identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03B06D5ABB3B662541554857FF28C7AD.text	03B06D5ABB3B662541554857FF28C7AD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phaseolus acutifolius A. Gray 1852	<div><p>Phaseolus acutifolius A. Gray (1852: 43) [clade B; sect. Acutifolii Freytag].</p><p>TYPE: C Wright 1311 (LECTOTYPE: GH-64068) .</p><p>In the original description (p. 44) there is no mention of a specimen. However, through the work of George Freeman (1912, p. 20 and following) and his interactions with the staff of the Gray Herbarium, it was possible to locate and identify the type. Although the contribution by Asa Gray has been accepted for publication in March 1850, it has been published in March 1852. Two wild forms are often recognized, var. acutifolius and var. tenuifolius (Delgado-Salinas 1985) (Fig. 1). There might be ecological characteristics that separate these two forms (Pratt &amp; Nabhan 1988), with var. tenuifolius being a colonizer of open sunny, slightly more xeric habitats. This variety often displays narrow leaflets (width 4–10 mm), the lateral ones often sublobate at base, with heteroblastic development being more expressed as the stem gets exposed to direct sun light. But studies using molecular markers (Muñoz et al. 2006; Blair et al. 2012) are less affirmative about the differences between the two forms, giving support to the synonymy proposed by Russ Buhrow (1983). Considered a member of the Vulgaris group (Delgado-Salinas et al. 2006), it is here maintained apart as the phylogenetic distance to common bean is not the same as for the other members of this group (Debouck 1999).</p><p>Specimens: U.S.A. Arizona: FS Coburn 1490 (ASU-292151). New Mexico: JT Columbus 1588 (NMCR). Texas: P Manning 4080 (K). MEXICO. Baja California Sur: R Moran 18725 (ARIZ-185218). Sonora: P Jenkins 88-329 (ARIZ-322133). Chihuahua: HS Gentry 2465 (ARIZ-300374). Coahuila: E Palmer 261 (PH). Sinaloa: J González-Ortega 870 (ENCB). Durango: P Tenorio-Lezama 9850 (TEX-272994). Nayarit: G Flores-Franco 1249 (MO-3851303). Jalisco: R Lépiz-Ildefonso 125 (IBUG-208586). Guanajuato: LI Aguilera G. 293 (CHAPA). Michoacán: JN Labat 1279 (P).</p></div>	https://treatment.plazi.org/id/03B06D5ABB3B662541554857FF28C7AD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Debouck, D. G.	Debouck, D. G. (2021): Phaseolus Beans (Leguminosae: Phaseoleae): A Checklist And Notes On Their Taxonomy And Ecology. Journal of the Botanical Research Institute of Texas 15 (1): 73-111, DOI: 10.17348/jbrit.v15.i1.1052, URL: https://doi.org/10.17348/jbrit.v15.i1.1052
03B06D5ABB3D6623415548CBFE06C766.text	03B06D5ABB3D6623415548CBFE06C766.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phaseolus albescens McVaugh ex R. RamIrez & A. Delgado 1999	<div><p>Phaseolus albescens McVaugh ex R. RamÍrez &amp; A. Delgado (1999: 638) [clade B; sect. Phaseoli DC. emend. Freytag].</p><p>TYPE: A Delgado-Salinas et al. 1705 (HOLOTYPE: MEXU n.v.; ISOTYPES: GH-423041, K-651820, NY-1104834).</p><p>It is a pluriannual vine in montane pine, fir, oak, ash forests on a few mountainous ridges of western Mexico (RamÍrez-Delgadillo &amp; Delgado-Salinas 1999) at 1300–2300 m. Apparently, it was Rogers McVaugh who first realized that this plant was not a P. coccineus nor a P. dumosus (McVaugh 1987, p. 654) . Primary bracts and bracteoles are smaller as compared to those of P.dumosus, corolla pale lavender to old rose contrasting with purple in P.dumosus (Fig. 2), and 4–5 seeds per pod (one less on average in wild P.dumosus). It is possibly at risk because of the logging in these old-growth forests, namely in the area of El Fresnito (county of Zapotiltic in Jalisco) as noted by the author in November 2003.</p><p>Specimens: MEXICO. Jalisco: T Kajita 95120310 (MEXU-1235511). Colima: L Rico 926 (IBUG-133696). Michoacán: C Medina G. 3601 (IEB-148134). Guerrero: E Velázquez 479 (K).</p></div>	https://treatment.plazi.org/id/03B06D5ABB3D6623415548CBFE06C766	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Debouck, D. G.	Debouck, D. G. (2021): Phaseolus Beans (Leguminosae: Phaseoleae): A Checklist And Notes On Their Taxonomy And Ecology. Journal of the Botanical Research Institute of Texas 15 (1): 73-111, DOI: 10.17348/jbrit.v15.i1.1052, URL: https://doi.org/10.17348/jbrit.v15.i1.1052
03B06D5ABB3C66224115480CFCB9C427.text	03B06D5ABB3C66224115480CFCB9C427.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phaseolus albicarminus Debouck & N. Chaves 2020	<div><p>Phaseolus albicarminus Debouck &amp; N. Chaves (2020: 3) [possibly in clade A and sect. Brevilegumeni Freytag].</p><p>TYPE: DG Debouck et al.3347 (HOLOTYPE: USJ-111997; ISOTYPES: BRIT, CR-282240, F, GH, K, MICH, MO, P, UC).</p><p>It is a recent addition to the flora of Costa Rica (Fig. 3), growing in premontane wet forest at 1600–2000 m, and known from three populations so far, from a small area south of mount Caraigres at the western end of the Talamanca mountain range (Debouck et al. 2020). The area is affected by the expansion of coffee plantations and pastures.</p><p>Specimens: COSTA RICA. San José: DG Debouck 3242 (CR-281568), DG Debouck 3342 (USJ-111444).</p></div>	https://treatment.plazi.org/id/03B06D5ABB3C66224115480CFCB9C427	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Debouck, D. G.	Debouck, D. G. (2021): Phaseolus Beans (Leguminosae: Phaseoleae): A Checklist And Notes On Their Taxonomy And Ecology. Journal of the Botanical Research Institute of Texas 15 (1): 73-111, DOI: 10.17348/jbrit.v15.i1.1052, URL: https://doi.org/10.17348/jbrit.v15.i1.1052
03B06D5ABB3C662241154906FF31C770.text	03B06D5ABB3C662241154906FF31C770.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phaseolus albiflorus Freytag & Debouck 2002	<div><p>Phaseolus albiflorus Freytag &amp; Debouck (2002: 177) [clade A; sect. Digitati Freytag].</p><p>TYPE: DG Debouck et al. 1510 (HOLOTYPE: US-3168862 and 3168864; ISOTYPES: COL-58795, G, K, M, MICH-1179903, SI, UNL n.v., UC, WIS).</p><p>It is a tall (more than 3 m) climbing plant, growing in oak and pine-oak forest at 1000–2200 m, with white flowers (fading dull ochre yellow after anthesis) (Fig. 4) and floral buds like fingertips, thus belonging to the Digitati section. The stems display heteroblastic development, with leaflets usually much lobed in the upper flowering parts. The keel is much deflected towards the left wing, the last coil being almost horizontal (Fig. 4). Recent collections have been done in northeastern Mexico (Acosta-DÍaz et al. 2019).</p><p>Specimens: MEXICO. Coahuila: JA Encina 591 (ANSM-74394). Nuevo León: DG Debouck 1507 (US-3168358). Tamaulipas: FO Zuloaga 9700 (ANSM-89228).</p></div>	https://treatment.plazi.org/id/03B06D5ABB3C662241154906FF31C770	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Debouck, D. G.	Debouck, D. G. (2021): Phaseolus Beans (Leguminosae: Phaseoleae): A Checklist And Notes On Their Taxonomy And Ecology. Journal of the Botanical Research Institute of Texas 15 (1): 73-111, DOI: 10.17348/jbrit.v15.i1.1052, URL: https://doi.org/10.17348/jbrit.v15.i1.1052
03B06D5ABB3F662141554FFBFD7AC560.text	03B06D5ABB3F662141554FFBFD7AC560.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phaseolus albinervus Freytag & Debouck 2002	<div><p>Phaseolus albinervus Freytag &amp; Debouck (2002: 127) [clade B; sect. Paniculati Freytag].</p><p>TYPE: M Lavin et al. 5426 (HOLOTYPE: TEX-371372; ISOTYPES: CAS-817320, F-2060227, MO-3412308, NY, UCR-46735).</p><p>This scandent pluriannual shrub seems to be known only from the type locality in W Chihuahua, Mexico.</p><p>This species has been validated as a member of the Paniculati (Delgado-Salinas et al. 2006); it seems to share with P. salicifolius and P. sonorensis the trait of blooming in spring.</p></div>	https://treatment.plazi.org/id/03B06D5ABB3F662141554FFBFD7AC560	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Debouck, D. G.	Debouck, D. G. (2021): Phaseolus Beans (Leguminosae: Phaseoleae): A Checklist And Notes On Their Taxonomy And Ecology. Journal of the Botanical Research Institute of Texas 15 (1): 73-111, DOI: 10.17348/jbrit.v15.i1.1052, URL: https://doi.org/10.17348/jbrit.v15.i1.1052
03B06D5ABB3F66214155484DFD07C406.text	03B06D5ABB3F66214155484DFD07C406.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phaseolus albiviolaceus Freytag & Debouck 2002	<div><p>Phaseolus albiviolaceus Freytag &amp; Debouck (2002: 179) [clade A; sect. Digitati Freytag].</p><p>TYPE: DG Debouck et al. 2063 (HOLOTYPE: US-3168863; ISOTYPES: BR, COL-235996, G, K, M, MICH-1179904, UNL n.v., WIS).</p><p>This species (Fig. 4) has been considered a synonym of P.neglectus (Delgado-Salinas et al. 2006); in the latter, on the type, the primary bracts are bigger (4–7 vs 2–2.5 mm), and the rachis of inflorescence is much shorter (2–4 vs 5–8 cm) as compared to the peduncle.Pending more material for study, this synonymy is kept on hold.</p><p>Specimen: MEXICO. Nuevo León: DG Debouck 2059 (BR, K, US-3168519).</p></div>	https://treatment.plazi.org/id/03B06D5ABB3F66214155484DFD07C406	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Debouck, D. G.	Debouck, D. G. (2021): Phaseolus Beans (Leguminosae: Phaseoleae): A Checklist And Notes On Their Taxonomy And Ecology. Journal of the Botanical Research Institute of Texas 15 (1): 73-111, DOI: 10.17348/jbrit.v15.i1.1052, URL: https://doi.org/10.17348/jbrit.v15.i1.1052
03B06D5ABB3F662141554966FC64C70D.text	03B06D5ABB3F662141554966FC64C70D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phaseolus altimontanus Freytag & Debouck 2002	<div><p>Phaseolus altimontanus Freytag &amp; Debouck (2002: 183) [clade A; sect. Digitati Freytag].</p><p>TYPE: DG Debouck et al. 1525 (HOLOTYPE: US-3168865; ISOTYPES: BR, G, K, M, SI, UNL n.v., UC-1716192).</p><p>This reclining vine with light purple standard and white wings (Fig. 4) is considered as endemic because only a few populations are known to date, from pine-oak forests of Santiago and Cola de Caballo south of Monterrey in Nuevo León and neighboring Tamaulipas, Mexico, at 1500–1900 m (also noted by Acosta-Díaz et al. 2015). The species is considered threatened (Acosta-Díaz et al. 2019).</p><p>Specimens: MEXICO. Nuevo León: S Rodríguez-Tijerina 138 (ANSM-63277). Tamaulipas: J Hinton 24657 (ANSM-71567).</p></div>	https://treatment.plazi.org/id/03B06D5ABB3F662141554966FC64C70D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Debouck, D. G.	Debouck, D. G. (2021): Phaseolus Beans (Leguminosae: Phaseoleae): A Checklist And Notes On Their Taxonomy And Ecology. Journal of the Botanical Research Institute of Texas 15 (1): 73-111, DOI: 10.17348/jbrit.v15.i1.1052, URL: https://doi.org/10.17348/jbrit.v15.i1.1052
03B06D5ABB3F662041554A7BFD8BC03D.text	03B06D5ABB3F662041554A7BFD8BC03D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phaseolus amabilis Standley 1940	<div><p>Phaseolus amabilis Standley (1940a: 28) [clade A; sect. Minkelersia (Martens &amp; Galeotti) Maréchal, Mascherpa, &amp; Stainier].</p><p>TYPE: HS Gentry 2471 (HOLOTYPE: F n.v.; ISOTYPES: DES, GH-64083, K-478471, MO-1163243, UC-581881).</p><p>It has larger flowers as compared to other species of the section ( P.pauciflorus being another case in this section but with a different floral morphology). It seems to be rare, but possibly not a hybrid as suggested elsewhere (Fishbein et al. 1998) because of the following collection found at different time and place.</p><p>Specimen: MEXICO. Chihuahua: T Lebgue 2481 (NMC-70649).</p></div>	https://treatment.plazi.org/id/03B06D5ABB3F662041554A7BFD8BC03D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Debouck, D. G.	Debouck, D. G. (2021): Phaseolus Beans (Leguminosae: Phaseoleae): A Checklist And Notes On Their Taxonomy And Ecology. Journal of the Botanical Research Institute of Texas 15 (1): 73-111, DOI: 10.17348/jbrit.v15.i1.1052, URL: https://doi.org/10.17348/jbrit.v15.i1.1052
03B06D5ABB3E662041154D2BFC76C0C0.text	03B06D5ABB3E662041154D2BFC76C0C0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phaseolus amblyosepalus (Piper) Morton 1944	<div><p>Phaseolus amblyosepalus (Piper) Morton (1944: 85) [clade A; sect. Minkelersia (Martens &amp; Galeotti) Maréchal, Mascherpa, &amp; Stainier]. TYPE: P Ibaña-García 456 (HOLOTYPE: US-1013387) .</p><p>The voucher 1013387 kept at US consists of two plants; the plant to the right is the type of P. amblyosepalus . This member of the Minkelersia can be easily identified by its large, rounded stipules. Its distribution seems to be the contact zone between Sinaloa and Durango in the Sierra Madre Occidental of Mexico, in pine forest at 2000–2700 m.</p><p>Specimens: MEXICO. Durango: Y Herrera 215 (ENCB), AC Sanders 21466 (UCR-105163), P Tenorio-Lezama 9685 (TEX-272982).</p></div>	https://treatment.plazi.org/id/03B06D5ABB3E662041154D2BFC76C0C0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Debouck, D. G.	Debouck, D. G. (2021): Phaseolus Beans (Leguminosae: Phaseoleae): A Checklist And Notes On Their Taxonomy And Ecology. Journal of the Botanical Research Institute of Texas 15 (1): 73-111, DOI: 10.17348/jbrit.v15.i1.1052, URL: https://doi.org/10.17348/jbrit.v15.i1.1052
03B06D5ABB3E662041154E2CFCF9C3E7.text	03B06D5ABB3E662041154E2CFCF9C3E7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phaseolus angucianae Debouck & Araya 2018	<div><p>Phaseolus angucianae Debouck &amp; Araya (2018: 508) [possibly in clade A and sect. Brevilegumeni Freytag].</p><p>TYPE: DG Debouck et al.3369 (HOLOTYPE: USJ-111488; ISOTYPES: BRIT-476561, CR-286418, GH, K, MICH, MO, NY, P).</p><p>This is an endemic species from SE Costa Rica (Fig. 3), the distribution of which seems to be restricted to the “Fila Costeña Sur” in lower montane rain forest at 1400–1600 m. Because of deforestation its habitat seems shrinking (Debouck et al. 2018).</p><p>Specimens: COSTA RICA. Puntarenas: N Chaves-Barrantes 6 (USJ-111737), M Grayum 10618 (MO-6103483).</p></div>	https://treatment.plazi.org/id/03B06D5ABB3E662041154E2CFCF9C3E7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Debouck, D. G.	Debouck, D. G. (2021): Phaseolus Beans (Leguminosae: Phaseoleae): A Checklist And Notes On Their Taxonomy And Ecology. Journal of the Botanical Research Institute of Texas 15 (1): 73-111, DOI: 10.17348/jbrit.v15.i1.1052, URL: https://doi.org/10.17348/jbrit.v15.i1.1052
03B06D5ABB3E662041154EC6FC45C58D.text	03B06D5ABB3E662041154EC6FC45C58D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phaseolus angustissimus , Asa Gray 1853	<div><p>Phaseolus angustissimus A. Gray (1853: 33) [clade B; sect. Rugosi Freytag].</p><p>TYPE: C Wright 68 (distributed as 951) (SYNTYPES: GH-64073; ISOSYNTYPES: BM, K-H922/76-1, GH-64074, MO-1731218).</p><p>The original material (no. 951 in Gray’s herbarium, with “n. sp.” handwritten on the label, possibly by Asa Gray himself; Padovani &amp; Cuccuini 2006 reported a series of handwritings) seems to refer to two gatherings, one made in New Mexico in July 1851 and another one made in Sonora in September 1851 (the plant to the left in GH-64073), thus the use of syntypes along Art. 9.5. of the Melbourne Code (2012). As noted by James Lackey (1983) this taxon has a tuberculate seed coat (Fig. 5), different from the raised lines radiating from the hilum which can be seen in wild P.lunatus or P. microcarpus . It is a widespread species, growing in pine, oak, ash or plane forest or chaparral at 800–2400 m, with distribution well documented in S Arizona, S New Mexico and the W end of Texas, but with few records from N Mexico (although claimed to be present: Hermann 1966; Correll &amp; Johnston 1970; Estrada &amp; MartÍnez 2000). In the original description of P. angustissimus, Asa Gray (1853, p. 33) wrote “between San Pedro and Santa Cruz, Sonora, Sept.”. Two specimens of collection 951 have been found in PH and TCD with the indication of Sonora.</p><p>Specimens: U.S.A. Arizona: D Atwood 18881 (NY). New Mexico: ND Atwood 30820 (DAV-175765). Texas: TL Steiger 927 (NY).</p></div>	https://treatment.plazi.org/id/03B06D5ABB3E662041154EC6FC45C58D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Debouck, D. G.	Debouck, D. G. (2021): Phaseolus Beans (Leguminosae: Phaseoleae): A Checklist And Notes On Their Taxonomy And Ecology. Journal of the Botanical Research Institute of Texas 15 (1): 73-111, DOI: 10.17348/jbrit.v15.i1.1052, URL: https://doi.org/10.17348/jbrit.v15.i1.1052
03B06D5ABB3E6620410A48FBFCE5C490.text	03B06D5ABB3E6620410A48FBFCE5C490.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phaseolus anisophyllus (Piper) Freytag & Debouck 2002	<div><p>Phaseolus anisophyllus (Piper) Freytag &amp; Debouck (2002: 155) [clade A; sect. Minkelersia (Martens &amp; Galeotti) Maréchal, Mascherpa, &amp; Stainier].</p><p>TYPE: EW Nelson 4911 (HOLOTYPE: US-332953).</p><p>It seems to be a too tall and vigorous plant to be considered merely as a variant of P.parvulus (see discussion by George Freytag, 2002, p. 155), and it has been validated as a separate species of this section (Delgado-Salinas et al.2006).It has been found above 2,000 m in oak woodland in the Sierra Madre Occidental from SW Chihuahua to S Durango.</p><p>Specimens: MEXICO. Chihuahua: A McDonald 2513 (ARIZ-340049). Durango: R Spellenberg 12747 (NMC-85870).</p></div>	https://treatment.plazi.org/id/03B06D5ABB3E6620410A48FBFCE5C490	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Debouck, D. G.	Debouck, D. G. (2021): Phaseolus Beans (Leguminosae: Phaseoleae): A Checklist And Notes On Their Taxonomy And Ecology. Journal of the Botanical Research Institute of Texas 15 (1): 73-111, DOI: 10.17348/jbrit.v15.i1.1052, URL: https://doi.org/10.17348/jbrit.v15.i1.1052
03B06D5ABB3E662F410A49FCFCBEC46F.text	03B06D5ABB3E662F410A49FCFCBEC46F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phaseolus augusti Harms 1921	<div><p>Phaseolus augusti Harms (1921: 503) [clade B; sect. Paniculati Freytag]. TYPE: A Weberbauer 6439 (HOLOTYPE: B n.v .; ISOTYPES: G, GH-64117, MOL, US-1496218).</p><p>The type was probably lost in Berlin during WWII; while the isotype at G consists only of fragments, the other three are complete blooming specimens. If there is any need (see Turland 2013, p. 64 and following), on the basis of the number of flowering racemes, the author wishes to indicate: “ Neotype designated here: A Weberbauer 6439 (GH-64117)”. Using this specimen and the specimen US-1058964 (= the type of Phaseolus bolivianus Piper), the author fully endorses the synonymy of the latter with P. augusti proposed by James Zarucchi and Alfonso Delgado-Salinas (1993, p. 508). Incidentally, it is quite strange that Charles Vancouver Piper (1926) did not mention P.augusti (thus creating a need for P.bolivianus) while he mentioned P.pachyrrhizoides (with one “r,” p. 700). The original population of P.bolivianus is probably gone by now due to the expanding city of Cochabamba; an alternative could be the population found NW of Quillacollo DG Debouck 3000 (DAV-167638, MO-5613159, NY) (Freyre et al. 1996). Along recommendation 60C.2 of the Code (McNeill et al. 2012, p. 134), the spelling of the current termination of the epithet can be used.</p><p>Phaseolus augusti is a species well distributed in the Andes in alder seasonal forest at 1400–3200 m, related to the Andean form of wild P. lunatus (Delgado-Salinas et al. 2006; Serrano-Serrano et al. 2010), and thus included into the Paniculati where the Lima bean is the reference species (Fig. 10).</p><p>Specimens: ECUADOR. Azuay: DG Debouck 2764 (QCA). Loja: L Ellemann 91733 (K). PERU. Piura: DG Debouck 2816 (COL-332611). Cajamarca: N Angulo 1609 (F-1746428). La Libertad: S Leiva 453 (HAO-3037). Huánuco: C Ochoa 1085 (MOL). Junín: DG Debouck 2146 (UC-1571124). Apurimac: C Vargas 8858 (CUZ-7418). Huancavelica: O Tovar 1919 (US-2490509). Cuzco: E Petersen 1535 (LIL-410643). Puno: J Soukup 726 (CUZ-7417). BOLIVIA. La Paz: SG Beck 8659 (BAB). Cochabamba: DG Debouck 2483 (UC-1716208). Norcinti: DG Debouck 2504 (LPB). Chuquisaca: DG Debouck 2490 (LPB). Tarija: JRI Wood 9539 (K). ARGENTINA. Jujuy: A Burkart 11591 (SI). Salta: DG Debouck 1774 (BAA). Tucumán: R Fortunato 3565 (BAB). Catamarca: M Cerana 318 (CORD).</p></div>	https://treatment.plazi.org/id/03B06D5ABB3E662F410A49FCFCBEC46F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Debouck, D. G.	Debouck, D. G. (2021): Phaseolus Beans (Leguminosae: Phaseoleae): A Checklist And Notes On Their Taxonomy And Ecology. Journal of the Botanical Research Institute of Texas 15 (1): 73-111, DOI: 10.17348/jbrit.v15.i1.1052, URL: https://doi.org/10.17348/jbrit.v15.i1.1052
03B06D5ABB31662E415549A3FE08C3ED.text	03B06D5ABB31662E415549A3FE08C3ED.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phaseolus pachyrrhizoides Harms 1921	<div><p>Phaseolus pachyrrhizoides Harms (1921: 504)[clade B; sect. Paniculati Freytag]. TYPE: A Weberbauer 1769 (HOLOTYPE: B n.v.) .</p><p>The type was probably lost in the fire of B in 1943; it is survived as a black-and-white photograph with number 2440 (seen by the author at G and US). It was probably a unicate because the author has not seen any other specimen with that field collection number during this study. August Weberbauer (1945) mentioned this taxon only once (and without any number) from the surroundings of Huacapistana, Junín, while Francis McBride (1943) mentioned a few collections from Junín and Huánuco. Zarucchi and co-workers (1993) reported it to be present also in Ayacucho and Huancavelica. Material that matches well the description given by Hermann Harms, namely in raceme size and size and shape of primary bracts and bracteoles, is DG Debouck 2149 (BR, COL-236000, MICH, UC-1571119, US-3168424, USM n.v.) found not far from Huacapistana in JunÍn, Peru. The following specimens give indication about the range of this taxon (note it is within the range of P. augusti reported here and by Zarucchi et al. 1993).</p><p>Specimens: PERU. Amazonas: R Ferreyra 15552 (USM-26770). Cajamarca: DG Debouck 2011 (MO-2828586). Huánuco: PG Smith s.n. (USM-26765). Junín: TB Croat 57694 (USM-56590). Apurimac: DG Debouck 2294 (UC-1716200).</p><p>While P.augusti and P. pachyrrhizoides share the same pollen type (Maréchal et al. 1978a), differences between populations of both taxa revealed by biochemical (Maquet et al. 1999) and molecular studies (Caicedo et al. 1999; Delgado-Salinas et al. 1999; Fofana et al. 1999) seem to be of the same magnitude as those between gene pools within wild Lima or common bean. Having studied in the field 36 populations of P.augusti and 24 of P. pachyrrhizoides, respectively, and in herbaria 112 and 40 specimens, respectively, the following synonymy can be proposed:</p><p>Phaseolus pachyrrhizoides Harms, based on A Weberbauer 1769 (B, as holotype,lost;photographed by Chicago Natural History Museum, formerly Field Museum of Natural History,through the Rockefeller Foundation Fund for Photographing Type Specimens, Types of the Berlin Herbarium, b/w photograph nr. 2440, in G and US),and originally published in:Notizbl. Bot. Gart. Mus. Berlin-Dahlem 7:504.1921.</p><p>= Phaseolus augusti Harms, based on A Weberbauer 6439 (GH-64117) as isotype (see above), and originally published in: Notizbl. Bot. Gart. Mus. Berlin-Dahlem 7:503.1921.</p></div>	https://treatment.plazi.org/id/03B06D5ABB31662E415549A3FE08C3ED	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Debouck, D. G.	Debouck, D. G. (2021): Phaseolus Beans (Leguminosae: Phaseoleae): A Checklist And Notes On Their Taxonomy And Ecology. Journal of the Botanical Research Institute of Texas 15 (1): 73-111, DOI: 10.17348/jbrit.v15.i1.1052, URL: https://doi.org/10.17348/jbrit.v15.i1.1052
03B06D5ABB30662E41154ED6FC36C29D.text	03B06D5ABB30662E41154ED6FC36C29D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phaseolus campanulatus Freytag & Debouck 2002	<div><p>Phaseolus campanulatus Freytag &amp; Debouck (2002: 224) [clade A; sect. Brevilegumeni Freytag].</p><p>TYPE: R McVaugh 16341 (HOLOTYPE: MICH-1210016).</p><p>This species known by only a few collections from western Mexico is close to P. oligospermus, but with differences such as leaf shape and number of seeds (five) per pod (Lépiz-Ildefonso &amp; RamÍrez-Delgadillo 2010). It grows in mixed broadleaf oak forest at 1300–1400 m.</p><p>Specimens: MEXICO. Jalisco: JA Solís-Magallanes 1961 (MEXU-330005). Nayarit: O Téllez-Valdés 9346 (IBUG-169372).</p></div>	https://treatment.plazi.org/id/03B06D5ABB30662E41154ED6FC36C29D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Debouck, D. G.	Debouck, D. G. (2021): Phaseolus Beans (Leguminosae: Phaseoleae): A Checklist And Notes On Their Taxonomy And Ecology. Journal of the Botanical Research Institute of Texas 15 (1): 73-111, DOI: 10.17348/jbrit.v15.i1.1052, URL: https://doi.org/10.17348/jbrit.v15.i1.1052
03B06D5ABB30662E41154F8BFC84C5A0.text	03B06D5ABB30662E41154F8BFC84C5A0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phaseolus carterae Freytag & Debouck 2002	<div><p>Phaseolus carterae Freytag &amp; Debouck (2002: 194) [clade B; sect. Rugosi Freytag].</p><p>TYPE: FW Gould 12181 (HOLOTYPE: MICH-1210182).</p><p>According to Recommendation 60C of the Melbourne Code (2012) and information provided by Turland (2013, p. 91–92), the termination of the epithet has been corrected. This species shows an infrequent first trifoliolate lobed leaf and rugose seed testa (Fig. 5); it needs additional collecting across the Peninsula of Baja California to confirm its status as endemic of southern Baja California Sur (Freytag &amp; Debouck 2002).</p><p>Specimens: MEXICO. Baja California Sur: TS Brandegee sn (UC-137617), JL LeÓn de la Luz 3751 (K).</p></div>	https://treatment.plazi.org/id/03B06D5ABB30662E41154F8BFC84C5A0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Debouck, D. G.	Debouck, D. G. (2021): Phaseolus Beans (Leguminosae: Phaseoleae): A Checklist And Notes On Their Taxonomy And Ecology. Journal of the Botanical Research Institute of Texas 15 (1): 73-111, DOI: 10.17348/jbrit.v15.i1.1052, URL: https://doi.org/10.17348/jbrit.v15.i1.1052
03B06D5ABB30662E410A488CFF29C48D.text	03B06D5ABB30662E410A488CFF29C48D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phaseolus chiapasanus Piper 1921	<div><p>Phaseolus chiapasanus Piper (1921: 41) [clade A; sect. Chiapasana A. Delgado].</p><p>TYPE: CA Purpus 6881 (HOLOTYPE: US-567182; ISOTYPES: F-415699, GH-64086, NY-26707, UC-172989).</p><p>This species stands alone in the genus because the large size of flowers, pods and seeds, and the black color of all plant parts upon drying. Its range at 700–1700 m goes beyond Chiapas, raising the question whether it enters into western Guatemala. Its lower montane deciduous forest habitat is shrinking because of the progress of coffee plantations (Rzedowski 1978).</p><p>Specimens: MEXICO. Chiapas: M Sousa-Sánchez 11410 (CAS-676163). Oaxaca: A Flores-Martínez 1685 (IEB-79142). Veracruz: S Avendano 554 (F-1974958).</p></div>	https://treatment.plazi.org/id/03B06D5ABB30662E410A488CFF29C48D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Debouck, D. G.	Debouck, D. G. (2021): Phaseolus Beans (Leguminosae: Phaseoleae): A Checklist And Notes On Their Taxonomy And Ecology. Journal of the Botanical Research Institute of Texas 15 (1): 73-111, DOI: 10.17348/jbrit.v15.i1.1052, URL: https://doi.org/10.17348/jbrit.v15.i1.1052
03B06D5ABB30662D410A49FBFF05C709.text	03B06D5ABB30662D410A49FBFF05C709.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phaseolus coccineus L. 1753	<div><p>Phaseolus coccineus L. (1753: 724) [clade B; sect. Phaseoli DC emend. Freytag].</p><p>TYPE (LECTOTYPE): according to Westphal (1974) and Delgado-Salinas (1985) the lectotype is the specimen 899.2 in LINN, a plant cultivated in the garden of Uppsala,behaving there as an annual and of unknown origin (McVaugh 1987) .</p><p>The wild form of the scarlet runner is rather polymorphic (Delgado-Salinas 1988), namely in pubescence (McVaugh 1987), also at molecular level (Mina-Vargas et al. 2016); not surprisingly it has received many names (Freytag &amp; Debouck 2002). Some populations are clearly distinct, but possibly because of outcrossed pollination interbreeding is frequent and many intermediate forms between taxonomic entities are found. The treatment of McVaugh (1987) for “ P. coccineus and its allies” (Piper 1926, p. 683) is followed here, excepting P. glabellus (Fig. 6). While wild P. coccineus extends from Chihuahua in Mexico down to Francisco Morazán in Honduras in very diverse but open habitats at 800–3000 m, feral forms exist in El Salvador (e.g., Chalatenango: D Rodríguez 1316 (LAGU)), Costa Rica (e.g., Alajuela: A Jiménez M. 2550 (CR-118121)), Colombia (e.g., Cundinamarca: H García-Barriga 12582 (COL-30058)) and Ecuador (e.g., Pichincha: JL Luteyn 8770 (QCA)). Phaseolus multiflorus Willdenow mentioned in the list of “Vascular Plants of the Americas” (Tropicos 2020) is a synonym. Many populations of the Central Valley of Mexico sampled by Eugène Bourgeau in 1865–1866 seem to be extinct.</p><p>Specimens: MEXICO. Chihuahua: RL Oliver 612 (DUKE-215258). Coahuila: JS Marroquín s.n. (ANSM-11433). Nuevo León: FG Meyer 2742 (BR). Tamaulipas: R Díaz 512 (RB-372771). Sinaloa: L McGill9574 (ASU-47963). Durango: S Acevedo 312 (NY). Zacatecas: JN Rose 2725 (K). San Luís Potosí: J Rzedowski 5107 (ENCB). Guanajuato: A Rubio-ChacÓn 118 (EBUM-10582). Queretaro: E Carranza 2731 (CHAPA). Hidalgo: R Hernández-Magaña 6788 (MO-3851282). Nayarit: B Bartholomew 2614 (NY). Jalisco: A Delgado-Salinas 1505 (BM). Michoacán: Ch Feddema 197 (NY). México: H Bravo H. 2006 (MA-263923). Distrito Federal: E Bourgeau 334 (G). Morelos: HS Gentry 22490 (NA). Tlaxcala: MA Martínez A. 1714 (NCU-501422). Puebla: IK Langman 2081 (PH-802749). Veracruz: E Bourgeau 3351 (P). Guerrero: DG Debouck 2359 (UC-1716201). Oaxaca: WR Anderson 13109 (DUKE-327868). Chiapas: DE Breedlove 6900 (ENCB). GUATEMALA. San Marcos: DG Debouck 1637 (M). Huehuetenango: GF Freytag 78-Guat-109 (K). Quezaltenango: DG Debouck 1675 (COL-324592). Sololá: M Nee 47366 (NY). Chimaltenango: DG Debouck 1625 (COL-324586). Sacatepéquez: WC Shannon 3683 (G). Alta Verapaz: WE Harmon 4290 (ENCB). Guatemala: A Molina R. 27546 (ENCB). El Progreso: DG Debouck 2419 (AGUAT). Santa Rosa: ET Heyde 4167 (K). Jalapa: DG Debouck 2437 (SI). HONDURAS. Ocotepeque: A Molina R. 22149 (G). Intibucá: D Dey 202 (MO-3626750). Francisco Morazán: J GÓmez 94 (BM).</p></div>	https://treatment.plazi.org/id/03B06D5ABB30662D410A49FBFF05C709	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Debouck, D. G.	Debouck, D. G. (2021): Phaseolus Beans (Leguminosae: Phaseoleae): A Checklist And Notes On Their Taxonomy And Ecology. Journal of the Botanical Research Institute of Texas 15 (1): 73-111, DOI: 10.17348/jbrit.v15.i1.1052, URL: https://doi.org/10.17348/jbrit.v15.i1.1052
03B06D5ABB33662C41554A67FE26C0A0.text	03B06D5ABB33662C41554A67FE26C0A0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phaseolus costaricensis Freytag & Debouck 1996	<div><p>Phaseolus costaricensis Freytag &amp; Debouck (1996: 157) [clade B; sect. Phaseoli DC emend. Freytag]. TYPE: DG Debouck et al.2135 (HOLOTYPE: US-3168869; ISOTYPES: BR, CR-125599, K, UC-1571112, USJ-30270).</p><p>This taxon was confused with P. coccineus in previous floras of Costa Rica and Panama (Standley 1937 and Lackey &amp; D’Arcy 1980, respectively), although several collectors noted the “bright crimson” or “brilliant fuschia” color of the corolla (Fig. 2). It thrives in alder or oak humid montane forest at 1400–2200 m. Some populations seem already extinct because of the expansion of urban areas and coffee plantations: the one at Ochomogo found in 1896 by A Tonduz 10914 (US-472031) or the one of San Isidro de Coronado found in 1937 by PH Allen 555 (F-920349). Urbanization and the raising of vegetable crops of temperate climate also affect some populations in the ChiriquÍ province of Panama (Debouck &amp; RodrÍguez-Quiel 2020).</p><p>Specimens: COSTA RICA. Alajuela: N Chaves-Barrantes 11 (CR-296552). San José: JF Morales 10156 (K). Cartago: DG Debouck 3365 (USJ-111614). PANAMA. Chiriquí: DG Debouck 3372 (UCH).</p></div>	https://treatment.plazi.org/id/03B06D5ABB33662C41554A67FE26C0A0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Debouck, D. G.	Debouck, D. G. (2021): Phaseolus Beans (Leguminosae: Phaseoleae): A Checklist And Notes On Their Taxonomy And Ecology. Journal of the Botanical Research Institute of Texas 15 (1): 73-111, DOI: 10.17348/jbrit.v15.i1.1052, URL: https://doi.org/10.17348/jbrit.v15.i1.1052
03B06D5ABB32662C41154D8CFCFDC3A7.text	03B06D5ABB32662C41154D8CFCFDC3A7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phaseolus dasycarpus Freytag & Debouck 2002	<div><p>Phaseolus dasycarpus Freytag &amp; Debouck (2002: 120) [clade A; sect. Pedicellati (Benth.) Freytag].</p><p>TYPE: R Hernández-Magaña 872 (HOLOTYPE: F-1845525; ISOTYPE: MEXU n.v.) .</p><p>This member of the Pedicellati (Mercado-Ruaro et al. 2009) seems to thrive only in the region of contact between the Sierra Madre Oriental and the Trans-Mexican Volcanic Belt in eastern central Mexico. It grows in pine-oak and montane forests with frequent mist at 1900–2200 m. The current number of populations—less than ten (Mercado-Ruaro et al. 2009; Debouck 2019b)—suggests considering this species as endemic.</p><p>Specimens: MEXICO. Hidalgo: O Alcántara-Ayala 2721 (MEXU-794839). Veracruz: M Nee 25183 (F-1916163).</p></div>	https://treatment.plazi.org/id/03B06D5ABB32662C41154D8CFCFDC3A7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Debouck, D. G.	Debouck, D. G. (2021): Phaseolus Beans (Leguminosae: Phaseoleae): A Checklist And Notes On Their Taxonomy And Ecology. Journal of the Botanical Research Institute of Texas 15 (1): 73-111, DOI: 10.17348/jbrit.v15.i1.1052, URL: https://doi.org/10.17348/jbrit.v15.i1.1052
03B06D5ABB32662C41154E86FF0FC530.text	03B06D5ABB32662C41154E86FF0FC530.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phaseolus debouckii A. Delgado 2017	<div><p>Phaseolus debouckii A. Delgado (2017: 260) [clade B; sect. Phaseoli DC emend. Freytag].</p><p>TYPE: DG Debouck et al. 2889 (HOLOTYPE: MEXU n.v.; ISOTYPE: P).</p><p>This is the interesting case of a species demonstrated to exist as such because of the progress in genomics (Rendón-Anaya et al. 2017a, b), while it was originally collected as “wild P.vulgaris ” (Fig. 7). Found by “accident” (the author was looking for the wild form of the large-seeded Lima bean), this species is distributed on the Pacific slope of the Andes in the Amotape-Huacabamba depression. When the author was consulted about this wild bean, the reply was “ancient and different”, and not less than 29 publications in the period 1992–2017 have precisely demonstrated that. The montane chaparral where it thrives at 1000–2000 m is at risk in many parts according to Michael Dillon (1994).</p><p>Specimens: ECUADOR. Chimborazo: JN Rose 23838 (US-1023151). Azuay: DG Debouck 2762 (QCA). PERU. Cajamarca: A Sagasteguí 15887 (HAO-5155).</p></div>	https://treatment.plazi.org/id/03B06D5ABB32662C41154E86FF0FC530	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Debouck, D. G.	Debouck, D. G. (2021): Phaseolus Beans (Leguminosae: Phaseoleae): A Checklist And Notes On Their Taxonomy And Ecology. Journal of the Botanical Research Institute of Texas 15 (1): 73-111, DOI: 10.17348/jbrit.v15.i1.1052, URL: https://doi.org/10.17348/jbrit.v15.i1.1052
03B06D5ABB32662C4115481CFE98C487.text	03B06D5ABB32662C4115481CFE98C487.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phaseolus dumosus Macfadyen 1837	<div><p>Phaseolus dumosus Macfadyen (1837: 279) [clade B; sect. Phaseoli DC emend. Freytag].</p><p>TYPE: not found so far (and no indication about it in the original publication), but a description without illustration. Therefore, George Freytag designated a neotype (GF Freytag 78-Guat-47: US-3168877; ISONEOTYPES: BR, MO) using a wild form that he collected with Nader Vakili in Sololá in October 1978 .</p><p>It is perhaps the only species of hybrid origin (Mina-Vargas et al. 2016). The wild form with lilac flowers (Fig. 2) has a restricted distribution mainly along the volcanic rim of Guatemala and SE Chiapas at 1400–1900 m. This is in contrast with the white-flowered feral form distributed from Veracruz (M Rosas 722 (P)) down to Cuzco (L Valenzuela 2760 (K)) (Debouck 2019c). Phaseolus harmsianus Diels (1937) from Ecuador (type: L Diels 1028: B n.v.; syntype: E Heinrichs 191: SI-2418) and P. polyanthus Greenman (1907) from Veracruz (type: CR Barnes 20: F-194750) mentioned in the list of “Vascular Plants of the Americas” (Tropicos 2020) are synonyms as indicated elsewhere (Freytag &amp; Debouck 2002, p. 45).</p><p>Specimens: MEXICO. Chiapas: E Ventura 2517 (MEXU-877627). GUATEMALA. San Marcos: JA Steyermark 37495 (F-1058746). Sololá: A Gentry 6516 (MO-2161807).</p></div>	https://treatment.plazi.org/id/03B06D5ABB32662C4115481CFE98C487	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Debouck, D. G.	Debouck, D. G. (2021): Phaseolus Beans (Leguminosae: Phaseoleae): A Checklist And Notes On Their Taxonomy And Ecology. Journal of the Botanical Research Institute of Texas 15 (1): 73-111, DOI: 10.17348/jbrit.v15.i1.1052, URL: https://doi.org/10.17348/jbrit.v15.i1.1052
03B06D5ABB32662B411549E6FE1EC2DB.text	03B06D5ABB32662B411549E6FE1EC2DB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phaseolus esperanzae Seaton 1893	<div><p>Phaseolus esperanzae Seaton (1893: 118) [clade A; sect. Pedicellati (Bentham) Freytag].</p><p>TYPE: HE Seaton 371 (HOLOTYPE: GH-64087; ISOTYPE: F-266758) .</p><p>A species of the Pedicellati distinguishable by torulose pods with thick sutures, spherical seeds and slightly coriaceous lobed leaflets. It seems to be distributed along the Mexican Trans-Volcanic Belt of central Mexico in oak or pine-oak woodland at 2000–2600 m.</p><p>Specimens: MEXICO. México: A Pineda R. 570 (ARIZ-178735). Distrito Federal: J Espinosa 1043 (ENCB). Hidalgo: A Ventura A. 318 (ITIC). Puebla: M Jiménez-Chimil 30220 (K).</p></div>	https://treatment.plazi.org/id/03B06D5ABB32662B411549E6FE1EC2DB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Debouck, D. G.	Debouck, D. G. (2021): Phaseolus Beans (Leguminosae: Phaseoleae): A Checklist And Notes On Their Taxonomy And Ecology. Journal of the Botanical Research Institute of Texas 15 (1): 73-111, DOI: 10.17348/jbrit.v15.i1.1052, URL: https://doi.org/10.17348/jbrit.v15.i1.1052
03B06D5ABB35662B41554835FEDCC585.text	03B06D5ABB35662B41554835FEDCC585.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phaseolus esquincensis Freytag 2002	<div><p>Phaseolus esquincensis Freytag (2002: 170) [clade A; sect. Xanthotricha A. Delgado].</p><p>TYPE: GF Freytag 78-Mex-75 (HOLOTYPE: US-3168870; ISOTYPES: BR, F-2107873, GH, K-262842, MICH-1210017, NA-27952) .</p><p>Apparently known only from the type material, this member of the Xanthotricha has been validated elsewhere (Delgado-Salinas et al. 2006). While the other members of this section have variously shaped tuberous taproots, this species seems to have a fibrous root; the terminal leaflets are hastately lobed, markedly towards the base (infrequent trait).</p></div>	https://treatment.plazi.org/id/03B06D5ABB35662B41554835FEDCC585	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Debouck, D. G.	Debouck, D. G. (2021): Phaseolus Beans (Leguminosae: Phaseoleae): A Checklist And Notes On Their Taxonomy And Ecology. Journal of the Botanical Research Institute of Texas 15 (1): 73-111, DOI: 10.17348/jbrit.v15.i1.1052, URL: https://doi.org/10.17348/jbrit.v15.i1.1052
03B06D5ABB35662B415548E0FE11C7BB.text	03B06D5ABB35662B415548E0FE11C7BB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phaseolus filiformis Bentham 1844	<div><p>Phaseolus filiformis Bentham (1844: 13) [clade B; sect. Rugosi Freytag].</p><p>TYPE: Hinds s.n. (LECTOTYPE: K-478487; ISOLECTOTYPE: K) .</p><p>There is no reference to a specimen in the original description.There is however in Kew Herbarium a specimen collected by Richard Brinsley Hinds in 1841, stamped “Herbarium Benthamianum 1854” and with a handwritten note “Bot.Sulph. p. 13. Bay of Magdalena ” that the author considers as the lectotype. The second collection of 1841 by Hinds kept at Kew and stamped “Herbarium Hookerianum 1867” is considered as isolectotype. It is the reference species of the section Rugosi (Fig. 5) with a huge distribution (from southern Arizona and New Mexico to trans-Pecos Texas to Baja California, including several islands of the Gulf of California, to coastal Sonora with a few spots in Chihuahua, Coahuila and Durango (Debouck 2019d). It is a component of the flora of the Sonoran Desert (Shreve &amp; Wiggins 1964) at 3–1900 m.</p><p>Specimens: U.S.A. Arizona: R Buhrow 30 (ARIZ-209019). New Mexico: EM Jercinovic 1415 (NMC-84700). Texas: WR Carr 10951 (TEX-18035). MEXICO. Chihuahua: B Barlow sn (F-409658). Coahuila: JA Villareal-Quintanilla 5326 (ANSM-53105). Durango: A García 2786 (ANSM-79586). (northern) Baja California: B Ertter 8062 (UC-1563744). Baja California Sur: P Tenorio-Lezama 12914 (MO-3599672). Sonora: GA Levin 2188 (UCR-84152).</p></div>	https://treatment.plazi.org/id/03B06D5ABB35662B415548E0FE11C7BB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Debouck, D. G.	Debouck, D. G. (2021): Phaseolus Beans (Leguminosae: Phaseoleae): A Checklist And Notes On Their Taxonomy And Ecology. Journal of the Botanical Research Institute of Texas 15 (1): 73-111, DOI: 10.17348/jbrit.v15.i1.1052, URL: https://doi.org/10.17348/jbrit.v15.i1.1052
03B06D5ABB34662A41154CBCFE24C2C0.text	03B06D5ABB34662A41154CBCFE24C2C0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phaseolus glabellus Piper 1926	<div><p>Phaseolus glabellus Piper (1926: 683) [clade A; not assigned to a sect.].</p><p>TYPE: CJW Schiede 229 (LECTOTYPE: HAL-54634; ISOLECTOTYPES: G, P-36703).</p><p>This taxon was published in 1926 as a new name within “ Phaseolus coccineus and its allies”.Charles Vancouver Piper faced the situation of two names: Phaseolus glaber Schlecht. and Phaseolus glaber Roxb., but on two different biological entities within the same genus! The former name was published in Linnaea (1838: 327) on two bean plants (n. 649 et n. 651) collected “in dumetis Jalapae et pr. S. Andres” by Christian Julius Wilhelm Schiede. The latter name was published in Flora Indica (1832: 291) on a legume from Mauritius (now Vigna glabrescens Maréchal, Mascherpa &amp; Stainier, 1978a, p. 212). The way out for Piper studying bean plants from the same region of Mexico was to rename the taxon. Because the plant is almost glabrous, but with the minute uncinate hairs namely at the nodes, a logical choice for the name was glabellus (a shorter synonym of glabriusculus, itself a diminutive of glaber or glabrous).</p><p>Piper (1926, p. 683) indicated that the type is in Paris, while Delgado-Salinas (1985, p. 207) indicated it is in Halle. When comparing the labels and handwritten notes on the herbarium sheets, it seems clear that the materials in G and P are specimens sent from elsewhere (Halle?). In HAL there are three specimens collected by C.J.W. Schiede. Bearing in mind Article 8.2. of the Code and applying some anteriority rule in the type description of P. glaber by Diederich Franz Leonhard von Schlechtendal (1838, p. 327–328), it seems that the lectotype is HAL-54634 (lectotype because there is none unequivocally indicated in the original publication; Art. 9.2. of the Code). It is a collection made in August 1828 near Jalapa (with number 229 becoming 649 in the herbarium, the latter being mentioned in the original publication), while specimen HAL-54633 is of May 1829 also “in dumetis Jalapae”. The third specimen HAL-54632 is collection 229 (too!but becoming 651 in the original publication) by C.J.W. Schiede made near (= prope) San Andres in August 1828, a syntype.</p><p>It is a taxon mimicking P. coccineus (note in Fig. 6 the hooded standard and shorter wings, brick red or dark orange rather than true scarlet) but quite isolated in the genus (Delgado-Salinas et al. 2006), with an intermediate distribution around the Gulf of Mexico in alder, oak, sweet gum montane forest at 700–1800 m (Schmit et al. 1996).</p><p>Specimens: MEXICO. Tamaulipas: S Rodríguez 83 (CICY-21373). San Luís Potosí: DG Debouck 2046 (BR). Hidalgo: L González-Quintero 1570 (ENCB). Queretaro: R Fernández 3115 (ENCB). Puebla: A Delgado-Salinas 1582 (K). Veracruz: E Bourgeau 2919 (G). Oaxaca: M Sousa-Sánchez 8886 (CR-91424). Chiapas: M Linden s.n. (P).</p></div>	https://treatment.plazi.org/id/03B06D5ABB34662A41154CBCFE24C2C0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Debouck, D. G.	Debouck, D. G. (2021): Phaseolus Beans (Leguminosae: Phaseoleae): A Checklist And Notes On Their Taxonomy And Ecology. Journal of the Botanical Research Institute of Texas 15 (1): 73-111, DOI: 10.17348/jbrit.v15.i1.1052, URL: https://doi.org/10.17348/jbrit.v15.i1.1052
03B06D5ABB34662A4115482CFC85C5D7.text	03B06D5ABB34662A4115482CFC85C5D7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phaseolus gladiolatus Freytag & Debouck 2002	<div><p>Phaseolus gladiolatus Freytag &amp; Debouck (2002: 167) [clade A; sect. Xanthotricha A. Delgado].</p><p>TYPE: DG Debouck 2057 (HOLOTYPE: US-3168871; ISOTYPE: CHAPA n.v.).</p><p>A member of the section Xanthotricha with fusiform pluriannual tuberous root, terminal leaflets like spades and lateral leaflets with a round external lobe, growing in oak-pine forests of central Mexico at 1500–1800 m, and validated by Delgado-Salinas and co-workers (2006).</p><p>Specimens: MEXICO. Jalisco: A Castro 4001 (IBUG-205436). Michoacán: VW Steinmann 4751 (NY).</p></div>	https://treatment.plazi.org/id/03B06D5ABB34662A4115482CFC85C5D7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Debouck, D. G.	Debouck, D. G. (2021): Phaseolus Beans (Leguminosae: Phaseoleae): A Checklist And Notes On Their Taxonomy And Ecology. Journal of the Botanical Research Institute of Texas 15 (1): 73-111, DOI: 10.17348/jbrit.v15.i1.1052, URL: https://doi.org/10.17348/jbrit.v15.i1.1052
03B06D5ABB34662A41154936FD23C76D.text	03B06D5ABB34662A41154936FD23C76D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phaseolus grayanus Wooton & Standley 1913	<div><p>Phaseolus grayanus Wooton &amp; Standley (1913: 139) [clade A; sect. Pedicellati (Bentham) Freytag].</p><p>TYPE: EA Mearns 2124 (HOLOTYPE: US-232982; no isotype indicated in the original publication nor seen during this study).</p><p>A widely distributed member of the Pedicellati in the oak-pine forest of the Chihuahuan Desert at 1400–2800 m. Once considered as variety of P.pedicellatus, it deserves the specific rank because of stronger trailing and then climbing stems, slightly coriaceous leaflets with squarish lobes, strong peduncles with 6–14 primary bracts. Phaseolus pyramidalis Freytag and P.teulensis Freytag mentioned in the list of “Vascular Plants of the Americas” (Tropicos 2020) seem to be synonyms of P.grayanus .</p><p>Specimens:U.S.A. Arizona: JC Blumer 1347 (MIN-141973). New Mexico: R Spellenberg 12579 (NMC-69787). Texas: AC Sanders 3152 (UCR-37304). MEXICO. Sonora: SS White 3419 (GH). Chihuahua: E Estrada-CastillÓn 2806 (BRIT). Coahuila: DH Riskind 2245 (TEX-273493). Nuevo León: JA Villareal-Quintanilla 9083 (ANSM-73611). Durango: HS Gentry 8376 (ARIZ-300362). Zacatecas: DG Debouck 331 (K). San Luís Potosí: CL Lundell 5521 (WIS). Jalisco: R Ramírez-Delgadillo 1708 (IBUG-107601).</p></div>	https://treatment.plazi.org/id/03B06D5ABB34662A41154936FD23C76D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Debouck, D. G.	Debouck, D. G. (2021): Phaseolus Beans (Leguminosae: Phaseoleae): A Checklist And Notes On Their Taxonomy And Ecology. Journal of the Botanical Research Institute of Texas 15 (1): 73-111, DOI: 10.17348/jbrit.v15.i1.1052, URL: https://doi.org/10.17348/jbrit.v15.i1.1052
03B06D5ABB37662941554CBCFD28C067.text	03B06D5ABB37662941554CBCFD28C067.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phaseolus hintonii A. Delgado 2000	<div><p>Phaseolus hintonii A. Delgado (2000: 431) [clade A; sect. Xanthotricha A. Delgado].</p><p>TYPE: GB Hinton 6728 (HOLOTYPE: K-118675; ISOTYPES: GH, POM-348563) .</p><p>A climbing vine resembling P.xanthotrichus from an almost cylindrical pluriannual tuberous root, differing in subcylindrical pods with more than ten seeds, thriving in oak woodlands of central western Mexico at 1200– 1900 m.</p><p>Specimens: MEXICO. Jalisco: EA Suárez-Muro 628 (IBUG-197086). México: A Delgado-Salinas 1567 (IBUG-198458). Michoacán: JC Soto-Núñez 4554 (IBUG-198014). Oaxaca: A Delgado-Salinas 1346 (IBUG-197386).</p></div>	https://treatment.plazi.org/id/03B06D5ABB37662941554CBCFD28C067	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Debouck, D. G.	Debouck, D. G. (2021): Phaseolus Beans (Leguminosae: Phaseoleae): A Checklist And Notes On Their Taxonomy And Ecology. Journal of the Botanical Research Institute of Texas 15 (1): 73-111, DOI: 10.17348/jbrit.v15.i1.1052, URL: https://doi.org/10.17348/jbrit.v15.i1.1052
03B06D5ABB37662941554D46FBD6C350.text	03B06D5ABB37662941554D46FBD6C350.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phaseolus hygrophilus Debouck 2011	<div><p>Phaseolus hygrophilus Debouck (2011: 54) [possibly in clade A and sect. Brevilegumeni Freytag].</p><p>TYPE: DG Debouck &amp; R Araya Villalobos 3172 (HOLOTYPE: INB-4026793; ISOTYPES: BRIT, CR, F, FI, G, GH-351974, INB, K-780075, MEXU, MICH-1229077, NY-1365122, P, UC, USJ) .</p><p>As indicated in the original publication (Salcedo-Castaño et al. 2011, p. 53), the series of herbarium specimens on which the holo- and isotypes were made was from a cutting grown in the Popayán station of CIAT in Colombia (Fig. 3). It seems to be an endemic species of the premontane rain forests of Costa Rica at 1300–1600 m (Debouck et al. 2020). It is not mentioned in “Vascular Plants of the Americas” (2020).</p><p>Specimens: COSTA RICA. San José: DG Debouck 3173 (USJ-88816), DG Debouck 3370 (USJ-111489), DG Debouck 3371 (USJ-111736).</p></div>	https://treatment.plazi.org/id/03B06D5ABB37662941554D46FBD6C350	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Debouck, D. G.	Debouck, D. G. (2021): Phaseolus Beans (Leguminosae: Phaseoleae): A Checklist And Notes On Their Taxonomy And Ecology. Journal of the Botanical Research Institute of Texas 15 (1): 73-111, DOI: 10.17348/jbrit.v15.i1.1052, URL: https://doi.org/10.17348/jbrit.v15.i1.1052
03B06D5ABB376629414A4EBCFDAFC26D.text	03B06D5ABB376629414A4EBCFDAFC26D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phaseolus jaliscanus Piper 1926	<div><p>Phaseolus jaliscanus Piper (1926: 697) [clade B; sect. Paniculati Freytag].</p><p>TYPE: EW Nelson 4030 (HOLOTYPE: US-763708; no isotypes seen nor mentioned in the original publication).</p><p>A species of the Paniculati found in pine or oak forest (with palms at lower altitude) at 500–2700 m, with a clear heteroblastic development, variegated leaflets and an internally striped banner (Fig. 8).</p><p>Specimens: MEXICO. Sinaloa: IL Wiggins 13184 (US-2365635). Nayarit: O Téllez-Valdés 9512 (IBUG-198882). Jalisco: JI Calzada 9488 (ENCB). Michoacán: A Delgado-Salinas 1137 (CAS-855044).</p></div>	https://treatment.plazi.org/id/03B06D5ABB376629414A4EBCFDAFC26D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Debouck, D. G.	Debouck, D. G. (2021): Phaseolus Beans (Leguminosae: Phaseoleae): A Checklist And Notes On Their Taxonomy And Ecology. Journal of the Botanical Research Institute of Texas 15 (1): 73-111, DOI: 10.17348/jbrit.v15.i1.1052, URL: https://doi.org/10.17348/jbrit.v15.i1.1052
03B06D5ABB37662941554F5BFD02C570.text	03B06D5ABB37662941554F5BFD02C570.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phaseolus juquilensis A. Delgado 2000	<div><p>Phaseolus juquilensis A. Delgado (2000: 421) [clade B; sect. Paniculati Freytag].</p><p>TYPE: M Sousa-Sánchez et al. 6353 (HOLOTYPE: MEXU n.v.; ISOTYPES: BM-898068 and 898069, CHAPA, ENCB).</p><p>A member of the Paniculati found in pine-oak forest with little disturbance, the distribution of which seems restricted to Oaxaca (in the Mixteca Alta and on the inner slope of Sierra de Miahuatlán), at 1800–2000 m. The few populations known to date invite to still consider it as endemic for the time being. The synonymy proposed by Delgado-Salinas and co-workers (2006) for P. acinaciformis Freytag &amp; Debouck makes sense to this author.</p><p>Specimen: MEXICO. Oaxaca: R. Hernández-Magaña 5143 (IBUG-197641).</p></div>	https://treatment.plazi.org/id/03B06D5ABB37662941554F5BFD02C570	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Debouck, D. G.	Debouck, D. G. (2021): Phaseolus Beans (Leguminosae: Phaseoleae): A Checklist And Notes On Their Taxonomy And Ecology. Journal of the Botanical Research Institute of Texas 15 (1): 73-111, DOI: 10.17348/jbrit.v15.i1.1052, URL: https://doi.org/10.17348/jbrit.v15.i1.1052
03B06D5ABB3766294155485CFDF9C437.text	03B06D5ABB3766294155485CFDF9C437.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phaseolus laxiflorus Piper 1926	<div><p>Phaseolus laxiflorus Piper (1926: 692) [clade A; sect. Pedicellati (Bentham) Freytag].</p><p>TYPE: CG Pringle 13690 (HOLOTYPE: US-462384; ISOTYPES: GH-64092, MICH-1104260) .</p><p>It is not mentioned in “Vascular Plants of the Americas” (2020); it seems, however, to be a valid species distributed in the eastern part of the Mexican Trans-Volcanic Belt at 2100 m.</p><p>Specimen: MEXICO. Veracruz: GF Freytag 78-Mex-29 (CS-77522).</p></div>	https://treatment.plazi.org/id/03B06D5ABB3766294155485CFDF9C437	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Debouck, D. G.	Debouck, D. G. (2021): Phaseolus Beans (Leguminosae: Phaseoleae): A Checklist And Notes On Their Taxonomy And Ecology. Journal of the Botanical Research Institute of Texas 15 (1): 73-111, DOI: 10.17348/jbrit.v15.i1.1052, URL: https://doi.org/10.17348/jbrit.v15.i1.1052
03B06D5ABB37662941554916FF36C721.text	03B06D5ABB37662941554916FF36C721.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phaseolus leptophyllus G. Don 1832	<div><p>Phaseolus leptophyllus G. Don (1832: 350) [clade A; sect. Revoluti Freytag].</p><p>TYPE: Sessé &amp; Mociño s.n. (LECTOTYPE: OXF; ISOLECTOTYPES: F-846796, FI-4903 (in Herbarium Webbianum no. 51912), MA-601675, OXF, P-36706).</p><p>Its unique foliar morphology suggests we consider it as in a separate section (Freytag &amp; Debouck 2002). Considering it close to P.acutifolius (Delgado-Salinas 1985) and next as a member of the Minkelersia (Delgado-Salinas et al. 2006) is a clear indication of the need for additional (and living) material. Some authors (Delgado-Salinas &amp; Gama-López 2015; Acosta-DÍaz et al. 2019) have considered it as extinct. It is known only from the type collected in Guerrero in 1789.It is not mentioned in the list of “Vascular Plants of the Americas” (Tropicos 2020).</p></div>	https://treatment.plazi.org/id/03B06D5ABB37662941554916FF36C721	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Debouck, D. G.	Debouck, D. G. (2021): Phaseolus Beans (Leguminosae: Phaseoleae): A Checklist And Notes On Their Taxonomy And Ecology. Journal of the Botanical Research Institute of Texas 15 (1): 73-111, DOI: 10.17348/jbrit.v15.i1.1052, URL: https://doi.org/10.17348/jbrit.v15.i1.1052
03B06D5ABB37662841554A0CFD0AC485.text	03B06D5ABB37662841554A0CFD0AC485.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phaseolus leptostachyus Bentham 1837	<div><p>Phaseolus leptostachyus Bentham (1837: 72) [clade B; sect. Falcati Freytag].</p><p>TYPE: WF Karwinski s.n. (LECTOTYPE: M; ISOLECTOTYPE: SI) .</p><p>The original publication about P.anisotrichos is of 1838 (von Schlechtendal 1838), therefore this name falls into synonymy. There would thus be no reason to maintain the latter as a separate species in the list of “Vascular Plants of the Americas” (Tropicos 2020). Note the specimen at the Darwinion is a small fragment of the original of M (from a loan?). It is a common species in openings of oak, pine-oak and tropical deciduous forests, from almost sea level to 2,500 m (Fig. 9). The synonymy proposed elsewhere (Delgado-Salinas 1985) for P. opacus Piper (1926) with P.leptostachyus makes sense. Five botanical varieties were recognized in this species (Freytag &amp; Debouck 2002), while in author’s view they represent the natural variation of this widespread species.</p><p>Specimens: MEXICO. Sonora: IL Wiggins 6995 (DS-264709). Tamaulipas: E Estrada-CastillÓn 2726 (CAS-794254). Chiapas: A Delgado-Salinas 823 (DS-719855). GUATEMALA. Huehuetenango: DG Debouck 3073 (AGUAT). Jalapa: PC Standley 76999 (F-1202124). HONDURAS. Francisco Morazán: LO Williams 17292 (MO-1737913). EL SALVADOR. Santa Ana: JL Linares 1070 (LAGU). La Libertad: AC del Cid 1681 (ITIC). NICARAGUA. Nueva Segovia: DG Debouck 3198 (O-V2147547). Granada: P Levy 1096 (P). COSTA RICA. Guanacaste: DG Debouck 3288 (USJ-111503). Puntarenas: F Oviedo-Brenes 2776 (USJ-111843).</p></div>	https://treatment.plazi.org/id/03B06D5ABB37662841554A0CFD0AC485	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Debouck, D. G.	Debouck, D. G. (2021): Phaseolus Beans (Leguminosae: Phaseoleae): A Checklist And Notes On Their Taxonomy And Ecology. Journal of the Botanical Research Institute of Texas 15 (1): 73-111, DOI: 10.17348/jbrit.v15.i1.1052, URL: https://doi.org/10.17348/jbrit.v15.i1.1052
03B06D5ABB366637411549E3FD7BC3C1.text	03B06D5ABB366637411549E3FD7BC3C1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phaseolus lignosus Britton 1918	<div><p>Phaseolus lignosus Britton (1918: 183) [clade B; sect. Paniculati Freytag].</p><p>TYPE (LECTOTYPE): Fig. 205 in “Flora of Bermuda”(from C Watlington 3: EPITYPE K-556376 and K-556377; ISOEPITYPES: BRIT, CAS, COL, F, FHO, FI, G, GH, MEXU, MICH, MO, NCU, NY, P; see Debouck 2015) .</p><p>Reasons were presented elsewhere (Debouck 2015) for choosing an epitype (and several duplicates from a single gathering), since the original type was an illustration. So, the “ Bermuda bean” was confused with wild P. lunatus as indicated on some labels (Debouck 2019a); the differences on living plants however are striking (Debouck 2015) (Fig. 10). It is not mentioned in the list of “Vascular Plants of the Americas” (Tropicos 2020), although in author’s view it is a valid species of the section Paniculati (also validated by Delgado-Salinas et al. 1999). This species is endemic to the Bermuda archipelago where 29 mature individuals thrive, explaining its status of critically endangered (Copeland et al. 2014).</p><p>Specimens: BERMUDA. Long Island: E Manuel 637 (A), C Watlington 3 (K-650154).</p></div>	https://treatment.plazi.org/id/03B06D5ABB366637411549E3FD7BC3C1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Debouck, D. G.	Debouck, D. G. (2021): Phaseolus Beans (Leguminosae: Phaseoleae): A Checklist And Notes On Their Taxonomy And Ecology. Journal of the Botanical Research Institute of Texas 15 (1): 73-111, DOI: 10.17348/jbrit.v15.i1.1052, URL: https://doi.org/10.17348/jbrit.v15.i1.1052
03B06D5ABB29663741554F2CFBDAC751.text	03B06D5ABB29663741554F2CFBDAC751.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phaseolus lunatus L. 1753	<div><p>Phaseolus lunatus L. (1753: 724) [clade B; sect. Paniculati Freytag].</p><p>TYPE: E Westphal 8622 (NEOTYPE, here designated: BR-18325706; isoneotypes K n.v., P n.v.).</p><p>Given the absence of a type in LINN and of indication about one in the catalogue produced by Carl August von Bergen in 1744 (p. 99, and referred to by both Carl von Linné, 1753, p. 724 and Bernard Verdcourt, 1971, p. 615), Egbert Westphal (1974, p. 146) used the cultivar “Didessa Mosaic” that he collected in Ethiopia and raised in a glasshouse in Wageningen, Holland, to select a neotype (the specimens claimed to be deposited in K and P have not been seen during this study). A variety var. silvester of possible Mexican origin (Le Marchand 2499, BR-839814) has been named (Baudet 1977).</p><p>It is perhaps the wild species of the genus with the widest range in (sub-)tropical America: from the Revillagigedo Islands off the Mexican Pacific coast, up to S Sonora and S Tamaulipas, through Central America, the Greater and Lesser Antilles down to the Chaco region of Argentina at 10–1900 m. The former distribution is that of the two lowland genepools (Motta-Aldana et al. 2010), which may have come from the northern Andes (Serrano-Serrano et al. 2010).</p><p>The Andean wild form is distinct (Serrano-Serrano et al. 2010), with a possible subcenter in the Colombian Andes (Chacón-Sánchez &amp; MartÍnez-Castillo 2017). The distribution of the Andean wild form, on the western slope of the Andes at 400–2000 m, is from SW Ecuador to NW Peru (down to the Jequetepeque Valley). This form is P.rosei Piper mentioned in the “Vascular Plants of the Americas” (Tropicos 2020) and can be considered as a synonym of P.lunatus (Freytag &amp; Debouck 2002, p. 103) . The following specimens are to indicate the extremes of ranges of the two wild forms.</p><p>Specimens: MEXICO. Revillagigedo Islands: FE Barkelew 211 (UC-137615). Sonora: JF Wiens 94-053 (ARIZ-324638). Tamaulipas: E Martínez-Ojeda 345 (ARIZ-207605). Quintana Roo: F May 2075 (CICY-54083). Chiapas: CP Cowan 5023 (CAS-834700). BELIZE. Orange Walk: C Whiteford 2675 (BM). GUATEMALA. Quezaltenango: L Rodríguez 1621 (P). HONDURAS. Francisco Morazán: JL Linares 2271 (MEXU-798503). EL SALVADOR. Ahuachapán: JM Rosales 2131 (LAGU). NICARAGUA. Estelí: DG Debouck 3209 (O-V2147454). COSTA RICA. Guanacaste: M Grayum 11508 (INB-3746751). PANAMA. Panamá: C Guerra 539 (PMA-88343). CUBA. Villa Clara: JG Jack 6725 (A). HAITI. Dept. du Nord: EC Leonard 8484 (US-1149957).TRINIDAD-AND-TOBAGO. Trinidad: RE Baker 14822 (K). COLOMBIA. Sucre: G Moreno 16 (COL-256468). VENEZUELA. Aragua: C Vogl 592 (DAV-59372). PERU. Cuzco: DG Debouck 2316 (US-3168369). BOLIVIA. El Bení: E Werdermann 2349 (MO-1000136). PARAGUAY. Alto Paraná (?): MS Bertoni 3649 (G). ARGENTINA. Chaco: AG Schulz 17027 (BAB). And for the Andean wild form: ECUADOR. Chimborazo: DG Debouck 2863 (QCA). PERU. Cajamarca: A Sagasteguí 16062 (HAO).</p></div>	https://treatment.plazi.org/id/03B06D5ABB29663741554F2CFBDAC751	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Debouck, D. G.	Debouck, D. G. (2021): Phaseolus Beans (Leguminosae: Phaseoleae): A Checklist And Notes On Their Taxonomy And Ecology. Journal of the Botanical Research Institute of Texas 15 (1): 73-111, DOI: 10.17348/jbrit.v15.i1.1052, URL: https://doi.org/10.17348/jbrit.v15.i1.1052
03B06D5ABB28663641154EF9FCA0C2B2.text	03B06D5ABB28663641154EF9FCA0C2B2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phaseolus macrolepis Piper 1926	<div><p>Phaseolus macrolepis Piper (1926: 698) [clade A; sect. Bracteati Freytag].</p><p>TYPE: O Salvin s.n. (HOLOTYPE: K-478468; no isotypes seen so far, nor mentioned in the original publication).</p><p>This species, reference of the Bracteati, grows in the now much logged old-growth pine forests, endemic of the volcanic axis of SW Guatemala at 2200–3000 m. It is not reported in the list of “Vascular Plants of the Americas” (Tropicos 2020).</p><p>Specimens: GUATEMALA. Chimaltenango: JC Montero-Castro 245 (NY). Sololá: DG Debouck 3095 (K).</p></div>	https://treatment.plazi.org/id/03B06D5ABB28663641154EF9FCA0C2B2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Debouck, D. G.	Debouck, D. G. (2021): Phaseolus Beans (Leguminosae: Phaseoleae): A Checklist And Notes On Their Taxonomy And Ecology. Journal of the Botanical Research Institute of Texas 15 (1): 73-111, DOI: 10.17348/jbrit.v15.i1.1052, URL: https://doi.org/10.17348/jbrit.v15.i1.1052
03B06D5ABB286636410A4F92FDBEC51C.text	03B06D5ABB286636410A4F92FDBEC51C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phaseolus maculatifolius Freytag & Debouck 2002	<div><p>Phaseolus maculatifolius Freytag &amp; Debouck (2002: 119) [clade B; sect. Paniculati Freytag].</p><p>TYPE: DG Debouck et al. 1509 (HOLOTYPE: US-3168873 and US-3168873A; ISOTYPES: BR, K-642355, M, UNL n.v., UC-1571140).</p><p>In addition to the type, five other populations have been reported recently, lowering the risk of extinction in the immediate future (Acosta-Díaz et al. 2015, 2019).</p></div>	https://treatment.plazi.org/id/03B06D5ABB286636410A4F92FDBEC51C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Debouck, D. G.	Debouck, D. G. (2021): Phaseolus Beans (Leguminosae: Phaseoleae): A Checklist And Notes On Their Taxonomy And Ecology. Journal of the Botanical Research Institute of Texas 15 (1): 73-111, DOI: 10.17348/jbrit.v15.i1.1052, URL: https://doi.org/10.17348/jbrit.v15.i1.1052
03B06D5ABB28663641154809FF39C722.text	03B06D5ABB28663641154809FF39C722.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phaseolus maculatus Scheele 1848	<div><p>Phaseolus maculatus Scheele (1848:465) [clade B;sect. Coriacei Freytag].</p><p>TYPE: F Lindheimer 263 (NEOTYPE: MO-1731163; ISONEOTYPES: BM, BRIT, CAN n.v., FI (in Herbarium Webbianum, no. 51931), G, GH-274635, K, MO-1731164, NA, NY, OXF, PH, TCD, UC-248307) .</p><p>In the large series of “Flora Texana exsiccata.367.Fasc.III” (Gillett &amp; Shchepanek 1977), the specimen with more information (i.e., “Cibolo, 15 miles west of New Braunfels, Texas.”) is the one out of three kept at Missouri Botanical Garden, and A. Delgado-Salinas (1985) was right in selecting it as neotype, since according to his interpretation the holotype was lost in Berlin during WWII.This sprawling legume component of the grasslands of the Chihuahuan Desert from Texas down to Puebla (Gentry 1957) at 1500–2800 m has often been confused with P.ritensis, a closely related species of the same section; apart from the seedlings (Fig. 11) the differences between them have reported elsewhere (Buhrow 1983; Delgado-Salinas 1985; Freytag &amp; Debouck 2002).</p><p>Specimens: U.S.A. Arizona: LA McGill 5683 (ASU-187971). New Mexico: JT Columbus 432 (NMCR). Texas: P Manning 4082 (K). MEXICO. Chihuahua: HS Gentry 8252 (RSA-411984). Coahuila: DH Riskind 2099 (ANSM-51545). Sonora: EA Mearns 1637 (DS-35348). Durango: S González 3195 (IEB-145721). Zacatecas: DG Debouck 346 (K-1041740). San Luís Potosí: JG Schaffner 252 (PH). Aguascalientes: JN Rose 6197 (US-396010). Guanajuato: MC Johnston 2625 B (TEX-273258). Jalisco: R Ramírez-Delgadillo 1285 (IBUG-99837). Queretaro: R Hernández-Magaña 12035 (IEB-173103). Hidalgo: JN Rose 8318 (US-451812). Tlaxcala: H Vibrans 2267 (TEX-273253). Puebla: A Delgado-Salinas 1400 (TEX-273252).</p></div>	https://treatment.plazi.org/id/03B06D5ABB28663641154809FF39C722	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Debouck, D. G.	Debouck, D. G. (2021): Phaseolus Beans (Leguminosae: Phaseoleae): A Checklist And Notes On Their Taxonomy And Ecology. Journal of the Botanical Research Institute of Texas 15 (1): 73-111, DOI: 10.17348/jbrit.v15.i1.1052, URL: https://doi.org/10.17348/jbrit.v15.i1.1052
03B06D5ABB28663541154A02FEE5C2F7.text	03B06D5ABB28663541154A02FEE5C2F7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phaseolus macvaughii A. Delgado 2000	<div><p>Phaseolus macvaughii A. Delgado (2000: 414) [clade B; sect. Falcati Freytag].</p><p>TYPE: A Delgado et al. 129 (HOLOTYPE: MEXU-200563; ISOTYPES: ANSM-8396, ARIZ-218681, CHAPA, ENCB, IBUG-52725, MICH-1180016, MO-2575900).</p><p>The “var. no. 1” of P.micranthus (McVaugh 1987, p. 660) eventually resulted in a new species of the Falcati, with chromosome number 2n = 20 (Mercado-Ruaro &amp; Delgado-Salinas 1998). This annual vine can be recognized by its fibrous narrow taproot, the unique dimorphism in pods and perhaps the smallest flower in the genus (Fig. 9). It grows in tropical dry forest and coastal dry scrub often close to the beach, from sea level up to 300 m. The author found it in 1978 in the dry bed of RÍo Salado in Colima, anticipating some salinity tolerance that was evidenced afterwards (Bayuelo-Jiménez et al. 2002).</p><p>Specimens: MEXICO. (northern) Baja California: P Tenorio-Lezama 10857 (ASU-179342). Sinaloa: RS Ferris 5199 (CAS-DS-149866). Jalisco: R Lépiz-Ildefonso 385 (IBUG-208597). Colima: DG Debouck 427 (US-3168522). Michoacán: E Martínez S. 4610 (IBUG-197388). Guerrero: J Acosta-Gallegos 211 (K).</p></div>	https://treatment.plazi.org/id/03B06D5ABB28663541154A02FEE5C2F7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Debouck, D. G.	Debouck, D. G. (2021): Phaseolus Beans (Leguminosae: Phaseoleae): A Checklist And Notes On Their Taxonomy And Ecology. Journal of the Botanical Research Institute of Texas 15 (1): 73-111, DOI: 10.17348/jbrit.v15.i1.1052, URL: https://doi.org/10.17348/jbrit.v15.i1.1052
03B06D5ABB2B663541554FD6FD2FC5DD.text	03B06D5ABB2B663541554FD6FD2FC5DD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phaseolus magnilobatus Freytag & Debouck 2002	<div><p>Phaseolus magnilobatus Freytag &amp; Debouck (2002: 167) [clade A; sect. Xanthotricha A. Delgado].</p><p>TYPE: GF Freytag 81-Mex-22 (HOLOTYPE: US-3386752) .</p><p>The type is material grown in screenhouse in Mayagüez from seed originally collected in SE Jalisco, Mexico; several isotypes not seen during this study were made from the same gathering. A couple of collections from S Durango complete the current range, thus suggest it as endemic. It has been proposed as a synonym of P.hintonii, a move that is not done here (because of the leaflet and seed traits) pending on additional collections and cross comparisons based on living materials.</p><p>Specimens: MEXICO. Durango: DG Debouck 402 (K), HS Gentry 22086 (NA).</p></div>	https://treatment.plazi.org/id/03B06D5ABB2B663541554FD6FD2FC5DD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Debouck, D. G.	Debouck, D. G. (2021): Phaseolus Beans (Leguminosae: Phaseoleae): A Checklist And Notes On Their Taxonomy And Ecology. Journal of the Botanical Research Institute of Texas 15 (1): 73-111, DOI: 10.17348/jbrit.v15.i1.1052, URL: https://doi.org/10.17348/jbrit.v15.i1.1052
03B06D5ABB2B6635414A48CBFF73C4E7.text	03B06D5ABB2B6635414A48CBFF73C4E7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phaseolus marechalii A. Delgado 2000	<div><p>Phaseolus marechalii A. Delgado (2000:427) [clade B; sect. Paniculati Freytag].</p><p>TYPE: A Delgado-Salinas 115 (HOLOTYPE: MEXU n.v.; ISOTYPES: GH-423042, K-651819).</p><p>Some differences with P. rotundatus and other related species have been presented elsewhere (Salcedo-Castaño et al. 2009) (note the striped standard in Fig. 8). Its range is mainly along the Mexican Trans-Volcanic Belt, at 2400–2500 m, in oak, pine-oak and pine forest.</p><p>Specimens: MEXICO. México: DG Debouck 2377 (MICH-1179985). Morelos: OW Norvell HM215 (UC-933928). Puebla: DG Debouck 2389 (IBUG-182551).</p></div>	https://treatment.plazi.org/id/03B06D5ABB2B6635414A48CBFF73C4E7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Debouck, D. G.	Debouck, D. G. (2021): Phaseolus Beans (Leguminosae: Phaseoleae): A Checklist And Notes On Their Taxonomy And Ecology. Journal of the Botanical Research Institute of Texas 15 (1): 73-111, DOI: 10.17348/jbrit.v15.i1.1052, URL: https://doi.org/10.17348/jbrit.v15.i1.1052
03B06D5ABB2B6634415549C6FE33C027.text	03B06D5ABB2B6634415549C6FE33C027.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phaseolus micranthus Hooker & Arnott 1841	<div><p>Phaseolus micranthus Hooker &amp; Arnott (1841: 287) [clade B; sect. Falcati Freytag].</p><p>TYPE: Lay &amp; Collie s.n. (LECTOTYPE: K-478473) .</p><p>In the book about Captain Beechey’s voyage (1841), there is no reference to a specimen. However, there is a specimen kept in the Kew Herbarium stamped “Herbarium Hookerianum 1867” and with a handwritten note directly on the sheet “ Phaseolus micranthus Hook. Mexico Beechey,” that the author considers as the nomenclatural type. It seems to be an annual plant with distribution in tropical deciduous forest and oak forest on the Pacific slopes of Sierra Madre Occidental and Sierra Madre del Sur from 20 up to 2,000 m (Fig. 9).</p><p>Specimens: MEXICO. Sonora: AC Sanders 14340 (UCR-113614). Sinaloa: JA Beltrán M. 418 (IBUG-97924). Nayarit: RS Ferris 5957 (US-1587247). Jalisco: TS Cochrane12169 (WIS). Michoacán: LM Villareal de Puga 10263 (IBUG-99743). Oaxaca: SH Salas M.4070 (IEB-161096). Chiapas: G Flores-Franco 5097 (MEXU-1232886).</p></div>	https://treatment.plazi.org/id/03B06D5ABB2B6634415549C6FE33C027	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Debouck, D. G.	Debouck, D. G. (2021): Phaseolus Beans (Leguminosae: Phaseoleae): A Checklist And Notes On Their Taxonomy And Ecology. Journal of the Botanical Research Institute of Texas 15 (1): 73-111, DOI: 10.17348/jbrit.v15.i1.1052, URL: https://doi.org/10.17348/jbrit.v15.i1.1052
03B06D5ABB2A663441154D06FC57C340.text	03B06D5ABB2A663441154D06FC57C340.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phaseolus microcarpus Martius 1829	<div><p>Phaseolus microcarpus Martius (1829: 18) [clade A; not assigned to a sect.].</p><p>TYPE (lectotype):Plate 12 (the illustration of a plant raised from seed in a greenhouse at München,Germany) .</p><p>It seems that the original seed, obtained from Mr. FW Keerl, was collected near Tlalpujahua [Michoacán], Mexico. It is an annual vine from dry (warm) (sub-) and tropical deciduous forests, with a wide distribution from Durango, Mexico down to NW Costa Rica (Debouck et al. 2019), in the range 0–2000 m. It is an isolated species in the genus, with no clear link with any current section (Delgado-Salinas et al. 1999).</p><p>Specimens: MEXICO. Durango: S González 2185 (ENCB). Guanajuato: LI Aguilera 279 (CHAPA). Nayarit: O Téllez-Valdés 9344 (RSA-482469). Jalisco: R Ramírez-Delgadillo 2270 (IBUG-109694). Morelos: G Flores-Franco 305 (BM). Puebla: F Miranda 2312 (MEXU-74655). Michoacán: JN Labat 1201 (P). Guerrero: JC Soto-Núñez 5966 (MO-3851263). Oaxaca: E Domínguez-Licona 1897 (K). Chiapas: DE Breedlove 46145 (CAS-651680). HONDURAS. Francisco Morazán: JL Linares 6995 (MEXU-1161332). EL SALVADOR. La Unión: G Cerén 972 (LAGU-6096). NICARAGUA. Carazo: A Grijalva 4131 (MO-5149166). COSTA RICA. Guanacaste: B Hammel 24516 (INB-4136116).</p></div>	https://treatment.plazi.org/id/03B06D5ABB2A663441154D06FC57C340	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Debouck, D. G.	Debouck, D. G. (2021): Phaseolus Beans (Leguminosae: Phaseoleae): A Checklist And Notes On Their Taxonomy And Ecology. Journal of the Botanical Research Institute of Texas 15 (1): 73-111, DOI: 10.17348/jbrit.v15.i1.1052, URL: https://doi.org/10.17348/jbrit.v15.i1.1052
03B06D5ABB2A6634410A4EACFD0AC287.text	03B06D5ABB2A6634410A4EACFD0AC287.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phaseolus mollis Hooker f. 1847	<div><p>Phaseolus mollis Hooker f. (1847: 228) [clade B; sect. Paniculati Freytag].</p><p>TYPE: C Darwin s.n. (LECTOTYPE: K) .</p><p>Among the plants collected by Charles Darwin during the voyage of H.M.Ship “Beagle,” Joseph Dalton Hooker described this bean plant; the specimen apparently unicate without collection number and stamped “Herbarium Hookerianum 1867” was found in Kew Herbarium. This is an endemic species of the Galapagos Islands (at 40–1200 m), possibly related to the Andean wild form of P. lunatus (Delgado-Salinas et al. 1999) . Similar relationships between biota of the Galapagos and the Pacific coast of South America have been noted for wild species of cotton and tomato (Wendel &amp; Percy 1990 and Alvarez et al. 2001, respectively).</p><p>Specimens: GALAPAGOS ISLANDS (ECUADOR). I. Isabela (= Albemarle I.): Ch Huttel 578 (QCA). I. Santa Cruz (= Indefatigable I.): A Adsersen 1599 (QCA). I. Santiago (= I. San Salvador = James I.): C Darwin s.n. (K).</p></div>	https://treatment.plazi.org/id/03B06D5ABB2A6634410A4EACFD0AC287	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Debouck, D. G.	Debouck, D. G. (2021): Phaseolus Beans (Leguminosae: Phaseoleae): A Checklist And Notes On Their Taxonomy And Ecology. Journal of the Botanical Research Institute of Texas 15 (1): 73-111, DOI: 10.17348/jbrit.v15.i1.1052, URL: https://doi.org/10.17348/jbrit.v15.i1.1052
03B06D5ABB2A6634410A4FE6FDF3C70D.text	03B06D5ABB2A6634410A4FE6FDF3C70D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phaseolus montanus Brandegee 1893	<div><p>Phaseolus montanus Brandegee (1893: 130) [possibly clade B; sect. Acutifolii Freytag].</p><p>TYPE: TS Brandegee 162 (LECTOTYPE: UC-82396) .</p><p>When Townshend Stith Brandegee reported about the flora of the Cape region of Baja California in 1893, he included his collection numbers, namely a “162. Phaseolus montanus,” but he did not make explicit that it was a new species nor that it was the type. That plant in the Brandegee Herbarium now at Berkeley, although labelled as type, was considered as synonym of P. acutifolius var. tenuifolius A. Gray (Delgado-Salinas 1985 p. 178), a treatment endorsed by George Freytag who did not see it. Comparing this plant with the holotype of P. parvifolius (US-3168875), the following synonymy can be indicated:</p><p>Phaseolus montanus Brandegee (based on TS Brandegee 162 as lectotype UC-82396 and published in: Proc. Calif. Acad. Sci. ser. 2, 3:130. 1893)</p><p>= Phaseolus parvifolius Freytag (based on GF Freytag &amp; J Sullivan 81-13 as holotype US-3168875; ISOTYPES: BR, EAP n.v., MEXU n.v., MO, and published in: Sida Bot.Misc.23:32.2002).</p><p>This taxon has consistently been evidenced as closely related to P.acutifolius (Fig. 1), but with differences that warrant the species status (Florez-Ramos et al. 2003; Muñoz et al. 2006; Gujaria-Verma et al. 2016).</p><p>Specimens: U.S.A. Arizona: RA Darrow 2261 (ARIZ-15237). MEXICO. Sonora: P Tenorio-Lezama 10254 (TEX-272992). Chihuahua: CH Muller 3443 (LL-273004). Baja California Sur: R Domínguez 313 (MEXU-431892). Sinaloa: A Delgado-Salinas 1042 (BM). Nayarit: OTéllez-Valdés 12306 (MEXU-490786). Jalisco: M Lavin 5072 (CHAPA). Colima: AC Sanders 11455 (RSA-554182). Michoacán: VW Steinmann 3860 (IEB-169341). Guerrero: DG Debouck 2354 (MICH-1180063). Oaxaca: M Sousa-Sánchez 9420 (NCU-531247). Chiapas: GF Freytag 78-Mex-77 (F-2107874). GUATEMALA. Jalapa: DG Debouck 2434 (MICH-1180058).</p></div>	https://treatment.plazi.org/id/03B06D5ABB2A6634410A4FE6FDF3C70D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Debouck, D. G.	Debouck, D. G. (2021): Phaseolus Beans (Leguminosae: Phaseoleae): A Checklist And Notes On Their Taxonomy And Ecology. Journal of the Botanical Research Institute of Texas 15 (1): 73-111, DOI: 10.17348/jbrit.v15.i1.1052, URL: https://doi.org/10.17348/jbrit.v15.i1.1052
03B06D5ABB2A663341154A7BFC23C07D.text	03B06D5ABB2A663341154A7BFC23C07D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phaseolus neglectus F. J. Hermann 1948	<div><p>Phaseolus neglectus F.J. Hermann (1948: 238) [clade A; sect. Digitati Freytag].</p><p>TYPE: CH Muller 2881 (HOLOTYPE: NA-227795, now US-2216220 since transfer in 1953) .</p><p>In the original publication Frederick Joseph Hermann noted the “corolla … pale salmon to light blue”. Alfonso Delgado-Salinas (1985, p. 198) defined two races, interestingly “not been found growing intermixed”.With the specimens available, field work and the cultivation of several materials of this section, the author now considers race A corresponding to the species described by Hermann, and race B to P. albiflorus . A couple of populations have been sampled recently (Acosta-Díaz et al. 2015).</p><p>Specimens: MEXICO. Nuevo León: R Rodríguez 377 (ANSM-5423). Tamaulipas: OL Briones s.n. (ANSM-31020).</p></div>	https://treatment.plazi.org/id/03B06D5ABB2A663341154A7BFC23C07D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Debouck, D. G.	Debouck, D. G. (2021): Phaseolus Beans (Leguminosae: Phaseoleae): A Checklist And Notes On Their Taxonomy And Ecology. Journal of the Botanical Research Institute of Texas 15 (1): 73-111, DOI: 10.17348/jbrit.v15.i1.1052, URL: https://doi.org/10.17348/jbrit.v15.i1.1052
03B06D5ABB2D6633414A4D6BFC52C0F0.text	03B06D5ABB2D6633414A4D6BFC52C0F0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phaseolus nelsonii Marechal, Mascherpa, & Stainier 1978	<div><p>Phaseolus nelsonii Maréchal, Mascherpa, &amp; Stainier (1978b: 199) [clade A; sect. Minkelersia (Martens &amp; Galeotti) Maréchal, Mascherpa, &amp; Stainier].</p><p>TYPE: EW Nelson 1362 (LECTOTYPE: US-764858,as Minkelersia pauciflora Rose (1897:142)) .</p><p>Specimens: MEXICO. Oaxaca: M Sousa-Sánchez 8540 (RSA-272960). Chiapas: DE Breedlove 46259 (CAS-651595).</p></div>	https://treatment.plazi.org/id/03B06D5ABB2D6633414A4D6BFC52C0F0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Debouck, D. G.	Debouck, D. G. (2021): Phaseolus Beans (Leguminosae: Phaseoleae): A Checklist And Notes On Their Taxonomy And Ecology. Journal of the Botanical Research Institute of Texas 15 (1): 73-111, DOI: 10.17348/jbrit.v15.i1.1052, URL: https://doi.org/10.17348/jbrit.v15.i1.1052
03B06D5ABB2D6633414A4DDCFC41C3B7.text	03B06D5ABB2D6633414A4DDCFC41C3B7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phaseolus nodosus Freytag & Debouck 2002	<div><p>Phaseolus nodosus Freytag &amp; Debouck (2002: 127) [clade B; sect. Paniculati Freytag].</p><p>TYPE: R McVaugh 13433 (HOLOTYPE: MICH-1210018) .</p><p>A member of the Paniculati with distribution in pine-oak forest of the western part of the Trans-Mexican Volcanic Belt (at 1900–2500 m), also with a striped standard (Fig. 8).</p><p>Specimens: MEXICO. Jalisco: A Delgado-Salinas 1181 (IBUG-198016). Colima: R Lépiz-Ildefonso 148 (IBUG-208602).</p></div>	https://treatment.plazi.org/id/03B06D5ABB2D6633414A4DDCFC41C3B7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Debouck, D. G.	Debouck, D. G. (2021): Phaseolus Beans (Leguminosae: Phaseoleae): A Checklist And Notes On Their Taxonomy And Ecology. Journal of the Botanical Research Institute of Texas 15 (1): 73-111, DOI: 10.17348/jbrit.v15.i1.1052, URL: https://doi.org/10.17348/jbrit.v15.i1.1052
03B06D5ABB2D6633414A4E96FD30C25D.text	03B06D5ABB2D6633414A4E96FD30C25D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phaseolus novoleonensis Debouck 2006	<div><p>Phaseolus novoleonensis Debouck (2006: 105) [clade B; sect. Coriacei Freytag].</p><p>TYPE: DG Debouck 2061 (HOLOTYPE: IBUG; ISOTYPES: ANSM, BR, CHAPA, COL, ENCB, F, G, K-478467, MEXU, MICH-1287074, MO, P, SI, UC, US).</p><p>A member of the Coriacei with rounded leaflets. One more population could have been found recently (Acosta-Díaz et al. 2015); the species is however considered as endangered because of its extremely small range (Acosta-Díaz et al. 2019).</p><p>Specimen: MEXICO. Nuevo León: AE Estrada-Castillón 633 (ANSM-33436).</p></div>	https://treatment.plazi.org/id/03B06D5ABB2D6633414A4E96FD30C25D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Debouck, D. G.	Debouck, D. G. (2021): Phaseolus Beans (Leguminosae: Phaseoleae): A Checklist And Notes On Their Taxonomy And Ecology. Journal of the Botanical Research Institute of Texas 15 (1): 73-111, DOI: 10.17348/jbrit.v15.i1.1052, URL: https://doi.org/10.17348/jbrit.v15.i1.1052
03B06D5ABB2D6633414A4F4BFBB4C540.text	03B06D5ABB2D6633414A4F4BFBB4C540.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phaseolus oaxacanus Rose 1903	<div><p>Phaseolus oaxacanus Rose (1903: 48) [tentatively in clade A; sect. Pedicellati (Benth.) Freytag].</p><p>TYPE: CG Pringle 5814a (LECTOTYPE: US-372011; ISOLECTOTYPES: ENCB, MEXU, MO-1731120).</p><p>This endemic species distributed in pine forests in the mountains surrounding the valley of Cd. Oaxaca at 2200–2700 m has been recently questioned as member of the section Pedicellati (Delgado-Salinas et al. 2006) . In a previous study (Delgado-Salinas et al. 1999) it was considered as related to this section, and also distinct enough to be seen as a valid species as compared to P.pedicellatus (a move from his varietal treatment of 1985). New data are needed to reassess the position of this taxon in the genus.</p><p>Specimens: MEXICO. Oaxaca: R Acevedo-Rodríguez 130 (ENCB), CG Pringle 5661 (GH), M Sousa-Sánchez 7816 (MEXU-209055).</p></div>	https://treatment.plazi.org/id/03B06D5ABB2D6633414A4F4BFBB4C540	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Debouck, D. G.	Debouck, D. G. (2021): Phaseolus Beans (Leguminosae: Phaseoleae): A Checklist And Notes On Their Taxonomy And Ecology. Journal of the Botanical Research Institute of Texas 15 (1): 73-111, DOI: 10.17348/jbrit.v15.i1.1052, URL: https://doi.org/10.17348/jbrit.v15.i1.1052
03B06D5ABB2D6633415548ACFC71C4B0.text	03B06D5ABB2D6633415548ACFC71C4B0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phaseolus oligospermus Piper 1926	<div><p>Phaseolus oligospermus Piper (1926: 698) [clade A; sect. Brevilegumeni Freytag].</p><p>TYPE: AS Oersted 30 (HOLOTYPE: K-478466; no isotypes seen so far nor mentioned in the original publication).</p><p>It is the reference species for the section. Sometimes confused with P.tuerckheimii (Fig. 3), it is known by scattered populations from Mexico east of the Isthmus of Tehuantepec down to central Costa Rica in premontane moist mixed (oak, pine, alder, sweet gum) forest at 450–2100 m.</p><p>Specimens: MEXICO. Oaxaca: R Torres-Colin 10589 (IBUG-169373). Chiapas: J Acosta-Gallegos 194 (K-1041773). HONDURAS. Francisco Morazán: JL Linares 6520 (MO-6132749). EL SALVADOR. Ahuachapán: JL Linares 3303 (LAGU). NICARAGUA. Estelí: CA Padilla-Gómez 40 (MO-6244283). Madríz: WD Stevens 16119 (MO-5149147). COSTA RICA. San José: DG Debouck 3333 (USJ-111445).</p></div>	https://treatment.plazi.org/id/03B06D5ABB2D6633415548ACFC71C4B0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Debouck, D. G.	Debouck, D. G. (2021): Phaseolus Beans (Leguminosae: Phaseoleae): A Checklist And Notes On Their Taxonomy And Ecology. Journal of the Botanical Research Institute of Texas 15 (1): 73-111, DOI: 10.17348/jbrit.v15.i1.1052, URL: https://doi.org/10.17348/jbrit.v15.i1.1052
03B06D5ABB2D66324155499CFBBEC047.text	03B06D5ABB2D66324155499CFBBEC047.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phaseolus parvulus Greene 1881	<div><p>Phaseolus parvulus Greene (1881:217) [clade A; sect. Minkelersia (Martens &amp; Galeotti) Maréchal, Mascherpa, &amp; Stainier].</p><p>TYPE: EL Greene s.n. (LECTOTYPE: NDG-27852; ISOLECTOTYPES: F-133379, G, K, MO-1731190, NA-27001, NY).</p><p>Edward Lee Greene presented the descriptions of “New plants of New Mexico and Arizona” based on specimens kept in his herbarium. For Phaseolus parvulus no specimen is mentioned, and there is no illustration in the paper. However, in the Greene Herbariun currently kept at the university of Notre Dame in Indiana, there is a specimen with three small plants stamped “Herbarium Greeneanum 32412” and labelled Phaseolus parvulus Greene n.sp. (handwritten by EL Greene?), that the author considers as the nomenclatural type. Because the original population had many plants (not uncommon in rainy years, from author’s field experience in 1978) from the same gathering Greene was able to distribute duplicates to several Herbaria. A small plant (up to 30–40 cm) from an almost spherical tuberous root (diam. 10–20 mm) of the section Minkelersia growing in altitude (1900–2900 m) pine-oak forests from New Mexico down the Sierra Madre Occidental as far south as the states of Jalisco and México!</p><p>Specimens: U.S.A. Arizona: D Atwood 17171 (NY). New Mexico: EH Roalson 1111 (NMCR). MEXICO. Sonora: AL Reina G. 99-546 (NMC-71768). Chihuahua: A Bénitez 2066 (ANSM-59059). Durango: S Acevedo 296 (IEB-79531). Zacatecas: JN Rose 2723 (US-301650). Sinaloa: AC Sanders 21202 (UCR-168974). Nayarit: JN Rose 3425 (US-302402). Jalisco: Luquín 160 (IBUG-32344). México: GB Hinton 4708 (GH).</p></div>	https://treatment.plazi.org/id/03B06D5ABB2D66324155499CFBBEC047	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Debouck, D. G.	Debouck, D. G. (2021): Phaseolus Beans (Leguminosae: Phaseoleae): A Checklist And Notes On Their Taxonomy And Ecology. Journal of the Botanical Research Institute of Texas 15 (1): 73-111, DOI: 10.17348/jbrit.v15.i1.1052, URL: https://doi.org/10.17348/jbrit.v15.i1.1052
03B06D5ABB2C6632410A4DA6FE64C240.text	03B06D5ABB2C6632410A4DA6FE64C240.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phaseolus pauciflorus G. Don 1832	<div><p>Phaseolus pauciflorus Sessé &amp; Mociño ex G. Don (1832:356) [clade A; sect. Minkelersia (Martens &amp; Galeotti) Maréchal, Mascherpa, &amp; Stainier].</p><p>TYPE: Sessé &amp; Mociño s.n. (LECTOTYPE: OXF) .</p><p>This taxon was briefly described by Martín de Sessé y Lacasta and José Mariano Mociño in 1791 (date of the unpublished manuscript, according to McVaugh (1977), but eventually published in 1887) on a specimen that ended up in the Lambert Herbarium (now in Oxford, England). As explained by Sousa-Sánchez and Delgado-Salinas (1981), George Don published the description in 1832, while he wanted to give credit to these botanists. On page 117 of “Plantas de Nueva España” by Sessé and Mociño (1887) and on page 356 of “A general system of gardening and botany” by Don (1832), no specimen is mentioned, but as aptly explained by Drs. Sousa and Delgado (1981) the type is the one-raceme specimen with the handwritten note “ Phaseolus linearis ” and stamped “Fielding Oxford Herbarium.” A sprawling vine usually less than 1 m long in pine or oak grassland (at 1300–2700 m), with clover-like leaves and erect often twin pods, from a pluriannual tuberous root.</p><p>Specimens: MEXICO. Sonora: M Fishbein 2471 (NY). Chihuahua: R Corral-Díaz 3964 (NMC-65116). Sinaloa: FW Pennell 20114 (PH-773300). Durango: S Acevedo 306 (ANSM-57382). Zacatecas: J Rzedowski 1022 (MO-2209626). Nayarit: G Flores-Franco 1169 (ENCB). Jalisco: P Carrillo-Reyes4324 (IBUG-169374). México: A Delgado-Salinas 1004 (CHAPA). Distrito Federal: E Matuda 21400 (K). Morelos: LH MacDaniels 269 (F-838137). Michoacán: CP Cowan 5668 (UCR-63757). Guerrero: I Wagenbreth 282 (MO-5043672). Oaxaca: H Galeotti 3175 (G).GUATEMALA. Quiché: JI Aguilar 1472 (F-1121845).</p></div>	https://treatment.plazi.org/id/03B06D5ABB2C6632410A4DA6FE64C240	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Debouck, D. G.	Debouck, D. G. (2021): Phaseolus Beans (Leguminosae: Phaseoleae): A Checklist And Notes On Their Taxonomy And Ecology. Journal of the Botanical Research Institute of Texas 15 (1): 73-111, DOI: 10.17348/jbrit.v15.i1.1052, URL: https://doi.org/10.17348/jbrit.v15.i1.1052
03B06D5ABB2C6632410A4FACFCF2C4BD.text	03B06D5ABB2C6632410A4FACFCF2C4BD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phaseolus pedicellatus Bentham 1837	<div><p>Phaseolus pedicellatus Bentham (1837: 73) [clade A; sect. Pedicellati (Benth.) Freytag].</p><p>TYPE: WF von Karwinski s.n. (LECTOTYPE: M; ISOLECTOTYPE: US-1326445) .</p><p>In the original description, George Bentham did not indicate any specimen. There is however a specimen kept in Munich that along the written indications on the sheet has been object of correspondence between Charles Vancouver Piper (at US) and Herman Ross (at M) in November 1924; it can be considered as the nomenclatural type, the specimen at US being a fragment obtained on loan. In spite of lack of data, it might be that the type material (a unicate) was collected in September 1827 near Mexico City during the first trip of Wilhelm Friedrich von Karwinski (McVaugh 1980). While it is abundant in Sierra Madre Oriental from Pico de Orizaba northwards and the Mexican Trans-Volcanic Belt, it is also known by a couple of collections from Sierra Madre Occidental (Nabhan 1990; Lépiz-Ildefonso &amp; Ramírez-Delgadillo 2010). It thrives in pine-oak, oak, mixed (oak, laurel, alder), cedar and fir forest at 1500–2800 m.</p><p>Specimens: MEXICO. Coahuila: E Palmer 2022 (K). Tamaulipas: OL Briones V s.n. (ENCB). Nuevo León: J Hinton 23424 (ANSM-61416). Guanajuato: R Santillán I 496 (ENCB). Queretaro: E Carranza 2729 (IEB-79444). Hidalgo: O Alcantará-Ayala 2354 (BM). Veracruz: F Ventura A 14780 (ENCB). Puebla: P Tenorio-Lezama 17126 (MEXU-522602). Distrito Federal: A Ventura A. 3566 (IEB-79463). Morelos: J García C. 180 (ENCB). México: H Bravo H. 2375 (MA-263930). Jalisco: JA Machuca-Núñez 6392 (IBUG-109697). Colima: R Lépiz-Ildefonso 147 (IBUG-208605). Michoacán: H Díaz-Barriga 1693 (IEB-79446). Guerrero: GB Hinton 11142 (RSA-411988).</p></div>	https://treatment.plazi.org/id/03B06D5ABB2C6632410A4FACFCF2C4BD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Debouck, D. G.	Debouck, D. G. (2021): Phaseolus Beans (Leguminosae: Phaseoleae): A Checklist And Notes On Their Taxonomy And Ecology. Journal of the Botanical Research Institute of Texas 15 (1): 73-111, DOI: 10.17348/jbrit.v15.i1.1052, URL: https://doi.org/10.17348/jbrit.v15.i1.1052
03B06D5ABB2C6632410A49ABFE32C730.text	03B06D5ABB2C6632410A49ABFE32C730.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phaseolus perplexus A. Delgado 2000	<div><p>Phaseolus perplexus A. Delgado (2000: 418) [clade A; sect. Minkelersia (Martens &amp; Galeotti) Maréchal, Mascherpa, &amp; Stainier].</p><p>TYPE: JA Solís-Magallanes 1965 (HOLOTYPE: MEXU n.v.; ISOTYPE: WIS-255906).</p><p>Specimens: MEXICO. Sinaloa: AC Sanders 21145 (UCR-105133). Jalisco: R Ramírez-Delgadillo 4571 (IBUG-198783). México: B Bartholomew 2951 (TEX-273390). Michoacán: JN Labat 406 (P).</p></div>	https://treatment.plazi.org/id/03B06D5ABB2C6632410A49ABFE32C730	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Debouck, D. G.	Debouck, D. G. (2021): Phaseolus Beans (Leguminosae: Phaseoleae): A Checklist And Notes On Their Taxonomy And Ecology. Journal of the Botanical Research Institute of Texas 15 (1): 73-111, DOI: 10.17348/jbrit.v15.i1.1052, URL: https://doi.org/10.17348/jbrit.v15.i1.1052
03B06D5ABB2C663141154A1CFC85C1E1.text	03B06D5ABB2C663141154A1CFC85C1E1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phaseolus persistentus Freytag & Debouck 2002	<div><p>Phaseolus persistentus Freytag &amp; Debouck (2002: 217) [clade B; sect. Phaseoli DC emend. Freytag].</p><p>TYPE: DG Debouck 1621 (HOLOTYPE: US-3168876; ISOTYPES: BR, K-478465, USCG n.v.).</p><p>It is known only from the type locality in the area of the volcanoes W of Guatemala City. This taxon has been moved from the section Falcati into the section Phaseoli by Delgado-Salinas and co-workers (2006). It is not reported in the list of “Vascular Plants of the Americas” (Tropicos 2020).</p></div>	https://treatment.plazi.org/id/03B06D5ABB2C663141154A1CFC85C1E1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Debouck, D. G.	Debouck, D. G. (2021): Phaseolus Beans (Leguminosae: Phaseoleae): A Checklist And Notes On Their Taxonomy And Ecology. Journal of the Botanical Research Institute of Texas 15 (1): 73-111, DOI: 10.17348/jbrit.v15.i1.1052, URL: https://doi.org/10.17348/jbrit.v15.i1.1052
03B06D5ABB2F663141554CCCFBE3C0CA.text	03B06D5ABB2F663141554CCCFBE3C0CA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phaseolus plagiocylix Harms 1921	<div><p>Phaseolus plagiocylix Harms (1921: 508) [clade A; sect. Minkelersia (Martens &amp; Galeotti) Maréchal, Mascherpa, &amp; Stainier].</p><p>TYPE: E Seler 1042 (HOLOTYPE: B n.v.; ISOTYPE: GH-64098).</p><p>The type was probably lost at Berlin during WWII, and in the original publication (p. 508) there is no mention about isotypes. There is however a specimen labelled “Ex Museo botanico Berolinensi” kept in the Gray Herbarium (GH-64098) and annotated by Hermann Harms “1042 Phaseolus plagiocylix Harms n.sp. ” that can be considered as the nomenclatural type. Known by few collections from Nuevo León, it has been reported from southern Coahuila (Carranza-P. &amp; Villareal-Quintanilla 1997), but no specimens are indicated (nor seen by the author during his visit at ANSM in 2011). Some authors (Acosta-DÍaz et al. 2019) consider it as extinct.</p></div>	https://treatment.plazi.org/id/03B06D5ABB2F663141554CCCFBE3C0CA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Debouck, D. G.	Debouck, D. G. (2021): Phaseolus Beans (Leguminosae: Phaseoleae): A Checklist And Notes On Their Taxonomy And Ecology. Journal of the Botanical Research Institute of Texas 15 (1): 73-111, DOI: 10.17348/jbrit.v15.i1.1052, URL: https://doi.org/10.17348/jbrit.v15.i1.1052
03B06D5ABB2F663141554E3BFE6FC227.text	03B06D5ABB2F663141554E3BFE6FC227.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phaseolus pluriflorus Marechal, Mascherpa, & Stainier 1978	<div><p>Phaseolus pluriflorus Maréchal, Mascherpa, &amp; Stainier (1978b: 199) [clade A; sect. Minkelersia (Martens &amp; Galeotti) Maréchal, Mascherpa, &amp; Stainier].</p><p>TYPE (as Minkelersia multiflora Rose (1897:142): CG Pringle 6471 (LECTOTYPE: US-304095; ISOLECTOTYPES: BM, G, ISC-41284, K, MO-3379357, UC-83969).</p><p>A 1–2 m climbing plant from a pluriannual tuberous root, often found among grasses and thickets on lava beds at 1400–2900 m in variously mixed pine-oak woodland.</p><p>Specimens: MEXICO. Sinaloa: HS Gentry 6307 (ARIZ-115055). Durango: S González 1803 (ANSM-59324). Nayarit: R Ramírez 944 (MEXU-541169). Jalisco: A Rodríguez-Contreras 28 (IBUG-42761). Guanajuato: A Rivas-Hernández 245 (EBUM-10619). Michoacán: J Espinosa-Garduño 1831 (ENCB). México: A Ventura A. 3289 (MO-3593725). Distrito Federal: E Bourgeau 576 (G). Tlaxcala: M Provance 2947 (UCR-151311). Morelos: JN Rose 7228 (US-450811).</p></div>	https://treatment.plazi.org/id/03B06D5ABB2F663141554E3BFE6FC227	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Debouck, D. G.	Debouck, D. G. (2021): Phaseolus Beans (Leguminosae: Phaseoleae): A Checklist And Notes On Their Taxonomy And Ecology. Journal of the Botanical Research Institute of Texas 15 (1): 73-111, DOI: 10.17348/jbrit.v15.i1.1052, URL: https://doi.org/10.17348/jbrit.v15.i1.1052
03B06D5ABB2F663141554F06FD91C4D1.text	03B06D5ABB2F663141554F06FD91C4D1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phaseolus polymorphus Sereno Watson 1882	<div><p>Phaseolus polymorphus Sereno Watson (1882: 346) [clade A; sect. Pedicellati (Benth.) Freytag].</p><p>TYPE: JG Schaffner 811 (SYNTYPES: GH-64100, CC Parry &amp; E Palmer 188 (MO-1731110)) .</p><p>Just above the original publication of this species, the specimen JG Schaffner 811 is also considered as Phaseolus heterophyllus Willd., that is, Macroptilium gibbosifolium (Ortega) A. Delgado ! As aptly noted by Alfonso Delgado-Salinas (1985, p. 281), the specimen GH-64100 is a mix including P. grayanus . The specimen JG Schaffner 811 kept in Kew (K-478482) consists of (mainly) P.grayanus, possibly a small stem of P. polymorphus and a small blooming stem of P. purpusii to the lower right. The specimen CC Parry &amp; E Palmer 186 kept in Harvard (GH-64101) and in Kew, and also cited as syntype, is P.grayanus, and thus not considered to represent this species. The specimen CC Parry &amp; E Palmer 188 kept in Saint-Louis (MO-1731110) is not a mixture and makes sense for P.polymorphus (different from P. filiformis Benth. and P.wrightii A. Gray sensu 1853) along the brief description provided by Watson. It is a climbing vine displaying heteroblastic development growing in pine-oak mixed forest and oak grassland at 900–2500 m.</p><p>Specimens: MEXICO. Chihuahua: J Passini 6654 (ENCB). Coahuila: Henrickson 22501 (ANSM-73568). Nuevo León: J Hinton 23421 (TEX-273397). Tamaulipas: JA Encina 1743 (ANSM-85810). Zacatecas: LA García-R. 758 (IBUG-167893). San Luís Potosí: DG Debouck 2056 (UC-1716191). Guanajuato: DG Debouck 2398 (US-3168517). Queretaro: L Paray 2480 (ENCB). Hidalgo: CA Purpus 1365 (POM-17092). Jalisco: LA García-R.650 (IBUG-163783).</p><p>It might be appropriate to finish here with a note about P. scabrellus Bentham ex S. Watson that is described in the same publication (1882: 346–347) just after P. polymorphus (and mentioned in the list of “Vascular Plants of the Americas” (Tropicos 2020)). Sereno Watson referred to two specimens (thus syntypes): Ed Palmer 2122 (GH-64106) and Th Coulter s.n. (GH-64105, K-478484). The author concurs with A. Delgado-Salinas (1985, p. 277) in considering this taxon as synonym of P.pedicellatus . Interestingly, Piper (1926, p. 691) wrote “ Palmer 2122, which really is P.pedicellatus Benth. ”.</p></div>	https://treatment.plazi.org/id/03B06D5ABB2F663141554F06FD91C4D1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Debouck, D. G.	Debouck, D. G. (2021): Phaseolus Beans (Leguminosae: Phaseoleae): A Checklist And Notes On Their Taxonomy And Ecology. Journal of the Botanical Research Institute of Texas 15 (1): 73-111, DOI: 10.17348/jbrit.v15.i1.1052, URL: https://doi.org/10.17348/jbrit.v15.i1.1052
03B06D5ABB2F663041554A3CFE13C0A0.text	03B06D5ABB2F663041554A3CFE13C0A0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phaseolus polystachyus (L.) Britton, Sterns, & Poggenburg 1888	<div><p>Phaseolus polystachyus (L.) Britton, Sterns, &amp; Poggenburg (1888: 15) [clade B; sect. Paniculati Freytag].</p><p>TYPE: J Clayton 568 (LECTOTYPE: BM-51643) .</p><p>In the “Preliminary Catalogue of Anthophyta … New York City” (1888, p. 15), Justus Ferdinand Poggenburg and his co-authors spelled the species as Phaseolus polystachyus; in line with the “golden rule” presented by Nicholas Turland (2013, p. 88) it is thus unclear to this author why this taxon is often changed into “ Phaseolus polystachios .” The “Flora Virginica” by Jan Fredrik Gronovius (1762, p. 106) was key to consider the collection 568 by John Clayton as the lectotype.</p><p>The “thicket bean” is a species with a huge distribution at 30–700 m in oak grassland and mixed deciduous woods of the eastern USA where its habitat is shrinking (Dohle et al. 2019). It has been reported from E Texas (Turner et al. 2003), goes as far north as New York and Connecticut (Hollick &amp; Britton 1895), and down to southern Florida (Small 1933). The following specimens just indicate the approximate extremes of the range.</p><p>Specimens: U.S.A. Connecticut: EB Harger 4148 (NEBC-643445). Florida: RP Wunderlin 5667 (POM-348542). Oklahoma: J Taylor 26884 (BRIT). Texas: WR Carr 18597 (TEX-201584).</p></div>	https://treatment.plazi.org/id/03B06D5ABB2F663041554A3CFE13C0A0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Debouck, D. G.	Debouck, D. G. (2021): Phaseolus Beans (Leguminosae: Phaseoleae): A Checklist And Notes On Their Taxonomy And Ecology. Journal of the Botanical Research Institute of Texas 15 (1): 73-111, DOI: 10.17348/jbrit.v15.i1.1052, URL: https://doi.org/10.17348/jbrit.v15.i1.1052
03B06D5ABB2E663041154D8CFDA4C387.text	03B06D5ABB2E663041154D8CFDA4C387.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phaseolus purpusii Brandegee 1912	<div><p>Phaseolus purpusii Brandegee (1912: 271) [clade A; sect. Pedicellati (Benth.) Freytag].</p><p>TYPE: CA Purpus 5196 (HOLOTYPE: UC-149841; ISOTYPES: BM, G, GH-64102, L-501280, MO-1731111, P).</p><p>Its deeply lobed narrow leaflets (with main nerves almost perpendicular) incite to maintain it separate from P. pedicellatus, while keeping it in the section Pedicellati. Few collections are known from W San LuÍs PotosÍ, Mexico. It is not mentioned in the “Vascular Plants of the Americas,” while P. pyramidalis Freytag belonging to the same section is cited. The author would agree on considering the latter as a synonym of P. grayanus (Delgado-Salinas et al. 2006) .</p><p>Specimen: MEXICO. San Luís Potosí: R Grether 782 (MEXU-274666).</p></div>	https://treatment.plazi.org/id/03B06D5ABB2E663041154D8CFDA4C387	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Debouck, D. G.	Debouck, D. G. (2021): Phaseolus Beans (Leguminosae: Phaseoleae): A Checklist And Notes On Their Taxonomy And Ecology. Journal of the Botanical Research Institute of Texas 15 (1): 73-111, DOI: 10.17348/jbrit.v15.i1.1052, URL: https://doi.org/10.17348/jbrit.v15.i1.1052
03B06D5ABB2E663041154EE6FC74C2AD.text	03B06D5ABB2E663041154EE6FC74C2AD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phaseolus reticulatus Freytag & Debouck 2002	<div><p>Phaseolus reticulatus Freytag &amp; Debouck (2002: 262) [clade B; sect. Coriacei Freytag].</p><p>TYPE: DG Debouck 409 (HOLOTYPE: US-3168878; ISOTYPES: CHAPA n.v., K-642353 and K-642354, M, MICH-1180182 ).</p><p>It is a member of the Coriacei with heteroblastic development, lobed leaflets with recurved nervation and variegation, that seems to be endemic of the SW Durango-Sinaloa border. Found in pine-oak grassland at 2000– 2500 m.</p><p>Specimens: MEXICO. Durango: M Lavin 5022 (GH), P Tenorio-Lezama 4282 (IEB-79482), P.Tenorio-Lezama 4310 (RSA-506258).</p></div>	https://treatment.plazi.org/id/03B06D5ABB2E663041154EE6FC74C2AD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Debouck, D. G.	Debouck, D. G. (2021): Phaseolus Beans (Leguminosae: Phaseoleae): A Checklist And Notes On Their Taxonomy And Ecology. Journal of the Botanical Research Institute of Texas 15 (1): 73-111, DOI: 10.17348/jbrit.v15.i1.1052, URL: https://doi.org/10.17348/jbrit.v15.i1.1052
03B06D5ABB2E6630410A4F9BFDAFC4B7.text	03B06D5ABB2E6630410A4F9BFDAFC4B7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phaseolus ritensis M. E. Jones 1908	<div><p>Phaseolus ritensis M.E. Jones (1908: 14) [clade B; sect. Coriacei Freytag].</p><p>TYPE: ME Jones s.n. (LECTOTYPE: POM-28786; ISOLECTOTYPES: DS-145645, GH-64079, NY-312364, POM-28788).</p><p>In 1908 Marcus Eugene Jones made the description of a bean plant from the Santa Rita mountains of Arizona; although no specimen is indicated, the botanist wrote (p. 12) that “the types of all species described are in my herbarium”. The ME Jones Herbarium was purchased in 1923 by the Pomona College, and now kept at the Rancho Santa Ana Botanic Garden, where the author saw the specimen (with barcode RSA 3602) in November 2010 (here considered as lectotype). P. ovatifolius reported in the list of “Vascular Plants of the Americas” (Tropicos 2020) is a synonym of this species (as also reported by Delgado-Salinas 1985). Phaseolus ritensis is a tall plant of the Coriacei with upper stems twining, often confused with P. maculatus (Fig. 11). It thrives on slopes in pine-oak forests from S Arizona down to Jalisco, mostly along the eastern slope of Sierra Madre Occidental, at 1200–2600 m. The presence in New Mexico and Texas is still to be documented, as the author has not seen specimens from these two states (neither reported by Wooton &amp; Standley 1915, nor by Turner et al. 2003, respectively), in contrast with P.maculatus .</p><p>Specimens: U.S.A. Arizona: JC Blumer 3360 (UC-153734). MEXICO. Chihuahua: R Corral-Díaz 2190 (NMC-2904). Durango: ME Maury 138 (IEB-79536). Jalisco: A Rodríguez-Contreras 1454 (IBUG-94444).</p></div>	https://treatment.plazi.org/id/03B06D5ABB2E6630410A4F9BFDAFC4B7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Debouck, D. G.	Debouck, D. G. (2021): Phaseolus Beans (Leguminosae: Phaseoleae): A Checklist And Notes On Their Taxonomy And Ecology. Journal of the Botanical Research Institute of Texas 15 (1): 73-111, DOI: 10.17348/jbrit.v15.i1.1052, URL: https://doi.org/10.17348/jbrit.v15.i1.1052
03B06D5ABB2E663F41154996FF77C18B.text	03B06D5ABB2E663F41154996FF77C18B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phaseolus rotundatus Freytag & Debouck 2002	<div><p>Phaseolus rotundatus Freytag &amp; Debouck (2002: 132) [clade B; sect. Paniculati Freytag].</p><p>TYPE: R Lépiz 410511.1 (HOLOTYPE: US-3386751; ISOTYPES: ARIZ-347215, CS-77509, F, MICH-1180113).</p><p>Because the planting of original seed resulted in a big flowering (Fig. 8) and fruiting plant in Palmira, Colombia, additional observations were made, as well as a good series of reference specimens (Salcedo-Castaño et al. 2009). It is a tall plant with scattered populations in W Mexico, the habitat of which is shrinking because of the recent expansion of agave planting. The author has doubts that the specimen HS Gentry 22509 (ARIZ-321197, NA-27957) belongs to P. rotundatus (see Freytag &amp; Debouck 2002, p. 134).</p><p>Specimens: MEXICO. Jalisco: MC de Anda Virgen 24 (IBUG-134932), HS Gentry 22188 (ARIZ-300442). Michoacán: R Lépiz-Ildefonso 496 (IBUG-208608).</p></div>	https://treatment.plazi.org/id/03B06D5ABB2E663F41154996FF77C18B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Debouck, D. G.	Debouck, D. G. (2021): Phaseolus Beans (Leguminosae: Phaseoleae): A Checklist And Notes On Their Taxonomy And Ecology. Journal of the Botanical Research Institute of Texas 15 (1): 73-111, DOI: 10.17348/jbrit.v15.i1.1052, URL: https://doi.org/10.17348/jbrit.v15.i1.1052
03B06D5ABB21663F41554CE5FC03C0AE.text	03B06D5ABB21663F41554CE5FC03C0AE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phaseolus salicifolius Piper 1926	<div><p>Phaseolus salicifolius Piper (1926: 692) [clade B; sect. Paniculati Freytag].</p><p>TYPE: J González-Ortega 4065 (HOLOTYPE: US-1083789; isotype: MEXU n.v.).</p><p>A species still considered as endemic (its range seems to be the Sierra Madre Occidental along the border of the state of Sinaloa in pine-oak woodlands at 1300–2500 m, pending on more data (Fig. 8). The author has doubts that the specimen Ed Palmer 142 (K, US-571163) belongs to P. salicifolius (see Delgado-Salinas 1985, p. 215).</p><p>Specimens: MEXICO. Durango: DG Debouck 411 (K-1041820). Sinaloa: DE Breedlove 1689 (DUKE-157400).</p></div>	https://treatment.plazi.org/id/03B06D5ABB21663F41554CE5FC03C0AE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Debouck, D. G.	Debouck, D. G. (2021): Phaseolus Beans (Leguminosae: Phaseoleae): A Checklist And Notes On Their Taxonomy And Ecology. Journal of the Botanical Research Institute of Texas 15 (1): 73-111, DOI: 10.17348/jbrit.v15.i1.1052, URL: https://doi.org/10.17348/jbrit.v15.i1.1052
03B06D5ABB21663F414A4D9FFC2BC374.text	03B06D5ABB21663F414A4D9FFC2BC374.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phaseolus scrobiculatifolius Freytag 2002	<div><p>Phaseolus scrobiculatifolius Freytag (2002: 126) [clade B; sect. Paniculati Freytag].</p><p>TYPE: CP Cowan 4902 (HOLOTYPE: TEX-371373; isotype MEXU n.v.).</p><p>This taxon has been put in synonymy of P.jaliscanus (Delgado-Salinas et al. 2006), a move that the author is not ready to make at this time, because it lacks the heteroblastic development, so common in P. jaliscanus .</p><p>Specimens: MEXICO. Michoacán: AE Estrada C. 2354 (MEXU-1109087), R Lépiz-Ildefonso 502 (IBUG-208610).</p></div>	https://treatment.plazi.org/id/03B06D5ABB21663F414A4D9FFC2BC374	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Debouck, D. G.	Debouck, D. G. (2021): Phaseolus Beans (Leguminosae: Phaseoleae): A Checklist And Notes On Their Taxonomy And Ecology. Journal of the Botanical Research Institute of Texas 15 (1): 73-111, DOI: 10.17348/jbrit.v15.i1.1052, URL: https://doi.org/10.17348/jbrit.v15.i1.1052
03B06D5ABB21663F41554E50FDF1C5B5.text	03B06D5ABB21663F41554E50FDF1C5B5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phaseolus sinuatus Nuttall ex Torrey & A. Gray 1838	<div><p>Phaseolus sinuatus Nuttall ex Torrey &amp; A. Gray (1838: 279) [clade B; sect. Paniculati Freytag].</p><p>TYPE: N Ware s.n. (SYNTYPE: PH-28331) .</p><p>John Torrey and Asa Gray mentioned two specimens for P.sinuatus (a name they have seen in a manuscript by Nathaniel Nuttall): one collected by Mr. Ware (studied at the Academy of Natural Sciences of Philadelphia and afore-mentioned) and another one by Dr. Burrows (not seen during this study). No strong arguments have been found to consider it as a variety or subspecies of P. polystachyus . It is worth remembering the footnote by Asa Gray (1852, p. 43): “ P.sinuatus Nutt., of which I have five specimens from Rugel’s Florida collection, No. 132, is widely different from P. perennis (since a synonym of P.polystachyus)”. The leaflets are distinctly (small round) lobed, the stems trailing instead of climbing and with heteroblastic development, and the perennial root system is much more globose. Further, it occupies a specific ecological niche of the SE United States: the longleaf and loblolly pine forests on sandy soils (Radford et al. 1968) at 100– 140 m.</p><p>Specimens: U.S.A. North Carolina: JA Duke 1768 (NCU-126435). South Carolina: CR Bell 11063 (NCU-174816). Georgia: LC Anderson 15917 (BRIT). Florida: EA Bessey sn (MSC), F Rugel 132 (NA).</p><p>The specimen collected by Charles Louis Pollard 1017 (F-48720, GH, NY) from Jackson Co. in Mississippi belongs to Rhynchosia cinerea Nash. While P. sinuatus has been claimed to be present in Alabama and Mississippi (Small 1903; Radford et al. 1968), no specimens from these states have been seen during this study (thus the indication of intermediate range in Table 2).</p></div>	https://treatment.plazi.org/id/03B06D5ABB21663F41554E50FDF1C5B5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Debouck, D. G.	Debouck, D. G. (2021): Phaseolus Beans (Leguminosae: Phaseoleae): A Checklist And Notes On Their Taxonomy And Ecology. Journal of the Botanical Research Institute of Texas 15 (1): 73-111, DOI: 10.17348/jbrit.v15.i1.1052, URL: https://doi.org/10.17348/jbrit.v15.i1.1052
03B06D5ABB21663F41554890FD24C4FB.text	03B06D5ABB21663F41554890FD24C4FB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phaseolus smilacifolius Pollard 1896	<div><p>Phaseolus smilacifolius Pollard (1896: 233) [clade B; sect. Paniculati Freytag].</p><p>TYPE: GV Nash 2505 (HOLOTYPE: US-252673; ISOTYPES: F-48642, G, GH-64080, K, MICH-1104263, MSC, P, PH).</p><p>The fact that this species has been found in another county of Florida 101 years after the type collection lessens seriously the hypothesis of this species being a natural hybrid between P. polystachyus and P. sinuatus (Isely 1990, p. 156) . Further, the planting of seeds with a progeny identical to the parents confirmed P.smilacifolius as a distinct species (Abbott &amp; Judd 2000, p. 497).With few collections from central Florida, it should be considered an endemic; from these few collections it is reported growing on hammock land in oak-juniper-palmetto association. According to Dohle and co-workers (2019), it seems that no accessions are currently kept in genebanks.</p><p>Specimens: U.S.A. Florida: JR Abbott 9557 (WIS), AS Hitchcock 483 (F-232856).</p></div>	https://treatment.plazi.org/id/03B06D5ABB21663F41554890FD24C4FB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Debouck, D. G.	Debouck, D. G. (2021): Phaseolus Beans (Leguminosae: Phaseoleae): A Checklist And Notes On Their Taxonomy And Ecology. Journal of the Botanical Research Institute of Texas 15 (1): 73-111, DOI: 10.17348/jbrit.v15.i1.1052, URL: https://doi.org/10.17348/jbrit.v15.i1.1052
03B06D5ABB21663E415549D5FF33C18B.text	03B06D5ABB21663E415549D5FF33C18B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phaseolus sonorensis Standley 1940	<div><p>Phaseolus sonorensis Standley (1940a: 31) [clade B; sect. Paniculati Freytag].</p><p>TYPE: HS Gentry 1396 (HOLOTYPE: F-770502; ISOTYPES: BM, DES, GH-64109, K, MICH-1104264, MO-1089160, UC-646354, WIS-3983).</p><p>A tall plant growing in pine-oak forest at 1100–1800 m with large showy magenta flowers. It blooms in spring as often does P. salicifolius and P.albinervus too. Pending additional field work, it is still considered as endemic (an opinion shared by Fishbein et al. 1998), but it might have a larger range in the Sierra Madre Occidental.</p><p>Specimens: MEXICO. Chihuahua: R Spellenberg 8081 (NMC-14198). Sinaloa: RA González 236 (MEXU-800335). Sonora: M Fishbein 1015 (ARIZ-304231).</p></div>	https://treatment.plazi.org/id/03B06D5ABB21663E415549D5FF33C18B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Debouck, D. G.	Debouck, D. G. (2021): Phaseolus Beans (Leguminosae: Phaseoleae): A Checklist And Notes On Their Taxonomy And Ecology. Journal of the Botanical Research Institute of Texas 15 (1): 73-111, DOI: 10.17348/jbrit.v15.i1.1052, URL: https://doi.org/10.17348/jbrit.v15.i1.1052
03B06D5ABB20663E41154CE5FC6CC08E.text	03B06D5ABB20663E41154CE5FC6CC08E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phaseolus talamancensis Debouck & Torres-Gonzalez 2001	<div><p>Phaseolus talamancensis Debouck &amp; Torres-González (2001: 280) [clade A; sect. Bracteati Freytag].</p><p>TYPE: Debouck et al. 2130 (HOLOTYPE: CR-224324; ISOTYPES: K-478463, MO-5260537).</p><p>An exploration carried out in 2003 in Costa Rica informed us that this species although uncommon can be found outside the Talamanca range sensu stricto: population #3182 was found in the county of Santa MarÍa de Dota.During the cultivation of population #3182 for seed conservation, the species eventually produced tuberous roots (Fig. 12). This taxon grows in openings of the lower montane oak forest at 1500–2000 m.</p><p>Specimens: COSTA RICA. Puntarenas: E Alfaro 2508 (CR-247762). San José: DG Debouck 3182 (CR-238742), JF Morales 8861 (K).</p></div>	https://treatment.plazi.org/id/03B06D5ABB20663E41154CE5FC6CC08E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Debouck, D. G.	Debouck, D. G. (2021): Phaseolus Beans (Leguminosae: Phaseoleae): A Checklist And Notes On Their Taxonomy And Ecology. Journal of the Botanical Research Institute of Texas 15 (1): 73-111, DOI: 10.17348/jbrit.v15.i1.1052, URL: https://doi.org/10.17348/jbrit.v15.i1.1052
03B06D5ABB20663E41154DFFFF1EC35B.text	03B06D5ABB20663E41154DFFFF1EC35B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phaseolus tenellus Piper 1926	<div><p>Phaseolus tenellus Piper (1926: 696) [clade A; sect. Minkelersia (Martens &amp; Galeotti) Maréchal, Mascherpa, &amp; Stainier].</p><p>TYPE: G Arsène 8310 (HOLOTYPE: US-1003657 n.v.; ISOTYPES: BM, F-48495, GH-64112, MO-541731, SI).</p><p>A member of the Minkelersia found in openings of pine-oak forest at 2100–2900 m.</p><p>Specimens: MEXICO. Zacatecas: CL Díaz-Luna 2352 (MICH-1180118). Michoacán: A Delgado-Salinas 1775 (K-1041785). México: GB Hinton 6528 (GH).</p></div>	https://treatment.plazi.org/id/03B06D5ABB20663E41154DFFFF1EC35B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Debouck, D. G.	Debouck, D. G. (2021): Phaseolus Beans (Leguminosae: Phaseoleae): A Checklist And Notes On Their Taxonomy And Ecology. Journal of the Botanical Research Institute of Texas 15 (1): 73-111, DOI: 10.17348/jbrit.v15.i1.1052, URL: https://doi.org/10.17348/jbrit.v15.i1.1052
03B06D5ABB20663E410A4EB5FC2CC27E.text	03B06D5ABB20663E410A4EB5FC2CC27E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phaseolus texensis A. Delgado & W. R. Carr 2007	<div><p>Phaseolus texensis A. Delgado &amp; W.R. Carr (2007: 13) [clade A; sect. Pedicellati (Bentham) Freytag].</p><p>TYPE: WR Carr 24232 (HOLOTYPE: TEX; ISOTYPE: MEXU n.v.).</p><p>This species seems to be endemic from the Edwards Plateau of SW Texas —an area noteworthy for its plant endemism (Johnston 1997)—and is included into the Pedicellati (Delgado-Salinas &amp; Carr 2007). It thrives in mixed woodland with oak, elm, ash, sycamore and black cherry at 170– 580 m.</p><p>Specimens: U.S.A. Texas. Kerr Co.: VL Cory 52444 (BRIT), GF Freytag 90-1 (CS-77512). Travis Co.: T Wendt 7379 (TEX).</p></div>	https://treatment.plazi.org/id/03B06D5ABB20663E410A4EB5FC2CC27E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Debouck, D. G.	Debouck, D. G. (2021): Phaseolus Beans (Leguminosae: Phaseoleae): A Checklist And Notes On Their Taxonomy And Ecology. Journal of the Botanical Research Institute of Texas 15 (1): 73-111, DOI: 10.17348/jbrit.v15.i1.1052, URL: https://doi.org/10.17348/jbrit.v15.i1.1052
03B06D5ABB20663E41154F6FFF34C528.text	03B06D5ABB20663E41154F6FFF34C528.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phaseolus trifidus Freytag 2002	<div><p>Phaseolus trifidus Freytag (2002: 181) [clade A; sect. Digitati Freytag].</p><p>TYPE: FG Meyer &amp; DJ Rogers 2899 (HOLOTYPE: MO-1597351; ISOTYPES: BM, G).</p><p>This species only known from the type specimens has been put in synonymy of P.neglectus (Delgado-Salinas et al. 2006), a move that is kept on hold, because the terminal leaflets are differently lobed, and the ratio peduncle/ floral rachis is different as compared to the former (in addition to the different morphology of primary bracts).</p></div>	https://treatment.plazi.org/id/03B06D5ABB20663E41154F6FFF34C528	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Debouck, D. G.	Debouck, D. G. (2021): Phaseolus Beans (Leguminosae: Phaseoleae): A Checklist And Notes On Their Taxonomy And Ecology. Journal of the Botanical Research Institute of Texas 15 (1): 73-111, DOI: 10.17348/jbrit.v15.i1.1052, URL: https://doi.org/10.17348/jbrit.v15.i1.1052
03B06D5ABB20663E41154805FD56C484.text	03B06D5ABB20663E41154805FD56C484.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phaseolus tuerckheimii Donnell-Smith 1913	<div><p>Phaseolus tuerckheimii Donnell-Smith (1913:54) [clade A; sect. Brevilegumeni Freytag].</p><p>TYPE: H von Tuerckheim II-1536 (LECTOTYPE: US-942247; ISOLECTOTYPES: BR, US-860832, US-942248) . SYNTYPE: H Pittier 10539 (US-942246) .</p><p>John Donnell Smith referred to two collections, the one by Hans von Türckheim (no. II-1536) and the one by Henri Pittier (no. 10539) in that order, without indicating the holotype. As there were several duplicates of no. II-1536 at the US National Herbarium, the selection of the voucher US-942247 by George Freytag as nomenclatural type seemed appropriate: it would have been impossible for JD Smith to write down a description of the floral parts of P.tuerckheimii without dissecting its flower! This species is a tall (6–10 m) vine growing in the moist montane forests of Central America at 1500–2600 m (Araya-Villalobos et al. 2001) (Fig. 3).</p><p>Specimens: MEXICO. Chiapas: U Bachem 398 (CHAPA). GUATEMALA. Huehuetenango: LO Williams 22323 (F-1656438). El Quiché: AJ Sharp 45191 (MEXU-136531). Quezaltenango: DG Debouck 3085 (AGUAT). Sololá: DG Debouck 3097 (AGUAT). Chimaltenango: DG Debouck 1624 (MICH-1180126). Sacatepéquez: M Véliz 14771 (MO-5871374). Guatemala: DG Debouck 2473 (US-3168469). Jalapa: JA Steyermark 32551 (F-1035022). El Progreso: DG Debouck 2418 (COL-324860). Zacapa: JA Steyermark 42471 (F-1202181). COSTA RICA. San José: JL Luteyn 4541 (DUKE-242717). PANAMA. Chiriquí: R Villareal 494 (PMA-108353).</p></div>	https://treatment.plazi.org/id/03B06D5ABB20663E41154805FD56C484	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Debouck, D. G.	Debouck, D. G. (2021): Phaseolus Beans (Leguminosae: Phaseoleae): A Checklist And Notes On Their Taxonomy And Ecology. Journal of the Botanical Research Institute of Texas 15 (1): 73-111, DOI: 10.17348/jbrit.v15.i1.1052, URL: https://doi.org/10.17348/jbrit.v15.i1.1052
03B06D5ABB20663D411549E0FE93C43D.text	03B06D5ABB20663D411549E0FE93C43D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phaseolus venosus Piper 1926	<div><p>Phaseolus venosus Piper (1926: 687) [clade B; sect. Coriacei Freytag].</p><p>TYPE: JN Rose 2688 (HOLOTYPE: US-301614; no isotype found during this study).</p><p>This species has been put in synonymy of P.maculatus (Delgado-Salinas et al. 2006) but differences in size of plant, leaflet nervation and acumen, shape, nervation and size of primary bracts as compared to the ones found in the “Metcalfe bean” prevent to make this move till more living material allows cross comparisons. It has been seen in pine-oak woodland at 2000–2200 m. This taxon is not reported in the list of “Vascular Plants of the Americas” (Tropicos 2020).</p><p>Specimens: MEXICO. Zacatecas: G Starr 672 (UCR-32721). Aguascalientes: S Gallegos G. 12 (IBUG-52731). Jalisco: R Ramírez-Delgadillo 1447 (IBUG-99829).</p></div>	https://treatment.plazi.org/id/03B06D5ABB20663D411549E0FE93C43D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Debouck, D. G.	Debouck, D. G. (2021): Phaseolus Beans (Leguminosae: Phaseoleae): A Checklist And Notes On Their Taxonomy And Ecology. Journal of the Botanical Research Institute of Texas 15 (1): 73-111, DOI: 10.17348/jbrit.v15.i1.1052, URL: https://doi.org/10.17348/jbrit.v15.i1.1052
03B06D5ABB23663D4155492BFF78C746.text	03B06D5ABB23663D4155492BFF78C746.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phaseolus viridis Piper 1926	<div><p>Phaseolus viridis Piper (1926: 693) [clade B; sect. Paniculati Freytag].</p><p>TYPE: H von Tuerckheim II-897 (= John Donnell-Smith 8510) (HOLOTYPE: US-576636 n.v.; ISOTYPES: F-185839, GH-64113, O, US-942171).</p><p>This species seems to be valid, namely because of flower, pod (Fig. 10) and seed number traits, in spite of previous transfers into synonymy of P. lunatus (Delgado-Salinas 1985, Freytag &amp; Debouck 2002). It is known by scattered populations across tropical Mexico and Guatemala and found in semi-evergreen seasonal forest at 10– 730 m. Studies with molecular markers on ITS/ 5.8S DNA and trnK cpDNA (Delgado-Salinas et al. 1999, 2006) have indicated a clear difference with P. lunatus . Phaseolus longiplacentifer Freytag would be a synonym of this species (Delgado-Salinas et al. 2006). Phaseolus longiplacentifer is reported in the list of “Vascular Plants of the Americas” (Tropicos 2020), while P.viridis is not.</p><p>Specimens: MEXICO. Veracruz: M Botteri 680 (P). Oaxaca: G Juárez-García 1086 (ARIZ-389122). Chiapas: E Palacios-Espinosa 1757 (CAS-852127).</p></div>	https://treatment.plazi.org/id/03B06D5ABB23663D4155492BFF78C746	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Debouck, D. G.	Debouck, D. G. (2021): Phaseolus Beans (Leguminosae: Phaseoleae): A Checklist And Notes On Their Taxonomy And Ecology. Journal of the Botanical Research Institute of Texas 15 (1): 73-111, DOI: 10.17348/jbrit.v15.i1.1052, URL: https://doi.org/10.17348/jbrit.v15.i1.1052
03B06D5ABB22663C41154CBCFFCAC5FA.text	03B06D5ABB22663C41154CBCFFCAC5FA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phaseolus vulgaris L. 1753	<div><p>Phaseolus vulgaris L. (1753: 723) [clade B; sect. Phaseoli DC emend. Freytag].</p><p>TYPE: C Linnaeus s.n. (LECTOTYPE: LINN-899.1) .</p><p>The type of the reference species of the section and of the genus is a plant cultivated at Uppsala, Sweden (Verdcourt 1971). Ironically, the most common (Linnaeus was right in naming it vulgaris !) species of the genus has its wild ancestor disclosed quite late: Arturo Burkart in Salta, NW Argentina in 1941 (Burkart 1941, 1952), and Felix Webster McBryde in W Guatemala in 1947, and not referring to each other’s finding! Keeping P.aborigineus Burkart as a separate species (as suggested in the List of Tropicos 2020) might result in hiding the biological link with the Andean cultivated form and its three races (Singh et al. 1991). The double independent domestication of the wild form of common bean distributed from Chihuahua (Nabhan 1985) down to Córdoba (Drewes 2008) is now widely agreed upon (Chacón-Sánchez et al. 2005; Kwak &amp; Gepts 2009; Bitocchi et al. 2013). There would be enough evidence (Tohme et al. 1996; Chacón-Sánchez et al. 2007; Ariani et al. 2017) to recognize five wild genetic pools: one W of Isthmus of Tehuantepec (Chihuahua down to Oaxaca), one including Chiapas, Guatemala, Honduras, El Salvador and NW Nicaragua, one including Costa Rica, one including the NE Andes of Colombia and W Andes of Venezuela, and one including the central and southeastern Andes of Peru (Huánuco down to Cuzco), eastern Andes of Bolivia (Cochabamba down to Tarija), and NW Andes of Argentina (Jujuy down to Córdoba; although Salinas Grandes might be another break). The significance of a formal treatment of botanical varieties, i.e., a var. mexicanus for the Mexican and Central American wild forms and a var. aborigineus for the Andean one (Delgado-Salinas et al. 1988), is pending on what the wild forms from Venezuela will reveal (Beebe &amp; Debouck 2019), while the recognition of a var. vulgaris for all cultivated forms (Baudet 1977) may blur their complex origins.The following specimens (different from the ones reported previously in Freytag &amp; Debouck 2002 p. 38–39) are wild forms and give an indication about the range.</p><p>Specimens: MEXICO. Durango: DG Debouck 276 (COL-324695). Zacatecas: JM González-Correa 30 (IBUG-29683). Nayarit: RS Ferris 5946 (DS-214469). Jalisco: JS Peñaflor-Cortés 9 (IBUG-36127). Colima: HS Gentry 22189 (ARIZ-300349). México: A Delgado-S. 1006 (CHAPA). Morelos: E Bourgeau 1375 (FI). Puebla: R Torres-C. 7544 (RSA-537099). Michoacán: DG Debouck 439 (COL-324743). Guerrero: GB Hinton 11551 (POM-348560). Oaxaca: B Bartholomew 3033 (CAS-747013). Chiapas: E Cabrera 5930 (CAS-819944). GUATEMALA. Sacatepéquez: DG Debouck 1610 (K). Santa Rosa: E Martínez-S.19230 (MEXU-526662). Jalapa: DG Debouck 2432 (COL-324875). HONDURAS. El Paraíso: JG Hawkes 2053 (K). Morazán: A Molina R. 731 (F-1262709). EL SALVADOR. Sonsonate: DG Debouck 3226 (LAGU). La Libertad: DG Debouck 3221 (LAGU). NICARAGUA. Madríz: DG Debouck 3202 (O-V2147425). Estelí: DG Debouck 3216 (O-V2147423). COSTA RICA. Alajuela: DG Debouck 3133 (CR-235446). Cartago: DG Debouck 3126 (CR-247283). San José: CO Morales 621 (USJ-48426). VENEZUELA. Mérida: O Berglund 26667 (SI). Lara: G Davidse 21172 (NY). COLOMBIA. Cundinamarca: JO Toro-Chica 498 (COL-520043). PERU. Huánuco: JF Macbride 3947 (G). Junín: R Ferreyra 14964 (USM-71284). Huancavelica: O Tovar 3910 (US-2490506). Apurimac: C Ochoa 2420 (MOL). Cuzco: OF Cook 1073 (US-604244). BOLIVIA. Cochabamba: R Caballero-Pardo s.n. (NY). Chuquisaca: DG Debouck 3007 (DAV-128327). Tarija: DG Debouck 3020 (MO-5613149). ARGENTINA. Jujuy: O Ahumada 4173 (SI). Salta: R Fortunato 3576 (BAB). Tucumán: M Lillo 1626 (LIL-62049). Catamarca: DG Debouck 645 (COL-324753). San Luís: R Batallanez s.n. (SI). Córdoba: T Stuckert 12831 (SI).</p></div>	https://treatment.plazi.org/id/03B06D5ABB22663C41154CBCFFCAC5FA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Debouck, D. G.	Debouck, D. G. (2021): Phaseolus Beans (Leguminosae: Phaseoleae): A Checklist And Notes On Their Taxonomy And Ecology. Journal of the Botanical Research Institute of Texas 15 (1): 73-111, DOI: 10.17348/jbrit.v15.i1.1052, URL: https://doi.org/10.17348/jbrit.v15.i1.1052
03B06D5ABB22663C411548EBFD97C4D0.text	03B06D5ABB22663C411548EBFD97C4D0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phaseolus xanthotrichus Piper 1926	<div><p>Phaseolus xanthotrichus Piper (1926: 698) [clade A; sect. Xanthotricha A. Delgado].</p><p>TYPE: ET Heyde &amp; E Lux 4171 (HOLOTYPE: US-247544 n.v.; ISOTYPES: GH-64114, M, US-942184).</p><p>While its habitat (lower montane moist forest with alder, oak, pine, cedar, at 1400–2400 m) has been invaded by Coffea arabica mostly over the last two centuries, it has been able to survive under the fences of coffee plantations where herbicide sprays are less frequent.</p><p>Specimens: MEXICO. Chiapas: DE Breedlove 53235 (CAS-668979). GUATEMALA. Sacatepéquez: M Véliz 7362 (MEXU-999506). Santa Rosa: DG Debouck 2438 (US-3168340). EL SALVADOR. Ahuachapán: EA Montalvo 4596 (ITIC). Santa Ana: DG Debouck 3229 (LAGU). Sonsonate: EA Montalvo 4926 (ITIC). COSTA RICA. San José: DG Debouck 3176 (CR-247291), DG Debouck 3358 (USJ-111605), B Hammel 23864 (INB-3974385). Cartago: DG Debouck 3125 (USJ-88822).</p></div>	https://treatment.plazi.org/id/03B06D5ABB22663C411548EBFD97C4D0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Debouck, D. G.	Debouck, D. G. (2021): Phaseolus Beans (Leguminosae: Phaseoleae): A Checklist And Notes On Their Taxonomy And Ecology. Journal of the Botanical Research Institute of Texas 15 (1): 73-111, DOI: 10.17348/jbrit.v15.i1.1052, URL: https://doi.org/10.17348/jbrit.v15.i1.1052
03B06D5ABB22663B41154A3CFDB8C1DD.text	03B06D5ABB22663B41154A3CFDB8C1DD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phaseolus xolocotzii A. Delgado 2000	<div><p>Phaseolus xolocotzii A. Delgado (2000: 429) [clade B; sect. Paniculati Freytag].</p><p>TYPE: R Torres-Colin 7700 (HOLOTYPE: MEXU n.v.) .</p><p>A member of the Paniculati known by scattered populations along the internal slope of W Sierra Madre del Sur up to the S slope of the Trans-Mexican Volcanic Belt, in dry pine-oak forest at 1200–1900 m. The author has doubts that the specimen DG Debouck 2346 (BR, COL-58966, M, SI, US-3168881) belongs to P. xolocotzii (see Freytag &amp; Debouck 2002, p. 137).</p><p>Specimen: MEXICO. México: GB Hinton 2484 (K-118678).</p></div>	https://treatment.plazi.org/id/03B06D5ABB22663B41154A3CFDB8C1DD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Debouck, D. G.	Debouck, D. G. (2021): Phaseolus Beans (Leguminosae: Phaseoleae): A Checklist And Notes On Their Taxonomy And Ecology. Journal of the Botanical Research Institute of Texas 15 (1): 73-111, DOI: 10.17348/jbrit.v15.i1.1052, URL: https://doi.org/10.17348/jbrit.v15.i1.1052
03B06D5ABB25663B41554CCBFEFCC0ED.text	03B06D5ABB25663B41554CCBFEFCC0ED.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phaseolus zimapanensis A. Delgado 2000	<div><p>Phaseolus zimapanensis A. Delgado (2000: 432) [clade A; sect. Xanthotricha A. Delgado].</p><p>TYPE: A Delgado-Salinas 1113 (HOLOTYPE: MEXU n.v.; ISOTYPES: K-651818, NY-1104835) .</p><p>A member of the section Xanthotricha with distribution in the central dry highlands of Mexico along the western slope of Sierra Madre Oriental, in oak, pine, pine-oak, cedar forest and mesquite scrub at 900–2400 m.</p><p>Specimens: MEXICO. Coahuila: JA Villareal-Quintanilla 3961 (ANSM-37738). Tamaulipas: TF Daniel 307 (MICH-1180184). Nuevo León: DG Debouck 1513 (UC-1716197). San Luís Potosí: OS Fearing 2042 (TEX-273467). Queretaro: UT Waterfall 14114 (IA-111313). Hidalgo: A Frias-Castro 701 (IBUG-182428).</p></div>	https://treatment.plazi.org/id/03B06D5ABB25663B41554CCBFEFCC0ED	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Debouck, D. G.	Debouck, D. G. (2021): Phaseolus Beans (Leguminosae: Phaseoleae): A Checklist And Notes On Their Taxonomy And Ecology. Journal of the Botanical Research Institute of Texas 15 (1): 73-111, DOI: 10.17348/jbrit.v15.i1.1052, URL: https://doi.org/10.17348/jbrit.v15.i1.1052
