identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
038887E6D810670133C5B97C0B65FEBD.text	038887E6D810670133C5B97C0B65FEBD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Araloselachus vorax (Le Hon 1871)	<div><p>Araloselachus vorax (Le Hon, 1871)</p><p>(Figs 4–12)</p><p>Referred material –12 teeth (V.61.915, V.61.917, V.61.921, V.62.396,V.63.412, V.69.1059, V.69.1100).</p><p>Description – The main cusp of anterior files is slender and pointed, showing a slight lingual curvature in profile view (Figs 4–9). Upper lateral tooth (V.62.396) has distally inclined main cusp, straight in profile view (Figs 10–12). While the examined anterior teeth are ×1.5–2 higher apicobasally than wide mesiodistally, the upper lateral tooth is nearly as wide as high. Th e cutting edge of all teeth is smooth and sharp, it runs from the tip to the crown-base. The labial crown face is weakly convex to flat, while the labial is strongly convex. The enameloid surface of the labial crown face is completely smooth. The root is bilobate, the lobes meet in a pronounced lingual protuberance, bearing a large nutrient groove. The lobes bear usually two pairs of sharp cusplets (some anterior files bear only one pair), of which the first pair is strongly hooked towards the main cusp in anterior teeth.</p><p>Remarks – This species was originally described as Lamna (Odontaspis) vorax by LE HON (1871). Later on, it was widely included in Carcharias (e.g. DE SCHUTTER 2011; see also synonym lists of REINECKE et al. 2011 and BOR et al. 2012), but BOR et al. (2012) re-assigned the species into the genus Araloselachus .</p><p>From the Miocene of Europe, large-sized odontaspidid teeth with smooth labial crown face are widely referred to as Araloselachus cuspidatus and A. vorax (e.g. REINECKE et al. 2011, 2014). The two species seem to be closely related, but the latter species has smaller, less-massive teeth, also less-developed root lobes, with multiple pairs of sharp and hooked lateral cusplets, of which the secondary ones are well developed, especially in anterior files (see DE SCHUTTER 2011; REINECKE et al. 2011; BOR et al. 2012: 23). A. vorax is distributed mainly in the North Sea Basin during the Burdigalian-Tortonian, but it seems to occur occasionally in the Burdigalian-Langhian deposits of the Central Paratethys exposed in Austria and southern Germany (DE SCHUTTER 2011: 179).</p><p>Teeth of C. acutissima and C. taurus, two hardly distinguishable species widespread in the Badenian of the Central Paratethys, are easily separable from A. vorax, as they have slender teeth with apicobasally striated lingual crown face (WARD &amp; BONAVIA 2001; REINECKE et al. 2011, 2014; SZABÓ &amp; KOCSIS 2016, table 2).</p></div>	https://treatment.plazi.org/id/038887E6D810670133C5B97C0B65FEBD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Szabó, Márton	Szabó, Márton (2019): Middle Miocene (Badenian) chondrichthyan and osteichthyan remains from St. Margarethen (eastern Austria) in the vertebrate palaeontological collection of the Hungarian Natural History Museum. Fragmenta Palaeontologica Hungarica 36: 53-90, DOI: 10.17111/FragmPalHung.2019.36.53, URL: https://doi.org/10.17111/fragmpalhung.2019.36.53
038887E6D812670133E0BCA70D9BFB7E.text	038887E6D812670133E0BCA70D9BFB7E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Carcharias crassidens (Agassiz 1838)	<div><p>cf. Carcharias crassidens (Agassiz, 1838)</p><p>(Figs 13–15)</p><p>Referred material – 1 tooth (V.69.1092).</p><p>Description – The single here referred tooth is nearly as wide mesiodistally as high apicobasally. The main cusp is upright, triangular, and narrow in labial and lingual views (Fig. 13), with a flattened labial and convex lingual face. Both faces and cutting edges are smooth, the cutting edges reach the crown base, and continue on the lateral cusplets. The main cusp has a weak lingual inclination in profile view (Fig. 14). The root is bilobate, the lobes are well separated, of which the better preserved one has a rounded extremity. The lobes meet in an obtuse angle. Both lobes bear one lateral cusplet, fused to the basal edges of the main cusp. The labial root face bears a characteristic, mesiodistally running ridge below the crown base (Fig. 15).</p><p>Remarks – Here I follow STONE &amp; SHIMADA (2019), who resurrected family Carchariidae for the genus Carcharias to separate it from family Odontaspididae . Tooth V.69.1092 is reminiscent of a figured type specimen of Carcharias crassidens (= Lamna crassidens, see AGASSIZ 1838, pl. 35, fig. 11, 11’). Since the validity of this species is doubtful, and its general tooth morphology is similar to that of the genus Araloselachus, here I refer tooth V.69.1092 provisionally as cf. Carcharias crassidens, until a revision of C. crassidens is done, or further specimens from St. Margarethen are identified.</p></div>	https://treatment.plazi.org/id/038887E6D812670133E0BCA70D9BFB7E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Szabó, Márton	Szabó, Márton (2019): Middle Miocene (Badenian) chondrichthyan and osteichthyan remains from St. Margarethen (eastern Austria) in the vertebrate palaeontological collection of the Hungarian Natural History Museum. Fragmenta Palaeontologica Hungarica 36: 53-90, DOI: 10.17111/FragmPalHung.2019.36.53, URL: https://doi.org/10.17111/fragmpalhung.2019.36.53
038887E6D812670B3305B8050E2EFDC2.text	038887E6D812670B3305B8050E2EFDC2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Carchariidae Muller et Henle 1841	<div><p>Carchariidae indet . or Odontaspididae indet.</p><p>(Figs 16 –65)</p><p>Referred material – a slightly disarticulated, partial dentition surrounded by placoid scales and suggested portions of cartilage (VER 2020.69.1.), from which 15 placoid scales (VER 2020.70.1.–84.1.), 1 piece of stone matrix embedding at least 5 placoid scales (VER 2020.85.1.), and 6 pieces of suggested cartilage tissue (VER 2020.86.1.–90.1.) were isolated.</p><p>Description – Based on the visible crown portions, specimen VER 2020.69.1. (Fig. 16) consists of at least 32 teeth, however, it is hard to tell how many teeth the broken root portions belong to. Some teeth are preserved in labial, while some in lingual aspect. Th e teeth represent the upper and lower anterior and lateral files, including the intermediate file of the upper dentition (Figs 17–25). No posterior files are visible. All teeth which are embedded in lingual aspect, possess a smooth lingual crown face. Based on the variety of the observed tooth positions, VER 2020.69.1. represents a slightly disarticulated mixture of the upper and lower dentition.</p><p>A large number of placoid scales are visible in the embedding limestone matrix. Placoid scales were isolated from three areas around the dentition, where the scales are densely aggregated (Fig. 26). It is noteworthy that placoid scales sparsely occur over a large part of the surface of the embedding piece of limestone. The isolated placoid scales represent four morphotypes. Morphotype 1 (isolated from areas I and III; see Figs 27–36, 49, 50) has drop- or arrowhead-shaped crown. Threefour anteriorly bulging, parallel folds are present on the apical surface, reaching from the anterior margin to the anteroposterior half of the crown. The crown of placoid scales representing Morphotype 2 (isolated from area II; see Figs 37–46) is circular in apical view. The anterior margin is weakly folded. Morphotype 3 (isolated from area III; see Figs 47, 48, 51–56) is similar to Morphotype 1 in having a pointed posterior edge, but the crown meets the base in a lower angle. This morphogroup seems to represent an intermediate form between the first and the fourth morphotype. Morphotype 4 (isolated from area III; see Figs 57, 58) has tricuspid crown, bearing an anteriorly bulging median keel, laterally flanked by parallel folds. The lateral folds diminish posteriorly, while the median keel reaches the posterior margin of the crown. The base of all morphotypes bears large foramina.</p><p>The suggested cartilage tissue of specimen VER 2020.69.1. is built up of mosaically adjacent units of irregular outline. The largest, and also the most well-preserved portions of the suggested cartilage are between the surface area I and III (Figs 59–61). Pieces of this structure were also isolated for scanning electron microscopic photography (Figs 62–65).</p><p>Remarks – Based on genetic studies, STONE &amp; SHIMADA (2019) resurrected Carchariidae for genus Carcharias to separate it from family Odontaspididae . However, tooth morphology of the here referred fossil taxa was a subject of many studies in the past (e.g. DE SCHUTTER 2011; REINECKE et al. 2011, 2014; BOR et al. 2012; CAPPETTA 2012), and differential tooth characteristics of the two families have not been detailed since the work of STONE &amp; SHIMADA (2019). Except for the less-developed and low number of lateral cusplets (usually one pair in any teeth of specimen VER 2020.69.1.), the dental characteristics are comparable with those of A. vorax (see DE SCHUTTER 2011; BOR et al. 2012). The notably small size of the teeth refers specimen VER 2020.69.1. (and its isolated portions) to a juvenile individual, with assumedly less-developed dental characteristics. However, two different shark species sharing similar dental characteristics, while inhabiting the same environment is highly unlikely from the biological point of view (after REINECKE et al. 2011: 21), following its ontogenetic stage, an assignment of specimen VER 2020.69.1. (and its isolated portions) to A. vorax would be quite hazardous. As both Carchariidae and Odontaspididae are reported from St. Margarethen (SCHMID et al. 2001; present study), here I find more appropriate to describe specimen VER 2020.69.1. with all of its isolated portions only as Carchariidae indet. or Odontaspididae indet. Differences in tooth morphology of these two, hardly separable families are not detailed yet, which is beyond the scope of the present study.</p><p>It is worth mentioning that placoid scales of Morphotype 2 are weakly reminiscent of scales/those of an embrional individual of the Oligocene-Miocene species C. gustrowensis (HOVESTADT &amp; HOVESTADT-EULER 2010, fi g. 6. 68.), in having a circular, anteriorly folded crown. REIF (1985) figured dermal denticles of the extant species Carcharias taurus, but they have a different morphology by having much stronger folds on the apical surface, reaching the posterior edge of the denticles.</p></div>	https://treatment.plazi.org/id/038887E6D812670B3305B8050E2EFDC2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Szabó, Márton	Szabó, Márton (2019): Middle Miocene (Badenian) chondrichthyan and osteichthyan remains from St. Margarethen (eastern Austria) in the vertebrate palaeontological collection of the Hungarian Natural History Museum. Fragmenta Palaeontologica Hungarica 36: 53-90, DOI: 10.17111/FragmPalHung.2019.36.53, URL: https://doi.org/10.17111/fragmpalhung.2019.36.53
038887E6D818670B33ECBFFF0B0BF9DB.text	038887E6D818670B33ECBFFF0B0BF9DB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cosmopolitodus hastalis (Agassiz 1838)	<div><p>Cosmopolitodus hastalis (Agassiz, 1838)</p><p>(Figs 66–71)</p><p>Referred material – 13 teeth (V.61.917, V.61.925, V.61.1278, V.61.1280, V.62.395, V.62.397, V.62.399, V.69.286, V.69.306, V.69.1010, V.69.1038, V.69.1092).</p><p>Description – The crown of upper anterior and anterolateral teeth (Figs 66, 67) is high, triangular and labiolingually flattened. The labial crown face is flat, while the lingual is weakly convex. Upper lateral teeth have a distally directed crown, rapidly widening basally, extending onto the shoulders of the root lobes, and sometimes forming a heel in upper distal files (see V.62.399, Figs 68, 69). The short root lobes of upper files are weakly pronounced lingual protuberance. Lower teeth (Figs 70, 71) have lingually weakly curved, narrow crown (compared to upper teeth), the labial crown face is weakly, while the lingual is strongly convex. The root of lower files is built up of well-separating lobes. The crown faces and cutting edges of all files are smooth. The lingual crown-root boundary is marked by a narrow dental band in all files.</p><p>Remarks – This species is widespread in the Badenian deposits of the Central Paratethys (SZABÓ &amp; KOCSIS 2016, table 2). Following the dental similarities of Cosmopolitodus and Carcharodon carcharias, some authors place the species into the genus Carcharodon (e.g. EHRET et al. 2012; TRIF &amp; CODREA 2017; KENT in GODFREY 2018; MAISCH et al. 2018). Since the evolutional relationship of Cosmopolitodus and Carcharodon is not fully resolved at present, here I follow CAPPETTA (2012), who places species hastalis in genus Cosmopolitodus .</p></div>	https://treatment.plazi.org/id/038887E6D818670B33ECBFFF0B0BF9DB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Szabó, Márton	Szabó, Márton (2019): Middle Miocene (Badenian) chondrichthyan and osteichthyan remains from St. Margarethen (eastern Austria) in the vertebrate palaeontological collection of the Hungarian Natural History Museum. Fragmenta Palaeontologica Hungarica 36: 53-90, DOI: 10.17111/FragmPalHung.2019.36.53, URL: https://doi.org/10.17111/fragmpalhung.2019.36.53
038887E6D81A670933CBBC0C0B73FC44.text	038887E6D81A670933CBBC0C0B73FC44.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Isurus oxyrinchus (Agassiz 1843)	<div><p>Isurus oxyrinchus (Agassiz, 1843)</p><p>(Fig. 72)</p><p>Referred material – 1 tooth (V.61.915).</p><p>Description – The only tooth referred here is embedded in a piece of Leitha Limestone, in labial aspect (Fig. 72). The crown is slender and pointed, it slightly inclines distally, with a lingual curvature the very tip reclines labially. Both cutting edges are smooth, the mesial is convex, while the distal is nearly straight. The distal cutting edge does not reach the crown base. The root is bilobate and slightly asymmetrical, the lobes meet in an acute angle. Based on the preserved dental characteristics, specimen V.61.915 is referred as an anterior tooth, most likely from the lower dentition (REINECKE et al. 2011, pl. 40 with all figurations; CAPPETTA 2012, fig. 203H, I).</p><p>Remarks – PURDY et al. (2001) noted that I. desori should be considered as junior synonym of I. oxyrinchus . BOR et al. (2012: 34) also suggest that there is no sufficient difference between Miocene teeth of the “ desori ” morphotype and teeth of extant I. oxyrinchus, which would support handling them under separate names. When treating the two species as synonyms, it can be concluded that I. oxyrinchus was widely distributed in the Central Paratethys during the Early and Middle Miocene (RADWAŃSKI 1965; SCHULTZ 1971; HOLEC et al. 1995; KOCSIS 2007).</p></div>	https://treatment.plazi.org/id/038887E6D81A670933CBBC0C0B73FC44	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Szabó, Márton	Szabó, Márton (2019): Middle Miocene (Badenian) chondrichthyan and osteichthyan remains from St. Margarethen (eastern Austria) in the vertebrate palaeontological collection of the Hungarian Natural History Museum. Fragmenta Palaeontologica Hungarica 36: 53-90, DOI: 10.17111/FragmPalHung.2019.36.53, URL: https://doi.org/10.17111/fragmpalhung.2019.36.53
038887E6D81A6709307CB91F0E82FA9E.text	038887E6D81A6709307CB91F0E82FA9E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lamnidae Muller et Henle 1838	<div><p>Lamnidae indet.</p><p>Referred material – 9 teeth (V.61.914, V.61.924, V.61.927, V.61.928, V.62.397, V.62.402, V.63.412, V.69.1057, V.69.1092).</p><p>Description – A few rootless and/or broken crowns and incomplete root remains, similar to those of C. hastalis and I. oxyrinchus are referred here. However, due to their poor preservation their assignment to any lamnid at St. Margarethen would be uncertain.</p></div>	https://treatment.plazi.org/id/038887E6D81A6709307CB91F0E82FA9E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Szabó, Márton	Szabó, Márton (2019): Middle Miocene (Badenian) chondrichthyan and osteichthyan remains from St. Margarethen (eastern Austria) in the vertebrate palaeontological collection of the Hungarian Natural History Museum. Fragmenta Palaeontologica Hungarica 36: 53-90, DOI: 10.17111/FragmPalHung.2019.36.53, URL: https://doi.org/10.17111/fragmpalhung.2019.36.53
038887E6D81A6717332DB8E00B70FD70.text	038887E6D81A6717332DB8E00B70FD70.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Otodus (Megaselachus) megalodon (Agassiz 1835)	<div><p>Otodus (Megaselachus) megalodon (Agassiz, 1835)</p><p>(Figs 73–78)</p><p>Referred material –8teeth(V.60.1741,V.61.940,V.61.944,V.61.1278,V.61.1282).</p><p>Description – The teeth are large and massive. The crown is triangular, both crown faces are smooth. The labial crown face is flat, while the lingual is convex.</p><p>Both cutting edges bear strong, regular serrations. The root-crown junction bears a well-developed and high, V-shaped chevron (= “neck”) on the lingual crown face. The root is bilobate, the lobes are equal in size. The most complete specimen (V.60.1741) is embedded in a piece of limestone in labial aspect (Figs 73, 74). This tooth shows a pathological deformity: the distal and mesial cutting edge undulates at the crown base (Figs 75–78).</p><p>Remarks – Here I follow the system of CAPPETTA (2012), however, his division of Otodus into subgenera Otodus, Carcharocles, and Megaselachus is not further supported by some authors (e.g. BOR et al. 2012). Otodus (Megaselachus) megalodon is the largest macropredatory shark ever lived. It was previously reported from St. Margarethen by SCHMID et al. (2001). The species is widely known from the Badenian of the Central Paratethys (SZABÓ &amp; KOCSIS 2016, table 2). SCHULTZ (1971, pl. 3, fig 17a, b) figures a tooth of Otodus (Megaselachus) megalodon with a pathological deformity similar to that of specimen V 60.1741.</p></div>	https://treatment.plazi.org/id/038887E6D81A6717332DB8E00B70FD70	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Szabó, Márton	Szabó, Márton (2019): Middle Miocene (Badenian) chondrichthyan and osteichthyan remains from St. Margarethen (eastern Austria) in the vertebrate palaeontological collection of the Hungarian Natural History Museum. Fragmenta Palaeontologica Hungarica 36: 53-90, DOI: 10.17111/FragmPalHung.2019.36.53, URL: https://doi.org/10.17111/fragmpalhung.2019.36.53
038887E6D804671733D9BE910E17FAC7.text	038887E6D804671733D9BE910E17FAC7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hemipristis serra Agassiz 1835	<div><p>Hemipristis serra Agassiz, 1835</p><p>(Figs 79, 80)</p><p>Referred material – 3 teeth (V.61.937, V.62.405, V.63.480).</p><p>Description – Specimens V.61.937 and V.62.405 are embedded in a piece of limestone matrix in labial aspect (Figs 79, 80). All of the here referred teeth have similar morphology. The crown is triangular, and distally inclined. Both cutting edges are coarsely serrated. The root is bilobate, the lobes are well-separated, with angled extremities. This morphology refers the teeth as upper files.</p><p>Remarks – The species is also widely reported from the Badenian deposits of the Central Paratethys (SZABÓ &amp; KOCSIS 2016, table 2). The genus is represented by a single extant species H. elongata, distributed in the Indo-West Pacific (EBERT et al. 2013).</p></div>	https://treatment.plazi.org/id/038887E6D804671733D9BE910E17FAC7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Szabó, Márton	Szabó, Márton (2019): Middle Miocene (Badenian) chondrichthyan and osteichthyan remains from St. Margarethen (eastern Austria) in the vertebrate palaeontological collection of the Hungarian Natural History Museum. Fragmenta Palaeontologica Hungarica 36: 53-90, DOI: 10.17111/FragmPalHung.2019.36.53, URL: https://doi.org/10.17111/fragmpalhung.2019.36.53
038887E6D804671633C5B8FD0B1DFDEB.text	038887E6D804671633C5B8FD0B1DFDEB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Galeocerdo aduncus (Agassiz 1835)	<div><p>Galeocerdo aduncus (Agassiz, 1835)</p><p>(Figs 81–85)</p><p>Referred material – 8 teeth (V.61.924, V.61.1281, V.69.1066, V.69.1089, V.69.1096).</p><p>Description – Both faces of the crown are smooth, the labial one is flat, while the lingual is convex. The convex mesial cutting edge bears slightly irregular serrations varying between fine and uniform. The straight to weakly convex distal cutting edge is shorter than the mesial edge, and finely serrated. A distal crown heel is present, which is separated from the distal cutting edge by a deep notch. The heel is convex in labial (and lingual) view, it declines to the distal crown margin. The root is apicobasally low and mesiodistally wide, the root lobes meet in an obtuse angle forming a weak lingual protuberance.</p><p>Remarks – This extinct tiger shark species is widely reported from the Badenian of the Central Paratethys (SZABÓ &amp; KOCSIS 2016, table 2).</p></div>	https://treatment.plazi.org/id/038887E6D804671633C5B8FD0B1DFDEB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Szabó, Márton	Szabó, Márton (2019): Middle Miocene (Badenian) chondrichthyan and osteichthyan remains from St. Margarethen (eastern Austria) in the vertebrate palaeontological collection of the Hungarian Natural History Museum. Fragmenta Palaeontologica Hungarica 36: 53-90, DOI: 10.17111/FragmPalHung.2019.36.53, URL: https://doi.org/10.17111/fragmpalhung.2019.36.53
038887E6D805671633AEBFA80EE0FAB3.text	038887E6D805671633AEBFA80EE0FAB3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Carcharhinus priscus (Agassiz 1843)	<div><p>Carcharhinus priscus (Agassiz, 1843)</p><p>(Figs 86–89)</p><p>Referred material – 8teeth (V.61.916, V.61.934, V.61.941, V.61.1279, V.63.482, V.63.483, V.69.883).</p><p>Description – A strong dignathic heterodonty is characteristic for the species. Upper teeth (Figs 86, 87) have triangular, labiolingually flattened crown with flat labial, and weakly convex lingual face. Th e cutting edges are serrated, they continue to the root lobes, forming serrated heels both mesially and distally. The root is mesiodistally wide, the lobes meet in an obtuse angle. Lower teeth (Figs 88, 89) have slender, upright cusp. The cutting edges and the heels are entire and smooth. The root is laterally expanded, the lobes are symmetrical with rounded extremities. The lingual protuberance bears a nutrient groove in all files.</p><p>Remarks – C. priscus is also widely reported from the Badenian of the Central Paratethys (SZABÓ &amp; KOCSIS 2016, table 2). This species has likely evolved from the Early Oligocene C. elongatus or the Late Oligocene C. gibbesi (BOR et al. 2012, and references therein).</p></div>	https://treatment.plazi.org/id/038887E6D805671633AEBFA80EE0FAB3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Szabó, Márton	Szabó, Márton (2019): Middle Miocene (Badenian) chondrichthyan and osteichthyan remains from St. Margarethen (eastern Austria) in the vertebrate palaeontological collection of the Hungarian Natural History Museum. Fragmenta Palaeontologica Hungarica 36: 53-90, DOI: 10.17111/FragmPalHung.2019.36.53, URL: https://doi.org/10.17111/fragmpalhung.2019.36.53
038887E6D80567143042B8490DE9FE28.text	038887E6D80567143042B8490DE9FE28.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Carcharhinidae (KOZUCH & FITZGERALD 1989)	<div><p>Carcharhinidae indet.</p><p>(Figs 90, 91)</p><p>Referred material – 1 vertebra (V.63.478).</p><p>Description – A single centrum, showing diagnostic features of those of Carcharhinidae (KOZUCH &amp; FITZGERALD 1989) is discussed here. Some portions around the edges are missing, the overall surface seems to be a bit eroded, and therefore no pores are visible on the sides. Th e concave articular surface shows circular annuli (Fig. 90). The dorsal foramina are oval, they do not extend the rims of the centrum (Fig. 91).</p><p>Remarks – Similar carcharhinid centra have been reported by PURDY et al. (2001, fig. 52e, f), BOR et al. (2012, pl. 57, figs 1, 2, 6), SZABÓ et al. (2017, fig. 8k- n) and SZABÓ &amp; KOCSIS (2020, pl. 10, figs 1–12). Due to the lack of characters, it is undefinable, whether this centrum belongs to G. aduncus or C. priscus, and it is identified here only as Carcharhinidae indet.</p></div>	https://treatment.plazi.org/id/038887E6D80567143042B8490DE9FE28	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Szabó, Márton	Szabó, Márton (2019): Middle Miocene (Badenian) chondrichthyan and osteichthyan remains from St. Margarethen (eastern Austria) in the vertebrate palaeontological collection of the Hungarian Natural History Museum. Fragmenta Palaeontologica Hungarica 36: 53-90, DOI: 10.17111/FragmPalHung.2019.36.53, URL: https://doi.org/10.17111/fragmpalhung.2019.36.53
038887E6D807671333B2BF3F0FEEFE57.text	038887E6D807671333B2BF3F0FEEFE57.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Aetobatus arcuatus (Agassiz 1843)	<div><p>Aetobatus arcuatus (Agassiz, 1843)</p><p>(Figs 92–95)</p><p>Referred material – 10 teeth (V.61.913, V.61.936, V.63.671).</p><p>Description – Only lower teeth are known in the HNHM collection. Lower teeth of A. arcuatus are of crescent shape, with distally curved lateral edges. The crown is low and straight with smooth occlusal surface. The crown is separated from the root by a lingual bulge. The root is lingually shifted, it is built up by long, distally inclined laminae (Figs 92–94). Specimen V.61.936 is a partial upper dentition, consisting of at least 3 teeth (Fig. 95).</p><p>Remarks – Dentition of genus Aetobatus consists of a single row of teeth, in a conveyor-like, horizontal arrangement both in the upper and lower jaw (BOR et al. 2012; HOVESTADT &amp; HOVESTADT-EULER 2013). Fossil species A. arcuatus is widely known from the Badenian Central Paratethys (SZABÓ &amp; KOCSIS 2016, table 2).</p></div>	https://treatment.plazi.org/id/038887E6D807671333B2BF3F0FEEFE57	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Szabó, Márton	Szabó, Márton (2019): Middle Miocene (Badenian) chondrichthyan and osteichthyan remains from St. Margarethen (eastern Austria) in the vertebrate palaeontological collection of the Hungarian Natural History Museum. Fragmenta Palaeontologica Hungarica 36: 53-90, DOI: 10.17111/FragmPalHung.2019.36.53, URL: https://doi.org/10.17111/fragmpalhung.2019.36.53
