taxonID	type	description	language	source
039A87F0FFF4FFD0FF07F7633B3EB1F3.taxon	materials_examined	Holotype: (♁) Crefeld 14.6.10 Puhlmann; Type ♁; Scopesis areolaris PFANK. ♁ (Berlin), type examined. D e s c r i p t i o n: ♁. Body length 9 mm. Flagellum with 40 flagellomeres; 1 st flagellar segment 6.0 × as long as wide and 1.8 × as long as 2 nd segment; 2 nd segment 3.3 × and 3 rd segment 2.8 × as long as wide, preapical segment c 2 × as long as wide. Temple moderately and roundly narrowed behind eye. OED 1.4 × ocellar diameter. Face granulated and with superficial punctures, dull. Clypeus distinctly convex, slightly bulging medially, with coarse dense punctures and some long setae; apical margin slightly rounded, with very small flat extensions laterally. Mandibular teeth of equal size. Mesoscutum finely granulate, ± shining. Notaulus impressed in frontal 0.3 of mesoscutum. Mesopleuron granulate and superficially punctate, with rather strong striae dorsally, dull. Speculum with microsculpture, ± shining. Area superomedia with incomplete divergent lateral carinae. Area petiolaris with complete carina. Lateral longitudinal carina of propodeum weak and partly obsolete; pleural carina distinct and complete. Hind femur 4.7 × as long as wide. Inner apical spur of hind tibia 0.65 × as long as hind metatarsus. Vein 2 rs-m 0.9 × as long as vein M between 2 rs-m and 2 m-cu. Vein 1 cu-a postfurcal by 2 × its width. Nervellus of hind wing inclivous, intercepted in its apical 0.7. 1 st tergite 2.0 × as long as wide, without latero-median carina. 2 nd tergite 1.2 × as wide as long. Color: black. Scape ventrally and flagellum reddish, basal flagellomeres blackish dorsally. Palps partly, mandible except teeth, clypeus, facial orbit and two paramedian dorsal spots on face yellowish. Tegula reddish-brown, subtegular ridge reddish. Apical 0.2 of 1 st tergite, 2 nd to 4 th tergites entirely and 5 th tergite dorsally red, laterally black. Coxae and trochanters blackish. Trochantelli reddish; legs otherwise red; hind femur red, black in apical 0.1; hind tibia yellowish, black in apical 0.4; hind tarsus black. Pterostigma chestnut-brown. ♀ and hosts unknown. D i s t r i b u t i o n: Only known from the German holotype. T a x o n o m i c a l r e m a r k: This species belongs to Scopesis (e. g. typical length of 1 st and 2 nd flagellar segments, form of areolet and structure of 1 st tergite) but is characterized by its unique facial color pattern and unusual structure of the clypeus which resembles Cuboscopesis fraternus (HOLMGREN). We did not find any ♀♀ with this form of clypeus in the studied Scopesis material, the ♀ of Sc. areolaris remains unknown.	en	Riedel, Matthias, H, Andres, Schmidt, Stefan (2024): Revision of the Palaearctic species of Scopesis FÖRSTER (Hymenoptera, Ichneumonidae, Ctenopelmatinae). Linzer biologische Beiträge 56 (1): 201-252, DOI: 10.5281/zenodo.14681817
039A87F0FFF7FFD1FF07F65938F4B6D0.taxon	materials_examined	Holotype: (♀) (Wroclaw), type examined. syn. Mesoleius praecatorius HOLMGREN, 1876: 48, ♀, synonymized by TEUNISSEN (1945: 220), type not examined. D e s c r i p t i o n: ♀. Body length 8.0 – 10.0 mm. Flagellum with 41 – 42 segments; 1 st flagellar segment 5.0 – 5.3 × as long as wide and 1.9 × as long as 2 nd segment; 2 nd segment 2.5 – 2.7 ×, 3 rd segment 2.3 – 2.4 × and preapical segment 1.5 × as long as wide. Temple moderately and roundly narrowed behind eye. OED 1.3 × ocellar diameter. Face granulated, at most with some sparse fine and superficial punctures. Clypeus slightly convex, not bulging medially, with sparse rather coarse punctures, shining; apical margin almost straight, with small flat extensions laterally. Ventral mandibular tooth slightly larger than dorsal tooth. Mesoscutum granulate, dull. Notaulus not distinctly impressed. Mesopleuron granulate or finely rugose or partly striate ventrally, with rather strong striae reaching frontal margin of mesopleuron dorsally. Speculum finely granulate, rather dull. Area superomedia sometimes with weak incomplete lateral carina. Area petiolaris usually with complete carina. Lateral longitudinal carina of propodeum sometimes weak or partly obsolete, pleural carina distinct and complete. Hind femur 4.5 – 4.7 × as long as wide. Inner apical spur of hind tibia 0.62 × as long as hind metatarsus. Vein 2 rs-m 0.65 – 0.85 × as long as vein M between 2 rs-m and 2 m-cu. Vein 1 cu-a interstitial or postfurcal by 1 – 2 × its width. Nervellus of hind wing slightly inclivous, intercepted in its apical 0.7 – 0.8. 1 st tergite 1.4 – 1.6 × as long as wide, without latero-median carina. 2 nd tergite 1.2 – 1.4 × as wide as long. Color: black. Flagellum reddish ventrally, brownish dorsally, black basally. Palps partly, spot on mandible, apical 2 / 3 of clypeus reddish. Tegula black or yellow. 1 st to 4 th tergites red, sometimes 1 st tergite ± black. Coxae, trochanters and trochantelli black; legs red; hind femur entirely black; hind tibia black basally and in apical 0.25; hind tarsomeres black, with paler bases. Pterostigma ochreous. The ♀♀ have two color patterns: a) tegula yellow: mid trochanter and mid and hind trochantelli with yellow spots; b) tegula black: all trochanters and trochantelli black. ♁. Body length 7.5 - 10.0 mm. Flagellum with 40 – 44 segments; 1 st flagellar segment 5.2 – 5.7 × as long as wide and 1.9 – 2.0 × as long as 2 nd segment; 2 nd segment 2.5 – 2.9 × and 3 rd segment 2.3 – 2.6 × as long as wide. Temple moderately and roundly narrowed behind eye. OED 1.4 × ocellar diameter. MI 0.3. Clypeus moderately convex, not bulging. Mesoscutum with fine granulation, often ± shining. Mesopleuron granulate or finely rugose or partly striate ventrally, with rather strong striae dorsally. Speculum finely granulate, rather dull. Mesopleuron granulate or finely rugose or partly striate ventrally, with rather strong striae reaching frontal margin of mesopleuron dorsally. Speculum finely granulate, rather dull. Hind femur 4.5 × as long as wide. Inner apical spur of hind tibia 0.59 × as long as hind metatarsus. 1 st tergite 1.8 – 2.2 × as long as wide. 2 nd tergite 0.9 – 1.1 × as long as wide. Color: black. Palps, mandible except teeth, clypeus, face, ventral spot at scape, often tegula tegula, subtegular ridge, sometimes apical spot of fore coxa, fore and mid trochanters partly and all trochantelli ± yellow (hind trochantellus sometimes black). 1 st tergite usually black. 2 nd to 4 th tergites red. Hind tibia yellowish, infuscate in apical 0.4. Host: Tenthredopsis spec. (Hym. Tenthredinidae) (ZSM). D i s t r i b u t i o n: West Palaearctic, studied material from Bulgaria, Germany, Italy, Spain, Switzerland, and United Kingdom. T a x o n o m i c a l r e m a r k: We have seen a ♁ from Kamchatka determined as bicolor (ROMAN 1931). It belongs to Scopesis sachalinensis (UCHIDA). Sc. bicolor has not been found in the Eastern Palaearctic region so far.	en	Riedel, Matthias, H, Andres, Schmidt, Stefan (2024): Revision of the Palaearctic species of Scopesis FÖRSTER (Hymenoptera, Ichneumonidae, Ctenopelmatinae). Linzer biologische Beiträge 56 (1): 201-252, DOI: 10.5281/zenodo.14681817
039A87F0FFF6FFDFFF07F6BF390DB390.taxon	materials_examined	Holotype: (♀) SW Georgia, Kintrishi NP, 1634 m asl, 41.44535308 ° N 41.05384144 ° E, MT 12, 10 - 24. VIII. 2018, leg. G. Japoshvili (Tbilisi). Paratypes: (♀) SW Georgia, Kintrishi NP, 1634 m asl, 41.44535308 ° N 41.05384144 ° E, MT 12, 10 - 24. VIII. 2018, leg. G. Japoshvili (coll. RIEDEL); (♀) SW Georgia, Kintrishi NP, 1264 m asl, 41.44388824 ° N 41.0502904 ° E, MT 4, 13 - 27. VII. 2018, leg. G. Japoshvili (Tbilisi); (♁) Georgien, Lagodekhi NP, H 4 - 5, 15 - 27. IX. 2014; [green] BC-ZSM-HYM- 27496 - E 01 (ZSM); (♁) Georgien, Lagodekhi NP, H 4 - 5, 15 - 27. IX. 2014; [green] BC-ZSM-HYM- 27496 - E 03 (ZSM); (♀ ♁) Georgien, Lagodekhi NP, H 4 - 5, 15 - 27. IX. 2014; [green] BC-ZSM-HYM- 27496 - D 08 (ZSM); (♁) Georgien, Lagodekhi NP, H 4 - 5, 15 - 25. VII. 2014; [green] BC-ZSM-HYM- 27496 - D 06 (ZSM). D e s c r i p t i o n: ♀. Body length 9.0 – 9.5 mm. Flagellum with 36 – 39 segments; 1 st flagellar segment 4.8 – 5.5 × as long as wide and 1.8 – 1.9 × as long as 2 nd segment; 2 nd segment 2.8 – 2.9 ×, 3 rd segment 2.3 – 2.5 × and preapical segment 1.6 × as long as wide. Temple slightly and roundly narrowed behind eye. OED 1.5 × ocellar diameter. Face granulated, with fine superficial punctures. Clypeus slightly convex, not bulging medially, with sparse punctures, shining; apical margin with small flat extensions laterally, slightly concave. Mandibular teeth of equal length. MI 0.4. Mesoscutum granulate and superficially punctate, dull. Notaulus weakly impressed in frontal half of mesoscutum. Mesopleuron finely rugose and ± punctate ventrally and usually with distinct striae dorsally. Speculum with microsculpture, strongly shining. Area superomedia sometimes with weak but incomplete lateral carina. Area petiolaris with complete carina. Lateral longitudinal carina and pleural carina of propodeum distinct and complete. Hind femur 4.3 – 4.9 × as long as wide. Hind tibia with numerous dark setae externally. Inner apical spur of hind tibia 0.60 × as long as hind metatarsus. Vein 2 rs-m 0.75 × as long as vein M between 2 rs-m and 2 m-cu. Vein 1 cu-a postfurcal by 1 – 2 × its width. Nervellus of hind wing vertical or slightly inclivous, intercepted in its apical 0.7 – 0.8. 1 st tergite 1.5 – 1.6 × as long as wide, without latero-median carina. 2 nd tergite 1.2 – 1.4 × as wide as long Color: black. Flagellum black, distal half ± reddish ventrally. Mandible black or reddish-brown centrally. Apical ½ – 3 / 4 of clypeus reddish or chestnut-brown. Tegula black. 1 st tergite ±, 2 nd to 4 th or 5 th tergites red, sometimes 6 th tergite reddish laterally. Coxae, trochanters and trochantelli black; legs otherwise red; hind femur red, not distinctly infuscate apically; hind tibia red, sometimes narrowly black basally, black in apical 0.2 – 0.3; apical spurs of hind tibia reddish or brown; hind tarsus black. Pterostigma dark ochreous or brown. ♁. Body length 6.5 – 8.5 mm. Flagellum with 34 – 37 segments; 1 st flagellar segment 5.5 – 5.9 × as long as wide and 1.9 × as long as 2 nd flagellomere; 2 nd segment 2.9 – 3.1 ×, 3 rd segment 2.6 × and preapical segment 1.8 × as long as wide. Temple parallel or slightly and roundly narrowed behind eye. OED 1.5 × and OOD 1.0 × ocellar diameter. Face granulated, without distinct punctures. Clypeus moderately and transversely convex, not bulging medially, with sparse punctures, shining; apical margin with small flat extensions laterally, slightly concave. Ventral mandibular tooth slightly larger than dorsal tooth. MI 0.5. Mesopleuron finely rugose and sometimes finely striate ventrally, usually with distinct striae dorsally. Speculum smooth or with microsculpture, strongly shining. Area superomedia sometimes with weak but incomplete lateral rugae in basal half. Area petiolaris with complete carina. Lateral longitudinal carina and pleural carina of propodeum distinct and complete. Hind femur 4.6 – 4.9 × as long as wide. Hind tibia with numerous setae externally. Inner apical spur of hind tibia 0.50 × as long as hind metatarsus. Vein 2 rs-m 0.7 – 0.8 × as long as vein M between 2 rs-m and 2 m-cu. Vein 1 cu-a postfurcal by 1 – 2 × its width. Nervellus of hind wing vertical or slightly inclivous, intercepted in its apical 0.7 – 0.8. 1 st tergite 1.8 – 2.0 × as long as wide, without latero-median carina. 2 nd tergite about as long as wide Color: black. Flagellum black, distal 2 / 3 ± reddish ventrally. Mandible except teeth reddish-yellowish. Clypeus and lateral spots or complete face cream-yellow. Mesosoma black. Tegula black. Tergites black, 2 nd and 3 rd tergites with ± extended reddish suffusion. Coxae, trochanters and trochantelli black; legs otherwise red; hind femur red, sometimes slightly infuscate apically; hind tibia red, sometimes narrowly black basally, black in apical 0.2 – 0.3; hind tarsus black. Pterostigma ochreous, with darker margins. D i s t r i b u t i o n: So far only known from Georgia. Taxonomical remarks: The ♀ of this new species resembles Scopesis rufolabris (ZETTERSTEDT), but differs by the darker coloration of flagellum, tegula and hind tibia, and numerous dark and stronger setae of hind tibia. The ♁ differs by the reduced red coloration of median tergites and cream-yellow facial coloration.	en	Riedel, Matthias, H, Andres, Schmidt, Stefan (2024): Revision of the Palaearctic species of Scopesis FÖRSTER (Hymenoptera, Ichneumonidae, Ctenopelmatinae). Linzer biologische Beiträge 56 (1): 201-252, DOI: 10.5281/zenodo.14681817
039A87F0FFF8FFDCFF07F3FF3B66B1DE.taxon	description	Lectotype: (♁) Uppsala Univ. Zool. Mus Thunbergsaml. nr. 25563 Ichneumon frontator Wg. By. Typ (Uppsala), examined and designated by HORSTMANN (2006: 81). syn. nov. Mesoleius (Scopesus) depressus THOMSON, 1894: 2030, ♀. Holotype: (♀) Col Ljgh; depressus (Lund), type examined by HORSTMANN. D e s c r i p t i o n: ♀. Body length (6.0) – 8.5 – 11 mm. Flagellum with (32) – 37 – 42 segments, stout; 1 st flagellar segment 4.4 – 5.3 × as long as wide and 1.9 – 2.3 × as long as 2 nd segment; 2 nd segment 2.0 – 2.5 ×, 3 rd segment 1.8 – 2.4 × and preapical segment 1.2 – 1.6 × as long as wide. Temple moderately and roundly narrowed. OED 1.2 – 1.5 × ocellar diameter. Face granulated, with superficial punctures, dull. Clypeus slightly transversely convex, with sparse punctures and shining, not bulging medially; apical margin slightly concave, without or with short flat extensions laterally. Ventral mandibular tooth equal to or slightly larger than dorsal tooth. MI 0.5 Mesoscutum granulate. Notaulus not or weakly impressed. Mesopleuron granulate or finely rugose, with superficial punctures and some fine striation ventrally, usually distinctly striate dorsally. Speculum with microsculpture, shining. Area superomedia without or with weak lateral carina. Area petiolaris usually with complete carina, sometimes obsolete medially. Lateral longitudinal carina and pleural carina of propodeum usually distinct and complete, sometimes weak. Hind femur 3.8 – 4.7 × as long as wide. Inner apical spur of hind tibia 0.62 × as long as hind metatarsus. Vein 2 rs-m 0.7 – 0.8 × as long as vein M between 2 rs-m and 2 m-cu. Vein 1 cu-a interstitial or postfurcal by 1 – 2 × its width. 1 st tergite 1.3 – 1.6 × as long as wide, without latero-median carina. 2 nd tergite 1.2 – 1.4 × as wide as long. Color: black. Flagellum red or ochreous, black to brownish basally and dorsally. Palps brown. Mandible except teeth and apical 1 / 2 – 3 / 4 of clypeus reddish or yellowish. Hind edge of pronotum brown. Tegula brown, rarely yellow. 1 st to 4 th tergites entirely red, 1 st tergite rarely ± black in basal 1 / 2 – 2 / 3. Coxae, trochanters and trochantelli black; legs otherwise red; hind femur entirely red; hind tibia reddish-yellow, sometimes narrowly infuscate basally, infuscate in apical 0.2 – 0.3; hind tarsus black. Pterostigma yellowish to ochreous. Var. ♀. One specimen from Haag, Bavaria (Hamburg): Coxae, trochanters and trochantelli black. Legs otherwise red; hind tibia reddish, not distinctly infuscate apically; hind tarsus reddish; distal tarsomeres of mid and hind tarsi brownish. ♁. Body length 8.0 – 11.5 mm. Flagellum with 36 – 41 segments; 1 st flagellar segment 4.3 – 5.5 × as long as wide and 1.9 – 2.1 × as long as 2 nd flagellomere; 2 nd segment 2.2 – 2.5 ×, 3 rd segment 2.0 – 2.3 × and preapical segment 1.7 × as long as wide. Clypeus slightly transversely convex; apical margin slightly concave and with short lateral extensions. Mesopleuron granulate and with superficial punctures ventrally, finely rugose medially, with fine striae dorsally, ± shining. Speculum with microsculpture, shining. Area superomedia usually with weak lateral carina in basal half. Area petiolaris with complete carina. Lateral longitudinal carina and pleural carina of propodeum distinct and complete. Hind femur 4.5 – 4.6 × as long as wide. 1 st tergite 1.7 – 2.0 × as long as wide, with latero-median carina in basal 0.2 – 0.4. Color: black. Flagellum black, sometimes ochreous in apical 2 / 3. Palps, mandible except teeth, clypeus, face and sometimes scape ventrally reddish-yellow or yellow. Hind edge of pronotum brown or narrowly yellowish. Tegula brown, with yellow spot frontally. 1 st tergite black, sometimes red apically, 2 nd to 4 th tergites entirely red. Coxae black, fore and mid coxae sometimes yellowish apically; fore and mid trochanters black dorsally, yellow ventrally; fore and mid trochantelli yellow; hind trochanter and trochantellus red or black; legs otherwise red; hind femur entirely red; hind tibia reddish-yellow, black in apical 0.4; hind tarsus black. Pterostigma ochreous centrally, margins darker. D i s t r i b u t i o n: Western Palaearctic; studied material from Bulgaria, Germany, Sweden and Turkey *. Taxonomical remarks: TEUNISSEN (1953) synonymized Scopesis bicolor (GRAVENHORST) with Scopesis frontator (THUNBERG) as a color variant with black hind femur, but these species also differ by the length of basal flagellomeres. The holotype of Mesoleius (Scopesus) depressus THOMSON was usually separated from Sc. frontator (THUNBERG) by the mainly black color of 1 st tergite and very short basal flagellomeres (2 nd flagellar segment 2.0 × and 3 rd segment 1.8 × as long as wide), but the coloration of the 1 st tergite and length of basal flagellar segments are variable in our material. We therefore synonymize Mesoleius depressus (THOMSON) with Sc. frontator (THUNBERG) here.	en	Riedel, Matthias, H, Andres, Schmidt, Stefan (2024): Revision of the Palaearctic species of Scopesis FÖRSTER (Hymenoptera, Ichneumonidae, Ctenopelmatinae). Linzer biologische Beiträge 56 (1): 201-252, DOI: 10.5281/zenodo.14681817
039A87F0FFFBFFDDFF07F1BC38F9B6A8.taxon	materials_examined	Holotype: (♁) (Uppsala), type not examined. syn. Tryphon nigricollis GRAVENHORST, 1829, synonymized by ROMAN (1912: 258), type not examined. syn. Tryphon longipes GRAVENHORST, 1829, synonymized by YU & HORSTMANN (1997: 437), type not examined. D e s c r i p t i o n: ♀. Body length 8 – 11 mm. Flagellum with 36 – 42 segments, slender; 1 st flagellar segment 4.6 – 5.0 × as long as wide and 1.8 × as long as 2 nd segment; 2 nd segment 2.4 – 2.8 ×, 3 rd segment 2.4 – 2.5 × and preapical segment c 1.8 × as long as wide. Temple distinctly and roundly narrowed. OED 1.4 – 1.5 × ocellar diameter. Face granulated, with superficial punctures, dull. Clypeus transversely convex and slightly bulging medially, with fine sparse punctures and some erected long setae, shining, with weak flat lateral extensions, apical margin slightly concave. Lower mandibular tooth slightly larger than upper tooth. MI 0.4. Mesoscutum finely granulate and superficially punctate, often ± shining. Notaulus weakly impressed frontally. Mesopleuron granulate and dull ventrally, with superficial punctures and ± shining dorsally, finely striate in dorsal 1 / 3, striae not reaching the frontal third of mesopleuron; speculum with microsculpture, shining. Area superomedia usually with strong lateral carina, divergent apically. Area petiolaris with complete carinae. Lateral longitudinal and pleural carinae of propodeum distinct and complete. Hind femur 4.4 – 5.2 × as long as wide. Inner apical spur of hind tibia 0.59 × as long as hind metatarsus. Vein 2 rs-m 0.8 – 0.9 × as long as vein M between 2 rs-m and 2 m-cu. Vein 1 cu-a postfurcal by 1 × its width. Nervellus of hind wing vertical, intercepted in its apical 0.7. 1 st tergite 1.4 – 1.6 × as long as wide, without latero-median carina. 2 nd tergite 1.1 – 1.4 × as wide as long. Color: black. Flagellum red, black basally. Palps, mandible except teeth, clypeus, often paramedian spots or stripes on face, sometimes spot on tegula yellow. Hind edge of pronotum, tegula and subtegular ridge and postscutellum sometimes reddish. 1 st to 4 th tergites yellow, 5 th tergite sometimes ± yellow or reddish. 2 nd to 4 th sternites yellow, 5 th sternite brown or black, hypopygium black. Coxae, trochanters and trochantelli black, trochanters and trochantelli often with yellow apical margins; fore and mid femora reddish, fore and mid tibiae and tarsi yellow; hind femur black, sometimes diffusely reddish-brown; hind tibia yellow, infuscate in apical 0.3, hind tarsus reddish or yellowish, ± brownish distally. Wings sometimes with yellowish tint. Pterostigma yellowish. ♁. Body length 9 – 10 mm. Flagellum with 35 – 40 segments; 1 st flagellar segment 4.8 – 6.0 × as long as wide and 1.9 – 2.0 × as long as 2 nd flagellomere; 2 nd segment 2.5 – 3.0 × and 3 rd segment 2.3 – 2.7 × longer than wide. OED 1.4 × ocellar diameter. Temple moderately and roundly narrowed behind eye. Mesopleuron granulate and superficially punctate, with few striae dorsally, ± shining. Hind femur 4.4 – 4.7 × as long as wide. 1 st tergite 1.7 – 2.0 × as long as wide, sometimes with weak latero-median carina in basal 0.4. 2 nd tergite 1.0 – 1.1 × as wide as long. Color: black. Flagellum ochreous or reddish, blackish basally. Palps, mandible except teeth, clypeus, face, ventral spot at scape, sometimes spot on pedicel, hind edge of pronotum and tegula, sometimes spot on subtegular ridge, anterolateral spots on mesoscutum and often spot on mesosternum yellow. 1 st to 4 th tergites usually yellowish or reddish-yellow, sometime 5 th tergite basally yellow. All coxae and hind trochanter black, often fore and mid coxae ± yellow; fore and mid trochanters and all trochantelli yellow. Legs yellowish, hind femur black, often narrowly yellow at base; hind tibia yellow, infuscate in apical 0.3, hind tarsus yellow or red. Pterostigma ochreous or reddish. D i s t r i b u t i o n: A Holarctic species; studied material from Austria, Bulgaria, France, Germany, Switzerland, and United Kingdom.	en	Riedel, Matthias, H, Andres, Schmidt, Stefan (2024): Revision of the Palaearctic species of Scopesis FÖRSTER (Hymenoptera, Ichneumonidae, Ctenopelmatinae). Linzer biologische Beiträge 56 (1): 201-252, DOI: 10.5281/zenodo.14681817
039A87F0FFFAFFDAFF07F69738A1B72E.taxon	description	Lectotype: (♁) Hlm / Bhn. (Stockholm), type examined by HORSTMANN. D e s c r i p t i o n: ♀. Body length 7.5 – 10.5 mm. Flagellum with 32 – 42 segments, slender; 1 st flagellar segment 5.0 – 6.3 × as long as wide and 1.8 – 2.0 × as long as 2 nd segment; 2 nd segment 2.7 – 3.2 ×, 3 rd segment 2.5 – 2.7 × and preapical segment 1.8 – 2.0 × as long as wide. Temple moderately and roundly narrowed. OED 1.5 × ocellar diameter. Face granulated and with superficial punctures. Clypeus slightly convex, not bulging medially, with sparse rather coarse punctures, with some erected setae, laterally with ± distinct flat extensions; apical margin ± concave. Mandibular teeth of equal length. MI 0.4 – 0.5. Mesoscutum granulate, with fine superficial punctures, dull. Notaulus impressed in frontal 0.3 of mesoscutum. Mesopleuron granulate, without or with superficial punctures, and weak fine striation dorsally; speculum with microsculpture, shining. Area superomedia usually without lateral carina, rarely with incomplete carination. Area petiolaris usually with obsolete carina medially. Lateral longitudinal carina of propodeum often weak and partly obsolete, pleural carina usually complete. Hind femur 3.9 – 4.7 × as long as wide. Inner apical spur of hind tibia 0.56 × as long as hind metatarsus. Vein 2 rs-m 0.7 – 1.0 × as long as vein M between 2 rs-m and 2 m-cu. Vein 1 cu-a postfurcal by 1 – 3 × its width. Nervellus of hind wing vertical or slightly inclivous, intercepted in its apical 0.6 – 0.7. 1 st tergite 1.5 – 1.6 × as long as wide, without latero-median carina. 2 nd tergite 1.2 – 1.4 × as wide as long. Color: black. Flagellum reddish or ochreous, black basally and / or dorsally. Palps, mandible except teeth and apical 1 / 2 – 3 / 4 of clypeus reddish. Tegula brown. Often 1 st tergite and usually 2 nd to 3 rd or 4 th tergites red (sometimes dark reddish). Coxae, trochanters and trochantelli black; legs otherwise reddish; fore and mid femora usually blackish basally; hind femur black; hind tibia yellowish, infuscate in apical 0.2 – 0.3; hind tarsus brown or black, hind metatarsus reddish basally. Pterostigma ochreous. Var. ♀. 1 specimen from United Kingdom (EDI) with very slender 1 st tergite, 1.9 × as long as wide. ♁. Body length 8 – 10 mm. Flagellum with 37 – 41 segments, slender; 1 st flagellar segment 5.4 – 6.0 × as long as wide and 1.8 – 2.0 × as long as 2 nd segment; 2 nd segment 2.8 – 3.2 × and 3 rd segment 2.3 – 2.9 × as long as wide. Temple distinctly roundly narrowed behind eye. OED 1.6 × ocellar diameter. MI 0.2 – 0.3. Mesopleuron granulate, with sparse punctures, dorsally usually with weak striae. Speculum with microsculpture, shining. Area superomedia usually without carina; area petiolaris sometimes closed medially. Hind femur 3.8 – 4.2 × as long as wide. Vein 2 rs-m 0.8 – 1.0 × as long as vein M between 2 rs-m and 2 m-cu. 1 st tergite 1.8 – 2.0 × as long as wide, without latero-median carina. 2 nd tergite 1.0 – 1.2 × as wide as long. Color: black. Flagellum yellow or reddish ventrally, ± brown dorsally. Palps, mandible except teeth, clypeus, face and large anterolateral spot on mesoscutum yellow. Tegula reddish or brown. 2 nd to 3 rd or 4 th tergites usually reddish. Coxae, trochanters and trochantelli black, sometimes fore coxa with reddish spot apically and / or fore and mid trochanters apically and trochantelli yellow. Legs otherwise red-yellow; mid femur basally and hind femur entirely black; hind tibia yellowish, infuscate in apical 0.3; hind tarsus blackish, metatarsus ochreous basally. Pterostigma yellow or ochreous. D i s t r i b u t i o n: Western Palaearctic, studied material from Austria, Bulgaria, Germany, Italy, Switzerland *, United Kingdom *.	en	Riedel, Matthias, H, Andres, Schmidt, Stefan (2024): Revision of the Palaearctic species of Scopesis FÖRSTER (Hymenoptera, Ichneumonidae, Ctenopelmatinae). Linzer biologische Beiträge 56 (1): 201-252, DOI: 10.5281/zenodo.14681817
039A87F0FFFDFFDBFF07F72C3B10B75E.taxon	description	Lectotype: (♀) Ostricourt (Lund), designated and examined by HORSTMANN (2006: 81). D e s c r i p t i o n: ♀. Body length 9.0 mm. Flagellum with 38 segments; 1 st flagellar segment 4.7 × as long as wide and 2.1 - 2.2 × as long as 2 nd segment; 2 nd segment 2.0 – 2.2 ×, 3 rd segment 1.8 – 2.0 × and preapical segment 1.3 × as long as wide. Temple moderately and roundly narrowed behind eye. OED 1.7 × ocellar diameter. Clypeus slightly convex, not bulging medially, with sparse punctures, laterally without apical extensions. Ventral mandibular tooth slightly larger than dorsal tooth. MI 0.5. Mesopleuron granulate and ± distinctly punctate ventrally, with fine striae dorsally. Speculum with microsculpture, shining. Area superomedia with weak lateral carina. Area petiolaris with complete carina. Hind femur 4.6 × as long as wide. 1 st tergite 1.5 – 1.6 × as long as wide, without latero-median carina. 2 nd tergite 1.4 – 1.5 × as wide as long. Color: black. Flagellum reddish ventrally, blackish basally, brownish dorsally. Palps brownish, mandible brownish or reddish. Apical 1 / 2 of clypeus reddish. Tegula brown. 1 st to 4 th tergites red, sometimes 1 st tergite ± black. Coxae, trochanters and trochantelli black; legs red; hind femur entirely black; hind tibia reddish-yellow, infuscate in apical 0.4; hind tarsomeres black. Pterostigma ochreous or yellowish. ♁. Body length 7.5 mm. Flagellum with 37 segments; 1 st flagellar segment 4.8 × as long as wide and 1.8 × as long as 2 nd flagellomere; 2 nd segment 2.6 × as long as wide. Temple distinctly roundly narrowed. OED 1.5 ×. Clypeus transversely convex and slightly bulging medially, with coarse punctures, granulate. Ventral mandibular tooth slightly larger than dorsal tooth. MI 0.5. Mesopleuron distinctly striate dorsally, with very superficial punctures, granulate but rather distinctly shining; speculum with microsculpture, shining. Area superomedia with weak lateral carina, sides slightly divergent apically. Area petiolaris with incomplete carina medially. Hind femur 4.6 × as long as wide. 1 st tergite 2.2 × as long as wide, without latero-median carina. 2 nd tergite 1.1 × as wide as long Color: black. Flagellum ochreous ventrally, brown dorsally. Palps, mandible except teeth, clypeus, face and scape ventrally yellow. Tegula black. 1 st tergite black, narrowly red apically; 2 nd to 4 th tergites red. Coxae, trochanters, trochantelli, mid femur basally and hind femur black. Hind tibia yellowish, infuscate in apical 0.4. Hind tarsus black. Pterostigma ochreous. Host: Tenthredo notha KLUG, 1817 (Hym. Tenthredinidae) (ZSM). D i s t r i b u t i o n: Western Palaearctic, studied material from Germany and Sweden. Taxonomical remarks: This species is closely related to Sc. bicolor (GRAVENHORST) and was synonymized with Sc. bicolor by SCHMIEDEKNECHT (1914). As already mentioned by HORSTMANN (2006), it differs by its stouter flagellum and slightly weaker structure of mesopleuron. Due to its rather short basal flagellomeres, it is not impossible that this species represents a variety of Sc. frontator (THUNBERG) or Sc. macropus (THOMSON) with black hind femur, but further genomic studies are necessary to solve this question.	en	Riedel, Matthias, H, Andres, Schmidt, Stefan (2024): Revision of the Palaearctic species of Scopesis FÖRSTER (Hymenoptera, Ichneumonidae, Ctenopelmatinae). Linzer biologische Beiträge 56 (1): 201-252, DOI: 10.5281/zenodo.14681817
039A87F0FFFCFFD8FF07F73C3931B41B.taxon	description	Lectotype: (♁) green label [= Ringsjön / Scåne] (Lund), examined and designated by HORSTMANN (2006: 82). D e s c r i p t i o n: ♀. Body length 8.5 – 10 mm. Flagellum with 40 – 44 segments; 1 st flagellar segment 4.2 – 4.3 × as long as wide and 2.0 × as long as 2 nd segment; 2 nd segment 2.1 – 2.3 ×, 3 rd segment 1.8 – 2.0 × and preapical segment 1.5 × as long as wide. Temple strongly and roundly narrowed behind eye. OED 1.3 × ocellar diameter. Face granulated and with superficial punctures. Clypeus slightly convex and slightly bulging medially, with sparse rather coarse punctures, shining; apical margin concave, with small flat extensions laterally. Mandibular teeth of equal length. MI 0.5. Mesoscutum finely granulate, dull. Notaulus weakly impressed. Mesopleuron finely rugose and superficially punctate ventrally, with rather strong striae reaching frontal margin of mesopleuron dorsally. Speculum finely granulate, rather dull. Area superomedia without lateral carina. Area petiolaris with medially obsolete carina. Lateral longitudinal and pleural carinae of propodeum complete. Hind femur 4.3 – 4.5 × as long as wide. Inner apical spur of hind tibia 0.58 × as long as hind metatarsus. Vein 2 rs-m 0.7 × as long as vein M between 2 rs-m and 2 m-cu. Vein 1 cu-a postfurcal by 1 × its width. Nervellus of hind wing slightly inclivous, intercepted in its apical 0.75. 1 st tergite 1.5 – 1.7 × as long as wide, without latero-median carina. 2 nd tergite 1.2 – 1.4 × as wide as long. Color: black. Flagellum reddish, basal flagellomeres blackish. Palps brown or reddish-yellow. Mandible medially and apical 2 / 3 of clypeus reddish-yellow. Scape usually red ventrally. Tegula yellow or brown. Hind edge of pronotum and subtegular ridge reddish or brown. 1 st to 4 th tergites red. Coxae and trochanters black; trochantelli sometimes ± yellowish; legs otherwise red; tibia reddish-yellow; hind tibia black in apical 0.3; hind tarsus black. Pterostigma yellowish. ♁. Body length 8.5 – 11.0 mm. Flagellum with 38 – 41 segments; 1 st flagellar segment 4.5 – 5.0 × as long as wide and 1.7 × as long as 2 nd segment, 2 nd segment 2.4 – 2.5 ×, 3 rd segment 2.4 × and preapical segment c 2 × as long as wide. OED 1.3 × ocellar diameter. Temple moderately and roundly narrowed behind eye. MI 0.3. Notaulus often ± distinct frontally. Mesopleuron punctate and granulate ventrally, partly and finely striate ventrally, ± shining; speculum almost smooth. Hind femur 4.5 – 4.6 × as long as wide. Vein 2 rs-m 0.6 – 0.9 × as long as vein M between vein 2 rs-m and vein 2 m-cu. 1 st tergite 1.9 – 2.2 × as long as wide. 2 nd tergite about as long as wide. Color: black. Flagellum reddish-yellow, brownish basally and / or dorsally. Palps, mandible except teeth, clypeus, face, ventral 1 / 2 of frontal orbit, ventral spots on scape and pedicel, sometimes spot on collar, wide hind edge and sometimes ventral edge of pronotum, sometimes anterolateral spot on mesoscutum, tegula, subtegular ridge, sometimes spot on ventral epicnemium, often mesepisternum, fore and mid coxae, apical margin of hind coxa, all trochanters and trochantelli yellow. Postpetiolus apically and 2 nd to 4 th tergites red, sometimes basal tergites mainly blackish, with ± distinct dark reddish shine. Legs reddish-yellow; hind tibia yellow, infuscate in apical 0.2 – 0.3; hind tarsus brownish. Pterostigma yellow or ochreous. D i s t r i b u t i o n: West Palaearctic, studied material from Bulgaria, Germany, Sweden, and Switzerland. Taxonomical remarks: As stated by HORSTMANN (2006: 82), the ♀ in coll. Thomson (" Madsr. 23.8.91 ") which had been designated as “ neotype ” by AUBERT (1966: 127) belongs to a different species, most probably to Sc. frontator. The differences between Sc. frontator and Sc. macropus are small in ♀♀ and sometimes variable. An unequivocal determination of ♀♀ is sometimes not possible.	en	Riedel, Matthias, H, Andres, Schmidt, Stefan (2024): Revision of the Palaearctic species of Scopesis FÖRSTER (Hymenoptera, Ichneumonidae, Ctenopelmatinae). Linzer biologische Beiträge 56 (1): 201-252, DOI: 10.5281/zenodo.14681817
039A87F0FFFEFFD9FF07F281380FB6FB.taxon	materials_examined	Holotype: (♀) 31. VII. 1960 zip. Brenner; [red] 1559 (ZSM). D e s c r i p t i o n: ♀. Body length 5.2 mm. Flagellum with 28 segments, stout; 1 st flagellar segment 3.9 × as long as wide and 1.9 × as long as 2 nd segment; 2 nd segment 2.0 ×, 3 rd segment 1.9 × and preapical segment 1.3 × as long as wide. Temple moderately and roundly narrowed behind eye. OED 1.5 × ocellar diameter. Face granulated, dull; inner eye margins slightly convergent ventrally. Clypeus moderately and transversely convex, with sparse punctures, shining, moderately bulging medially, with distinct flat lateral extensions; apical margin slightly concave. Mandibular teeth of equal size. MI 0.65. Mesoscutum granulate, without punctures, dull. Notaulus impressed in frontal 0.5 of mesoscutum. Mesopleuron granulate, dull, without punctures, with few irregular rugae dorsally; speculum with microsculpture, shining. Hind femur 4.4 × as long as wide. Inner apical spur of hind tibia slightly longer than outer spur, 0.53 × as long as hind metatarsus. Area superomedia with weak lateral carina and moderately widened apically; area petiolaris with complete carina. Lateral longitudinal and pleural carinae of propodeum complete. Vein 2 rs-m 0.7 × as long as vein M between veins 2 rs-m and 2 - mcu. Vein 2 m-cu almost straight. Vein 1 cu-a interstitial. Nervellus of hind wing inclivous, intercepted in its apical 0.8. Discoidella of hind wing vestigial. Metasoma stout; 1 st tergite 1.3 × as long as wide, without latero-median carina. 2 nd tergite 1.6 × and 3 rd tergite 2.1 × as wide as long. Color: black. Flagellum reddish-brown; 1 st flagellar segment blackish. Palps brown. Median spot at mandible and clypeus reddish. Hind edge of pronotum and tegula brownish. 2 nd to 3 rd tergites reddish-brown, with diffuse brownish transverse bands medially. Coxae, trochanters and trochantelli black; legs otherwise reddish; hind femur weakly infuscate apically; hind tibia narrowly infuscate basally and apically; hind tarsomeres reddish, with brownish tips. Pterostigma brown, paler proximally. ♁ and hosts unknown. D i s t r i b u t i o n: Only known from Austria. Taxonomical remarks: This new species is characterized by its small size and low number of flagellar segments, stout flagellum, stout metasoma, granulate mesopleuron without distinct striation, bulging clypeus medially and dark color of palps and tegula. The form of clypeus resembles members of the genus Campodorus, but the 1 st tergite of Sc. minuta has no latero-median carina.	en	Riedel, Matthias, H, Andres, Schmidt, Stefan (2024): Revision of the Palaearctic species of Scopesis FÖRSTER (Hymenoptera, Ichneumonidae, Ctenopelmatinae). Linzer biologische Beiträge 56 (1): 201-252, DOI: 10.5281/zenodo.14681817
039A87F0FFFEFFC7FF07F74138A7B372.taxon	description	herewith designated, synonymized by SCHMIEDEKNECHT (1914: 2923), high resolution photos examined. syn. Mesoleius imitator HOLMGREN, 1857: 148, ♀ ♁. Lectotype: (♀) Lp. in; Bhn; Mesoleius imitator HLGR. ♀ Lectotypus R. Hinz 1962 (Stockholm), designated by TOWNES et al. (1965: 259), synonymized by ROMAN (1931: 24), high resolution photos examined. D e s c r i p t i o n: ♀. Body length (6.0) – 8.5 – 10.5 mm. Flagellum with 33 – 40 segments; 1 st flagellar segment 4.2 – 5.0 × as long as wide and 1.7 – 2.0 × as long as 2 nd segment; 2 nd segment 2.3 – 2.7 ×, 3 rd segment 2.2 – 2.5 × and preapical segment 1.4 × as long as wide. Temple moderately to distinctly and roundly narrowed behind eye. OED 1.2 – 1.5 × ocellar diameter. Face granulated, with superficial punctures, inner eye margins parallel. Clypeus moderately convex, not bulging medially, slightly extended laterally, with sparse setiferous punctures; apical margin almost straight. MI 0.4. Mesoscutum granulate, dull. Notaulus not impressed. Mesopleuron granulate, sometimes finely rugose and with superficial punctures, with ± distinct striae in dorsal half; speculum with microsculpture, shining. Lateral carinae of area superomedia sometimes partly present. Area petiolaris with carina, obsolete or complete medially. Lateral longitudinal and pleural carinae of propodeum often ± obsolete, sometimes completely lacking. Hind femur 4.0 – 4.5 × as long as wide. Inner apical spur of hind tibia 0.64 × as long as hind metatarsus. Vein 2 rs-m 0.6 – 0.9 × as long as vein M between 2 rs-m and 2 m-cu. Vein 1 cu- a interstitial or postfurcal by 1 × its width. Nervellus of hind wing vertical or slightly inclivous, intercepted in its apical 0.6 – 0.7. 1 st tergite 1.4 – 1.6 × as long as wide, without latero-median carina. 2 nd tergite 1.2 – 1.5 × as wide as long. Color: black. Flagellum reddish or reddish-yellow, ± black basally. Palps brown or ochreous. Mandible partly yellow. Clypeus reddish-brown or yellow, sometimes with narrow yellowish facial transverse stripe above clypeus. Scape sometimes reddish ventrally. Tegula brown or yellowish. Coxae, trochanters and trochantelli black. Legs red; hind tibia reddish-yellow, infuscate in apical 0.3 - 0.4; hind tarsus black. Metasoma black or brown, 2 nd and / or 3 rd tergites often with some dark reddish shine. Pterostigma ochreous. ♁. Body length 8.0 – 11.0 mm. Flagellum with 32 – 41 segments; 1 st flagellar segment 4.5 – 5.3 × as long as wide and 1.8 × as long as 2 nd segment; 2 nd segment 2.8 – 3.2 × and 3 rd segment 2.5 – 2.7 × longer than wide. OED 1.1 – 1.5 × ocellar diameter. MI 0.2 – 0.3. Mesopleuron granulate and with superficial punctures, ± striate dorsally. Area superomedia often with ± distinct lateral carina. Area petiolaris with complete carina medially. Hind femur 3.9 – 5.0 × as long as wide. Inner apical spur of hind tibia 0.53 × as long as hind metatarsus. 1 st tergite 1.8 – 2.1 × as long as wide, without latero-median carina. 2 nd tergite 1.0 × as long as wide. Color: black. Flagellum ochreous, ± infuscate basally. Palps, mandible, clypeus, often ventral gena, face, ventral 1 / 3 of frontal orbit, ventral spots on scape and pedicel, sometimes collar, hind edge of pronotum, tegula, often subtegular ridge, often anterolateral spot on mesoscutum and often spot on scutellum and postscutellum yellow; sometimes mesosoma except yellow hind edge of pronotum and tegula entirely black. Coxae black, apical spots on fore and mid coxae, usually fore and mid trochanters and trochantelli and often hind trochantellus yellow. legs reddish; hind femur sometimes slightly infuscate apically; hind tibia reddish-yellow, infuscate in apical 0.4 – 0.5; hind tarsus black. Metasoma black; 2 nd and 3 rd tergites often with ± extended dark reddish shine. Pterostigma yellow or brownish. D i s t r i b u t i o n: Transpalaearctic, studied material from Austria, Georgia *, Germany, Sweden, Russia Far East (Kamtchatka), United Kingdom *. Taxonomical remarks: This species has a large range of body size and color patterns, especially in ♁♁. The lectotype of Sc. polita is an exceptionally small ♀ with a body length of 6.0 mm and 33 flagellar segments.	en	Riedel, Matthias, H, Andres, Schmidt, Stefan (2024): Revision of the Palaearctic species of Scopesis FÖRSTER (Hymenoptera, Ichneumonidae, Ctenopelmatinae). Linzer biologische Beiträge 56 (1): 201-252, DOI: 10.5281/zenodo.14681817
039A87F0FFE0FFC4FF07F3D938C2B35E.taxon	description	Lectotype: (♀) ♀; det. Schmiedekn.; Scopesis rubrotinctus SCHMDKN. (Vienna), type not examined. D e s c r i p t i o n: ♀. Body length 7.0 – 9.0 mm. Flagellum with 33 – 35 segments; 1 st flagellar segment 5.0 × as long as wide and 1.8 – 1.9 × as long as 2 nd segment; 2 nd segment 2.3 – 2.4 ×, 3 rd segment 2.1 – 2.2 × and preapical segment 1.8 × longer than wide. Temple distinctly and roundly narrowed behind eye. OED 1.3 – 1.5 × ocellar diameter. Face granulated, with fine superficial punctures. Clypeus moderately convex, not bulging medially, with fine sparse punctures and some long and erected setae, shining, with rather distinct flat extensions laterally; apical margin slightly concave. Mandibular teeth equal or lower tooth slightly larger than upper tooth. MI 0.3 – 0.4. Mesoscutum granulate, with superficial punctures, dull. Notaulus impressed in frontal 0.3. Mesopleuron finely granulate, with superficial punctures, ± shining; dorsally sometimes with few superficial striae. Speculum with microsculpture, shining. Area superomedia without lateral carina. Area petiolaris with short lateral carina, widely open medially. Lateral longitudinal and pleural carinae of propodeum often ± obsolete. Hind femur 3.9 – 4.3 × as long as wide. Inner apical spur of hind tibia 0.62 × as long as hind metatarsus. Vein 2 rs-m 0.5 – 0.7 × as long as vein M between 2 rs-m and 2 m-cu. Vein 1 cu- a interstitial. Nervellus of hind wing vertical, intercepted in its apical 0.6. 1 st tergite 1.5 – 1.7 × as long as wide, without latero-median carina. 2 nd tergite 1.3 × as wide as long. Color: black. Flagellum reddish, infuscate basally. Palps, mandible except teeth, clypeus, often ventral spot at scape, spot on scutellum and postscutellum red. Tegula brown. 1 st tergite except base and 2 nd to 4 th or 5 th tergites entirely red. Coxae, trochanters and trochantelli black; legs red; hind tibia reddish, infuscate in apical 0.3; hind tarsomeres black, with ± paler bases. Pterostigma ochreous. ♁. Body length 8.5 – 10.0 mm. Flagellum with 34 – 35 segments; 1 st flagellar segment 5.0 – 5.2 × as long as wide and 1.7 – 1.8 × as long as 2 nd segment; 2 nd segment 2.7 × and 3 rd segment 2.4 × as long as wide. OED 1.1 – 1.4 × ocellar diameter. MI 0.2 – 0.3. Mesopleuron with smoothened superficial structure, superficially punctate and granulate, weakly striate dorsally, strongly shining. Area superomedia sometimes with incomplete weak lateral carina. Area petiolaris with short lateral carina, widely open medially. Hind femur 3.8 – 4.0 × as long as wide. 1 st tergite 1.9 – 2.0 × as long as wide. 2 nd tergite about as long as wide. Color: black. Flagellum reddish or brown, ± infuscate basally. Palps, mandible except teeth, clypeus, face, lower 1 / 2 of frontal orbit, ventral spots on scape and pedicel, collar, anterolateral spot on mesoscutum, hind edge of pronotum, tegula, subtegular ridge, spot on mesosternum, spot on scutellum and postscutellum, fore and mid coxae largely, fore and mid trochanters and trochantelli and spot on hind trochantellus yellow. Legs otherwise red; hind tibia reddish-yellow, infuscate in apical 0.4; hind tarsus black. Sometimes postpetiolus apically and 2 nd to 4 th tergites entirely red. Pterostigma ochreous or chestnut-brown. D i s t r i b u t i o n: Transpalaearctic, studied material from Bulgaria *, Germany, Norway *, Poland and East Siberia * (Irkutsk environment).	en	Riedel, Matthias, H, Andres, Schmidt, Stefan (2024): Revision of the Palaearctic species of Scopesis FÖRSTER (Hymenoptera, Ichneumonidae, Ctenopelmatinae). Linzer biologische Beiträge 56 (1): 201-252, DOI: 10.5281/zenodo.14681817
039A87F0FFE3FFC5FF07F33C39CAB605.taxon	description	Lectotype: (♀) no label (Lund), synonymized with Sc. frontator by ROMAN (1912: 256), type examined by Horstmann. syn. nov. Mesoleius rufonotatus HOLMGREN, 1876: 31, ♀ ♁. Lectotype: (♀) sc. / Bhn. (Stockholm), type examined. syn. nov. Scopesus thomsoni HABERMEHL, 1925: 180. ♀ ♁, synonymized with Sc. rufonotata by TEUNISSEN (1953: 35), type not examined. D e s c r i p t i o n: ♀. Body length (6.0) – 7.5 – 9.5 mm. Flagellum with 36 – 44 segments; 1 st flagellar segment 5.0 – 6.3 × as long as wide and 1.7 – 2.0 × as long as 2 nd segment; 2 nd segment 2.5 – 3.0 ×, 3 rd segment 2.2 – 2.8 × and preapical segment 1.5 – 1.7 × as long as wide. Temple moderately and roundly narrowed. OED 1.4 – 1.7 × ocellar diameter. Face granulated and with fine superficial punctures, dull. Clypeus moderately convex, ± bulging medially, with sparse rather coarse punctures; apical margin slightly concave, with small flat extensions laterally. Mandibular teeth equal or ventral mandibular tooth slightly larger than dorsal tooth. MI 0.4 – 0.5. Mesoscutum finely granulate, dull. Notaulus usually impressed in frontal 0.3 of mesoscutum. Mesopleuron granulate, without clear punctures ventrally and with ± distinct striae dorsally. Speculum with microsculpture, shining. Area superomedia sometimes with weak but incomplete lateral carina. Area petiolaris with complete carina, sometimes obsolete medially. Lateral longitudinal and pleural carinae of propodeum sometimes ± obsolete. Hind femur 4.5 – 5.3 × as long as wide. Vein 2 rs-m 0.6 × as long as vein M between 2 rs-m and 2 m-cu. Vein 1 cu-a interstitial. Nervellus of hind wing inclivous, intercepted in its apical 0.7 – 0.8. 1 st tergite 1.4 – 1.8 × as long as wide, without latero-median carina. 2 nd tergite 1.3 – 1.6 × as wide as long Color: black. Flagellum red or ochreous, ± black basally. Palps brown. Mandible centrally and apical ½ – 3 / 4 of clypeus reddish-yellow. Hind edge of pronotum brown. Tegula brown or reddish, sometimes yellow. Often 1 st tergite ±, 2 nd and 3 rd tergites mainly or entirely and sometimes 4 th tergite red or reddish-brown. Coxae, trochanters and trochantelli black; legs otherwise red; hind femur often slightly infuscate basally and / or apically; hind tibia red or reddish-yellow, rarely yellowish, sometimes narrowly infuscate basally and usually infuscate in apical 0.2 – 0.3; spurs of hind tibia ochreous or yellowish; hind tarsus black. Pterostigma yellowish to ochreous. Var. ♀: 1 specimen from Mt. Canigou / France (ZSM): Flagellum very slender; 1 st flagellar segment 6.0 ×, 2 nd segment 2.9 × and 3 rd segment 2.8 × as long as wide. Hind femur 5.3 × as long as wide. Metasoma black, 2 nd and 3 rd tergites with dark reddish shine. ♁. Body length 8.0 – 8.5 mm. Flagellum with 35 – 37 segments; 1 st flagellar segment 5.1 – 6.2 × as long as wide and 1.7 – 1.8 × as long as 2 nd segment; 2 nd segment 2.6 – 3.0 × and 3 rd segment 2.5 × as long as wide. Temple distinctly and roundly narrowed. OED 1.5 ×. Clypeus distinctly convex and bulging forwards medially, with sparse punctures, laterally with flat extensions. Lower mandibular tooth slightly larger. MI 0.3. Mesopleuron granulate, without punctures and distinct striae; speculum almost smooth, shining. Area superomedia with ± distinct lateral carina. Area petiolaris with complete carina. Hind femur 4.6 – 5.5 × as long as wide. 1 st tergite 1.8 – 2.0 × as long as wide, without latero-median carina. 2 nd tergite 1.1 – 1.2 × as wide as long. Color: black. Flagellum red, black basally. Palps, mandible, clypeus, face except brown median stripe, sometimes ventral half of frontal orbit and ventral spot at scape, sometimes spot on propleuron, hind edge of pronotum and tegula yellow. 2 nd to 4 th tergites red, rarely metasoma ± infuscate with dark reddish suffusion. Coxae black, apically yellow or fore and mid coxae entirely yellow; hind coxa sometimes yellow apically. Trochanters and trochantelli yellow, hind trochanter ± black dorsally. Legs red; hind femur sometimes slightly infuscate basally and usually apically; hind tibia reddish or reddish-yellow, infuscate in apical 0.3, hind tarsomeres black, with pale bases. Pterostigma brownish. Host: Tenthredo ferruginea SCHRANK, 1776 (Hym. Tenthredinidae) (ZSM). D i s t r i b u t i o n: Probably Transpalaearctic, but the report from Irkutsk environment (MOCSARY & SZÉPLIGETI 1901) is questionable and unconfirmed, studied material from Bulgaria *, Finland, France, Georgia *, Germany, Italy, Sweden, Switzerland, Turkey *, United Kingdom. Taxonomical remarks: Scopesis rufolabris was usually regarded as a synonym of Sc. frontator since ROMAN (1912: 256) had synonymized both species (see HORSTMANN 2007: 81), but the comparison of the type material of both species shows some differences. The lectotype of Sc. rufolabris has longer basal flagellar segment (e. g. 3 rd segment 2.5 × as long as wide) than Sc. frontator (3 rd flagellar segment 2.0 × as long as wide). In Sc. rufolabris, the structure of mesopleuron is weaker and the red hind femur is slightly infuscate apically, whereas the mesopleuron has a coarser structure and the hind femur is entirely red in Sc. frontator. Due to these differences, we separate both species here. Sc. bicolor differs from Sc. rufolabris mainly by its entirely black hind femur.	en	Riedel, Matthias, H, Andres, Schmidt, Stefan (2024): Revision of the Palaearctic species of Scopesis FÖRSTER (Hymenoptera, Ichneumonidae, Ctenopelmatinae). Linzer biologische Beiträge 56 (1): 201-252, DOI: 10.5281/zenodo.14681817
039A87F0FFE2FFC2FF07F6043B8FB0DE.taxon	materials_examined	Holotype: (♀) 1. VII. 74; L. X. 1973 Alfeld; ex Tenthredopsis annuligera (Tenthr.); frontator ♀ (ZSM). D e s c r i p t i o n: ♀. Body length 6.0 mm. Flagellum with 37 segments; 1 st flagellar segment 4.4 × as long as wide and 2.0 × as long as 2 nd segment; 2 nd segment 2.3 ×, 3 rd segment 2.1 × and preapical segment 1.5 × as long as wide. Temple distinctly and almost linearly narrowed behind eye; dorsally 0.7 × as long as eye. OED 1.4 × ocellar diameter. Face granulated; inner orbits slightly convergent ventrally. Clypeus weakly convex, not bulging medially, almost straight apically, without distinct extensions laterally. Lower mandibular tooth slightly longer than upper one. MI 0.3. Mesoscutum granulate, dull. Notaulus impressed in frontal 0.3 of mesoscutum. Mesopleuron irregularly rugose-striate, without distinct punctures, dull. Speculum with microsculpture, slightly shining. Hind femur 4.5 × as long as wide. Inner apical spur of hind tibia 0.67 × as long as hind metatarsus. Area superomedia without lateral carina; area petiolaris widely interrupted medially. Lateral longitudinal and pleural carinae of propodeum weak but complete. Vein 2 rs-m 0.6 × as long as vein M between veins 2 rs-m and 2 - mcu. Vein 1 cu-a postfurcal by 1 × its width. Nervellus of hind wing vertical, intercepted in its apical 0.7. 1 st tergite 1.8 × as long as wide, without dorsal carina. 2 nd tergite 1.1 × as wide as long. Color: black. Flagellum ochreous, infuscate basally. Palps, mandible except teeth, clypeus, hind edge of pronotum and tegula yellow. 1 st tergite in apical 0.25 and 2 nd to 4 th tergites entirely red. Coxae, trochanters and trochantelli black; mid trochantellus ± yellowish; legs otherwise red; hind femur slightly infuscate apically, hind tibia cream-yellow, black in apical 0.3; hind tarsomeres blackish. Pterostigma hyaline-yellow. ♁ unknown. Host: Tenthredopsis tischbeinii (FRIVALDSKY, 1876) (Hym. Tenthredinidae) (ZSM). D i s t r i b u t i o n: Only known from the German holotype. Taxonomical remarks: This specimen was determined as Sc. frontator by HINZ, but seems to be closely related to Sc. macropus (THOMSON). Due its small size, characteristic structure of mesopleuron and long 1 st tergite, it is described as a separate species here, but might represent a very small (host-depended?) specimen of Scopesis macropus (THOMSON). For a definitive decision, more material, host records and genomic studies are necessary in the future.	en	Riedel, Matthias, H, Andres, Schmidt, Stefan (2024): Revision of the Palaearctic species of Scopesis FÖRSTER (Hymenoptera, Ichneumonidae, Ctenopelmatinae). Linzer biologische Beiträge 56 (1): 201-252, DOI: 10.5281/zenodo.14681817
039A87F0FFE5FFC3FF07F0BC3E24B1A8.taxon	description	Lectotype: (♀) Saghalin, Adachi, Issiki (Sapporo), type examined. syn. nov. Scopesis alpivagans HEINRICH, 1949: 89, ♀ ♁. No syntypes but material determined by Heinrich examined. D e s c r i p t i o n: ♀. Body length 7.5 – 10.0 mm. Flagellum with 36 – 40 segments; 1 st flagellar segment 4.4 – 5.5 × as long as wide and 1.8 – 2.2 × as long as 2 nd segment; 2 nd segment 2.4 – 2.8 ×, 3 rd segment 2.1 – 2.5 × and preapical segment 1.3 – 1.5 × as long as wide. Temple moderately and roundly narrowed behind eye. OED 1.5 – 1.6 × ocellar diameter. Face wide, granulate, inner eye margins parallel or slightly convergent. Clypeus slightly convex, shining, not bulging medially, with fine sparse punctures, apical margin with small flat extensions laterally. Ventral mandibular tooth equal to or slightly larger than dorsal tooth. MI 0.2 – 0.4. Mesoscutum granulate, without distinct punctures, dull. Notaulus usually impressed in frontal 0.3 of mesoscutum. Mesopleuron finely granulate, dull, without or with very superficial punctures ventrally; without or with fine striae dorsally. Speculum with microsculpture, but ± shining. Propodeum granulated. Area superomedia without lateral carina; area petiolaris usually with obsolete carina medially. Lateral longitudinal and pleural carinae of propodeum usually complete. Hind femur 4.3 – 4.7 × as long as wide. Inner apical spur of hind tibia 0.56 × as long as hind metatarsus. Vein 2 rs-m 0.8 – 0.9 × as long as vein M between veins 2 rs-m and 2 - mcu. Vein 1 cu-a postfurcal by 1 – 3 × its width. Nervellus of hind wing vertical, intercepted in its apical 0.6. 1 st tergite 1.5 – 1.9 × as long as wide, without latero-median carina. 2 nd tergite 1.0 – 1.4 × as wide as long. Color: black. Flagellum ochreous or red, ± brownish dorsally. Palps mainly, mandible except teeth, clypeus and sometimes ventral margin of face narrowly yellow. Tegula usually brown or black, rarely reddish or yellow. 1 st tergite usually black, sometimes ± reddish, 2 nd to 3 rd tergites entirely and often 4 th tergite basally red. Coxae, trochanters and trochantelli black; legs otherwise red; fore and mid femora often basally black; hind femur entirely black; hind tibia reddish or yellowish, black at narrow base and in apical 0.2 – 0.4; sometimes distal mid tarsomere and hind tarsus except yellowish base of metatarsus black. Pterostigma yellow or ochreous. ♁. Body length 8.5 mm. Flagellum with 36 – 39 segments; 1 st flagellar segment 5.3 – 5.9 × as long as wide and 1.7 – 2.0 × as long as 2 nd segment; 2 nd segment 2.8 – 2.9 × and 3 rd segment 2.5 × as long as wide. Temple slightly or moderately and roundly narrowed behind eye. OED 1.6 - 1.7 × ocellar diameter. MI 0.5. Mesopleuron granulate ventrally, with fine rugae in dorsal half. Speculum with microsculpture, shining. Area superomedia sometimes with lateral carina. Hind femur 4.4 – 4.8 × as long as wide. 1 st tergite 1.7 – 2.1 × as long as wide, without latero-median carina. 2 nd tergite 1.0 – 1.2 × as wide as long. Color: black. Palps, mandible, clypeus, face, ventral spot of scape and sometimes tegula yellow. Often apical margin of 1 st tergite, 2 nd tergite ±, 3 rd tergite entirely and 4 th tergite ± red. Coxae black, fore and mid coxae sometimes with apical yellow spots. Fore and mid trochanters and all trochantelli ± yellow. Legs otherwise red; fore and mid femora usually ± infuscate basally; hind femur black; hind tibia yellowish, infuscate in apical 0.3 - 0.4. Distal tarsomeres sometimes and hind tarsus brownish. Pterostigma ochreous. Hosts: unknown. D i s t r i b u t i o n: Transpalaearctic, studied material from Austria, France *, Germany, Italy *, Siberia (Irkutsk region), Sakhalin, United Kingdom *. Taxonomical remarks: The hind tibiae of European ♀♀ are mainly reddish, the Eastern Palaearctic ♀♀ have yellowish hind tibiae, but we could not find other differences between the Western and Eastern Palaearctic material and therefore synonymized Scopesis alpivagans HEINRICH with Scopesis sachalinenis (UCHIDA) here.	en	Riedel, Matthias, H, Andres, Schmidt, Stefan (2024): Revision of the Palaearctic species of Scopesis FÖRSTER (Hymenoptera, Ichneumonidae, Ctenopelmatinae). Linzer biologische Beiträge 56 (1): 201-252, DOI: 10.5281/zenodo.14681817
039A87F0FFE4FFC0FF07F1973B60B148.taxon	materials_examined	Holotype: (♀) Steiermark, Admont, Kreuzkogel, 1700 m, 28.7.50 (ZSM), type examined. D e s c r i p t i o n: ♀. Body length 7.0 – 9.0 mm. Flagellum with 35 – 38 segments; 1 st flagellar segment 5.0 – 5.6 × as long as wide and 1.7 – 1.8 × as long as 2 nd segment; 2 nd segment 2.9 – 3.3 × and 3 rd segment 2.7 – 3.0 × as long as wide, preapical segment 1.7 × as long as wide. Temple distinctly and roundly narrowed behind eye, dorsally 0.85 × as long as eye. OED 1.4 – 1.5 × ocellar diameter. Face granulated, dull. Clypeus moderately and transversely convex, smooth and shining, slightly bulging medially, with sparse fine punctures and with distinct flat lateral extensions; apical margin distinctly concave. Ventral mandibular tooth slightly larger than dorsal tooth. MI 0.3 – 0.4. Mesoscutum granulate, without punctures, dull. Notaulus ± impressed in frontal 0.4 of mesoscutum. Mesopleuron granulate, dull, without distinct punctures or striae; speculum with microsculpture, shining. Hind femur 5.6 – 5.9 × as long as wide. Inner apical tibial spur 0.56 × as long as hind metatarsus. Area superomedia with ± complete lateral carina, slightly widened apically; area petiolaris with complete carina. Vein 2 rs-m 0.8 – 0.9 × as long as vein M between veins 2 rs-m and 2 - mcu. Vein 1 cu-a interstitial. Nervellus of hind wing vertical or slightly inclivous, intercepted in its apical 0.7 – 0.8. 1 st tergite slender, 2.1 – 2.2 × as long as wide, without latero-median carina. 2 nd tergite 1.0 – 1.1 × as wide as long. Color: black. Flagellum reddish-yellow, infuscate basally. Palps, mandible except teeth, clypeus and V-shaped spot at face, often small anterolateral spot on mesoscutum, hind edge of pronotum, tegula and often small spot on subtegular ridge yellow. Scutellum sometimes with dark reddish shine. 2 nd to 4 th tergites reddish, often apical tergites laterally or diffusely reddish. 2 nd and 3 rd sternites yellow, 4 th sternite red, 5 th sternite and hypopygium brown. Coxae black, fore and mid coxae with apical yellow spots; all trochanters and trochantelli yellow; legs red; hind femur except narrow yellowish base black; hind tibia reddish or reddish-yellow, black in apical 0.2 – 0.3; hind tarsus yellowish. Pterostigma yellow. ♁. Body length 10 mm. Flagellum with 35 segments; 1 st flagellar segment 5.8 × as long as wide and 1.85 × as long as 2 nd segment; 2 nd segment 3.1 × and 3 rd segment 3.0 × as long as wide. OED 1.5 × ocellar diameter. MI 0.3. Hind femur 5.3 × as long as wide. 1 st tergite 2.6 × as long as wide; 2 nd tergite about as long as wide. Structure otherwise as in ♀. Color: black. Flagellum ochreous ventrally, brown dorsally. Palps, mandible except teeth, clypeus, face and orbit opposite antennal groove, ventral spot at scape, large anterolateral spot on mesoscutum, hind edge of pronotum, tegula and spot on subtegular ridge yellow. 2 nd to 4 th tergites red. Fore coxa entirely and mid coxa mainly yellow; hind coxa black; legs otherwise as in ♀. Pterostigma ochreous Host: Sciapteryx costalis (FABRICIUS, 1775) (Hym. Tenthredinidae) (ZSM). D i s t r i b u t i o n: Western Palaearctic, studied material from Austria, Germany *, and Switzerland *.	en	Riedel, Matthias, H, Andres, Schmidt, Stefan (2024): Revision of the Palaearctic species of Scopesis FÖRSTER (Hymenoptera, Ichneumonidae, Ctenopelmatinae). Linzer biologische Beiträge 56 (1): 201-252, DOI: 10.5281/zenodo.14681817
039A87F0FFE7FFC1FF07F1373929B670.taxon	materials_examined	Holotype: (♀) L. 24. X. 1962 Alfeld Han.; 1962 / LXIX 11.6.63; ex Tenthredopsis tarsata F. (Tenthr.) (ZSM), type examined. D e s c r i p t i o n: ♀. Body length 8.0 – 9.0 mm. Flagellum with 39 – 42 segments; 1 st flagellar segment 5.0 – 5.3 × as long as wide and 1.9 – 2.0 × as long as 2 nd segment; 2 nd segment 2.5 – 2.8 ×, 3 rd segment 2.1 – 2.4 × and preapical segment 1.4 – 1.5 × as long as wide. Temple distinctly and roundly narrowed behind eye. OED 1.5 – 1.8 × ocellar diameter. Face granulated, with ± distinct punctures dorsally. Clypeus moderately convex, not bulging, with sparse punctures, shining, with small flat extensions laterally; apical margin ± concave. Mandibular teeth equal or ventral tooth slightly larger than dorsal tooth. MI 0.4. Mesoscutum granulate, dull. Notaulus impressed in frontal 0.3 of mesoscutum. Mesopleuron granulate, with rather distinct punctures and partly with rugae or striae ventrally, distinctly striate dorsally. Speculum with microsculpture, shining. Area superomedia without lateral carina; area petiolaris with complete or sometimes obsolete carina medially. Lateral longitudinal and pleura carinae of propodeum usually complete. Hind femur 4.4 – 4.5 × as long as wide. Inner apical spur of hind tibia 0.62 × as long as hind metatarsus. Vein 2 rs-m 0.7 – 0.8 × as long as vein M between veins 2 rs-m and 2 - mcu. Vein 1 cu-a interstitial. Nervellus of hind wing ± inclivous, intercepted in its apical 0.6 – 0.7. 1 st tergite 1.6 – 1.8 × as long as wide, without latero-median carina. 2 nd tergite 1.1 – 1.2 × as wide as long. Color: black. Flagellum reddish, infuscate basally. Palps, mandible except teeth, clypeus (often with baso-median dark spot), sometimes ventral margin and / or oblique paramedi- an spots on face, often hind edge of pronotum and tegula yellow. Scape often reddish ventrally. 1 st tergite except base and 2 nd to 4 th tergites entirely red. Coxae, trochanters and trochantelli black (mid trochantellus sometimes yellowish); legs red; fore and mid tibiae and tarsi ± yellowish; hind femur red, slightly infuscate apically, hind tibia yellow, black in apical 0.2; hind tarsus black. Pterostigma yellow. ♁. Body length 7.0 - 9.0 mm. Flagellum with 39 – 42 segments; 1 st flagellar segment 5.2 × as long as wide and 1.9 × as long as 2 nd segment; 2 nd segment 2.8 ×, 3 rd segment 2.5 × and preapical segment c 2 × as long as wide. Temple distinctly and roundly narrowed behind eye. OED 1.2 – 1.3 ×. Clypeus moderately convex, with small flat extension laterally. Lower mandibular tooth slightly larger than upper tooth. MI 0.25. Mesopleuron granulate and with superficial punctures, ± shining, with some fine striae dorsally. Speculum with microsculpture, shining. Hind femur 4.5 – 4.8 × longer than wide. Area superomedia without lateral carina; area petiolaris with complete carina. 1 st tergite 2.2 – 2.5 × as long as wide, without latero-median carina. 2 nd tergite 1.1 – 1.2 × as long as wide. Color: black. Flagellum ochreous, ± infuscate basally and dorsally. Palps, mandible except teeth, clypeus, malar space, face, scape and pedicel ventrally, rarely anterolateral spot of mesoscutum, wide hind edge of pronotum and tegula, usually fore and mid coxae entirely and all trochanters and trochantelli yellow. 2 nd to 4 th tergites red. Hind coxa black, rarely with reddish spot dorsally; femora, tibiae and tarsi reddish; hind femur usually red, slightly infuscate apically or sometimes entirely black; hind tibia yellow or reddish-yellow (rarely reddish), infuscate in apical 0.3; hind tarsus black. Pterostigma yellow. Var. ♁: A few ♁♁ have black coxae and resemble the ♁♁ of Scopesis bicolor. They can be differentiated by the ± yellow hind edge of pronotum. Host: Tenthredopsis tarsata (FABRICIUS, 1804) (Hym., Tenthredinidae) (ZSM). D i s t r i b u t i o n: Transpalaearctic, studied material from Germany, Norway *, Siberia * (Irkutsk environment), Sweden *, United Kingdom *.	en	Riedel, Matthias, H, Andres, Schmidt, Stefan (2024): Revision of the Palaearctic species of Scopesis FÖRSTER (Hymenoptera, Ichneumonidae, Ctenopelmatinae). Linzer biologische Beiträge 56 (1): 201-252, DOI: 10.5281/zenodo.14681817
039A87F0FFE6FFCFFF07F6DF3929B313.taxon	description	Lectotype: (♀) Hlm; De V (Lund), type examined by Horstmann. D e s c r i p t i o n: ♀. Body length 8.5 – 9.5 mm. Flagellum with 40 – 44 segments; 1 st flagellar segment 5.5 – 5.7 × as long as wide and 1.9 – 2.0 × as long as 2 nd segment; 2 nd segment 2.6 – 2.8 ×, 3 rd segment 2.5 × and preapical segment 1.5 × as long as wide. Temple distinctly and roundly narrowed behind eye. OED 1.4 – 1.5 × ocellar diameter. Face granulated, with fine superficial punctures. Clypeus slightly convex, not bulging medially, with sparse punctures and very small flat extension laterally; apical margin almost straight or slightly concave. Ventral mandibular tooth equal to or slightly larger than dorsal tooth. MI 0.4. Mesoscutum granulate and with fine superficial punctures, dull. Notaulus impressed in frontal 0.25 of mesoscutum. Mesopleuron granulate and finely punctate, with some fine striae ventrally, dorsally with strong striae. Speculum with microsculpture, rather dull. Area superomedia without distinct carina; area petiolaris with obsolete carina medially. Lateral longitudinal and pleural carinae of propodeum usually complete. Hind femur 4.5 – 4.6 × as long as wide. Inner apical spur of hind tibia 0.65 × as long as hind metatarsus. Vein 2 rs-m 0.7 – 0.8 × as long as vein M between veins 2 rs-m and 2 - mcu. Vein 1 cu-a slightly postfurcal by 1 × its width. Nervellus of hind wing ± inclivous, intercepted in its apical 0.7. 1 st tergite 1.6 – 1.7 × as long as wide, without latero-median carina. 2 nd tergite 1.2 – 1.3 × as wide as long. Color: Flagellum ochreous, black basally. Palps brown or yellow. Mandible except teeth, clypeus (sometimes reaching ventral margin of face) and tegula yellow. Hind edge of pronotum and spot on subtegular ridge reddish. 1 st to 4 th tergites red. Coxae, trochanters and trochantelli black; legs otherwise red; hind femur entirely brown or black; hind tibia reddish-yellow, infuscate in apical 0.2; hind tarsus black. Pterostigma yellow. ♁. Body length 9.0 – 10.0 mm. Flagellum with 40 – 43 segments; 1 st flagellar segment 5.3 – 5.6 × as long as wide and 1.7 – 1.9 × as long as 2 nd segment; 2 nd segment 3.0 ×, 3 rd segment 2.5 – 2.9 × and preapical segment c 2 × as long as wide. Temple moderately roundly narrowed. OED 1.3 × ocellar diameter. Clypeus transversely convex, not bulging medially, with sparse punctures. Ventral mandibular tooth about as long as dorsal tooth. MI 0.4 – 0.5. Mesopleuron finely striate dorsally, granulate and with superficial punctures ventrally; speculum with microsculpture or almost smooth. Area superomedia sometimes with weak lateral carina. Area petiolaris with complete carina. Hind femur 4.4 – 5.2 × as long as wide. Vein 1 cu-a postfurcal by 1 × its width. 1 st tergite (2.0) – 2.3 – 2.6 × as long as wide, without latero-median carina. 2 nd tergite 1.0 – 1.1 × as long as wide. Color: black. Flagellum brownish, rarely ochreous. Palps, mandible except teeth, clypeus, face, often scape and pedicel ventrally, sometimes collar, wide hind edge of pronotum, tegula and sometimes subtegular ridge yellow. Postscutellum sometimes reddish. 1 st tergite black; 2 nd to 4 th tergites red. Fore and mid coxae with apical yellow spots, fore and mid trochanters and trochantelli mainly yellow; hind trochanter black; legs red; hind femur black; hind tibia yellowish, infuscate in apical 0.3 – 0.4. Hind tarsus black. Pterostigma ochreous. Hosts unknown. Var. ♁: One specimen from Llangristiolus, Anglesey / United Kingdom (EDI): Flagellum with 40 segments; 1 st flagellar segment 6.3 × as long as wide and 1.8 × as long as 2 nd segment; 2 nd segment 3.1 × and 3 rd segment 2.8 × as long as wide. Hind femur 4.7 × as long as wide. 1 st tergite 2.0 × and 2 nd tergite 1.25 × as long as wide. Color: Fore and mid femora brown. Fore and mid tibiae and tarsi cream-yellow, with brown external stripes. Hind femur black; hind tibia brownish, with ivory internal stripe; hind tarsus black. Metasoma black; 2 nd tergite with diffuse dark reddish shine apically. Palps except brown distal palpomeres, mandible, clypeus, face, scape ventrally, tegula, hind edge of pronotum, spot on subtegular ridge, apical marks on fore and mid coxae and fore and mid trochanters and trochantelli entirely yellow. This specimen probably represents a melanistic variant of Sc. tegularis. Hosts unknown. D i s t r i b u t i o n: Transpalaearctic, studied material from France, Germany, Sweden, Russia Far East (Kamtchatka) and United Kingdom.	en	Riedel, Matthias, H, Andres, Schmidt, Stefan (2024): Revision of the Palaearctic species of Scopesis FÖRSTER (Hymenoptera, Ichneumonidae, Ctenopelmatinae). Linzer biologische Beiträge 56 (1): 201-252, DOI: 10.5281/zenodo.14681817
039A87F0FFE8FFCFFF07F3793860B7E7.taxon	materials_examined	Holotype: (♀) Crefeld Hafen, leg. Ulbricht, 26. IX. (Krefeld), type not examined. D e s c r i p t i o n: ♀. Body length 9.0 – 9.5 mm. Flagellum with 37 – 39 segments; 1 st flagellar segment 3.9 – 4.3 × as long as wide and 1.8 – 1.9 × as long as 2 nd segment; 2 nd segment 2.0 – 2.4 ×, 3 rd segment 1.7 – 2.1 × and preapical segment 1.5 × as long as wide. Head covered with rather long erected brownish setae. Temple slightly and roundly narrowed behind eye. OED 1.8 – 1.9 × ocellar diameter. Face granulated, with ± distinct punctuation. Clypeus slightly convex and slightly bulging medially, with sparse rather coarse punctures, shining; apical margin with short flat extensions laterally. Ventral mandibular tooth ± larger than dorsal tooth. MI 0.6 – 0.7. Mesoscutum granulate and with fine punctures, structure ± smoothened on lateral lobe and ± shining. Notaulus weakly impressed frontally. Mesopleuron entirely and coarsely rugose, mesosternum ± punctate. Speculum smooth and shining. Metapleuron and propodeum often finely rugose. Area superomedia usually finely rugose, with indistinct lateral carina. Area petiolaris with complete carina and often with median longitudinal ridge. Lateral longitudinal and pleural carinae of propodeum complete. Hind femur 4.0 – 4.4 × as long as wide. Inner apical spur of hind tibia rather short, 0.47 – 0.52 × as long as hind metatarsus. Vein 2 rs-m 0.9 - 1.0 × as long as vein M between veins 2 rs-m and 2 - mcu. Vein 1 cu-a postfurcal by 2 – 3 × its width. Nervellus of hind ± inclivous, intercepted in its apical 0.7. 1 st tergite 1.3 – 1.4 × as long as wide, sometimes with weak latero-median carina in basal third. 2 nd tergite 1.5 – 1.6 × as wide as long. Color: black. Flagellum black. Head and mesosoma entirely black, sometimes clypeus reddish-brown. 2 nd to 3 rd tergites entirely red. Coxae, trochanters and trochantelli black; legs otherwise red; hind tibia black in apical 0.3 – 0.4; hind tarsus blackish. Pterostigma ochreous. ♁. Body length 8.5 – 9.5 mm. Flagellum with 37 – 38 segments; 1 st flagellar segment 4.5 – 4.8 × as long as wide and 1.7 – 1.9 × as long as 2 nd segment; 2 nd segment 2.5 – 2.8 × and 3 rd segment 2.3 × as long as wide. Temple moderately and roundly narrowed behind eye. OED 1.6 – 1.8 × ocellar diameter. Mesopleuron coarsely rugose, speculum shining. Propodeum granulate and ± finely rugose. Hind femur 4.5 × as long as wide. 1 st tergite 1.5 – 1.6 × as long as wide, without latero-median carina. 2 nd tergite 1.1 – 1.2 × as wide as long. Color: black. Flagellum black. Palps, mandible centrally and apical 1 / 2 of clypeus reddish-brown. 2 nd and 3 rd tergites entirely and sometimes 4 th tergite ± red. Coxae, trochanters and trochantelli black; legs otherwise red; hind tibia reddish-yellow, black in apical 0.3; hind tarsus blackish. Pterostigma ochreous. Hosts unknown. D i s t r i b u t i o n: West Palaearctic, studied material from Austria *, Germany, Italy * (ZSM) and United Kingdom * (EDI).	en	Riedel, Matthias, H, Andres, Schmidt, Stefan (2024): Revision of the Palaearctic species of Scopesis FÖRSTER (Hymenoptera, Ichneumonidae, Ctenopelmatinae). Linzer biologische Beiträge 56 (1): 201-252, DOI: 10.5281/zenodo.14681817
039A87F0FFEBFFCCFF07F34F387BB046.taxon	description	The type was studied and transferred to Scopesus [recte: Scopesis] by PFANKUCH (1906: 221), but is lost now (TOWNES 1959: 77). Taxonomical remarks (by PFANKUCH 1906, translated): The type is not complete, head and several legs missing. Close to nigricollis [recte: gesticulator], but differs by: flagellum entirely reddish-yellow, scutellum and postscutellum red, hind coxa reddish-brown, hind femur reddish-yellow, propodeum with lateral red spots. Sculpture and coloration otherwise as in nigricollis. Body length 10 mm. We have not seen any ♁ of Scopesis correlating with the description of immatura, the identity of this species remains doubtful.	en	Riedel, Matthias, H, Andres, Schmidt, Stefan (2024): Revision of the Palaearctic species of Scopesis FÖRSTER (Hymenoptera, Ichneumonidae, Ctenopelmatinae). Linzer biologische Beiträge 56 (1): 201-252, DOI: 10.5281/zenodo.14681817
039A87F0FFEBFFCDFF07F0F13E1EB610.taxon	materials_examined	Type lost (PFANKUCH 1906: 21), transferred to Scopesis by HORSTMANN (2007: 319). syn. Scopesus Sziladyi KISS, 1924 Holotype: (♀) Zengg 909. VII. 13 (Budapest), examined and synonymized by HORSTMANN (2007: 319). D e s c r i p t i o n: ♀. Body length 8.5 – 9.0 mm. Flagellum with 40 segments, very slender; 1 st flagellar segment 6.4 × as long as wide and 1.6 × as long as 2 nd segment; 2 nd segment 3.6 ×, 3 rd segment 3.2 × and preapical segment 1.8 × as long as wide. Temple distinctly and roundly narrowed behind eye. Ocelli large, OED 1.0 × ocellar diameter. Face granulated, dull. Clypeus moderately convex, not bulging medially, with sparse punctures; apical margin with short lateral extensions, almost straight. Mandibular teeth equally sized, sometimes ventral tooth slightly larger than dorsal tooth. MI 0.5. Mesoscutum finely granulate, dull. Notaulus ± distinct in frontal 0.5 of mesoscutum. Mesopleuron granulate, without punctures, dull; with few fine striae below subtegular ridge. Speculum large, almost smooth and strongly shining. Propodeum granulated. Are superomedia without lateral carina. Area petiolaris with very short lateral carina, widely absent medially. Lateral longitudinal and pleural carinae of propodeum complete. Legs slender; hind femur 5.4 × as long as wide. Inner apical spur of hind tibia 0.6 × as long as hind metatarsus. Areolet in fore wing closed, e. g. vein 3 rs-m present but usually unpigmented. Vein 2 m-cu reaching areolet close to vein 3 rs-m. Vein 1 cu-a postfurcal by 2 × its width. Nervellus of hind wing slightly inclivous, intercepted in its apical 0.7. 1 st tergite slender, 2.2 × as long as wide, without latero-median carina. 2 nd tergite 1.1 × as wide as long. Color: black. Antenna yellowish. Palps, mandible except teeth, clypeus, face (except small dorso-median brown spot), large hook-shaped anterolateral spot on mesoscutum, hind edge of pronotum, tegula, subtegular ridge, mesepisternum, scutellum and postscutellum yellow. 1 st tergite apically, 2 nd tergite except two paramedian brown spots and 3 rd to 4 th tergites entirely red. Fore and mid coxae, all trochanters and trochantelli reddish-yellow; hind coxa red; legs otherwise red; hind tibia yellowish, reddish-brown in apical 0.1; hind tarsus ochreous. Pterostigma yellow. ♁. Body length 8.0 – 8.5 mm. Flagellum with 39 – 42 segments, slender; 1 st flagellar segment 5.0 – 5.3 × as long as wide and 1.5 – 1.6 × as long as 2 nd segment; 2 nd segment 3.0 – 3.3 ×, 3 rd segment 2.5 – 2.8 × and preapical segment c 2.0 × as long as wide. Temple moderately and roundly narrowed behind eye. Ocelli large, OED 1.1 – 1.2 × ocellar diameter. Clypeus moderately convex, not bulging medially, with sparse punctures; apical margin with short lateral extensions. Mandibular teeth equally sized. MI 0.3. Mesopleuron granulate, without punctures, dull; sometimes with few fine irregular striae dorsally. Speculum with microsculpture, shining. Legs slender; hind femur 4.9 – 5.3 × as long as wide. Propodeum granulated. Area superomedia with fine almost parallel lateral carinae. Area petiolaris with complete carina. Areolet in fore wing closed, e. g. vein 3 rsm present but unpigmented. Vein 2 m-cu reaching areolet close to vein 3 rs-m. Vein 1 cu-a postfurcal by 2 × its width. 1 st tergite 1.8 – 2.2 × as long as wide, without latero-median carina. 2 nd tergite 1.0 – 1.1 × as wide as long. Color: black. Antenna yellowish. Palps, mandible except teeth, clypeus, face, malar space, collar, large hook-shaped anterolateral spot on mesoscutum, ventral and hind edge of pronotum, tegula, subtegular ridge, mesepisternum, scutellum and postscutellum yellow. 1 st tergite apically, 2 nd tergite except large basal brown spot, 3 rd tergite entirely and 4 th to 5 th tergites ± red. Fore and mid coxae, all trochanters and trochantelli yellowish; hind coxa red; legs otherwise red; hind tibia yellowish, infuscate in apical 0.2; hind tarsus brownish or ochreous. Pterostigma yellowish. Host: Aglaostigma aucupariae (KLUG, 1817) (Hym., Tenthredinidae) (ZSM). D i s t r i b u t i o n: West Palaearctic, studied material from Germany. Taxonomical remarks: This species was transferred to the genus Scopesis by HORSTMANN (2007) with question mark. He noted that the equally sized teeth speak against an inclusion into the genus Alcochera FÖRSTER, but the relation of the basal two flagellomeres, presence of vein 3 rs-m in fore wing and color pattern argue against the genus Scopesis and fit better with Alcochera. We transfer this species to Alcochera here.	en	Riedel, Matthias, H, Andres, Schmidt, Stefan (2024): Revision of the Palaearctic species of Scopesis FÖRSTER (Hymenoptera, Ichneumonidae, Ctenopelmatinae). Linzer biologische Beiträge 56 (1): 201-252, DOI: 10.5281/zenodo.14681817
039A87F0FFEAFFCAFF07F67E3E37B6DD.taxon	description	flagellar segment 4.5 – 4.7 × as long as wide and 1.5 – 1.6 × as long as 2 nd segment; 2 nd segment 2.9 × and 3 rd segment 2.5 × as long as wide. Temple short, dorsally 0.45 – 0.6 × as long as eye, distinctly and roundly narrowed. OED 1.3 – 1.4 × ocellar diameter. Face granulated, dull. Clypeus moderately convex, with sparse punctures, shining, moderately bulging medially, slightly and narrowly extended laterally. Mandibular teeth of equal size. MI 0.3 – 0.5. Mesoscutum finely granulate, dull. Notaulus impressed in frontal 0.3 – 0.4 of mesoscutum. Mesopleuron granulate, sometimes slightly rugose ventrally, finely striate dorsally; speculum smooth and shining. Area superomedia with distinct lateral carina, ± widened apically. Area petiolaris large, with strong carina. Lateral longitudinal and pleural carinae of propodeum complete. Hind femur 4.5 – 5.3 × as long as wide. Inner apical spur of hind tibia 0.58 – 0.67 × as long as hind metatarsus. Vein 2 rs-m 0.7 – 0.9 × as long as vein M between 2 rs-m and 2 m-cu. Vein 1 cu-a postfurcal by 1 – 3 × its width. Nervellus subvertical, intercepted in its apical 0.6. 1 st tergite 1.3 – 1.8 × as long as wide, with latero-median carina in basal 0.5. 2 nd tergite 1.2 – 1.5 × as wide as long. Color: black. Flagellum brown, ± ochreous ventrally. Palps, mandible, clypeus and hind edge of pronotum yellow. Tegula ivory. Coxae and trochanters black, trochanters apically and usually all trochantelli entirely yellow. Legs reddish-yellow; hind femur red, sometimes infuscate apically; hind tibia yellow, basally and in apical 0.2 – 0.3 black; hind tarsus black. 2 nd to 4 th tergites red-yellow to reddish-brown. Pterostigma brown. ♁. Body length 6.5 – 7.0 mm. Flagellum with 35 – 37 segments; 1 st flagellar segment 4.3 – 4.5 × as long as wide and 1.5 – 1.7 × as long as 2 nd segment; 2 nd segment 2.7 – 2.8 × and 3 rd segment 2.3 – 2.5 × as long as wide. Temple distinctly roundly narrowed behind eye. OED 1.2 × ocellar diameter. Face wide, inner eye margin slightly convergent ventrally. Clypeus moderately convex and bulging medially. MI 0.2 – 0.3. Mesopleuron finely rugose-striate ventrally and finely striate dorsally. Speculum with fine microsculpture, shining. Area superomedia with distinct lateral carina, moderately widened apically. Area petiolaris large, with strong carina. Hind femur 4.3 – 4.4 × as long as wide. Vein 2 rsm 0.6 × as long as vein M between 2 rs-m and 2 m-cu. Vein 1 cu-a postfurcal by 3 × its width. Color: black. Flagellum brown. Palps, mandible except teeth, clypeus, face, hind edge of pronotum, tegula, apical spots on fore and mid coxae, fore and mid trochanters, hind trochanter apically and all trochantelli yellow. 2 nd and 3 rd tergites ± red. Femora, tibiae and tarsi red; hind tibia yellow, infuscate basally and in apical 0.2; hind tarsus brown. Pterostigma brown. Host: Pachynematus clitellatus (SERVILLE, 1823) (Hym. Tenthredinidae) (ZSM). D i s t r i b u t i o n: West Palaearctic, studied material from Bulgaria *, Germany and Sweden. Taxonomical remarks: This species was transferred to Scopesus (recte: Scopesis) by THOMSON (1894) and included in this genus by all following authors. It differs from Scopesis by a shorter 1 st flagellar segment in relation to the 2 nd segment and the presence of a latero-median carina in the basal half of 1 st tergite. The clypeus is moderately convex and bulging medially. We transfer this species to the genus Campodorus here (nov. comb.)	en	Riedel, Matthias, H, Andres, Schmidt, Stefan (2024): Revision of the Palaearctic species of Scopesis FÖRSTER (Hymenoptera, Ichneumonidae, Ctenopelmatinae). Linzer biologische Beiträge 56 (1): 201-252, DOI: 10.5281/zenodo.14681817
