taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
0397DE379E056646623FF98070C0F836.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15655723/files/figure.png	https://doi.org/10.5281/zenodo.15655723	FIG. 2.— Photographs of live animals of the Gymnodoris nigricolor species complex: A, B, Gymnodoris nigricolor Baba, 1960, specimen MNHN-IM-2019-26208, isolate JD67 (A), specimen MNHN-IM-2019-26209, isolate JD27 (B); C-J, Gymnodoris boussionae n. sp. specimen MNHN-IM-2019-26204, isolate JD23 (C), specimen MNHN-IM-2019-26203, isolate JD35 (D), specimen MNHN-IM-2019-26206, isolate JD41 (E), specimen MNHN-IM-2019-26200, isolate JD43 (F), specimen MNHN-IM-2019-26205, isolate JD39 (G), specimen MNHN-IM-2019-26299, isolate JD51 (H), specimen MNHN-IM-2019-26207, isolate JD44 (I), specimen MNHN-IM-2019-26201, isolate JD10 (J); K, Gymnodoris nagoensis n. sp., specimen from Sunabe,Okinawa Prefecture,Japan,NSMT-Mo 79541,photo H.Tokura.	FIG. 2.— Photographs of live animals of the Gymnodoris nigricolor species complex: A, B, Gymnodoris nigricolor Baba, 1960, specimen MNHN-IM-2019-26208, isolate JD67 (A), specimen MNHN-IM-2019-26209, isolate JD27 (B); C-J, Gymnodoris boussionae n. sp. specimen MNHN-IM-2019-26204, isolate JD23 (C), specimen MNHN-IM-2019-26203, isolate JD35 (D), specimen MNHN-IM-2019-26206, isolate JD41 (E), specimen MNHN-IM-2019-26200, isolate JD43 (F), specimen MNHN-IM-2019-26205, isolate JD39 (G), specimen MNHN-IM-2019-26299, isolate JD51 (H), specimen MNHN-IM-2019-26207, isolate JD44 (I), specimen MNHN-IM-2019-26201, isolate JD10 (J); K, Gymnodoris nagoensis n. sp., specimen from Sunabe,Okinawa Prefecture,Japan,NSMT-Mo 79541,photo H.Tokura.	2025-06-12	Souza-Canal, Jade De;Ángel, Rie Nakano;Valdés, Ángel		Zenodo	biologists	Souza-Canal, Jade De;Ángel, Rie Nakano;Valdés, Ángel			
0397DE379E056646623FF98070C0F836.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15655725/files/figure.png	https://doi.org/10.5281/zenodo.15655725	FIG. 3.— SEM of the radula of the Gymnodoris nigricolor species complex: A, B, Gymnodoris nigricolor Baba, 1960, specimen MNHN-IM-2019-26209, isolate JD27, innermost teeth (A) and outer lateral teeth (B); C-D, Gymnodoris boussionae n. sp., specimen MNHN-IM-2019-26203, isolate JD35, innermost teeth (C) and outer lateral teeth (D); E-F, Gymnodoris nagoensis n. sp., specimen from Sunabe, Okinawa Prefecture, Japan, NSMT-Mo 79541, innermost teeth (E) and outer lateral teeth (F). Scale bars: A, 30 μm; B, D, 20 μm; C, 30 μm; E, F, 10 μm.	FIG. 3.— SEM of the radula of the Gymnodoris nigricolor species complex: A, B, Gymnodoris nigricolor Baba, 1960, specimen MNHN-IM-2019-26209, isolate JD27, innermost teeth (A) and outer lateral teeth (B); C-D, Gymnodoris boussionae n. sp., specimen MNHN-IM-2019-26203, isolate JD35, innermost teeth (C) and outer lateral teeth (D); E-F, Gymnodoris nagoensis n. sp., specimen from Sunabe, Okinawa Prefecture, Japan, NSMT-Mo 79541, innermost teeth (E) and outer lateral teeth (F). Scale bars: A, 30 μm; B, D, 20 μm; C, 30 μm; E, F, 10 μm.	2025-06-12	Souza-Canal, Jade De;Ángel, Rie Nakano;Valdés, Ángel		Zenodo	biologists	Souza-Canal, Jade De;Ángel, Rie Nakano;Valdés, Ángel			
0397DE379E056646623FF98070C0F836.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15655729/files/figure.png	https://doi.org/10.5281/zenodo.15655729	FIG. 4.— Drawings of the reproductive system of the Gymnodoris nigricolor species complex: A, B, Gymnodoris nigricolor Baba, 1960, specimen MNHN- IM-2019-26209, isolate JD27, reproductive system (A) and penial spines (B); C, D, Gymnodoris boussionae n. sp., specimen MNHN-IM-2019-26203, isolate JD35, reproductive system (C) and penial spines (D); E, F, Gymnodoris nagoensis n. sp. specimen from Sunabe, Okinawa Prefecture, Japan, NSMT-Mo 79541, reproductive system (E) and penial spines (F). Scale bars: A, C, E, 1 mm; B, D, F, 100 μm. Abbreviations: see Material and methods.	FIG. 4.— Drawings of the reproductive system of the Gymnodoris nigricolor species complex: A, B, Gymnodoris nigricolor Baba, 1960, specimen MNHN- IM-2019-26209, isolate JD27, reproductive system (A) and penial spines (B); C, D, Gymnodoris boussionae n. sp., specimen MNHN-IM-2019-26203, isolate JD35, reproductive system (C) and penial spines (D); E, F, Gymnodoris nagoensis n. sp. specimen from Sunabe, Okinawa Prefecture, Japan, NSMT-Mo 79541, reproductive system (E) and penial spines (F). Scale bars: A, C, E, 1 mm; B, D, F, 100 μm. Abbreviations: see Material and methods.	2025-06-12	Souza-Canal, Jade De;Ángel, Rie Nakano;Valdés, Ángel		Zenodo	biologists	Souza-Canal, Jade De;Ángel, Rie Nakano;Valdés, Ángel			
0397DE379E0866456213FB867178FA50.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15655723/files/figure.png	https://doi.org/10.5281/zenodo.15655723	FIG. 2.— Photographs of live animals of the Gymnodoris nigricolor species complex: A, B, Gymnodoris nigricolor Baba, 1960, specimen MNHN-IM-2019-26208, isolate JD67 (A), specimen MNHN-IM-2019-26209, isolate JD27 (B); C-J, Gymnodoris boussionae n. sp. specimen MNHN-IM-2019-26204, isolate JD23 (C), specimen MNHN-IM-2019-26203, isolate JD35 (D), specimen MNHN-IM-2019-26206, isolate JD41 (E), specimen MNHN-IM-2019-26200, isolate JD43 (F), specimen MNHN-IM-2019-26205, isolate JD39 (G), specimen MNHN-IM-2019-26299, isolate JD51 (H), specimen MNHN-IM-2019-26207, isolate JD44 (I), specimen MNHN-IM-2019-26201, isolate JD10 (J); K, Gymnodoris nagoensis n. sp., specimen from Sunabe,Okinawa Prefecture,Japan,NSMT-Mo 79541,photo H.Tokura.	FIG. 2.— Photographs of live animals of the Gymnodoris nigricolor species complex: A, B, Gymnodoris nigricolor Baba, 1960, specimen MNHN-IM-2019-26208, isolate JD67 (A), specimen MNHN-IM-2019-26209, isolate JD27 (B); C-J, Gymnodoris boussionae n. sp. specimen MNHN-IM-2019-26204, isolate JD23 (C), specimen MNHN-IM-2019-26203, isolate JD35 (D), specimen MNHN-IM-2019-26206, isolate JD41 (E), specimen MNHN-IM-2019-26200, isolate JD43 (F), specimen MNHN-IM-2019-26205, isolate JD39 (G), specimen MNHN-IM-2019-26299, isolate JD51 (H), specimen MNHN-IM-2019-26207, isolate JD44 (I), specimen MNHN-IM-2019-26201, isolate JD10 (J); K, Gymnodoris nagoensis n. sp., specimen from Sunabe,Okinawa Prefecture,Japan,NSMT-Mo 79541,photo H.Tokura.	2025-06-12	Souza-Canal, Jade De;Ángel, Rie Nakano;Valdés, Ángel		Zenodo	biologists	Souza-Canal, Jade De;Ángel, Rie Nakano;Valdés, Ángel			
0397DE379E0866456213FB867178FA50.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15655725/files/figure.png	https://doi.org/10.5281/zenodo.15655725	FIG. 3.— SEM of the radula of the Gymnodoris nigricolor species complex: A, B, Gymnodoris nigricolor Baba, 1960, specimen MNHN-IM-2019-26209, isolate JD27, innermost teeth (A) and outer lateral teeth (B); C-D, Gymnodoris boussionae n. sp., specimen MNHN-IM-2019-26203, isolate JD35, innermost teeth (C) and outer lateral teeth (D); E-F, Gymnodoris nagoensis n. sp., specimen from Sunabe, Okinawa Prefecture, Japan, NSMT-Mo 79541, innermost teeth (E) and outer lateral teeth (F). Scale bars: A, 30 μm; B, D, 20 μm; C, 30 μm; E, F, 10 μm.	FIG. 3.— SEM of the radula of the Gymnodoris nigricolor species complex: A, B, Gymnodoris nigricolor Baba, 1960, specimen MNHN-IM-2019-26209, isolate JD27, innermost teeth (A) and outer lateral teeth (B); C-D, Gymnodoris boussionae n. sp., specimen MNHN-IM-2019-26203, isolate JD35, innermost teeth (C) and outer lateral teeth (D); E-F, Gymnodoris nagoensis n. sp., specimen from Sunabe, Okinawa Prefecture, Japan, NSMT-Mo 79541, innermost teeth (E) and outer lateral teeth (F). Scale bars: A, 30 μm; B, D, 20 μm; C, 30 μm; E, F, 10 μm.	2025-06-12	Souza-Canal, Jade De;Ángel, Rie Nakano;Valdés, Ángel		Zenodo	biologists	Souza-Canal, Jade De;Ángel, Rie Nakano;Valdés, Ángel			
0397DE379E0866456213FB867178FA50.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15655729/files/figure.png	https://doi.org/10.5281/zenodo.15655729	FIG. 4.— Drawings of the reproductive system of the Gymnodoris nigricolor species complex: A, B, Gymnodoris nigricolor Baba, 1960, specimen MNHN- IM-2019-26209, isolate JD27, reproductive system (A) and penial spines (B); C, D, Gymnodoris boussionae n. sp., specimen MNHN-IM-2019-26203, isolate JD35, reproductive system (C) and penial spines (D); E, F, Gymnodoris nagoensis n. sp. specimen from Sunabe, Okinawa Prefecture, Japan, NSMT-Mo 79541, reproductive system (E) and penial spines (F). Scale bars: A, C, E, 1 mm; B, D, F, 100 μm. Abbreviations: see Material and methods.	FIG. 4.— Drawings of the reproductive system of the Gymnodoris nigricolor species complex: A, B, Gymnodoris nigricolor Baba, 1960, specimen MNHN- IM-2019-26209, isolate JD27, reproductive system (A) and penial spines (B); C, D, Gymnodoris boussionae n. sp., specimen MNHN-IM-2019-26203, isolate JD35, reproductive system (C) and penial spines (D); E, F, Gymnodoris nagoensis n. sp. specimen from Sunabe, Okinawa Prefecture, Japan, NSMT-Mo 79541, reproductive system (E) and penial spines (F). Scale bars: A, C, E, 1 mm; B, D, F, 100 μm. Abbreviations: see Material and methods.	2025-06-12	Souza-Canal, Jade De;Ángel, Rie Nakano;Valdés, Ángel		Zenodo	biologists	Souza-Canal, Jade De;Ángel, Rie Nakano;Valdés, Ángel			
0397DE379E0966426218FA607219FE2D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15655723/files/figure.png	https://doi.org/10.5281/zenodo.15655723	FIG. 2.— Photographs of live animals of the Gymnodoris nigricolor species complex: A, B, Gymnodoris nigricolor Baba, 1960, specimen MNHN-IM-2019-26208, isolate JD67 (A), specimen MNHN-IM-2019-26209, isolate JD27 (B); C-J, Gymnodoris boussionae n. sp. specimen MNHN-IM-2019-26204, isolate JD23 (C), specimen MNHN-IM-2019-26203, isolate JD35 (D), specimen MNHN-IM-2019-26206, isolate JD41 (E), specimen MNHN-IM-2019-26200, isolate JD43 (F), specimen MNHN-IM-2019-26205, isolate JD39 (G), specimen MNHN-IM-2019-26299, isolate JD51 (H), specimen MNHN-IM-2019-26207, isolate JD44 (I), specimen MNHN-IM-2019-26201, isolate JD10 (J); K, Gymnodoris nagoensis n. sp., specimen from Sunabe,Okinawa Prefecture,Japan,NSMT-Mo 79541,photo H.Tokura.	FIG. 2.— Photographs of live animals of the Gymnodoris nigricolor species complex: A, B, Gymnodoris nigricolor Baba, 1960, specimen MNHN-IM-2019-26208, isolate JD67 (A), specimen MNHN-IM-2019-26209, isolate JD27 (B); C-J, Gymnodoris boussionae n. sp. specimen MNHN-IM-2019-26204, isolate JD23 (C), specimen MNHN-IM-2019-26203, isolate JD35 (D), specimen MNHN-IM-2019-26206, isolate JD41 (E), specimen MNHN-IM-2019-26200, isolate JD43 (F), specimen MNHN-IM-2019-26205, isolate JD39 (G), specimen MNHN-IM-2019-26299, isolate JD51 (H), specimen MNHN-IM-2019-26207, isolate JD44 (I), specimen MNHN-IM-2019-26201, isolate JD10 (J); K, Gymnodoris nagoensis n. sp., specimen from Sunabe,Okinawa Prefecture,Japan,NSMT-Mo 79541,photo H.Tokura.	2025-06-12	Souza-Canal, Jade De;Ángel, Rie Nakano;Valdés, Ángel		Zenodo	biologists	Souza-Canal, Jade De;Ángel, Rie Nakano;Valdés, Ángel			
0397DE379E0966426218FA607219FE2D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15655725/files/figure.png	https://doi.org/10.5281/zenodo.15655725	FIG. 3.— SEM of the radula of the Gymnodoris nigricolor species complex: A, B, Gymnodoris nigricolor Baba, 1960, specimen MNHN-IM-2019-26209, isolate JD27, innermost teeth (A) and outer lateral teeth (B); C-D, Gymnodoris boussionae n. sp., specimen MNHN-IM-2019-26203, isolate JD35, innermost teeth (C) and outer lateral teeth (D); E-F, Gymnodoris nagoensis n. sp., specimen from Sunabe, Okinawa Prefecture, Japan, NSMT-Mo 79541, innermost teeth (E) and outer lateral teeth (F). Scale bars: A, 30 μm; B, D, 20 μm; C, 30 μm; E, F, 10 μm.	FIG. 3.— SEM of the radula of the Gymnodoris nigricolor species complex: A, B, Gymnodoris nigricolor Baba, 1960, specimen MNHN-IM-2019-26209, isolate JD27, innermost teeth (A) and outer lateral teeth (B); C-D, Gymnodoris boussionae n. sp., specimen MNHN-IM-2019-26203, isolate JD35, innermost teeth (C) and outer lateral teeth (D); E-F, Gymnodoris nagoensis n. sp., specimen from Sunabe, Okinawa Prefecture, Japan, NSMT-Mo 79541, innermost teeth (E) and outer lateral teeth (F). Scale bars: A, 30 μm; B, D, 20 μm; C, 30 μm; E, F, 10 μm.	2025-06-12	Souza-Canal, Jade De;Ángel, Rie Nakano;Valdés, Ángel		Zenodo	biologists	Souza-Canal, Jade De;Ángel, Rie Nakano;Valdés, Ángel			
0397DE379E0966426218FA607219FE2D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15655729/files/figure.png	https://doi.org/10.5281/zenodo.15655729	FIG. 4.— Drawings of the reproductive system of the Gymnodoris nigricolor species complex: A, B, Gymnodoris nigricolor Baba, 1960, specimen MNHN- IM-2019-26209, isolate JD27, reproductive system (A) and penial spines (B); C, D, Gymnodoris boussionae n. sp., specimen MNHN-IM-2019-26203, isolate JD35, reproductive system (C) and penial spines (D); E, F, Gymnodoris nagoensis n. sp. specimen from Sunabe, Okinawa Prefecture, Japan, NSMT-Mo 79541, reproductive system (E) and penial spines (F). Scale bars: A, C, E, 1 mm; B, D, F, 100 μm. Abbreviations: see Material and methods.	FIG. 4.— Drawings of the reproductive system of the Gymnodoris nigricolor species complex: A, B, Gymnodoris nigricolor Baba, 1960, specimen MNHN- IM-2019-26209, isolate JD27, reproductive system (A) and penial spines (B); C, D, Gymnodoris boussionae n. sp., specimen MNHN-IM-2019-26203, isolate JD35, reproductive system (C) and penial spines (D); E, F, Gymnodoris nagoensis n. sp. specimen from Sunabe, Okinawa Prefecture, Japan, NSMT-Mo 79541, reproductive system (E) and penial spines (F). Scale bars: A, C, E, 1 mm; B, D, F, 100 μm. Abbreviations: see Material and methods.	2025-06-12	Souza-Canal, Jade De;Ángel, Rie Nakano;Valdés, Ángel		Zenodo	biologists	Souza-Canal, Jade De;Ángel, Rie Nakano;Valdés, Ángel			
0397DE379E0966426218FA607219FE2D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15655719/files/figure.png	https://doi.org/10.5281/zenodo.15655719	FIG. 1.— A, histogram representing the distance plot for the ASAP analysis using the COI gene showing pairwise p-distances; B, consensus phylogenetic tree of specimens sequenced based on Bayesian analysis of the three concatenated loci. Posterior probabilities from Bayesian inference (BI) are given above branches and percentages corresponding to maximum likelihood bootstrap support are given below branches. Supported nodes are highlighted in blue, not supported nodes are highlighted in orange. Results of two species delimitation analyses are indicated by vertical bars.	FIG. 1.— A, histogram representing the distance plot for the ASAP analysis using the COI gene showing pairwise p-distances; B, consensus phylogenetic tree of specimens sequenced based on Bayesian analysis of the three concatenated loci. Posterior probabilities from Bayesian inference (BI) are given above branches and percentages corresponding to maximum likelihood bootstrap support are given below branches. Supported nodes are highlighted in blue, not supported nodes are highlighted in orange. Results of two species delimitation analyses are indicated by vertical bars.	2025-06-12	Souza-Canal, Jade De;Ángel, Rie Nakano;Valdés, Ángel		Zenodo	biologists	Souza-Canal, Jade De;Ángel, Rie Nakano;Valdés, Ángel			
