taxonID	type	description	language	source
03AF5F29AB68FF9C1784A8C2E9B39ADD.taxon	description	Boulenophrys palpebralespinosa (Bourret, 1937) (Fig. 3 C): Two specimens were found at night on leaves, at ca. 30 – 90 cm above the ground near a stream, and many other individuals were observed along streams at night in evergreen forest.	en	Pham, Anh Van, Nguyen, Truong Quang, Nguyen, Tao Thien, Le, Minh Duc, Ziegler, Thomas, Tran, Cuong Thien (2024): The herpetofauna of Copia Nature Reserve, Vietnam. Amphibian & Reptile Conservation (e 331) 18 (1): 20-47
03AF5F29AB68FF9C1784A8C2E9B39ADD.taxon	description	Leptobrachium masatakasatoi Matsui, 2013 (Fig. 3 H): Three specimens were found at night on the ground near a stream in evergreen forest. Ophryophryne pachyproctus Kou, 1985 (Fig. 3 I): Two specimens and many others were found at night on leaves, at ca. 50 – 120 cm above the ground near streams, and many other individuals were observed on leaves along streams at night in evergreen forest. Xenophrys maosonensis (Bourret, 1937) (Fig. 3 J): Two specimens were found at night on the ground near a stream, and many other individuals were observed along streams at night in evergreen forest. This is a new record for Copia NR.	en	Pham, Anh Van, Nguyen, Truong Quang, Nguyen, Tao Thien, Le, Minh Duc, Ziegler, Thomas, Tran, Cuong Thien (2024): The herpetofauna of Copia Nature Reserve, Vietnam. Amphibian & Reptile Conservation (e 331) 18 (1): 20-47
03AF5F29AB68FF9C141EA9E2E8AB999D.taxon	description	Microhyla butleri Boulenger, 1900 (Fig. 3 L): Two specimens were found at night on the ground in meadowlands, and other individuals were observed at night in croplands and small puddle edges near the rice fields and forest edges. Microhyla heymonsi Vogt, 1911 (Fig. 3 M): Two specimens were found at night on the ground in croplands, and other individuals were observed at night on the ground in croplands in meadowlands, and along forest trails near forest edges and inside the forest. Microhyla mukhlesuri Hasan, Islam, Kuramoto, Kurabayashi, and Sumida, 2014 (Fig. 3 N): Two specimens were found at night on the ground near small puddle edges near a road, and other individuals were observed at night on the ground near small puddle edges in rice fields, meadowlands, and croplands. This is a new record for Copia NR. Microhyla pulchra (Hallowell, 1861) (Fig. 3 O): Three specimens were found at night on the ground in meadowlands near rice fields, and other individuals were observed at night on the ground around small puddle edges and in meadowlands near rice fields. Micryletta menglienica (Yang and Su, 1980) (Fig. 3 P): Two specimens were found at night on the ground in meadowlands near limestone mountains. This is a new record for Copia NR.	en	Pham, Anh Van, Nguyen, Truong Quang, Nguyen, Tao Thien, Le, Minh Duc, Ziegler, Thomas, Tran, Cuong Thien (2024): The herpetofauna of Copia Nature Reserve, Vietnam. Amphibian & Reptile Conservation (e 331) 18 (1): 20-47
03AF5F29AB68FF81141EAC62EAAE9D9E.taxon	description	Hoplobatrachus chinensis (Osbeck, 1765) (Fig. 3 R): Five individuals were observed at night on the ground, and at pond edges and rice field edges. Limnonectes bannaensis Ye, Fei, Xie, and Jiang, 2007 (Fig. 4 A): Two specimens were found at night on the ground near a stream, and many other individuals were observed at night on the ground near streams or water edges of streams in evergreen forest. This is a new record for Copia NR. Nanorana aenea (Smith, 1922) (Fig. 4 B): Two specimens were found at night on the ground near a stream, and many other individuals were observed at night on the ground near streams in evergreen forest. Quasipaa verrucospinosa (Bourret, 1937) (Fig. 4 C): Individuals were observed at night on rocks in streams or near waterfalls in evergreen forest.	en	Pham, Anh Van, Nguyen, Truong Quang, Nguyen, Tao Thien, Le, Minh Duc, Ziegler, Thomas, Tran, Cuong Thien (2024): The herpetofauna of Copia Nature Reserve, Vietnam. Amphibian & Reptile Conservation (e 331) 18 (1): 20-47
03AF5F29AB75FF811784A862EA579ADE.taxon	description	Amolops vitreus (Bain, Stuart, and Orlov, 2006) (Fig. 4 E): Two specimens were found at night on tree branches near a stream, and six other individuals were observed at night on tree branches near streams in evergreen forest. Odorrana chapaensis (Bourret, 1937) (Fig. 4 F): Two specimens were found at night on a rock near a waterfall in evergreen forest. This is a new record for Copia NR. Odorrana chloronota (Günther, 1876) (Fig. 4 G): Two specimens were found at night on a rock near a stream, and many other individuals were observed at night on tree branches or on rocks near streams in evergreen forest. This is a new record for Copia NR. Odorrana graminea Boulenger, 1900 (Fig. 4 H): One specimen was found at night on a rock near a waterfall in evergreen forest. This is a new record for Copia NR. Odorrana jingdongensis Fei, Ye, and Li, 2001 (Fig. 4 I): Two specimens were found at night on a tree branch near a stream, and many other individuals were observed at night on tree branches or on rocks at ca. 0 – 80 cm above the ground near streams in evergreen forest. This is a new record for Copia NR. Odorrana nasica (Boulenger, 1903) (Fig. 4 J): Two specimens were found at night on a tree branch near a waterfall, and many other individuals were observed at night on tree branches or on rocks at ca. 0 – 90 cm above the ground near waterfalls in evergreen forest. This is a new record for Copia NR. Sylvirana guentheri (Boulenger, 1882) (Fig. 4 K): Individuals were observed at night on the ground near pond edges and streams. The surrounding habitat was rice fields. Sylvirana nigrovittata (Blyth, 1856) (Fig. 4 L): Two specimens were found at night on the ground near a stream, and other individuals were observed at night on the ground, and on stones near streams or at the water edge in streams. The surrounding habitat was evergreen forest. The call concerts of this species were regularly heard in the evening.	en	Pham, Anh Van, Nguyen, Truong Quang, Nguyen, Tao Thien, Le, Minh Duc, Ziegler, Thomas, Tran, Cuong Thien (2024): The herpetofauna of Copia Nature Reserve, Vietnam. Amphibian & Reptile Conservation (e 331) 18 (1): 20-47
03AF5F29AB75FF83141EA9E2EAB3991E.taxon	description	Gracixalus quangi Rowley, Dau, Nguyen, Cao, and Nguyen, 2011 (Fig. 4 O): Two specimens were found at night on leaves near a stream, and other individuals were observed at night while sitting on leaves near streams at ca. 0.8 – 1.5 m above the ground. The surrounding habitat was mixed evergreen forest of small hardwoods, bamboo, and shrubs.	en	Pham, Anh Van, Nguyen, Truong Quang, Nguyen, Tao Thien, Le, Minh Duc, Ziegler, Thomas, Tran, Cuong Thien (2024): The herpetofauna of Copia Nature Reserve, Vietnam. Amphibian & Reptile Conservation (e 331) 18 (1): 20-47
03AF5F29AB75FF83141EA9E2EAB3991E.taxon	description	Theloderma albopunctatum (Liu and Hu, 1962) (Fig. 5 D): One specimen was found at night on edge of the waterhole and six individuals were observed at night at the same perch sites in evergreen forest. Theloderma bicolor (Bourret, 1937) (Fig. 5 E): One specimen was found at night on edge of a water filled tree hole in evergreen forest. Theloderma corticale (Boulenger, 1903) (Fig. 5 F): One specimen was found at night on edge of a water filled tree hole in evergreen forest.	en	Pham, Anh Van, Nguyen, Truong Quang, Nguyen, Tao Thien, Le, Minh Duc, Ziegler, Thomas, Tran, Cuong Thien (2024): The herpetofauna of Copia Nature Reserve, Vietnam. Amphibian & Reptile Conservation (e 331) 18 (1): 20-47
03AF5F29AB77FF831784AC82E88E9CBE.taxon	description	Calotes emma Gray, 1845 (Fig. 6 B): One specimen was found in the morning on a forest path in evergreen forest. This is a new record for Copia NR. Calotes versicolor (Daudin, 1802) (Fig. 6 C): Individuals were observed during the daytime near cultivated lands or bushes in gardens. Draco maculatus (Gray, 1845) (Fig. 6 D): A road-killed individual was found on Road 108. The surrounding habitat was mixed evergreen forest of hardwoods and shrubs. Pseudocalotes brevipes (Werner, 1904) (Fig. 6 E): One specimen was found at night on a tree branch, and six individuals were observed at night while sitting on the same perch sites at ca. 1 – 2 m above the ground in evergreen forest.	en	Pham, Anh Van, Nguyen, Truong Quang, Nguyen, Tao Thien, Le, Minh Duc, Ziegler, Thomas, Tran, Cuong Thien (2024): The herpetofauna of Copia Nature Reserve, Vietnam. Amphibian & Reptile Conservation (e 331) 18 (1): 20-47
03AF5F29AB77FF831410A882E8389E9E.taxon	description	Hemidactylus frenatus Duméril and Bibron, 1836 (Fig. 6 G): Individuals were observed at night on a wall near a light bulb in a residential area. Hemidactylus garnotii Duméril and Bibron, 1836 (Fig. 6 H): Two specimens were found and other individuals were observed at night on limestone karst outcrops at 1 – 3 m above the ground in secondary forest.	en	Pham, Anh Van, Nguyen, Truong Quang, Nguyen, Tao Thien, Le, Minh Duc, Ziegler, Thomas, Tran, Cuong Thien (2024): The herpetofauna of Copia Nature Reserve, Vietnam. Amphibian & Reptile Conservation (e 331) 18 (1): 20-47
03AF5F29AB77FF851410AD62EA329CFE.taxon	description	Eutropis multifasciatus (Kuhl, 1820) (Fig. 6 J): Individuals were observed during the daytime on the ground at road edges, and on shrubs near cultivated land. This is a new record for Copia NR. Plestiodon cf. tamdaoensis (Bourret, 1937) (Fig. 6 K): One specimen was found during the daytime on the ground at road edges near cultivated land. This is a new record for Copia NR. Scincella devorator Darevsky, Orlov, and Ho, 2004 (Fig. 6 L): One specimen was found in the afternoon on a forest path in evergreen forest. Sphenomorphus indicus (Gray, 1853) (Fig. 6 M): One specimen was found in the afternoon on a forest path and other individuals observed on the ground in evergreen forest. Tropidophorus baviensis Bourret, 1939 (Fig. 6 N): Two specimens were found under a carpet of fallen leaves, at a forest edge near evergreen forest. Fig. 6. Lizard species recorded in Copia Nature Reserve, Vietnam. (A) Acanthosaura lepidogaster, (B) Calotes emma, (C) C. versicolor, (D) Draco maculatus, (E) Pseudocalotes brevipes, (F) Gekko reevesii, (G) Hemidactylus frenatus, (H) H. garnotii, (I) Eutropis longicaudatus, (J) E. multifasciatus, (K) Plestiodon cf. tamdaoensis, (L) Scincella devorator, (M) Sphenomorphus indicus, (N) Tropidophorus baviensis, (O) Dopasia harti, and (P) D. ludovici. Dopasia harti (Boulenger, 1899) (Fig. 6 O): One specimen was found during the daytime on the ground at road edges near evergreen forest. Dopasia ludovici (Mocquard, 1905) (Fig. 6 P): Two specimens were found during the daytime on the ground at road edges near evergreen forest. This is a new record for Son La Province.	en	Pham, Anh Van, Nguyen, Truong Quang, Nguyen, Tao Thien, Le, Minh Duc, Ziegler, Thomas, Tran, Cuong Thien (2024): The herpetofauna of Copia Nature Reserve, Vietnam. Amphibian & Reptile Conservation (e 331) 18 (1): 20-47
03AF5F29AB71FF851784AA42EF639BDD.taxon	description	Boiga multomaculata (Boie, 1827) (Fig. 7 D): Three individuals were observed at night on tree branches at ca. 1 – 3 m above the ground in evergreen forest. Calamaria pavimentata Duméril, Bibron, and Duméril, 1854 (Fig. 7 E): One specimen was found at night on the ground while moving across a forest trail near evergreen forest. This is a new record for Copia NR. Coelognathus radiatus (Boie, 1827) (Fig. 7 F): Two individuals were observed in the afternoon while moving across a road near agricultural areas. Dendrelaphis ngansonensis (Bourret, 1935) (Fig. 7 G): One specimen was found during the daytime on the ground on a road edge near the evergreen forest. This is a new record for Copia NR. Dendrelaphis pictus (Gmelin, 1789) (Fig. 7 H): Two specimens were found during the daytime on tree branches at road edges near evergreen forest. Elaphe moellendorffi (Boettger, 1886) (Fig. 7 I): One individual was observed at 1805 h near the entrance of a cave at a forest edge. This is a new record for Copia NR. Elaphe taeniura (Cope, 1861) (Fig. 7 J): Two individuals were observed in the afternoon at the entrance of a cave near a forest edge and agricultural area. This is a new record for Copia NR. Euprepiophis mandarinus (Cantor, 1842) (Fig. 7 K): One individual was observed at night on the ground near a stream in evergreen forest. This is a new record for Copia NR.	en	Pham, Anh Van, Nguyen, Truong Quang, Nguyen, Tao Thien, Le, Minh Duc, Ziegler, Thomas, Tran, Cuong Thien (2024): The herpetofauna of Copia Nature Reserve, Vietnam. Amphibian & Reptile Conservation (e 331) 18 (1): 20-47
03AF5F29AB71FF851784AA42EF639BDD.taxon	description	Liopeltis frenata (Günther, 1858) (Fig. 7 N): A road-killed specimen was found in the morning on a road near evergreen forest. Lycodon fasciatus (Anderson, 1879) (Fig. 7 O): One specimen was found at night on the ground while catching prey near a road, and five other individuals were observed at night on the ground near streams or forest trails in evergreen forest. This is a new record for Copia NR. Lycodon futsingensis (Pope, 1928) (Fig. 7 P): One specimen was found at night on the ground near a stream in evergreen forest. This is a new record for Copia NR. Oligodon catenatus (Blyth, 1854) (Fig. 7 Q): One specimen was found at night on the ground near a forest trail in evergreen forest, and a road-killed specimen was found on the road. Oligodon fasciolatus (Günther, 1864) (Fig. 7 R): One specimen was found at night on the ground in evergreen forest. This is a new record for Copia NR. Oreocryptophis porphyraceus (Cantor, 1839) (Fig. 8 A): One individual was observed at night on the ground while moving near a forest trail in evergreen forest. Ptyas korros (Schlegel, 1837) (Fig. 8 B): Two individuals were observed during the daytime on the ground at the roadside near a forest edge.	en	Pham, Anh Van, Nguyen, Truong Quang, Nguyen, Tao Thien, Le, Minh Duc, Ziegler, Thomas, Tran, Cuong Thien (2024): The herpetofauna of Copia Nature Reserve, Vietnam. Amphibian & Reptile Conservation (e 331) 18 (1): 20-47
03AF5F29AB71FF84141FAFA1E8799A05.taxon	description	Fig. 7. Snake species recorded in Copia Nature Reserve, Vietnam. (A) Indotyphlops braminus, (B) Xenopeltis unicolor, (C) Ahaetulla prasina, (D) Boiga multomaculata, (E) Calamaria pavimentata, (F) Coelognathus radiatus, (G) Dendrelaphis ngansonensis, (H) D. pictus, (I) Elaphe moellendorffi, (J) Elaphe taeniura, (K) Euprepiophis mandarinus, (L) Gonyosoma coeruleum, (M) G. frenatum, (N) Liopeltis frenata, (O) Lycodon fasciatus, (P) L. futsingensis, (Q) Oligodon catenatus, and (R) O. fasciolatus.	en	Pham, Anh Van, Nguyen, Truong Quang, Nguyen, Tao Thien, Le, Minh Duc, Ziegler, Thomas, Tran, Cuong Thien (2024): The herpetofauna of Copia Nature Reserve, Vietnam. Amphibian & Reptile Conservation (e 331) 18 (1): 20-47
03AF5F29AB73FF871784A9E2EB8C991D.taxon	description	Sinomicrurus macclellandi (Reinhardt, 1844) (Fig. 8 H): One individual was observed at night on the ground while moving near a forest trail in evergreen forest.	en	Pham, Anh Van, Nguyen, Truong Quang, Nguyen, Tao Thien, Le, Minh Duc, Ziegler, Thomas, Tran, Cuong Thien (2024): The herpetofauna of Copia Nature Reserve, Vietnam. Amphibian & Reptile Conservation (e 331) 18 (1): 20-47
03AF5F29AB73FF871784AC22E8E59C7D.taxon	description	Hebius boulengeri (Gressitt, 1937) (Fig. 8 K): Two specimens were found at night on the ground near a stream in evergreen forest. This is a new record for Copia NR. Hebius chapaensis (Bourret, 1934) (Fig. 8 L): One specimen was found at night in a stream in evergreen forest. Rhabdophis subminiatus (Schlegel, 1837) (Fig. 8 M): Three individuals were observed during the daytime on grass near the roadside in an agricultural cultivated area. Trimerodytes percarinatus (Boulenger, 1899) (Fig. 8 N): Two individuals were observed at night in streams in evergreen forest. This is a new record for Copia NR.	en	Pham, Anh Van, Nguyen, Truong Quang, Nguyen, Tao Thien, Le, Minh Duc, Ziegler, Thomas, Tran, Cuong Thien (2024): The herpetofauna of Copia Nature Reserve, Vietnam. Amphibian & Reptile Conservation (e 331) 18 (1): 20-47
03AF5F29AB73FF87141EA842EE759E5E.taxon	description	Pareas hamptoni (Boulenger, 1905) (Fig. 8 P): One specimen was found at night while sitting on a tree branch, and many other individuals were observed at night on tree branches at ca. 0.6 – 1.5 m above the ground in evergreen forest. Pareas margaritophorus (Jan, 1866) (Fig. 8 Q): Two specimens were found at night while sitting on tree branches at ca. 0.6 – 1.0 m in evergreen forest.	en	Pham, Anh Van, Nguyen, Truong Quang, Nguyen, Tao Thien, Le, Minh Duc, Ziegler, Thomas, Tran, Cuong Thien (2024): The herpetofauna of Copia Nature Reserve, Vietnam. Amphibian & Reptile Conservation (e 331) 18 (1): 20-47
03AF5F29AB73FF87141EAD62E8029B7D.taxon	description	Ovophis monticola (Günther, 1864) (Fig. 9 B): Two specimens were found at night on the ground near streams in evergreen forest. Trimeresurus albolabris Gray, 1842 (Fig. 9 C): Individuals were observed during the daytime on tree branches at ca. 0.5 – 1.2 m above the ground near secondary forest and cultivated agricultural areas.	en	Pham, Anh Van, Nguyen, Truong Quang, Nguyen, Tao Thien, Le, Minh Duc, Ziegler, Thomas, Tran, Cuong Thien (2024): The herpetofauna of Copia Nature Reserve, Vietnam. Amphibian & Reptile Conservation (e 331) 18 (1): 20-47
03AF5F29AB73FF89141EAE62E85A9E13.taxon	description	Fig. 8. Snake and turtle species recorded in Copia Nature Reserve, Vietnam. (A) Oreocryptophis porphyraceus, (B) Ptyas korros, (C) P. multicincta, (D) Sibynophis collaris, (E) Bungarus fasciatus, (F) B. wanghaotingi, (G) Naja atra, (H) Sinomicrurus macclellandi, (I), Psammodynastes pulverulentus, (J) Fowlea flavipunctatus, (K) Hebius boulengeri, (L) Hebius chapaensis, (M) Rhabdophis subminiatus, (N) Trimerodytes percarinatus, (O) Pareas carinatus, (P) P. hamptoni, (Q) P. margaritophorus, and (R) Pseudoxenodon macrops. Fig. 9. Snake and turtle species recorded in Copia Nature Reserve, Vietnam. (A) Ovophis makazayazaya, (B) O. monticola, (C) Trimeresurus albolabris, (D) Platysternon megacephalum, and (E) Manouria impressa.	en	Pham, Anh Van, Nguyen, Truong Quang, Nguyen, Tao Thien, Le, Minh Duc, Ziegler, Thomas, Tran, Cuong Thien (2024): The herpetofauna of Copia Nature Reserve, Vietnam. Amphibian & Reptile Conservation (e 331) 18 (1): 20-47
