taxonID	type	description	language	source
03A57F5D0779432E12BBFA21C716F894.taxon	distribution	Distribution. Palaearctic: China (Inner Mongolia), Japan, Finland, Norway, Sweden; Nearctic: Canada, USA (Soós & Papp, 1986 c; Wang et al., 2023; https: // www. gbif. org).	en	Shi, Li, Wang, Zhiwei, Ma, Rui, Wang, Yanwei, Yang, Yingang (2025): A survey of the Calyptratae (Diptera) in the Northern Primitive Forest Region of Greater Khingan Mountains, Inner Mongolia with a checklist of fifty-five species. Zootaxa 5683 (4): 451-491, DOI: 10.11646/zootaxa.5683.4.1, URL: https://doi.org/10.11646/zootaxa.5683.4.1
03A57F5D0779432D12BBF80DC65DFE34.taxon	distribution	Distribution. Palaearctic: China (Inner Mongolia *, Heilongjiang, Jilin, Shanxi), Austria, Belarus, Czech Republic, Finland, Denmark, France, Germany, Hungary, Italy, Lithuania, Netherlands, Norway, Romania, Slovakia, Sweden, Switzerland, The United Kingdom, Russia, Ukraine; Nearctic: Canada, USA (Yang et al., 2020 b; Soós & Papp, 1986 c; https: // www. gbif. org).	en	Shi, Li, Wang, Zhiwei, Ma, Rui, Wang, Yanwei, Yang, Yingang (2025): A survey of the Calyptratae (Diptera) in the Northern Primitive Forest Region of Greater Khingan Mountains, Inner Mongolia with a checklist of fifty-five species. Zootaxa 5683 (4): 451-491, DOI: 10.11646/zootaxa.5683.4.1, URL: https://doi.org/10.11646/zootaxa.5683.4.1
03A57F5D077A432D12BBFE14C5EDFD70.taxon	distribution	Distribution. Palaearctic: China (Inner Mongolia) (Wang et al., 2023); Nearctic: Canada, USA, Greenland (https: // www. gbif. org).	en	Shi, Li, Wang, Zhiwei, Ma, Rui, Wang, Yanwei, Yang, Yingang (2025): A survey of the Calyptratae (Diptera) in the Northern Primitive Forest Region of Greater Khingan Mountains, Inner Mongolia with a checklist of fifty-five species. Zootaxa 5683 (4): 451-491, DOI: 10.11646/zootaxa.5683.4.1, URL: https://doi.org/10.11646/zootaxa.5683.4.1
03A57F5D077B432C12BBFA13C07DF916.taxon	distribution	Distribution. Widely distributed all over the world (Soós & Papp, 1986 c).	en	Shi, Li, Wang, Zhiwei, Ma, Rui, Wang, Yanwei, Yang, Yingang (2025): A survey of the Calyptratae (Diptera) in the Northern Primitive Forest Region of Greater Khingan Mountains, Inner Mongolia with a checklist of fifty-five species. Zootaxa 5683 (4): 451-491, DOI: 10.11646/zootaxa.5683.4.1, URL: https://doi.org/10.11646/zootaxa.5683.4.1
03A57F5D077B432C12BBFF59C77EFE10.taxon	description	(Figs 1 ‒ 2) Specimens examined. CHINA: 3 ♂, Inner Mongolia, Genhe City, Mangui Town, Northern Primitive Forest Region of Greater Khingan Mountains, Changliangbeishan, Houdu, burned area in 2002, Pinus pumila, Malaise trap 3, 120 º 54 ʹ 40.10 ʺE, 52 º 25 ʹ 31.24 ʺN, 1009 m, 14. vii. 2022, L. SHI, Z. W. WANG, R. MA.	en	Shi, Li, Wang, Zhiwei, Ma, Rui, Wang, Yanwei, Yang, Yingang (2025): A survey of the Calyptratae (Diptera) in the Northern Primitive Forest Region of Greater Khingan Mountains, Inner Mongolia with a checklist of fifty-five species. Zootaxa 5683 (4): 451-491, DOI: 10.11646/zootaxa.5683.4.1, URL: https://doi.org/10.11646/zootaxa.5683.4.1
03A57F5D077B432C12BBFF59C77EFE10.taxon	distribution	Distribution. Palaearctic: China #, Norway, Sweden, Finland, Russia; Nearctic: Canada, USA, Greenland (Soós & Papp, 1986 c; Xue et al., 2009; https: // www. gbif. org).	en	Shi, Li, Wang, Zhiwei, Ma, Rui, Wang, Yanwei, Yang, Yingang (2025): A survey of the Calyptratae (Diptera) in the Northern Primitive Forest Region of Greater Khingan Mountains, Inner Mongolia with a checklist of fifty-five species. Zootaxa 5683 (4): 451-491, DOI: 10.11646/zootaxa.5683.4.1, URL: https://doi.org/10.11646/zootaxa.5683.4.1
03A57F5D077B432B12BBF933C630FF68.taxon	distribution	Distribution. China (Inner Mongolia *, Heilongjiang, Liaoning, Beijing); Japan, Albania, Austria, Belgium, Czech Republic, Denmark, Estonia, Finland, Germany, France, Hungary, Italy, Netherlands, Norway, Poland, Portugal, Romania, Slovakia, Spain, Sweden, Switzerland, The United Kingdom, Russia; (Yang et al., 2020 b; Soós & Papp, 1986 c; https: // www. gbif. org).	en	Shi, Li, Wang, Zhiwei, Ma, Rui, Wang, Yanwei, Yang, Yingang (2025): A survey of the Calyptratae (Diptera) in the Northern Primitive Forest Region of Greater Khingan Mountains, Inner Mongolia with a checklist of fifty-five species. Zootaxa 5683 (4): 451-491, DOI: 10.11646/zootaxa.5683.4.1, URL: https://doi.org/10.11646/zootaxa.5683.4.1
03A57F5D077D432A12BBFD35C5D9FB88.taxon	description	(Figs 3 ‒ 5) Specimens examined. CHINA: 6 ♂ 6 ♀, Inner Mongolia, Genhe City, Mangui Town, Northern Primitive Forest Region of Greater Khingan Mountains, Changliangbeishan, Houdu, burned area in 2002, Larix gmelinii, Malaise trap 2, 120 ° 54 ' 15.76 " E, 52 ° 25 ' 31.24 " N, 1032 m, 14. vii. 2022, L. SHI, Z. W. WANG, R. MA.	en	Shi, Li, Wang, Zhiwei, Ma, Rui, Wang, Yanwei, Yang, Yingang (2025): A survey of the Calyptratae (Diptera) in the Northern Primitive Forest Region of Greater Khingan Mountains, Inner Mongolia with a checklist of fifty-five species. Zootaxa 5683 (4): 451-491, DOI: 10.11646/zootaxa.5683.4.1, URL: https://doi.org/10.11646/zootaxa.5683.4.1
03A57F5D077D432A12BBFD35C5D9FB88.taxon	distribution	Distribution. Palaearctic: China #, Japan, Russia, Austria, Denmark, Finland, Germany, Sweden, Norway, Latvia, Stwitzerland, The United Kingdom; Nearctic: Canada, USA (Soós & Papp, 1986 c; https: // www. gbif. org; http: // www. diptera. org /).	en	Shi, Li, Wang, Zhiwei, Ma, Rui, Wang, Yanwei, Yang, Yingang (2025): A survey of the Calyptratae (Diptera) in the Northern Primitive Forest Region of Greater Khingan Mountains, Inner Mongolia with a checklist of fifty-five species. Zootaxa 5683 (4): 451-491, DOI: 10.11646/zootaxa.5683.4.1, URL: https://doi.org/10.11646/zootaxa.5683.4.1
03A57F5D077D432A12BBFF11C7B0FDA4.taxon	distribution	Distribution. Palaearctic: China (Inner Mongolia, Heilongjiang, Liaoning, Xinjiang, Sichuan, Tibet), Ireland, Croatia, Czech Republic, Slovakia, Albania, Algeria, Bulgaria, Denmark, United Kingdom, France, Germany, Italy, Malta, Norway, Finland, Russia, Serbia, Syria, Sweden; Morocco, Tunisia; Oriental: China (Taiwan) (Xue & Zhao, 1996; Yang et al., 2020 b; https: // www. gbif. org; http: // www. diptera. org /).	en	Shi, Li, Wang, Zhiwei, Ma, Rui, Wang, Yanwei, Yang, Yingang (2025): A survey of the Calyptratae (Diptera) in the Northern Primitive Forest Region of Greater Khingan Mountains, Inner Mongolia with a checklist of fifty-five species. Zootaxa 5683 (4): 451-491, DOI: 10.11646/zootaxa.5683.4.1, URL: https://doi.org/10.11646/zootaxa.5683.4.1
03A57F5D077F432812BBFF59C0B1FE34.taxon	distribution	Distribution. Palaearctic: China (Inner Mongolia) (Wang et al., 2023), Germany, Sweden, Russia; Nearctic: Canada, USA, Greenland (Soós & Papp, 1986 c; https: // www. gbif. org; http: // www. diptera. org /).	en	Shi, Li, Wang, Zhiwei, Ma, Rui, Wang, Yanwei, Yang, Yingang (2025): A survey of the Calyptratae (Diptera) in the Northern Primitive Forest Region of Greater Khingan Mountains, Inner Mongolia with a checklist of fifty-five species. Zootaxa 5683 (4): 451-491, DOI: 10.11646/zootaxa.5683.4.1, URL: https://doi.org/10.11646/zootaxa.5683.4.1
03A57F5D0770432712BBFF11C4BAFDA4.taxon	distribution	Distribution. Palaearctic: China (Inner Mongolia, Heilongjiang, Hebei), Japan, Austria, Czech Republic, Slovakia, Germany, Denmark, France, The United Kingdom, Italy, Norway, Romania, Sweden, Finland, Russia; Nearctic: Canada, USA; Neotropical: Madagascar (Yang et al., 2020 b; Soós & Papp, 1986 c; https: // www. gbif. org).	en	Shi, Li, Wang, Zhiwei, Ma, Rui, Wang, Yanwei, Yang, Yingang (2025): A survey of the Calyptratae (Diptera) in the Northern Primitive Forest Region of Greater Khingan Mountains, Inner Mongolia with a checklist of fifty-five species. Zootaxa 5683 (4): 451-491, DOI: 10.11646/zootaxa.5683.4.1, URL: https://doi.org/10.11646/zootaxa.5683.4.1
03A57F5D0770432712BBFA33C110F88A.taxon	description	(Figs 6 ‒ 7) Specimens examined. CHINA: 2 ♂, Inner Mongolia, Genhe City, Mangui Town, Northern Primitive Forest Region of Greater Khingan Mountains, Abba River in first branch, unburned area, Pinus pumila, Malaise trap 44, 121 ° 30 ' 10.80 " E, 52 ° 12 ' 48.70 " N, 897 m, 14. vii. 2022, L. SHI, Z. W. WANG, R. MA.	en	Shi, Li, Wang, Zhiwei, Ma, Rui, Wang, Yanwei, Yang, Yingang (2025): A survey of the Calyptratae (Diptera) in the Northern Primitive Forest Region of Greater Khingan Mountains, Inner Mongolia with a checklist of fifty-five species. Zootaxa 5683 (4): 451-491, DOI: 10.11646/zootaxa.5683.4.1, URL: https://doi.org/10.11646/zootaxa.5683.4.1
03A57F5D0770432712BBFA33C110F88A.taxon	distribution	Distribution. Palaearctic: China #, Japan, Austria, Czech Republic, Slovakia, Denmark, Finland, France, Germany, Hungary, Netherlands, Norway, Iceland, Poland, Spain, Sweden, The United Kingdom, Russia; Nearctic: Canada, USA, Greenland (Soós & Papp, 1986 c; Michelsen, 2014, 2015; https: // www. gbif. org; http: // www. diptera. org /).	en	Shi, Li, Wang, Zhiwei, Ma, Rui, Wang, Yanwei, Yang, Yingang (2025): A survey of the Calyptratae (Diptera) in the Northern Primitive Forest Region of Greater Khingan Mountains, Inner Mongolia with a checklist of fifty-five species. Zootaxa 5683 (4): 451-491, DOI: 10.11646/zootaxa.5683.4.1, URL: https://doi.org/10.11646/zootaxa.5683.4.1
03A57F5D0771432612BBFF59C0A7FE10.taxon	description	(Figs 8 ‒ 9) Specimens examined. CHINA: 4 ♂, Inner Mongolia, Genhe City, Mangui Town, Northern Primitive Forest Region of Greater Khingan Mountains, Changliangbeishan, Liaochang, unburned area, Pinus pumila, Malaise trap 10, 121 ° 02 ' 40.50 " E, 52 ° 28 ' 55.60 " N, 937 m, 14. vii. 2022, L. SHI, Z. W. WANG, R. MA.	en	Shi, Li, Wang, Zhiwei, Ma, Rui, Wang, Yanwei, Yang, Yingang (2025): A survey of the Calyptratae (Diptera) in the Northern Primitive Forest Region of Greater Khingan Mountains, Inner Mongolia with a checklist of fifty-five species. Zootaxa 5683 (4): 451-491, DOI: 10.11646/zootaxa.5683.4.1, URL: https://doi.org/10.11646/zootaxa.5683.4.1
03A57F5D0771432612BBFF59C0A7FE10.taxon	distribution	Distribution. Palaearctic: China #, Czech Republic, Denmark, Finland, France, Germany, Italy, Norway, Poland, Slovakia, Sweden, The United Kingdom, Russia; Nearctic: Canada, USA (Soós & Papp, 1986 c).	en	Shi, Li, Wang, Zhiwei, Ma, Rui, Wang, Yanwei, Yang, Yingang (2025): A survey of the Calyptratae (Diptera) in the Northern Primitive Forest Region of Greater Khingan Mountains, Inner Mongolia with a checklist of fifty-five species. Zootaxa 5683 (4): 451-491, DOI: 10.11646/zootaxa.5683.4.1, URL: https://doi.org/10.11646/zootaxa.5683.4.1
03A57F5D0772432512BBF90CC038F82C.taxon	distribution	Distribution. Widely distributed all over the world (Prado e Castro et al., 2016).	en	Shi, Li, Wang, Zhiwei, Ma, Rui, Wang, Yanwei, Yang, Yingang (2025): A survey of the Calyptratae (Diptera) in the Northern Primitive Forest Region of Greater Khingan Mountains, Inner Mongolia with a checklist of fifty-five species. Zootaxa 5683 (4): 451-491, DOI: 10.11646/zootaxa.5683.4.1, URL: https://doi.org/10.11646/zootaxa.5683.4.1
03A57F5D0772432512BBFBF5C5C1FACB.taxon	description	(Figs. 10 ‒ 11) Specimens examined. CHINA: 2 ♂, Inner Mongolia, Genhe City, Mangui Town, Northern Primitive Forest Region of Greater Khingan Mountains, Yanling Road at 5 km, unburned area, Pinus pumila, Malaise trap 24, 121 ° 06 ' 30.15 " E, 52 ° 53 ' 18.40 " N, 843 m, 13. vii. 2022, L. SHI, Z. W. WANG, R. MA.	en	Shi, Li, Wang, Zhiwei, Ma, Rui, Wang, Yanwei, Yang, Yingang (2025): A survey of the Calyptratae (Diptera) in the Northern Primitive Forest Region of Greater Khingan Mountains, Inner Mongolia with a checklist of fifty-five species. Zootaxa 5683 (4): 451-491, DOI: 10.11646/zootaxa.5683.4.1, URL: https://doi.org/10.11646/zootaxa.5683.4.1
03A57F5D0772432512BBFBF5C5C1FACB.taxon	distribution	Distribution. Palaearctic: China #, Japan, Austria, Czech Republic, Denmark, Finland, France, Germany, Netherlands, Norway, Sweden, The United Kingdom, Russia; Nearctic: Canada, USA, Greenland (Soós & Papp, 1986 c; Michelsen, 2015).	en	Shi, Li, Wang, Zhiwei, Ma, Rui, Wang, Yanwei, Yang, Yingang (2025): A survey of the Calyptratae (Diptera) in the Northern Primitive Forest Region of Greater Khingan Mountains, Inner Mongolia with a checklist of fifty-five species. Zootaxa 5683 (4): 451-491, DOI: 10.11646/zootaxa.5683.4.1, URL: https://doi.org/10.11646/zootaxa.5683.4.1
03A57F5D0773432412BBF9EAC052F8E3.taxon	distribution	Distribution. Palaearctic: China (Inner Mongolia, Heilongjiang, Jilin, Liaoning, Hebei, Beijing, Tianjin, Hubei, Shanxi, Shandong, Henan, Shaanxi, Ningxia, Gansu, Qinghai, Anhui,), Mongolia, Japan, North Korea, South Korea, Russia; Oriental: China (Jiangsu, Shanghai, Zhejiang, Jiangxi, Hunan, Sichuan, Chongqing, Guizhou, Fujian, Yunnan) (Yang et al., 2020 b; Soós & Papp, 1986 b; https: // www. gbif. org).	en	Shi, Li, Wang, Zhiwei, Ma, Rui, Wang, Yanwei, Yang, Yingang (2025): A survey of the Calyptratae (Diptera) in the Northern Primitive Forest Region of Greater Khingan Mountains, Inner Mongolia with a checklist of fifty-five species. Zootaxa 5683 (4): 451-491, DOI: 10.11646/zootaxa.5683.4.1, URL: https://doi.org/10.11646/zootaxa.5683.4.1
03A57F5D0774432312BBFB7EC5E6F98F.taxon	distribution	Distribution. Palaearctic: China (Inner Mongolia, Heilongjiang, Jilin, Liaoning, Tianjin, Hebei, Shanxi, Shandong, Shaanxi, Ningxia, Gansu, Qinghai, Xinjiang), Mongolia, North Korea, South Korea, Japan, Turkey, Austria, Czech Republic, Denmark, Estonia, Finland, France, Germany, Greece, Hungary, Ireland, Italy, Luxembourg, Netherlands, Norway, Poland, Portugal, Romania, Spain, Sweden, The United Kingdom, Russia, Ukraine; Nearctic: USA; Oriental: China (Sichuan, Jiangsu, Hainan, Macao, Hong Kong) (Yang et al., 2020 b; Soós & Papp, 1986 b; https: // www. gbif. org).	en	Shi, Li, Wang, Zhiwei, Ma, Rui, Wang, Yanwei, Yang, Yingang (2025): A survey of the Calyptratae (Diptera) in the Northern Primitive Forest Region of Greater Khingan Mountains, Inner Mongolia with a checklist of fifty-five species. Zootaxa 5683 (4): 451-491, DOI: 10.11646/zootaxa.5683.4.1, URL: https://doi.org/10.11646/zootaxa.5683.4.1
03A57F5D0774432212BBF93EC772FEFC.taxon	distribution	Distribution. Palaearctic: China (Inner Mongolia, Heilongjiang, Jilin, Liaoning, Hebei, Beijing, Tianjin, Shanxi, Shandong, Henan, Shaanxi, Ningxia, Gansu, Qinghai, Xinjiang, Tibet), Japan, North Korea, Austria, Finland, France, Germany, Netherlands, Norway, Slovakia, Spain, Sweden, The United Kingdom, Russia; Nearctic: Canada, USA, Greenland; Oriental: China (Jiangsu, Shanghai, Sichuan); Neotropical: Chile; Afrotropical: Madagascar (Yang et al., 2020 b; Soós & Papp, 1986 b; https: // www. gbif. org).	en	Shi, Li, Wang, Zhiwei, Ma, Rui, Wang, Yanwei, Yang, Yingang (2025): A survey of the Calyptratae (Diptera) in the Northern Primitive Forest Region of Greater Khingan Mountains, Inner Mongolia with a checklist of fifty-five species. Zootaxa 5683 (4): 451-491, DOI: 10.11646/zootaxa.5683.4.1, URL: https://doi.org/10.11646/zootaxa.5683.4.1
03A57F5D0775432212BBFE15C159FD70.taxon	description	(Figs 12 ‒ 13) Specimens examined. CHINA: 5 ♂, Inner Mongolia, Genhe City, Mangui Town, Northern Primitive Forest Region of Greater Khingan Mountains, Abba River in first branch, unburned area, Pinus sylvestris var. mongholica, Malaise trap 40, 121 ° 22 ' 46.80 " E, 52 ° 13 ' 23.30 " N, 742 m, 14. vii. 2022, L. SHI, Z. W. WANG, R. MA.	en	Shi, Li, Wang, Zhiwei, Ma, Rui, Wang, Yanwei, Yang, Yingang (2025): A survey of the Calyptratae (Diptera) in the Northern Primitive Forest Region of Greater Khingan Mountains, Inner Mongolia with a checklist of fifty-five species. Zootaxa 5683 (4): 451-491, DOI: 10.11646/zootaxa.5683.4.1, URL: https://doi.org/10.11646/zootaxa.5683.4.1
03A57F5D0775432212BBFE15C159FD70.taxon	distribution	Distribution. Palaearctic: China #; Nearctic: Canada, USA (Stone et al., 1983; https: // www. gbif. org).	en	Shi, Li, Wang, Zhiwei, Ma, Rui, Wang, Yanwei, Yang, Yingang (2025): A survey of the Calyptratae (Diptera) in the Northern Primitive Forest Region of Greater Khingan Mountains, Inner Mongolia with a checklist of fifty-five species. Zootaxa 5683 (4): 451-491, DOI: 10.11646/zootaxa.5683.4.1, URL: https://doi.org/10.11646/zootaxa.5683.4.1
03A57F5D0776432112BBFC07C12AFB62.taxon	distribution	Distribution. Palaearctic: China (Inner Mongolia *, Beijing), Japan, Norway, Sweden, Finland, France, Denmark, Germany, Poland, Czech Republic, Slovakia, Switzerland, Austria, Hungary, Romania; Nearctic: Canada, USA; Oriental: China (Taiwan) (Yang et al., 2020 b; Soós & Papp, 1986 a; Stone et al., 1983; https: // www. gbif. org).	en	Shi, Li, Wang, Zhiwei, Ma, Rui, Wang, Yanwei, Yang, Yingang (2025): A survey of the Calyptratae (Diptera) in the Northern Primitive Forest Region of Greater Khingan Mountains, Inner Mongolia with a checklist of fifty-five species. Zootaxa 5683 (4): 451-491, DOI: 10.11646/zootaxa.5683.4.1, URL: https://doi.org/10.11646/zootaxa.5683.4.1
03A57F5D0776432112BBFAFFC675F9D6.taxon	distribution	Distribution. Palaearctic: China (Inner Mongolia, Liaoning, Shaanxi), Albania, Austria, Belarus, Bulgaria, Czech Republic, Germany, Finland, France, Hungary, Italy, Norway, Poland, Romania, Siberia, Slovakia, Spain, Sweden, The United Kingdom, Russia; Nearctic: Canada, USA (Yang et al., 2020 b; Soós & Papp, 1986 a; Stone et al., 1983; https: // www. gbif. org).	en	Shi, Li, Wang, Zhiwei, Ma, Rui, Wang, Yanwei, Yang, Yingang (2025): A survey of the Calyptratae (Diptera) in the Northern Primitive Forest Region of Greater Khingan Mountains, Inner Mongolia with a checklist of fifty-five species. Zootaxa 5683 (4): 451-491, DOI: 10.11646/zootaxa.5683.4.1, URL: https://doi.org/10.11646/zootaxa.5683.4.1
03A57F5D0776432112BBF973C1EDF84A.taxon	distribution	Distribution. Palaearctic: China (Inner Mongolia *, Liaoning, Hebei, Shanxi, Ningxia), Mongolia, Japan, Latvia, Romania, Siberia, Austria, Belgium, Bulgaria, Czech Republic, Denmark, Finland, France, Germany, Hungary, Italy, Ireland, Netherlands, Norway, Poland, Spain, Sweden, Switzerland, The United Kingdom, Russia; Nearctic: Canada, USA; Oriental: China (Sichuan, Taiwan) (Yang et al., 2020 b; Soós & Papp, 1986 a; https: // www. gbif. org).	en	Shi, Li, Wang, Zhiwei, Ma, Rui, Wang, Yanwei, Yang, Yingang (2025): A survey of the Calyptratae (Diptera) in the Northern Primitive Forest Region of Greater Khingan Mountains, Inner Mongolia with a checklist of fifty-five species. Zootaxa 5683 (4): 451-491, DOI: 10.11646/zootaxa.5683.4.1, URL: https://doi.org/10.11646/zootaxa.5683.4.1
03A57F5D0777432012BBFF59C552FE10.taxon	distribution	Distribution. Palaearctic: China (Inner Mongolia *, Jilin), Japan, Czech Republic, Slovakia, Denmark, Norway, Sweden, Finland, France, Switzerland, The United Kingdom, Germany, Italy; Nearctic: USA and Canada (Zhang et al., 2011).	en	Shi, Li, Wang, Zhiwei, Ma, Rui, Wang, Yanwei, Yang, Yingang (2025): A survey of the Calyptratae (Diptera) in the Northern Primitive Forest Region of Greater Khingan Mountains, Inner Mongolia with a checklist of fifty-five species. Zootaxa 5683 (4): 451-491, DOI: 10.11646/zootaxa.5683.4.1, URL: https://doi.org/10.11646/zootaxa.5683.4.1
03A57F5D0777432012BBFE31C593FC8E.taxon	distribution	Distribution. Palaearctic: China (Inner Mongolia *, Liaoning, Shanxi, Hebei, Gansu), Japan, Belarus, Austria, Belgium, Switzerland, Czech Republic, Slovakia, Germany, Denmark, France, The United Kingdom, Hungary, Italy, Ireland, Norway, Netherlands, Poland, Romania, Sweden, Finland, Russia (Yang et al., 2020 b; Soós & Papp, 1986 a; https: // www. gbif. org).	en	Shi, Li, Wang, Zhiwei, Ma, Rui, Wang, Yanwei, Yang, Yingang (2025): A survey of the Calyptratae (Diptera) in the Northern Primitive Forest Region of Greater Khingan Mountains, Inner Mongolia with a checklist of fifty-five species. Zootaxa 5683 (4): 451-491, DOI: 10.11646/zootaxa.5683.4.1, URL: https://doi.org/10.11646/zootaxa.5683.4.1
03A57F5D0777432012BBFC4DC6A8FB24.taxon	distribution	Distribution. Palaearctic: China (Inner Mongolia, Heilongjiang, Jilin, Liaoning, Shanxi), Japan, Siberia, Tajikistan, Austria, Germany, Finland, Norway, Poland, Slovakia, Sweden, Russia; Nearctic: Canada, USA (Stone et al., 1983; Yang et al., 2020 b; Barták et al., 2021).	en	Shi, Li, Wang, Zhiwei, Ma, Rui, Wang, Yanwei, Yang, Yingang (2025): A survey of the Calyptratae (Diptera) in the Northern Primitive Forest Region of Greater Khingan Mountains, Inner Mongolia with a checklist of fifty-five species. Zootaxa 5683 (4): 451-491, DOI: 10.11646/zootaxa.5683.4.1, URL: https://doi.org/10.11646/zootaxa.5683.4.1
03A57F5D0777432012BBFB25C56EF998.taxon	distribution	Distribution. Palaearctic: China (Inner Mongolia *, Liaoning, Xinjiang), Japan, Estonia, Latvia, Albania, Austria, France, Czech Republic, Denmark, Estonia, Finland, Germany, Hungary, Ireland, Italy, The United Kingdom, Netherlands, Norway, Poland, Romania, Slovakia, Spain, Switzerland, Sweden; Nearctic: Canada, USA (Yang et al., 2020 b).	en	Shi, Li, Wang, Zhiwei, Ma, Rui, Wang, Yanwei, Yang, Yingang (2025): A survey of the Calyptratae (Diptera) in the Northern Primitive Forest Region of Greater Khingan Mountains, Inner Mongolia with a checklist of fifty-five species. Zootaxa 5683 (4): 451-491, DOI: 10.11646/zootaxa.5683.4.1, URL: https://doi.org/10.11646/zootaxa.5683.4.1
03A57F5D0777433F12BBF881C6AEFEA0.taxon	distribution	Distribution. Palaearctic: China (Inner Mongolia *, Shanxi, Xinjiang), Mongolia, Japan, Afghanistan, Kyrgyzstan, Siberia, Tajikistan, Turkey, Austria, Bulgaria, Czech Republic, Denmark, Finland, France, Germany, Italy, Norway, Poland, Romania, Slovakia, Spain, Switzerland, Sweden, The United Kingdom, Russia (Yang et al., 2020 b; Soós & Papp, 1986 a; https: // www. gbif. org).	en	Shi, Li, Wang, Zhiwei, Ma, Rui, Wang, Yanwei, Yang, Yingang (2025): A survey of the Calyptratae (Diptera) in the Northern Primitive Forest Region of Greater Khingan Mountains, Inner Mongolia with a checklist of fifty-five species. Zootaxa 5683 (4): 451-491, DOI: 10.11646/zootaxa.5683.4.1, URL: https://doi.org/10.11646/zootaxa.5683.4.1
03A57F5D0768433E12BBF885C768FEA0.taxon	distribution	Distribution. Palaearctic: China (Inner Mongolia *, Shanxi, Qinghai), Japan, Belarus, Estonia, Latvia, Siberia, Turkey, Austria, Belgium, Czech Republic, Denmark, Finland, France, Germany, Hungary, Italy, Ireland, Netherlands, Norway, Poland, Romania, Slovakia, Spain, Sweden, The United Kingdom, Russia; Nearctic: Canada, USA (Yang et al., 2020 b; Soós & Papp, 1986 a; https: // www. gbif. org).	en	Shi, Li, Wang, Zhiwei, Ma, Rui, Wang, Yanwei, Yang, Yingang (2025): A survey of the Calyptratae (Diptera) in the Northern Primitive Forest Region of Greater Khingan Mountains, Inner Mongolia with a checklist of fifty-five species. Zootaxa 5683 (4): 451-491, DOI: 10.11646/zootaxa.5683.4.1, URL: https://doi.org/10.11646/zootaxa.5683.4.1
03A57F5D0769433E12BBFE31C088FD4C.taxon	description	(Figs 14 ‒ 15) Specimens examined. CHINA: 4 ♂, Inner Mongolia, Genhe City, Mangui Town, Northern Primitive Forest Region of Greater Khingan Mountains, Abba River in third branch, burned area in 2018, Larix gmelinii, Malaise trap 31, 121 ° 25 ' 38.40 " E, 52 ° 19 ' 54.70 " N, 651 m, 14. vii. 2022, L. SHI, Z. W. WANG, R. MA.	en	Shi, Li, Wang, Zhiwei, Ma, Rui, Wang, Yanwei, Yang, Yingang (2025): A survey of the Calyptratae (Diptera) in the Northern Primitive Forest Region of Greater Khingan Mountains, Inner Mongolia with a checklist of fifty-five species. Zootaxa 5683 (4): 451-491, DOI: 10.11646/zootaxa.5683.4.1, URL: https://doi.org/10.11646/zootaxa.5683.4.1
03A57F5D0769433E12BBFE31C088FD4C.taxon	distribution	Distribution. Palaearctic: China #, Russia; Nearctic: USA (Huckett, 1965; https: // www. gbif. org).	en	Shi, Li, Wang, Zhiwei, Ma, Rui, Wang, Yanwei, Yang, Yingang (2025): A survey of the Calyptratae (Diptera) in the Northern Primitive Forest Region of Greater Khingan Mountains, Inner Mongolia with a checklist of fifty-five species. Zootaxa 5683 (4): 451-491, DOI: 10.11646/zootaxa.5683.4.1, URL: https://doi.org/10.11646/zootaxa.5683.4.1
03A57F5D0769433E12BBF91DC031F878.taxon	description	(Figs 16 ‒ 17) Specimens examined. CHINA: 4 ♂, Inner Mongolia, Genhe City, Mangui Town, Northern Primitive Forest Region of Greater Khingan Mountains, Wulonggan Forestry Center 4207, burned area in 2007, Pinus sylvestris var. mongholica, Malaise trap 18, 120 ° 58 ' 55.80 " E, 52 ° 51 ' 53.10 " N, 823 m, 13. vii. 2022, L. SHI, Z. W. WANG, R. MA.	en	Shi, Li, Wang, Zhiwei, Ma, Rui, Wang, Yanwei, Yang, Yingang (2025): A survey of the Calyptratae (Diptera) in the Northern Primitive Forest Region of Greater Khingan Mountains, Inner Mongolia with a checklist of fifty-five species. Zootaxa 5683 (4): 451-491, DOI: 10.11646/zootaxa.5683.4.1, URL: https://doi.org/10.11646/zootaxa.5683.4.1
03A57F5D0769433E12BBF91DC031F878.taxon	distribution	Distribution. Palaearctic: China #, Russia (Hennig, 1961; https: // www. gbif. org).	en	Shi, Li, Wang, Zhiwei, Ma, Rui, Wang, Yanwei, Yang, Yingang (2025): A survey of the Calyptratae (Diptera) in the Northern Primitive Forest Region of Greater Khingan Mountains, Inner Mongolia with a checklist of fifty-five species. Zootaxa 5683 (4): 451-491, DOI: 10.11646/zootaxa.5683.4.1, URL: https://doi.org/10.11646/zootaxa.5683.4.1
03A57F5D076A433D12BBF9F5C0A7F824.taxon	distribution	Distribution. Palaearctic: China (Inner Mongolia, Heilongjiang, Jilin, Liaoning, Shanxi, Shaanxi, Xinjiang, Sichuan), Mongolia, Armenia, Azerbaijan, Belarus, Estonia, Georgia, Kazakhstan, Kyrgyzstan, Latvia, Turkey, Ukraine, Austria, Albania, Belgium, Bulgaria, Czech Republic, Denmark, Finland, France, Germany, Hungary, Italy, Ireland, Netherlands, Norway, Poland, Romania, Sweden, Switzerland, Slovakia, Spain, The United Kingdom, Russia; Nearctic: Canada, USA (Yang et al., 2020 b; Soós & Papp, 1986 a; https: // www. gbif. org).	en	Shi, Li, Wang, Zhiwei, Ma, Rui, Wang, Yanwei, Yang, Yingang (2025): A survey of the Calyptratae (Diptera) in the Northern Primitive Forest Region of Greater Khingan Mountains, Inner Mongolia with a checklist of fifty-five species. Zootaxa 5683 (4): 451-491, DOI: 10.11646/zootaxa.5683.4.1, URL: https://doi.org/10.11646/zootaxa.5683.4.1
03A57F5D076B433C12BBFBA2C6E6FAE3.taxon	distribution	Distribution. Palaearctic: China (Inner Mongolia *, Jilin), Finland, Norway, Sweden, Russia; Nearctic: Canada, USA (Sorokina, 2023; https: // www. gbif. org).	en	Shi, Li, Wang, Zhiwei, Ma, Rui, Wang, Yanwei, Yang, Yingang (2025): A survey of the Calyptratae (Diptera) in the Northern Primitive Forest Region of Greater Khingan Mountains, Inner Mongolia with a checklist of fifty-five species. Zootaxa 5683 (4): 451-491, DOI: 10.11646/zootaxa.5683.4.1, URL: https://doi.org/10.11646/zootaxa.5683.4.1
03A57F5D076B433C12BBF9D2C497F8EB.taxon	distribution	Distribution. Palaearctic: China (Inner Mongolia *, Qinghai), Mongolia, Belarus, Latvia, Kazakhstan, Siberia, Austria, Czech Republic, Slovakia, Denmark, Finland, France, Germany, Hungary, Italy, Netherlands, Norway, Poland, Sweden, The United Kingdom, Russia; Nearctic: Canada, USA (Soós & Papp, 1986 a; Vikhrev, 2015, 2020).	en	Shi, Li, Wang, Zhiwei, Ma, Rui, Wang, Yanwei, Yang, Yingang (2025): A survey of the Calyptratae (Diptera) in the Northern Primitive Forest Region of Greater Khingan Mountains, Inner Mongolia with a checklist of fifty-five species. Zootaxa 5683 (4): 451-491, DOI: 10.11646/zootaxa.5683.4.1, URL: https://doi.org/10.11646/zootaxa.5683.4.1
03A57F5D076C433B12BBFF59C6D4FE6C.taxon	description	(Figs 18 ‒ 19) Specimens examined. CHINA: 6 ♂, Inner Mongolia, Genhe City, Mangui Town, Northern Primitive Forest Region of Greater Khingan Mountains, Changliangbeishan, Houdu, burned area in 2002, Pinus sylvestris var. mongholica, Malaise trap 6, 120 ° 56 ' 53.20 " E, 52 ° 26 ' 01.00 " N, 985 m, 14. vii. 2022, L. SHI, Z. W. WANG, R. MA.	en	Shi, Li, Wang, Zhiwei, Ma, Rui, Wang, Yanwei, Yang, Yingang (2025): A survey of the Calyptratae (Diptera) in the Northern Primitive Forest Region of Greater Khingan Mountains, Inner Mongolia with a checklist of fifty-five species. Zootaxa 5683 (4): 451-491, DOI: 10.11646/zootaxa.5683.4.1, URL: https://doi.org/10.11646/zootaxa.5683.4.1
03A57F5D076C433B12BBFF59C6D4FE6C.taxon	distribution	Distribution. Palaearctic: China #, Estonia, Latvia, Siberia, Austria, Belgium, Czech Republic, Slovakia, Denmark, Finland, France, Germany, Ireland, Iceland, Norway, Poland, Sweden, The United Kingdom, Russia; Nearctic: Canada, USA (Soós & Papp, 1986 a).	en	Shi, Li, Wang, Zhiwei, Ma, Rui, Wang, Yanwei, Yang, Yingang (2025): A survey of the Calyptratae (Diptera) in the Northern Primitive Forest Region of Greater Khingan Mountains, Inner Mongolia with a checklist of fifty-five species. Zootaxa 5683 (4): 451-491, DOI: 10.11646/zootaxa.5683.4.1, URL: https://doi.org/10.11646/zootaxa.5683.4.1
03A57F5D076D433A12BBFAD6C5E6F9EF.taxon	description	(Figs 20 ‒ 21) Specimens examined. CHINA: 3 ♂, Inner Mongolia, Genhe City, Mangui Town, Northern Primitive Forest Region of Greater Khingan Mountains, main line between Qiqian and Yonganshan at 126 km, unburned area, Pinus sylvestris var. mongholica, Malaise trap 12, 121 ° 02 ' 45.50 " E, 52 ° 29 ' 32.80 " N, 856.7 m, 14. vii. 2022, L. SHI, Z. W. WANG, R. MA.	en	Shi, Li, Wang, Zhiwei, Ma, Rui, Wang, Yanwei, Yang, Yingang (2025): A survey of the Calyptratae (Diptera) in the Northern Primitive Forest Region of Greater Khingan Mountains, Inner Mongolia with a checklist of fifty-five species. Zootaxa 5683 (4): 451-491, DOI: 10.11646/zootaxa.5683.4.1, URL: https://doi.org/10.11646/zootaxa.5683.4.1
03A57F5D076D433A12BBFAD6C5E6F9EF.taxon	distribution	Distribution. Palaearctic: China #, Sweden, The United Kingdom; Nearctic: Canada (Soós & Papp, 1986 a; https: // www. gbif. org).	en	Shi, Li, Wang, Zhiwei, Ma, Rui, Wang, Yanwei, Yang, Yingang (2025): A survey of the Calyptratae (Diptera) in the Northern Primitive Forest Region of Greater Khingan Mountains, Inner Mongolia with a checklist of fifty-five species. Zootaxa 5683 (4): 451-491, DOI: 10.11646/zootaxa.5683.4.1, URL: https://doi.org/10.11646/zootaxa.5683.4.1
03A57F5D076D433912BBF8FAC6BCFF68.taxon	distribution	Distribution. Palaearctic: China (Inner Mongolia, Heilongjiang, Jilin), South Korea, Russia; Neactic: Canada, USA (Soós & Papp, 1986 a; https: // www. gbif. org).	en	Shi, Li, Wang, Zhiwei, Ma, Rui, Wang, Yanwei, Yang, Yingang (2025): A survey of the Calyptratae (Diptera) in the Northern Primitive Forest Region of Greater Khingan Mountains, Inner Mongolia with a checklist of fifty-five species. Zootaxa 5683 (4): 451-491, DOI: 10.11646/zootaxa.5683.4.1, URL: https://doi.org/10.11646/zootaxa.5683.4.1
03A57F5D076E433912BBF98CC4BAF8E4.taxon	distribution	Distribution. Palaearctic: China (Inner Mongolia *, Xinjiang) (Yang et al., 2020 b), Japan, Austria, Bulgaria, Czech Republic, Slovakia, France, Romania, Sweden, Finland, Russia (Soós & Papp, 1986 a; https: // www. gbif. org).	en	Shi, Li, Wang, Zhiwei, Ma, Rui, Wang, Yanwei, Yang, Yingang (2025): A survey of the Calyptratae (Diptera) in the Northern Primitive Forest Region of Greater Khingan Mountains, Inner Mongolia with a checklist of fifty-five species. Zootaxa 5683 (4): 451-491, DOI: 10.11646/zootaxa.5683.4.1, URL: https://doi.org/10.11646/zootaxa.5683.4.1
03A57F5D076F433812BBF9FAC6D7F82F.taxon	distribution	Distribution. Palaearctic: China (Inner Mongolia *, Heilongjiang, Liaoning, Xinjiang) (Yang et al., 2020 b), Mongolia, North Korea, Japan, Iran, Belarus, Estonia, Kirghizistan, Kazakhstan, Lithuania, Austria, Belgium, Bulgaria, Switzerland, Czech Republic, Slovakia, Germany, Spain, France, Ireland, The United Kingdom, Hungary, Italy, Norway, Netherlands, Poland, Romania, Sweden, Finland, Moldavia, Russia, Ukraine (Soós & Papp, 1986 a); Nearctic: Canada, USA (https: // www. gbif. org)	en	Shi, Li, Wang, Zhiwei, Ma, Rui, Wang, Yanwei, Yang, Yingang (2025): A survey of the Calyptratae (Diptera) in the Northern Primitive Forest Region of Greater Khingan Mountains, Inner Mongolia with a checklist of fifty-five species. Zootaxa 5683 (4): 451-491, DOI: 10.11646/zootaxa.5683.4.1, URL: https://doi.org/10.11646/zootaxa.5683.4.1
03A57F5D076F433812BBFB66C095FA27.taxon	distribution	Distribution. Palaearctic: China (Inner Mongolia *, Heilongjiang, Jilin, Liaoning), Monglia, Japan, Norway, Sweden, Finland, Russia (Soós & Papp, 1986 a); Afrotropical: Madagascar (https: // www. gbif. org).	en	Shi, Li, Wang, Zhiwei, Ma, Rui, Wang, Yanwei, Yang, Yingang (2025): A survey of the Calyptratae (Diptera) in the Northern Primitive Forest Region of Greater Khingan Mountains, Inner Mongolia with a checklist of fifty-five species. Zootaxa 5683 (4): 451-491, DOI: 10.11646/zootaxa.5683.4.1, URL: https://doi.org/10.11646/zootaxa.5683.4.1
03A57F5D0760433712BBFEE9C5E6FDDC.taxon	description	(Figs 22 ‒ 23) Specimens examined. CHINA: 4 ♂, Inner Mongolia, Genhe City, Mangui Town, Northern Primitive Forest Region of Greater Khingan Mountains, Abba River in first branch, unburned are, Pinus pumila, Malaise trap 44, 121 ° 30 ' 10.80 " E, 52 ° 12 ' 48.70 " N, 897 m, 14. vii. 2022, L. SHI, Z. W. WANG, R. MA.	en	Shi, Li, Wang, Zhiwei, Ma, Rui, Wang, Yanwei, Yang, Yingang (2025): A survey of the Calyptratae (Diptera) in the Northern Primitive Forest Region of Greater Khingan Mountains, Inner Mongolia with a checklist of fifty-five species. Zootaxa 5683 (4): 451-491, DOI: 10.11646/zootaxa.5683.4.1, URL: https://doi.org/10.11646/zootaxa.5683.4.1
03A57F5D0760433712BBFEE9C5E6FDDC.taxon	distribution	Distribution. Palaearctic: China #, Mongolia, Japan, Austria, Bulgaria, Switzerland, Germnay, Denmark, France, Hungary, Ireland, Norway, Poland, Sweden, Finland, The United Kingdom, Russia (Soós & Papp, 1986 a; https: // www. gbif. org).	en	Shi, Li, Wang, Zhiwei, Ma, Rui, Wang, Yanwei, Yang, Yingang (2025): A survey of the Calyptratae (Diptera) in the Northern Primitive Forest Region of Greater Khingan Mountains, Inner Mongolia with a checklist of fifty-five species. Zootaxa 5683 (4): 451-491, DOI: 10.11646/zootaxa.5683.4.1, URL: https://doi.org/10.11646/zootaxa.5683.4.1
03A57F5D0761433612BBFB13C086F9A5.taxon	description	(Figs 24 ‒ 25) Specimens examined. CHINA: 3 ♂, Inner Mongolia, Genhe City, Mangui Town, Northern Primitive Forest Region of Greater Khingan Mountains, Abba River in third branch, burned area in 2018, Larix gmelinii, Malaise trap 31, 121 ° 25 ' 38.40 " E, 52 ° 19 ' 54.70 " N, 651 m, 14. VII. 2022, L. SHI, Z. W. WANG, R. MA.	en	Shi, Li, Wang, Zhiwei, Ma, Rui, Wang, Yanwei, Yang, Yingang (2025): A survey of the Calyptratae (Diptera) in the Northern Primitive Forest Region of Greater Khingan Mountains, Inner Mongolia with a checklist of fifty-five species. Zootaxa 5683 (4): 451-491, DOI: 10.11646/zootaxa.5683.4.1, URL: https://doi.org/10.11646/zootaxa.5683.4.1
03A57F5D0761433612BBFB13C086F9A5.taxon	distribution	Distribution. Palaearctic: China #, Finland (Kahanpää & Haarto, 2014), Bulgaria, Spain, Hungary, Norway, Sweden, The United Kingdom (Soós & Papp, 1986 a), Belarus (Makovetskaya & Vikhrev, 2019), Poland, Russia (http: // www. diptera. org), Pakistan (https: // www. gbif. org); Nearctic: Canada (https: // www. gbif. org).	en	Shi, Li, Wang, Zhiwei, Ma, Rui, Wang, Yanwei, Yang, Yingang (2025): A survey of the Calyptratae (Diptera) in the Northern Primitive Forest Region of Greater Khingan Mountains, Inner Mongolia with a checklist of fifty-five species. Zootaxa 5683 (4): 451-491, DOI: 10.11646/zootaxa.5683.4.1, URL: https://doi.org/10.11646/zootaxa.5683.4.1
03A57F5D0761433612BBF9A7C74EF819.taxon	distribution	Distribution. Palaearctic: China (Inner Mongolia *, Heilongjiang, Qinghai), Japan, Mongolia, Austria, Albania, Belgium, Czech Republic, Slovakia, Finland, Germany, Denmark, Spain, France, Hungary, Italy, Ireland, Iceland, Norway, Netherlands, Poland, Romania, Switzerland, The United Kingdom, Russia; Nearctic: Canada, USA (Yang et al., 2020 b; https: // www. gbif. org; http: // www. diptera. org).	en	Shi, Li, Wang, Zhiwei, Ma, Rui, Wang, Yanwei, Yang, Yingang (2025): A survey of the Calyptratae (Diptera) in the Northern Primitive Forest Region of Greater Khingan Mountains, Inner Mongolia with a checklist of fifty-five species. Zootaxa 5683 (4): 451-491, DOI: 10.11646/zootaxa.5683.4.1, URL: https://doi.org/10.11646/zootaxa.5683.4.1
03A57F5D0762433512BBF9EBC712F8DD.taxon	distribution	Distribution. Palaearctic: China (Inner Mongolia *, Heilongjiang, Xinjiang, Qinghai, Tibet), Mongolia, Austria, Bulgaria, Czech Republic, Slovakia, Finland, Germany, Spain, Italy, Norway, Poland, Romania, Sweden, Switzerland; Oriental: China (Sichuan, Yunnan); Nearctic: Canada, USA; Afrotropical: Madagascar (Sorokina & Ovtshinnikova, 2023; Yang et al., 2020 b; https: // www. gbif. org).	en	Shi, Li, Wang, Zhiwei, Ma, Rui, Wang, Yanwei, Yang, Yingang (2025): A survey of the Calyptratae (Diptera) in the Northern Primitive Forest Region of Greater Khingan Mountains, Inner Mongolia with a checklist of fifty-five species. Zootaxa 5683 (4): 451-491, DOI: 10.11646/zootaxa.5683.4.1, URL: https://doi.org/10.11646/zootaxa.5683.4.1
03A57F5D0763433412BBF9A7C534F856.taxon	distribution	Distribution. Palaearctic: China (Inner Mongolia, Heilongjiang, Jilin, Liaoning, Hebei, Beijing, Shanxi, Shaanxi, Xinjiang, Tibet), Mongolia, Korea, Japan, Kyrgyzstan, Tajikistan, Uzbekistan, Kazakhstan, Pakistan, Russia, Syria, Israel, Turkey, Austria, Albania, Bulgaria, Switzerland, Czech Republic, Slovakia, Cyprus, Germany, Denmark, Spain, France, The United Kingdom, Greece, Hungary, Italy, Ireland, Norway, Netherlands, Portugal, Poland, Romania, Sweden, Finland; Nearctic: Canada, USA, Mexico (Yang et al., 2020 b; Falk et al., 2024; https: // www. gbif. org).	en	Shi, Li, Wang, Zhiwei, Ma, Rui, Wang, Yanwei, Yang, Yingang (2025): A survey of the Calyptratae (Diptera) in the Northern Primitive Forest Region of Greater Khingan Mountains, Inner Mongolia with a checklist of fifty-five species. Zootaxa 5683 (4): 451-491, DOI: 10.11646/zootaxa.5683.4.1, URL: https://doi.org/10.11646/zootaxa.5683.4.1
03A57F5D0763433412BBFB83C0A2FA3A.taxon	distribution	Distribution. Palaearctic: China (Inner Mongolia *), Japan, South Korea, Austria, Czech Republic, Slovakia, Denmark, Finland, Germany, Hungary, Italy, Ireland, Netherlands, Norway, Poland, Romania, Sweden, The United Kingdom, Russia; Oriental: China (Guizhou, Taiwan) (Yang et al., 2020 b; https: // www. gbif. org).	en	Shi, Li, Wang, Zhiwei, Ma, Rui, Wang, Yanwei, Yang, Yingang (2025): A survey of the Calyptratae (Diptera) in the Northern Primitive Forest Region of Greater Khingan Mountains, Inner Mongolia with a checklist of fifty-five species. Zootaxa 5683 (4): 451-491, DOI: 10.11646/zootaxa.5683.4.1, URL: https://doi.org/10.11646/zootaxa.5683.4.1
03A57F5D0764433312BBFE79C6E9FC84.taxon	distribution	Distribution. Palaearctic: China (Inner Mongolia, Heilongjiang, Jilin, Liaoning, Hebei, Shanxi, Shaanxi, Xinjiang, Tibet), Mongolia, Japan, Russia, Kyrgyzstan, Finland, Germany, Norway, Poland, Sweden, Netherlands, The United Kingdom, Kazakhstan, Armenia, Azerbaijan, Austria, Belarus, Belgium, Bulgaria, Czech Republic, Slovakia, Denmark, Estonia, France, Greece, Georgia, Hungary, Ireland, Italy, Lithuania, Moldova, Romania, Spain, Ukraine; Nearctic: Canada, USA; Oriental: China (Sichuan, Fujian, Yunnan), India (Yang et al., 2020 b; https: // www. gbif. org; http: // www. diptera. org).	en	Shi, Li, Wang, Zhiwei, Ma, Rui, Wang, Yanwei, Yang, Yingang (2025): A survey of the Calyptratae (Diptera) in the Northern Primitive Forest Region of Greater Khingan Mountains, Inner Mongolia with a checklist of fifty-five species. Zootaxa 5683 (4): 451-491, DOI: 10.11646/zootaxa.5683.4.1, URL: https://doi.org/10.11646/zootaxa.5683.4.1
03A57F5D0765433212BBFC51C05AFB08.taxon	description	(Figs 26 ‒ 27) Specimens examined. CHINA: 3 ♂, Inner Mongolia, Genhe City, Mangui Town, Northern Primitive Forest Region of Greater Khingan Mountains, Yanling Road at 5 km, unburned area, Pinus pumila, Malaise trap 24, 121 ° 06 ' 30.15 " E, 52 ° 53 ' 18.40 " N, 843 m, 13. vii. 2022, L. SHI, Z. W. WANG, R. MA.	en	Shi, Li, Wang, Zhiwei, Ma, Rui, Wang, Yanwei, Yang, Yingang (2025): A survey of the Calyptratae (Diptera) in the Northern Primitive Forest Region of Greater Khingan Mountains, Inner Mongolia with a checklist of fifty-five species. Zootaxa 5683 (4): 451-491, DOI: 10.11646/zootaxa.5683.4.1, URL: https://doi.org/10.11646/zootaxa.5683.4.1
03A57F5D0765433212BBFC51C05AFB08.taxon	distribution	Distribution. Palaearctic: China #, Austria Czech Republic, Slovakia (Šifner & Bernasconi, 2018), Finland, Norway, Sweden, The United Kingdom; Nearctic: Canada (https: // www. gbif. org).	en	Shi, Li, Wang, Zhiwei, Ma, Rui, Wang, Yanwei, Yang, Yingang (2025): A survey of the Calyptratae (Diptera) in the Northern Primitive Forest Region of Greater Khingan Mountains, Inner Mongolia with a checklist of fifty-five species. Zootaxa 5683 (4): 451-491, DOI: 10.11646/zootaxa.5683.4.1, URL: https://doi.org/10.11646/zootaxa.5683.4.1
03A57F5D0765433212BBFEA1C045FDC0.taxon	distribution	Distribution. Palaearctic: China (Inner Mongolia, Heilongjiang, Liaoning, Jilin, Beijing, Shanxi, Shaanxi, Gansu, Henan, Sichuan), North Korea, Japan, Mongolia, Russia, Azerbaijan, Belarus, Bulgaria, Poland, Germany, Czech Republic, Hungary, Netherlands (Yang et al., 2020 b; https: // www. gbif. org).	en	Shi, Li, Wang, Zhiwei, Ma, Rui, Wang, Yanwei, Yang, Yingang (2025): A survey of the Calyptratae (Diptera) in the Northern Primitive Forest Region of Greater Khingan Mountains, Inner Mongolia with a checklist of fifty-five species. Zootaxa 5683 (4): 451-491, DOI: 10.11646/zootaxa.5683.4.1, URL: https://doi.org/10.11646/zootaxa.5683.4.1
03A57F5D0766433112BBFF11C6B7FE48.taxon	description	(Figs 28 ‒ 29) Specimens examined. CHINA: 2 ♂, Inner Mongolia, Genhe City, Mangui Town, Northern Primitive Forest Region of Greater Khingan Mountains, Wulonggan Forestry Center, unburned area, Pinus sylvestris var. mongholica, Malaise trap 19, 120 ° 55 ' 03.45 " E, 52 ° 47 ' 47.98 " N, 816 m, 13. vii. 2022, L. SHI, Z. W. WANG, R. MA.	en	Shi, Li, Wang, Zhiwei, Ma, Rui, Wang, Yanwei, Yang, Yingang (2025): A survey of the Calyptratae (Diptera) in the Northern Primitive Forest Region of Greater Khingan Mountains, Inner Mongolia with a checklist of fifty-five species. Zootaxa 5683 (4): 451-491, DOI: 10.11646/zootaxa.5683.4.1, URL: https://doi.org/10.11646/zootaxa.5683.4.1
03A57F5D0766433112BBFF11C6B7FE48.taxon	distribution	Distribution. Palaearctic: China #, Mongolia, Russia, Norway, Sweden (Becker, 1894; Ozerov, 2014 b; de Jong, 2016; Bernasconi, 2021; http: // www. diptera. org).	en	Shi, Li, Wang, Zhiwei, Ma, Rui, Wang, Yanwei, Yang, Yingang (2025): A survey of the Calyptratae (Diptera) in the Northern Primitive Forest Region of Greater Khingan Mountains, Inner Mongolia with a checklist of fifty-five species. Zootaxa 5683 (4): 451-491, DOI: 10.11646/zootaxa.5683.4.1, URL: https://doi.org/10.11646/zootaxa.5683.4.1
03A57F5D0767433012BBFA22C499F82F.taxon	distribution	Distribution. Palaearctic: China (Inner Mongolia, Heilongjiang, Jilin, Liaoning, Beijing, Hebei, Shanxi, Shaanxi, Ningxia, Xinjiang, Tibet), Mongolia, Pakistan, South Korea, Japan, Russia, Austria, Belarus, Belgium, Czech Republic, Denmark, Finland, France, Germany, Greece, Hungary, Israel, Lithuania, Netherlands, Norway, Poland, Spain, Sweden, Switzerland, The United Kingdom, Ukraine (Fleming et al., 2017; https: // www. gbif. org); Nearctic: Canada, USA (https: // www. gbif. org); Neotropical: Brazil (https: // www. gbif. org); Australian / Oceanian: Australia, Papua New Guinea; Oriental: China (Sichuan, Yunnan, Guangxi, Taiwan), India, Nepal (Yang et al., 2020 b).	en	Shi, Li, Wang, Zhiwei, Ma, Rui, Wang, Yanwei, Yang, Yingang (2025): A survey of the Calyptratae (Diptera) in the Northern Primitive Forest Region of Greater Khingan Mountains, Inner Mongolia with a checklist of fifty-five species. Zootaxa 5683 (4): 451-491, DOI: 10.11646/zootaxa.5683.4.1, URL: https://doi.org/10.11646/zootaxa.5683.4.1
03A57F5D0758430F12BBFEE9C57CFD80.taxon	description	(Figs 30 ‒ 31) Specimens examined. CHINA: 2 ♂, Inner Mongolia, Genhe City, Mangui Town, Northern Primitive Forest Region of Greater Khingan Mountains, Abba River in first branch, unburned area, Pinus sylvestris var. mongholica, Malaise trap 39, 121 ° 22 ' 46.80 " E, 52 ° 13 ' 23.30 " N, 742 m, 14. vii. 2022, L. SHI, Z. W. WANG, R. MA.	en	Shi, Li, Wang, Zhiwei, Ma, Rui, Wang, Yanwei, Yang, Yingang (2025): A survey of the Calyptratae (Diptera) in the Northern Primitive Forest Region of Greater Khingan Mountains, Inner Mongolia with a checklist of fifty-five species. Zootaxa 5683 (4): 451-491, DOI: 10.11646/zootaxa.5683.4.1, URL: https://doi.org/10.11646/zootaxa.5683.4.1
03A57F5D0758430F12BBFEE9C57CFD80.taxon	distribution	Distribution. Palaearctic: China #, Norway, Sweden (https: // www. gbif. org); Nearctic: Canada, USA (O'Hara et al., 2004).	en	Shi, Li, Wang, Zhiwei, Ma, Rui, Wang, Yanwei, Yang, Yingang (2025): A survey of the Calyptratae (Diptera) in the Northern Primitive Forest Region of Greater Khingan Mountains, Inner Mongolia with a checklist of fifty-five species. Zootaxa 5683 (4): 451-491, DOI: 10.11646/zootaxa.5683.4.1, URL: https://doi.org/10.11646/zootaxa.5683.4.1
03A57F5D0759430E12BBFBBBC607FAD5.taxon	distribution	Distribution. Palaearctic: China (Inner Mongolia, Jilin, Liaoning, Beijing) (Yang et al., 2020 b), South Korea (Lee & Han, 2008), Japan (Shima, 1988;).	en	Shi, Li, Wang, Zhiwei, Ma, Rui, Wang, Yanwei, Yang, Yingang (2025): A survey of the Calyptratae (Diptera) in the Northern Primitive Forest Region of Greater Khingan Mountains, Inner Mongolia with a checklist of fifty-five species. Zootaxa 5683 (4): 451-491, DOI: 10.11646/zootaxa.5683.4.1, URL: https://doi.org/10.11646/zootaxa.5683.4.1
03A57F5D0759430E12BBF987C5EDF8E2.taxon	distribution	Distribution. Palaearctic: China (Inner Mongolia *, Heilongjiang, Henan), Mongolia, Japan, Kazakhstan, Russia (Yang et al., 2020 b), Finland, Norway, Sweden, The United Kingdom; Nearctic: Canada; Afrotropical: Congo (https: // www. gbif. org).	en	Shi, Li, Wang, Zhiwei, Ma, Rui, Wang, Yanwei, Yang, Yingang (2025): A survey of the Calyptratae (Diptera) in the Northern Primitive Forest Region of Greater Khingan Mountains, Inner Mongolia with a checklist of fifty-five species. Zootaxa 5683 (4): 451-491, DOI: 10.11646/zootaxa.5683.4.1, URL: https://doi.org/10.11646/zootaxa.5683.4.1
03A57F5D075A430D12BBFF11C001FE6C.taxon	description	(Figs 32 ‒ 33) Specimens examined. CHINA: 5 ♂, Inner Mongolia, Genhe City, Mangui Town, Northern Primitive Forest Region of Greater Khingan Mountains, Wulonggan Forestry Center 4217, unburned area, Larix gmelinii, Malaise trap 13, 120 º 53 ' 40.10 " E, 52 º 47 ' 05.26 " N, 789 m, 13. vii. 2022, L. SHI, Z. W. WANG, R. MA.	en	Shi, Li, Wang, Zhiwei, Ma, Rui, Wang, Yanwei, Yang, Yingang (2025): A survey of the Calyptratae (Diptera) in the Northern Primitive Forest Region of Greater Khingan Mountains, Inner Mongolia with a checklist of fifty-five species. Zootaxa 5683 (4): 451-491, DOI: 10.11646/zootaxa.5683.4.1, URL: https://doi.org/10.11646/zootaxa.5683.4.1
03A57F5D075A430D12BBFF11C001FE6C.taxon	distribution	Distribution. Palaearctic: China #, Finland, Norway, Sweden (https: // www. gbif. org).	en	Shi, Li, Wang, Zhiwei, Ma, Rui, Wang, Yanwei, Yang, Yingang (2025): A survey of the Calyptratae (Diptera) in the Northern Primitive Forest Region of Greater Khingan Mountains, Inner Mongolia with a checklist of fifty-five species. Zootaxa 5683 (4): 451-491, DOI: 10.11646/zootaxa.5683.4.1, URL: https://doi.org/10.11646/zootaxa.5683.4.1
03A57F5D075D430A12BBFD89C030FC84.taxon	description	(Figs 36 ‒ 37) Specimens examined. CHINA: 2 ♂, Inner Mongolia, Genhe City, Mangui Town, Northern Primitive Forest Region of Greater Khingan Mountains, Wulonggan Forestry Center, unburned area, Pinus sylvestris var. mongholica, Malaise trap 20, 120 º 55 ' 03.45 " E, 52 º 47 ' 47.98 " N, 816 m, 13. vii. 2022, L. SHI, Z. W. WANG, R. MA.	en	Shi, Li, Wang, Zhiwei, Ma, Rui, Wang, Yanwei, Yang, Yingang (2025): A survey of the Calyptratae (Diptera) in the Northern Primitive Forest Region of Greater Khingan Mountains, Inner Mongolia with a checklist of fifty-five species. Zootaxa 5683 (4): 451-491, DOI: 10.11646/zootaxa.5683.4.1, URL: https://doi.org/10.11646/zootaxa.5683.4.1
03A57F5D075D430A12BBFD89C030FC84.taxon	distribution	Distribution. Palaearctic: China #; Nearctic: Canada, USA (O'Hara et al., 2004).	en	Shi, Li, Wang, Zhiwei, Ma, Rui, Wang, Yanwei, Yang, Yingang (2025): A survey of the Calyptratae (Diptera) in the Northern Primitive Forest Region of Greater Khingan Mountains, Inner Mongolia with a checklist of fifty-five species. Zootaxa 5683 (4): 451-491, DOI: 10.11646/zootaxa.5683.4.1, URL: https://doi.org/10.11646/zootaxa.5683.4.1
03A57F5D075D430A12BBFF59C738FE10.taxon	description	(Figs 34 ‒ 35) Specimens examined. CHINA: 2 ♂, Inner Mongolia, Genhe City, Mangui Town, Northern Primitive Forest Region of Greater Khingan Mountains, Changliangbeishan, Houdu, burned area in 2002, Pinus sylvestris var. mongholica, Malaise trap 6, 120 ° 56 ' 53.20 " E, 52 ° 26 ' 01.00 " N, 985 m, 28. vii. 2022, R. MA, Q. J. R. LIU.	en	Shi, Li, Wang, Zhiwei, Ma, Rui, Wang, Yanwei, Yang, Yingang (2025): A survey of the Calyptratae (Diptera) in the Northern Primitive Forest Region of Greater Khingan Mountains, Inner Mongolia with a checklist of fifty-five species. Zootaxa 5683 (4): 451-491, DOI: 10.11646/zootaxa.5683.4.1, URL: https://doi.org/10.11646/zootaxa.5683.4.1
03A57F5D075D430A12BBFF59C738FE10.taxon	distribution	Distribution. Palaearctic: China #, Mongolia, Austria, Denmark, Finland, France, Poland, Sweden, Netherlands, Norway, The United Kingdom, Russia (https: // www. gbif. org).	en	Shi, Li, Wang, Zhiwei, Ma, Rui, Wang, Yanwei, Yang, Yingang (2025): A survey of the Calyptratae (Diptera) in the Northern Primitive Forest Region of Greater Khingan Mountains, Inner Mongolia with a checklist of fifty-five species. Zootaxa 5683 (4): 451-491, DOI: 10.11646/zootaxa.5683.4.1, URL: https://doi.org/10.11646/zootaxa.5683.4.1
