taxonID	type	description	language	source
03AD991AFF9F5741FFDED3B5CE82D082.taxon	distribution	Distribution in Yemen. Lahj and 12 km NW Manakhah (van Loon & Soliman, 2023) Distribution in the Arabian Peninsula. Yemen. Saudi Arabia (van Loon & Soliman, 2023).	en	Rosa, Paolo (2024): Contributions to the cuckoo wasp fauna (Hymenoptera: Chrysididae) of Yemen with description of five new species and updated checklist. Journal of Insect Biodiversity and Systematics 10 (4): 995-1031, DOI: 10.61186/jibs.10.4.995, URL: https://doi.org/10.61186/jibs.10.4.995
03AD991AFF9F5741FFDED184CBD7D57F.taxon	distribution	Distribution in the Arabian Peninsula. Saudi Arabia (Farasan) (Strumia & Dawah, 2019 as Hedychridium flavipes (Eversmann, 1858 )). Extralimital distribution. Eastern Mediterranean, Middle East and Western Asia: Cyprus, Iran, Türkiye, Central Asia, Egypt (Linsenmaier, 1999; Rosa, 2017).	en	Rosa, Paolo (2024): Contributions to the cuckoo wasp fauna (Hymenoptera: Chrysididae) of Yemen with description of five new species and updated checklist. Journal of Insect Biodiversity and Systematics 10 (4): 995-1031, DOI: 10.61186/jibs.10.4.995, URL: https://doi.org/10.61186/jibs.10.4.995
03AD991AFF9F5741FFDED184CBD7D57F.taxon	discussion	Remarks. Strumia & Dawah (2019) included Yemen and the United Arab Emirates in the distribution of this subspecies, citing localities referenced in Strumia et al. (2016). However, Yemen and United Arab Emirates are not mentioned in the latter article. Therefore, a confirmation of the occurrence of Colpopyga in Yemen is still needed.	en	Rosa, Paolo (2024): Contributions to the cuckoo wasp fauna (Hymenoptera: Chrysididae) of Yemen with description of five new species and updated checklist. Journal of Insect Biodiversity and Systematics 10 (4): 995-1031, DOI: 10.61186/jibs.10.4.995, URL: https://doi.org/10.61186/jibs.10.4.995
03AD991AFF9F5741FFDED676CB06D746.taxon	materials_examined	Material examined. 1 ♀, Yemen: Lahj, III. - V. 2002, Malaise trap, leg. A. van Harten & A. Sallum (RMNH).	en	Rosa, Paolo (2024): Contributions to the cuckoo wasp fauna (Hymenoptera: Chrysididae) of Yemen with description of five new species and updated checklist. Journal of Insect Biodiversity and Systematics 10 (4): 995-1031, DOI: 10.61186/jibs.10.4.995, URL: https://doi.org/10.61186/jibs.10.4.995
03AD991AFF9F5741FFDED676CB06D746.taxon	distribution	Distribution in Yemen. Lahj (new record). Distribution in the Arabian Peninsula. Yemen (new record). Extralimital distribution. Cameroon, Kenya, South Africa, Tanzania (Tanzania, Zanzibar), Uganda (Madl & Rosa, 2012).	en	Rosa, Paolo (2024): Contributions to the cuckoo wasp fauna (Hymenoptera: Chrysididae) of Yemen with description of five new species and updated checklist. Journal of Insect Biodiversity and Systematics 10 (4): 995-1031, DOI: 10.61186/jibs.10.4.995, URL: https://doi.org/10.61186/jibs.10.4.995
03AD991AFF9F5741FFDED520CF28D6F9.taxon	distribution	Distribution in Yemen. Yemen (with no specific locality) (Strumia, 2014). Distribution in the Arabian Peninsula. Saudi Arabia (Linsenmaier, 1994 as Omalus (Elampus) albipennis; Strumia & Dawah, 2010, 2012, 2019), Oman (Strumia & Dawah, 2019), United Arab Emirates (Linsenmaier, 1994 as Omalus (Elampus) albipennis; Howarth & Gillett, 2008 as Omalus albipennis; Strumia, 2008, 2014). Extralimital distribution. South-East Europe, Middle East, Türkiye, Southern Russia, Central Asia to Mongolia (Kimsey & Bohart, 1991; Linsenmaier, 1994, Strumia & Dawah, 2019; Rosa et al., 2020).	en	Rosa, Paolo (2024): Contributions to the cuckoo wasp fauna (Hymenoptera: Chrysididae) of Yemen with description of five new species and updated checklist. Journal of Insect Biodiversity and Systematics 10 (4): 995-1031, DOI: 10.61186/jibs.10.4.995, URL: https://doi.org/10.61186/jibs.10.4.995
03AD991AFF985744FF2FD223C94FD3C2.taxon	materials_examined	Material examined. Holotype ♀, Yemen: Lahj, VII-IX. 2001, Malaise trap, leg. A. van Harten & A. Sallum (RMNH). Paratypes: 2 ♂♂, Yemen: Ibb province, 20 km S Ta’izz, 1200 m, 24. X. 2005, leg. J. Halada (MHC, PRC); 1 ♂, Saudi Arabia: Jizan province, Farasan Island, Sajid Island: 16 ° 51 ' 52 " N 41 ° 56 ' 04 " E, 16. IX. 2023, leg. J. Halada (MHC); 1 ♀, Saudi Arabia, Jizan province: N of Ayban, Smad, 17 ° 20 ' 42 " N 43 ° 02 ' 20 " E, 1000 m, 26. II. 2024, leg. B. Halada (MHC).	en	Rosa, Paolo (2024): Contributions to the cuckoo wasp fauna (Hymenoptera: Chrysididae) of Yemen with description of five new species and updated checklist. Journal of Insect Biodiversity and Systematics 10 (4): 995-1031, DOI: 10.61186/jibs.10.4.995, URL: https://doi.org/10.61186/jibs.10.4.995
03AD991AFF985744FF2FD223C94FD3C2.taxon	diagnosis	Diagnosis. Hedychridium eudaimon sp. nov. is closely related to only one species in the Western Palaearctic, Hedychridium chakouri du Buysson, 1907 (Figs 2 E, 2 F). Both species belong to the subgenus incorrectly named Acrotoma Mocsáry, 1902 (nec Boettger, 1881) by Linsenmaier (1959, 1999). These two species can be readily separated from other northern African and Arabian Hedychridium by the median notch on the apical margin of the third tergum. Compared to the other Afrotropical species in this group, they are both small ranging 3.0 to 4.5 mm. H. eudaimon sp. nov. can be differentiated from H. chakouri by the large, sparse punctures on the mesosoma (Fig. 2 B) and on the second metasomal tergum (Figs 2 D, 3 B) in contrast to H. chakouri (Figs 2 E, 2 F) which has smaller, denser punctures. The colour pattern is also distinct: H. eudaimon sp. nov. is bluish medially, consistent with the colour pattern of several Afrotropical species (Fig. 3 A), while H. chakouri has a colour pattern more similar to the widespread Palaearctic H. monochroum du Buysson, 1888, with darker median area of the mesoscutum and a darker disk on the second metasomal tergum (Fig. 2 E). H. eudaimon sp. nov. can be further distinguished from other Afrotropical species traditionally included in “ Acrotoma ” by the following characteristics: H. arnoldii (Edney, 1940) and H. braunsii (Mocsáry, 1902) have the apical margin of the third tergum with two distinct teeth and a the carinate margin of the second metasomal tergum; H. discrepans (Edney, 1940) has a similar apical margin, but is a larger species (6.5 mm) with a different colour pattern, being entirely red dorsally, with the face, sides of the mesosoma and propodeal angles greenish, while the subspecies H. discrepans candida (Edney, 1940) is entirely flame red; H. dybowskii has two small apical teeth on the apical margin and dense punctation, with very small punctures similar to H. chakouri (Fig. 2 F); H. heymonsi Bischoff, 1910 is similar to H. dybowskii but has a thin hyaline rim along the apical margin of the third tergum and a distinct angle between the apical tooth and the tergal base.	en	Rosa, Paolo (2024): Contributions to the cuckoo wasp fauna (Hymenoptera: Chrysididae) of Yemen with description of five new species and updated checklist. Journal of Insect Biodiversity and Systematics 10 (4): 995-1031, DOI: 10.61186/jibs.10.4.995, URL: https://doi.org/10.61186/jibs.10.4.995
03AD991AFF985744FF2FD223C94FD3C2.taxon	description	Description. Holotype ♀. (Fig. 2 A – D). body length 4.4 mm; fore wing length 2.8 mm. Head. Vertex and frons finely and densely punctate, with small (0.3 MOD) subcontiguous punctures; scapal basin with smaller, dense punctures, each puncture bearing a whitish seta, altogether covering the side of face; scapal basin transversally slightly wrinkled medially; ocellar triangle isosceles; OOL = 2.8 × MOD; POL = 2.4 × MOD; relative length of P: F 1: F 2: F 3 = 1.0: 1.2: 0.9: 0.8. Mesosoma. Pronotal antero-median line shallow; punctation double with deep, small punctures (0.2 – 0.3 × MOD) intermixed with smaller dots on interspaces; median area of mesoscutum with larger punctures (up to 0.5 × MOD), without interspaces, becoming smaller and more spaced (up to 1 puncture diameter) at sides; mesoscutum largely polished postero-medially; lateral areas of mesoscutum with deep, large punctures, spaced and with small deep punctures on interspaces; notauli as fine lines, barely visible among punctures, parapsidal furrow deep and elongate; mesoscutellum with large, sparse punctures, separated up to 1.5 puncture diameter, and polished interspaces, punctures smaller laterally and posteriorly (Fig. 2 B); metanotum with foveate-reticulate punctures; mesopleuron with large, round punctures, separate by polished interspaces with small punctures. Propodeal projection like acute triangle, pointing downwards; fore femur slightly carinate ventrally, hind femur only partially metallic; hind tarsus unmodified, with second and third tarsomeres subequal in length. Tarsal claws with sharp submedian tooth. Metasoma. Metasomal terga with deep punctures, denser apico-medially on second tergum becoming very sparse on the second half of the segment, with polished interspaces up to 2 – 3 puncture diameters (Fig. 2 D); third tergum with spaced puncture becoming smaller toward apex and slightly bulging before the apical margin; apical margin with narrow hyaline rim and medially notched (Figs 2 C, 3 B, 3 C). Colour. Body green to blue; dark blue to purplish on ocellar area, median area of mesoscutum, anteromedially on second and third tergum; sterna entirely metallic green (Fig. 3 C). Pedicel and flagellomeres brown; tegula metallic green; legs metallic green; tarsi brownish to light brown. Vestiture. Erect, whitish short setae on head (less than 1 MOD); longer ventrally and on coxae; pronotum and mesoscutum with short and appressed, erect on scutellum and metanotum; metasoma with short and adpressed setae. Male. Similar to female (Fig. 3 A – D).	en	Rosa, Paolo (2024): Contributions to the cuckoo wasp fauna (Hymenoptera: Chrysididae) of Yemen with description of five new species and updated checklist. Journal of Insect Biodiversity and Systematics 10 (4): 995-1031, DOI: 10.61186/jibs.10.4.995, URL: https://doi.org/10.61186/jibs.10.4.995
03AD991AFF985744FF2FD223C94FD3C2.taxon	etymology	Etymology. The specific epithet is derived from the Old Greek name of Yemen Εὐδαίμων Ἀραβία (Eudaemon Arabia), Latinized into Eudaimon, better known as Arabia Felix, name attributed to Eratosthenes of Cyrene (c. 276 BC – c. 195 / 194 BC), which has the meaning of both fertile and blessed, being this area well irrigated in the Arabian Peninsula.	en	Rosa, Paolo (2024): Contributions to the cuckoo wasp fauna (Hymenoptera: Chrysididae) of Yemen with description of five new species and updated checklist. Journal of Insect Biodiversity and Systematics 10 (4): 995-1031, DOI: 10.61186/jibs.10.4.995, URL: https://doi.org/10.61186/jibs.10.4.995
03AD991AFF985744FF2FD223C94FD3C2.taxon	distribution	Distribution. Saudi Arabia (Jizan), Yemen (Lahj, Ibb).	en	Rosa, Paolo (2024): Contributions to the cuckoo wasp fauna (Hymenoptera: Chrysididae) of Yemen with description of five new species and updated checklist. Journal of Insect Biodiversity and Systematics 10 (4): 995-1031, DOI: 10.61186/jibs.10.4.995, URL: https://doi.org/10.61186/jibs.10.4.995
03AD991AFF9A5744FF2FD1A3C8E9D556.taxon	distribution	Distribution in Yemen. Yemen (with no specific locality) (Strumia, 2008, 2014). Distribution in the Arabian Peninsula. Oman, United Arab Emirates (Strumia, 2008, 2014), Yemen (Strumia, 2008, 2014). Extralimital distribution. Palestine (Linsenmaier, 1999); Egypt, Iran, Pakistan (Kimsey & Bohart, 1991; Strumia, 2008, 2014; Strumia et al., 2016).	en	Rosa, Paolo (2024): Contributions to the cuckoo wasp fauna (Hymenoptera: Chrysididae) of Yemen with description of five new species and updated checklist. Journal of Insect Biodiversity and Systematics 10 (4): 995-1031, DOI: 10.61186/jibs.10.4.995, URL: https://doi.org/10.61186/jibs.10.4.995
03AD991AFF9B5745FFDED3B5CBCDD353.taxon	materials_examined	Material examined. 1 ♀, Yemen: N Yemen Hadda 30. V. 1967 leg. Mühle / Hedychridium monochroum Buyss. det. Linsenmaier 1989 / NML _ ENT GBIF _ Chr 0009715 (NMLU).	en	Rosa, Paolo (2024): Contributions to the cuckoo wasp fauna (Hymenoptera: Chrysididae) of Yemen with description of five new species and updated checklist. Journal of Insect Biodiversity and Systematics 10 (4): 995-1031, DOI: 10.61186/jibs.10.4.995, URL: https://doi.org/10.61186/jibs.10.4.995
03AD991AFF9B5745FFDED3B5CBCDD353.taxon	distribution	Distribution in Yemen. Hadda (Linsenmaier, 1994). Distribution in the Arabian Peninsula. United Arab Emirates (Strumia, 2014). Extralimital distribution. widespread in the Palaearctic Region; also known from the Oriental Region (Kimsey & Bohart, 1991).	en	Rosa, Paolo (2024): Contributions to the cuckoo wasp fauna (Hymenoptera: Chrysididae) of Yemen with description of five new species and updated checklist. Journal of Insect Biodiversity and Systematics 10 (4): 995-1031, DOI: 10.61186/jibs.10.4.995, URL: https://doi.org/10.61186/jibs.10.4.995
03AD991AFF9B5745FFDED134CF02D282.taxon	materials_examined	Material examined. 1 ♀, Yemen: Al-Lahima, 16. X. - 31. XI. 2000, Malaise trap, leg. A. van Harten (RMNH); 1 ♀, Al-Lahima, 9. IV. - 5. VI. 2001, Malaise trap, leg. A. van Harten (RMNH); 1 ♀, Yemen: 12 km NW of Manakhah, 21. VIII. - 29. X. 2002, Malaise trap, leg. A. van Harten (RMNH).	en	Rosa, Paolo (2024): Contributions to the cuckoo wasp fauna (Hymenoptera: Chrysididae) of Yemen with description of five new species and updated checklist. Journal of Insect Biodiversity and Systematics 10 (4): 995-1031, DOI: 10.61186/jibs.10.4.995, URL: https://doi.org/10.61186/jibs.10.4.995
03AD991AFF9B5745FFDED134CF02D282.taxon	distribution	Distribution in Yemen. Al-Lahima, Manakhah (new record). Distribution in the Arabian Peninsula. Yemen (new record); United Arab Emirates (Strumia, 2014).	en	Rosa, Paolo (2024): Contributions to the cuckoo wasp fauna (Hymenoptera: Chrysididae) of Yemen with description of five new species and updated checklist. Journal of Insect Biodiversity and Systematics 10 (4): 995-1031, DOI: 10.61186/jibs.10.4.995, URL: https://doi.org/10.61186/jibs.10.4.995
03AD991AFF9B5745FFDED7A1CBFBD4E0.taxon	distribution	Distribution in Yemen. Yemen (with no specific locality) (Strumia, 2008). Distribution in the Arabian Peninsula. Arabia (with no specific locality, Linsenmaier, 1999), Oman (Strumia, 2014), Saudi Arabia, United Arab Emirates (Linsenmaier, 1994; Howarth & Gillett, 2008 as Hedychridium alfierii).	en	Rosa, Paolo (2024): Contributions to the cuckoo wasp fauna (Hymenoptera: Chrysididae) of Yemen with description of five new species and updated checklist. Journal of Insect Biodiversity and Systematics 10 (4): 995-1031, DOI: 10.61186/jibs.10.4.995, URL: https://doi.org/10.61186/jibs.10.4.995
03AD991AFF9B5745FFDED5FACF3BD6D8.taxon	materials_examined	Material examined. 2 ♀♀, Yemen: Al-Lahima, 16. X. - 31. XII. 2000, Malaise trap, leg. A. v. Harten & A. M. Hager (RMNH); 3 ♀♀, Al-Lahima, 1. I. - 9. IV. 2001, Malaise trap, leg. A. van Harten & A. M. Hajer (RMNH); 4 ♀♀, Al-Lahima, 9. IV. - 5. VI. 2001, Malaise trap, leg. A. van Harten (RMNH).	en	Rosa, Paolo (2024): Contributions to the cuckoo wasp fauna (Hymenoptera: Chrysididae) of Yemen with description of five new species and updated checklist. Journal of Insect Biodiversity and Systematics 10 (4): 995-1031, DOI: 10.61186/jibs.10.4.995, URL: https://doi.org/10.61186/jibs.10.4.995
03AD991AFF9B5745FFDED5FACF3BD6D8.taxon	distribution	Distribution in Yemen. Al-Lahima (new record). Distribution in the Arabian Peninsula. Yemen (new record). Saudi Arabia (Linsenmaier, 1994).	en	Rosa, Paolo (2024): Contributions to the cuckoo wasp fauna (Hymenoptera: Chrysididae) of Yemen with description of five new species and updated checklist. Journal of Insect Biodiversity and Systematics 10 (4): 995-1031, DOI: 10.61186/jibs.10.4.995, URL: https://doi.org/10.61186/jibs.10.4.995
03AD991AFF945748FF2ED602C845D721.taxon	materials_examined	Material examined. Holotype ♀, Yemen: Al-Lahima, 16. X. - 31. XII. 2000, Malaise trap, leg. A. v. Harten & A. M. Hager (RMNH); Paratypes 2 ♀♀, same data and collectors of the holotype (RMNH).	en	Rosa, Paolo (2024): Contributions to the cuckoo wasp fauna (Hymenoptera: Chrysididae) of Yemen with description of five new species and updated checklist. Journal of Insect Biodiversity and Systematics 10 (4): 995-1031, DOI: 10.61186/jibs.10.4.995, URL: https://doi.org/10.61186/jibs.10.4.995
03AD991AFF945748FF2ED602C845D721.taxon	diagnosis	Diagnosis. Hedychrum harteni sp. nov. may initially resemble a species related to H. rutilans Dahlbom, 1854 due to its elongate pronotum (longer than the combined mesonotum and metanotum) and the coloration of the mesosoma, with a light green pronotum and mesonotum contrasting with a blue metanotum and propodeum. However, the shape of the apical margin of the female third sternum (Fig. 4 F) is typical of species closely related to H. alfierii Trautmann, 1926. Hedychrum harteni sp. nov. shares several characteristics with H. alfierii (Fig. 4), which ranges from Egypt to Palestine and Arabia. These shared features include its small size, 5 mm, the apical margin of the female third sternum, which is apically digitate on each hemisternum and medially sinuate along the midline, and the discoloured legs and sterna (Fig. 4 A). However, H. harteni sp. nov. can be distinguished from H. alfierii by the following traits: an elongate pronotum (slightly longer than the scutellum in H. alfierii); dense, small punctures on the pronotum without polished interspaces (Fig. 5 C) (round and separated punctures on mesosoma, with polished interspaces in H. alfierii, Fig. 4 C); first tergum densely punctate basally (Fig. 5 D) (largely polished in H. alfierii, Fig. 4 E); metallic blue outer tibiae (non-metallic yellow with a weak rose hue in H. alfierii, Fig. 4 A); and contrasting green and blue coloration on the mesosoma (head and mesosoma unicolored green in H. alfierii, Figs 4 A, 5 C).	en	Rosa, Paolo (2024): Contributions to the cuckoo wasp fauna (Hymenoptera: Chrysididae) of Yemen with description of five new species and updated checklist. Journal of Insect Biodiversity and Systematics 10 (4): 995-1031, DOI: 10.61186/jibs.10.4.995, URL: https://doi.org/10.61186/jibs.10.4.995
03AD991AFF945748FF2ED602C845D721.taxon	description	Description. Holotype ♀ (Fig. 5 A – F). Body length 5.0 mm. Head. Vertex, frons and temples densely punctate, with small (0.3 – 0.4 × MOD) subcontiguous punctures; without polish spaces; large impunctate area (1 × MOD) laterad posterior ocelli; scapal basin deep, with sharp wrinkles; clypeus medially impunctate to slightly wrinkled, laterally with dense, rugose sculpture and undefined punctures; subantennal distance short (0.5 × MOD); clypeal margin with median dark brown thickened apex, laterally with expanded hyaline rim; OOL = 2.4 × MOD; POL = 2.2 × MOD; MW = 0.7 MOD; relative length of P: F 1: F 2: F 3 = 1.0: 1.2: 0.9: 0.8. Mesosoma. Pronotum elongate, as long as scutellum and metanotum together (Fig. 5 A); pronotal anteromedian line shallow; punctation even, deep, subcontiguous, with small punctures (0.3 – 0.4 × MOD) (Fig. 5 B); median area of mesoscutum with similar punctures but with larger, polished interspaces, becoming narrower on lateral sides of mesoscutum; notauli as fine lines, barely visible between punctures, parapsidal furrow deeper, more visible than notauli and elongate; mesoscutellum with smaller and sparser puncture, with polished interspaces; punctures on scutellum larger and subcontinuous laterally and posteriorly; metanotum with large, foveate-reticulate punctures, as large as 1 × MOD; mesopleuron with large, polygonal punctures, subcontinuous without polished interspaces. Propodeal projection as acute triangle, pointing outwards; tarsal claws with pointed submedian tooth. Metasoma. Metasomal first tergum with relatively small punctures dorsally, becoming larger and deeper at sides; punctures smaller and dense basally on second tergum, becoming very sparse on second half of segment, with polished interspaces up to 2 – 3 puncture diameter (Figs 5 D, 5 E); on third tergum with spaced and relatively shallow puncture; third tergum regularly slightly bulging before the apical margin; margin with narrow hyaline rim and two lateral evident angles. Second sternum densely punctate at sides and sparsely punctate medially; third sternum densely micropunctate, with scattered large punctures; apical margin of the third sternum digitate on each hemisternum, sinuate along the midline before the apical teeth (Fig. 5 F). Colour. Head and mesosoma blue, with a light green hue on pronotum and mesonotum contrasting with blue metanotum and propodeum; metasomal sterna brown without metallic reflections; scape iridescent, pedicel and flagellomeres brown; tegula brown; legs brown on ventral side, with metallic colour on outer side; tarsi brownish to light brown; wings brownish. Vestiture. Head and mesosoma with short (0.5 × MOD), appressed and whitish setae, longer on propodeum laterally, 1.0 × MOD. Scattered longer setae on mid and hind femora (1.0 - 3.0 × MOD) (Fig. 5 B) and scattered, erect setae on mid and hind tibiae; thick and longer setae on first and second terga laterally and on the third posterolaterally (1.0 × MOD). Male. Unknown.	en	Rosa, Paolo (2024): Contributions to the cuckoo wasp fauna (Hymenoptera: Chrysididae) of Yemen with description of five new species and updated checklist. Journal of Insect Biodiversity and Systematics 10 (4): 995-1031, DOI: 10.61186/jibs.10.4.995, URL: https://doi.org/10.61186/jibs.10.4.995
03AD991AFF945748FF2ED602C845D721.taxon	etymology	Etymology. The specific epithet harteni is dedicated to Antonius van Harten, who has made significant contributions to the collection of the material examined in this study and to the broader knowledge of the entomological fauna of the Arabian Peninsula over the past three decades.	en	Rosa, Paolo (2024): Contributions to the cuckoo wasp fauna (Hymenoptera: Chrysididae) of Yemen with description of five new species and updated checklist. Journal of Insect Biodiversity and Systematics 10 (4): 995-1031, DOI: 10.61186/jibs.10.4.995, URL: https://doi.org/10.61186/jibs.10.4.995
03AD991AFF945748FF2ED602C845D721.taxon	discussion	Remarks. The previous identifications of Hedychrum alfierii in Palestine and Arabia need to be reassessed due to existing uncertainties regarding this species (types examined, Fig. 4), for example Kimsey & Bohart (1991) classified this species in the genus Hedychridium Abeille de Perrin, 1878.	en	Rosa, Paolo (2024): Contributions to the cuckoo wasp fauna (Hymenoptera: Chrysididae) of Yemen with description of five new species and updated checklist. Journal of Insect Biodiversity and Systematics 10 (4): 995-1031, DOI: 10.61186/jibs.10.4.995, URL: https://doi.org/10.61186/jibs.10.4.995
03AD991AFF945748FF2ED602C845D721.taxon	distribution	Distribution. Yemen (Al-Lahima).	en	Rosa, Paolo (2024): Contributions to the cuckoo wasp fauna (Hymenoptera: Chrysididae) of Yemen with description of five new species and updated checklist. Journal of Insect Biodiversity and Systematics 10 (4): 995-1031, DOI: 10.61186/jibs.10.4.995, URL: https://doi.org/10.61186/jibs.10.4.995
03AD991AFF965748FF2ED504CF28D66B.taxon	materials_examined	Material examined. 1 ♀, Yemen: Al-Lahima, 16. X. 2000 - 31. XII. 2000, Malaise trap, leg. A. van Harten & A. M. Hager (RMNH); 1 ♀, Al-Lahima, I. - 9. IV. 2001, Malaise trap, leg. A. van Harten & A. M. Hojer (RMNH); 2 ♂♂, 1 ♀, Al-Lahima, 9. IV. - 5. VI. 2001, Malaise trap, leg. A. van Harten (RMNH).	en	Rosa, Paolo (2024): Contributions to the cuckoo wasp fauna (Hymenoptera: Chrysididae) of Yemen with description of five new species and updated checklist. Journal of Insect Biodiversity and Systematics 10 (4): 995-1031, DOI: 10.61186/jibs.10.4.995, URL: https://doi.org/10.61186/jibs.10.4.995
03AD991AFF965748FF2ED504CF28D66B.taxon	distribution	Distribution in Yemen. Al-Lahima (new record). Distribution in the Arabian Peninsula. Yemen (new record). Saudi Arabia (Linsenmaier, 1994).	en	Rosa, Paolo (2024): Contributions to the cuckoo wasp fauna (Hymenoptera: Chrysididae) of Yemen with description of five new species and updated checklist. Journal of Insect Biodiversity and Systematics 10 (4): 995-1031, DOI: 10.61186/jibs.10.4.995, URL: https://doi.org/10.61186/jibs.10.4.995
03AD991AFF975749FFDED154CB94D2DF.taxon	materials_examined	Material examined. 1 ♂, Yemen: Sana’a, I. 1991, leg. A. van Harten, Holophris coriaceus Dahlbom ♂ det. F. Strumia (RMNH); 1 ♂ and 2 ♀♀, Sana’a, XII. 2002, yellow warer traps, leg. A. van Harten (RMNH); 1 ♂, Al-Lahima, 1. I. - 9. IV. 2001, Malaise trap, leg. A. van Harten (RMNH); 1 ♀, Al-Lahima, 9. IV. - 5. VI. 2001, Malaise trap, leg. A. van Harten (RMNH); 2 ♀♀, 12 km NW of Manakhah, 3. VII. - 21. VIII. 2001, leg. A. van Harten (RMNH); 2 ♀♀, Ar Rujum, 9. IV. 5. VI. 2001, leg. A. van Harten (RMNH).	en	Rosa, Paolo (2024): Contributions to the cuckoo wasp fauna (Hymenoptera: Chrysididae) of Yemen with description of five new species and updated checklist. Journal of Insect Biodiversity and Systematics 10 (4): 995-1031, DOI: 10.61186/jibs.10.4.995, URL: https://doi.org/10.61186/jibs.10.4.995
03AD991AFF975749FFDED154CB94D2DF.taxon	distribution	Distribution in Yemen. Sana’a, Al-Lahima, Ar Rujum, Manakhah (new record). Distribution in the Arabian Peninsula. Yemen (new record). Extralimital distribution. Guinea Bissau, Kenya, Lesotho, Rwanda, Senegal, South Africa, Tanzania, Uganda (Madl & Rosa, 2012).	en	Rosa, Paolo (2024): Contributions to the cuckoo wasp fauna (Hymenoptera: Chrysididae) of Yemen with description of five new species and updated checklist. Journal of Insect Biodiversity and Systematics 10 (4): 995-1031, DOI: 10.61186/jibs.10.4.995, URL: https://doi.org/10.61186/jibs.10.4.995
03AD991AFF975749FFDED0A9C8CED42E.taxon	materials_examined	Material examined. 2 ♀♀, Yemen: Al-Lahima, 9. IV. - 5. VI. 2001, Malaise trap, leg. A. van Harten (RMNH); 1 ♀, Al Kowd, 27. X. - 15. XI. 1992, Malaise trap, leg. A. van Harten, Holophris coriaceus Dahlbom ♀ det. F. Strumia (RMNH).	en	Rosa, Paolo (2024): Contributions to the cuckoo wasp fauna (Hymenoptera: Chrysididae) of Yemen with description of five new species and updated checklist. Journal of Insect Biodiversity and Systematics 10 (4): 995-1031, DOI: 10.61186/jibs.10.4.995, URL: https://doi.org/10.61186/jibs.10.4.995
03AD991AFF975749FFDED0A9C8CED42E.taxon	distribution	Distribution in Yemen. Al-Lahima, Al Kowd (Strumia & Dawah, 2012, 2019 as Holophris mochianus). Distribution in the Arabian Peninsula. Yemen (Strumia & Dawah, 2012, 2019). Saudi Arabia (Strumia & Dawah, 2012, 2019 as Holophris mochianus).	en	Rosa, Paolo (2024): Contributions to the cuckoo wasp fauna (Hymenoptera: Chrysididae) of Yemen with description of five new species and updated checklist. Journal of Insect Biodiversity and Systematics 10 (4): 995-1031, DOI: 10.61186/jibs.10.4.995, URL: https://doi.org/10.61186/jibs.10.4.995
03AD991AFF90574EFF2FD224CBF5D2A8.taxon	materials_examined	Material examined. 1 ♂, 1 ♀, Yemen: Usaifira, 1 mile N of Ta'izz ca. 4,500 ft 21. XII. 1937 / B. M. Exp. to S. W. Arabia H. Scott E. B. Britton, B. M. 1938 - 246 / Paratypes Holopyga arabica Lins. Dett. Linsenmaier 1989 / NML _ ENT GBIF _ Chr 0005374, 5375 (NMLU); 1 ♂, 1 ♀, N Yemen Wadi Siham 7. VI. 1987 leg. Mühle / Paratypes Holopyga arabica Lins. Dett. Linsenmaier 1989 / NML _ ENT GBIF _ Chr 0005376, 5377 (NMLU); 1 ♀, 12 km NW of Manakhah, 21. VIII. - 29. X. 2002, Malaise trap, leg. A. van Harten (RMNH); 1 ♀, Al-Lahima, 9. IV. - 5. VI. 2001, Malaise trap, leg. A. van Harten (RMNH); 1 ♂, Lahj, 1. IV. - 17. V. 2000, Malaise trap, leg. A. van Harten & A. Sallam (RMNH); Lahj, III. - V. 2002, Malaise trap, leg. A. van Harten (RMNH).	en	Rosa, Paolo (2024): Contributions to the cuckoo wasp fauna (Hymenoptera: Chrysididae) of Yemen with description of five new species and updated checklist. Journal of Insect Biodiversity and Systematics 10 (4): 995-1031, DOI: 10.61186/jibs.10.4.995, URL: https://doi.org/10.61186/jibs.10.4.995
03AD991AFF90574EFF2FD224CBF5D2A8.taxon	distribution	Distribution in Yemen. Al-Lahima, Manakhah, Wadi Siham; Usaifira; Yemen (Linsenmaier, 1994, present study). Distribution in the Arabian Peninsula. Oman (Linsenmaier, 1994), Saudi Arabia (Linsenmaier, 1994; Strumia & Dawah, 2010, 2012, 2019), United Arab Emirates (Linsenmaier, 1994; Howarth & Gillett, 2008; Strumia, 2014).	en	Rosa, Paolo (2024): Contributions to the cuckoo wasp fauna (Hymenoptera: Chrysididae) of Yemen with description of five new species and updated checklist. Journal of Insect Biodiversity and Systematics 10 (4): 995-1031, DOI: 10.61186/jibs.10.4.995, URL: https://doi.org/10.61186/jibs.10.4.995
03AD991AFF90574EFF2FD74BC8C6D5CE.taxon	distribution	Distribution in Yemen. Yemen (with no specific locality) (Strumia, 2008, 2014; Strumia & Dawah, 2019). Distribution in the Arabian Peninsula. Oman, Saudi Arabia (Strumia & Dawah, 2012), United Arab Emirates (Strumia, 2014; Farhad et al., 2017).	en	Rosa, Paolo (2024): Contributions to the cuckoo wasp fauna (Hymenoptera: Chrysididae) of Yemen with description of five new species and updated checklist. Journal of Insect Biodiversity and Systematics 10 (4): 995-1031, DOI: 10.61186/jibs.10.4.995, URL: https://doi.org/10.61186/jibs.10.4.995
03AD991AFF90574EFF2FD66ACE78D4E8.taxon	distribution	Distribution in Yemen. Yemen (Socotra) (Kohl, 1906). Distribution in the Arabian Peninsula. Yemen (Socotra) (Kohl, 1906).	en	Rosa, Paolo (2024): Contributions to the cuckoo wasp fauna (Hymenoptera: Chrysididae) of Yemen with description of five new species and updated checklist. Journal of Insect Biodiversity and Systematics 10 (4): 995-1031, DOI: 10.61186/jibs.10.4.995, URL: https://doi.org/10.61186/jibs.10.4.995
03AD991AFF90574EFF2FD5F8CB6CD6FB.taxon	materials_examined	Material examined. 1 ♀: Yemen: Al Kowd, 27. X. - 15. XI. 1992, Malaise trap, leg. A. Van Harten, Holopyga subglabrata Linsenmaier? det. F. Strumia (RMNH).	en	Rosa, Paolo (2024): Contributions to the cuckoo wasp fauna (Hymenoptera: Chrysididae) of Yemen with description of five new species and updated checklist. Journal of Insect Biodiversity and Systematics 10 (4): 995-1031, DOI: 10.61186/jibs.10.4.995, URL: https://doi.org/10.61186/jibs.10.4.995
03AD991AFF90574EFF2FD5F8CB6CD6FB.taxon	distribution	Distribution in Yemen. Al Kowd (new record). Distribution in the Arabian Peninsula. Yemen (new record). Oman (Strumia, 2008, 2014), Saudi Arabia (Strumia & Dawah, 2012), United Arab Emirates (Linsenmaier, 1994; Howarth & Gillett, 2008; Strumia, 2008, 2014).	en	Rosa, Paolo (2024): Contributions to the cuckoo wasp fauna (Hymenoptera: Chrysididae) of Yemen with description of five new species and updated checklist. Journal of Insect Biodiversity and Systematics 10 (4): 995-1031, DOI: 10.61186/jibs.10.4.995, URL: https://doi.org/10.61186/jibs.10.4.995
03AD991AFF91574FFFDED3B5CEE6D367.taxon	materials_examined	Material examined. 1 ♀, Yemen: Lahj, XI. - XII. 1999, Malaise trap, leg. A. van Harten & A. Sallum (RMNH); 1 ♂, Lahj, III. 2000, Malaise trap, leg. A. van Harten (RMNH); 1 ♂, Lahj, 1. IV. - 17. V. 2000, Malaise trap, leg. A. van Harten & A. Sallum (RMNH); 1 ♂, Lahj, 1. X. - 17. XII. 2001, Malaise trap, leg. A. van Harten & A. Sallum (RMNH); 2 ♂♂, 1 ♀, Lahj, III. - V. 2002, Malaise trap, leg. A. van Harten & A. Sallum (RMNH).	en	Rosa, Paolo (2024): Contributions to the cuckoo wasp fauna (Hymenoptera: Chrysididae) of Yemen with description of five new species and updated checklist. Journal of Insect Biodiversity and Systematics 10 (4): 995-1031, DOI: 10.61186/jibs.10.4.995, URL: https://doi.org/10.61186/jibs.10.4.995
03AD991AFF91574FFFDED3B5CEE6D367.taxon	distribution	Distribution in Yemen. Lahj (new record). Distribution in the Arabian Peninsula. Yemen (new record). Oman, Saudi Arabia (Linsenmaier, 1994), United Arab Emirates (Linsenmaier, 1994; Howarth & Gillett, 2008; Strumia, 2014).	en	Rosa, Paolo (2024): Contributions to the cuckoo wasp fauna (Hymenoptera: Chrysididae) of Yemen with description of five new species and updated checklist. Journal of Insect Biodiversity and Systematics 10 (4): 995-1031, DOI: 10.61186/jibs.10.4.995, URL: https://doi.org/10.61186/jibs.10.4.995
03AD991AFF91574FFFDED002CB17D405.taxon	materials_examined	Material examined. 1 ex., Yemen: Near Ta'izz, field on road to Mocha, ca 4,100 ft 16. XII. 1937 / B. M. Exp. to S. W. Arabia H. Scott & E. B. Britton. B. M. 1938 - 246 / NML _ ENT GBIF _ Chr 0001612 / Omalus (Philoctetes) deflexus Ab. det. Linsenmaier 1964 (NMLU).	en	Rosa, Paolo (2024): Contributions to the cuckoo wasp fauna (Hymenoptera: Chrysididae) of Yemen with description of five new species and updated checklist. Journal of Insect Biodiversity and Systematics 10 (4): 995-1031, DOI: 10.61186/jibs.10.4.995, URL: https://doi.org/10.61186/jibs.10.4.995
03AD991AFF91574FFFDED002CB17D405.taxon	distribution	Distribution in Yemen. Ta'izz (Linsenmaier, 1994 as Omalus (Philoctetes) deflexus; present study); no specific locality (Strumia & Dawah, 2019) Distribution in the Arabian Peninsula. Saudi Arabia (Linsenmaier, 1994 as Omalus (Philoctetes) deflexus; Strumia & Dawah, 2019), United Arab Emirates (Strumia, 2014). Extralimital distribution. Greece, Northern Africa, Sudan, Palestine, Syria (Kimsey & Bohart, 1991; Linsenmaier, 1994).	en	Rosa, Paolo (2024): Contributions to the cuckoo wasp fauna (Hymenoptera: Chrysididae) of Yemen with description of five new species and updated checklist. Journal of Insect Biodiversity and Systematics 10 (4): 995-1031, DOI: 10.61186/jibs.10.4.995, URL: https://doi.org/10.61186/jibs.10.4.995
03AD991AFF91574FFFDED60FCEB2D77F.taxon	materials_examined	Material examined. 1 ♂, Yemen: N Jemen Wadi Siham 7. VI. 1987 leg. Mühle / Type Omalus (Philoctetes) jemenensis Lins det. Linsenmaier 1989 / NML _ ENT GBIF _ Chr 0001694 (NMLU).	en	Rosa, Paolo (2024): Contributions to the cuckoo wasp fauna (Hymenoptera: Chrysididae) of Yemen with description of five new species and updated checklist. Journal of Insect Biodiversity and Systematics 10 (4): 995-1031, DOI: 10.61186/jibs.10.4.995, URL: https://doi.org/10.61186/jibs.10.4.995
03AD991AFF91574FFFDED60FCEB2D77F.taxon	distribution	Distribution in Yemen. Wadi Siham (Linsenmaier, 1994 as Omalus (Philoctetes) jemenensis, present study). Distribution in the Arabian Peninsula. Yemen. United Arab Emirates (Strumia, 2014).	en	Rosa, Paolo (2024): Contributions to the cuckoo wasp fauna (Hymenoptera: Chrysididae) of Yemen with description of five new species and updated checklist. Journal of Insect Biodiversity and Systematics 10 (4): 995-1031, DOI: 10.61186/jibs.10.4.995, URL: https://doi.org/10.61186/jibs.10.4.995
03AD991AFF91574CFFDED4BECFE3D09B.taxon	materials_examined	Material examined. 1 ♀, Yemen: Al-Lahima, 16. X. - 31. XII. 2000, Malaise trap, leg. A. van Harten & A. M. Hager (RMNH).	en	Rosa, Paolo (2024): Contributions to the cuckoo wasp fauna (Hymenoptera: Chrysididae) of Yemen with description of five new species and updated checklist. Journal of Insect Biodiversity and Systematics 10 (4): 995-1031, DOI: 10.61186/jibs.10.4.995, URL: https://doi.org/10.61186/jibs.10.4.995
03AD991AFF91574CFFDED4BECFE3D09B.taxon	distribution	Distribution in Yemen. Al-Lahima (new record). Distribution in the Arabian Peninsula. Yemen (new record). Saudi Arabia (Strumia & Dawah, 2012).	en	Rosa, Paolo (2024): Contributions to the cuckoo wasp fauna (Hymenoptera: Chrysididae) of Yemen with description of five new species and updated checklist. Journal of Insect Biodiversity and Systematics 10 (4): 995-1031, DOI: 10.61186/jibs.10.4.995, URL: https://doi.org/10.61186/jibs.10.4.995
03AD991AFF92574CFF2FD12EC81ED24C.taxon	distribution	Distribution in Yemen. Aden (Mocsáry, 1912). Distribution in the Arabian Peninsula. Yemen (Mocsáry, 1912; Rosa et al., 2017); Oman (Strumia, 2008), United Arab Emirates (Linsenmaier, 1994 as Chrysis (Cornuchrysis) adenica; Howarth & Gillett, 2008; Strumia, 2008, 2014; Rosa et al., 2017).	en	Rosa, Paolo (2024): Contributions to the cuckoo wasp fauna (Hymenoptera: Chrysididae) of Yemen with description of five new species and updated checklist. Journal of Insect Biodiversity and Systematics 10 (4): 995-1031, DOI: 10.61186/jibs.10.4.995, URL: https://doi.org/10.61186/jibs.10.4.995
03AD991AFF92574CFF2FD754C8A9D45B.taxon	materials_examined	Material examined. 1 ♀, Yemen: Al-Lahima, 16. X. - 31. XII. 2000, Malaise trap, leg. A. van Harten & A. M. Hager (RMNH).	en	Rosa, Paolo (2024): Contributions to the cuckoo wasp fauna (Hymenoptera: Chrysididae) of Yemen with description of five new species and updated checklist. Journal of Insect Biodiversity and Systematics 10 (4): 995-1031, DOI: 10.61186/jibs.10.4.995, URL: https://doi.org/10.61186/jibs.10.4.995
03AD991AFF92574CFF2FD754C8A9D45B.taxon	distribution	Distribution in Yemen. Al-Lahima, Yemen (Strumia, 2008, 2014). Distribution in the Arabian Peninsula. Oman, Saudi Arabia, United Arab Emirates (Linsenmaier, 1994, 1999; Howarth & Gillett, 2008; Strumia, 2014).	en	Rosa, Paolo (2024): Contributions to the cuckoo wasp fauna (Hymenoptera: Chrysididae) of Yemen with description of five new species and updated checklist. Journal of Insect Biodiversity and Systematics 10 (4): 995-1031, DOI: 10.61186/jibs.10.4.995, URL: https://doi.org/10.61186/jibs.10.4.995
03AD991AFF92574CFF2ED6FECE6CD779.taxon	distribution	Distribution in Yemen. Yemen (Kimsey & Bohart, 1991; Rosa et al., 2017). Distribution in the Arabian Peninsula. Arabia (with no specific locality, Mocsáry, 1911 as Chrysis (Holochrysis) arabica), Yemen (Kimsey & Bohart, 1991; Rosa et al., 2017).	en	Rosa, Paolo (2024): Contributions to the cuckoo wasp fauna (Hymenoptera: Chrysididae) of Yemen with description of five new species and updated checklist. Journal of Insect Biodiversity and Systematics 10 (4): 995-1031, DOI: 10.61186/jibs.10.4.995, URL: https://doi.org/10.61186/jibs.10.4.995
03AD991AFF925752FF2FD59CCBEFD902.taxon	materials_examined	Material examined. Holotype ♀, Yemen: Lahj, III. - V. 2002, Malaise trap, A. van Harten & A. Sallum (RMNH).	en	Rosa, Paolo (2024): Contributions to the cuckoo wasp fauna (Hymenoptera: Chrysididae) of Yemen with description of five new species and updated checklist. Journal of Insect Biodiversity and Systematics 10 (4): 995-1031, DOI: 10.61186/jibs.10.4.995, URL: https://doi.org/10.61186/jibs.10.4.995
03AD991AFF925752FF2FD59CCBEFD902.taxon	diagnosis	Diagnosis. Chrysis bilqis sp. nov. belongs to the leachii group. Species in this group are very small to small (2.5 – 5.0 mm), usually brightly colored, with the apex of third metasomal tergum either edentate or with short median tooth, sometimes lanceolate; the transverse frontal carina is faint; the malar spaces are short and strongly convergent; the scapal basin is broadly microridged in both sexes; the black spots on the second sternum vary from medium (covering about half the segment length) to large (covering two-thirds of the segment length), and are subsquare or subrectangular, normally separated by a narrow metallic line. Aside from a few dark species endemic to the CanaryIslands, only two species have both sexes with a completely green to blue body colour and are distributed in Africa and Central Asia: Chrysis nilensis Linsenmaier, 1959 and C. infantula Semenov-Tian-Shanskij, 1967 (Rosa, 2021). Chrysis bilqis sp. nov. can be distinguished from the females of C. nilensis and C. infantula by the following characteristics: even punctation with polished interspaces becoming larger on the second half of the second tergum (vs. coarse, dense punctation, without distinct polished interspaces in the other two species, Fig. 8 A); black spots on the second sternum entirely fused medially, laterally and basally, extending under the first sternum (Fig. 7 F) (in C. nilensis female, the spots are largely separated medially and basally, Fig. 8 B; while in C. infantula the spots are separated by a narrow metallic line and are expanded below the first tergum).	en	Rosa, Paolo (2024): Contributions to the cuckoo wasp fauna (Hymenoptera: Chrysididae) of Yemen with description of five new species and updated checklist. Journal of Insect Biodiversity and Systematics 10 (4): 995-1031, DOI: 10.61186/jibs.10.4.995, URL: https://doi.org/10.61186/jibs.10.4.995
03AD991AFF925752FF2FD59CCBEFD902.taxon	description	Description. Holotype ♀ (Fig. 7 A – F). Body length: 4.7 mm. Head. Frons between scapal basin and anterior ocellus with dense, polygonal, large (about 0.6 – 0.7 × MOD) and contiguous punctures, without polished interspaces; punctures arranged radially around anterior ocellus with their size regularly decreasing towards ocellus; head posterior to ocelli with smaller punctures; impunctate postero-laterally to ocelli; transverse frontal carina faint; scapal basin deep, with median sharp wrinkles; face between scapal basin and eye margin with large, contiguous punctures like those on frons (Fig. 7 B); genal carina straight, sharp, fully developed from temples to mandibular insertion; malar spaces relatively long (1.4 × MOD); subantennal space short, 0.6 × MOD; apex of clypeus with thin brown rim. OOL 1.9 × MOD; POL 2.8 × MOD; MS 1.4 × MOD; relative length of P: F 1: F 2: F 3 = 1.0: 1.3: 0.8: 0.7. Mesosoma. Medial pronotal line large and elongate, reaching two-thirds of pronotal length; pronotum as long as mesoscutellum, with deep, large punctures, as large as those on frons, rounded and with narrow, polished interspaces; mesoscutum with double punctation, larger punctures on median area; punctures separated by polished interspaces up to 1 puncture diameter; notauli formed by deep, subrectangular foveae, larger at base, their width smaller than larger adjacent punctures; parapsidal signum as deep line; mesoscutellum with shallow punctures, slightly smaller than those on mesoscutum with wider interspaces; scutellar-metanotal suture deep; metanotum antero-medially with large and deep fovea (Fig. 7 C); metanotal punctures denser, without large, polished interspaces; posterior propodeal projections slightly divergent; mesopleuron with punctures similar to those at sides of propodeum; episternal sulcus formed by row of round to subsquare foveae, partially confluent with other punctures. Metasoma. First tergum densely micropunctate on interspaces; punctures larger dorsally, distinctly smaller and aligned along apical margin (Fig. 7 D); largest punctures smaller than those on mesoscutellum; antero-median concavity deep and wide; second tergum with dense, double punctation on disc, with small punctures on interspaces; even, large interspaces between punctures, up to 1 puncture diameter, on remaining tergum, with only scattered small punctures on interspaces; median longitudinal carina indicated by punctate line, composed by small punctures; punctation on third tergum denser, double; apical area without slight pre pit swelling; pits of pit row large, deep, as large as 2 – 3 puncture diameters (Fig. 7 E); apical margin with blunt median tooth (Fig. 7 E); apico-median margin bordered by thin, hyaline rim; black spots on second sternum fused medially, forming subrectangular black stripe (Fig. 7 F). Colour. Body entirely blue; scape, pedicel and first flagellomere metallic blue, remaining flagellomeres brown; tegula metallic blue; wings hyaline with brown nervures; legs blue on femur and outer tibia, tarsi brown; sterna metallic blue. Vestiture. Head and mesosoma with short, whitish setae (up to 1 × MOD); metasoma laterally and posteriorly with long, erected setae, as long as 2 × MOD. Femora and tibiae with scattered, elongate and whitish setae, as long as 2 × MOD (Fig. 7 F). Male. Unknown.	en	Rosa, Paolo (2024): Contributions to the cuckoo wasp fauna (Hymenoptera: Chrysididae) of Yemen with description of five new species and updated checklist. Journal of Insect Biodiversity and Systematics 10 (4): 995-1031, DOI: 10.61186/jibs.10.4.995, URL: https://doi.org/10.61186/jibs.10.4.995
03AD991AFF925752FF2FD59CCBEFD902.taxon	etymology	Etymology. The specific epiteth belqis is dedicated to Belqis, the Yemenite and Islamic name for the Queen of Sheba, who has become the subject of one of the most widespread and fertile cycles of legends in Asia and Africa. Sheba was the ancient South Arabian kingdom of Saba’a.	en	Rosa, Paolo (2024): Contributions to the cuckoo wasp fauna (Hymenoptera: Chrysididae) of Yemen with description of five new species and updated checklist. Journal of Insect Biodiversity and Systematics 10 (4): 995-1031, DOI: 10.61186/jibs.10.4.995, URL: https://doi.org/10.61186/jibs.10.4.995
03AD991AFF925752FF2FD59CCBEFD902.taxon	distribution	Distribution. Yemen.	en	Rosa, Paolo (2024): Contributions to the cuckoo wasp fauna (Hymenoptera: Chrysididae) of Yemen with description of five new species and updated checklist. Journal of Insect Biodiversity and Systematics 10 (4): 995-1031, DOI: 10.61186/jibs.10.4.995, URL: https://doi.org/10.61186/jibs.10.4.995
03AD991AFF8D5753FFDED049C9A3D55E.taxon	materials_examined	Material examined. 1 ♀, Yemen: N Yemen Hadda 30. V. 87 leg. Mühle / 459 / Type Chrysis convexianalis Lins. det. Linsenmaier 1990 / NML _ ENT GBIF _ Chr 0019414 (NMLU); 1 ♀, paratype, same data of holotype, NML _ ENT GBIF _ Chr 0019412 (NMLU); 1 ♀, N Yemen Wadi Siham 7. VI. 1987 leg. Mühle / Paratype Chrysis convexianalis Lins. det. Linsenmaier 1990 / NML _ ENT GBIF _ Chr 0019413 (NMLU).	en	Rosa, Paolo (2024): Contributions to the cuckoo wasp fauna (Hymenoptera: Chrysididae) of Yemen with description of five new species and updated checklist. Journal of Insect Biodiversity and Systematics 10 (4): 995-1031, DOI: 10.61186/jibs.10.4.995, URL: https://doi.org/10.61186/jibs.10.4.995
03AD991AFF8D5753FFDED049C9A3D55E.taxon	distribution	Distribution in Yemen. Hadda, Wadi Siham (Linsenmaier, 1994; present study). Distribution in the Arabian Peninsula. Yemen (Linsenmaier, 1994).	en	Rosa, Paolo (2024): Contributions to the cuckoo wasp fauna (Hymenoptera: Chrysididae) of Yemen with description of five new species and updated checklist. Journal of Insect Biodiversity and Systematics 10 (4): 995-1031, DOI: 10.61186/jibs.10.4.995, URL: https://doi.org/10.61186/jibs.10.4.995
03AD991AFF8D5753FFDED7FDC8CDD4A6.taxon	materials_examined	Material examined. 1 ♀, Yemen: Al-Lahima, 16. X. - 31. XII. 2000, Malaise trap, leg. A. van Harten & A. M. Hager (RMNH); 1 ♀, Lahj, III. - V. 2002, Lalaise trap, A. van Harten & A. Sallum (RMNH).	en	Rosa, Paolo (2024): Contributions to the cuckoo wasp fauna (Hymenoptera: Chrysididae) of Yemen with description of five new species and updated checklist. Journal of Insect Biodiversity and Systematics 10 (4): 995-1031, DOI: 10.61186/jibs.10.4.995, URL: https://doi.org/10.61186/jibs.10.4.995
03AD991AFF8D5753FFDED7FDC8CDD4A6.taxon	distribution	Distribution in Yemen. Al-Lahima, Lahj (new record). Distribution in the Arabian Peninsula. Saudi Arabia (Linsenmaier, 1994; Strumia & Dawah, 2010, 2012), United Arab Emirates (Strumia, 2014).	en	Rosa, Paolo (2024): Contributions to the cuckoo wasp fauna (Hymenoptera: Chrysididae) of Yemen with description of five new species and updated checklist. Journal of Insect Biodiversity and Systematics 10 (4): 995-1031, DOI: 10.61186/jibs.10.4.995, URL: https://doi.org/10.61186/jibs.10.4.995
03AD991AFF8D5750FFDED554CFF2D6E3.taxon	materials_examined	Material examined. Holotype ♀, Yemen: 12 km NW of Manakhah, 3. VII. - 21. VIII. 2001, Malaise trap, leg. A. van Harten (RMNH). Paratypes 1 ♀, same data (RMNH); 1 ♂, Al-Lahima, I. 2009 - 9. IV. 2001, Malaise trap, leg. A. van Harten & A. M. Hajer (RMNH).	en	Rosa, Paolo (2024): Contributions to the cuckoo wasp fauna (Hymenoptera: Chrysididae) of Yemen with description of five new species and updated checklist. Journal of Insect Biodiversity and Systematics 10 (4): 995-1031, DOI: 10.61186/jibs.10.4.995, URL: https://doi.org/10.61186/jibs.10.4.995
03AD991AFF8D5750FFDED554CFF2D6E3.taxon	diagnosis	Diagnosis. Chrysis felix sp. nov. belongs to the succincta group. It is closely related to the Arabian species Chrysis maidaquensis Strumia, 2014, which was originally described from United Arab Emirates and later reported from Iran (Tavasoli & Fallahzadeh, 2015, Farhard et al., 2015; Rosa et al., 2024 a). The female of Chrysis felix sp. nov. can be easily distinguished by the following combination of characters: frons without transverse frontal carina (Fig. 9 B) (vs. double carina in C. maidaquensis); apical margin of the third tergum with a blunt, large median tooth and two lateral blunt corners (Fig. 9 E) (vs. three long and pointed teeth in C. maidaquensis); black spots on the second sternum rounded and separated medially (Fig. 9 F), covering about two-thirds of sternal length (vs. black spots fused in a transverse rectangle, covering about half sternal length). The male shares a similar habitus, colour pattern, and sculpture; the median apical tooth of the third tergum is more of an undulation rather than a distinct lobe or pointed tooth (Figs 10 E, 10 G); the genital capsule has the inner margin of gonocoxa forming a right angle with the apical margin (Fig. 10 F) (the male of Chrysis maidaquensis is still unknown).	en	Rosa, Paolo (2024): Contributions to the cuckoo wasp fauna (Hymenoptera: Chrysididae) of Yemen with description of five new species and updated checklist. Journal of Insect Biodiversity and Systematics 10 (4): 995-1031, DOI: 10.61186/jibs.10.4.995, URL: https://doi.org/10.61186/jibs.10.4.995
03AD991AFF8D5750FFDED554CFF2D6E3.taxon	description	Description. Holotype ♀ (Fig. 9 A – F). Body length: 4.0 mm (female paratype body length: 3.6 mm). Head. Frons with dense small to medium punctures (0.3 – 0.5 × MOD), subcontiguous without interspaces; frontal carina faint (Fig. 9 B); scapal basin deep, fully wrinkled; area between scapal basin and eye with large punctures from frons to malar space; malar space long (1.4 × MOD) covered by dense, large punctures (Fig. 9 C); subantennal space short (0.7 × MOD); genal carina sharp, fully developed from temple to mandible insertion; clypeal apex with thin brown rim; mandible simple. Distance from anterior ocellus to scapal basin 2.8 × MOD; OOL 1.6 × MOD; POL 2.3 × MOD; MS 1.4 × MOD. Holotype missing both flagella; relative length of P: F 1: F 2 measured on paratype = 1.0: 1.3: 0.9. Mesosoma. Medial pronotal line deep and elongate, as long as two thirds of pronotal length; pronotum as long as scutellum; pronotal punctation double with punctures of variable size, with narrow interspaces; punctures smaller along basal margin; mesonotum with double punctation, larger punctures situated medially on mesoscutum and mesoscutellum; punctures separated by polished interspaces; notauli formed by subsquare foveae, relatively small (Fig. 9 D); parapsidal signum as deep line; scutellar-metanotal suture deep, formed by longitudinally elongate foveae, longer than larger punctures on mesonotum; metanotal punctures denser, with large anteromedian fovea; posterior propodeal projections subparallel, downwardly directed; mesepimeron with punctures similar to those at sides of propodeum, mesepisteron with smaller punctures; episternal sulcus formed by relatively small irregular foveae, partially confluent each other, as large as punctures on mesepimeron. Metasoma. First tergum with even, relatively small punctures, smaller than those on mesoscutum, with micropunctate interspaces; second tergum with denser, larger, geminate punctation, becoming shallower, smaller and sparser on the second half of tergum; third tergum with denser punctures, as large as those at base of second tergum, with corrugate interspaces; median longitudinal carina weak, on second tergum formed by aligned micropunctures; pits of pit row deep, round, and black (Fig. 9 E); apical margin of third tergum bordered by thick hyaline margin; median tooth as a large lobe or convexity, lateral teeth like blunt angles (Fig. 9 E); black spots on second sternum large, covering about two thirds of sternum length and separated medially (Fig. 9 F). Colour. Head blue, green on frons and face; pronotum blue with anterolateral green band; mesoscutum pinkish to purplish, metanotum and propodeum green; notauli, axillae and sutures blue; metasomal first tergum olive green, second and third terga purplish with apical margin green to blue; sterna golden-green. Scape golden-red, pedicel and flagellomeres non-metallic brown; wings hyaline with brown nervures. Male (Fig. 10). Similar to female for habitus, colour pattern, and sculpture. Sexual dimorphism can be observed in metallic pedicel, longer POL 2.4 × MOD, apical margin of third tergum with three undulations, being the median tooth aligned to lateral ones. Inner margin of gonocoxa fully developed, forming a right angle with apical margin (Fig. 10 F).	en	Rosa, Paolo (2024): Contributions to the cuckoo wasp fauna (Hymenoptera: Chrysididae) of Yemen with description of five new species and updated checklist. Journal of Insect Biodiversity and Systematics 10 (4): 995-1031, DOI: 10.61186/jibs.10.4.995, URL: https://doi.org/10.61186/jibs.10.4.995
03AD991AFF8D5750FFDED554CFF2D6E3.taxon	etymology	Etymology. The specific epithet felix is derived from the Old Latin name of Yemen Arabia Felix, one of the three regions into which the Romans subdivided the Arabian Peninsula: Arabia Deserta, Arabia Felix, and Arabia Petraea. The meaning of Felix is both fertile but also happy, fortunate or blessed, being this area well irrigated in the Arabian Peninsula with significant vegetation and river beds (wadis).	en	Rosa, Paolo (2024): Contributions to the cuckoo wasp fauna (Hymenoptera: Chrysididae) of Yemen with description of five new species and updated checklist. Journal of Insect Biodiversity and Systematics 10 (4): 995-1031, DOI: 10.61186/jibs.10.4.995, URL: https://doi.org/10.61186/jibs.10.4.995
03AD991AFF8D5750FFDED554CFF2D6E3.taxon	distribution	Distribution. Yemen (Al-Lahima, Manakhah).	en	Rosa, Paolo (2024): Contributions to the cuckoo wasp fauna (Hymenoptera: Chrysididae) of Yemen with description of five new species and updated checklist. Journal of Insect Biodiversity and Systematics 10 (4): 995-1031, DOI: 10.61186/jibs.10.4.995, URL: https://doi.org/10.61186/jibs.10.4.995
03AD991AFF8D5750FFDED554CFF2D6E3.taxon	discussion	Remarks. The holotype is damaged missing both flagella, fore and mid legs, right hind leg. These damages are likely due to the prolonged stay in the Malaise trap. The paratype is conversely complete. The choice to designate the damaged specimen as holotype is due to the preparation that allows the study of diagnostic characters, such as face and black spots on second sternum which are not visible clearly on the paratype, being prepared rolled up. All the other diagnostic characters like sculpture, relative lengths, colour and shape of apical margin is anyway conforming between the two females.	en	Rosa, Paolo (2024): Contributions to the cuckoo wasp fauna (Hymenoptera: Chrysididae) of Yemen with description of five new species and updated checklist. Journal of Insect Biodiversity and Systematics 10 (4): 995-1031, DOI: 10.61186/jibs.10.4.995, URL: https://doi.org/10.61186/jibs.10.4.995
03AD991AFF8F5756FFDED4DDCEB3D6CA.taxon	materials_examined	Material examined. 1 ♀, Yemen: Hadramaut Coll. Linsenmaier / 460 / Type Chrysis hadramauta Lins. det. Linsenmaier 1992 / NML _ ENT GBIF _ Chr 0023418 (NMLU).	en	Rosa, Paolo (2024): Contributions to the cuckoo wasp fauna (Hymenoptera: Chrysididae) of Yemen with description of five new species and updated checklist. Journal of Insect Biodiversity and Systematics 10 (4): 995-1031, DOI: 10.61186/jibs.10.4.995, URL: https://doi.org/10.61186/jibs.10.4.995
03AD991AFF8F5756FFDED4DDCEB3D6CA.taxon	distribution	Distribution in Yemen. Hadramaut (Linsenmaier, 1994). Distribution in the Arabian Peninsula. Saudi Arabia (Strumia & Dawah, 2010, 2019).	en	Rosa, Paolo (2024): Contributions to the cuckoo wasp fauna (Hymenoptera: Chrysididae) of Yemen with description of five new species and updated checklist. Journal of Insect Biodiversity and Systematics 10 (4): 995-1031, DOI: 10.61186/jibs.10.4.995, URL: https://doi.org/10.61186/jibs.10.4.995
03AD991AFF895757FFDED3B6CB20D363.taxon	materials_examined	Material examined. 3 ♂♂ and 1 ♀, Yemen: Lahj, III. - V. 2002, Malaise trap, leg. A. van Harten & A. Sallum (RMNH); 1 ♀, Seiyun, light trap, 12. - 14. VII. 2002, leg. A. van Harten (RMNH).	en	Rosa, Paolo (2024): Contributions to the cuckoo wasp fauna (Hymenoptera: Chrysididae) of Yemen with description of five new species and updated checklist. Journal of Insect Biodiversity and Systematics 10 (4): 995-1031, DOI: 10.61186/jibs.10.4.995, URL: https://doi.org/10.61186/jibs.10.4.995
03AD991AFF895757FFDED3B6CB20D363.taxon	distribution	Distribution in Yemen. Lahj, Seiyun (present study); Aden (Linsenmaier, 1994 as Chrysis (Hexachrysis) jousseaumei). Distribution in the Arabian Peninsula. Yemen. United Arab Emirates (Linsenmaier, 1994 as Chrysis (Hexachrysis) jousseaumei; Howarth & Gillett, 2008; Strumia, 2008; Madl, 2018). Extralimital distribution. Northern Africa, Somalia to South Africa (Kimsey & Bohart, 1991; Linsenmaier, 1999; Madl, 2018).	en	Rosa, Paolo (2024): Contributions to the cuckoo wasp fauna (Hymenoptera: Chrysididae) of Yemen with description of five new species and updated checklist. Journal of Insect Biodiversity and Systematics 10 (4): 995-1031, DOI: 10.61186/jibs.10.4.995, URL: https://doi.org/10.61186/jibs.10.4.995
03AD991AFF895757FFDED1C7CE4AD29A.taxon	materials_examined	Material examined. 2 ♀♀, Yemen: Wadi Higgan, 7. VIII. 1936 (EFC).	en	Rosa, Paolo (2024): Contributions to the cuckoo wasp fauna (Hymenoptera: Chrysididae) of Yemen with description of five new species and updated checklist. Journal of Insect Biodiversity and Systematics 10 (4): 995-1031, DOI: 10.61186/jibs.10.4.995, URL: https://doi.org/10.61186/jibs.10.4.995
03AD991AFF895757FFDED1C7CE4AD29A.taxon	distribution	Distribution in Yemen. Wadi Higgan (Rosa et al., 2024 b; present study). Distribution in the Arabian Peninsula. Oman, United Arab Emirates (Linsenmaier, 1994 as C. (Pyria) lyncea; Howarth & Gillett, 2008; Strumia, 2008), Yemen (Rosa et al., 2024 b).	en	Rosa, Paolo (2024): Contributions to the cuckoo wasp fauna (Hymenoptera: Chrysididae) of Yemen with description of five new species and updated checklist. Journal of Insect Biodiversity and Systematics 10 (4): 995-1031, DOI: 10.61186/jibs.10.4.995, URL: https://doi.org/10.61186/jibs.10.4.995
03AD991AFF895757FFDED080CEA2D5BF.taxon	materials_examined	Material examined. 1 ♀, Yemen: Seiyun, 12. - 14. VII. 2002, light trap, leg. A. van Harten (RMNH).	en	Rosa, Paolo (2024): Contributions to the cuckoo wasp fauna (Hymenoptera: Chrysididae) of Yemen with description of five new species and updated checklist. Journal of Insect Biodiversity and Systematics 10 (4): 995-1031, DOI: 10.61186/jibs.10.4.995, URL: https://doi.org/10.61186/jibs.10.4.995
03AD991AFF895757FFDED080CEA2D5BF.taxon	distribution	Distribution in Yemen. Seiyun (new record). Distribution in the Arabian Peninsula. Yemen, Saudi Arabia (Strumia & Dawah, 2012).	en	Rosa, Paolo (2024): Contributions to the cuckoo wasp fauna (Hymenoptera: Chrysididae) of Yemen with description of five new species and updated checklist. Journal of Insect Biodiversity and Systematics 10 (4): 995-1031, DOI: 10.61186/jibs.10.4.995, URL: https://doi.org/10.61186/jibs.10.4.995
03AD991AFF895757FFDED6C7CE52D7BF.taxon	materials_examined	Material examined. 1 ♀, Yemen: Wadi Hirran, 28. iv. 1936 (EFC); 1 ♂, Sana’a University Campus, 2300 m, 3. XI. 2005, leg. J. Halada (MHC); 1 ♂, Wadi Sudd 10 km W Marib, 1120 m, 15 ° 24 ’ N, 45 ° 16 ’ E, 8. X. 2005, leg. J. Halada (MHC).	en	Rosa, Paolo (2024): Contributions to the cuckoo wasp fauna (Hymenoptera: Chrysididae) of Yemen with description of five new species and updated checklist. Journal of Insect Biodiversity and Systematics 10 (4): 995-1031, DOI: 10.61186/jibs.10.4.995, URL: https://doi.org/10.61186/jibs.10.4.995
03AD991AFF895757FFDED6C7CE52D7BF.taxon	distribution	Distribution in Yemen. Hajjah Governorate, Sana’a, Wadi Sudd (Rosa et al., 2024 b). Distribution in the Arabian Peninsula. Oman, Yemen (Rosa et al., 2024 b).	en	Rosa, Paolo (2024): Contributions to the cuckoo wasp fauna (Hymenoptera: Chrysididae) of Yemen with description of five new species and updated checklist. Journal of Insect Biodiversity and Systematics 10 (4): 995-1031, DOI: 10.61186/jibs.10.4.995, URL: https://doi.org/10.61186/jibs.10.4.995
03AD991AFF895754FFDED45EC80CD93B.taxon	materials_examined	Material examined. 1 ♂, Yemen: Lahj, III. - V. 2002, Malaise trap, leg. A. van Harten & A. Sallum (RMNH); 1 ♂, Seiyun, 12. - 14. VIII. 2002, light trap, leg. A. van Harten (RMNH); 1 ♀, Al Kowd, I. 2000, light trap, leg. A. van Harten & S. Al Haruri (RMNH).	en	Rosa, Paolo (2024): Contributions to the cuckoo wasp fauna (Hymenoptera: Chrysididae) of Yemen with description of five new species and updated checklist. Journal of Insect Biodiversity and Systematics 10 (4): 995-1031, DOI: 10.61186/jibs.10.4.995, URL: https://doi.org/10.61186/jibs.10.4.995
03AD991AFF895754FFDED45EC80CD93B.taxon	distribution	Distribution in Yemen. Al Kowd, Lahj, Seiyun (present study). Distribution in the Arabian Peninsula. Yemen (Madl, 2018). Oman, Saudi Arabia (Linsenmaier, 1994; Strumia & Dawah, 2019), United Arab Emirates (Linsenmaier, 1994; Howarth & Gillett, 2008; Strumia, 2008, 2014). Extralimital distribution. Central Asia, Northern Africa, widespread in sub-Saharan Africa (Kimsey & Bohart, 1991; Linsenmaier, 1994, 1999).	en	Rosa, Paolo (2024): Contributions to the cuckoo wasp fauna (Hymenoptera: Chrysididae) of Yemen with description of five new species and updated checklist. Journal of Insect Biodiversity and Systematics 10 (4): 995-1031, DOI: 10.61186/jibs.10.4.995, URL: https://doi.org/10.61186/jibs.10.4.995
03AD991AFF8B5755FFDED3B5C815D314.taxon	materials_examined	Material examined. 1 ♀, Yemen: Sana’a gov., 5 km SW of Matnah vill., stream valley, 15 ° 13 ' 48 " N 43 ° 59 ' 42 " E, 2750 m, 19. xi. 2010, leg. J. Hájek (MNHP).	en	Rosa, Paolo (2024): Contributions to the cuckoo wasp fauna (Hymenoptera: Chrysididae) of Yemen with description of five new species and updated checklist. Journal of Insect Biodiversity and Systematics 10 (4): 995-1031, DOI: 10.61186/jibs.10.4.995, URL: https://doi.org/10.61186/jibs.10.4.995
03AD991AFF8B5755FFDED3B5C815D314.taxon	distribution	Distribution in Yemen. Sana’s (present study), Yemen (Mocsáry, 1912) Distribution in the Arabian Peninsula. Saudi Arabia (Farasan) (Strumia & Dawah, 2019); Arabia (with no specific locality, Linsenmaier, 1999).	en	Rosa, Paolo (2024): Contributions to the cuckoo wasp fauna (Hymenoptera: Chrysididae) of Yemen with description of five new species and updated checklist. Journal of Insect Biodiversity and Systematics 10 (4): 995-1031, DOI: 10.61186/jibs.10.4.995, URL: https://doi.org/10.61186/jibs.10.4.995
03AD991AFF8B5755FFDED100CF3AD26B.taxon	materials_examined	Material examined. 1 ♀, Yemen: Lahj, III. - V. 2002, Malaise trap, leg. A. van Harten & A. Sallum (RMNH).	en	Rosa, Paolo (2024): Contributions to the cuckoo wasp fauna (Hymenoptera: Chrysididae) of Yemen with description of five new species and updated checklist. Journal of Insect Biodiversity and Systematics 10 (4): 995-1031, DOI: 10.61186/jibs.10.4.995, URL: https://doi.org/10.61186/jibs.10.4.995
03AD991AFF8B5755FFDED100CF3AD26B.taxon	distribution	Distribution in Yemen. Lahj (new record). Distribution in the Arabian Peninsula. Yemen (new record). Oman (Linsenmaier, 1994), Saudi Arabia (Linsenmaier, 1994; Strumia & Dawah, 2010, 2019), United Arab Emirates (Strumia, 2008, 2014).	en	Rosa, Paolo (2024): Contributions to the cuckoo wasp fauna (Hymenoptera: Chrysididae) of Yemen with description of five new species and updated checklist. Journal of Insect Biodiversity and Systematics 10 (4): 995-1031, DOI: 10.61186/jibs.10.4.995, URL: https://doi.org/10.61186/jibs.10.4.995
03AD991AFF8B5755FFDED08ECF75D589.taxon	distribution	Distribution in Yemen. Yemen (with no locality cited) (Strumia, 2014). Distribution in the Arabian Peninsula. Yemen (Strumia, 2014). Arabia (with no specific locality, Linsenmaier, 1999), Qatar (Linsenmaier, 1994), United Arab Emirates (Strumia, 2008, 2014).	en	Rosa, Paolo (2024): Contributions to the cuckoo wasp fauna (Hymenoptera: Chrysididae) of Yemen with description of five new species and updated checklist. Journal of Insect Biodiversity and Systematics 10 (4): 995-1031, DOI: 10.61186/jibs.10.4.995, URL: https://doi.org/10.61186/jibs.10.4.995
03AD991AFF8B5755FFDED7ADCBC9D4F8.taxon	materials_examined	Material examined. 1 ♀, Yemen: Wadi Hirran, 28. IV. 1930 (EFC).	en	Rosa, Paolo (2024): Contributions to the cuckoo wasp fauna (Hymenoptera: Chrysididae) of Yemen with description of five new species and updated checklist. Journal of Insect Biodiversity and Systematics 10 (4): 995-1031, DOI: 10.61186/jibs.10.4.995, URL: https://doi.org/10.61186/jibs.10.4.995
03AD991AFF8B5755FFDED7ADCBC9D4F8.taxon	distribution	Distribution in Yemen. Hajjah Governorate (Rosa et al., 2024 b). Distribution in the Arabian Peninsula. Yemen (Rosa et al., 2024 b). Saudi Arabia (Linsenmaier, 1994 as Chrysis (Pyria) simillima).	en	Rosa, Paolo (2024): Contributions to the cuckoo wasp fauna (Hymenoptera: Chrysididae) of Yemen with description of five new species and updated checklist. Journal of Insect Biodiversity and Systematics 10 (4): 995-1031, DOI: 10.61186/jibs.10.4.995, URL: https://doi.org/10.61186/jibs.10.4.995
03AD991AFF8B5755FFDED51CC873D6EF.taxon	materials_examined	Material examined. 1 ♀, Yemen: Gabal El Mogash (s. w. Sana’a), 31. v. 1936 (EFC); 1 ♂, Wadi Sharis, 1936 (EFC); 3 ♀, Wadi Higgan, 7. viii. 1936 (Manahla-Hajila) (EFC).	en	Rosa, Paolo (2024): Contributions to the cuckoo wasp fauna (Hymenoptera: Chrysididae) of Yemen with description of five new species and updated checklist. Journal of Insect Biodiversity and Systematics 10 (4): 995-1031, DOI: 10.61186/jibs.10.4.995, URL: https://doi.org/10.61186/jibs.10.4.995
03AD991AFF8B5755FFDED51CC873D6EF.taxon	distribution	Distribution in Yemen. Gabal El Mogash, Wadi Sharis, Wadi Higgan (Rosa et al., 2024 b); Yemen (no specific locality) (Madl, 2018). Distribution in the Arabian Peninsula. Yemen (Madl, 2018). Kuwait (Al-Houty, 1989 as Stilbum splendidum), Oman, Saudi Arabia (Linsenmaier, 1994 as Chrysis (Pyria) stilboides), United Arab Emirates (Strumia, 2014). Extralimital distribution. Widespread in sub-Saharan Africa to South Asia (Kimsey & Bohart, 1991; Linsenmaier, 1994; Madl, 2018).	en	Rosa, Paolo (2024): Contributions to the cuckoo wasp fauna (Hymenoptera: Chrysididae) of Yemen with description of five new species and updated checklist. Journal of Insect Biodiversity and Systematics 10 (4): 995-1031, DOI: 10.61186/jibs.10.4.995, URL: https://doi.org/10.61186/jibs.10.4.995
03AD991AFF845758FF2FD3B4C84DD070.taxon	materials_examined	Material examined. Holotype ♀, Yemen: Al-Lahima, I. 2001 – 9. IV. 2001, in Malaise trap, A. van Harten & A. M. Hajer, RMNH Leiden ex collection ZMAN (RMNH).	en	Rosa, Paolo (2024): Contributions to the cuckoo wasp fauna (Hymenoptera: Chrysididae) of Yemen with description of five new species and updated checklist. Journal of Insect Biodiversity and Systematics 10 (4): 995-1031, DOI: 10.61186/jibs.10.4.995, URL: https://doi.org/10.61186/jibs.10.4.995
03AD991AFF845758FF2FD3B4C84DD070.taxon	diagnosis	Diagnosis. Chrysis yemenita sp. nov. belongs to the succincta group and is closely related to the Central Asian Chrysis irenes Semenov-Tian-Shanskij & Nikol’skaya, 1954, known from Tajikistan. The type is illustrated in Rosa et al. (2017, plate 67). Chrysis yemenita sp. nov. can be distinguished by its colour pattern, an important diagnostic character in this species group, as well as by the shape of the apical margin of the third tergum, the pit row and the body punctation. The body colour of Chrysis yemenita sp. nov. is blue with red to golden-green areas on the mesosoma and metasoma rather than green with red to purplish areas (Fig. 12 A). However, this coloration may have been altered due to prolonged preservation in ethanol from a Malaise trap. Nonetheless, the colour pattern remains a key character for this species: the mesoscutum, scutellum and metascutum and propodeum are red to greenish, contrasting with the blue colour of the basal pronotum, tegula, and mesopleuron (whereas C. irenes has only the red mesoscutum contrasting with green scutellum, metascutum and propodeum); pronotum largely greenish anteriorly and laterally (almost entirely green in C. irenes); sterna blue (Fig. 12 F) (flame red in C. irenes); black spots on second sternum separated medially (Fig. 12 F) (fused in C. irenes); transverse frontal carina fully developed, sharp (Fig. 12 B) (faint to barely visible in C. irenes); metasomal punctation denser, with larger and deeper punctures (Fig. 12 E); third tergum with long, pointed teeth and small, shallow pits of the pit row (short and blunt apical teeth, deep and elongate pits of the pit row in C. irenes). It is also similar to C. chamrosh Rosa, 2024 [in Rosa et al., 2024 a] but immediately distinguishable by the blue colour of the sterna instead of red.	en	Rosa, Paolo (2024): Contributions to the cuckoo wasp fauna (Hymenoptera: Chrysididae) of Yemen with description of five new species and updated checklist. Journal of Insect Biodiversity and Systematics 10 (4): 995-1031, DOI: 10.61186/jibs.10.4.995, URL: https://doi.org/10.61186/jibs.10.4.995
03AD991AFF845758FF2FD3B4C84DD070.taxon	description	Description. Holotype ♀ (Fig. 12 A – F). Body length: 6.2 mm. Head. Frons with dense small punctures (0.2 – 0.4 MOD), larger and more spaced on vertex between lateral ocelli and eyes, denser and smaller on ocelli area, occipital area and temples; punctures between scapal basin and transverse frontal carina longitudinally elongate; lateral ocelli with deep lateral fovea; frontal carina sharp, slightly downcurved medially and with curved lateral branches arriving at declivity of scapal basin (Fig. 12 B); scapal basin deep, impunctate medially with dense, small to medium punctures laterally; malar space long (1.1 × MOD) covered by this dense punctation; subantennal space short, half as long as malar space; genal carina sharp, fully developed from temple to mandible insertion; apex of clypeus with thin brown rim. OOL 1.8 × MOD; POL 2.3 × MOD; MS 1.1 × MOD; relative length of P: F 1: F 2: F 3 = 1.0: 1.2: 0.8: 0.7. Mesosoma. Medial pronotal line weak and elongate, reaching one third of pronotal length; pronotum as long as scutellum, with spaced, small to medium punctures, as large as those on vertex, rounded and with polished interspaces; mesoscutum with larger punctures on median area; punctures separated by polished interspaces up to 1 puncture diameter; notauli formed by deep foveae subrectangular at base, small and rounded towards apex (Fig. 12 C), parapsidal signum as deep line; mesoscutellum with spaced and shallow punctures, slightly larger than those on mesoscutum; scutellar-metanotal suture deep, formed by longitudinally elongate foveae; metanotal punctures denser, with anteromedian polished area; posterior propodeal projections slightly divergent; mesopleuron with punctures similar to those at sides of propodeum; episternal sulcus formed by subsquare foveae, partially confluent each other. Metasoma. First and second terga with even punctures, smaller than those on mesoscutum; on first tergum denser, on second tergum with small punctures on interspaces; punctures relatively smaller and sparser only apically and laterally; punctures on third tergum larger than on second tergum; median longitudinal carina weak, on first tergum marked by a black line on the first half, on third tergum more distinct; pits of the pit row deep, black and elongate (Fig. 12 E); apical teeth long, sharp with hyaline apical tip; black spots on second sternum separated medially (Fig. 12 F), covering half sternum length. Colour. Head blue with golden-greenish area on frons, between anterior ocellus and eyes, clypeus; pronotum blue with extended golden-green area anteromedially and laterally; mesoscutum, scutellum, metanotum and propodeum red to golden-greenish; tegula, mesopleuron, legs, notauli, axillae and sutures blue; metasoma golden-green with apical margin of all tergite blue; sterna blue; wings hyaline with brown nervures. Male. Unknown.	en	Rosa, Paolo (2024): Contributions to the cuckoo wasp fauna (Hymenoptera: Chrysididae) of Yemen with description of five new species and updated checklist. Journal of Insect Biodiversity and Systematics 10 (4): 995-1031, DOI: 10.61186/jibs.10.4.995, URL: https://doi.org/10.61186/jibs.10.4.995
03AD991AFF845758FF2FD3B4C84DD070.taxon	etymology	Etymology. The specific epiteth yemenita is derived from the country name.	en	Rosa, Paolo (2024): Contributions to the cuckoo wasp fauna (Hymenoptera: Chrysididae) of Yemen with description of five new species and updated checklist. Journal of Insect Biodiversity and Systematics 10 (4): 995-1031, DOI: 10.61186/jibs.10.4.995, URL: https://doi.org/10.61186/jibs.10.4.995
03AD991AFF845758FF2FD3B4C84DD070.taxon	distribution	Distribution. Yemen (Al-Lahima)	en	Rosa, Paolo (2024): Contributions to the cuckoo wasp fauna (Hymenoptera: Chrysididae) of Yemen with description of five new species and updated checklist. Journal of Insect Biodiversity and Systematics 10 (4): 995-1031, DOI: 10.61186/jibs.10.4.995, URL: https://doi.org/10.61186/jibs.10.4.995
03AD991AFF865758FF2FD2D2CA91D369.taxon	distribution	Distribution in Yemen. Aden (du Buysson, 1896). Distribution in the Arabian Peninsula. Yemen (du Buysson, 1896) Saudi Arabia (Farasan) (Strumia & Dawah, 2019); Arabia (with no specific locality, Linsenmaier, 1968, 1994 as Chrysis (Cornuchrysis) zobeida).	en	Rosa, Paolo (2024): Contributions to the cuckoo wasp fauna (Hymenoptera: Chrysididae) of Yemen with description of five new species and updated checklist. Journal of Insect Biodiversity and Systematics 10 (4): 995-1031, DOI: 10.61186/jibs.10.4.995, URL: https://doi.org/10.61186/jibs.10.4.995
03AD991AFF865758FF2FD03CCA80D5D5.taxon	materials_examined	Material examined. 1 ♀: Yemen: Al-Lahima, 16. X. - 31. XII. 2000, Malaise trap, leg. A. van Harten & A. M. Hager, ex coll. ZMAN (RMNH); 1 ♀: Yemen, Al-Lahima, 5. VI. 2001, Malaise trap, leg. A. van Harten & A. M. Hager (RMNH). Both specimens are paratypes.	en	Rosa, Paolo (2024): Contributions to the cuckoo wasp fauna (Hymenoptera: Chrysididae) of Yemen with description of five new species and updated checklist. Journal of Insect Biodiversity and Systematics 10 (4): 995-1031, DOI: 10.61186/jibs.10.4.995, URL: https://doi.org/10.61186/jibs.10.4.995
03AD991AFF865758FF2FD03CCA80D5D5.taxon	distribution	Distribution in Yemen. Al-Lahima (Soliman et al., 2022). Distribution in the Arabian Peninsula. Yemen (Soliman et al., 2022). Saudi Arabia, Oman (Soliman et al., 2022).	en	Rosa, Paolo (2024): Contributions to the cuckoo wasp fauna (Hymenoptera: Chrysididae) of Yemen with description of five new species and updated checklist. Journal of Insect Biodiversity and Systematics 10 (4): 995-1031, DOI: 10.61186/jibs.10.4.995, URL: https://doi.org/10.61186/jibs.10.4.995
03AD991AFF865758FF2FD6F0C990D740.taxon	distribution	Distribution in Yemen. Jebel Jihaf (Linsenmaier, 1994 as Chrysis (Praestochrysis) amplifera). Distribution in the Arabian Peninsula. Yemen (Linsenmaier, 1994).	en	Rosa, Paolo (2024): Contributions to the cuckoo wasp fauna (Hymenoptera: Chrysididae) of Yemen with description of five new species and updated checklist. Journal of Insect Biodiversity and Systematics 10 (4): 995-1031, DOI: 10.61186/jibs.10.4.995, URL: https://doi.org/10.61186/jibs.10.4.995
03AD991AFF865758FF2FD462CFBBD6F2.taxon	distribution	Distribution in Yemen. Yemen (Strumia, 2008, 2014). Distribution in the Arabian Peninsula. Oman, Saudi Arabia, UAE (Linsenmaier, 1994; Strumia, 2008, 2014).	en	Rosa, Paolo (2024): Contributions to the cuckoo wasp fauna (Hymenoptera: Chrysididae) of Yemen with description of five new species and updated checklist. Journal of Insect Biodiversity and Systematics 10 (4): 995-1031, DOI: 10.61186/jibs.10.4.995, URL: https://doi.org/10.61186/jibs.10.4.995
03AD991AFF875759FFDED3B5CF2FD0EF.taxon	distribution	Distribution in Yemen. Yemen (no specific locality) (Strumia, 2008, 2014). Distribution in the Arabian Peninsula. “ Arabia deserta ” (Mocsáry, 1889); Oman (Linsenmaier, 1994 as Chrysis (Spintharina) integerrima; Strumia, 2008), Saudi Arabia (Linsenmaier, 1994 as Chrysis (Spintharina) integerrima; Strumia & Dawah, 2010, 2012, 2019), United Arab Emirates (Linsenmaier, 1994 as Chrysis (Spintharina) integerrima; Howarth & Gillett, 2008 as Chrysis integerrima; Strumia, 2014).	en	Rosa, Paolo (2024): Contributions to the cuckoo wasp fauna (Hymenoptera: Chrysididae) of Yemen with description of five new species and updated checklist. Journal of Insect Biodiversity and Systematics 10 (4): 995-1031, DOI: 10.61186/jibs.10.4.995, URL: https://doi.org/10.61186/jibs.10.4.995
03AD991AFF875759FFDED1BACB23D5D0.taxon	materials_examined	Material examined. 1 ♀, Yemen: Sana'a, ca. 7900 ft Dr. Carl Rathjens 17. - 18. IX. 1937 B. M. 1938 - 396 (NMLU).	en	Rosa, Paolo (2024): Contributions to the cuckoo wasp fauna (Hymenoptera: Chrysididae) of Yemen with description of five new species and updated checklist. Journal of Insect Biodiversity and Systematics 10 (4): 995-1031, DOI: 10.61186/jibs.10.4.995, URL: https://doi.org/10.61186/jibs.10.4.995
03AD991AFF875759FFDED1BACB23D5D0.taxon	distribution	Distribution in Yemen. Sana'a (present study). Yemen (Kirby, 1903, Socotra; Kohl, 1906, including Socotra; Linsenmaier, 1994; Madl, 2018). Distribution in the Arabian Peninsula. Yemen (Kirby, 1903, Socotra; Kohl, 1906, including Socotra; Linsenmaier, 1994; Madl, 2018). Kuwait (Walker & Pittaway, 1987; Al-Houty, 1989, 2011), Oman (Walker & Pittaway, 1987; Linsenmaier, 1994; Madl, 2018), Saudi Arabia (Walker & Pittaway, 1987; Linsenmaier, 1994; Strumia & Dawah, 2012; Madl, 2018), United Arab Emirates (Gillett & Gillett, 2005; Howarth & Gillett, 2008; Strumia, 2008, 2014; Madl, 2018), Extralimital distribution. Afrotropical, Australian, Indo-Malaysian, Palaearctic (Kimsey & Bohart, 1991; Madl, 2018).	en	Rosa, Paolo (2024): Contributions to the cuckoo wasp fauna (Hymenoptera: Chrysididae) of Yemen with description of five new species and updated checklist. Journal of Insect Biodiversity and Systematics 10 (4): 995-1031, DOI: 10.61186/jibs.10.4.995, URL: https://doi.org/10.61186/jibs.10.4.995
03AD991AFF875759FFDED7B3CEE7D71A.taxon	materials_examined	Material examined. 1 ♀, Socotra: Hadibo Plain. S. L. 19. III. 1967 K. Guichard / B. M. 1967 - 455 / Paratypes Stilbum cyanurum sokotranum Lins det. Linsenmaier 1973 / NML _ ENT GBIF _ Chr 0045804, 45805 (NMLU).	en	Rosa, Paolo (2024): Contributions to the cuckoo wasp fauna (Hymenoptera: Chrysididae) of Yemen with description of five new species and updated checklist. Journal of Insect Biodiversity and Systematics 10 (4): 995-1031, DOI: 10.61186/jibs.10.4.995, URL: https://doi.org/10.61186/jibs.10.4.995
03AD991AFF875759FFDED7B3CEE7D71A.taxon	distribution	Distribution in Yemen. Socotra (Linsenmaier, 1987, 1994). Distribution in the Arabian Peninsula. Yemen (Socotra) (Linsenmaier, 1987, 1994).	en	Rosa, Paolo (2024): Contributions to the cuckoo wasp fauna (Hymenoptera: Chrysididae) of Yemen with description of five new species and updated checklist. Journal of Insect Biodiversity and Systematics 10 (4): 995-1031, DOI: 10.61186/jibs.10.4.995, URL: https://doi.org/10.61186/jibs.10.4.995
03AD991AFF87575EFFDED5ADCEC2D6E9.taxon	materials_examined	Material examined. 1 ♀, Yemen: 12 km NW of Manakhah, 6. VII. - 21. VIII. 2002, Malaise trap, leg. A. van Harten (RMNH); 2 ♀♀, Suq Bani Mansour, Malaise trap, 28. VIII. - 14. XI. 2001, leg. A. van Harten (RMNH); 1 ♀, Al-Lahima, 9. IV. - 5. VI. 2001, Malaise trap, leg. A. van Harten (RMNH); 1 ♂, Lahj, 1. IV. - 17. V. 200, Malaise trap, leg. A. van Harten & A. Sallum (RMNH); 1 ♀, Lahj, VII. - XI. 2001, Malaise trap, leg. A. van Harten & A. Sallum (RMNH); 5 ♂♂, 4 ♀♀, Lahj, III. - V. 2002, Malaise trap, leg. A. van Harten & A. Sallum (RMNH); 1 ♂, Seiyun, 12 - 14. VIII. 2002, leg. A. van Harten, light trap (RMNH).	en	Rosa, Paolo (2024): Contributions to the cuckoo wasp fauna (Hymenoptera: Chrysididae) of Yemen with description of five new species and updated checklist. Journal of Insect Biodiversity and Systematics 10 (4): 995-1031, DOI: 10.61186/jibs.10.4.995, URL: https://doi.org/10.61186/jibs.10.4.995
03AD991AFF87575EFFDED5ADCEC2D6E9.taxon	distribution	Distribution in Yemen. Manakhah, Suq Bani Mansour, Al-Lahima, Lahj, Seiyun (present study). Distribution in the Arabian Peninsula. Yemen (Mocsáry, 1912 as Chrysis (Trichrysis) longispina; Strumia, 2009). Oman (Linsenmaier, 1994 as Chrysis (Trichrysis) longispina; Strumia, 2009), Saudi Arabia (Strumia & Dawah, 2010, 2012, 2019), United Arab Emirates (Strumia, 2014; Rosa et al., 2017).	en	Rosa, Paolo (2024): Contributions to the cuckoo wasp fauna (Hymenoptera: Chrysididae) of Yemen with description of five new species and updated checklist. Journal of Insect Biodiversity and Systematics 10 (4): 995-1031, DOI: 10.61186/jibs.10.4.995, URL: https://doi.org/10.61186/jibs.10.4.995
03AD991AFF81575FFFDED0E4CEFAD596.taxon	distribution	Distribution in Yemen. Aden (Mocsáry, 1913 as Parnopes arabs; Linsenmaier, 1994 as Parnopes (Cephaloparnops) denticulatus; Rosa et al., 2017). Distribution in the Arabian Peninsula. Yemen (Mocsáry, 1913; Rosa et al., 2017).	en	Rosa, Paolo (2024): Contributions to the cuckoo wasp fauna (Hymenoptera: Chrysididae) of Yemen with description of five new species and updated checklist. Journal of Insect Biodiversity and Systematics 10 (4): 995-1031, DOI: 10.61186/jibs.10.4.995, URL: https://doi.org/10.61186/jibs.10.4.995
03AD991AFF81575FFFDED630CE5AD775.taxon	materials_examined	Material examined. 1 ♀, Yemen: Süd Yemen Wadi Tiban coll. Linsenmaier / Type Parnopes grandior jemenensis Lins. det. Linsenmaier 1977 / NML _ ENT GBIF _ Chr 0046204 (NMLU).	en	Rosa, Paolo (2024): Contributions to the cuckoo wasp fauna (Hymenoptera: Chrysididae) of Yemen with description of five new species and updated checklist. Journal of Insect Biodiversity and Systematics 10 (4): 995-1031, DOI: 10.61186/jibs.10.4.995, URL: https://doi.org/10.61186/jibs.10.4.995
03AD991AFF81575FFFDED630CE5AD775.taxon	distribution	Distribution in Yemen. Wadi Tiban (Linsenmaier, 1987, 1994). Distribution in the Arabian Peninsula. Yemen (Linsenmaier, 1987, 1994).	en	Rosa, Paolo (2024): Contributions to the cuckoo wasp fauna (Hymenoptera: Chrysididae) of Yemen with description of five new species and updated checklist. Journal of Insect Biodiversity and Systematics 10 (4): 995-1031, DOI: 10.61186/jibs.10.4.995, URL: https://doi.org/10.61186/jibs.10.4.995
03AD991AFF81575FFFDED29FCBC6D211.taxon	distribution	Distribution in Yemen. Aden (Linsenmaier, 1994 as Chrysis (Trichrysis) scioensis); no specific locality (Madl, 2018). Distribution in the Arabian Peninsula. Yemen (Linsenmaier, 1994; Madl, 2018). Oman (Strumia, 2009; Madl, 2018), Saudi Arabia (Strumia & Dawah, 2010; Madl, 2018). Extralimital distribution. Cameroon, Djibouti, Egypt, Ethiopia, Tanzania to South Africa (Kimsey & Bohart, 1991; Madl, 2018).	en	Rosa, Paolo (2024): Contributions to the cuckoo wasp fauna (Hymenoptera: Chrysididae) of Yemen with description of five new species and updated checklist. Journal of Insect Biodiversity and Systematics 10 (4): 995-1031, DOI: 10.61186/jibs.10.4.995, URL: https://doi.org/10.61186/jibs.10.4.995
