taxonID	type	description	language	source
03B487EDFFEDF52A96DD84A9FA610CFA.taxon	type_taxon	Type species. Empoasca polyphemus Matsumura, 1931, by original designation of Ishiharella Dworakowska, 1970.	en	Wang, Ran, Zhang, Yalin, Cao, Yanghui (2025): Review of the genus Ishiharella Dworakowska (Hemiptera: Cicadellidae: Typhlocybinae), with description of two new species. Zootaxa 5604 (2): 176-184, DOI: 10.11646/zootaxa.5604.2.7, URL: https://doi.org/10.11646/zootaxa.5604.2.7
03B487EDFFEDF52A96DD84A9FA610CFA.taxon	description	Description. Body robust, cylindrical. General color sordid ochre to fuscous. Head with a central dark patch at the transition from vertex to face and pair of slim, bracket-like depressions on each side, sometimes with a dark patch in the center of crown posterior margin. Anteclypeus dark apically. Thorax with pair of slim, sinuated depressions on each side. Ocelli present. Face broad, width across eyes nearly equal to length, frontoclypeus swollen. Profile of transition to face rounded. Crown short and wide, rounded anteriorly, in dorsal view distinctly shorter than width between eyes, anterior and posterior margins subparallel, coronal suture present, usually indistinct. Pronotum large, lateral part general with a sinuate transverse depression on each side. Forewing narrow, rounded apically, 2 nd apical cell broad at base and narrowed towards apex, veins RP, MP’ stalked, RP, MP’ and MP’ ’ + CuA’ arise from m cell; c and r cells narrower than m and cua cells. Hindwing with CuA unbranched. Coloration of female same as male except abdomen brownish red, ovipositor brown. Front femur intercalary row with 7 – 9 fine setae and 1 relatively large basal seta more distad, extended nearly half length of femur; row with 9 – 14 posteroventral setae (PV) situated on the whole tibia in a row from short to long and 1 pair of setae situated on apex of tibia; tarsus elongate, approximately one third length of tibia. Middle femur row PV slim, with 20 – 25 short, small setae in ventral view; tarsus almost same as tarsus in fore leg. Hind femur with macrosetal formula 2 + 1 + 1, tibia anteroventral row (AV) with 6 preapical macrosetae; anterodorsal setae (AD) and posterodorsal setae (PD) numerous, almost same size, both longer than posteroventral setae (PV), and row AV with 6 – 9 macrosetae near apex; tarsus longer than front and middle tarsus. Male basal abdominal apodemes not greatly enlarged. Male pygofer elongated, with ventral margins folded inward and forming a small process distally, with macrosetae caudally, ventral appendage absent. Dorsal bridge of pygofer short, about 1 / 3 as the length of pygofer lobe. Subgenital plate partly to completely fused, lateral margins infolded on both sides, with row of sublateral macrosetae. Basal group setae absent. Paramere elongate, apex variable interspecifically, spirally twisted, or bifurcated at apex and bearing a tooth basad of bifurcation. Aedeagus shaft tubular, structure complex, with processes basally. Anal tube process developed or short and truncated apically. Connective lamellate caudally and crimped basally.	en	Wang, Ran, Zhang, Yalin, Cao, Yanghui (2025): Review of the genus Ishiharella Dworakowska (Hemiptera: Cicadellidae: Typhlocybinae), with description of two new species. Zootaxa 5604 (2): 176-184, DOI: 10.11646/zootaxa.5604.2.7, URL: https://doi.org/10.11646/zootaxa.5604.2.7
03B487EDFFEDF52A96DD84A9FA610CFA.taxon	discussion	Notes. Previous studies indicated that the coronal suture was absent, but the samples examined which were boiled the whole body in NaOH showed that the coronal suture can be observed.	en	Wang, Ran, Zhang, Yalin, Cao, Yanghui (2025): Review of the genus Ishiharella Dworakowska (Hemiptera: Cicadellidae: Typhlocybinae), with description of two new species. Zootaxa 5604 (2): 176-184, DOI: 10.11646/zootaxa.5604.2.7, URL: https://doi.org/10.11646/zootaxa.5604.2.7
03B487EDFFEDF52A96DD84A9FA610CFA.taxon	distribution	Distribution. China (Anhui, Chongqing, Guangdong, Guangxi, Guizhou, Hunan, Hubei, Shaanxi, Yunnan, Zhejiang); Thailand; Japan; Russia.	en	Wang, Ran, Zhang, Yalin, Cao, Yanghui (2025): Review of the genus Ishiharella Dworakowska (Hemiptera: Cicadellidae: Typhlocybinae), with description of two new species. Zootaxa 5604 (2): 176-184, DOI: 10.11646/zootaxa.5604.2.7, URL: https://doi.org/10.11646/zootaxa.5604.2.7
03B487EDFFEEF52B96DD86A1FE180971.taxon	distribution	Distribution. China (Shaanxi)	en	Wang, Ran, Zhang, Yalin, Cao, Yanghui (2025): Review of the genus Ishiharella Dworakowska (Hemiptera: Cicadellidae: Typhlocybinae), with description of two new species. Zootaxa 5604 (2): 176-184, DOI: 10.11646/zootaxa.5604.2.7, URL: https://doi.org/10.11646/zootaxa.5604.2.7
03B487EDFFEEF52B96DD8711FDB109E1.taxon	distribution	Distribution. China (Yunnan, Guizhou)	en	Wang, Ran, Zhang, Yalin, Cao, Yanghui (2025): Review of the genus Ishiharella Dworakowska (Hemiptera: Cicadellidae: Typhlocybinae), with description of two new species. Zootaxa 5604 (2): 176-184, DOI: 10.11646/zootaxa.5604.2.7, URL: https://doi.org/10.11646/zootaxa.5604.2.7
03B487EDFFEEF52B96DD8781FE150A51.taxon	distribution	Distribution. Japan (Ryukyus)	en	Wang, Ran, Zhang, Yalin, Cao, Yanghui (2025): Review of the genus Ishiharella Dworakowska (Hemiptera: Cicadellidae: Typhlocybinae), with description of two new species. Zootaxa 5604 (2): 176-184, DOI: 10.11646/zootaxa.5604.2.7, URL: https://doi.org/10.11646/zootaxa.5604.2.7
03B487EDFFEEF52B96DD84F1FD6E0AC1.taxon	distribution	Distribution. Thailand (Nakhon, Phetchabun)	en	Wang, Ran, Zhang, Yalin, Cao, Yanghui (2025): Review of the genus Ishiharella Dworakowska (Hemiptera: Cicadellidae: Typhlocybinae), with description of two new species. Zootaxa 5604 (2): 176-184, DOI: 10.11646/zootaxa.5604.2.7, URL: https://doi.org/10.11646/zootaxa.5604.2.7
03B487EDFFEEF52B96DD8561FE160BB1.taxon	distribution	Distribution. China (Guizhou)	en	Wang, Ran, Zhang, Yalin, Cao, Yanghui (2025): Review of the genus Ishiharella Dworakowska (Hemiptera: Cicadellidae: Typhlocybinae), with description of two new species. Zootaxa 5604 (2): 176-184, DOI: 10.11646/zootaxa.5604.2.7, URL: https://doi.org/10.11646/zootaxa.5604.2.7
03B487EDFFEEF52B96DD85D1FDD70C21.taxon	distribution	Distribution. Thailand (Phetchabun)	en	Wang, Ran, Zhang, Yalin, Cao, Yanghui (2025): Review of the genus Ishiharella Dworakowska (Hemiptera: Cicadellidae: Typhlocybinae), with description of two new species. Zootaxa 5604 (2): 176-184, DOI: 10.11646/zootaxa.5604.2.7, URL: https://doi.org/10.11646/zootaxa.5604.2.7
03B487EDFFEEF52B96DD8241FE220C91.taxon	distribution	Distribution. China (Yunnan)	en	Wang, Ran, Zhang, Yalin, Cao, Yanghui (2025): Review of the genus Ishiharella Dworakowska (Hemiptera: Cicadellidae: Typhlocybinae), with description of two new species. Zootaxa 5604 (2): 176-184, DOI: 10.11646/zootaxa.5604.2.7, URL: https://doi.org/10.11646/zootaxa.5604.2.7
03B487EDFFEEF52B96DD82B1FDBA0D65.taxon	distribution	Distribution. China (Zhejiang, Hunan)	en	Wang, Ran, Zhang, Yalin, Cao, Yanghui (2025): Review of the genus Ishiharella Dworakowska (Hemiptera: Cicadellidae: Typhlocybinae), with description of two new species. Zootaxa 5604 (2): 176-184, DOI: 10.11646/zootaxa.5604.2.7, URL: https://doi.org/10.11646/zootaxa.5604.2.7
03B487EDFFEEF52B96DD83FDFE210DD5.taxon	distribution	Distribution. Thailand (Loei)	en	Wang, Ran, Zhang, Yalin, Cao, Yanghui (2025): Review of the genus Ishiharella Dworakowska (Hemiptera: Cicadellidae: Typhlocybinae), with description of two new species. Zootaxa 5604 (2): 176-184, DOI: 10.11646/zootaxa.5604.2.7, URL: https://doi.org/10.11646/zootaxa.5604.2.7
03B487EDFFEEF52B96DD806DFD5D0E45.taxon	distribution	Distribution. China (Zhejiang, Fujian, Jiangxi)	en	Wang, Ran, Zhang, Yalin, Cao, Yanghui (2025): Review of the genus Ishiharella Dworakowska (Hemiptera: Cicadellidae: Typhlocybinae), with description of two new species. Zootaxa 5604 (2): 176-184, DOI: 10.11646/zootaxa.5604.2.7, URL: https://doi.org/10.11646/zootaxa.5604.2.7
03B487EDFFEEF52B96DD80DDFCCA0F19.taxon	distribution	Distribution. Russia (Caucasus), Japan (Hokkaido, Honshu)	en	Wang, Ran, Zhang, Yalin, Cao, Yanghui (2025): Review of the genus Ishiharella Dworakowska (Hemiptera: Cicadellidae: Typhlocybinae), with description of two new species. Zootaxa 5604 (2): 176-184, DOI: 10.11646/zootaxa.5604.2.7, URL: https://doi.org/10.11646/zootaxa.5604.2.7
03B487EDFFEEF52B96DD8129FDE50FA5.taxon	distribution	Distribution. India (Uttar Pradesh)	en	Wang, Ran, Zhang, Yalin, Cao, Yanghui (2025): Review of the genus Ishiharella Dworakowska (Hemiptera: Cicadellidae: Typhlocybinae), with description of two new species. Zootaxa 5604 (2): 176-184, DOI: 10.11646/zootaxa.5604.2.7, URL: https://doi.org/10.11646/zootaxa.5604.2.7
03B487EDFFEFF52A96DD86E8FE1608E5.taxon	distribution	Distribution. China (Guangxi)	en	Wang, Ran, Zhang, Yalin, Cao, Yanghui (2025): Review of the genus Ishiharella Dworakowska (Hemiptera: Cicadellidae: Typhlocybinae), with description of two new species. Zootaxa 5604 (2): 176-184, DOI: 10.11646/zootaxa.5604.2.7, URL: https://doi.org/10.11646/zootaxa.5604.2.7
03B487EDFFEFF52A96DD877CFDFA0971.taxon	distribution	Distribution. China (Chongqing)	en	Wang, Ran, Zhang, Yalin, Cao, Yanghui (2025): Review of the genus Ishiharella Dworakowska (Hemiptera: Cicadellidae: Typhlocybinae), with description of two new species. Zootaxa 5604 (2): 176-184, DOI: 10.11646/zootaxa.5604.2.7, URL: https://doi.org/10.11646/zootaxa.5604.2.7
03B487EDFFEFF52D96DD8363FDD00F96.taxon	description	(Figs 1 – 4, 9 – 10, 12 – 15, 17 – 28)	en	Wang, Ran, Zhang, Yalin, Cao, Yanghui (2025): Review of the genus Ishiharella Dworakowska (Hemiptera: Cicadellidae: Typhlocybinae), with description of two new species. Zootaxa 5604 (2): 176-184, DOI: 10.11646/zootaxa.5604.2.7, URL: https://doi.org/10.11646/zootaxa.5604.2.7
03B487EDFFEFF52D96DD8363FDD00F96.taxon	materials_examined	Type material: Holotype. ♂ (NWAFU), China, Guangxi, Hechi, Huanjiang Maonan Autonomous County, 23 - VI- 2022, Ran Wang. Paratypes. 13 ♂♂ (NWAFU), same data as holotype.	en	Wang, Ran, Zhang, Yalin, Cao, Yanghui (2025): Review of the genus Ishiharella Dworakowska (Hemiptera: Cicadellidae: Typhlocybinae), with description of two new species. Zootaxa 5604 (2): 176-184, DOI: 10.11646/zootaxa.5604.2.7, URL: https://doi.org/10.11646/zootaxa.5604.2.7
03B487EDFFEFF52D96DD8363FDD00F96.taxon	description	Description. Length: male 4.0 – 4.3 mm. General color red brown. Area between eyes with two pigmented depressions and yellowish patches, posterior margin with median brownish patch. Eyes black. Ocelli white. Face whitish yellow, anteclypeus with apex black. Pronotum dark centrally, with black sinuate transverse depression on each side. Scutellum yellow, apex black, scutoscutellar sulcus black brownish. Abdomen slightly brown infuscated in midline dorsally. Forewing brown, hind wing hyaline. Legs sordid except claws black (Figs 1 – 4). Male 2 S abdominal apodemes reaching basal of segment IV, basal apodemes broad, apically narrowed (Figs 13, 15). Male pygofer brown, strongly narrowed in apical half, with filamentous setae on outer face near dorsal margin, apex pointed and curved upward, caudoventral protrusion of pygofer short, in lateral view not surpassing the end of pygofer side, in dorsal aspect skeletonized and incised subbasally (Figs 9, 12, 14, 17, 19). Subgenital plate surpassing pygofer lobe, separated in basal 1 / 3, slightly expanded basally, apex rounded; ca. 24 macrosetae uniseriate in most part; arising near basal 1 / 4 of plate with 6 – 7 feeble macrosetae in alignment, reaching apex (Figs 9, 12, 14, 17 – 18). Aedeagal shaft slim and long, with even width from base to apex, gonopore vterminal; dorsoatrium shorter than shaft, near the apex of the dorsally protrusion bifurcated, spread out to the sides extended, with a slightly curved apex; basoventral protrusion of aedeagus much longer and wider than shaft, broadened basally and tapering to apex in lateral view, basally with a broad cavity which is connected with the connective basoventrally (Figs 9, 10, 12, 14, 20, 21). Connective lamellate caudally and crimped basally, caudal margin concave medially (Figs 9, 26). Paramere almost the same length with pygofer, subbasally adorned with 3 – 4 teeth laterally, bifurcated at nearly apex and bearing a lamellar process basad of bifurcation (Figs 9, 27). Anal tube appendage developed, skeletonized, curved toward apex (Figs 9, 28).	en	Wang, Ran, Zhang, Yalin, Cao, Yanghui (2025): Review of the genus Ishiharella Dworakowska (Hemiptera: Cicadellidae: Typhlocybinae), with description of two new species. Zootaxa 5604 (2): 176-184, DOI: 10.11646/zootaxa.5604.2.7, URL: https://doi.org/10.11646/zootaxa.5604.2.7
03B487EDFFEFF52D96DD8363FDD00F96.taxon	discussion	Notes. This new species is similar to I. paradentata Lu & Qin, but differs in male genitalia (features of I. paradentata in parentheses): dorsoatrium of aedeagus skeletonized with bifurcates and curved apically (dorsoatrium slightly radian, skeletonized basally and membranous apically) (Figs 20 – 21); aedeagal shaft glabrate, without small teeth (aedeagal shaft with 4 – 5 small teeth apically) (Figs 20 – 21); caudoventral protrusion of pygofer short and not surpassing the end of pygofer side, curved dorsad and without teeth (caudoventral protrusion of pygofer significantly surpassing the end of pygofer side, ventrally bearing a tooth medially) (Figs 17, 19). It is noteworthy that the variation of aedeagal dorsoatrium is evident (Figs 20 – 25) although the other structures of male genitalia are nearly identical among samples. The aedeagal dorsoatrium of one paratype bifurcates at the base with a dorsally protruding curved apex turning left in dorsal view (Figs 22 – 23). In another paratype, although dorsoatrium bifurcates near the apex, apicad of which is strongly curved (Figs 10, 14, 24 – 25).	en	Wang, Ran, Zhang, Yalin, Cao, Yanghui (2025): Review of the genus Ishiharella Dworakowska (Hemiptera: Cicadellidae: Typhlocybinae), with description of two new species. Zootaxa 5604 (2): 176-184, DOI: 10.11646/zootaxa.5604.2.7, URL: https://doi.org/10.11646/zootaxa.5604.2.7
03B487EDFFEFF52D96DD8363FDD00F96.taxon	etymology	Etymology. The name is derived from the Latin word “ trifurcatus ” (branch), referring to the trifurcate aedeagal processes of the type specimens.	en	Wang, Ran, Zhang, Yalin, Cao, Yanghui (2025): Review of the genus Ishiharella Dworakowska (Hemiptera: Cicadellidae: Typhlocybinae), with description of two new species. Zootaxa 5604 (2): 176-184, DOI: 10.11646/zootaxa.5604.2.7, URL: https://doi.org/10.11646/zootaxa.5604.2.7
03B487EDFFEFF52D96DD8363FDD00F96.taxon	distribution	Distribution. China (Guangxi).	en	Wang, Ran, Zhang, Yalin, Cao, Yanghui (2025): Review of the genus Ishiharella Dworakowska (Hemiptera: Cicadellidae: Typhlocybinae), with description of two new species. Zootaxa 5604 (2): 176-184, DOI: 10.11646/zootaxa.5604.2.7, URL: https://doi.org/10.11646/zootaxa.5604.2.7
03B487EDFFE9F52F96DD8278FDB40EE2.taxon	description	(Figs 5 – 8, 11, 16, 29 – 36)	en	Wang, Ran, Zhang, Yalin, Cao, Yanghui (2025): Review of the genus Ishiharella Dworakowska (Hemiptera: Cicadellidae: Typhlocybinae), with description of two new species. Zootaxa 5604 (2): 176-184, DOI: 10.11646/zootaxa.5604.2.7, URL: https://doi.org/10.11646/zootaxa.5604.2.7
03B487EDFFE9F52F96DD8278FDB40EE2.taxon	materials_examined	Type material: Holotype. ♂ (NWAFU), China, Chongqing, Wuxi County, 17 - VIII- 2023, Qingquan Xue. Paratypes. 3 ♂♂ (NWAFU), same data as holotype.	en	Wang, Ran, Zhang, Yalin, Cao, Yanghui (2025): Review of the genus Ishiharella Dworakowska (Hemiptera: Cicadellidae: Typhlocybinae), with description of two new species. Zootaxa 5604 (2): 176-184, DOI: 10.11646/zootaxa.5604.2.7, URL: https://doi.org/10.11646/zootaxa.5604.2.7
03B487EDFFE9F52F96DD8278FDB40EE2.taxon	description	Description. Length: male 3.8 – 4.0 mm. Crown yellowish orange. Vertex with the central subcircular black patch at base blackish, submedially with a small brownish spot on each side. Eyes dark, ocelli white. Face yellow, except anteclypeus black apically. Pronotum laterally with sinuate transverse depressions black brown on each side. Scutellum yellow, apex slightly black, scutoscutellar sulcus brownish. Abdomen dark brown infuscated in dorsally. Forewing brown, hind wing hyaline. Legs yellowish except claws black (Figs 5 – 8). Male 2 S abdominal apodemes reaching the posterior margin of segment IIV, basal apodemes broad, apically narrowed (Fig. 29). Male pygofer brown, elongated in lateral view, narrowed in apical 1 / 5, terminally with 10 rigid microsetae on each side of lobe, with ca. 8 filamentous setae on outer face near dorsal margin, caudoventral protrusion of pygofer short, in lateral view not surpassing the end of pygofer side and incised subbasally (Figs 11, 16, 30). Subgenital plate surpassing pygofer lobe, fused almost to apex; ca. 40 microsetae uniseriate in most part; arising near middle of plate with 6 – 7 feeble macrosetae in alignment, reaching apex (Figs 11, 16, 30 – 31). Aedeagal shaft slight long, broad, ovoid, width of base nearly equal to width subapically, gonopore terminal, basoventral protrusion of aedeagus slender, sinuate, slightly shorter than shaft, in ventral aspect bifurcated at the middle, parallel extended with pointed apex, without dorsoatrium, (Figs 11, 16, 32 – 33). Connective lamellate caudally and crimped basally, caudal margin concave medially (Figs 11, 16, 34). Paramere surpassing pygofer in lateral view, almost the same length with subgenital, curved medially and bifurcated near apically, bearing a rigid tooth basad of bifurcation (Figs 11, 16, 35). Anal tube appendage developed, skeletonized, curved toward apex (Figs 11, 16, 36).	en	Wang, Ran, Zhang, Yalin, Cao, Yanghui (2025): Review of the genus Ishiharella Dworakowska (Hemiptera: Cicadellidae: Typhlocybinae), with description of two new species. Zootaxa 5604 (2): 176-184, DOI: 10.11646/zootaxa.5604.2.7, URL: https://doi.org/10.11646/zootaxa.5604.2.7
03B487EDFFE9F52F96DD8278FDB40EE2.taxon	discussion	Notes. This new species is similar to I. iochoui Dworakowska and I. falcata Yu, Yang & Dietrich in having the aedeagal process branched, but differs in the structure of the paramere and pygofer. It can be distinguished from I. iochoui by the subgenital plate fused almost to apex (Fig. 31) and anal tube appendage developed (Fig. 36). It also differs from I. falcata by the paramere branched near apically and the position, structure of the branching of aedeagal basoventral protrusion (Figs 33, 35).	en	Wang, Ran, Zhang, Yalin, Cao, Yanghui (2025): Review of the genus Ishiharella Dworakowska (Hemiptera: Cicadellidae: Typhlocybinae), with description of two new species. Zootaxa 5604 (2): 176-184, DOI: 10.11646/zootaxa.5604.2.7, URL: https://doi.org/10.11646/zootaxa.5604.2.7
03B487EDFFE9F52F96DD8278FDB40EE2.taxon	etymology	Etymology. The new species is named after its type locality: Wuxi County, Chongqing Municipality.	en	Wang, Ran, Zhang, Yalin, Cao, Yanghui (2025): Review of the genus Ishiharella Dworakowska (Hemiptera: Cicadellidae: Typhlocybinae), with description of two new species. Zootaxa 5604 (2): 176-184, DOI: 10.11646/zootaxa.5604.2.7, URL: https://doi.org/10.11646/zootaxa.5604.2.7
03B487EDFFE9F52F96DD8278FDB40EE2.taxon	distribution	Distribution. China (Chongqing).	en	Wang, Ran, Zhang, Yalin, Cao, Yanghui (2025): Review of the genus Ishiharella Dworakowska (Hemiptera: Cicadellidae: Typhlocybinae), with description of two new species. Zootaxa 5604 (2): 176-184, DOI: 10.11646/zootaxa.5604.2.7, URL: https://doi.org/10.11646/zootaxa.5604.2.7
03B487EDFFEAF52F96DD8168FDA20FC6.taxon	discussion	Notes: Sohi & Mann (1990) described this species in Daluana from India. We here transfer it to Ishiharella mainly based on the characters of the body and especially the configuration of the male genitalia.	en	Wang, Ran, Zhang, Yalin, Cao, Yanghui (2025): Review of the genus Ishiharella Dworakowska (Hemiptera: Cicadellidae: Typhlocybinae), with description of two new species. Zootaxa 5604 (2): 176-184, DOI: 10.11646/zootaxa.5604.2.7, URL: https://doi.org/10.11646/zootaxa.5604.2.7
03B487EDFFEAF52F96DD8168FDA20FC6.taxon	distribution	Distribution: India (Uttar Pradesh).	en	Wang, Ran, Zhang, Yalin, Cao, Yanghui (2025): Review of the genus Ishiharella Dworakowska (Hemiptera: Cicadellidae: Typhlocybinae), with description of two new species. Zootaxa 5604 (2): 176-184, DOI: 10.11646/zootaxa.5604.2.7, URL: https://doi.org/10.11646/zootaxa.5604.2.7
