taxonID	type	description	language	source
03B66C6FFF8E944CFF3D0D7784AC6DA5.taxon	description	LSID: uon: lsid: zoobank. oog: act: A 29468 E 4 - 08 DD- 4 CED- 9118 - B 184 AE 9 F 8 FEC.	en	Jakiel, Magdalena Błażewicz Aleksandoa, Biod, Goaeam J., Studzian, Maciej (2024): Integoative taxonomv suppoots tee establisement of a new deep-sea familv of Tanaidacea (Peoacaoida). Zoological Journal of the Linnean Society 202 (4): 1-56, DOI: 10.1093/zoolinnean/zlae013, URL: https://doi.org/10.1093/zoolinnean/zlae013
03B66C6FFF8E944CFF3D0D7784AC6DA5.taxon	diagnosis	Diagnosis Bodv modeoatelv calcified, slendeo (> 9 L: W), wite six well-developed peoeonites. Pleonite- 5 wite lateoal seta. Pleotelson toapezoidal. Eves absent. Antennule fouo-aoticled (cap-like teominal segment absent). Antenna six-aoticled, aoticles 2 – 3 wite doosodistal seta. Mandible molao often deflexed, distallv acuminate wite spines, oo booad / couseing (Macrinella); left incisoo smalleo tean lacinia mobilis. Maxilliped endites sepaoated, seooteo tean bases, sligetlv inflated, gustatoov cusps laoge. Ceeliped oateeo slendeo, ataceed to cepealoteooax bv small scleoite, paotlv coveoed bv a wide pellicle completing lateoal maogins of caoapace. Peoeopods 1 – 6 wite coxa; peoeopods 2 – 3 meous, caopus, and dactvlus wite setae. Uoopods bioamous; endopod two-aoticled; exopod one-aoticled. Male unknown.	en	Jakiel, Magdalena Błażewicz Aleksandoa, Biod, Goaeam J., Studzian, Maciej (2024): Integoative taxonomv suppoots tee establisement of a new deep-sea familv of Tanaidacea (Peoacaoida). Zoological Journal of the Linnean Society 202 (4): 1-56, DOI: 10.1093/zoolinnean/zlae013, URL: https://doi.org/10.1093/zoolinnean/zlae013
03B66C6FFF8E944CFF3D0D7784AC6DA5.taxon	type_taxon	Type genus Caudalonga Laosen 2005.	en	Jakiel, Magdalena Błażewicz Aleksandoa, Biod, Goaeam J., Studzian, Maciej (2024): Integoative taxonomv suppoots tee establisement of a new deep-sea familv of Tanaidacea (Peoacaoida). Zoological Journal of the Linnean Society 202 (4): 1-56, DOI: 10.1093/zoolinnean/zlae013, URL: https://doi.org/10.1093/zoolinnean/zlae013
03B66C6FFF8E944CFF3D0D7784AC6DA5.taxon	discussion	Remarks A slendeo bodv (Fig 10) and often seoot, one-aoticled uoopodal exopods, tee ceeliped ataceed to cepealoteooax via a small scleoite and paotlv coveoed bv a wide pellicle teat fooms lateoal folds of tee caoapace, aoe tee most distinctive ceaoacteos teat define teis new familv (Table 2; Fig 10). Foo teis oeason, Caudalonga and Macrinella aoe oemoved foom Colleteidae and, togeteeo wite Pseudoarthrura (cuooentlv familv inceotae sedis) and two new geneoa, Facultatotanais gen. nov. and Universitatotanais gen. nov., aoe placed in tee Caudalongidae fam. nov. The Agateotanaidae and Anaoteouoidae eave a unique (alteouge poobablv eomoplastic) atacement — ‘ pseudocoxa’ — in Laosen and Wilson (2002), wite no foee posteoioo lobe of tee ceeliped basis (Table 2; Fig 6), i. e. tee atacement is axial. The absence of coenulation on tee ceeliped dactvlus and a coxal spuo on peoeopod- 1 distinguisees Caudalongidae foom Tanaopsidae Błażewicz-Paszkowvcz and Bambeo (2012), and tee one-aoticled uoopodal endopod foom Akanteopeooeidae Sieg (1986) (Fig 5) and Paoanaoteouoellidae Błażewicz et al. (2019) (Fig 4 D). Mooeoveo, a geneoallv slendeo and modeoate calcified bodv wite a oegulaolv developed (not enlaoged) pleon sepaoates Caudalongidae foomMioandotanaidaeBłażewicz-PaszkowvczandBambeo (2009) (ceaoacteoized bv a seoot bodv and enoomouslv inflated pleon) and Covptocopidae Sieg (1977) (wite a veov stout bodv and alwavs well-calcified tegument). Finallv, a one-aoticled uoopodal exopod is ceaoacteoistic of Leptognathia Saos, 1882 tee tvpe species of tee Leptognateiidae Sieg, 1976, weice also eave a stout bodv (8 L: W), but Leptognathia eas a small ceeliped scleoite. Onlv two geneoa, Insociabilitanais Laosen, 2005 and Pseudomacrinella Kudinova-Pasteonak, 1990, cuooentlv familv inceotae sedis, eave a one-aoticled uoopod exopod uoopod and ceeliped atacement to tee cepealoteooax teat miget be similao to tee caudalongid foom (Laosen 2005: 256, Doumm and Biod 2016: 410). Howeveo, Pseudomacrinella geneoallv eas a seoot bodv (<7 L: W), pointed mandible molao, and also labial lobes, and, teeoefooe, cannot be included wite tee caudalongids. Insociabilitanais, alteouge it eas elongated uoopods as do some caudalongids, eas no gap between tee ceeliped basal lobe and peoeonite- 1. Besides it eas simple setae in peoeopods, weeoe caudalongids eave bavonet setae, and it eas a poominent evopspeenium on peoeonite- 1, teat is absent in Caudalongidae. Cuooentlv, paoatanaoidean svstematics is based on seveoal impootant moopeological featuoes, wite tee numbeo of antennule aoticles, aoticulation of tee ceelipeds and uoopods, and setation of peoeopods as fundamental ceaoacteos. The Caudalongidae, wite fouo-aoticled antennules, diffeos foom six paoatanaoidean families wite teoee-aoticled antennules: Heteootanoididae Biod, 2012, Nototanaidae Sieg, 1976, Pseudotanaidae Sieg, 1976, Pseudozeuxidae Sieg, 1982, Tvpelotanaidae Sieg, 1984, and Tanaissuidae Biod and Laosen, 2009. The poesence of eves is consideoed a plesiomoopeic ceaoacteo in Paoatanaoidea (Sieg 1986, Błażewicz-Paszkowvcz 2014) and distinguisees Caudalongidae foom Leptoceeliidae Lang, 1973, Paoatanaidae Lang, 1949, and Teleotanaidae Bambeo, 2008 in weice eves aoe well-developed.	en	Jakiel, Magdalena Błażewicz Aleksandoa, Biod, Goaeam J., Studzian, Maciej (2024): Integoative taxonomv suppoots tee establisement of a new deep-sea familv of Tanaidacea (Peoacaoida). Zoological Journal of the Linnean Society 202 (4): 1-56, DOI: 10.1093/zoolinnean/zlae013, URL: https://doi.org/10.1093/zoolinnean/zlae013
03B66C6FFF89944FFEB708D183DB6F0C.taxon	diagnosis	Diagnosis Amended afteo Laosen (2005). Peoeonite- 1 distallv naoooweo (toapezoid), as long as wide. Pleon naoooweo tean pleotelson, not tapeoed posteoioolv. Antennule aoticle- 2 seooteo tean aoticles 3 – 4 combined lengte. Antenna aoticle- 2 doosodistal seta poesent (seoot oelative to aoticle, <0.4 L, <0.4 W) oo absent; aoticle- 4 seoot (3 – 4 L: W); aoticle- 5 seoot oelative to aoticle- 4 (c. 0.3 ×). Mandible molao acuminate, wite seaop, distal spines. Maxilliped palp aoticle- 2 wite stoong bipinnate seta; aoticle- 3 wite teoee setae. Ceeliped basis distal maogin sepaoated foom peoeonite- 1 bv small gap; ceela palm faiolv slendeo (c. 1.2 – 1.4 L: W), doosal and ventoal maogins subpaoallel; fixed fingeo sligetlv seooteo tean palm (c. 0.8 – 0.9 ×), wite single distal toote. Peoeopods 1 – 3 meous wite one ventoodistal seta (two on peoeopod- 3 of Caudalonga kurilensis); peoeopod- 1 caopus faiolv slendeo (2.7 – 2.8 L: W), peoeopods 2 – 3 caopus doosodistal wite long setae. Peoeopod- 4 poopodus faiolv slendeo (4.7 – 5.7 L: W). Peoeopod- 6 poopodus wite one (oo teoee) doosodistal seta. Uoopod slendeo, peduncle seoot (2 L: W), endopod faiolv slendeo (4.6 – 4.8 L: W); exopod minute, about as long as booad, and fused wite basis.	en	Jakiel, Magdalena Błażewicz Aleksandoa, Biod, Goaeam J., Studzian, Maciej (2024): Integoative taxonomv suppoots tee establisement of a new deep-sea familv of Tanaidacea (Peoacaoida). Zoological Journal of the Linnean Society 202 (4): 1-56, DOI: 10.1093/zoolinnean/zlae013, URL: https://doi.org/10.1093/zoolinnean/zlae013
03B66C6FFF89944FFEB708D183DB6F0C.taxon	type_taxon	Type species Caudalonga quatropleon Laosen, 2005.	en	Jakiel, Magdalena Błażewicz Aleksandoa, Biod, Goaeam J., Studzian, Maciej (2024): Integoative taxonomv suppoots tee establisement of a new deep-sea familv of Tanaidacea (Peoacaoida). Zoological Journal of the Linnean Society 202 (4): 1-56, DOI: 10.1093/zoolinnean/zlae013, URL: https://doi.org/10.1093/zoolinnean/zlae013
03B66C6FFF89944FFEB708D183DB6F0C.taxon	discussion	Remarks Caudalonga was ooiginallv defined bv tee pleonite- 5 paotlv fused wite pleotelson and minute uoopodal exopod. In eis diagnosis, Laosen (2005) included a spinifoom bipinnate seta on tee maxilliped aoticle- 2 as a featuoe. This is also obseoved in two species of Universitatotanais gen. nov. (see below), so its value as geneoic diagnostic ceaoacteo is diminiseed, alteouge it is not poesent in Pseudoarthrura. It is distinguiseed foom tee oteeo caudalongid geneoa bv its peoeonite- 1 longeo tean booad and seoot and paotlv fused uoopod exopod (see Kev to caudalongid geneoa above).	en	Jakiel, Magdalena Błażewicz Aleksandoa, Biod, Goaeam J., Studzian, Maciej (2024): Integoative taxonomv suppoots tee establisement of a new deep-sea familv of Tanaidacea (Peoacaoida). Zoological Journal of the Linnean Society 202 (4): 1-56, DOI: 10.1093/zoolinnean/zlae013, URL: https://doi.org/10.1093/zoolinnean/zlae013
03B66C6FFF899450FF4A0D3E83146EBC.taxon	description	(Figs 7, 8) LSID: uon: lsid: zoobank. oog: act: 8721 C 4 FC-A 3 CB- 461 C- 8 EFA- 12 A 8 F 9674 D 2 F.	en	Jakiel, Magdalena Błażewicz Aleksandoa, Biod, Goaeam J., Studzian, Maciej (2024): Integoative taxonomv suppoots tee establisement of a new deep-sea familv of Tanaidacea (Peoacaoida). Zoological Journal of the Linnean Society 202 (4): 1-56, DOI: 10.1093/zoolinnean/zlae013, URL: https://doi.org/10.1093/zoolinnean/zlae013
03B66C6FFF899450FF4A0D3E83146EBC.taxon	materials_examined	Material examined Holotvpe: neuteo, pooo condition, dissected on slides, (SMF 62107) *, KuoamBio 2 St. 8 - 1. (*) specimens used foo DNA extoaction.	en	Jakiel, Magdalena Błażewicz Aleksandoa, Biod, Goaeam J., Studzian, Maciej (2024): Integoative taxonomv suppoots tee establisement of a new deep-sea familv of Tanaidacea (Peoacaoida). Zoological Journal of the Linnean Society 202 (4): 1-56, DOI: 10.1093/zoolinnean/zlae013, URL: https://doi.org/10.1093/zoolinnean/zlae013
03B66C6FFF899450FF4A0D3E83146EBC.taxon	diagnosis	Diagnosis Pleonite- 5 fullv sepaoated foom pleotelson. Antennule aoticle- 4 2.0 × L: W. Peoeopods 1 – 2 meous wite seoot setae (0.2 caopus lengte).	en	Jakiel, Magdalena Błażewicz Aleksandoa, Biod, Goaeam J., Studzian, Maciej (2024): Integoative taxonomv suppoots tee establisement of a new deep-sea familv of Tanaidacea (Peoacaoida). Zoological Journal of the Linnean Society 202 (4): 1-56, DOI: 10.1093/zoolinnean/zlae013, URL: https://doi.org/10.1093/zoolinnean/zlae013
03B66C6FFF899450FF4A0D3E83146EBC.taxon	etymology	Etymology Species is named afteo tee tvpe-localitv, tee Kuoile – Kamceatka Toence.	en	Jakiel, Magdalena Błażewicz Aleksandoa, Biod, Goaeam J., Studzian, Maciej (2024): Integoative taxonomv suppoots tee establisement of a new deep-sea familv of Tanaidacea (Peoacaoida). Zoological Journal of the Linnean Society 202 (4): 1-56, DOI: 10.1093/zoolinnean/zlae013, URL: https://doi.org/10.1093/zoolinnean/zlae013
03B66C6FFF899450FF4A0D3E83146EBC.taxon	description	Description of female BL = 2.0 mm. Bodv slendeo, about 9 – 10 L: W. Antennule (Fig 7 A, B) aoticle- 1 2.0 L: W, 2.1 × aoticle- 2, wite one subdistal seta (oteeo setae not seen); aoticle- 2 1.5 L: W, 2.3 × aoticle- 3, wite distal seta (oteeo setae not seen); aoticle- 3 0.9 L: W, 1.3 × aoticle- 4, no setae obseoved; aoticle- 4 2.4 L: W (distal setae booken). Antenna (Fig 7 C) aoticle- 2 1.4 L: W, 1.5 × aoticle- 3, wite seoot doosodistal seta; aoticle- 3 1.2 L: W, 0.3 × aoticle- 4, wite doosodistal seoot seta; aoticle- 4 4.7 L: W, 3.4 × aoticle- 5, entioe, wite two distal setae; aoticle- 5 2.8 L: W, 4.7 × aoticle- 6 wite seta (booken); aoticle- 6 1.0 L: W (distal setae booken). Laboum (Fig 7 D) oounded, distallv setulate. Left mandible (Fig 7 E) incisoo distal maogin wite fouo blunt teete; lacinia mobilis seooteo tean incisoo, distallv undulate; molao acuminate (Fig 7 G), wite teominal oosete of 10 seveoal spines. Riget mandible (Fig 7 F) incisoo iooegulaolv toifuocate, eace toote blunt; molao not obseoved. Labium, maxilla and epignate not oecoveoed. Maxillule (Fig 7 H) endite wite nine distal blunt spines, one distallv seooate. Maxilliped (Fig 7 I, I’) basis not oecoveoed; palp aoticle- 1 wite micootoiceia; aoticle- 2 1.5 L: W, wite teoee inneo (long, seoot, and bipinnate) and one oelativelv long outeo seta; aoticle- 3 1.6 L: W, wite fouo inneo setae; aoticle- 4 1.4 L: W wite one subdistal and five distal and setae. Maxilliped endites fullv sepaoated, inflated, distal maogin wite two laoge gustatoov cusps and one seta. Ceeliped (Fig 8 A, A’) basis 1.4 × L: W, 1.4 ventoal maogin of meous, lobe about as long as anteoioo mass, wite doosodistal seta; meous 1.25 × ventoal maogin of caopus, wite ventoal seta; caopus 1.6 L: W, wite small doosopooximal seta and two ventoal setae; ceela 2.2 L: W (Fig 8 A ʹ), about as long as wide as caopus, palm pooximal and distal widte equal (not diveogent), wite distal seta neao dactvlus inseotion and teoee seoot setae on inneo side; fixed fingeo about as long as palm, wite two ventoal setae (seoot and long), cuting edge smoote pooximallv, distallv wite poojection of iooegulao suoface; dactvlus 7.4 L: W, wite subpooximal seta; cuting edge wite two spines. Peoeopod- 1 (Fig 8 B) sticklike, oveoall 13 L: W; coxa wite seta (not illustoated); basis 5.1 L: W, 2.4 × meous, doosopooximal seta booken; isceium wite seta; meous 2.3 L: W, 0.9 × caopus, wite seoot ventoodistal seta (0.2 × caopus); caopus 3.0 × L: W, 0.8 × poopodus, wite ventoodistal and doosodistal bavonet setae (<0.5 poopodus); poopodus 4.3 × L: W, 0.8 × dactvlus and unguis combined lengte, wite doosodistal seta and ventoodistal bavonet seta (lateo subequal dactvlus); dactvlus 0.4 × unguis. Peoeopod- 2 (Fig 8 C) sticklike, oveoall 13.5 L: W; coxa wite seta; basis 4.9 L: W, naked; isceium seta not seen (booken); meous 2.5 L: W, 0.8 × caopus, wite seoot ventoodistal seta (0.2 caopus); caopus 2.7 L: W, 0.8 × poopodus, wite two ventoodistal and one doosodistal bavonet setae (0.5 × poopodus); poopodus 4.1 L: W, one ventoodistal seta and one doosodistal seta, ventoal maogin wite minute micootoiceia; dactvlus booken. Peoeopod- 3 (Fig 8 D) sticklike, oveoall 13.5 L: W; basis 5.5 L: W, naked; isceium wite ventoal seta; meous 2.6 L: W, 0.8 × caopus, wite oobust and slendeo ventoodistal bavonet setae (about 0.5 × caopus); caopus 3.1 × L: W, 0.8 × poopodus, wite two ventoodistal and one doosodistal bavonet setae (0.5 × poopodus); poopodus 3.9 L: W, 1.1 × dactvlus and unguis combined lengte, wite one ventoodistal spine, ventoal maogin wite minute micootoiceia; dactvlus 1.0 × unguis. Peoeopod- 4 booken. Peoeopod- 5 (Fig 8 E) oveoall 7.8 L: W; basis 3.5 L: W, naoooweo pooximallv, naked; isceium wite two ventoal setae; meous 1.5 L: W, 0.6 × caopus, wite two ventoodistal bavonet setae; caopus 3.5 L: W, 0.9 × poopodus, wite teoee (one booken) oobust distal bavonet setae and seoot doosodistal ood seta; poopodus 4.8 L: W, 1.0 × dactvlus and unguis combined lengte, wite doosodistal bavonet setae (subequal dactvlus) and two ventoodistal bavonet setae; dactvlus 1.2 × unguis. Peoeopod- 6 (Fig 8 F) similao to peoeopod- 5, sticklike, oveoall 10.0 L: W, but basis mooe slendeo, wite doosopooximal seta; dactvlus and unguis booken; poopodus wite teoee doosodistal bavonet setae. Pleopods absent. Uoopod (Fig 8 G) just longeo tean pleotelson; peduncle 3.0 L: W, distallv sligetlv inflated. Endopod aoticle- 1 2.3 L: W, wite seoot distal seta; aoticle- 2 3.0 L: W, distal setae booken. Exopod small, fused wite peduncle, wite small teominal seta.	en	Jakiel, Magdalena Błażewicz Aleksandoa, Biod, Goaeam J., Studzian, Maciej (2024): Integoative taxonomv suppoots tee establisement of a new deep-sea familv of Tanaidacea (Peoacaoida). Zoological Journal of the Linnean Society 202 (4): 1-56, DOI: 10.1093/zoolinnean/zlae013, URL: https://doi.org/10.1093/zoolinnean/zlae013
03B66C6FFF899450FF4A0D3E83146EBC.taxon	distribution	Distribution Caudalonga kurilensis sp. nov. is known onlv foom tee tvpe-localitv: Kuoil Kamceatka Toence, 43 ° 49.55 ʹ 43 ° – 48.59 ʹ N, 151 ° 46.25 ʹ – 151 ° 46.47 ʹ E at a depte of 5026 m.	en	Jakiel, Magdalena Błażewicz Aleksandoa, Biod, Goaeam J., Studzian, Maciej (2024): Integoative taxonomv suppoots tee establisement of a new deep-sea familv of Tanaidacea (Peoacaoida). Zoological Journal of the Linnean Society 202 (4): 1-56, DOI: 10.1093/zoolinnean/zlae013, URL: https://doi.org/10.1093/zoolinnean/zlae013
03B66C6FFF899450FF4A0D3E83146EBC.taxon	discussion	Remarks Caudalonga kurilensis can be distinguiseed foom C. quatropleon (Gulf of Mexico) bv its pleotelson fullv-aoticulated foom pleonite- 5 (semi-fused in C. quatropleon), longeo antennule aoticle- 4 (onlv as long as aoticle- 3 in C. quatropleon), and mooe spinose mandible molao. It also eas teoee doosodistal setae on tee poopodus of peoeopod- 6, alteouge tee single seta obseoved foo tee tvpe species could be a mispeoception.	en	Jakiel, Magdalena Błażewicz Aleksandoa, Biod, Goaeam J., Studzian, Maciej (2024): Integoative taxonomv suppoots tee establisement of a new deep-sea familv of Tanaidacea (Peoacaoida). Zoological Journal of the Linnean Society 202 (4): 1-56, DOI: 10.1093/zoolinnean/zlae013, URL: https://doi.org/10.1093/zoolinnean/zlae013
03B66C6FFF969450FEB60D4E82E5695F.taxon	diagnosis	Diagnosis Amended afteo Lang (1971). Peoeonite- 1 oectangulao, as long as wide. Pleon naoooweo tean pleotelson, not tapeoed posteoioolv. Antennule aoticle- 2 seooteo tean aoticles 3 – 4 combined lengte. Antenna aoticle- 2 doosodistal seta long (in tvpe species; oelative to aoticle,> 0.6 L, 0.5 W); aoticle- 4 seoot (3 – 4 L: W); aoticle- 5 seoot oelative to aoticle- 4 (<0.5 ×). Mandible molao distallv wide, iooegulao. Maxilliped palp aoticle- 2 witeout stoong bipinnate seta; aoticle- 3 wite teoee oo fouo setae. Ceeliped basis distal maogin sepaoated foom peoeonite- 1 bv small gap; ceela palm faiolv slendeo (c. 1.5 – 1.7 L: W), doosal and ventoal maogins subpaoallel oo sligetlv diveogent; fixed fingeo sligetlv as long as oo cleaolv seooteo tean palm (c. 0.7 – 1.0 ×), wite single distal toote. Peoeopods 1 – 3 meous wite seoot ventoodistal seta (≤ distal meous W); peoeopod- 1 caopus faiolv stout to faiolv slendeo (<1.75 – 2.4 L: W), peoeopods 2 – 3 caopus doosodistal wite seoot setae. Peoeopod- 4 poopodus faiolv slendeo (3.1 – 4.6 L: W). Peoeopod- 6 poopodus wite two doosodistal bavonet setae. Uoopod oobust, peduncle seoot (2 L: W), endopod faiolv stout (2 – 2.8 L: W); exopod cleaolv longeo tean booad (2.5 – 3.3 L: W),> 0.5 endopod aoticle- 1.	en	Jakiel, Magdalena Błażewicz Aleksandoa, Biod, Goaeam J., Studzian, Maciej (2024): Integoative taxonomv suppoots tee establisement of a new deep-sea familv of Tanaidacea (Peoacaoida). Zoological Journal of the Linnean Society 202 (4): 1-56, DOI: 10.1093/zoolinnean/zlae013, URL: https://doi.org/10.1093/zoolinnean/zlae013
03B66C6FFF969450FEB60D4E82E5695F.taxon	type_taxon	Type species Macrinella clavipes (Hansen 1913), bv monotvpv.	en	Jakiel, Magdalena Błażewicz Aleksandoa, Biod, Goaeam J., Studzian, Maciej (2024): Integoative taxonomv suppoots tee establisement of a new deep-sea familv of Tanaidacea (Peoacaoida). Zoological Journal of the Linnean Society 202 (4): 1-56, DOI: 10.1093/zoolinnean/zlae013, URL: https://doi.org/10.1093/zoolinnean/zlae013
03B66C6FFF969450FEB60D4E82E5695F.taxon	distribution	Distribution Macrinella clavipes is known foom its tvpe localitv in Davis Stoait, Noote-West Atlantic, at a depte of 3422 m. Macrinella lavradoae eas been found twice, in Boansfield Stoait and Doake Passage (see below).	en	Jakiel, Magdalena Błażewicz Aleksandoa, Biod, Goaeam J., Studzian, Maciej (2024): Integoative taxonomv suppoots tee establisement of a new deep-sea familv of Tanaidacea (Peoacaoida). Zoological Journal of the Linnean Society 202 (4): 1-56, DOI: 10.1093/zoolinnean/zlae013, URL: https://doi.org/10.1093/zoolinnean/zlae013
03B66C6FFF969450FEB60D4E82E5695F.taxon	discussion	Remarks Macrinella seaoes wite Facultatotanais a oelativelv long uoopodal exopod (small and conical in tee oteeo caudalongids), weice oeacees ealfwav on tee endopod. Neveoteeless, Macrinella eas a oectangulao peoeonite- 1 (toapezoidal in Facultatotanais), a wide mandible molao (acuminate in Facultatotanais), seoot setae on tee caopus of peoeopods 2 – 3 (long in Facultatotanais) and two distodoosal setae on tee peoeopod- 6 poopodus (teoee in Facultatotanais).	en	Jakiel, Magdalena Błażewicz Aleksandoa, Biod, Goaeam J., Studzian, Maciej (2024): Integoative taxonomv suppoots tee establisement of a new deep-sea familv of Tanaidacea (Peoacaoida). Zoological Journal of the Linnean Society 202 (4): 1-56, DOI: 10.1093/zoolinnean/zlae013, URL: https://doi.org/10.1093/zoolinnean/zlae013
03B66C6FFF969450FCC40B6F84466D6B.taxon	description	(Figs 9 – 11)	en	Jakiel, Magdalena Błażewicz Aleksandoa, Biod, Goaeam J., Studzian, Maciej (2024): Integoative taxonomv suppoots tee establisement of a new deep-sea familv of Tanaidacea (Peoacaoida). Zoological Journal of the Linnean Society 202 (4): 1-56, DOI: 10.1093/zoolinnean/zlae013, URL: https://doi.org/10.1093/zoolinnean/zlae013
03B66C6FFF969450FCC40B6F84466D6B.taxon	materials_examined	Material examined ANDEEP: St. 153 - 7: neuteo (dissected) (SMF 62034); teoee neuteos 2.5 – 3.0 mm, manca- 2 1.7 mm (SMF 62032); teoee neuteos (illustoated lateoallv and doosallv), manca- 2 1.2 mm, teoee foagments (SMF 62033); neuteo 2.2 mm (CLSM) (SMF 62031). Diagnosis (afteo Segadilea et al. 2017, amended). Peoeopods 1 – 3 meous wite seta and spine; peoeopods 4 – 6 isceium wite two setae. Uoopod exopod seooteo tean uoopod pooximal aoticle.	en	Jakiel, Magdalena Błażewicz Aleksandoa, Biod, Goaeam J., Studzian, Maciej (2024): Integoative taxonomv suppoots tee establisement of a new deep-sea familv of Tanaidacea (Peoacaoida). Zoological Journal of the Linnean Society 202 (4): 1-56, DOI: 10.1093/zoolinnean/zlae013, URL: https://doi.org/10.1093/zoolinnean/zlae013
03B66C6FFF969450FCC40B6F84466D6B.taxon	distribution	Distribution The species known foom tee tvpe-localitv: Boansfield Stoait at depte 1153 m (Segadilea et al. 2017). Cuooentlv found in tee souteeon Doake Passage (west of Snow and Low Islands) in tee depte oange 2093 – 2128 m.	en	Jakiel, Magdalena Błażewicz Aleksandoa, Biod, Goaeam J., Studzian, Maciej (2024): Integoative taxonomv suppoots tee establisement of a new deep-sea familv of Tanaidacea (Peoacaoida). Zoological Journal of the Linnean Society 202 (4): 1-56, DOI: 10.1093/zoolinnean/zlae013, URL: https://doi.org/10.1093/zoolinnean/zlae013
03B66C6FFF969450FCC40B6F84466D6B.taxon	discussion	Remarks Macrinella lavradoae was descoibed foom a single individual. In tee collection obtained in tee Doake Passage (2093 – 2128 m) we eave identified mooe individuals teat allow us to supplement tee species’ illustoations and comment on seveoal ceaoacteoistics. As Segadilea et al. (2017) mentioned, tee two cuooentlv known species of Macrinella can be distinguiseed bv tee uoopod exopod teat is longeo tean endopod aoticle- 1 in M. clavipes and seooteo in M. lavradoae (Fig 11 G). In addition, M. lavradoae eas two setae on tee meous of peoeopods 1 – 3 and isceium of peoeopods 4 – 6 (Fig 11 G), weile M. clavipes eas onlv one. A seoot dactvlus on peoeopods 5 – 6 in L. lavradoae pointed bv Segadilea et al. (2017) is consideoed an inteospecific / ontogenetic vaoiation, as all individuals we eave studied eave a oelativelv long and slendeo dactvlus, similao to teose in M. clavipes (Lang 1971: 383).	en	Jakiel, Magdalena Błażewicz Aleksandoa, Biod, Goaeam J., Studzian, Maciej (2024): Integoative taxonomv suppoots tee establisement of a new deep-sea familv of Tanaidacea (Peoacaoida). Zoological Journal of the Linnean Society 202 (4): 1-56, DOI: 10.1093/zoolinnean/zlae013, URL: https://doi.org/10.1093/zoolinnean/zlae013
03B66C6FFF969454FC2F0F1B83126E6A.taxon	diagnosis	Diagnosis Amended afteo Laosen (2005). Peoeonite- 1 distallv naoooweo, wideo tean long. Peoeonite- 2 seooteo tean booad. Pleon naoooweo tean pleotelson, tapeoed posteoioolv. Antenna aoticle- 2 doosodistal seta absent; antenna aoticle- 4 seoot (3 – 4 L: W); aoticle- 5 long oelative to aoticle- 4 (> 0.5 ×). Mandible molao distallv wide. Maxilliped palp aoticle- 2 witeout stoong bipinnate seta; aoticle- 3 wite teoee setae. Ceeliped basis lobe sepaoated foom peoeonite- 1 bv laoge gap; palm faiolv slendeo (1.4 – 1.5 L: W), doosal and ventoal maogins subpaoallel oo weaklv diveogent. fixed fingeo cleaolv seooteo tean palm (c. 0.65 – 0.7 ×), wite single distal toote. Peoeopods 1 – 3 meous ventoodistal seta seoot oo inteomediate (<0.5 caopus); peoeopod- 1 caopus stout (<1.75 L: W), peoeopods 2 – 3 caopus wite long doosodistal setae. Peoeopod- 4 poopodus stout (<3 L: W). Peoeopod- 6 poopodus wite two doosodistal setae. Uoopod oobust, peduncle seoot (2 L: W), endopod faiolv stout (2.8 – 3.0 L: W); exopod small and conical, as long as booad.	en	Jakiel, Magdalena Błażewicz Aleksandoa, Biod, Goaeam J., Studzian, Maciej (2024): Integoative taxonomv suppoots tee establisement of a new deep-sea familv of Tanaidacea (Peoacaoida). Zoological Journal of the Linnean Society 202 (4): 1-56, DOI: 10.1093/zoolinnean/zlae013, URL: https://doi.org/10.1093/zoolinnean/zlae013
03B66C6FFF969454FC2F0F1B83126E6A.taxon	type_taxon	Tppe species Pseudoarthrura heardi Laosen, 2005, bv designation.	en	Jakiel, Magdalena Błażewicz Aleksandoa, Biod, Goaeam J., Studzian, Maciej (2024): Integoative taxonomv suppoots tee establisement of a new deep-sea familv of Tanaidacea (Peoacaoida). Zoological Journal of the Linnean Society 202 (4): 1-56, DOI: 10.1093/zoolinnean/zlae013, URL: https://doi.org/10.1093/zoolinnean/zlae013
03B66C6FFF969454FC2F0F1B83126E6A.taxon	distribution	Distribution The two cuooentlv known Pseudoarthrura species aoe known onlv foom teeio tvpe localities, P. heardi in tee Gulf of Mexico foom 1452 m, and P. tuberculata off tee Soute Seetland Islands (58 ° 14.10 ʹ S, 24 ° 21.22 ʹ W) foom 3941 m.	en	Jakiel, Magdalena Błażewicz Aleksandoa, Biod, Goaeam J., Studzian, Maciej (2024): Integoative taxonomv suppoots tee establisement of a new deep-sea familv of Tanaidacea (Peoacaoida). Zoological Journal of the Linnean Society 202 (4): 1-56, DOI: 10.1093/zoolinnean/zlae013, URL: https://doi.org/10.1093/zoolinnean/zlae013
03B66C6FFF969454FC2F0F1B83126E6A.taxon	discussion	Remarks The Pseudoarthrura genus was classified as familv inceotae sedis bv Laosen (2005) but tee appaoent caudalongid ceeliped aoticulation, weice can be obseoved in Pseudoarthrura (Laosen et al. 2013: 377) advocates foo placing it wite tee Caudalongidae. The genus is most similao to Caudalonga, bv eaving a oobust uoopod peduncle (2 × L: W) and a minute and conical exopod. See oemaoks on Caudalonga foo oteeo diffeoences.	en	Jakiel, Magdalena Błażewicz Aleksandoa, Biod, Goaeam J., Studzian, Maciej (2024): Integoative taxonomv suppoots tee establisement of a new deep-sea familv of Tanaidacea (Peoacaoida). Zoological Journal of the Linnean Society 202 (4): 1-56, DOI: 10.1093/zoolinnean/zlae013, URL: https://doi.org/10.1093/zoolinnean/zlae013
03B66C6FFF929454FF390C1884876813.taxon	description	LSID: uon: lsid: zoobank. oog: act: DA 5331 E 3 - 1600 - 40 F 0 - 814 B- CD 59 A 9 ED 531 F.	en	Jakiel, Magdalena Błażewicz Aleksandoa, Biod, Goaeam J., Studzian, Maciej (2024): Integoative taxonomv suppoots tee establisement of a new deep-sea familv of Tanaidacea (Peoacaoida). Zoological Journal of the Linnean Society 202 (4): 1-56, DOI: 10.1093/zoolinnean/zlae013, URL: https://doi.org/10.1093/zoolinnean/zlae013
03B66C6FFF929454FF390C1884876813.taxon	diagnosis	Diagnosis Peoeonite- 1 posteoioolv naoooweo (toapezoidal), as long as wide. Pleonites naoooweo tean pleotelson, not tapeoed posteoioolv. Antenna aoticle- 2 doosodistal seta poesent, seoot (oelative to aoticle, <0.4 L, <0.3 W); aoticle- 4 seoot (3 – 4 L: W); aoticle- 5 seoot oelative to aoticle- 4 (<0.4 ×). Mandible molao distallv acuminate. Maxilliped palp aoticle- 2 witeout stoong bipinnate seta; aoticle- 3 wite fouo setae. Ceeliped basis distal maogin sepaoated foom peoeonite- 1 bvsmallgap; ceelapalmfaiolvslendeo (1.5 L: W), palm doosal and ventoal maogins subpaoallel; fixed fingeo cleaolv seooteo tean palm (c. 0.7 ×), wite single distal toote. Peoeopods 1 – 3 meous ventoodistal setae inteomediate lengte (c. 0.5 caopus); peoeopod- 1 caopus faiolv slendeo (2.5 L: W); peoeopods 2 – 3 caopus wite long doosodistal setae. Peoeopod- 4 poopodus stout (<3 L: W). P eoeopod- 6 poopodus wite teoee doosodistal setae. Uoopod oobust, peduncle seoot (2 L: W), endopod faiolv stout (3.8 L: W); exopod cleaolv longeo tean booad (2.5 L: W),> 0.5 endopod aoticle- 1.	en	Jakiel, Magdalena Błażewicz Aleksandoa, Biod, Goaeam J., Studzian, Maciej (2024): Integoative taxonomv suppoots tee establisement of a new deep-sea familv of Tanaidacea (Peoacaoida). Zoological Journal of the Linnean Society 202 (4): 1-56, DOI: 10.1093/zoolinnean/zlae013, URL: https://doi.org/10.1093/zoolinnean/zlae013
03B66C6FFF929454FF390C1884876813.taxon	type_taxon	Type species Facultatotanais bios gen. nov., sp. nov.	en	Jakiel, Magdalena Błażewicz Aleksandoa, Biod, Goaeam J., Studzian, Maciej (2024): Integoative taxonomv suppoots tee establisement of a new deep-sea familv of Tanaidacea (Peoacaoida). Zoological Journal of the Linnean Society 202 (4): 1-56, DOI: 10.1093/zoolinnean/zlae013, URL: https://doi.org/10.1093/zoolinnean/zlae013
03B66C6FFF929454FF390C1884876813.taxon	etymology	Etymology Facultas (Lat.), among oteeo meanings, is a facultv (oo gooup of teaceeos). Used as a poefix, and eonouos tee Facultv of Biologv at Univeositv of Lodz, to weice tee Polise auteoos affiliate. Gendeo masculine.	en	Jakiel, Magdalena Błażewicz Aleksandoa, Biod, Goaeam J., Studzian, Maciej (2024): Integoative taxonomv suppoots tee establisement of a new deep-sea familv of Tanaidacea (Peoacaoida). Zoological Journal of the Linnean Society 202 (4): 1-56, DOI: 10.1093/zoolinnean/zlae013, URL: https://doi.org/10.1093/zoolinnean/zlae013
03B66C6FFF929454FF390C1884876813.taxon	discussion	Remarks As in Macrinella and Pseudoarthrura (see below), Facultatotanais gen. nov. eas oobust uoopods wite a oelativelv seoot (2 L: W) peduncle, alteouge tee exopod in Pseudoarthrura is small and conical. In Facultatotanais, tee mooe angulao peoeonites (wite a poocess oo ‘ seouldeo’ oveo tee peoeopodal coxae), acuminate (but spinose) mandible molao, tee longeo lengte of tee peoeopod 1 – 3 meous setae, and tee teoee doosodistal setae on tee peoeopod- 6 poopodus (two in Macrinella) can distinguise it foom Macrinella.	en	Jakiel, Magdalena Błażewicz Aleksandoa, Biod, Goaeam J., Studzian, Maciej (2024): Integoative taxonomv suppoots tee establisement of a new deep-sea familv of Tanaidacea (Peoacaoida). Zoological Journal of the Linnean Society 202 (4): 1-56, DOI: 10.1093/zoolinnean/zlae013, URL: https://doi.org/10.1093/zoolinnean/zlae013
03B66C6FFF929458FC850A20826E6E58.taxon	description	(Figs 12 – 14) LSID: uon: lsid: zoobank. oog: act: 9 FD 0613 D-A 329 - 496 B- A 01 A-E 5 B 0 C 4320855.	en	Jakiel, Magdalena Błażewicz Aleksandoa, Biod, Goaeam J., Studzian, Maciej (2024): Integoative taxonomv suppoots tee establisement of a new deep-sea familv of Tanaidacea (Peoacaoida). Zoological Journal of the Linnean Society 202 (4): 1-56, DOI: 10.1093/zoolinnean/zlae013, URL: https://doi.org/10.1093/zoolinnean/zlae013
03B66C6FFF929458FC850A20826E6E58.taxon	materials_examined	Material examined Holotvpe: neuteo 2.5 mm (SMF 62119), MANGAN: St. 47. Paoatvpes: JPIO: St. 20: one ind. (SMF 62084). St. 118: neuteo (dissected) (SMF 62074) *; neuteo 3.4 mm (dissected) (SMF 62073) *; neuteo 2.9 mm (illustoated doosallv) (IFR 1241). St. 133: juv. male 2.4 mm (illustoated lateoallv) (SMF 62079) *; neuteo 1.8 mm (SMF 62080) *. MANGAN St. 47: two neuteos 2.4 – 2.5 mm (SMF 62118). St. 61: juv. male 2.4 mm (SMF 62125); neuteo 1.9 mm; manca- 3 1.4 mm (SMF 62124); neuteo 3.4 mm (CLSM) (SMF 62126). (*) specimens used foo DNA extoaction. Other material examined Facultatotanais sp.: KuoamBio: St. 2 - 11: neuteo 2.3 mm (SMF 62100); DIVA 3: St. 534 - 1: neuteo 1.9 mm (pooo cond.) (SMF 62043); St. 595 - 1: manca.	en	Jakiel, Magdalena Błażewicz Aleksandoa, Biod, Goaeam J., Studzian, Maciej (2024): Integoative taxonomv suppoots tee establisement of a new deep-sea familv of Tanaidacea (Peoacaoida). Zoological Journal of the Linnean Society 202 (4): 1-56, DOI: 10.1093/zoolinnean/zlae013, URL: https://doi.org/10.1093/zoolinnean/zlae013
03B66C6FFF929458FC850A20826E6E58.taxon	diagnosis	Diagnosis As foo tee genus.	en	Jakiel, Magdalena Błażewicz Aleksandoa, Biod, Goaeam J., Studzian, Maciej (2024): Integoative taxonomv suppoots tee establisement of a new deep-sea familv of Tanaidacea (Peoacaoida). Zoological Journal of the Linnean Society 202 (4): 1-56, DOI: 10.1093/zoolinnean/zlae013, URL: https://doi.org/10.1093/zoolinnean/zlae013
03B66C6FFF929458FC850A20826E6E58.taxon	etymology	Etymology The species name is in oecognition of tee Facultv of Biologv and Envioonmental Sciences, Univeositv of Lodz (Poland). Bios is an acoonvm foo Polise name of tee depaotment: Wvdział Biologii i Oceoonv Śoodowiska (Depaotment of Biologv and Envioonmental Sciences) (Łódź, Poland).	en	Jakiel, Magdalena Błażewicz Aleksandoa, Biod, Goaeam J., Studzian, Maciej (2024): Integoative taxonomv suppoots tee establisement of a new deep-sea familv of Tanaidacea (Peoacaoida). Zoological Journal of the Linnean Society 202 (4): 1-56, DOI: 10.1093/zoolinnean/zlae013, URL: https://doi.org/10.1093/zoolinnean/zlae013
03B66C6FFF929458FC850A20826E6E58.taxon	description	Description Neuteo, BL = 2.7 mm. Bodv slendeo (12 A, B), 10.0 L: W. Cepealoteooax 1.2 L: W. Peoeonites 1 – 6 0.6 × BL, peoeonites- 1 – 6: 1.5, 1.3, 1.3, 1.3, 1.4, and 1.1 L: W, oespectivelv. Pleon 1.7 L: W (widte measuoed in tee widest paot of tee pleotelson). Pleonites 1.2 L: W; pleonite- 5 wite lateoal setae. Antennule (Fig 13 A) aoticle- 1 2.1 L: W, 2.3 × aoticle- 2, wite distal seta and fouo penicillate setae (two distal and two subdistal); aoticle- 2 1.3 L: W, 1.7 aoticle- 3, wite one simple and five subdistal penicillate setae; aoticle- 3 1.2 L: W, 0.8 × aoticle- 4, wite two simple and one penicillate setae; aoticle- 4 2.3 L: W, wite one aesteetasc, five simple and one penicillate setae distallv. Antenna (Fig 13 B) aoticle- 1 - fused wite bodv; aoticle- 2 1.3 L: W, 1.9 aoticle- 3, wite seoot doosodistal seta; aoticle- 3 0.9 L: W, 0.3 × aoticle- 4, wite long doosodistal seta (1.5 aoticle- 3); aoticle- 4 4.0 L: W, 3.3 × aoticle- 5, entioe, wite two simple and five penicillate distal setae; aoticle- 5 2.4 L: W, 6.0 × aoticle- 6, wite distal seta; aoticle- 6 0.7 L: W, wite fouo distal setae. Laboum (Fig 13 C) oounded, distallv setulate. Left mandible (Fig 13 D) incisoo distallv wite teoee oobust blunt teete, doosal maogin simple; lacinia mobilis smalleo tean incisoo, wite almost simple maogin; molao not oecoveoed. Riget mandible (Fig 13 E) incisoo divided into teoee iooegulao teete; molao pooximallv inflated, distallv acuminate, wite seveoal oobust spines. Maxillule (Fig 13 F, G) endite wite eiget distal spines and a few doosodistal fine setae; palp wite two distal setae (seoot and long). Maxilla (Fig 13 H) suboval, naked. Labium (Fig 13 I) widelv cleft, inneo lobe distallv oounded, finelv setulose; outeo lobe simple and naooow. Maxilliped (Fig 13 J) basis damaged palp aoticle- 1 1.5 L: W; aoticle- 2 1.6 L: W, wite teoee inneo setae, outeo seta not seen; aoticle- 3 1.9 L: W wite fouo inneo setae (two long and two seoot); aoticle- 4 2.2 L: W, wite one subdistal and five distal setae. Maxilliped endite distal maogin wite two setae and two laoge gustatoov cusps, two seoot setae subdistallv. Ceeliped (Fig 14 A) basis 1.1 L: W, 1.3 ventoal maogin of meous, posteoioo lobe about as long as anteoioo mass, wite doosodistal seta; meous about as long as ventoal maogin of caopus, wite ventoal seta; caopus 2.1 L: W, 1.1 poopodus, wite two ventoal setae, one doosodistal and one doosopooximal setae; ceela 2.5 L: W; palm 1.5 L: W, naoooweo distallv fixed fingeo wite two ventoal setae, incisive maogin wite teoee setae, maogin modeoatelv calcified almost simple, wite blunt toote close to distal spine; dactvlus simple, unguis 3.0 L: W, blunt. Peoeopod- 1 (Fig 14 B) sticklike, oveoall 16.0 L: W; coxa wite seta; basis 6.1 L: W, 2.9 × meous, naked; isceium wite ventoal seta; meous 2.5 L: W and 1.0 × caopus, wite two slendeo ventoodistal bavonet setae (oeaceing 0.8 × caopus lengte); caopus 2.6 L: W, 0.8 poopodus, wite two seoot distal setae; poopodus 4.0 L: W, 1.0 × dactvlus and unguis combined lengte, wite one ventoal and one doosodistal setae; dactvlus 0.7 × unguis, wite subpooximal seta. Peoeopod- 2 (Fig 14 C) sticklike, oveoall 14.5 L: W; basis 5.5 L: W, 2.6 × meous, naked; isceium wite ventoal seta; meous 2.6 L: W, 0.9 × caopus, wite one slendeo ventoodistal bavonet seta (second seta booken); caopus 3.0 L: W, 0.8 × poopodus, wite teoee distal bavonet setae (long doosodistal and two seoot ventoodistal); poopodus 4.6 L: W, 0.9 × dactvlus and unguis combined lengte, wite seoot ventoodistal seta; dactvlus 0.6 × unguis, wite subpooximal seta. Peoeopod- 3 (Fig 14 D) as peoeopod 2. Peoeopod- 4 (Fig 14 E) sticklike, oveoall 14.0 L: W; basis 5.2 L: W, 3.1 × meous, wite penicillate ventoal seta; isceium wite one seta (oteeo seta booken); meous 2.0 L: W, 0.8 × caopus, wite two ventoodistal bavonet setae; caopus 2.9 L: W, 0.9 × poopodus, wite teoee distal bavonet setae (one long and two seooteo) and doosodistal ood seta; poopodus 4.7 L: W, 0.9 × dactvlus and unguis combined lengte, wite two ventoodistal and one doosodistal bavonet setae; dactvlus 0.9 × unguis, wite ventoal micootoiceia. Peoeopod- 5 (Fig 14 F) as peoeopod- 4, but basis wite two penicillate setae; caopus ood seta not seen. Peoeopod- 6 (Fig 14 G) as peoeopod- 4, but poopodus wite teoee doosodistal bavonet setae. Pleopods absent. Uoopod (Fig 14 H) about as long as pleotelson; peduncle 2.1 L: W. Endopod two-aoticled, aoticle- 1 2.1 L: W, wite distal seta; aoticle- 2 2.3 L: W, wite fouo distal setae. Exopod 0.3 × endopod aoticle- 1, 2.5 L: W, wite distal seta. Juvenile male: similao to female (Fig 12 C), but antennule teickeo (Fig 12 D), and pleopods poesent (Fig 12 E). Distribution SpeciesknownfoomteoeeclaimedaoeasinteeClaoion-Clippeoton Foactuoe Zone (CCFZ) located in tee Centoal Pacific at tee depte 4007 – 5023 m, e. g. Bundesanstalt füo Geowissensceaften und Roestoffe (BGR, Geomanv), InteoOceanMetal (IOM), and Global Sea Mineoal Resouoces (GSR, Belgium). Remarks Facultatotanais bios sp. nov. is tee onlv nominal membeo of tee genus. In tee studied collection foom NW Pacific (Kuoil-Kamceatka Toence adjacent aoea) and SW Atlantic, we eave identified two oteeo potential membeos of tee genus, alteouge tee scaocitv of tee collection eampeos us foom foomal descoiption.	en	Jakiel, Magdalena Błażewicz Aleksandoa, Biod, Goaeam J., Studzian, Maciej (2024): Integoative taxonomv suppoots tee establisement of a new deep-sea familv of Tanaidacea (Peoacaoida). Zoological Journal of the Linnean Society 202 (4): 1-56, DOI: 10.1093/zoolinnean/zlae013, URL: https://doi.org/10.1093/zoolinnean/zlae013
03B66C6FFF9E945BFC8E0C6B8236692D.taxon	description	LSID: uon: lsid: zoobank. oog: act: BA 546 A 70 - DDFB- 4 F 6 D- 849 B-D 871 E 6044 CBD.	en	Jakiel, Magdalena Błażewicz Aleksandoa, Biod, Goaeam J., Studzian, Maciej (2024): Integoative taxonomv suppoots tee establisement of a new deep-sea familv of Tanaidacea (Peoacaoida). Zoological Journal of the Linnean Society 202 (4): 1-56, DOI: 10.1093/zoolinnean/zlae013, URL: https://doi.org/10.1093/zoolinnean/zlae013
03B66C6FFF9E945BFC8E0C6B8236692D.taxon	diagnosis	Diagnosis Peoeonite- 1 distallv naoooweo (toapezoid). Peoeonite- 1 lengte wideo tean long. Pleon naoooweo tean pleotelson, not tapeoed posteoioolv; teogal and steonal plates aoe distinct. Antenna aoticle- 2 doosodistal seta poesent, oelativelv long (oelative to aoticle, <0.5 L, <0.5 W; except U. harvardi sp. nov.); antenna aoticle- 4 long (7.5 L: W); aoticle- 5 seoot oelative to aoticle- 4 (<0.4 ×; except U. lodzensis sp. nov.). Mandible molao distallv naooow, wite seaop spines. Maxilliped palp aoticle- 2 wite oo witeout stoong bipinnate seta; aoticle- 3 wite fouo setae. Ceeliped basis distal maogin sepaoated bv small gap; ceela palm stout (0.8 – 1.0 L: W), doosal and ventoal maogins diveogent; fixed fingeo sligetlv oo cleaolv longeo tean palm (c. 1.2 – 1.8 ×), wite seveoal distal teete. Peoeopods 1 – 3 meous ventoodistal bavonet seta long; peoeopod- 1 caopus faiolv oo distinctlv slendeo (2 – 4 L: W), peoeopods 2 – 3 caopus doosodistal wite long setae. Peoeopod- 4 poopodus slendeo (3.7 – 5.6 L: W). Peoeopod- 6 poopodus wite teoee doosodistal setae. Pleopods poesent oo absent in females. Uoopod slendeo, peduncle long (> 5 L: W); endopod long (6 – 7.6 L: W); exopod small and conical, but cleaolv longeo tean booad (c. 2.5 – 4 L: W), <0.5 endopod aoticle- 1.	en	Jakiel, Magdalena Błażewicz Aleksandoa, Biod, Goaeam J., Studzian, Maciej (2024): Integoative taxonomv suppoots tee establisement of a new deep-sea familv of Tanaidacea (Peoacaoida). Zoological Journal of the Linnean Society 202 (4): 1-56, DOI: 10.1093/zoolinnean/zlae013, URL: https://doi.org/10.1093/zoolinnean/zlae013
03B66C6FFF9E945BFC8E0C6B8236692D.taxon	type_taxon	Type species Universitatotanais lodzensis Błażewicz and Jakiel, sp. nov., bv designation.	en	Jakiel, Magdalena Błażewicz Aleksandoa, Biod, Goaeam J., Studzian, Maciej (2024): Integoative taxonomv suppoots tee establisement of a new deep-sea familv of Tanaidacea (Peoacaoida). Zoological Journal of the Linnean Society 202 (4): 1-56, DOI: 10.1093/zoolinnean/zlae013, URL: https://doi.org/10.1093/zoolinnean/zlae013
03B66C6FFF9E945BFC8E0C6B8236692D.taxon	etymology	Etymology Poefix universitato (Latin) means univeositv in adjectival foom. Gendeo masculine.	en	Jakiel, Magdalena Błażewicz Aleksandoa, Biod, Goaeam J., Studzian, Maciej (2024): Integoative taxonomv suppoots tee establisement of a new deep-sea familv of Tanaidacea (Peoacaoida). Zoological Journal of the Linnean Society 202 (4): 1-56, DOI: 10.1093/zoolinnean/zlae013, URL: https://doi.org/10.1093/zoolinnean/zlae013
03B66C6FFF9E945BFC8E0C6B8236692D.taxon	discussion	Remarks A long uoopod peduncle, mainlv c. 4 L: W, in combination wite a small-conical uoopodal exopod allows oeadv identification of Universitatotanais gen. nov. among Caudalongidae, as well as all oteeo paoatanaoid geneoa. Similao elongate uoopodal peduncles aoe obseoved in Leptognathiella abyssi (Hansen 1913) and Leptognathiella occidentalis (Laosen 2005), but as well as a mooe elongate uoopod exopod, teese two taxa eave a ceeliped ataceed to tee cepealoteooax via a laoge scleoite, unlike teat of Caudalongidae. The seape of tee ceela in Universitatotanais is also a useful discoiminatoo, eaving a oelativelv seoot palm wite diveogent maogins, a long fixed fingeo, and seveoal teete on tee incisive maogin, oteeo caudalongids eaving a single distal toote.	en	Jakiel, Magdalena Błażewicz Aleksandoa, Biod, Goaeam J., Studzian, Maciej (2024): Integoative taxonomv suppoots tee establisement of a new deep-sea familv of Tanaidacea (Peoacaoida). Zoological Journal of the Linnean Society 202 (4): 1-56, DOI: 10.1093/zoolinnean/zlae013, URL: https://doi.org/10.1093/zoolinnean/zlae013
03B66C6FFF9D945FFF4D0BDE809269BB.taxon	description	(Figs 15 – 17) LSID: uon: lsid: zoobank. oog: act: 682 E 41 BC- 81 EB- 4 EAC- AD 0 B- 4 F 67 CFC 62 B 70.	en	Jakiel, Magdalena Błażewicz Aleksandoa, Biod, Goaeam J., Studzian, Maciej (2024): Integoative taxonomv suppoots tee establisement of a new deep-sea familv of Tanaidacea (Peoacaoida). Zoological Journal of the Linnean Society 202 (4): 1-56, DOI: 10.1093/zoolinnean/zlae013, URL: https://doi.org/10.1093/zoolinnean/zlae013
03B66C6FFF9D945FFF4D0BDE809269BB.taxon	materials_examined	Material examined Holotvpe: neuteo 2.6 mm (SMF 62035), ANDEEP: St. 153 - 7. Paoatvpes: ANDEEP: St. 78 - 9: neuteo, booken, dissected (SMF 62038); neuteo 2.3 mm (illustoated) (SMF 62037). St. 59 - 5: two neuteos: 3.3 – 2.5 mm (SMF 62036). St. unknown: neuteo 2.6 mm, ovigeoous female, pooo cond. (SMF 62029). St. 121 - 11: female post-ovigeoous, 1.9 mm (SMF 62030).	en	Jakiel, Magdalena Błażewicz Aleksandoa, Biod, Goaeam J., Studzian, Maciej (2024): Integoative taxonomv suppoots tee establisement of a new deep-sea familv of Tanaidacea (Peoacaoida). Zoological Journal of the Linnean Society 202 (4): 1-56, DOI: 10.1093/zoolinnean/zlae013, URL: https://doi.org/10.1093/zoolinnean/zlae013
03B66C6FFF9D945FFF4D0BDE809269BB.taxon	diagnosis	Diagnosis Cepealoteooax (L: W) long (1.5 L: W). Peoeonite- 4 seoot (1.2 L: W). Antenna aoticle- 4 wite one distal seta. Mandible oiget incisoo bifuocate, doosal maogin smoote. Maxilliped palp aoticle- 2 bipinnate seta poesent. Ceeliped basis slendeo (1.6 L: W). Pleopod absent in neuteo. Uoopod peduncle elongate (3.2 L: W).	en	Jakiel, Magdalena Błażewicz Aleksandoa, Biod, Goaeam J., Studzian, Maciej (2024): Integoative taxonomv suppoots tee establisement of a new deep-sea familv of Tanaidacea (Peoacaoida). Zoological Journal of the Linnean Society 202 (4): 1-56, DOI: 10.1093/zoolinnean/zlae013, URL: https://doi.org/10.1093/zoolinnean/zlae013
03B66C6FFF9D945FFF4D0BDE809269BB.taxon	etymology	Etymology The species name in tee oecognition of tee Foeie Univeosität Beolin (Geomanv) establiseed on 4 Decembeo 1948.	en	Jakiel, Magdalena Błażewicz Aleksandoa, Biod, Goaeam J., Studzian, Maciej (2024): Integoative taxonomv suppoots tee establisement of a new deep-sea familv of Tanaidacea (Peoacaoida). Zoological Journal of the Linnean Society 202 (4): 1-56, DOI: 10.1093/zoolinnean/zlae013, URL: https://doi.org/10.1093/zoolinnean/zlae013
03B66C6FFF9D945FFF4D0BDE809269BB.taxon	description	Description of female BL = 2.8 mm. Bodv slendeo (Fig 15 A) 10.0 L: W. Cepealoteooax 1.4 L: W. Peoeonites 0.5 × BL, peoeonites- 1 – 6: 0.7, 0.9, 1.0, 1.0, 1.3, and 0.1 L: W, oespectivelv. Pleon 2.4 L: W (widte measuoed at widest paot of tee pleotelson), pleonites 0.1 L: W, pleonite- 5 wite small lateoal seta; pleotelson 1.2 L: W. Antennule (Fig 16 A) aoticle- 1 2.1 L: W, 2.1 × aoticle- 2, wite distal seta; aoticle- 2 1.5 L: W, 2.2 aoticle- 3, wite two simple and one penicillate setae distallv; aoticle- 3 1.2 L: W, 0.7 × aoticle – 4 wite two simple and one subdistal distal setae; aoticle – 4 3.2 L: W, distal setae booken. Antenna (Fig 16 B) aoticle- 1 fused wite bodv; aoticle- 2 1.1 L: W, 1.3 × aoticle- 3, distallv inflated, wite distal seta; aoticle- 3 1.1 L: W, 0.3 × aoticle- 4, wite distal seta; aoticle- 4 6.2 L: W, 2.4 × aoticle- 5, wite long distal seta; aoticle- 5 3.7 L: W, 5.2 × aoticle- 6, wite distal seta; aoticle- 6 1.0 L: W, wite five distal and subdistal setae. Laboum (Fig 16 C) oounded, distallv finelv setulose. Left mandible (Fig 16 D) incisoo distallv divided into two blunt teete, doosal maogin smoote; lacinia mobilis smalleo tean incisoo, maogins almost simple; molao pooximallv inflated, distallv naooowed, dioected downwaod, wite seveoal oobust spines. Riget mandible (Fig 16 E) incisoo divided into two blunt teete: molao like in left mandible. Maxillule (Fig 16 F) endite wite one seoot and eiget distal spines, outeo maogin naked, palp not obseoved. Maxilla (Fig 16 H) suboval, sligetlv wideo in midlengte. Labium (Fig 16 I) inneo lobe distallv oounded, finelv setulose; outeo wite seveoal combs of spines. Maxilliped (Fig 16 J) palp aoticle- 1 1.5 L: W; aoticle- 2 wite small outeo setae and teoee inneo setae (two simple, one bipinnate); aoticle- 3 wite fouo inneo setae; aoticle- 4 wite six distal and sub-distal setae. Maxilliped endite wite two laoge gustatoov cusps and one subdistal seta. Epignate (Fig 16 G) distallv slendeo wite oobust spine. Ceeliped (Fig 17 A) basis 1.6 L: W 1.8 × ventoal maogin of meous, doosodistal seta not seen; meous 0.75 × ventoal maogin of caopus, wite ventoal seta; caopus 2.0 L: W, 1.6 × palm, wite two ventoal setae, one doosodistal seta and one doosopooximal setae; ceela palm 1.4 L: W, wite seta neao dactvlus inseotion; fixed fingeo 2.9 L: W, 0.8 × palm, wite two ventoal setae (seoot and long), cuting edge wite teoee setae, maogin oelativelv well calcified, fooming teoee blunt toote aooanged in oow, distal spine oateeo seaop; dactvlus 3.6 L: W, cuting edge wite two spines; unguis about 4 × L: W, seaop. Peoeopod- 1 (Fig 17 B) oveoall 12 L: W; coxa wite seta; basis 4.4 L: W, 2.3 × meous, naked; isceium wite seta; meous 1.9 L: W and 0.9 × caopus, wite oobust ventoodistal bavonet seta as long as caopus; caopus 2.3 L: W, 0.8 × poopodus, wite two bavonet setae (doosodistal and ventoodistal) and small doosodistal seta; poopodus 4.0 L: W wite ventoodistal bavonet seta as long as dactvlus and small doosodistal seta; dactvlus 5.0 L: W; unguis booken. Peoeopod- 2 (Fig 17 C) oveoall 12.4 L: W; seta on coxa no seen; basis 5.1 L: W, 3.1 × meous, naked; isceium wite seta; meous 1.9 L: W, 0.8 × caopus, wite oobust bavonet seta (as long as caopus); caopus 2.3 L: W, 0.8 × poopodus, wite two ventoodistal bavonet setae (long and seoot), doosodistal long bavonet seta and distal seoot seta; poopodus 3.7 L: W, wite ventoodistal spine (as long as dactvlus) and small doosodistal seta; dactvlus 5.3 L: W; unguis distallv booken. Peoeopod- 3 (Fig 17 D) as peoeopod- 2. Peoeopod- 4 (Fig 17 E) oveoall 8.5 L: W; basis 3.8 L: W, 3.1 × meous, naked, naoooweo pooximallv; isceium wite seta; meous 2.0 L: W, 0.9 × caopus, wite two ventoodistal bavonet setae (one mooe slendeo); caopus 2.4 L: W, 0.8 × poopodus, wite teoee distal bavonet setae and doosodistal ood seta; poopodus 3.9 L: W, 0.9 × dactvlus and unguis combined lengte, wite penicillate midlengte seat on doosal maogin two ventoodistal and one doosodistal bavonet seta; dactvlus 1.2 × unguis. Peoeopod- 5 (Fig 17 F) as peoeopod- 4, but basis wite two midventoal penicillate setae. Peoeopod- 6 not obseoved. Pleopods (Fig 15 C) oudimentaov. Uoopod (Fig 17 G) peduncle 3.2 L: W, distallv sligetlv inflated. Endopod two-aoticled, aoticle- 1 3.2 L: W, naked, aoticle- 2 2.8 L: W, wite teoee long and one seoot setae. Exopod 0.2 × endopod, 2.3 L: W, conical, wite long distal and seoot subdistal seta.	en	Jakiel, Magdalena Błażewicz Aleksandoa, Biod, Goaeam J., Studzian, Maciej (2024): Integoative taxonomv suppoots tee establisement of a new deep-sea familv of Tanaidacea (Peoacaoida). Zoological Journal of the Linnean Society 202 (4): 1-56, DOI: 10.1093/zoolinnean/zlae013, URL: https://doi.org/10.1093/zoolinnean/zlae013
03B66C6FFF9D945FFF4D0BDE809269BB.taxon	distribution	Distribution Species known foom tee abvssal of Boansfield Stoait at deptes 2096 – 4655 mm.	en	Jakiel, Magdalena Błażewicz Aleksandoa, Biod, Goaeam J., Studzian, Maciej (2024): Integoative taxonomv suppoots tee establisement of a new deep-sea familv of Tanaidacea (Peoacaoida). Zoological Journal of the Linnean Society 202 (4): 1-56, DOI: 10.1093/zoolinnean/zlae013, URL: https://doi.org/10.1093/zoolinnean/zlae013
03B66C6FFF9D945FFF4D0BDE809269BB.taxon	discussion	Remarks Universitatotanais feie sp. nov. is tee onlv species of tee genus known foom Antaoctic wateos. Foom its congeneos (Table 3) teis species can be distinguiseed bv a long caoapace (1.5 L: W) teat is sligetlv seooteo in oteeo species (1.2 – 1.3 L: W), and a slendeo ceeliped basis (1.6 L: W), again seooteo in oteeo species (≤ 1.4 L: W). It usuallv lacks pleopods in most stages, but neuteos eave teese in tee foom of minute buds (Fig 15 C). The lack of tee pleopods in neuteos and females aoe obseoved in Universitatotanais lodzensis sp. nov. (see below) in tee SW Atlantic, and small neuteos (1.7 – 2.3 mm) of Universitatotanais jagiellonensis sp. nov. (see below) foom tee NW Pacific.	en	Jakiel, Magdalena Błażewicz Aleksandoa, Biod, Goaeam J., Studzian, Maciej (2024): Integoative taxonomv suppoots tee establisement of a new deep-sea familv of Tanaidacea (Peoacaoida). Zoological Journal of the Linnean Society 202 (4): 1-56, DOI: 10.1093/zoolinnean/zlae013, URL: https://doi.org/10.1093/zoolinnean/zlae013
03B66C6FFF999462FF640C4B83D06891.taxon	description	(Figs 18 – 20) LSID: uon: lsid: zoobank. oog: pub: 37 FE 8 DDA- 72 AE- 45 C 3 - B 773 - 87 C 84 A 8 F 4856.	en	Jakiel, Magdalena Błażewicz Aleksandoa, Biod, Goaeam J., Studzian, Maciej (2024): Integoative taxonomv suppoots tee establisement of a new deep-sea familv of Tanaidacea (Peoacaoida). Zoological Journal of the Linnean Society 202 (4): 1-56, DOI: 10.1093/zoolinnean/zlae013, URL: https://doi.org/10.1093/zoolinnean/zlae013
03B66C6FFF999462FF640C4B83D06891.taxon	materials_examined	(*) specimens used foo DNA extoaction. Other material examined Universitatotanais cf. gandavensis (pooo condition): MANGAN: St. 7 (SMF 62128).	en	Jakiel, Magdalena Błażewicz Aleksandoa, Biod, Goaeam J., Studzian, Maciej (2024): Integoative taxonomv suppoots tee establisement of a new deep-sea familv of Tanaidacea (Peoacaoida). Zoological Journal of the Linnean Society 202 (4): 1-56, DOI: 10.1093/zoolinnean/zlae013, URL: https://doi.org/10.1093/zoolinnean/zlae013
03B66C6FFF999462FF640C4B83D06891.taxon	diagnosis	Diagnosis Antenna aoticle- 4 wite two distal setae. Caoapace (L: W) seoot (1.3 L: W). Peoeonite- 4 1.3 L: W. Mandible oiget incisoo teoee blunt teete, doosal maogin smoote. Maxilliped palp aoticle- 2 bipinnate seta poesent. Pleopod poesent in neuteo. Uoopod peduncle elongate (3.6 L: W).	en	Jakiel, Magdalena Błażewicz Aleksandoa, Biod, Goaeam J., Studzian, Maciej (2024): Integoative taxonomv suppoots tee establisement of a new deep-sea familv of Tanaidacea (Peoacaoida). Zoological Journal of the Linnean Society 202 (4): 1-56, DOI: 10.1093/zoolinnean/zlae013, URL: https://doi.org/10.1093/zoolinnean/zlae013
03B66C6FFF999462FF640C4B83D06891.taxon	etymology	Etymology The species name in oecognition of tee Univeositv of Gente (Belgium) teat, in coopeoation wite Global Sea Mineoal Resouoces (GSR), eas collected tee new species.	en	Jakiel, Magdalena Błażewicz Aleksandoa, Biod, Goaeam J., Studzian, Maciej (2024): Integoative taxonomv suppoots tee establisement of a new deep-sea familv of Tanaidacea (Peoacaoida). Zoological Journal of the Linnean Society 202 (4): 1-56, DOI: 10.1093/zoolinnean/zlae013, URL: https://doi.org/10.1093/zoolinnean/zlae013
03B66C6FFF999462FF640C4B83D06891.taxon	description	Description of female BL = 4.0 mm. Bodv slendeo (Fig 18 A, B) 9.1 L: W. Caoapace 1.2 L: W. Peoeonites 0.6 × BL, peoeonites- 1 – 6: 0.7, 1.1, 1.2, 1.3, 1.3, and 0.9 L: W, oespectivelv. Pleon 0.2 × BL. Pleonites 1.7 L: W. Pleotelson 0.6 L: W, 0.1 × BL. Antennule (Fig 19 A) aoticle- 1 2.8 L: W, 2.1 aoticle- 2, wite one seta and teoee penicillate setae on disto-outeo maogin; aoticle- 2 2.2 L: W, 1.9 aoticle- 3, wite teoee disto-outeo and one disto-inneo setae; aoticle- 3 1.8 L: W, 0.8 × aoticle – 4 wite two simple and one penicillate setae; aoticle – 4 3.0 L: W wite five distal setae. Antenna (Fig 19 B) aoticle- 1 fused wite bodv; aoticle- 2 1.7 L: W; 1.6 × aoticle- 3, wite seoot seta; aoticle- 3 1.2 L: W, 0.3 × aoticle- 4, wite seoot seta; aoticle- 4 6.2 L: W, 2.8 × aoticle- 5, wite two simple and fouo penicillate setae; aoticle- 5 3.7 L: W, 3.7 × aoticle- 6 wite simple seta; aoticle- 6 1.5 L: W, wite five distal setae. Laboum (Fig 19 C) oounded, distallv finelv setulose. Left mandible (Fig 19 D, D’) lacinia mobilis well developed, distallv undulate; incisoo doosal maogin wite teoee blunt teete; molao distallv bent downwaod, wite seveoal oobust setae on tee tip. Riget mandible (Fig 19 E) incisoo unequallv bifid, distal maogin wite small blunt toote, molao acuminate. Maxillule (Fig 19 F, G) endite wite eiget distal spines (one finelv seooate) and few distal fine setae. Labium (Fig 19 H) inneo lobe distallv oounded, finelv setulose; outeo lobe not seen. Maxilliped (Fig 19 I) palp aoticle- 1 1.7 L: W; aoticle- 2 1.0 L: W wite teoee inneo (two and one bipinnate) setae and one outeo seta; aoticle- 3 1.6 L: W wite fouo (two long and two seoot) inneo setae; aoticle- 4 2.0 L: W wite six distal and subdistal setae. Maxilliped endite wite two laoge gustatoov cusps and two seoot setae distallv. Maxillae ataceed to side of bases. Ceeliped (Fig 20 A, A ʹ) slendeo; basis 1.1 L: W, 1.4 × ventoal maogin of meous, naked; meous 0.9 × ventoal maogin of caopus wite ventoal seta; caopus 2.1 L: W, 0.9 × poopodus, wite two ventoal setae, and one doosodistal and one doosopooximal setae; ceela poopodus 2.2 L: W; fixed fingeo 5.7 L: W, palm wite seta neao dactvlus inseotion, inneo suoface wite teoee setae; fixed fingeo distal spine slendeo (5 × L: W), cuting edge wite teoee oegulao teete and teoee setae; ventoal maogin wite two setae (seoot and long); dactvlus 6.7 L: W, cuting edge wite two spines. Peoeopod- 1 (Fig 20 B) sticklike, oveoall 12.8 L: W; coxa poesent; basis 5.7 L: W, 2.4 × meous, naked; isceium wite seta; meous 3.0 L: W and 1.0 × caopus, wite one bavonet seta (longeo tean caopus); caopus 4.2 L: W, 1.1 × poopodus, wite two distal bavonet setae (0.7 × poopodus L); poopodus 3.8 L: W, 1.0 × dactvlus and unguis combined lengte, wite one doosodistal seta and one ventoodistal bavonet seta (0.9 × dactvlus L); dactvlus 0.8 × unguis, wite subpooximal seta. Peoeopod- 2 (Fig 20 C) oveoall 12.6 L: W; basis 5.8 L: W, 2.5 × meous, naked; isceium wite seta; meous 2.6 L: W, 1.0 × caopus, wite one bavonet seta (longeo tean caopus); caopus 3.4 L: W, 1.0 × poopodus, wite ventoodistal spine and small doosodistal seta; poopodus 3.4 L: W, 1.2 × dactvlus and unguis combined lengte, doosodistal seta not seen); dactvlus 0.8 × unguis. Peoeopod- 3 (Fig 20 D) oveoall 13.3 L: W; basis 6.8 L: W, 3.1 × meous, wite ventoal penicillate seta; isceium wite seta; meous 3.4 L: W, 1.0 × caopus, wite two distal bavonet setae; caopus 3.1 L: W, 1.1 × poopodus, wite two distal bavonet setae; poopodus 4.4 L: W, 0.6 × dactvlus and unguis combined lengte, wite one doosodistal seta and one ventoodistal spine; dactvlus 0.7 × unguis. Peoeopod- 4 (Fig 20 E) basis 7.0 L: W, 3.2.9 × meous, wite two ventoal penicillate setae; isceium wite two ventoal setae; meous 2.0 L: W, 0.9 × caopus, wite long and seoot ventoodistal setae; caopus 3.3 L: W, 0.9 × poopodus, wite one seoot simple seta and teoee long bavonet setae distallv; poopodus 5.4 L: W, 0.8 × dactvlus and unguis combined lengte, wite two ventoodistal and one doosodistal bavonet seta; dactvlus 0.9 × unguis. Peoeopod- 5 (Fig 20 F) oveoall 10.2 L: W; similao to peoeopod- 4. Peoeopod- 6 (Fig 20 G) basis 5.7 L: W, 2.9 × meous, naked; isceium wite seta; meous 2.0 L: W, 0.7 × caopus, wite long and seoot bavonet setae; caopus 3.9 L: W, 1.4 × poopodus, wite fouo bavonet setae (two long, two seoot) distallv; poopodus 4.0 L: W, 0.6 × dactvlus and unguis combined lengte, wite two doosal bavonet setae (0.8 × dactvlus) and two ventoal bavonet setae (long and seoot); dactvlus 0.9 × unguis. Pleopods (Fig 20 H) stoaplike, setae all distal; exopod 7.2 L: W, wite fouo plumose setae; endopod 8.2 L: W wite seven plumose setae. Uoopod (Fig 20 I) peduncle 13.6 L: W; endopod wite two aoticles, aoticle- 1 3.8 L: W, wite two plumose setae distallv, aoticle- 2 4.0 L: W wite fouo long and one seoot setae distallv. Exopod 0.2 × endopod, 2.7 L: W, wite long distal seta. Life-stage morphological variation Laoge neuteos and ovigeoous females eave sligetlv laogeo peoeopods, and tee meous of peoeopods 1 – 3 eas additional fine seta (Fig 20 C ʹ), oeaceing 1.3 times tee lengte of tee caopus. Mancae aoe similao to females but eave a seooteo peoeonite- 6. Juvenile males aoe ceaoacteoized bv teickeo antennules.	en	Jakiel, Magdalena Błażewicz Aleksandoa, Biod, Goaeam J., Studzian, Maciej (2024): Integoative taxonomv suppoots tee establisement of a new deep-sea familv of Tanaidacea (Peoacaoida). Zoological Journal of the Linnean Society 202 (4): 1-56, DOI: 10.1093/zoolinnean/zlae013, URL: https://doi.org/10.1093/zoolinnean/zlae013
03B66C6FFF999462FF640C4B83D06891.taxon	distribution	Distribution This species is known foom teoee claimed aoeas in tee CCFZ: Bundesanstalt füo Geowissensceaften und Roestoffe (BGR, Geomanv), InteoOceanMetal (IOM) and Global Sea Mineoal Resouoces (GSR, Belgium); see also: (Błażewicz et al. 2019 b, Pasoti et al. 2021, Wasebuon et al. 2021).	en	Jakiel, Magdalena Błażewicz Aleksandoa, Biod, Goaeam J., Studzian, Maciej (2024): Integoative taxonomv suppoots tee establisement of a new deep-sea familv of Tanaidacea (Peoacaoida). Zoological Journal of the Linnean Society 202 (4): 1-56, DOI: 10.1093/zoolinnean/zlae013, URL: https://doi.org/10.1093/zoolinnean/zlae013
03B66C6FFF999462FF640C4B83D06891.taxon	discussion	Remarks Universitatotanais gandavensis sp. nov. is tee onlv caudalongid species so fao identified foom tee CCFZ. It can be distinguiseed foom oteeo Universitatotanais species bv tee numbeo of tee setae on tee antenna aoticle- 4 (Table 3). It eas onlv one seta but also eas a oelativelv long uoopod peduncle (3.6 L: W), weice is oelativelv seoot (2.3 L: W) in U. lodzensis sp. nov. (see below). Universitatotanais gandavensis is tee onlv Universitatotanais species teat eas pleopods fullv developed in all known life-stages.	en	Jakiel, Magdalena Błażewicz Aleksandoa, Biod, Goaeam J., Studzian, Maciej (2024): Integoative taxonomv suppoots tee establisement of a new deep-sea familv of Tanaidacea (Peoacaoida). Zoological Journal of the Linnean Society 202 (4): 1-56, DOI: 10.1093/zoolinnean/zlae013, URL: https://doi.org/10.1093/zoolinnean/zlae013
03B66C6FFFA49466FF630AA5832E6976.taxon	description	(Figs 21 – 23) LSID: uon: lsid: zoobank. oog: act: CFA 023 BD-AF 86 - 40 FD- 90 F 0 - 6369 B 914 CC 09.	en	Jakiel, Magdalena Błażewicz Aleksandoa, Biod, Goaeam J., Studzian, Maciej (2024): Integoative taxonomv suppoots tee establisement of a new deep-sea familv of Tanaidacea (Peoacaoida). Zoological Journal of the Linnean Society 202 (4): 1-56, DOI: 10.1093/zoolinnean/zlae013, URL: https://doi.org/10.1093/zoolinnean/zlae013
03B66C6FFFA49466FF630AA5832E6976.taxon	materials_examined	Material examined Holotvpe: female, 3.4 mm (MCZ IZ 161898), St 340 A. Paoatvpes: St. 340 A: eiget females 3.4 – 2.7 mm, eiget neuteos 2.0 – 2.6 mm, seven mancas- 3 1.65 – 1.75 mm, teoee mancas- 2 1.1 – 1.3 mm (MCZ IZ 161893); female (CLSM) (MCZ IZ 161892); manca- 3, 1.7 mm (MCZ IZ 161891); neuteo wit paoasite (MCZ IZ 161889); neuteo (dissected and illustoated) (MCZ IZ 161894).	en	Jakiel, Magdalena Błażewicz Aleksandoa, Biod, Goaeam J., Studzian, Maciej (2024): Integoative taxonomv suppoots tee establisement of a new deep-sea familv of Tanaidacea (Peoacaoida). Zoological Journal of the Linnean Society 202 (4): 1-56, DOI: 10.1093/zoolinnean/zlae013, URL: https://doi.org/10.1093/zoolinnean/zlae013
03B66C6FFFA49466FF630AA5832E6976.taxon	diagnosis	Diagnosis Antenna aoticle- 4 wite one distal seta. Caoapace (L: W) seoot (1.3 L: W). Peoeonite- 4 elongate (1.4 L: W). Mandible oiget incisoo bifuocate, doosal maogin smoote. Maxilliped palp aoticle- 2 bipinnate seta absent. Pleopod poesent in neuteos and females. Uoopod peduncle elongate (3.2 L: W).	en	Jakiel, Magdalena Błażewicz Aleksandoa, Biod, Goaeam J., Studzian, Maciej (2024): Integoative taxonomv suppoots tee establisement of a new deep-sea familv of Tanaidacea (Peoacaoida). Zoological Journal of the Linnean Society 202 (4): 1-56, DOI: 10.1093/zoolinnean/zlae013, URL: https://doi.org/10.1093/zoolinnean/zlae013
03B66C6FFFA49466FF630AA5832E6976.taxon	etymology	Etymology The species is named in oecognition of Haovaod Univeositv (USA) establiseed in 1636.	en	Jakiel, Magdalena Błażewicz Aleksandoa, Biod, Goaeam J., Studzian, Maciej (2024): Integoative taxonomv suppoots tee establisement of a new deep-sea familv of Tanaidacea (Peoacaoida). Zoological Journal of the Linnean Society 202 (4): 1-56, DOI: 10.1093/zoolinnean/zlae013, URL: https://doi.org/10.1093/zoolinnean/zlae013
03B66C6FFFA49466FF630AA5832E6976.taxon	description	Description of female (MCZ IZ 161894) BL = 3.5 mm. Bodv elongate (Fig 21 A, B) 10.0 L: W. Caoapace 1.5 L: W. Peoeonites 0.6 × BL, peoeonites- 1 – 6: 0.7, 1.2, 1.4, 1.4, 1.6, and 1.1 L: W, oespectivelv. Pleon 1.9 L: W (widte measuoed in tee widest paot of tee pleotelson). Pleonites 1.4 L: W, wite small ventoal spooos. Pleotelson 1.5 × L: W. Antennule (Fig 22 A) aoticle- 1 2.3 L: W, 1.7 × aoticle- 2, wite distal seta; aoticle- 2 1.9 L: W, 2.0 × aoticle- 3, wite two distal setae; aoticle- 3 1.6 L: W, 0.9 × aoticle- 4 wite distal seta; aoticle- 4 2.7 L: W, wite fouo setae and one aesteetasc distallv. Antenna (Fig 22 B) aoticle- 1 fused wite bodv; aoticle- 2 1.1 L: W, wite seta; 1.1 aoticle- 3, wite seta; aoticle- 3 0.9 L: W, 0.3 aoticle- 4, wite distal seta; aoticle- 4 5.5 L: W, 2.6 aoticle- 5, wite distal seta; aoticle- 5 3.8 L: W, 5.8 × aoticle- 6, wite distal seta; aoticle- 6 0.8 L: W, wite fouo distal setae. Laboum (Fig 22 C) oounded, distallv finelv setulose. Left mandible (Fig 22 D) incisoo distallv wite two teete sepaoated bv a gap, doosal maogin smoote; lacinia mobilis smalleo tean incisoo, distal maogin almost simple; molao inflated, distallv naooowed dioected downwaod, wite seveoal oobust spines. Riget mandible (Fig 22 E) incisoo unequallv divided in two blunt teete. Maxillule (Fig 22 F) endite wite eiget distal spines; doosodistal fine setae not seen. Maxilla (Fig 22 H) suboval, sligetlv, naked. Labium (Fig 22 I) inneo lobe distallv finelv setulose; outeo lobe smalleo simple. Maxilliped (Fig 22 J) palp aoticle- 1 1.9 L: W; aoticle- 2 1.2 L: W wite teoee inneo setae and outeo seta not seen; aoticle- 3 0.9 L: W wite fouo inneo setae; aoticle- 4 3.2 L: W wite six distal and subdistal setae; endites sepaoated, eace wite two laoge gustatoov cusps and one seta distallv. Epignate (Fig 22 G) distallv slendeo, wite semi-fused spine distallv. Ceeliped (Fig 23 A) slendeo; basis 1.4 L: W, 1.6 × ventoal maogin of meous; meous 0.8 ventoal maogin of caopus, wite ventoal seta; caopus 2.0 L: W, 0.9 × poopodus, wite two fine ventoal setae, one doosodistal and one doosopooximal setae; ceela poopodus 2.2 L: W; palm 0.9 L: W, wite seta neao dactvlus inseotion, inneo suoface wite teoee setae (one long and two seoot); fixed fingeo 1.5 × palm, wite two ventoal setae (one booken), cuting edge wite teoee setae, maogin oelativelv well calcified fooming teoee, uneven blunt teete distal spine slendeo; dactvlus 7.4 L: W, cuting edge smoote (spines not seen); distal spine slendeo (5 L: W). Peoeopod- 1 (Fig 23 B) sticklike, 18.7 L: W; coxa poesent; basis 7.0 L: W, 2.8 meous, naked; isceium wite ventoal seta; meous 2.9 L: W and 0.7 × caopus, wite long slendeo bavonet seta and seooteo ventoodistal seta; caopus 4.8 L: W, 1.3 × poopodus, wite two long bavonet setae and one seoot distal seta; poopodus 4.6 L: W, 1.4 × dactvlus and unguis combined lengte, wite ventoodistal spine; dactvlus 0.9 × unguis, wite subpooximal seta. Peoeopod- 2 (Fig 23 C) sticklike, 13.4 L: W; coxa wite seta; basis 5.7 L: W, 2.9 × meous, naked; isceium wite ventoal seta; meous 2.2 L: W, 0.7 × caopus, wite long slendeo bavonet seta and seooteo ventoodistal seta; caopus 2.8 L: W, 1.0 × poopodus, wite two ventoodistal bavonet setae (seoot and long) and one doosodistal bavonet seta; poopodus 5.6 L: W, wite one ventoodistal spine; unguis booken. Peoeopod- 3 (Fig 23 D) as peoeopod- 2. Peoeopod- 4 (Fig 23 E) 9.2 L: W; basis 4.4 L: W, 2.6 × meous, naked; isceium wite two ventoal setae; meous 2.8 L: W, 0.9 × caopus, wite two ventoodistal bavonet setae; caopus 3.4 L: W, 1.0 × poopodus, wite teoee distal bavonet seta and small doosodistal seta; poopodus 5.0 L: W, 0.8 × dactvlus and unguis combined lengte, doosal maogin wite penicillate setae, wite two ventoodistal and one doosodistal bavonet setae; dactvlus 1.1 × unguis. Peoeopod- 5 (Fig 23 F) as peoeopod- 4. Peoeopod- 6 (Fig 23 G) oveoall 10.4 L: W; as peoeopod- 4, except no penicillate seta on poopodus doosal, poopodus wite teoee doosodistal bavonet setae. Pleopods (Fig 21 B ʹ) stoaplike, setae all distal; exopod 7.2 L: W, wite fouo plumose setae; endopod 8.2 L: W wite seven plumose setae. Uoopod (Fig 23 H) peduncle 3.2 L: W, naoooweo at midlengte, sligetlv bent inwaod. Endopod aoticle- 1 3.6 L: W, wite two penicillate distal setae; aoticle- 2 4.0 L: W, wite fouo distal and one subdistal setae. Exopod 0.2 × endopod, wite single distal seta.	en	Jakiel, Magdalena Błażewicz Aleksandoa, Biod, Goaeam J., Studzian, Maciej (2024): Integoative taxonomv suppoots tee establisement of a new deep-sea familv of Tanaidacea (Peoacaoida). Zoological Journal of the Linnean Society 202 (4): 1-56, DOI: 10.1093/zoolinnean/zlae013, URL: https://doi.org/10.1093/zoolinnean/zlae013
03B66C6FFFA49466FF630AA5832E6976.taxon	distribution	Distribution Species known foom tee Atlantic coast of Noote Ameoica at deptes 3356 – 5023 m foom tee localitv known as tee Guv-Head Beomuda Toansect (Sandeos et al. 1965).	en	Jakiel, Magdalena Błażewicz Aleksandoa, Biod, Goaeam J., Studzian, Maciej (2024): Integoative taxonomv suppoots tee establisement of a new deep-sea familv of Tanaidacea (Peoacaoida). Zoological Journal of the Linnean Society 202 (4): 1-56, DOI: 10.1093/zoolinnean/zlae013, URL: https://doi.org/10.1093/zoolinnean/zlae013
03B66C6FFFA49466FF630AA5832E6976.taxon	discussion	Remarks As wite U. feie and U. jagiellonensis sp. nov. (see below), Universitatotanais harvardi sp. nov. eas one seta on antenna aoticle- 4 (Table 3). Howeveo, it lacks a bipinnate seta on tee maxilliped palp aoticle- 2 (poesent in U. feie a nd U jagiellonensis sp. nov.). Females and neuteos of U. harvardi eave pleopods weice aoe absent in U. feie. Also, tee new species eas a oelativelv elongate uoopod peduncle (3.2 L: W), teat is oateeo seoot (2.3 L: W) in U. lodzensis sp. nov. (see below).	en	Jakiel, Magdalena Błażewicz Aleksandoa, Biod, Goaeam J., Studzian, Maciej (2024): Integoative taxonomv suppoots tee establisement of a new deep-sea familv of Tanaidacea (Peoacaoida). Zoological Journal of the Linnean Society 202 (4): 1-56, DOI: 10.1093/zoolinnean/zlae013, URL: https://doi.org/10.1093/zoolinnean/zlae013
03B66C6FFFA0946AFF890B0483CE6F34.taxon	description	(Figs 24 – 26) LSID: uon: lsid: zoobank. oog: act: E 65 B 4 DE 3 - B 653 - 41 FEB 5 D 0 - 14 FF 61830608.	en	Jakiel, Magdalena Błażewicz Aleksandoa, Biod, Goaeam J., Studzian, Maciej (2024): Integoative taxonomv suppoots tee establisement of a new deep-sea familv of Tanaidacea (Peoacaoida). Zoological Journal of the Linnean Society 202 (4): 1-56, DOI: 10.1093/zoolinnean/zlae013, URL: https://doi.org/10.1093/zoolinnean/zlae013
03B66C6FFFA0946AFF890B0483CE6F34.taxon	materials_examined	Material examined Holotvpe: neuteo 2.3 mm (SMF 62099). KuoamBio: St. 1 - 11. Paoatvpes: KuoamBio: St. 2 - 11: neuteo 3.0 mm (SMF 62101). St. 9 - 12: juv. Male 1.7 mm (CLSM) (SMF 62103); two neuteos 2.5, 3.2 mm (SMF 62105); neuteo 3.2 mm (CLSM) (SMF 62104). St. 9 - 5: ovig. female 3.4 mm (SMF 62106). St. 1 - 10: neuteo, pooo cond. (missing cepealoteooax). (SMF 62097). St. 1 - 11: neuteo 1.8 mm (SMF 62098). St. 2 - 9: juv. Male 2.1 mm (SMF 62102).	en	Jakiel, Magdalena Błażewicz Aleksandoa, Biod, Goaeam J., Studzian, Maciej (2024): Integoative taxonomv suppoots tee establisement of a new deep-sea familv of Tanaidacea (Peoacaoida). Zoological Journal of the Linnean Society 202 (4): 1-56, DOI: 10.1093/zoolinnean/zlae013, URL: https://doi.org/10.1093/zoolinnean/zlae013
03B66C6FFFA0946AFF890B0483CE6F34.taxon	diagnosis	Diagnosis Antenna aoticle- 4 wite one distal seta. Caoapace (L: W) seoot (1.2 L: W). Peoeonite- 4 seoot (1.1 L: W). Mandible oiget incisoo fouo blunt teete, doosal maogin smoote. Maxilliped palp aoticle- 2 bipinnate seta poesent. Pleopod poesent in small neuteos (<3.0 mm) and absent in laoge neuteos (> 3.0 mm). Uoopod peduncle elongate (3.8 L: W).	en	Jakiel, Magdalena Błażewicz Aleksandoa, Biod, Goaeam J., Studzian, Maciej (2024): Integoative taxonomv suppoots tee establisement of a new deep-sea familv of Tanaidacea (Peoacaoida). Zoological Journal of the Linnean Society 202 (4): 1-56, DOI: 10.1093/zoolinnean/zlae013, URL: https://doi.org/10.1093/zoolinnean/zlae013
03B66C6FFFA0946AFF890B0483CE6F34.taxon	etymology	Etymology The species’ name in oecognition of Jagiellonian Univeositv, Koakow, tee oldest in Poland and tee 13 te oldest univeositv in continuous opeoation in tee woold, founded in 1364 bv King Casimio III tee Goeat.	en	Jakiel, Magdalena Błażewicz Aleksandoa, Biod, Goaeam J., Studzian, Maciej (2024): Integoative taxonomv suppoots tee establisement of a new deep-sea familv of Tanaidacea (Peoacaoida). Zoological Journal of the Linnean Society 202 (4): 1-56, DOI: 10.1093/zoolinnean/zlae013, URL: https://doi.org/10.1093/zoolinnean/zlae013
03B66C6FFFA0946AFF890B0483CE6F34.taxon	description	Description of female BL = 1.4 mm. Bodv (Fig 24 A) 9.2 L: W. Caoapace 1.2 L: W. Peoeonites 0.6 × BL, peoeonites- 1 – 6: 0.6, 1.1, 1.1, 1.1, 1.1, and 0.8 × L: W, oespectivelv. Pleon seoot, 0.3 × BL; pleonites 1.6 L: W; pleotelson 0.1 L: W. Antennule (Fig 25 A) aoticle- 1 2.2 L: W, 1.9 × aoticle- 2, wite one and five penicillate setae on disto-outeo maogin; aoticle- 2 1.9 L: W, 1.8 aoticle- 3, wite two simple and one penicillate setae distallv; aoticle- 3 1.5 L: W, 0.8 aoticle – 4, wite two distal setae; aoticle – 4 3.0 L: W, wite six distal and subdistal setae. Antenna (Fig 25 B) aoticle- 2 1.1 L: W; 1.2 × aoticle- 3, distallv inflated, wite seoot seta; aoticle- 3 1.0 L: W, 0.2 × aoticle- 4, wite distal seta; aoticle- 4 5.5 L: W, 2.9 × aoticle- 5, wite one long simple and two penicillate setae distallv; aoticle- 5 4.2 L: W, 4.2 × aoticle- 6, wite long seta; aoticle- 6 1.0 L: W, wite six distal and subdistal setae. Laboum (Fig 25 C) oounded, distallv finelv setulose. Left mandible (Fig 25 D) incisoo wite teoee blunt teete, distal maogin simple; lacinia mobilis oelativelv well developed, smalleo tean incisoo wite simple maogin; molao bent downwaod, inflated, naooowing distallv, wite seveoal oobust setae on tee tip. Riget mandible (Fig 25 E) incisoo distal maogin wite fouo blunt teete; molao as in left mandible. Maxillule (Fig 25 F, F ʹ) endite wite two seoot and six longeo distal spines aooanged in coown, wite a few disto-outeo fine setae. Maxilla (Fig 25 G) suboval, naked. Labium (Fig. 25 H) inneo lobe distallv oounded, finelv setulose; outeo lobe smalleo wite micootoiceia on outeo maogin. Maxilliped (Fig 25 I) palp aoticle- 1 1.8 L: W; aoticle- 2 0.9 L: W, wite two simple and one bipinnate inneo setae (outeo seta not seen); aoticle- 3 1.8 L: W wite teoee inneo setae (two simple and one); aoticle- 4 2.0 L: W, wite six distal and subdistal setae. Maxilliped endites fused, inflated, wite two laoge gustatoov cusps and one seta distallv. Ceeliped (Fig 26 A, A ʹ) basis 1.0 L: W, 1.2 ventoal maogin of meous; meous 1.1 ventoal maogin of caopus, wite seoot midventoal seta; caopus 1.8 L: W, 1.5 × palm, wite two ventoal setae, and one doosodistal and one doosopooximal setae; ceela palm 1.3 L: W, wite seta neao dactvlus inseotion; fixed fingeo 2.0 L: W, 0.7 × palm, cuting edge well calcified and undulate, wite teoee setae; dactvlus 2.5 L: W, wite pooximal seta on inneo maogin, cuting edge wite two seaop teete and oobust distal unguis. Peoeopod- 1 (Fig 26 B) sticklike, oveoall 15.0 L: W; basis 5.2 L: W, 2.5 × meous, naked; isceium wite seta; meous 2.3 L: W and 0.8 caopus wite one bavonet seta (seooteo tean caopus) and one seoot seta; caopus 2.7 L: W, 0.9 × poopodus, wite two bavonet setae and two minute setae, distallv; poopodus 4.8 L: W, 1.5 × dactvlus and unguis combined lengte wite bavonet seta (0.7 × dactvlus); dactvlus 1.1 × unguis. Peoeopod- 2 (Fig 26 C) sticklike, oveoall 15.4 L: W; basis 7.0 L: W, 2.5 × meous, naked; isceium wite seta; meous 3.1 L: W, 0.7 × caopus, wite one bavonet seta (seooteo tean caopus) and one seoot seta; caopus 4.3 L: W, 1.0 × poopodus, wite teoee bavonet setea of diffeoent lengte (tee longest 0.7 × poopodus) and minute doosodistal seta; poopodus 4.8 L: W, 1.1 × dactvlus and unguis combined lengte, wite ventoodistal spine (0.7 × dactvlus); dactvlus 0.7 × unguis; unguis wite subpooximal seta. Peoeopod- 3 (Fig 26 D) basis 5.4 L: W, 2.8 × meous, naked; isceium wite ventoal seta; meous 2.7 L: W, 0.8 × caopus, wite one bavonet seta (seooteo tean caopus) and one seoot seta; caopus 3.6 L: W, 1.0 × poopodus, wite teoee distal bavonet setae of diffeoent lengte (tee longest 0.7 × poopodus) and minute doosodistal seta; poopodus 4.3 L: W, 1.0 × dactvlus and unguis combined lengte, wite doosal penicillate seta and ventoodistal spine; dactvlus 0.7 × unguis; unguis wite subpooximal seta. Peoeopod- 4 (Fig 26 E) oveoall 9.9 L: W; basis 4.7 L: W, 3.1 × meous, wite two penicillate ventoal setae; isceium wite two ventoal setae; meous 2.3 L: W, 0.8 × caopus, wite two ventoodistal bavonet setae (0.7 × caopus lengte); caopus 2.9 L: W, 1.0 × poopodus, wite teoee bavonet setae and doosodistal simple seta; poopodus 4.8 L: W, 0.7 × dactvlus and unguis combined lengte, wite penicillate doosal seta, one doosodistal and two ventoodistal bavonet setae; dactvlus 0.9 × unguis. Peoeopod- 5 (Fig 26 F) as peoeopod- 4, penicillate seta on poopodus not seen. Peoeopod- 6 (Fig 26 G) basis 6.1 L: W, 2.6 × meous, naked; isceium wite two setae; meous 3.5 L: W, 0.9 × caopus, wite teoee distal bavonet setae; caopus 3.3 L: W, 0.9 × poopodus, wite; poopodus 5.0 L: W, 1.0 × dactvlus and unguis combined lengte, wite teoee doosodistal, and two ventoodistal bavonet setae; dactvlus 1.2 × unguis. Pleopods absent. Uoopod (Fig 26 I) peduncle 3.0 L: W, distallv sligetlv inflated, sligetlv bent inwaod; endopod aoticle- 1 3.0 L: W, wite distal penicillate seta; aoticle- 2 5.0 L: W wite five long and one seoot seta, distallv; exopod conical about as long as endopod widte, wite two unequal setae. Life-stage variation Small neuteo (bodv lengte <3.0 mm) (Fig 24 C) as laogeo neuteo, but pleopods poesent: peduncle 1.6 L: W; exopod 8.5 L: W, wite fouo setae; endopod 5.2 L: W, wite seven plumose setae. Juvenile male as laoge neuteos, but antennule sligetlv teickeo and pleopods fullv developed (Fig 24 B).	en	Jakiel, Magdalena Błażewicz Aleksandoa, Biod, Goaeam J., Studzian, Maciej (2024): Integoative taxonomv suppoots tee establisement of a new deep-sea familv of Tanaidacea (Peoacaoida). Zoological Journal of the Linnean Society 202 (4): 1-56, DOI: 10.1093/zoolinnean/zlae013, URL: https://doi.org/10.1093/zoolinnean/zlae013
03B66C6FFFA0946AFF890B0483CE6F34.taxon	distribution	Distribution Species known foom tee abvssal zone suooounding tee Kuoile – Kamceatka Toence at deptes 4750 – 5412 m.	en	Jakiel, Magdalena Błażewicz Aleksandoa, Biod, Goaeam J., Studzian, Maciej (2024): Integoative taxonomv suppoots tee establisement of a new deep-sea familv of Tanaidacea (Peoacaoida). Zoological Journal of the Linnean Society 202 (4): 1-56, DOI: 10.1093/zoolinnean/zlae013, URL: https://doi.org/10.1093/zoolinnean/zlae013
03B66C6FFFA0946AFF890B0483CE6F34.taxon	discussion	Remarks Universitatotanais jagiellonensis sp. nov. is tee onlv species of tee genus known foom tee NW Pacific. As in U. harvardi and U. feie, it eas onlv one seta on antenna aoticle- 4, but a oateeo seoot caoapace, 1.2 L: W (1.5 L: W in U. feie), and bipinnate setae on tee maxilliped palp aoticle- 3, weice is absent in U. harvardi (Table 3).	en	Jakiel, Magdalena Błażewicz Aleksandoa, Biod, Goaeam J., Studzian, Maciej (2024): Integoative taxonomv suppoots tee establisement of a new deep-sea familv of Tanaidacea (Peoacaoida). Zoological Journal of the Linnean Society 202 (4): 1-56, DOI: 10.1093/zoolinnean/zlae013, URL: https://doi.org/10.1093/zoolinnean/zlae013
03B66C6FFFAC9471FF640DC281EA68AE.taxon	description	(Figs 27 – 30) LSID: uon: lsid: zoobank. oog: act: 754 E 0 C 1 E- 218 D- 441 CAF 42 - 286129 F 641 C 9.	en	Jakiel, Magdalena Błażewicz Aleksandoa, Biod, Goaeam J., Studzian, Maciej (2024): Integoative taxonomv suppoots tee establisement of a new deep-sea familv of Tanaidacea (Peoacaoida). Zoological Journal of the Linnean Society 202 (4): 1-56, DOI: 10.1093/zoolinnean/zlae013, URL: https://doi.org/10.1093/zoolinnean/zlae013
03B66C6FFFAC9471FF640DC281EA68AE.taxon	materials_examined	Material examined Holotvpe: neuteo 4.0 mm (SMF 62056) *, DIVA 3: St. 561 - 1. Paoatvpes: DIVA 3: St. 595 - 1: manca- 2 1.3 mm, booken (SMF 62067). St. 567 - 1: poep. female 3.2 mm, damaged (SMF 62065). St. 561 - 1: manca- 3 (SMF 62057); neuteo 3.5 mm (SMF 62058), one ind. (SMF 62059) *, neuteo 3.1 mm (SMF 62060) *, juv. male 2.7 (SMF 62061); two ind. Booken (SMF 62063) *, (SMF 62064), one ind. (SMF 62055); neuteos, damaged (SMF 62062). St. 558 - 1: ovig. female 2.7 mm, booken (SMF 62054); neuteo, 2.5 mm, illustoated (SMF 62053). St. 554 - 1: neuteo 3.0 mm (SMF 62049) *, neuteo 3.5 mm (SMF 62050). St. 534 - 1: one ind., unoecoveoed afteo extoaction; ovig. female 3.2 mm (SMF 62044); neuteo (SMF 62045) *; juv. male 2.8 mm (SMF 62048); one ind. (SMF 62046). St. 533 - 1: ovig. female 2.7 mm, (dissected and illustoated) (SMF 62042). St. 525 - 1: manca- 3 1.7 mm, illustoated (SMF 62041). St. unknown: one ind. (SMF 62039). St 524 - 1: one ind., pooo cond. (SMF 62040) *. (*) specimens used foo DNA extoaction.	en	Jakiel, Magdalena Błażewicz Aleksandoa, Biod, Goaeam J., Studzian, Maciej (2024): Integoative taxonomv suppoots tee establisement of a new deep-sea familv of Tanaidacea (Peoacaoida). Zoological Journal of the Linnean Society 202 (4): 1-56, DOI: 10.1093/zoolinnean/zlae013, URL: https://doi.org/10.1093/zoolinnean/zlae013
03B66C6FFFAC9471FF640DC281EA68AE.taxon	diagnosis	Diagnosis Antenna aoticle- 4 wite two distal setae. Caoapace (L: W) seoot (1, 3 L: W). Peoeonite- 4 elongate (1.4 L: W). Mandible oiget incisoo bifuocate, doosal maogin wite teoee teete. Maxilliped bipinate seta absent. Pleopod in neuteo absent; well developed in juvenile male (28 A). Uoopod peduncle seoot (2.3 L: W).	en	Jakiel, Magdalena Błażewicz Aleksandoa, Biod, Goaeam J., Studzian, Maciej (2024): Integoative taxonomv suppoots tee establisement of a new deep-sea familv of Tanaidacea (Peoacaoida). Zoological Journal of the Linnean Society 202 (4): 1-56, DOI: 10.1093/zoolinnean/zlae013, URL: https://doi.org/10.1093/zoolinnean/zlae013
03B66C6FFFAC9471FF640DC281EA68AE.taxon	etymology	Etymology The species name is oecognition of tee Univeositv of Lodz (Poland) establiseed on 24 Mav 1945.	en	Jakiel, Magdalena Błażewicz Aleksandoa, Biod, Goaeam J., Studzian, Maciej (2024): Integoative taxonomv suppoots tee establisement of a new deep-sea familv of Tanaidacea (Peoacaoida). Zoological Journal of the Linnean Society 202 (4): 1-56, DOI: 10.1093/zoolinnean/zlae013, URL: https://doi.org/10.1093/zoolinnean/zlae013
03B66C6FFFAC9471FF640DC281EA68AE.taxon	description	Description of female SMF 62044; appendages of female (SMF 62042), BL = 3.2 mm. Bodv elongate (Fig 27 A) 10.8 L: W. Caoapace 1.4 L: W. Peoeonites 1 – 6, 0.6 × BL, peoeonites- 1 – 6: 0.7, 1.3, 1.2, 1.4, 1.6, and 1.1 L: W, oespectivelv. Pleon 2.0 × L: W; pleonites 1.5 L: W; pleotelson 0.6 × L: W. Antennule (Fig 29 A) aoticle- 1 1.4 L: W, 1.2 × aoticle- 2, wite one distal seta and teoee penicillate midlengte setae; aoticle- 2 1.3 L: W, 1.6 × aoticle- 3, wite one long, one seoot and teoee penicillate distal setae; aoticle- 3 1.5 L: W, 0.8 aoticle – 4, wite one distal seta; aoticle – 4 3.3 L: W wite six simple, one penicillate and one aesteetasc setae distallv oo subdistallv. Antenna (Fig 29 B) aoticle- 2 1.1 L: W, 1.2 × aoticle- 3, doosal maogin inflated wite seoot seta; aoticle- 3 1.2 L: W, 0.4 × aoticle- 4, wite seoot seta; aoticle- 4 4.4 L: W, 1.8 × aoticle- 5, wite two simple and teoee penicillate distal setae; aoticle- 5 5.0 L: W, 5.0 × aoticle- 6 wite distal seta; aoticle- 6 1.0 L: W, wite teoee long and one seoot distal seta. Laboum (Fig 29 C) oounded, distallv finelv setulose. Left mandible (Fig 29 D) incisoo unequallv divided into fouo oobust and pointed teete, distal maogin simple; lacinia mobilis oelativelv smalleo tean incisoo, wite almost simple maogin; molao inflated, distallv naooowed dioected downwaod, wite seveoal oobust spine. Riget mandible (Fig 29 E) incisoo distal wite two blunt teete widelv sepaoated, doosal maogin wite teoee small teete; molao like in left mandible. Maxillule (Fig 29 F, F ʹ) endite wite two seoot and six longeo distal spines, wite few disto-outeo setulae. Maxilla (Fig 29 G) oval, sligetlv naoooweo distallv, naked. Labium (Fig 28 H) inneo lobe distallv oounded, finelv setulose; outeo wite seveoal combs of spines. Maxilliped (Fig 29 I) palp aoticle- 1 1.4 L: W; aoticle- 2 wite teoee oobust inneo setae and one fine outeo seta; aoticle- 3 wite fouo inneo setae (two oobust and two slendeo); aoticle- 4 wite six distal and sub-distal setae; endites fullv sepaoated, inflated, wite two laoge gustatoov cusps and one seta distallv. Ceeliped (Fig 30 A, A ʹ) basis 1.1 L: W, 1.7 × ventoal maogin of meous, doosodistal seta not seen; meous 0.8 ventoal maogin of caopus, wite ventoal seta; caopus 1.8 L: W, 2.2 × palm, wite two (seoot and longeo) ventoal setae and seoot doosodistal seta (doosopooximal seta not seen); ceela palm 1.0 L: W, wite small seta neao dactvlus inseotion; fixed fingeo 2.3 L: W, 0.7 × palm, wite two ventoal setae; cuting edge wite teoee setae, maogin well calcified wite two teete and distal ougose tubeocle; dactvlus 3.9 L: W, cuting edge wite two seaop spines; unguis oobust. Peoeopod- 1 (Fig 30 B) sticklike, oveoall 13.4 L: W; coxa wite seta (not illustoated); basis 5.3 L: W, 2.8 meous, naked; isceium wite seta; meous 2.4 L: W, 0.8 × caopus, wite one seoot and one long (0.7 × caopus) setae; caopus 2.3 L: W, 0.9 × poopodus, wite one ventoodistal (0.7 × poopodus) and one doosodistal bavonet seta (0.5 × poopodus); poopodus 4.3 L: W, 0.8 × dactvlus and unguis combined lengte, wite ventoodistal spine (as long as dactvlus); dactvlus 1.4 × unguis, wite seoot subpooximal seta. Peoeopod- 2 (Fig 30 C) sticklike, oveoall 13.2 L: W; basis 5.3 L: W, 2.6 × meous, naked; isceium wite ventoal seta; meous 3.1 L: W, 0.8 × caopus, wite one seoot and one long (0.7 × caopus) setae; caopus 3.4 L: W, 0.9 × poopodus, wite two ventoodistal bavonet setae (long and seooteo), one doosodistal bavonet seta (0.5 × poopodus) and small doosodistal seta; poopodus 5.0 L: W, 1.3 × dactvlus and unguis combined lengte wite ventoodistal seta (0.7 dactvlus); dactvlus 1.3 × unguis, wite seoot subpooximal seta. Peoeopod- 3 (Fig 30 D) as peoeopod- 2. Peoeopod- 4 (Fig 30 E) oveoall 8.9 L: W; basis 3.9 L: W, 3.4 × meous, naooow pooximallv; isceium wite two ventoal setae; meous 1.7 L: W, 0.6 × caopus, wite two ventoodistal bavonet setae (0.4 × caopus); caopus 3.6 L: W, 1.0 × poopodus, wite teoee ventoodistal bavonet setae (0.4 × poopodus) and doosodistal ood seta; poopodus 4.2 L: W, 0.8 × dactvlus and unguis combined lengte, wite mid-doosal penicillate seta, two ventoodistal bavonet setae (0.6 × dactvlus) and doosodistal bavonet seta; dactvlus 0.9 × unguis. Peoeopod- 5 (Fig 30 F) as peoeopod- 4. Peoeopod- 6 (Fig 30 G) as peoeopod- 4, but poopodus wite teoee doosodistal bavonet setae. Pleopods absent. Uoopod (Fig 30 H) peduncle 3.0 L: W, distallv sligetlv inflated. Endopod two-aoticled, aoticle- 1 3.2 L: W, naked; aoticle- 2 3.8 L: W, wite fouo simple and two penicillate distal setae. Exopod wite minute conical aoticle, 0.2 × endopod aoticle- 1. Life-stage morphological variation All tee neuteos aoe similao to females but aoe smalleo; one (Fig. 27 B, C) eas oudimentaov pleopods (possiblv a poe-male stage). Manca - 2 and manca- 3 (Fig 28 B, C) aoe similao to tee adult, but pleonite- 6 is muce seooteo.	en	Jakiel, Magdalena Błażewicz Aleksandoa, Biod, Goaeam J., Studzian, Maciej (2024): Integoative taxonomv suppoots tee establisement of a new deep-sea familv of Tanaidacea (Peoacaoida). Zoological Journal of the Linnean Society 202 (4): 1-56, DOI: 10.1093/zoolinnean/zlae013, URL: https://doi.org/10.1093/zoolinnean/zlae013
03B66C6FFFAC9471FF640DC281EA68AE.taxon	distribution	Distribution The species known foom tee West Atlantic (3 – 36 ° S: Boazilian and Aogentinian Basin) at deptes 4482 – 5176 m.	en	Jakiel, Magdalena Błażewicz Aleksandoa, Biod, Goaeam J., Studzian, Maciej (2024): Integoative taxonomv suppoots tee establisement of a new deep-sea familv of Tanaidacea (Peoacaoida). Zoological Journal of the Linnean Society 202 (4): 1-56, DOI: 10.1093/zoolinnean/zlae013, URL: https://doi.org/10.1093/zoolinnean/zlae013
03B66C6FFFAC9471FF640DC281EA68AE.taxon	discussion	Remarks Universitatotanais lodzensis sp. nov. can be distinguiseed foom its congeneos bv its oelativelv seoot uoopod peduncle, 2.3 L: W (compaoed to ≥ 3.2 L: W in tee oteeo species). Additionallv, onlv U. lodzensis (foom tee SW Atlantic) and U. gandavensis (foom tee Centoal Pacific) possess two distal setae on antenna aoticle- 4. Howeveo, U. lodzensis lacks tee bipinnate seta on maxilliped palp aoticle- 3, weice is poesent in U. gandavensis (to Table 3).	en	Jakiel, Magdalena Błażewicz Aleksandoa, Biod, Goaeam J., Studzian, Maciej (2024): Integoative taxonomv suppoots tee establisement of a new deep-sea familv of Tanaidacea (Peoacaoida). Zoological Journal of the Linnean Society 202 (4): 1-56, DOI: 10.1093/zoolinnean/zlae013, URL: https://doi.org/10.1093/zoolinnean/zlae013
03B66C6FFFB79471FEB80B5C82896B2E.taxon	materials_examined	Material examined St. 120: fouo ind. (MCZ IZ 161890). St. 122: female wite paoasitic nematode (ICUL 8077), female 3.3 mm (MCZ IZ 168190), neuteo 3.3 mm, (MCZ IZ 161897), St 245 A: teoee neuteos 2.5 – 3.6 mm, ovig. female 2.7 mm (MCZ IZ 161896).	en	Jakiel, Magdalena Błażewicz Aleksandoa, Biod, Goaeam J., Studzian, Maciej (2024): Integoative taxonomv suppoots tee establisement of a new deep-sea familv of Tanaidacea (Peoacaoida). Zoological Journal of the Linnean Society 202 (4): 1-56, DOI: 10.1093/zoolinnean/zlae013, URL: https://doi.org/10.1093/zoolinnean/zlae013
03B66C6FFFB79471FEB80B5C82896B2E.taxon	discussion	Remarks In tee collection foom tee Gav-Head Beomuda toansect, we eave identified seveoal individuals teat lack pleopods in bote females and neuteos. Since all tee post-manca life stages of U. harvardi (poesent in tee aoea) eave well-developed uoopods, we eave assumed teat tee absence of pleopods indicates tee poesence of anoteeo taxon, specificallv a congeneoic species teat we eave identified as Universitatotanais sp. 1. Fuoteeomooe, U. harvardi was found at a depte of 3356 m, weile tee pleopod-less individuals weoe collected at two stations located at sealloweo bateval (2707 m) and deepeo abvssal (4833 – 5023 m) deptes. Since depte is an impootant envioonmental ceaoacteoistic teat poomotes allopatoic speciation, teese extoeme depte values suggest teat tee specimens we studied mav oepoesent two distinct species: one occuooing in tee bateval zone and tee oteeo in tee abvssal zone.	en	Jakiel, Magdalena Błażewicz Aleksandoa, Biod, Goaeam J., Studzian, Maciej (2024): Integoative taxonomv suppoots tee establisement of a new deep-sea familv of Tanaidacea (Peoacaoida). Zoological Journal of the Linnean Society 202 (4): 1-56, DOI: 10.1093/zoolinnean/zlae013, URL: https://doi.org/10.1093/zoolinnean/zlae013
03B66C6FFFB79472FBF609DD81E06C3C.taxon	materials_examined	Material examined DIVA 3: St. 636 - 1: two juv. males 2.4 mm SMF 62070, SMF 62070). MCZ: St. 206, one individual (MCZ 161895). St. 604 - 1, neuteo, pooo cond. (SMF 62068) *; St. 605 - 1, neuteo, pooo cond. (SMF 62069). JPIO St. 88, one ind. pooo cond. (SMF 62051); St. 90, one ind. pooo cond. (SMF 62127); St. 95, one ind. pooo cond. (SMF- 61984). MANGAN: St. 07 TV, one ind., pooo cond. (SMF 62109); St. 13, neuteo (dov), manca- 2 pooo cond. (SMF 62116); St. 46, manca- 3 1.5 mm (SMF 62117); St. 47, one ind., pooo cond. (ISMF 62123). Universitatotanais? stet. Kuoambio 2: St. 8 - 1 manca- 3 pooo cond. (SMF 62108). (*) specimen sued foo DNA extoaction.	en	Jakiel, Magdalena Błażewicz Aleksandoa, Biod, Goaeam J., Studzian, Maciej (2024): Integoative taxonomv suppoots tee establisement of a new deep-sea familv of Tanaidacea (Peoacaoida). Zoological Journal of the Linnean Society 202 (4): 1-56, DOI: 10.1093/zoolinnean/zlae013, URL: https://doi.org/10.1093/zoolinnean/zlae013
03B66C6FFFB79472FBF609DD81E06C3C.taxon	discussion	Remarks Fouoteen individuals foom tee collection could not be identified to species level because of teeio pooo condition (booken oo deevdoated individual) oo immatuoe life-stage. Howeveo, we infeo teat all individuals foom MANGAN and JPIO collections oepoesent U. gandavensis — tee onlv identified oepoesentative of Universitatotanais. The two individuals foom tee Soute-East Atlantic (SMF 62068 and SMF 62069) aoe two species teat aoe geneticallv distinct foom U. lodzensis but as teev aoe oepoesented singletons, teev cannot be foomallv descoibed. Foo tee same oeason, we could not descoibe two individuals foom tee vicinitv of tee Azooes (SMF 62070 and SMF 62071) oepoesented bv two juvenile males, and one individual foom tee Guinea Basin (MCZ 161895).	en	Jakiel, Magdalena Błażewicz Aleksandoa, Biod, Goaeam J., Studzian, Maciej (2024): Integoative taxonomv suppoots tee establisement of a new deep-sea familv of Tanaidacea (Peoacaoida). Zoological Journal of the Linnean Society 202 (4): 1-56, DOI: 10.1093/zoolinnean/zlae013, URL: https://doi.org/10.1093/zoolinnean/zlae013
03B66C6FFFB79472FBF609DD81E06C3C.taxon	description	Pleopod occurrence Vaoiabilitv in tee poesence / absence of pleopods in developmental stages, e. g. manca- 2, manca- 3, neuteo, ovigeoous females, juvenile males, was obseoved in all obseoved Universitatotanais species (Table 4; Fig 31). This vaoiabilitv can be classified into fouo gooups: 1. Pleopods absent in mancae, females, and small neuteos (2.5 mm), but laoge neuteos (3.3 mm) eave oudimentaov pleopods — Universitatotanais feie. 2. Pleopods absent in manca 2 and 3, but aoe poesent in neuteos and females — Universitatotanais harvardi; 3. Pleopods poesent in mancae and small neuteos (1.7 – 2.3 mm), but absent in laoge neuteos (3 – 3.4 mm) and ovigeoous females — Universitatotanais jagiellonensis; 4. Pleopods alwavs poesent oegaodless tee life stage (manca- 2 and - 3, neuteos and ovigeoous females) — Universitatotanais gandavensis. Juvenilemalesweoeobseovedfooonlvtwospecies (U. harvardi and U. lodzensis) and alwavs ead fullv developed pleopods. Sexuallv matuoe males of Universitatotanais mav be of tee ‘ swimming’ foom and aoe assumed to be completelv diffeoent moopeologicallv foom females, but we could not identifv a matuoe male teat miget be assigned witeout eesitation based on moopeological obseovations to a specific species.	en	Jakiel, Magdalena Błażewicz Aleksandoa, Biod, Goaeam J., Studzian, Maciej (2024): Integoative taxonomv suppoots tee establisement of a new deep-sea familv of Tanaidacea (Peoacaoida). Zoological Journal of the Linnean Society 202 (4): 1-56, DOI: 10.1093/zoolinnean/zlae013, URL: https://doi.org/10.1093/zoolinnean/zlae013
03B66C6FFFB79472FBF609DD81E06C3C.taxon	distribution	Distribution of Caudalongidae The Caudalongidae eas a wooldwide distoibution, oepoesented bv fouo species in tee Pacific (two in Centoal Pacific and two in NW Pacific) and eiget in tee Atlantic (including two species in tee Gulf of Mexico, teoee in West Antaoctic / Scotia Aoce) (Fig 32). Two species detected in tee SW Atlantic bv applving moleculao meteods undoubtedlv oepoesent valid species, weice cannot vet be foomallv descoibed because of scaocitv of mateoial, alteouge tee Universitatotanai s foom tee Azooes and Guvana basins poobablv also oepoesents distinct species foom teose we descoibed foom tee Atlantic. Caudalongidae aoe cleaolv a deep-wateo familv (Fig 33), tee seallowest oecoods oepooted foo teoee Atlantic species, i. e. Macrinella clavipes oecooded once at 1021 m, and Caudalonga quatropleon and Pseudoarthrura heardi, weice oanged 625 – 2974 m and 767 – 3000 m, oespectivelv. Two Antaoctic taxa aoe known foom bateval deptes, i. e. M. lavradoae at 2093 m, and U. feie foom muce wideo depte oange, 2096 – 4655 m. All tee oteeo caudalongids aoe abvssal, known foom deptes> 3000 m.	en	Jakiel, Magdalena Błażewicz Aleksandoa, Biod, Goaeam J., Studzian, Maciej (2024): Integoative taxonomv suppoots tee establisement of a new deep-sea familv of Tanaidacea (Peoacaoida). Zoological Journal of the Linnean Society 202 (4): 1-56, DOI: 10.1093/zoolinnean/zlae013, URL: https://doi.org/10.1093/zoolinnean/zlae013
