identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
0398CE51FFACFF9FFF6BFCD6FD5365CB.text	0398CE51FFACFF9FFF6BFCD6FD5365CB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	(Ocreatomurus) Yu 2023	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Subgenus  Ocreatomurus Yu, 2023 in Yu et al. 2023 </p>
            <p> Common characters shared by the new species: Eyes 6+6. Both dorsal and ventral sides of head scaled. Mouthparts normal for  Tomocerus . Labral formula 4/5, 5, 4, distal edge of labrum with four papillae ending in curved spines. Mentum with five chaetae, submentum with numerous chaetae. Mandibular head asymmetrical, the left side with four teeth and the right side with five teeth, left molar plate distally with a tapered tooth. Maxillary lamella 5 without beard-like appendage. Maxillary outer lobe with trifurcate palp, one basal chaeta and four sublobal hairs. Both dorsal and ventral sides of Ant. I and Ant. II scaled, Ant. IV unscaled. Cephalic dorsal macrochaetotaxy: anterior area: 2 (A2), 4 (A3, A5); interocular area: 2 (M2), 7 (S0, S2, S5, S6); postocular area: 2+2 (Pa5, Pa6); posterior area: 3+3 (Pa2, Pp3, Pe5). </p>
            <p>Macrochaetae densely arranged along anterior margin of Th. II and sparsely on each tergum, most mesochaetae present laterally and posteriorly on terga. Number of macrochaetae or large mesochaetae in posterior row as 3 (p2, p3, p4), 3 (p1, p3, p5)/3 (m2, m3, m4), 3 (m2, m3, m4), 4 (p1, p3, p6, p7), 2 (p6, p7), 4 (m2, m3, m5, m6) on Th. II–III/Abd. I–V respectively. Th. II behind anterior marginal macrochaetae cluster with a file of macrochaetae, including a3, a4, and a4ae; central macrochaetae a2, a5, m1, m2 and m3 arranged approximately in triangle, m4 lateral to m2. Th. III with anterior macrochaeta a4; Abd. III with two anterior macrochaetae m3 and m6; Abd. IV with antero-lateral macrochaeta m6. Bothriotricha 2 (a6, m6), 1 (m6)/ 0, 0, 1 (a5), 2 (a2, a5), 0, 0 on Th. II–III/ Abd. I–VI. Tergal pseudopores 1, 1/1, 1, 1, 1, 0, 0 from Th. II–III/Abd. I–VI near body axis.</p>
            <p> Trochantero-femoral organ with 1, 1 slender chaetae. Each tibiotarsus with distal whorl of 11 chaetae, among which the six ventral (inner) chaetae as ordinary chaetae, the five dorsal (outer) chaetae modified into a central clavate tenent hair, a pair of short accessory chaetae and a pair of slender guard chaetae. Unguis slender, with basointernal ridges about 1/4–1/3 distant from base; lateral teeth pointed, of moderate size. Inner edge of unguis with basal tooth and 4–6 more distal teeth, sub-basal tooth subequal to others or slightly larger. Unguiculus lanceolate. Rami of tenaculum with 4 + 4 teeth. Manubrium ventrally scaled without chaetae; laterally with large round scales and a row of strong chaetae; dorsally with two longitudinal strips of chaetae, including 2 + 2 pointed prominent chaetae at 1/3–2/5 and 3/4–4/5 the length of manubrium from base, respectively. Inner edge of each dorsal chaetal strip with a narrow strip of scales. Dens basally with a pointed prominent dorsal chaeta, without modified outer strong chaetae of  Plutomurus - type or differentiated inner large scales of  Pogonognathellus - type. Dental spines compound, with numerous small denticles on surface. Distal end of dental spine array with a single large spine. Dens dorsally with ordinary chaetae and plumose chaetae, ventrally with dense scales and several apical chaetae. Mucro elongated, with numerous chaetae; apical tooth subequal to or slightly larger than subapical tooth; two dorsal lamellae running from proximal edge of subapical tooth, outer lamella ending in distal edge of inner basal tooth, inner lamella ending at the base of mucro, intermediate teeth located on outer lamella; both basal teeth with proximal lamellae, outer basal tooth with a toothlet. </p>
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	https://treatment.plazi.org/id/0398CE51FFACFF9FFF6BFCD6FD5365CB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yu, Daoyuan;Xue, Huiying;Zhang, Feng	Yu, Daoyuan, Xue, Huiying, Zhang, Feng (2025): Two new species of Tomocerus (Ocreatomurus) Yu from Xizang, China (Collembola, Tomoceridae). Zootaxa 5570 (2): 371-379, DOI: 10.11646/zootaxa.5570.2.8, URL: https://doi.org/10.11646/zootaxa.5570.2.8
0398CE51FFAFFF9AFF6BFF03FCB5652C.text	0398CE51FFAFFF9AFF6BFF03FCB5652C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tomocerus (Ocreatomurus) gyirongensis Yu & Xue & Zhang 2025	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Tomocerus (Ocreatomurus) gyirongensis sp. nov.</p>
            <p>Figs 1–14, Table 1</p>
            <p>
                 Type material.   Holotype: male on slide, coniferous and broad-leaved mixed forest,  
                <a title="Search Plazi for locations around (long 85.25235/lat 28.4392)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=85.25235&amp;materialsCitation.latitude=28.4392">Gyirong Valley</a>
                 , Gyirong County,  
                <a title="Search Plazi for locations around (long 85.25235/lat 28.4392)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=85.25235&amp;materialsCitation.latitude=28.4392">Xigaze</a>
                 , Xizang Autonomous Region, China, 85°15.141′E, 28°26.352′N (WGS84), alt. 2965 m, 6.viii.2018, leg. Daoyuan Yu, Yiming Wei and Chunyan Qin (Sample code 18XZ13)  .   Paratypes: one female, two males and one juvenile on slides, 19 subadults and juveniles in ethanol, same data as holotype; four subadults in ethanol, under a wood board in  
                <a title="Search Plazi for locations around (long 85.32822/lat 28.375383)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=85.32822&amp;materialsCitation.latitude=28.375383">Jipu Village</a>
                 , Gyirong County,  
                <a title="Search Plazi for locations around (long 85.32822/lat 28.375383)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=85.32822&amp;materialsCitation.latitude=28.375383">Xigaze</a>
                 , Xizang Autonomous Region, China, 85°19.693′E, 28°22.523′N, alt. 2765 m, 7.viii.2018, leg. Daoyuan Yu, Yiming Wei and Chunyan Qin (Sample code 18XZ15). All types deposited in NJAU  . 
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            <p>Description. Body length 3.0– 4.3 mm (average 3.8, five specimens). Body ground color pale yellow. Ant. III and IV purple. Diffuse reddish-purple pigment present mainly on Ant. I and II, head, coxae, tibiotarsi, ventral tube, and manubrium. In juveniles (Fig. 1), pigment on head distributed more evenly, and light reddish-purple pigment also around bothriotricha on terga, forming diffuse patches on Th. II and spots on other segments. In subadults (Fig. 2) and adults (Fig. 3), pigment on head concentrating anteriorly around mouth and antennal bases, and the patterns around bothriotricha becoming indistinct. In fully grown adults (Fig. 3), purple pigment more intense, becoming brownish.</p>
            <p>Intact four-segmented antennae 0.75–0.81 × length of body (average 0.78, two specimens). Length ratio of Ant I: II: III: IV = 1.0: 1.6: 8.5–10.1: 0.9–1.2. Ant. III basally with 3–8 scales. Cephalic dorsal macrochaetotaxy as in Fig. 4. Posterior margin of head with 35–40 chaetae on each side.</p>
            <p>Pattern of body macrochaetotaxy as in Fig. 5.</p>
            <p>Single chaeta of femoral organ longer than that of trochanteral organ (Fig. 6). Tibiotarsi I, II, III ventrally with 8–10, 8–9, 7–9 strong chaetae, 5–8, 7–8, 7–9 of them blunt and spine-like, respectively (Fig. 7). Tenent hair 0.8–1.0 × the length of inner edge of unguis; accessory chaetae on tibiotarsi I and II almost equal to each other, and subequal to or shorter than pretarsal chaetae; on tibiotarsus III, anterior accessory chaeta longer than pretarsal chaetae, while posterior accessory chaeta subequal to pretarsal chaetae; guard chaetae 0.8–0.9 × the length of tenent hair. Unguiculus about 0.56–0.64 × the length of unguis (average 0.58, four specimens), its inner edge with 0–1 tooth (Fig. 8).</p>
            <p>Ventral tube scaled on both faces; anterior face with 28–50 (average 38, three specimens) chaetae on each side; posterior face with 82–122 (average 103, three specimens) chaetae; each lateral flap with 137–170 (average 149, three specimens) chaetae and 2–8 scales. Anterior face of tenaculum unscaled, with 14–16 (average 15, three specimens) chaetae (Fig. 9). Ratio manubrium: dens: mucro = 3.3–3.9: 4.4–5.0: 1.0. Manubrium laterally with 9–10 strong chaetae; dorsally with 211–301 (average 252, three specimens) chaetae of different sizes on each dorsal chaetal strip, 6–12 (average eight, four specimens) pseudopores almost arranging in a file and scale strip running from base to about 3/4 the length of manubrium (Fig. 10). External distal corner chaeta as large as or shorter than medium-sized mesochaetae in chaetal strip (Fig. 11). Dental spine formula sensu Folsom (1913) as 4–8/3–5, 1, basal spines arranged in two rows (Fig. 12). Denticles almost evenly distributed on dental spines except for tips (Fig. 13). Mucro with 4–6 (average five, four specimens) intermediate teeth (Fig. 14).</p>
            <p>Etymology. Named after the type locality: Gyirong Valley.</p>
            <p>Habitat. In leaf litter, rotten woods and live mosses in a coniferous and broad-leaved mixed forest; also under a wood board beside a cottage in a village.</p>
            <p> Remarks. Within  Tomocerus (Ocreatomurus) ,  T. (O.) gyirongensis sp. nov. belongs to a species group with only one large distal dental spine. This group currently contains  T. (O.) spinulus Chen &amp; Christiansen, 1998 ,  T. (O.) leyensis Yu &amp; Deharveng, 2018 ,  T. (O.) pseudospinulus Gong, Qin &amp; Yu, 2018 and  T. (O.) paraspinulus Gong, Qin &amp; Yu, 2018 . Among them,  T. (O.) gyirongensis sp. nov. is most similar to  T. (O.) spinulus in the presence of pointed prominent chaetae on manubrium and dens, and the small size of denticles on dental spines, but the new species can be distinguished from the later by the following characters (Table 1): first,  T. (O.) gyirongensis sp. nov. has a larger number of tibiotarsal strong inner chaetae than  T. (O.) spinulus ; second,  T. (O.) gyirongensis sp. nov. has 2 + 2 prominent chaetae on manubrium, while  T. (O.) spinulus has only 1 + 1 of them (sensu Yu et al. 2018); third, in  T. (O.) gyirongensis sp. nov. the proximal dental spines are arranged strictly in two rows, while in  T. (O.) spinulus these spines are arranged in a single row. In addition to these major differences, the relative length of antennae in  T. (O.) gyirongensis sp. nov. is approximately 20–40 % (average 30 %) longer than that in  T. (O.) spinulus (sensu Yu et al. 2018). Likely due to its larger body size,  T. (O.) gyirongensis sp. nov. also has more chaetae on the ventral tube, tenaculum, and dorsal chaetal strips of the manubrium. </p>
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	https://treatment.plazi.org/id/0398CE51FFAFFF9AFF6BFF03FCB5652C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yu, Daoyuan;Xue, Huiying;Zhang, Feng	Yu, Daoyuan, Xue, Huiying, Zhang, Feng (2025): Two new species of Tomocerus (Ocreatomurus) Yu from Xizang, China (Collembola, Tomoceridae). Zootaxa 5570 (2): 371-379, DOI: 10.11646/zootaxa.5570.2.8, URL: https://doi.org/10.11646/zootaxa.5570.2.8
0398CE51FFA8FF96FF6BFDE8FD0862E4.text	0398CE51FFA8FF96FF6BFDE8FD0862E4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tomocerus (Ocreatomurus) oppositus Yu & Xue & Zhang 2025	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Tomocerus (Ocreatomurus) oppositus sp. nov.</p>
            <p>Figs 15–28, Table 1</p>
            <p>
                 Type material.   Holotype: male on slide, coniferous and broad-leaved mixed forest,  
                <a title="Search Plazi for locations around (long 85.363014/lat 28.368)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=85.363014&amp;materialsCitation.latitude=28.368">Gyirong Valley</a>
                 , Gyirong County,  
                <a title="Search Plazi for locations around (long 85.363014/lat 28.368)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=85.363014&amp;materialsCitation.latitude=28.368">Xigaze</a>
                 , Xizang Autonomous Region, China, 85°21.781′E, 28°22.080′N (WGS84), alt. 2500 m, 7.viii.2018, leg. Daoyuan Yu, Yiming Wei and Chunyan Qin (Sample code 18XZ16)  .   Paratypes: three females and one juvenile on slides, 30 in ethanol, same data as holotype; one female on slide, 18 in ethanol, under a wood board in  
                <a title="Search Plazi for locations around (long 85.32822/lat 28.375383)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=85.32822&amp;materialsCitation.latitude=28.375383">Jipu Village</a>
                 , Gyirong County,  
                <a title="Search Plazi for locations around (long 85.32822/lat 28.375383)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=85.32822&amp;materialsCitation.latitude=28.375383">Xigaze</a>
                 , Xizang Autonomous Region, China, 85°19.693′E, 28°22.523′N, alt. 2765 m, 7.viii.2018, leg. Daoyuan Yu, Yiming Wei and Chunyan Qin (Sample code 18XZ15). All types deposited in NJAU  . 
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            <p>Description. Body length 3.4–5.1 mm (average 4.3, five specimens). In subadults and small adults, head, body, and coxae deep purple; trochanteres, ventral tube, and manubrium with diffuse purple pigment; Ant. I–II, femora, tibiotarsi, and dens pale or bright yellow; Ant. III basally pale and gradually darker towards apex; Ant. IV purple (Fig. 15). In large adults, purple pigment more intense, becoming dark brown, and extending to Ant. I–II, femora, and dens (Fig. 16).</p>
            <p>Intact four-segmented antennae 0.7–0.8 × length of body (average 0.75, three specimens). Length ratio of A nt I: II: III: IV = 1.0: 1.4–1.6: 7.3–8.5: 1.0–1.6. Ant. III occasionally with 1–2 scales at base. Cephalic dorsal macrochaetotaxy as in Fig. 17. Posterior margin of head with 35–45 chaetae on each side.</p>
            <p>Pattern of body macrochaetotaxy as in Fig. 18.</p>
            <p>Tibiotarsi I, II, III ventrally with 8–9, 8–10, 6–10 strong chaetae, 5–6, 6–8, 6–9 of them blunt and spine-like, respectively (Fig. 19). Tenent hair 0.79–0.86 × the length of inner edge of unguis; accessory chaetae on tibiotarsus I equal to each other and shorter than pretarsal chaetae; on tibiotarsi II and III, anterior accessory chaeta subequal to or slightly longer than pretarsal chaetae, while posterior accessory chaeta shorter than pretarsal chaetae (Fig. 20); guard chaetae as long as tenent hair. Unguiculus about 0.57–0.65 × the length of unguis (average 0.61, four specimens), its inner edge with one tooth (Fig. 21).</p>
            <p>Ventral tube scaled on both faces; anterior face with 35–55 (average 46, five specimens) chaetae on each side; posterior face with 75–100 (average 89, five specimens) chaetae; each lateral flap with 90–180 (average 144, five specimens) chaetae and occasionally 1–3 scales. Anterior face of tenaculum with 10–17 (average 13, five specimens) chaetae and 0–3 scales (Fig. 22). Ratio manubrium: dens: mucro = 3.4–4.1: 4.2–5.2: 1.0. Manubrium laterally with 7–11 strong chaetae; dorsally with 170–280 (average 237, five specimens) chaetae of different sizes on each chaetal strip, 12–21 (average 16, five specimens) pseudopores almost arranging in a file and scale strip running from base to about 2/3 length of manubrium (Fig. 23). External distal corner chaeta as large as medium-sized mesochaetae in chaetal strip (Fig. 24). Dental spine formula sensu Folsom (1913) as 4–6/4–8, 1, basal spines arranged either in one row (Fig. 25) or two rows (Fig. 26). Denticles on dental spines distributed more densely on ventral side than on dorsal side of spine (Fig. 27). Mucro with 4–7 (average six, five specimens) intermediate teeth (Fig. 28).</p>
            <p> Etymology. The Latin word  oppositus (masculine adjective) means “opposite”, referencing the unique color pattern of the new species with dark ground and pale appendages, opposite to most other species of the subgenus with pale ground and darker appendages. </p>
            <p>Habitat. In ground litter, dead logs and live mosses on steep slopes in a closed forest; also under a wood board beside a cottage in a village.</p>
            <p> Remarks. At first glance, this new species resembles many others of the subgenus  Tomocerus (Striatomurus) for its dark body color. However, other characters, particularly the compound dental spines, put it in  Ocreatomurus . Within the subgenus,  T. (O.) oppositus sp. nov. possesses a unique color pattern with intensely dark ground and pale appendages, which opposes to most other species with pale ground and darker appendages. A few other species of  Ocreatomurus , such as  T. (O.) postantennalis Yu, Zhang &amp; Deharveng, 2014 ,  T. (O.) dong Yu &amp; Li, 2016 ,  T. (O.) deharvengi Yu &amp; Li, 2016 and  T. (O.) cthulhu Yu &amp; Li, 2016 , also have dark pigment covering almost the whole body, however, pigment in these species is diffusely distributed, and the whole animals are from light to dark grey, not as intensely dark as  T. (O.) oppositus sp. nov. In spite of the color pattern, this new species is most similar to  T. (O.) spinulus in the absence of blunt chaetae on manubrium, presence of single large distal dental spine, and small size of denticles on dental spines, but is different from the later mainly in the larger body size, relatively longer antennae (10 %–38 % longer, average 20 %), larger number of blunt inner chaetae on tibiotarsi, and number and position of manubrial prominent chaetae (Table 1). </p>
            <p> A few variations in macrochaetotaxy exist in  T. (O.) oppositus sp. nov. On the head, macrochaeta A3 is absent from the right side in paratype #5, and macrochaeta S0 is absent in paratype #4. On Th. II, macrochaeta m2 is absent from one side in paratypes #2 and #4, whereas chaeta ap5, which is a microchaeta in most types, becomes a small macrochaeta on the left side in paratype #6. </p>
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	https://treatment.plazi.org/id/0398CE51FFA8FF96FF6BFDE8FD0862E4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yu, Daoyuan;Xue, Huiying;Zhang, Feng	Yu, Daoyuan, Xue, Huiying, Zhang, Feng (2025): Two new species of Tomocerus (Ocreatomurus) Yu from Xizang, China (Collembola, Tomoceridae). Zootaxa 5570 (2): 371-379, DOI: 10.11646/zootaxa.5570.2.8, URL: https://doi.org/10.11646/zootaxa.5570.2.8
