taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
03AC87DDFF8AFF86FCDDC046FC5AD2B0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15701403/files/figure.png	https://doi.org/10.5281/zenodo.15701403	Figs. 1–4. Onthophagus etlaensis and O. anthracinus habitus. 1, Onthophagus etlaensis dorsal view; 2, O. etlaensis anterolateral view; 3, O. anthracinus dorsal view; 4, O. anthracinus anterolateral view.	Figs. 1–4. Onthophagus etlaensis and O. anthracinus habitus. 1, Onthophagus etlaensis dorsal view; 2, O. etlaensis anterolateral view; 3, O. anthracinus dorsal view; 4, O. anthracinus anterolateral view.	2019-12-16	Arriaga-Jiménez, Alfonsina;Escobar-Hernández, Fernando;Rös, Matthias;Kohlmann, Bert		Zenodo	biologists	Arriaga-Jiménez, Alfonsina;Escobar-Hernández, Fernando;Rös, Matthias;Kohlmann, Bert			
03AC87DDFF8AFF86FCDDC046FC5AD2B0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15701405/files/figure.png	https://doi.org/10.5281/zenodo.15701405	Fig. 5. Map of the known distribution of Onthophagus etlaensis (blue dots). Orography of Oaxaca is shown, based on the digital elevation model downloaded from www.inegi.org.mx. Grey areas show the extent of Mexico, see map in the upper left for the location of the state of Oaxaca within Mexico.	Fig. 5. Map of the known distribution of Onthophagus etlaensis (blue dots). Orography of Oaxaca is shown, based on the digital elevation model downloaded from www.inegi.org.mx. Grey areas show the extent of Mexico, see map in the upper left for the location of the state of Oaxaca within Mexico.	2019-12-16	Arriaga-Jiménez, Alfonsina;Escobar-Hernández, Fernando;Rös, Matthias;Kohlmann, Bert		Zenodo	biologists	Arriaga-Jiménez, Alfonsina;Escobar-Hernández, Fernando;Rös, Matthias;Kohlmann, Bert			
03AC87DDFF8AFF86FCDDC046FC5AD2B0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15701407/files/figure.png	https://doi.org/10.5281/zenodo.15701407	Fig. 6. Typical habitat of Onthophagus etlaensis: San Mateo Río Hondo, Sierra Sur, Pinus-Quercus forest.	Fig. 6. Typical habitat of Onthophagus etlaensis: San Mateo Río Hondo, Sierra Sur, Pinus-Quercus forest.	2019-12-16	Arriaga-Jiménez, Alfonsina;Escobar-Hernández, Fernando;Rös, Matthias;Kohlmann, Bert		Zenodo	biologists	Arriaga-Jiménez, Alfonsina;Escobar-Hernández, Fernando;Rös, Matthias;Kohlmann, Bert			
03AC87DDFF8AFF86FCDDC046FC5AD2B0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15701413/files/figure.png	https://doi.org/10.5281/zenodo.15701413	Figs 13–15. Species in the Onthophagus anthracinus species complex. 13, Male pronotum of Onthophagus etlaensis; 14, male pronotum of Onthophagus anthracinus; 15, dorsal view of a male Onthophagus durangoensis.	Figs 13–15. Species in the Onthophagus anthracinus species complex. 13, Male pronotum of Onthophagus etlaensis; 14, male pronotum of Onthophagus anthracinus; 15, dorsal view of a male Onthophagus durangoensis.	2019-12-16	Arriaga-Jiménez, Alfonsina;Escobar-Hernández, Fernando;Rös, Matthias;Kohlmann, Bert		Zenodo	biologists	Arriaga-Jiménez, Alfonsina;Escobar-Hernández, Fernando;Rös, Matthias;Kohlmann, Bert			
03AC87DDFF8AFF86FCDDC046FC5AD2B0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15701415/files/figure.png	https://doi.org/10.5281/zenodo.15701415	Figs 16–20. Aedeagi of the Onthophagus anthracinus species complex. 16, Onthopagus alluvius; 17, O. anthracinus; 18, O. durangoensis; 19, O. etlaensis; 20, O. mextexus.	Figs 16–20. Aedeagi of the Onthophagus anthracinus species complex. 16, Onthopagus alluvius; 17, O. anthracinus; 18, O. durangoensis; 19, O. etlaensis; 20, O. mextexus.	2019-12-16	Arriaga-Jiménez, Alfonsina;Escobar-Hernández, Fernando;Rös, Matthias;Kohlmann, Bert		Zenodo	biologists	Arriaga-Jiménez, Alfonsina;Escobar-Hernández, Fernando;Rös, Matthias;Kohlmann, Bert			
03AC87DDFF8AFF86FCDDC046FC5AD2B0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15701417/files/figure.png	https://doi.org/10.5281/zenodo.15701417	Figs 21–22. Lamella copulatrix and accesory lamellae. 21, Onthophagus anthracinus; 22, O. etlaensis.	Figs 21–22. Lamella copulatrix and accesory lamellae. 21, Onthophagus anthracinus; 22, O. etlaensis.	2019-12-16	Arriaga-Jiménez, Alfonsina;Escobar-Hernández, Fernando;Rös, Matthias;Kohlmann, Bert		Zenodo	biologists	Arriaga-Jiménez, Alfonsina;Escobar-Hernández, Fernando;Rös, Matthias;Kohlmann, Bert			
03AC87DDFF81FF86FC3CC447FD04D76F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15701411/files/figure.png	https://doi.org/10.5281/zenodo.15701411	Figs 7–12. Species in the Onthophagus anthracinus species complex. 7, Pygidium of male Onthophagus alluvius; 8, pronotum of a male Onthophagus alluvius; 9, pygidium of male Onthophagus mextexus; 10, pronotum of a male of Onthophagus mextexus; 11, metasternal midline of a male Onthophagus alluvius; 12, metasternal midline of a male Onthophagus durangoensis.	Figs 7–12. Species in the Onthophagus anthracinus species complex. 7, Pygidium of male Onthophagus alluvius; 8, pronotum of a male Onthophagus alluvius; 9, pygidium of male Onthophagus mextexus; 10, pronotum of a male of Onthophagus mextexus; 11, metasternal midline of a male Onthophagus alluvius; 12, metasternal midline of a male Onthophagus durangoensis.	2019-12-16	Arriaga-Jiménez, Alfonsina;Escobar-Hernández, Fernando;Rös, Matthias;Kohlmann, Bert		Zenodo	biologists	Arriaga-Jiménez, Alfonsina;Escobar-Hernández, Fernando;Rös, Matthias;Kohlmann, Bert			
03AC87DDFF81FF86FC3CC447FD04D76F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15701415/files/figure.png	https://doi.org/10.5281/zenodo.15701415	Figs 16–20. Aedeagi of the Onthophagus anthracinus species complex. 16, Onthopagus alluvius; 17, O. anthracinus; 18, O. durangoensis; 19, O. etlaensis; 20, O. mextexus.	Figs 16–20. Aedeagi of the Onthophagus anthracinus species complex. 16, Onthopagus alluvius; 17, O. anthracinus; 18, O. durangoensis; 19, O. etlaensis; 20, O. mextexus.	2019-12-16	Arriaga-Jiménez, Alfonsina;Escobar-Hernández, Fernando;Rös, Matthias;Kohlmann, Bert		Zenodo	biologists	Arriaga-Jiménez, Alfonsina;Escobar-Hernández, Fernando;Rös, Matthias;Kohlmann, Bert			
03AC87DDFF81FF87FC3CC271FDCBD008.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15701403/files/figure.png	https://doi.org/10.5281/zenodo.15701403	Figs. 1–4. Onthophagus etlaensis and O. anthracinus habitus. 1, Onthophagus etlaensis dorsal view; 2, O. etlaensis anterolateral view; 3, O. anthracinus dorsal view; 4, O. anthracinus anterolateral view.	Figs. 1–4. Onthophagus etlaensis and O. anthracinus habitus. 1, Onthophagus etlaensis dorsal view; 2, O. etlaensis anterolateral view; 3, O. anthracinus dorsal view; 4, O. anthracinus anterolateral view.	2019-12-16	Arriaga-Jiménez, Alfonsina;Escobar-Hernández, Fernando;Rös, Matthias;Kohlmann, Bert		Zenodo	biologists	Arriaga-Jiménez, Alfonsina;Escobar-Hernández, Fernando;Rös, Matthias;Kohlmann, Bert			
03AC87DDFF81FF87FC3CC271FDCBD008.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15701413/files/figure.png	https://doi.org/10.5281/zenodo.15701413	Figs 13–15. Species in the Onthophagus anthracinus species complex. 13, Male pronotum of Onthophagus etlaensis; 14, male pronotum of Onthophagus anthracinus; 15, dorsal view of a male Onthophagus durangoensis.	Figs 13–15. Species in the Onthophagus anthracinus species complex. 13, Male pronotum of Onthophagus etlaensis; 14, male pronotum of Onthophagus anthracinus; 15, dorsal view of a male Onthophagus durangoensis.	2019-12-16	Arriaga-Jiménez, Alfonsina;Escobar-Hernández, Fernando;Rös, Matthias;Kohlmann, Bert		Zenodo	biologists	Arriaga-Jiménez, Alfonsina;Escobar-Hernández, Fernando;Rös, Matthias;Kohlmann, Bert			
03AC87DDFF81FF87FC3CC271FDCBD008.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15701415/files/figure.png	https://doi.org/10.5281/zenodo.15701415	Figs 16–20. Aedeagi of the Onthophagus anthracinus species complex. 16, Onthopagus alluvius; 17, O. anthracinus; 18, O. durangoensis; 19, O. etlaensis; 20, O. mextexus.	Figs 16–20. Aedeagi of the Onthophagus anthracinus species complex. 16, Onthopagus alluvius; 17, O. anthracinus; 18, O. durangoensis; 19, O. etlaensis; 20, O. mextexus.	2019-12-16	Arriaga-Jiménez, Alfonsina;Escobar-Hernández, Fernando;Rös, Matthias;Kohlmann, Bert		Zenodo	biologists	Arriaga-Jiménez, Alfonsina;Escobar-Hernández, Fernando;Rös, Matthias;Kohlmann, Bert			
03AC87DDFF81FF87FC3CC271FDCBD008.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15701417/files/figure.png	https://doi.org/10.5281/zenodo.15701417	Figs 21–22. Lamella copulatrix and accesory lamellae. 21, Onthophagus anthracinus; 22, O. etlaensis.	Figs 21–22. Lamella copulatrix and accesory lamellae. 21, Onthophagus anthracinus; 22, O. etlaensis.	2019-12-16	Arriaga-Jiménez, Alfonsina;Escobar-Hernández, Fernando;Rös, Matthias;Kohlmann, Bert		Zenodo	biologists	Arriaga-Jiménez, Alfonsina;Escobar-Hernández, Fernando;Rös, Matthias;Kohlmann, Bert			
03AC87DDFF80FF80FCDDC597FDB1D277.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15701411/files/figure.png	https://doi.org/10.5281/zenodo.15701411	Figs 7–12. Species in the Onthophagus anthracinus species complex. 7, Pygidium of male Onthophagus alluvius; 8, pronotum of a male Onthophagus alluvius; 9, pygidium of male Onthophagus mextexus; 10, pronotum of a male of Onthophagus mextexus; 11, metasternal midline of a male Onthophagus alluvius; 12, metasternal midline of a male Onthophagus durangoensis.	Figs 7–12. Species in the Onthophagus anthracinus species complex. 7, Pygidium of male Onthophagus alluvius; 8, pronotum of a male Onthophagus alluvius; 9, pygidium of male Onthophagus mextexus; 10, pronotum of a male of Onthophagus mextexus; 11, metasternal midline of a male Onthophagus alluvius; 12, metasternal midline of a male Onthophagus durangoensis.	2019-12-16	Arriaga-Jiménez, Alfonsina;Escobar-Hernández, Fernando;Rös, Matthias;Kohlmann, Bert		Zenodo	biologists	Arriaga-Jiménez, Alfonsina;Escobar-Hernández, Fernando;Rös, Matthias;Kohlmann, Bert			
03AC87DDFF80FF80FCDDC597FDB1D277.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15701413/files/figure.png	https://doi.org/10.5281/zenodo.15701413	Figs 13–15. Species in the Onthophagus anthracinus species complex. 13, Male pronotum of Onthophagus etlaensis; 14, male pronotum of Onthophagus anthracinus; 15, dorsal view of a male Onthophagus durangoensis.	Figs 13–15. Species in the Onthophagus anthracinus species complex. 13, Male pronotum of Onthophagus etlaensis; 14, male pronotum of Onthophagus anthracinus; 15, dorsal view of a male Onthophagus durangoensis.	2019-12-16	Arriaga-Jiménez, Alfonsina;Escobar-Hernández, Fernando;Rös, Matthias;Kohlmann, Bert		Zenodo	biologists	Arriaga-Jiménez, Alfonsina;Escobar-Hernández, Fernando;Rös, Matthias;Kohlmann, Bert			
03AC87DDFF80FF80FCDDC597FDB1D277.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15701415/files/figure.png	https://doi.org/10.5281/zenodo.15701415	Figs 16–20. Aedeagi of the Onthophagus anthracinus species complex. 16, Onthopagus alluvius; 17, O. anthracinus; 18, O. durangoensis; 19, O. etlaensis; 20, O. mextexus.	Figs 16–20. Aedeagi of the Onthophagus anthracinus species complex. 16, Onthopagus alluvius; 17, O. anthracinus; 18, O. durangoensis; 19, O. etlaensis; 20, O. mextexus.	2019-12-16	Arriaga-Jiménez, Alfonsina;Escobar-Hernández, Fernando;Rös, Matthias;Kohlmann, Bert		Zenodo	biologists	Arriaga-Jiménez, Alfonsina;Escobar-Hernández, Fernando;Rös, Matthias;Kohlmann, Bert			
03AC87DDFF87FF81FCB0C2C5FB50D180.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15701411/files/figure.png	https://doi.org/10.5281/zenodo.15701411	Figs 7–12. Species in the Onthophagus anthracinus species complex. 7, Pygidium of male Onthophagus alluvius; 8, pronotum of a male Onthophagus alluvius; 9, pygidium of male Onthophagus mextexus; 10, pronotum of a male of Onthophagus mextexus; 11, metasternal midline of a male Onthophagus alluvius; 12, metasternal midline of a male Onthophagus durangoensis.	Figs 7–12. Species in the Onthophagus anthracinus species complex. 7, Pygidium of male Onthophagus alluvius; 8, pronotum of a male Onthophagus alluvius; 9, pygidium of male Onthophagus mextexus; 10, pronotum of a male of Onthophagus mextexus; 11, metasternal midline of a male Onthophagus alluvius; 12, metasternal midline of a male Onthophagus durangoensis.	2019-12-16	Arriaga-Jiménez, Alfonsina;Escobar-Hernández, Fernando;Rös, Matthias;Kohlmann, Bert		Zenodo	biologists	Arriaga-Jiménez, Alfonsina;Escobar-Hernández, Fernando;Rös, Matthias;Kohlmann, Bert			
03AC87DDFF87FF81FCB0C2C5FB50D180.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15701413/files/figure.png	https://doi.org/10.5281/zenodo.15701413	Figs 13–15. Species in the Onthophagus anthracinus species complex. 13, Male pronotum of Onthophagus etlaensis; 14, male pronotum of Onthophagus anthracinus; 15, dorsal view of a male Onthophagus durangoensis.	Figs 13–15. Species in the Onthophagus anthracinus species complex. 13, Male pronotum of Onthophagus etlaensis; 14, male pronotum of Onthophagus anthracinus; 15, dorsal view of a male Onthophagus durangoensis.	2019-12-16	Arriaga-Jiménez, Alfonsina;Escobar-Hernández, Fernando;Rös, Matthias;Kohlmann, Bert		Zenodo	biologists	Arriaga-Jiménez, Alfonsina;Escobar-Hernández, Fernando;Rös, Matthias;Kohlmann, Bert			
03AC87DDFF87FF81FCB0C2C5FB50D180.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15701415/files/figure.png	https://doi.org/10.5281/zenodo.15701415	Figs 16–20. Aedeagi of the Onthophagus anthracinus species complex. 16, Onthopagus alluvius; 17, O. anthracinus; 18, O. durangoensis; 19, O. etlaensis; 20, O. mextexus.	Figs 16–20. Aedeagi of the Onthophagus anthracinus species complex. 16, Onthopagus alluvius; 17, O. anthracinus; 18, O. durangoensis; 19, O. etlaensis; 20, O. mextexus.	2019-12-16	Arriaga-Jiménez, Alfonsina;Escobar-Hernández, Fernando;Rös, Matthias;Kohlmann, Bert		Zenodo	biologists	Arriaga-Jiménez, Alfonsina;Escobar-Hernández, Fernando;Rös, Matthias;Kohlmann, Bert			
03AC87DDFF87FF81FCB0C2C5FB50D180.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15701417/files/figure.png	https://doi.org/10.5281/zenodo.15701417	Figs 21–22. Lamella copulatrix and accesory lamellae. 21, Onthophagus anthracinus; 22, O. etlaensis.	Figs 21–22. Lamella copulatrix and accesory lamellae. 21, Onthophagus anthracinus; 22, O. etlaensis.	2019-12-16	Arriaga-Jiménez, Alfonsina;Escobar-Hernández, Fernando;Rös, Matthias;Kohlmann, Bert		Zenodo	biologists	Arriaga-Jiménez, Alfonsina;Escobar-Hernández, Fernando;Rös, Matthias;Kohlmann, Bert			
