taxonID	type	description	language	source
0397A14B5F7F5A3BFDBFFA61FC0237B4.taxon	materials_examined	Type material Holotype. 1 ♂, China: Shaanxi Province: Xi’an City, Huyi District, Taiping National Forest Park, 33.905 ° N, 108.647 ° E, 5 April 2020, leg. Li Wei-Hai and Yan Yan-Hua (HIST). General colour dark brown. Hemitergal lobe yellow, short, generally triangular with widely notched edge. Epiproct roughly rod-like, apex hooked. Aedeagus membranous, slender and tubular in lateral Description Adult habitus. (Figure 1 (a )). General body colour dark brown. Head black, a trapezoidal brownish originating between the posterior ocelli and terminating anterior to the pronotum; a yellow brown marking present behind compound eye. M-line shallow and brown. Triocellate, anterior ocellus Antennae brown, basal segments darker, palpi brown. Pronotum dark brown, disc with obscure and narrow pale medial stripe (Figure 1 (a )). Legs tri-coloured: femur black with yellow-brown distal tibia brown with basal and distal portion darker, tarsal segments brown. Wings subhyaline, veins Male. (Figures 1 (b), 2). Fore wing length 10.4 mm, hind wing length 8.7 mm (N = 1). Abdominal terga brownish and terga 1 – 2 less pigmented. Tergum 9 bearing two elevated paramedial patches of sensilla long hairs on the posterior margin, the patches separated by a narrow bald mesial area (Figure 1 (b )). 10 bilobate, hemitergal lobe yellow, short, generally triangular in dorsal view, with the inner edge HISTORY connected to the epiproct apically (Figure 1 (c )). Epiproct roughly rod-like, apex slightly enlarged a downcurved hook, heavily sclerotised ventrolaterally, apex with an enlarged anterior membranous Lateral stylet blade-like and oriental anteriorly, apex sharp and slightly downcast (Figures 1 (c, d )). mostly membranous, basal two-thirds robust with two lateral extended lobes before tapering apical where brown spinules occur. In lateral view the aedeagus slender and tubular, apical portion slightly dorsally (Figure 2 (a – c )). Female. Unknown. Etymology The specific name refers to the hemitergal lobe being yellow, which differs dramatically hemiterga. Remarks The only Chinese species of Skwala, S. compacta (McLachlan, 1872), is also a dark species a short hemitergal lobe, but the new species has an unusual narrower pronotal band than S. compacta. The Japanese species, Skwala natorii Chino, 1999, also has an anteriorly hooked Distribution The new species is currently known only from Shaanxi Province, China.	en	Zhang, Yao, Li, Wen-Liang, Introduction, Wei-Hai Li (2025): Two new species of Skwala (Plecoptera: Perlodidae) from. Journal of Natural History 59 (33 - 36): 2127-2132, DOI: 10.1080/00222933.2025.2536602, URL: https://doi.org/10.1080/00222933.2025.2536602
0397A14B5F7C5A3CFDCDFF54FAE13721.taxon	materials_examined	Type material Holotype. 1 ♂, China: Jilin Province: Linjiang County, 338 m, 41.817 ° N, 126.902 ° E, 26 April 1955 Unknown (IZCAS). Paratype. 2 ♂♂, the same data as the holotype.	en	Zhang, Yao, Li, Wen-Liang, Introduction, Wei-Hai Li (2025): Two new species of Skwala (Plecoptera: Perlodidae) from. Journal of Natural History 59 (33 - 36): 2127-2132, DOI: 10.1080/00222933.2025.2536602, URL: https://doi.org/10.1080/00222933.2025.2536602
0397A14B5F7C5A3CFDCDFF54FAE13721.taxon	diagnosis	Diagnosis General colour dark brown. Hemitergal lobe elongate, medially constricted, apex swollen and small setae. Epiproct with an enlarged anterior membranous lobe near apex. Aedeagus with two dorsolateral lobes at base, apex tapering and ventrally hooked.	en	Zhang, Yao, Li, Wen-Liang, Introduction, Wei-Hai Li (2025): Two new species of Skwala (Plecoptera: Perlodidae) from. Journal of Natural History 59 (33 - 36): 2127-2132, DOI: 10.1080/00222933.2025.2536602, URL: https://doi.org/10.1080/00222933.2025.2536602
0397A14B5F7C5A3CFDCDFF54FAE13721.taxon	description	Description Adult habitus. (Figure 3 (a )). General colour dark brown. Head mostly brown, with a brown triangular originating from between posterior ocelli extending to centre of occiput. M-line brown. Antennae palpi yellow brown. Pronotum disc with dark brown rugosities and a wide, brown, longitudinal disjunct with thin black midline, lateral margins paler than disc (Figure 3 (a )). Legs brown. Wings veins brown. Cerci brown. Male. (Figure 3 (b – f )). Fore wing length 14.7 – 14.9 mm, hind wing length 13.6 – 13.8 mm (N = 2). bearing two patches of sensilla on the posterior margin, separated by a triangular medial notched area covers half length of tergum 9 (Figure 3 (b )). Tergum 10 divided into two hemiterga. Hemitergal lobe in view short, extended posteriorly, medially constricted, apex swollen and covered by small setae, in view strongly curved upward, the lobe seemingly very elongate after being cleared (Figure Paragenital plate band-like and flat (Figure 3 (c )). Basal anchor large and triangular in dorsal view and ventrally after the cowl being everted. Lateral arm large and arrow-like in dorsal view (Figure 3 (c )). stout and gently curved and connected to the epiproct apically (Figure 3 (c, d )). Epiproct mostly sclerotised, apex with an enlarged anterior membranous lobe. Lateral stylet sclerotised, band-like with apex. Aedeagus mostly membranous with two rounded, dorsolateral lobes at base, medially dorsal view then tapering to a ventrally curved apex which is hooked in lateral view; the area posterior dorsolateral lobe mostly covered by dense, dark spinules of the same size. Female. Unknown.	en	Zhang, Yao, Li, Wen-Liang, Introduction, Wei-Hai Li (2025): Two new species of Skwala (Plecoptera: Perlodidae) from. Journal of Natural History 59 (33 - 36): 2127-2132, DOI: 10.1080/00222933.2025.2536602, URL: https://doi.org/10.1080/00222933.2025.2536602
0397A14B5F7C5A3CFDCDFF54FAE13721.taxon	etymology	Etymology The species epithet refers to the elongate hemitergal lobe which can be used to distinguish this new from its congeners and is presumed to be atypical for this genus.	en	Zhang, Yao, Li, Wen-Liang, Introduction, Wei-Hai Li (2025): Two new species of Skwala (Plecoptera: Perlodidae) from. Journal of Natural History 59 (33 - 36): 2127-2132, DOI: 10.1080/00222933.2025.2536602, URL: https://doi.org/10.1080/00222933.2025.2536602
0397A14B5F7C5A3CFDCDFF54FAE13721.taxon	discussion	Remarks The new species is similar to Skwala americana (Klapálek, 1912). However, the epiproct of S. americana a bilobed anterior membranous area (see figs. 19 – 23 in Frison 1936), while in the new species, the membranous lobe is not divided. The elongate hemitergal lobe of the new species is easily distinguishable from those of other congeners. In addition, the lobe and the shape of the aedeagus are reminiscent new species of Arcynopteryx dichroa (McLachlan, 1872). However, Skwala longiloba lacks a slender needle-like stylet of the epiproct which is present in A. dichroa (see Teslenko 2012, figs. 2 and 5). HISTORY	en	Zhang, Yao, Li, Wen-Liang, Introduction, Wei-Hai Li (2025): Two new species of Skwala (Plecoptera: Perlodidae) from. Journal of Natural History 59 (33 - 36): 2127-2132, DOI: 10.1080/00222933.2025.2536602, URL: https://doi.org/10.1080/00222933.2025.2536602
