László, Gyula M. (2025): Integrative taxonomic review of the Meganola monofascia van Son species group sensu Hacker et al. (Lepidoptera, Noctuoidea, Nolidae, Nolinae, Nolini). Ecologica Montenegrina 87: 135-191, DOI: 10.37828/em.2025.87.7, URL: https://doi.org/10.37828/em.2025.87.7
038A879EFFBD8C6DFF67FBD756A2FFDB.taxon	materials_examined	Holotype. Male, “ GABON 430 m / Mikongo (Rougier), Monts / de Cristal (Secondary Forest) / 0 ° 29 ' 47 " N, 11 ° 10 ' 42 " E / 28. vii- 12. viii. 2019 MV LightTrap / Albert, J-L., Aristophanous, M., / Bie Mba, J., Dérozier, V., / Moretto, P. Leg. / ANHRT: 2019.17 ” // “ ANHRTUK / 00165572 ” // “ SAMPLED FOR: / DNA (2022) ” // “ Gen. slide No. / LGN A 1576 ♂ / prep. by Gy. M. Laszlo ” (ANHRT). Paratypes (all in coll. ANHRT). Côte d’Ivoire. 2 males, Tai NP., Tai Research Station, 174 m, 05 ° 49 ' 59.8 " N, 07 ° 20 ' 32.0 W, 14 – 23. xi. 2015, light trap, leg. Aristophanous, M., Moretto, P., Ruzzier, E., gen. slide Nos: LGNA 144, LGNA 228. Gabon. 1 male, same site and collectors as in the holotype, but collected at LepiLED light trap, gen. slide No.: LGNA 1579; 1 female, Nyonie (Lowland forest), 10 m, 0 ° 2 ' 22 " S, 9 ° 20 ' 25 " E, 23 – 28. viii. 2019, MV light trap, leg. Albert, J-L., Aristophanous, M., Bie Mba, J., Dérozier, V., Moretto, P., gen. slide No.: LGNA 1155. Liberia. 1 male, 1 female, Lofa County, Foya Proposed Protected Area, 530 m, 7 ° 56 ' 36 " N, 10 ° 16 ' 36 " W, 10 – 19. xi. 2017, MV light trap (125 W), leg. Aristophanous, M., Sáfián, Sz., Simonics, G. & Smith, L., gen. slide Nos: LGNA 861 (male), LGNA 539 (female); 1 male, 2 females, Lofa County, Wologizi Mts, base camp forest, 611 m, 8 ° 07 ' 17 " N, 9 ° 57 ' 42 " W, 20. xi – 01. xii. 2017, MV light trap (125 W), leg. Aristophanous, M., Sáfián, Sz., Simonics, G., Smith, L., gen. slide No: LGNA 1176 (male), LGNA 587, LGNA 864 (females); 4 females, Lofa County, 8 ° 07 ' 10 " N, 9 ° 57 ' 11 " W, 865 m, 24 – 29. xi. 2017, light trap (blended bulb 250 W) and Cold Cathode light bucket, leg. Aristophanous, M., Sáfián, Sz., Simonics, G. & Smith, L., gen. slide Nos: LGNA 541, LGNA 542, LGNA 1167, LGNA 1166; 1 female, Nimba Mts, ENNR, Nimba county (Cellcom road), 1000 – 1100 m, 7 ° 32 ' 45.88 " N, 8 ° 31 ' 21.04 " W, 16 – 28. xii. 2018, Cold Cathode UV Light Trap (8 W), leg. Sáfián, Sz., Simonics, G., gen. slide No.: LGNA 1284.	en	László, Gyula M. (2025): Integrative taxonomic review of the Meganola monofascia van Son species group sensu Hacker et al. (Lepidoptera, Noctuoidea, Nolidae, Nolinae, Nolini). Ecologica Montenegrina 87: 135-191, DOI: 10.37828/em.2025.87.7, URL: https://doi.org/10.37828/em.2025.87.7
038A879EFFBD8C6DFF67FBD756A2FFDB.taxon	diagnosis	Diagnosis. Forewing length 7 – 8 mm in males, 9 – 11 in females. Meganola spherographata sp. n. is reminiscent externally of M. furvitincta furvitincta, but it can be distinguished by the lighter greyish forewing ground colour, where the medial area has only a large costal patch including a rounded discal spot, whilst in M. furvitincta, a larger proportion of the medial area displays a darker colouration. The postmedial line in the new species is finer, but often more contrasting, and the terminal forewing area is less darkened in M. spherographata sp. n. than in its allied species. As the two species occur sympatrically in western and central Africa, accurate identification often necessitates genitalia dissection. In the male genitalia, the clasping apparatus of M. spherographata sp. n. has a somewhat shorter valva, a noticeably narrower and more elongate harpe, and a shorter vinculum. The most distinct differences between this new species and M. furvitincta are expressed in the configuration of the aedeagus, where M. spherographata sp. n. has a considerably narrower and apically tapering flap-like carinal process, which does not reach over the tip of the aedeagus, whilst in both subspecies of M. furvitincta, the apically dilated carinal process extends well over the tip of the aedeagus. The length of the carinal process is nearly identical in both species; however, the post-carinal distal section of the aedeagus in M. spherographata sp. n. is ca. twice as long as in M. furvitincta. In the female genitalia, the antrum is noticeably shorter and narrower with nearly straight lateral margins and a deeper, semicircular distal concavity marking the ostium bursae in M. spherographata sp. n. compared to the considerably larger, goblet-shaped antrum with a V-shaped notch of the ostium bursae in M. furvitincta. Further differences between the two species are the shorter, medially more constricted eighth tergite, the slightly shorter and broader anterior apophysis, the ca. 10 % longer tubular section of the corpus bursae and the less extensive scobinated plate of the signum bursae of M. spherographata sp. n. compared to its congener. Meganola spherographata sp. n. also closely resembles M. aarviki, but the latter species can be distinguished by its creamy sheen of the forewing, in contrast to the matte grey ground colour of the new species. In the male genitalia, compared to M. spherographata sp. n., M. aarviki has a ca. 10 % shorter aedeagus with a markedly shorter flap-like carinal process that ends well before the tip of the aedeagus and has a sharply truncate tip, unlike the tapered-rounded tip of the carinal process found in M. spherographata sp. n. The configuration of the female genitalia is nearly identical in M. spherographata sp. n. and M. aarviki; however, the latter has considerably larger anterior lobes of the eighth sternite and a noticeably shorter antrum.	en	László, Gyula M. (2025): Integrative taxonomic review of the Meganola monofascia van Son species group sensu Hacker et al. (Lepidoptera, Noctuoidea, Nolidae, Nolinae, Nolini). Ecologica Montenegrina 87: 135-191, DOI: 10.37828/em.2025.87.7, URL: https://doi.org/10.37828/em.2025.87.7
038A879EFFBD8C6DFF67FBD756A2FFDB.taxon	etymology	Etymology. The specific epithet is a modified form of “ spherographa ”, an adjective of Greek origin, referring to the species M. spherographa, which has sunk into synonymy with M. furvitincta due to the designation of a misidentified female holotype (Hacker et al. 2012). Genetic information. Meganola spherographata sp. n. has been assigned the BIN URI: BOLD AAP 3225. The species is BIN-sharing with M. furvitincta furvitincta, M. furvitincta poliographa and M. aarviki, diverging by 0.46 – 2.19 %, 0.00 – 2.51 % and 0.00 – 2.33 %, respectively. The intraspecific divergence of the COI 5 P sequences calculated from seven samples is 0.00 – 2.33 %. This species is also closely related to M. fontainei and M. trigonographa, diverging by 2.57 – 3.31 % and 2.67 – 3.15 %, respectively. Taxonomic note. This species was illustrated by Hacker et al. (2012) as M. spherographa Hacker, 2012. However, since Hacker designated a female holotype for this taxon which has been proven to be conspecific with M. furvitincta (as mentioned above), M. spherographa syn. n. was placed in synonymy with the former taxon. Consequently, the otherwise distinct species required a new description based on a male holotype, along with a correct matching of male and female sexes through DNA barcode analyses.	en	László, Gyula M. (2025): Integrative taxonomic review of the Meganola monofascia van Son species group sensu Hacker et al. (Lepidoptera, Noctuoidea, Nolidae, Nolinae, Nolini). Ecologica Montenegrina 87: 135-191, DOI: 10.37828/em.2025.87.7, URL: https://doi.org/10.37828/em.2025.87.7
038A879EFFBD8C6DFF67FBD756A2FFDB.taxon	distribution	Distribution. A male specimen of this new species was previously identified as M. spherographa syn. n. from Nigeria (Hacker et al. 2012). In the current study, the species has been confirmed in Gabon, Côte d’Ivoire, and Liberia. Additionally, the female specimens illustrated in Hacker et al. (2012) as M. spherographa syn. n. from Nigeria and Sierra Leone actually belong to M. furvitincta.	en	László, Gyula M. (2025): Integrative taxonomic review of the Meganola monofascia van Son species group sensu Hacker et al. (Lepidoptera, Noctuoidea, Nolidae, Nolinae, Nolini). Ecologica Montenegrina 87: 135-191, DOI: 10.37828/em.2025.87.7, URL: https://doi.org/10.37828/em.2025.87.7
038A879EFF838C52FF67FF5F537CFECA.taxon	description	(Figs 61 – 64, 123 – 125, 157)	en	László, Gyula M. (2025): Integrative taxonomic review of the Meganola monofascia van Son species group sensu Hacker et al. (Lepidoptera, Noctuoidea, Nolidae, Nolinae, Nolini). Ecologica Montenegrina 87: 135-191, DOI: 10.37828/em.2025.87.7, URL: https://doi.org/10.37828/em.2025.87.7
038A879EFF838C52FF67FF5F537CFECA.taxon	materials_examined	Material examined (all in coll. ANHRT). Côte d’Ivoire. 1 male, Tai NP. Tai Research Station (SRET), 174 m, 05 ° 50 ' 00 " N, 07 ° 20 ' 32.0 W, 25. iii – 17. iv. 2017, MV light trap, leg. Aristophanous, A., Aristophanous, M., Geiser, M., Moretto, P., gen. slide No.: LGNA 980. Republic of Congo. Long series of both sexes from the following localities: Sangha Prov., Nouabale-Ndoki National Park, Mbeli camp, (Gilbertiodendron forest), 372 m, 02 ° 14 ' 23.8 " N, 16 ° 23 ' 52.1 " E, (1 – 10. x. 2022, leg. Dérozier, V., Fouka, B., Kirk-Spriggs, A., Takano, H., 14 – 20. ii. 2023, leg. Bakala, N. M., Dérozier, V., Kirk-Spriggs, A., László, G.); Nouabale-Ndoki National Park, Bomassa camp, 341 m, 02 ° 12 ' 36.9 " N, 16 ° 11 ' 30.2 " E, (20 – 21. ii. 2023, leg. Bakala, N. M., Dérozier, V., Kirk-Spriggs, A., László, G., 16 – 23. ix. 2022, leg. Dérozier, V., Fouka, B., Kirk-Spriggs, A., Takano, H.); Nouabale-Ndoki National Park, Mondika camp, 365 m, 02 ° 21 ' 50.63 " N, 16 ° 16 ' 25.82 " E, (07 – 14. ii. 2023, leg. Bakala N., M., Dérozier, V., Kirk-Spriggs, A., László, G., 27. iv – 06. v. 2023, leg. Dérozier, V., Kirk-Spriggs, A., László, G., Mvouende, S.); Sangha Prov., Noubale-Ndoki National Park, Ndoki formation (Secondary forest), 352 m, 02 ° 12 ' 47.7 " N, 16 ° 23 ' 45.8 " E, 29. ix – 1. x. 2022, leg. Dérozier, V., Fouka, B., Kirk-Spriggs, A., Takano, H.; Odzala-Kokoua National Park, Imbalanga Camp, 480 m, 00 ° 45 ' 47 " N, 15 ° 15 ' 39 " E, 05 – 11. iv. 2024, leg. Bashford, M., László, G., Talani, M., Yaba Ngouma, S.; Odzala-Kokoua National Park, Mbomo HQ, 570 m, 00 ° 26 ' 13 '' N, 14 ° 42 ' 01 '' E, (19 – 22. iv. 2024, leg. Bashford, M., László, G., Talani, M., Yaba Ngouma S., 28. ix – 01. x. 2024, leg. Bashford, M., László, G., Talani, M., Volynkin, A., Yaba Ngouma S.), gen. slide Nos: ANHRT 02257, ANHRT 2513, ANHRT 02524, ANHRT 02544, ANHRT 02547, ANHRT 02557, ANHRT 02574, ANHRT 02577, ANHRT 02576, ANHRT 02579, ANHRT 02580, ANHRT 02588, ANHRT 02218, ANHRT 02219, ANHRT 02220, ANHRT 02230, ANHRT 02247, ANHRT 02231, ANHRT 02258, ANHRT 02260, ANHRT 02263, ANHRT 02264, ANHRT 02265, ANHRT 02266, ANHRT 02268, ANHRT 02269, ANHRT 02270, ANHRT 02299, ANHRT 02304, ANHRT 03228, ANHRT 03256, ANHRT 03260, ANHRT 03276, ANHRT 03279, ANHRT 03289, LGNA 1355, LGNA 1357, LGNA 1359, LGNA 1378, LGNA 1379, LGNA 1380, LGNA 1391, LGNA 1825 (males), ANHRT 02539, ANHRT 02483, ANHRT 02556, ANHRT 02589, ANHRT 02552, ANHRT 02310, LGNA 1382 (females). Genetic information. Meganola aarviki has been assigned the BIN URI: BOLD AAP 3225, and the species is BIN-sharing with both subspecies of M. furvitincta and M. spherographata sp. n. As only one sample was included in the genetic analyses, the intraspecific divergence could not be calculated. The PWD from M. furvitincta furvitincta, M. furvitincta poliographa and M. spherographata sp. n. is 1.86 – 2.02 %, 2.17 % and 0.00 – 2.33 %, respectively. The pairwise genetic distances between this species and M. fontainei and M. trigonographa are 2.90 – 3.21 % and 2.99 – 3.29 %, respectively.	en	László, Gyula M. (2025): Integrative taxonomic review of the Meganola monofascia van Son species group sensu Hacker et al. (Lepidoptera, Noctuoidea, Nolidae, Nolinae, Nolini). Ecologica Montenegrina 87: 135-191, DOI: 10.37828/em.2025.87.7, URL: https://doi.org/10.37828/em.2025.87.7
038A879EFF838C52FF67FF5F537CFECA.taxon	distribution	Distribution. The species was described from Kisangani (DRC). A female paratype from Odzala National Park, illustrated in Hacker et al. (2012), belongs to M. trigonographa Hacker, 2012. However, the identity of a second paratype specimen from Kenya, which is not illustrated, remains uncertain. This species is relatively abundant in northern Congo, where a long series was collected in the Nouabale-Ndoki and Odzala-Kokoua National Parks. Additionally, a single, somewhat worn male specimen from Tai National Park in Côte d’Ivoire can be confidently identified as M. aarviki based on its genital characteristics.	en	László, Gyula M. (2025): Integrative taxonomic review of the Meganola monofascia van Son species group sensu Hacker et al. (Lepidoptera, Noctuoidea, Nolidae, Nolinae, Nolini). Ecologica Montenegrina 87: 135-191, DOI: 10.37828/em.2025.87.7, URL: https://doi.org/10.37828/em.2025.87.7
038A879EFF838C53FF67F88F5020F9CB.taxon	description	(Figs 65 – 66, 126 – 127)	en	László, Gyula M. (2025): Integrative taxonomic review of the Meganola monofascia van Son species group sensu Hacker et al. (Lepidoptera, Noctuoidea, Nolidae, Nolinae, Nolini). Ecologica Montenegrina 87: 135-191, DOI: 10.37828/em.2025.87.7, URL: https://doi.org/10.37828/em.2025.87.7
038A879EFF838C53FF67F88F5020F9CB.taxon	materials_examined	Material examined (all in coll. ANHRT). Zambia. 2 males, Muchinga Prov., Benyanga village, 1250 m, 10 ° 40 ' 41 " S, 33 ° 27 ' 45 " E, 07 – 12. xii. 2023, MV light trap, leg. László, G., Morgan, L., Volynkin, A., gen. slide Nos: LGNA 1673, LGNA 1967. Genetic information. This species has not been sequenced in this study.	en	László, Gyula M. (2025): Integrative taxonomic review of the Meganola monofascia van Son species group sensu Hacker et al. (Lepidoptera, Noctuoidea, Nolidae, Nolinae, Nolini). Ecologica Montenegrina 87: 135-191, DOI: 10.37828/em.2025.87.7, URL: https://doi.org/10.37828/em.2025.87.7
038A879EFF838C53FF67F88F5020F9CB.taxon	distribution	Distribution. Meganola stigmatographa was previously known only from the vicinity of Songea in southwestern Tanzania. The specimens collected in a Miombo woodland in northeastern Zambia represent a new country record.	en	László, Gyula M. (2025): Integrative taxonomic review of the Meganola monofascia van Son species group sensu Hacker et al. (Lepidoptera, Noctuoidea, Nolidae, Nolinae, Nolini). Ecologica Montenegrina 87: 135-191, DOI: 10.37828/em.2025.87.7, URL: https://doi.org/10.37828/em.2025.87.7
038A879EFF828C50FF67FD885197FBC1.taxon	description	(Figs 67 – 70, 128 – 129, 158 – 160)	en	László, Gyula M. (2025): Integrative taxonomic review of the Meganola monofascia van Son species group sensu Hacker et al. (Lepidoptera, Noctuoidea, Nolidae, Nolinae, Nolini). Ecologica Montenegrina 87: 135-191, DOI: 10.37828/em.2025.87.7, URL: https://doi.org/10.37828/em.2025.87.7
038A879EFF828C50FF67FD885197FBC1.taxon	materials_examined	Material examined (all in coll. ANHRT). Côte d’Ivoire. Long series of both sexes, Mt. Tonkoui Peak, 1171 m, 07 ° 27 ' 15.2 " N, 07 ° 38 ' 12.5 " W, 19 – 27. xi. 2019, actinic, LepiLED and MV light trap, leg. Aristophanous, M., Dérozier, V., Moretto, P., Ouattara, S., gen. slide Nos: LGNA 1205, LGNA 1536, LGNA 1702 (males), LGNA 1169, LGNA 1206, LGNA 1577, LGNA 1578, LGNA 1704 (females); long series of both sexes from the same site, 1 – 8. xi. 2015, light trap, leg. Aristophanous, M., Moretto, P., Ruzzier, E., gen. slide Nos: LGNA 39, LGNA 41, LGNA 148, LGNA 166, LGNA 235, LGNA 236, LGNA 238 (males), LGNA 40, LGNA 87, LGNA 150, LGNA 165, LGNA 167, LGNA 168, LGNA 240, LGNA 252, LGNA 254 (females); 1 female, same site and collectors, 12 – 18. viii. 2015, gen. slide No.: LGNA 24; 2 males, Tai NP. Tai Research Station, 174 m, 05 ° 49 ' 59.8 " N, 07 ° 20 ' 32.0 W, 14 – 23. xi. 2015, light trap, leg. Aristophanous, M., Moretto, P., Ruzzier, E., gen. slide Nos: LGNA 227, LGNA 234; 3 males, 2 females, same site, 25. iii – 17. iv. 2017, leg. Aristophanous, A., Aristophanous, M., Geiser, M., Moretto, P., gen. slide Nos: LGNA 394, LGNA 398 (males), LGNA 1285, LGNA 983 (PT of M. taiana László, 2020) (females); 1 male, Denguele Classified Forest, 479 m, 09 ° 30 ' 0.6 " N, 07 ° 40 ' 51.1 " W, 11 – 18. xi. 2019, actinic light trap, leg. Aristophanous, M., Dérozier, V., Moretto, P., Outtara, S. Gabon. 1 female, Mikongo, (Rougier), Monts de Cristal (Anthropised Forest), 430 m, 0 ° 29 ' 47 " N, 11 ° 10 ' 42 " E, 28. vii – 12. viii. 2019, actinic light trap, leg. Albert, J-L., Aristophanous, M., Bie Mba, J., Dérozier, V., Moretto, P. Guinea. 1 male, Guineé Forestiére Monts Nimba UNESCO World Heritage Site, Mont Richard Molard camp and ridge (High-Altitude Grassland & Forest), 1400 – 1752 m, 07 ° 36 ' 19 " N, 08 ° 25 ' 30 " W, 1 – 7. vii. 2019, 250 W blended light trap, leg. Dérozier, V., Miles, W., Sáfián, S.; 1 male, Nimba Mts, SMFG concession area (Societe des Mines de Fer de Guinee) 600 Forest, 1500 m, 7 ° 39 ' 49.93 " N, 8 ° 22 ' 22.19 " W, 21 – 30. viii. 2017, Cold Cathode UV light trap (8 W), Sáfián, Sz., Simonics, G., gen. slide No.: LGNA 831; 2 females, Seredou camp, Foret Classee de Ziama (upland forest), 872 m, 8 ° 21 ' 14.53 " N, 9 ° 19 ' 31.93 " W, 24. ii – 06. iii. 2019, general coll., leg. Sáfián, Sz., Simonics, G., Florczyk, K., Koivogui, S. Liberia. 6 males, 1 female, Nimba Mts camp, ENNR, Nimba county, 1165 m, 7 ° 31 ' 45 " N, 8 ° 31 ' 37 " W, 03 – 13. xii. 2017, light trap (blended bulb 250 W), leg. Aristophanous, M., Sáfián, Sz., Simonics, G., Smith, L., gen. slide Nos: LGNA 544, LGNA 478 (males); 1 male, 1 female, Nimba county, ENNR, Nimba Mts, main ridge (montane forest), 1327 m, 7 ° 31 ' 1.3 " N, 8 ° 31 ' 1.0 " W, 9. xii. 2017, light trap (blended bulb 250 W), leg. Aristophanous, M., Sáfián, Sz., Simonics, G., Smith, L., gen. slide No.: LGNA 538; 2 males, 2 females, ENNR, Nimba Mts, Cellcom road, little stream, 1250 – 1350 m, 7 ° 31 ' 28 " N, 8 ° 30 ' 39.9 " W, 14 – 15. iii. 2017, 250 W blended bulb and 8 W Cold Cathode light bucket trap, leg. Sáfián, Sz., Simonics, G.; 1 female, Mount Nimba Main Ridge, ENNR, Nimba County, 1300 m, 7 ° 31 ' 1.53 " N, 8 ° 31 ' 1.24 " W, 14 – 28. xii. 2017, light trap blended bulb (250 W), leg. Sáfián, Sz., Simonics, G., gen. slide No.: LGNA 1700; 2 females, same site and collectors, 31. iii – 1. iv. 2017; 2 males, 1000 – 1100 m, Nimba Mts, ENNR, Nimba county, Cellcom road, 7 ° 32 ' 45.88 " N, 8 ° 31 ' 21.04 " W, 02 – 14. xii. 2017, Cold Cathode light bucket, leg. Aristophanous, M., Sáfián, Sz., Simonics, G., Smith, L.; 1 male, Nimba County, Nimba Mts, ENNR, Cellcom road, 750 m, 7 ° 33 ' 3.78 " N, 8 ° 31 ' 46.49 " W, 16 – 28. xii. 2018 Cold Cathode UV Light Trap (8 W), Sáfián, Sz., Simonics, G.; 1 male, Nimba Mts, Mt. Gangra Summit, 978 m, 7 ° 32 ' 45.82 " N, 8 ° 38 ' 9.36 " W, 17 – 25. iii. 2017, light trap (250 w blended bulb) & cold cathode UV light bucket trap (8 w), leg. Sáfián, Sz., Simonics, G.; 3 males, Lofa County, Wologizi Mts, base camp forest, 611 m, 8 ° 07 ' 17 " N, 9 ° 57 ' 42 " W, 20. xi – 01. xii. 2017, MV light trap (125 W), leg. Aristophanous, M., Sáfián, Sz., Simonics, G. & Smith, L.; 1 female, Lofa County, Zuwulor Village School, 230 m, 7 ° 54 ' 52 " N, 9 ° 31 ' 08 " W, 8. xi. 2017, light trap (blended bulb 250 W), leg. Aristophanous, M., Sáfián, Sz., Simonics, G. & Smith, L., gen. slide No.: LGNA 543; 2 males, Krahn-Bassa Reserve, Sinoe County, Juboe River, 7.5 km South West Pellokon Town, 140 m, 5 ° 39 ' 4 " N, 8 ° 39 ' 4 " W, 14 – 21. i. 2018, light trap (blended bulb 250 W), leg. Geiser, M., Sáfián, Sz., Simonics, G. Republic of Congo. 1 male, Nouabale-Ndoki National Park, Mombongo camp (Moist mixed forest), 352 m, 02 ° 10 ' 30.7 " N, 16 ° 8 ' 37.7 " E, 28. vi. 2022, MV light trap, leg. Dérozier, V., gen. slide No.: ANHRT 02232; 2 males, 1 female, Odzala-Kokoua National Park, Kokoua base, 540 m, 00 ° 28 ' 39 " N, 15 ° 16 ' 42 " E, 05 – 13. ix. 2024, MV light trap, leg. Bashford, M., László, G., Talani, M., Volynkin, A., gen. slide Nos: LGNA 1486, LGNA 1775 (males), LGNA 1778 (female). Tanzania. 1 female, Mizimu, Mwanihana, Udzungwa Mountains N. P., 850 m, 07 ° 48 ' 21.8 " S, 36 ° 51 ' 09.5 " E, 3 – 6. viii. 2010, light trap leg. Smith, R. & Takano, H., gen. slide No.: LGNA 259. Genetic information. Meganola palaeographa has been assigned the BIN URI: BOLD ABW 4980. The intraspecific divergence calculated from 13 samples is 0.00 – 0.73 %. The genetically closest species are M. fontainei, M. trigonographa and M. dicranographa diverging by 2.71 – 3.47 %, 2.66 – 3.75 % and 3.30 – 3.80 %, respectively.	en	László, Gyula M. (2025): Integrative taxonomic review of the Meganola monofascia van Son species group sensu Hacker et al. (Lepidoptera, Noctuoidea, Nolidae, Nolinae, Nolini). Ecologica Montenegrina 87: 135-191, DOI: 10.37828/em.2025.87.7, URL: https://doi.org/10.37828/em.2025.87.7
038A879EFF828C50FF67FD885197FBC1.taxon	distribution	Distribution. This species was initially known only from the Udzungwa Mountains in Tanzania (Hacker et al. 2012, Hacker 2014). László (2020) reported records of the species in West Africa, including Côte d’Ivoire and Liberia. In the current study, M. palaeographa was also found in Gabon, Guinea, and the Republic of Congo, indicating a wide equatorial distribution.	en	László, Gyula M. (2025): Integrative taxonomic review of the Meganola monofascia van Son species group sensu Hacker et al. (Lepidoptera, Noctuoidea, Nolidae, Nolinae, Nolini). Ecologica Montenegrina 87: 135-191, DOI: 10.37828/em.2025.87.7, URL: https://doi.org/10.37828/em.2025.87.7
038A879EFF818C51FF67FB9250CBF923.taxon	description	(Figs 71 – 74, 130 – 131, 161 – 162)	en	László, Gyula M. (2025): Integrative taxonomic review of the Meganola monofascia van Son species group sensu Hacker et al. (Lepidoptera, Noctuoidea, Nolidae, Nolinae, Nolini). Ecologica Montenegrina 87: 135-191, DOI: 10.37828/em.2025.87.7, URL: https://doi.org/10.37828/em.2025.87.7
038A879EFF818C51FF67FB9250CBF923.taxon	materials_examined	Material examined (all in coll. ANHRT). Cameroon. 2 males, 3 females, North Region, Wack (La Falaise), 900 m, 07 ° 40 ' 16.5 " N, 13 ° 33 ' 18.4 " E, 2 – 21. x. 2018, Cold Cathode UV and LepiLED light trap, leg. Sáfián, Sz., Simonics, G., gen. slide Nos: LGNA 1159 (male), LGNA 1160 (female). Côte d’Ivoire. 2 males, 3 females, Denguélé Classified Forest, 479 m, 09 ° 30 ' 0.6 " N, 07 ° 40 ' 51.1 " W, 11 – 18. xi. 2019, actinic and MV light trap, leg. Aristophanous, M., Dérozier, V., Moretto, P., Outtara, S., gen. slide Nos: LGNA 1204 (male), LGNA 1246, LGNA 1275 (females). Zambia. 1 female, Zambezi Rapids (Miombo / Riverine forest mosaic), 1205 m, 11 ° 7 ' 30 " S, 24 ° 11 ' 6 " E, 4 – 9. xi. 2018, actinic light trap, leg. Aristophanous, M., Dérozier, V., László, G., Oram, D., gen. slide No.: LGNA 1966. Genetic information. This species has been assigned the BIN URI: BOLD AEG 3087; BIN-sharing with M. fontainei. The intraspecific divergence calculated from six samples is 0.00 – 0.16 %. The genetically closest species are M. fontainei, M. trigonographa and M. palaeographa, diverging by 0.82 – 1.10 %, 1.49 – 1.86 % and 3.30 – 3.80 %, respectively. Taxonomic note. This study has identified the male and female sexes of M. dicranographa through DNA barcode analyses. It has been revealed that the true female of this species is conspecific with M. parastrigula Hacker, 2012 (which was previously known only from female specimens). Furthermore, the female previously illustrated as M. dicranographa actually belongs to M. trigonographa Hacker, 2012. As a result, M. parastrigula syn. n. is hereby synonymized with M. dicranographa.	en	László, Gyula M. (2025): Integrative taxonomic review of the Meganola monofascia van Son species group sensu Hacker et al. (Lepidoptera, Noctuoidea, Nolidae, Nolinae, Nolini). Ecologica Montenegrina 87: 135-191, DOI: 10.37828/em.2025.87.7, URL: https://doi.org/10.37828/em.2025.87.7
038A879EFF818C51FF67FB9250CBF923.taxon	distribution	Distribution. This species was previously known from Burkina Faso (type locality) and Nigeria (also as M. parastrigula) (Hacker et al. 2012). The current study confirmed the presence of M. dicranographa in Cameroon, Côte d’Ivoire, and NW Zambia.	en	László, Gyula M. (2025): Integrative taxonomic review of the Meganola monofascia van Son species group sensu Hacker et al. (Lepidoptera, Noctuoidea, Nolidae, Nolinae, Nolini). Ecologica Montenegrina 87: 135-191, DOI: 10.37828/em.2025.87.7, URL: https://doi.org/10.37828/em.2025.87.7
038A879EFF808C56FF67FEF75662FA52.taxon	description	(Figs 75 – 80, 132 – 134, 163)	en	László, Gyula M. (2025): Integrative taxonomic review of the Meganola monofascia van Son species group sensu Hacker et al. (Lepidoptera, Noctuoidea, Nolidae, Nolinae, Nolini). Ecologica Montenegrina 87: 135-191, DOI: 10.37828/em.2025.87.7, URL: https://doi.org/10.37828/em.2025.87.7
038A879EFF808C56FF67FEF75662FA52.taxon	materials_examined	Material examined (all in coll. ANHRT). Gabon. Long series of both sexes, Mikongo (Rougier), Monts de Cristal (Secondary Forest), 430 m, 0 ° 29 ' 47 " N, 11 ° 10 ' 42 " E, 28. vii – 12. viii. 2019, actinic, LepiLED and MV light trap, leg. Albert, J-L., Aristophanous, M., Bie Mba, J., Dérozier, V., Moretto, P., gen. slide Nos: LGNA 1575, LGNA 1580, LGNA 1855 (males), LGNA 1574, LGNA 1581, LGNA 1584, LGNA 1872 (females). Republic of Congo. 1 male, Odzala-Kokoua National Park, Kokoua base, 540 m, 00 ° 28 ' 39 " N, 15 ° 16 ' 42 " E, 05 – 13. ix. 2024, MV light trap, leg. Bashford, M., László, G., Talani, M., Volynkin, A, Yaba Ngouma, S., gen. slide No.: LGNA 1772; 1 male, Odzala-Kokoua National Park, Mbomo HQ, 570 m, 00 ° 26 ' 13 '' N, 14 ° 42 ' 01 '' E, 28. ix – 01. x. 2024, MV light trap, leg. Bashford, M., László, G., Volynkin, A., gen. slide No.: LGNA 1492, LGNA 1506. Zambia. Long series of both sexes, Kasanka River Pontoon, Kasanka N. P., 1191 m, 12 ° 34 ' 23 " S, 30 ° 14 ' 05 " E, 2 – 4. xii. 2012, light trap, leg. Smith, R., Takano, H.; Kabwe, Kasanka N. P., 1187 m, 12 ° 32 ' 28 " S, 30 ° 12 ' 42 " E, 14 – 17. v. 2013, light trap, leg. Smith, Takano & Oram; Chisimbe Falls, Kasama, 1320 m, 10 ° 06 ' 31 " S, 30 ° 55 ' 02 " E, 27 – 29. iv. 2013, light trap, leg. Smith, Takano & Oram; Lukulu River, Lavushi Manda N. P., 1285 m, 12 ° 15 ' 05 " S, 30 ° 53 ' 43 " E, 27 – 29. xi. 2012, light trap, leg. Smith, R., Takano, H.; Kalene Hill (Miombo woodland), 1440 m, 11 ° 11 ' 11 " S, 24 ° 12 ' 5 " E, 6. xi. 2018, actinic light trap, leg. Aristophanous, M., Dérozier, V., László, G., Oram, D.; Lungu Forest Reserve, 20 km N of Mbala, Northern Province, 1452 m, 08 ° 42 ’ 27 ” S, 31 ° 16 ’ 34 ” E, 6 – 9. v. 2019, actinic light trap, leg. Dérozier, V., László, G., Miles, W.; Nyangombe Falls (Miombo / Riverine forest mosaic), 1300 m, 11 ° 48 ' 25 " S, 24 ° 32 ' 12 " E, 15 – 17. xi. 2018, actinic light trap, leg. Aristophanous, M., Dérozier, V., László, G., Oram, D.; Ikelenge, Hillwood, (Miombo / Riverine forest mosaic), 1400 m, 11 ° 16 ' 02 " S, 24 ° 18 ' 59 ” E, 5 – 11. vi. 2021, MV light trap, leg. Chizuwa, D., Choongo, W.; Kalungu, North of Isoka, 1280 m, 9 ° 40 ' 52 " S, 32 ° 42 ' 50 " E, 30. iv – 1. v. 2019, actinic light trap, leg. Dérozier, V., László, G., Miles, W.; Jiwundu Swamp (Miombo / Riverine forest mosaic), 1340 m, 11 ° 51 ' 54 " S, 25 ° 33 ' 20 " E, 29. x – 4. xi. 2018, actinic light trap, leg. Aristophanous, M., Dérozier, V., László, G., Oram, D.; Bruce-Miller Farm Nansai (Southern miombo woodland), 1179 m, 16 ° 38 ' 32 " S, 27 ° 01 ' 47 " E, 25. iv – 1. v. 2022, MV light trap, leg. Miles, W., Mulvaney, L.; Muchinga Province, 30 km N of Mpika, Danger Hill, 1684 m, 11 ° 37 ' 38 " S, 31 ° 33 ' 56 " E, 13 – 15. xii. 2023, MV light trap, leg. László, G., Morgan, L., Volynkin, A.; Chitunta Plain (Miombo / Dambo mosaic), 1396 m, 11 ° 29 ' 12 " S, 24 ° 24 ' 18 " E, 29. xi – 4. xii. 2019, actinic light trap, leg. Bashford, M., Miles, W., Mulvaney, L.; Senka Hill, Mukulizi Forest Reserve, Muchinga Province, 1566 m, 09 ° 05 ' 43 " S, 32 ° 05 ' 06 " E, 1 – 6. v. 2019, MV light trap, leg. Dérozier, V., László, G., Miles, W.; Zambezi Rapids (Miombo / Riverine forest mosaic), 1205 m, 11 ° 7 ' 30 " S, 24 ° 11 ' 6 " E, 4 – 9. xi. 2018, MV light trap, leg. Aristophanous, M., Dérozier, V., László, G., Oram, D., gen. slide Nos: LGNA 1, 2, 4, 9, 10, 11, 16, 75, 76, 77, 693, 754, 1031, 1593, LGNA 1975, LGNA 1976, LGNA 1977, LGNA 1978, LGNA 1979, LGNA 1982, LGNA 1983, (males), LGNA 3, 12, 13, 15, 62, 71, 72, 73, 74, 114, 262, 266, 1981, LGNA 1984 (females). Genetic information. Meganola fontainei has been assigned the BIN URI: BOLD AEG 3087, BINsharing with M. dicranographa. The intraspecific divergence calculated from 13 samples is 0.00 – 0.16 %. The genetically closest taxa are M. dicranographa, M. trigonographa and the M. furvitincta-spherographata sp. n. species pair, diverging by 0.82 – 1.10 %, 1.26 – 1.73 % and 2.40 – 3.04 %, respectively. The pairwise distance between M. fontainei and M. palaeographa is in the range of 2.71 – 3.47 %. Taxonomic note. Hacker et al. (2012) illustrated a female specimen and its genitalia from Côte d’Ivoire. However, László (2020) expressed doubts about the accuracy of this specimen's locality, noting that no other records of M. fontainei have been found in West Africa. After closely examining the image of the female genitalia presented by Hacker et al. (2012: 450, fig. G), it is clear that the inverse trapezoidal shape of the antrum, with its straight margins, is characteristic of M. palaeographa. In contrast, M. fontainei is known to have arcuate antrum margins. This suggests that the record of M. fontainei from Côte d’Ivoire is a misidentification and likely refers to M. palaeographa instead.	en	László, Gyula M. (2025): Integrative taxonomic review of the Meganola monofascia van Son species group sensu Hacker et al. (Lepidoptera, Noctuoidea, Nolidae, Nolinae, Nolini). Ecologica Montenegrina 87: 135-191, DOI: 10.37828/em.2025.87.7, URL: https://doi.org/10.37828/em.2025.87.7
038A879EFF808C56FF67FEF75662FA52.taxon	distribution	Distribution. Confirmed records of M. fontainei have been documented from DRC-Lubumbashi (type locality), the Republic of Congo, Burundi (Hacker et al. 2012), and Zambia (Hacker 2014). The specimens reported from Gabon in the current study represent a new country record. The species exhibits a broad distribution throughout central Africa, extending to the Zambian plateau.	en	László, Gyula M. (2025): Integrative taxonomic review of the Meganola monofascia van Son species group sensu Hacker et al. (Lepidoptera, Noctuoidea, Nolidae, Nolinae, Nolini). Ecologica Montenegrina 87: 135-191, DOI: 10.37828/em.2025.87.7, URL: https://doi.org/10.37828/em.2025.87.7
038A879EFF878C57FF67FD0753BDF9E9.taxon	description	(Figs 81 – 86, 135 – 137, 165)	en	László, Gyula M. (2025): Integrative taxonomic review of the Meganola monofascia van Son species group sensu Hacker et al. (Lepidoptera, Noctuoidea, Nolidae, Nolinae, Nolini). Ecologica Montenegrina 87: 135-191, DOI: 10.37828/em.2025.87.7, URL: https://doi.org/10.37828/em.2025.87.7
038A879EFF878C57FF67FD0753BDF9E9.taxon	materials_examined	Material examined (all in coll. ANHRT). Côte d’Ivoire. 1 male, Tai NP, Tai Research Station (SRET), 174 m, 05 ° 50 ' 00 " N, 07 ° 20 ' 32.0 " W, 25. iii – 17. iv. 2017, MV Light Trap, Aristophanous, A., Aristophanous, M., Geiser, M., Moretto, P. Gabon. 1 male, Nyonie (Lowland forest), 10 m, 0 ° 2 ' 22 " S, 9 ° 20 ' 25 " E, 23 – 28. viii. 2019, MV light trap, Albert, J-L., Aristophanous, M., Bie Mba, J., Dérozier, V., Moretto, P., gen. slide No.: LGNA 1170. Liberia. 1 male, 1 female, Nimba Mts, ENNR, Nimba county (Cellcom road), 1000 – 1100 m, 7 ° 32 ' 45.88 " N, 8 ° 31 ' 21.04 " W, 02 – 14. xii. 2017, Cold Cathode light bucket, leg. Aristophanous, M., Sáfián, Sz., Simonics, G., Smith, L., gen. slide No.: LGNA 536 (male); 2 males, 2 females, Nimba Mts camp, ENNR, Nimba county, 1165 m, 7 ° 31 ' 45 " N, 8 ° 31 ' 37 " W, 03 – 13. xii. 2017, MV (125 W) and blended bulb (250 W) light trap, leg. Aristophanous, M., Sáfián, Sz., Simonics, G., Smith, L., gen. slide Nos: LGNA 862, LGNA 477 (males), LGNA 588, LGNA 1245 (females); 1 male, ENNR, Nimba Mts, Cellcom road, little stream, 1250 – 1350 m, 7 ° 31 ' 28 " N, 8 ° 30 ' 39.9 " W, 14 – 15. iii. 2017, 250 W blended bulb and 8 W Cold Cathode light bucket trap, leg. Sáfián, Sz., Simonics, G.; 1 male, Lofa County, Wologizi Mts, Ridge Camp, 883 m, 2, 8 ° 07 ' 20.79 " N, 9 ° 56 ' 50.75 " W, 22 – 30. xi. 2018, Light Trap (blended bulb 250 W), Sáfián, Sz., Simonics, G.; 1 male, Lofa County, Wologizi Mts, base camp forest, 611 m, 8 ° 07 ' 17 " N, 9 ° 57 ' 42 " W, 20. xi – 01. xii. 2017, MV light trap (125 W), leg. Aristophanous, M., Sáfián, Sz., Simonics, G. & Smith, L., gen. slide No.: LGNA 537. Republic of Congo. 1 male, Nouabale-Ndoki National Park, Mombongo camp (Moist mixed forest), 352 m, 02 ° 10 ' 30.7 " N, 16 ° 8 ' 37.7 " E, 28. vi. 2022, MV light trap, leg. Dérozier, V., gen. slide No.: ANHRT 02233; 1 female, Nouabale-Ndoki National Park, Mondika camp, 365 m, 02 ° 21 ' 50.63 " N, 16 ° 16 ' 25.82 " E, 07 – 14. ii. 2023, MV light trap, Bakala N., M., Dérozier, V., Kirk-Spriggs, A., László, G., gen. slide No.: ANHRT 02538; 1 female, Odzala-Kokoua National Park, Lobo Research Camp, 390 m, 00 ° 35 ' 04 " N, 14 ° 53 ' 12 " E, 12 – 19. iv. 2024, actinic light trap, Bashford, M., László, G., Talani, M., Yaba Ngouma, S., gen. slide No.: LGNA 1692. Zambia. 1 male, 1400 m, Hillwood, Ikelenge, S 11 ° 16 ' 02 ", E 24 ° 18 ' 59 ", 17 – 24. iii. 2013, light trap, leg. Smith, R., Takano, H., gen. slide No.: LGNA 14; 1 male, same site and collectors, 19 – 21. viii. 2013, gen. slide No.: LGNA 115; 1 male, 1 female, same site, 9 – 16. v. 2015, leg. Smith, R., Takano, H., Aristophanous, M., gen. slide No.: LGNA 753 (male); 2 males, 2 females, same site, 5 – 11. vi. 2021, leg. Chizuwa, D., Choongo, W., gen. slide Nos: LGNA 1708, LGNA 1709 (males); 1 male, same site and collectors, 28. v – 4. vi. 2020, gen. slide No.: LGNA 1974; 1 male, same site, 23 – 30. xi. 2019, leg. Bashford, M., Miles, W., Mulvaney, L., Smith, R., gen. slide No: 1701; 3 females, Nkwaji, Mwinilunga, 1316 m, 11 ° 36 ' 22 " S, 24 ° 33 ' 17 " E, 21 – 22. viii. 2013, light trap, leg. Smith, R. & Takano, H., gen. slide Nos: LGNA 173, LGNA 265, LGNA 267; 1 male, Zambezi Rapids (Miombo / Riverine forest mosaic), 1205 m, 11 ° 7 ' 30 " S, 24 ° 11 ' 6 " E, 4 – 9. xi. 2018, actinic light trap, leg. Aristophanous, M., Dérozier, V., László, G., Oram, D., gen. slide No.: LGNA 1965. Genetic information. This species has been assigned the BIN URI: BOLD AAQ 3396. The intraspecific divergence calculated from eight samples is 0.00 – 0.31 %. The genetically closest taxa are M. fontainei (PWD = 1.26 – 1.73 %), M. dicranographa (PWD = 1.49 – 1.86 %) and the M. furvitincta-spherographata sp. n. species pair (PWD = 2.49 – 3.29 %).	en	László, Gyula M. (2025): Integrative taxonomic review of the Meganola monofascia van Son species group sensu Hacker et al. (Lepidoptera, Noctuoidea, Nolidae, Nolinae, Nolini). Ecologica Montenegrina 87: 135-191, DOI: 10.37828/em.2025.87.7, URL: https://doi.org/10.37828/em.2025.87.7
038A879EFF878C57FF67FD0753BDF9E9.taxon	distribution	Distribution. Meganola trigonographa was previously known only from Burkina Faso (Hacker et al. 2012). The current research indicates that this species is more widespread, extending from West Africa to southern Central Africa, with confirmed records from Côte d'Ivoire, Liberia, Gabon, the Republic of Congo, and Zambia.	en	László, Gyula M. (2025): Integrative taxonomic review of the Meganola monofascia van Son species group sensu Hacker et al. (Lepidoptera, Noctuoidea, Nolidae, Nolinae, Nolini). Ecologica Montenegrina 87: 135-191, DOI: 10.37828/em.2025.87.7, URL: https://doi.org/10.37828/em.2025.87.7
038A879EFF868C54FF67FDAA5098FC48.taxon	description	(Figs 87 – 90, 138 – 140, 166)	en	László, Gyula M. (2025): Integrative taxonomic review of the Meganola monofascia van Son species group sensu Hacker et al. (Lepidoptera, Noctuoidea, Nolidae, Nolinae, Nolini). Ecologica Montenegrina 87: 135-191, DOI: 10.37828/em.2025.87.7, URL: https://doi.org/10.37828/em.2025.87.7
038A879EFF868C54FF67FDAA5098FC48.taxon	materials_examined	Material examined (all in coll. ANHRT). Gabon. 1 male, Mikongo (Rougier), Monts de Cristal (Secondary Forest), 430 m, 0 ° 29 ' 47 " N, 11 ° 10 ' 42 " E, 28. vii – 12. viii. 2019, MV light trap, leg. Albert, J-L., Aristophanous, M., Bie Mba, J., Dérozier, V., Moretto, P., gen. slide No.: LGNA 1278. Mozambique. 1 male, Zambezia Province, Mt. Namuli, SW slopes near Mucunha village, (forest remnant), 1245 m, 15 ° 22 ' 33 " S, 37 ° 05 ' 05.7 " E, 15 – 16. viii. 2018, actinic light trap, leg. László, G., Miles, W., Vetina, A., gen. slide No.: LGNA 636. Zambia. 4 males, Source of the Zambezi, 1494 m, 11 ° 22 ' 09 " S, 24 ° 18 ' 30 " E, 12 – 13. v. 2015, light trap, leg. Smith, R., Takano, H., Aristophanous, M.; 4 males, Hillwood, Ikelenge (Miombo / Riverine forest mosaic), 1400 m, S 11 ° 16 ' 02 ", E 24 ° 18 ' 59 ', 23 – 30. xi. 2019, actinic and MV light trap, leg. Bashford, M., Miles, W., Mulvaney, L., Smith, R.; 1 male, same site, 28. v – 4. vi. 2020, leg. Chizuwa, D., Choongo, W.; 3 males, 1 female, Chitunta Plain (Miombo / Dambo mosaic), 1396 m, 11 ° 29 ' 12 " S, 24 ° 24 ' 18 " E, 4 – 12. vi. 2020, actinic light trap, leg. Chizuwa, D., Choongo, W.; 5 males, 1 female, same site, 29. xi – 4. xii. 2019, leg. Bashford, M., Miles, W., Mulvaney, L. leg.; 1 male, Kalene Hill (Miombo woodland), 1440 m, 11 ° 11 ' 11 " S, 24 ° 12 ' 5 " E, 12 – 15. vi. 2020, MV light trap, leg. Chizuwa, D., Choongo, W.; 1 female, Nyangombe Falls, 1300 m, 11 ° 48 ' 25 " S, 24 ° 32 ' 12 " E, 17 – 23. xi. 2019, Actinic light trap, leg. Bashford, M., Miles, W., Mulvaney, L., Smith, R.; 1 male, Zambezi Rapids (Miombo / Riverine forest mosaic), 1205 m, S 11 ° 7 ' 30 ", E 24 ° 11 ' 6 ", 4 – 9. xi. 2018, actinic light trap, leg. Aristophanous, M., Dérozier, V., László, G., Oram, D.; 2 males, 1 female, Nkwaji, Mwinilunga, 1316 m, 11 ° 36 ' 22 " S, 24 ° 33 ' 17 " E, 21 – 22. viii. 2013, light trap, leg. Smith, R & Takano, H., gen. slide Nos: LGNA 8 (male), LGNA 20 (female); 1 female, same site and collectors, 27. viii. 2013, gen. slide No.: LGNA 117; 1 male, Kambishi, Jiwundu, 1340 m, 11 ° 54 ' 39 " S, 25 ° 29 ' 05 " E, 18 – 19. x. 2014 Light Trap leg. Smith, Takano & Oram, gen. slide No.: LGNA 1045; 1 male, Chilambwe Falls, Kafubu River, 1420 m, 09 ° 50 ' 13 " S; 30 ° 43 ' 35 " E, 8 – 12. ii. 2019, MV light trap, leg. Dérozier, V., Mulvaney, L., Smith, R., Takano, H.; 1 male, Ntumbachushi Falls, Ngona River, Luapula Prov., 1166 m, 09 ° 51 ' 12 " S, 28 ° 56 ' 40 " E, 12 – 13. v. 2013, light trap, leg. Smith, Takano & Oram, gen. slide No.: LGNA 118; 2 males, Kankonde Camp, Mutinondo Stream, 1400 m, S 12 ° 23 ' 29 ", E 31 ° 19 ' 24 ", 12 – 15. xi. 2014, light trap, leg. Smith, R. & Takano, H., gen. slide Nos: LGNA 282, LGNA 283; 5 males, 3 females, Mutinondo Wilderness Area, Northern Province, 1460 m, 12 ° 27 ' 06 " S, 31 ° 17 ' 30 " E, 17 – 20. v. 2019, actinic and MV light trap, leg. Dérozier, V., Imakando, M., László, G., Miles, W.; 1 female, same site, 16 – 20. iii. 2017, leg. Oram, D., Miles, W., Smith, L.; 1 male, Muchinga Province, 30 km N of Mpika, Danger Hill, 1684 m, 11 ° 37 ' 38 " S, 31 ° 33 ' 56 " E, 13 – 15. xii. 2023, MV light trap, leg. László, G., Morgan, L., Volynkin, A.; 1 female, Senka Hill, Mukulizi Forest Reserve, Muchinga Province, 1566 m, 09 ° 05 ' 43 " S, 32 ° 05 ' 06 " E, 1 – 6. v. 2019 MV light trap, leg. Dérozier, V., László, G., Miles, W.; 1 female, Kalambo Falls, 30 km N of Mbala, Northern Province, 1189 m, 08 ° 35 ' 50 " S, 31 ° 14 ' 26 " E, 9 – 11. v. 2019, actinic light trap, leg. Dérozier, V., László, G., Miles, W., gen. slide No.: LGNA 1033; 3 males, Muchinga Prov., Benyanga village, 1250 m, 10 ° 40 ' 41 " S, 33 ° 27 ' 45 " E, 07 – 12. xii. 2023, actinic and MV light trap, leg. László, G., Morgan, L., Volynkin, A.; 1 male, Muchinga Province, Jombo village, 1400 m, 10 ° 27 ' 01 " S, 33 ° 14 ' 30 " E, 30. xi – 05. xii. 2023, actinic light trap, Bashford, M., Collins, A., László, G., Morgan, L., Volynkin, A. Genetic information. All but one sample (n = 5) of this species belong to the BIN URI: BOLD AEJ 7130. One specimen from northeastern Zambia is assigned the BIN URI: BOLD AEJ 3272. Despite lacking any obvious morphological differences, this specimen diverges from the others by 2.82 – 3.00 %. The samples under AEJ 7130 exhibit significant intraspecific divergence in the range of 0.00 – 0.46 %; however, the PWD between a Gabonese and a Mozambican sample is 1.40 %, along with distinct morphological differences (as discussed below). Meganola ochromorpha does not appear to have any particularly close relatives, as shown in the ML tree, where this species forms a distinct cluster. The genetically closest taxon is M. trigonographa, with a divergence of 3.76 – 4.92 %. Taxonomic notes. A notably small specimen from Gabon exhibited a considerably smaller mediocostal patch and noticeably paler transverse lines compared to specimens from Zambia and Mozambique. However, when examining the genitalia, the Gabonese specimen did not display any distinct characteristics that differentiate it from the southeastern African specimens. It falls within the same BIN URI as M. ochromorpha (BOLD AEJ 7130), but it is situated on a distinct branch of the ML tree. Given its unique external appearance, it cannot be ruled out that the Gabonese population of M. ochromorpha may represent a distinct subspecies; however, a taxonomic decision can only be made when more specimens from the region are available. Interestingly, a female specimen collected from Kalambo Falls in northeastern Zambia also appeared on a distinct branch and has been assigned a different BIN URI: BOLD AEJ 3273. Since this specimen does not exhibit any distinctive genital or external features, it is considered a genetically divergent individual of M. ochromorpha.	en	László, Gyula M. (2025): Integrative taxonomic review of the Meganola monofascia van Son species group sensu Hacker et al. (Lepidoptera, Noctuoidea, Nolidae, Nolinae, Nolini). Ecologica Montenegrina 87: 135-191, DOI: 10.37828/em.2025.87.7, URL: https://doi.org/10.37828/em.2025.87.7
038A879EFF868C54FF67FDAA5098FC48.taxon	distribution	Distribution. This species was previously recorded from Lubumbashi (DRC, type locality), as well as from Burundi and Malawi (Hacker et al. 2012). However, the record from Malawi was based on a misidentification of a female specimen, which actually belongs to M. pyrrhomorpha Hacker, 2012 (see Hacker et al. 2012: 408, fig. G). The specimens examined in this study from Gabon, Mozambique, and Zambia represent new country records.	en	László, Gyula M. (2025): Integrative taxonomic review of the Meganola monofascia van Son species group sensu Hacker et al. (Lepidoptera, Noctuoidea, Nolidae, Nolinae, Nolini). Ecologica Montenegrina 87: 135-191, DOI: 10.37828/em.2025.87.7, URL: https://doi.org/10.37828/em.2025.87.7
038A879EFF858C58FF67FB09573EF941.taxon	description	(Figs 91 – 92, 141 – 143, 167)	en	László, Gyula M. (2025): Integrative taxonomic review of the Meganola monofascia van Son species group sensu Hacker et al. (Lepidoptera, Noctuoidea, Nolidae, Nolinae, Nolini). Ecologica Montenegrina 87: 135-191, DOI: 10.37828/em.2025.87.7, URL: https://doi.org/10.37828/em.2025.87.7
038A879EFF858C58FF67FB09573EF941.taxon	description	Meganola taiana László, 2020, syn. n., Zootaxa 4853 (2): 164. TL: Côte d’Ivoire, Tai NP, Tai Research Station. Holotype, male, in coll. ANHRT.	en	László, Gyula M. (2025): Integrative taxonomic review of the Meganola monofascia van Son species group sensu Hacker et al. (Lepidoptera, Noctuoidea, Nolidae, Nolinae, Nolini). Ecologica Montenegrina 87: 135-191, DOI: 10.37828/em.2025.87.7, URL: https://doi.org/10.37828/em.2025.87.7
038A879EFF858C58FF67FB09573EF941.taxon	materials_examined	Material examined (all in coll. ANHRT). Côte d’Ivoire. 3 males, Tai NP. Tai Research Station, 174 m, 05 ° 49 ' 59.8 " N, 07 ° 20 ' 32.0 W, 14 – 23. xi. 2015, light trap, leg. Aristophanous, M., Moretto, P., Ruzzier, E., gen. slide Nos: LGNA 142 (holotype of M. taiana), LGNA 147, LGNA 209 (paratypes of M. taiana); 2 females, same site, 25. iii – 17. iv. 2017, actinic and MV light trap, leg. Aristophanous, A., Aristophanous, M., Geiser, M., Moretto, P., gen. slide Nos: LGNA 400, LGNA 981. Guinea. 1 female, Geipa Camp, Foret de Diecke, 435 m, 7 ° 26 ' 7.06 " N, 8 ° 50 ' 47.87 " W, 05 – 14. iv. 2019, light trap, blended bulb (250 W), leg. Sáfián, Sz., Koivogui, S., gen. slide No.: LGNA 987. Liberia. 1 female, Krahn-Bassa Reserve, Sinoe County, Juboe River, 7.5 km South West Pellokon Town, 140 m, 5 ° 39 ' 4 " N, 8 ° 39 ' 4 " W, 14 – 21. i. 2018, Cold Cathode UV bucket light trap, leg. Geiser, M., Sáfián, Sz., Simonics, G., gen. slide No.: LGNA 985; 1 female, Welezu Camp, Wonegizi Nature Reserve, Lofa County, 551 m, 8 ° 4 ' 57.11 " N, 9 ° 34 ' 47.86 " W, 19 – 27. iii. 2019, Cold Cathode UV light trap (8 W), leg. Sáfián, Sz., Koivogui, S., gen. slide No.: LGNA 1260; 1 male, Lofa County, Foya Proposed Protected Area, 530 m, 7 ° 56 ' 36 " N, 10 ° 16 ' 36 " W, 10 – 19. xi. 2017, MV light trap (125 W), leg. Aristophanous, M., Sáfián, Sz., Simonics, G. & Smith, L., gen. slide No.: LGNA 551 (paratype of M. taiana); 1 male, Nimba Mts camp, ENNR, Nimba county, 1165 m, 7 ° 31 ' 45 " N, 8 ° 31 ' 37 " W, 03 – 13. xii. 2017, light trap (blended bulb 250 W), leg. Aristophanous, M., Sáfián, Sz., Simonics, G., Smith, L., gen. slide No.: LGNA 487 (paratype of M. taiana). Sierra Leone. 4 males, Tiwai Island, Moa River, 120 m, 07 ° 33 ' 00 " N, 11 ° 21 ' 09 " W, 17 – 22. vi. 2016, light trap, leg. Takano, Miles & Goff, gen. slide Nos: LGNA 409, LGNA 515, LGNA 564 (paratypes of M. taiana), LGNA 1252. Genetic information. This species has been assigned the BIN URI: BOLD AEJ 4832. The intraspecific divergence calculated from five samples is 0.00 %. Genetic analyses indicate that M. taiana László, 2020 syn. n., is conspecific with M. canocolorata, which was described based on a female holotype. This species is genetically distinct from all other taxa in the complex and has a similar PDW to M. fontainei (4.18 – 4.42 %), M. bispermutata cryptochra (4.09 – 4.41 %), M. trigonographa (4.25 – 4.42 %), and M. dicranographa (4.41 – 4.42 %), making these its closest relatives. Taxonomic notes. Meganola canocolorata was described based on a single female specimen from Côte d’Ivoire. While the external appearance of this species is not particularly distinctive, the configuration of the female genitalia is relatively unique. It is characterized by a short and thick ductus bursae with asymmetrical laterodistal sclerotisation and a relatively large, ovoid corpus bursae that features two flat signa. The only other species exhibiting a similar female genitalia configuration is M. odzala Hacker, 2012. In the collections of the ANHRT, several female specimens from West African localities were found with genital structures that closely match the illustration of M. canocolorata in Hacker et al. (2012). Thanks to large-scale DNA barcoding of the Nolinae material at ANHRT, the male of M. canocolorata has been identified from Sierra Leone and, as part of the paratype series of M. taiana László, 2020, from Liberia. The ML analysis revealed that the female specimens included in the paratype series of M. taiana actually belong to M. palaeographa, as previously discussed. Consequently, based on the genetic identity of morphologically identified females of M. canocolorata and the male type specimens of M. taiana, this latter species is synonymised here with the former: M. taiana syn. n. Conclusions This study aimed to revise the taxonomy of a diverse and complex group of species that resemble Meganola monofascia and establish a consistent species group classification. The taxa of these species groups are often externally indistinguishable from one another, and, at the same time, several taxa display remarkable polymorphism, further complicating their identification. The new species grouping system combines diagnostic genital morphology with genetic analyses based on DNA barcodes. This integrative approach has led to a morphology-based classification that identifies three species groups, aligning perfectly with the genetic findings: each morphological group corresponds to a monophyletic cluster in the phylogenetic tree. The molecular studies also clarified male and female individuals for several species, correcting previous taxonomic errors that mistakenly treated different sexes as distinct species or incorrectly paired non-conspecific males and females within a single species. The M. monofascia-furvitincta-bispermutata complexes represent one of the most taxonomically challenging groups within the genus. Preliminary analyses have uncovered additional discrepancies in previous classifications concerning other Meganola species groups as well. This indicates the need for further integrative studies on the remaining Afrotropical Meganola taxa (László in prep.). It is important to emphasize that, despite the publication of numerous comprehensive works over the past decade, our understanding of Afrotropical Nolini at the species level remains incomplete, with many taxa still awaiting description. Hacker et al. (2012) estimated that 70 to 90 species are yet to be discovered in the Afrotropics. Based on material currently being studied at ANHRT, more than 100 Nolini taxa have already been identified as undescribed. Over half of these are from the previously neglected northern region of the Republic of Congo, while the remaining new species are expected to be found in West Africa and the Zambian plateau (László in prep.). Acknowledgements The author is indebted to Alberto Zilli and Geoff Martin (NHMUK) for facilitating access to the Nolini type material under their care. Image of the lectotype of M. furvitincta used with permission and are copyright of the Trustees of the Natural History Museum, London, made available under Creative Commons License 4.0 (https: // creativecommons. org / licenses / by / 4.0 /). Anthony Kingston (Albrighton, UK) is thanked for the loan of Nolinae specimens for this study. Paul D. N. Hebert (CBG, Guelph, Canada), Evgeny Zakharov (CCDB, Guelph, Canada) and their competent teams are thanked for their continuous support in DNA barcoding of Nolidae. The genetic analyses would not have been possible without Sujeevan Ratnasingham (CBG, Guelph, Canada), who developed the BOLD database. Oleg Pekarsky (Budapest, Hungary) prepared several genital slides used in this study. The author is especially grateful to Gabriella László (ANHRT) for her exceptional skill in mounting thousands of Nolini specimens. The following collaborative partners are thanked for facilitating ANHRT field work and issuing the necessary research and export permits. Côte d'Ivoire: the Ministère de l'Enseignement Supérieur et de la Recherche Scientifique; the Office Ivoirien des Parcs et Réserves (OIPR) and the Société de Développement des Forêts (SODEFOR). Gabon. Scientific research was authorised by the Centre National de la Recherche Scientifique (CENAREST) in collaboration with the Université des Sciences et Techniques de Masuku (USTM). Rougier Gabon is thanked for allowing access to concession forest. Guinea: Mamadou Diawara, (Guineé Ecologie); Colonel Layaly Camara, Cece Papa Konde (Ministère de l’Environment et aux Eaux et Forêts); Jamison Suter (Societe des Mines de Fer de Guineé). Liberia. Annika Hiller (Wild Chimpanzee Foundation), Jerry Garteh (Society for the Conservation of Nature), Darlington Tuaben, Mike C. Doryen and Kederick F. Johnson (Forestry Department Authority, Liberia). Mozambique: Bartolomeu Soto, Mateus Mutemba and Antony Chilengue (Administração Nacional das Areas de Conservação, Maputo); Lucilia Chuquela, Kisimenda Muambalo, Anacleto Cossa and Alvaro Vetina (Museu de História Natural de Maputo); Majka Burhardt and Marcos Assane (LEGADO Namuli, Gurue). Republic of Congo: Richard Malonga, Morgane Cournarie, Vittoria Estienne, Ben Evans, Onesi Samba and Yako Valentin (WCS, Parc National de Nouabalé-Ndoki); Jonas Erickson, Gwili Gibbon, Sylvain Yaba Ngouma, Adrich Mouamana (African Parks, Parc National d’Odzala-Kokoua), Merlia Talani (Marien Ngouabi University, Brazzaville), the Ministère de l’Enseignement Superieur de la Recherche Scientifique et de l’Innovation Technologique, Joseph Moumbouilou and Jean Bosco Nganongo (Direction de la Faune et des aires protégées, DEFAP), Joseph Goma-Tchimbakala, Victor Mamonekene (Institut National de Recherche en Sciences Exactes et Naturelles, IRSEN). Zambia: Rhoda Kachali (Department of National Parks and Wildlife – ZAWA, Lusaka), Claire Mateke and Martha Imakando (Livingstone Museum, Livingstone). The author declares that, to the best of his knowledge, he conforms to the national regulations and meets with the conditions and requirements of International Conventions concerning collecting / export and handling of the specimens presented in this Article.	en	László, Gyula M. (2025): Integrative taxonomic review of the Meganola monofascia van Son species group sensu Hacker et al. (Lepidoptera, Noctuoidea, Nolidae, Nolinae, Nolini). Ecologica Montenegrina 87: 135-191, DOI: 10.37828/em.2025.87.7, URL: https://doi.org/10.37828/em.2025.87.7
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