taxonID	type	description	language	source
03BDA235A4766F7C59CBA3E08EF2F880.taxon	description	Plants very small, soft, forming small flat mats, green or yellowish green, silky. Stems fragile, prostrate, irregularly branched, branches short, mostly densely foliate, julaceous; central strand and hyalodermis not differentiated, all cells ± thin-walled; rhizoids clastered near branch bases and developed on dorsal side of costa in lower part of leaves; paraphyllia absent; juvenile branch leaves narrow lanceolate or linear, sutuated on branch bases. Leaves loosely appressed when dry, occasionally slightly secund, spreading when moist, concave, ovate, ovate-lanceolate or lanceolate, abruptly or gradually acuminate, often piliferous; margins plane, dentate, denticulate or subentire, marginal teeth unicellular, short or long, or large, multicellular; costa slender, ending at midleaf; lamina unistratose, smooth; leaf cells rhomboidal or long-rhomboidal, with moderately thickened, non-porose walls; alar cells differentiated, quadrate and short rectangular, numerous. Autoicous. Perichaetial leaves broader than vegetative leaves. Setae erect, slender, yellowish or brownish, smooth. Capsules erect, ovoid, pyriform or cylindrical; annulus absent; exothecial cells quasrate and short rectangular, with wavy walls, stomata present, with elongate pore; peristome single, consisting of 16 exostome teeth usually fused in pairs, broadly lanceolate, blunt, light brown or dark red-brown, densely papillose-striolate on both surfaces, without trabeculae on both sides, usually reflexed when dry and appressed to the capsule wall, fragile, often broken in opened capsules. Opercula low conic or almost flat, mammillate or obliquely rostrate. Spores small, smooth or finely papillose.	en	Ignatova, Elena A., Kuznetsova, Oxana I., Milyutina, Irina A., Fedosov, Vladimir E., Ignatov, Michael S. (2017): The genus Fabronia (Fabroniaceae, Bryophyta) in Russia. Arctoa 26 (1): 11-34, DOI: 10.15298/arctoa.26.02, URL: https://doi.org/10.15298/arctoa.26.02
03BDA235A4766F7D59C3A7E58E12F887.taxon	materials_examined	Type: Russia, Altai Republic, Ulagan District, left slope to Chulyshman River valley ca. 1 km downstream from Chulcha River mouth, ca. 51 ° 05 ’ N, 87 ° 59 ’ E, 550 m a. s. l., at cliff base in Rhododendron thickets, on rocks covered with thin layer of soil, 15. VIII. 2012, coll. M. Ignatov & E. Ignatova # 12 - 747 (Holotype MHA, isotypes MW, LE, H, S).	en	Ignatova, Elena A., Kuznetsova, Oxana I., Milyutina, Irina A., Fedosov, Vladimir E., Ignatov, Michael S. (2017): The genus Fabronia (Fabroniaceae, Bryophyta) in Russia. Arctoa 26 (1): 11-34, DOI: 10.15298/arctoa.26.02, URL: https://doi.org/10.15298/arctoa.26.02
03BDA235A4766F7D59C3A7E58E12F887.taxon	description	Illustrations: Figs. 12, 5: 7 – 11, 6: 6 – 10, 8: 7 – 12; 11: 14 – 17.	en	Ignatova, Elena A., Kuznetsova, Oxana I., Milyutina, Irina A., Fedosov, Vladimir E., Ignatov, Michael S. (2017): The genus Fabronia (Fabroniaceae, Bryophyta) in Russia. Arctoa 26 (1): 11-34, DOI: 10.15298/arctoa.26.02, URL: https://doi.org/10.15298/arctoa.26.02
03BDA235A4766F7D59C3A7E58E12F887.taxon	diagnosis	Diagnosis: this species is close to Fabronia ciliaris and differs from it in lanceolate vs. ovate leaves, gradually vs. abruptly acuminate, longer leaf cells: 30 – 50 µm vs. 20 – 35 µm long, and ovate-cylindrical vs. ovate capsules. Plants small, silky-green, glossy, in dense patches. Stems creeping, ascending at upper part, ca. 5 mm long, irregularly branching; branches 2 – 3 mm long, erect or arcuate, densely foliate. Stem and branch leaves similar, loosely appressed when dry, straight or ± secund, spreading when moist, ovate-lanceolate or lanceolate, gradually long acuminate, 0.65 – 0.90 u 0.25 – 0.30 mm; leaf margins dentate, marginal teeth unicellular, moderate in size (20 – 30 µm long); costa slender, smooth, ending at mid-leaf; lamina smooth, upper and median laminal cells elongate-rhomboidal, (29 –) 35 – 50 (– 75) u 7 – 9 µm, with length / width ratio 4 – 6: 1 and cell length, µm / leaf length, mm ratio 42.7 – 70.9; apical leaf cell to 100 µm long; alar cells quadrate to short-rectangular, forming a weakly delimited rectangular group 3 – 4 cells wide and 7 – 11 cells along leaf margin. Autoicous, usually with sporophytes. Perigonia bud-like. Perichaetial leaves ca. 0.8 mm long and 0.4 mm wide, with oblong base and abruptly attenuate narrow acumen; costa ending in the acumen. Sporophytes single in perichaetium. Seta ca. 4 mm long, straight, flexuose when dry, yellow. Capsule ovate-cylindrical, 0.6 – 0.9 mm long and ca. 0.3 mm wide, slightly constricted below mouth when open, with short neck, smooth, pale brown, with dark-brown rim; annulus absent. Peristome single, consisting of 16 exostome teeth fused in pair, dark-brown, obtuse, ca. 150 µm long; outer surface densely papillose and striolate, inner surface with less prominent ornamentation, smooth in lower part of teeth and covered with low oblique ridges and occasionally with scarce papillae in distal part. Spores 13 – 15 µm, finely papillose. Operculum low conic, with short oblique beak.	en	Ignatova, Elena A., Kuznetsova, Oxana I., Milyutina, Irina A., Fedosov, Vladimir E., Ignatov, Michael S. (2017): The genus Fabronia (Fabroniaceae, Bryophyta) in Russia. Arctoa 26 (1): 11-34, DOI: 10.15298/arctoa.26.02, URL: https://doi.org/10.15298/arctoa.26.02
03BDA235A4766F7D59C3A7E58E12F887.taxon	materials_examined	Specimens examined: RUSSIA: ASIAN RUSSIA: Altai Republic: Ulagan District: Chulyshman River valley 1 km downstream Chulcha River mouth, 550 m alt., 15. VIII. 2012, Ignatov & Ignatova 12 - 745 (MHA); Shebalino District: Katun River valley near Ust-Sema, 580 m alt., 28. VII. 1991, Ignatov & Ignatova 24 / 62 (MHA); Chemal District: Katun River valley 10 km upstream from Chemal, 450 m alt., 7. VII. 1993, Ignatov & Ignatova 34 / 24 (MHA); Edikhta Creek (Aedigan Creek tributary), 1100 m alt., 8. VII. 1993, Ignatov & Ignatova 34 / 111 (MHA); Ongudai District: Malyj Yaloman Settl., 900 m alt., 30. VII. 1991, Ignatov & Ignatova 25 / 151 (MHA); Malyj Yaloman Settl., 950 m alt., 30. VII. 1991, Ignatov & Ignatova 25 / 158 (MHA); Malyj Yaloman Creek 8.5 km upstream from Katun River, 1100 m alt., 31. VII. 1991, Ignatov & Ignatova 25 / 53 (MHA); CAUCASUS: Dagestan Republic: Gunib District, Gunib settlement surroundings: 1550 m alt., 22. V. 2009, Ignatov & Ignatova 09 - 725 (MHA, MW); 1600 m alt., 21. V. 2009, Ignatov & Ukrainskaya 09 - 475 (MHA); 1560 m alt., 22. V. 2009, Ignatov et al. 14095 & 14098 (LE); Republic of Ingushetia: Aramkhi, 1300 m alt., 12. VII. 2005, Bersanova s. n. (MHA).? ITALY: Como, 8. IX. 1896 & 31. X. 1898, F. A. Artaria s. n. (LE). Plants from rich collection with many duplicates (seen in LE, H and S) from the northern Italy (Lombardy) is indistingishable from F. altaica by morphology. However, some specimens of F. major from Switzerland (proved by DNA) had laminal cells also more or less similar to F. altaica (cf. Fig. 7). Considering this, and also the absence of any other specimens of F. altaica from Central Europe, admitting that this specimen may represent an ultimate modification of F. major. In the illustrations it is shown under the name F. cf. altaica.	en	Ignatova, Elena A., Kuznetsova, Oxana I., Milyutina, Irina A., Fedosov, Vladimir E., Ignatov, Michael S. (2017): The genus Fabronia (Fabroniaceae, Bryophyta) in Russia. Arctoa 26 (1): 11-34, DOI: 10.15298/arctoa.26.02, URL: https://doi.org/10.15298/arctoa.26.02
03BDA235A4766F7D59C3A7E58E12F887.taxon	discussion	Differentiation. Fabronia altaica is rather variable in leaf shape and size (cf. Fig. 5: 7 – 11); however, there is rather clear difference in shape from F. ciliaris: ovate-lanceolate or lanceolate vs. ovate, and more gradually narrowed to the acumen vs. abruptly acuminate. There is a statistically proved difference in cell length between these species (Fig. 7). Fertile plants of F. altaica are readily distinguished from F. ciliaris by the shape of capsules: oblong, ovate-cylindrical vs. ovate. Peristome of F. altaica are usually darker colored, dark-brown vs. light-brown in F. ciliaris, but this difference may be caused by longer remaining color of peristome in the former species, while in young, deoperculate capsules they are of the same color. Opercula were rarely seen in F. altaica, but it seems that they are also different, conic and with short oblique beak, while in specimens of F. ciliaris we observed capsules mainly with mammillate opercula (though some exceptions were also seen: at least one specimen of F. ciliaris from Kabardino-Balkaria has capsules with obliquely rostrate opercula). In leaf shape, F. altaica resembles F. major, and in the beginning of the present study we did not separate them from each other. However, the latter species has longer cells, (36 –) 50 – 70 (– 95) µm (with few exceptions: three specimens from Switzerland had shorter cells, mainly 43 – 50 µm) but long marginal teeth. See also above the discussion about the specimen from northern Italy which resembles F. altaica in cell shape but more likely represents a deviating specimens of F. major. Capsules were found only in few collctions of F. major; they resemble F. altaica rather than F. ciliaris (see Fig. 11: 8 – 10, 14 – 17 and 1 – 7).	en	Ignatova, Elena A., Kuznetsova, Oxana I., Milyutina, Irina A., Fedosov, Vladimir E., Ignatov, Michael S. (2017): The genus Fabronia (Fabroniaceae, Bryophyta) in Russia. Arctoa 26 (1): 11-34, DOI: 10.15298/arctoa.26.02, URL: https://doi.org/10.15298/arctoa.26.02
03BDA235A4776F67599DA71A89E1FD6B.taxon	materials_examined	Holotype. “ Schleicher Botaunte Fabronia octoblepharis, Fabronia ciliaris Bryol. Univ. Hypnum [sericeulum?] Helvetia, 1807 ” (B 31 0677 01), http: // ww 2. bgbm. org / Herbarium / specimen. cfm? Barcode = B 31067701 (Röpert, 2000). Putative isotype in LE: “ Pterigynandrum obctoblepharis Schl., am Nov. genus?, ex Helvet ” [another later hardwriting at label bottom “ Fabronia ”].	en	Ignatova, Elena A., Kuznetsova, Oxana I., Milyutina, Irina A., Fedosov, Vladimir E., Ignatov, Michael S. (2017): The genus Fabronia (Fabroniaceae, Bryophyta) in Russia. Arctoa 26 (1): 11-34, DOI: 10.15298/arctoa.26.02, URL: https://doi.org/10.15298/arctoa.26.02
03BDA235A4776F67599DA71A89E1FD6B.taxon	description	Illustrations: Figs. 5: 1 – 6, 6: 1 – 5, 11: 1 – 7; Ignatov & Ignatova (2004): Fig 429 on page 647. Plants small, silky-green or yellow-green, glossy, in dense patches. Stems creeping, ca. 5 mm long, irregularly branching; branches 2 – 3 mm long, erect, densely foliate. Stem and branch leaves similar, loosely appressed when dry, straight, often with slightly recurved apices, spreading when moist, ovate, abruptly long acuminate, 0.5 – 0.9 u 0.25 – 0.3 mm; leaf margins denticulate, marginal teeth unicellular, small (ca. 0.1 mm long), or, rarely, consisting of 2 – 3 cells; costa slender, smooth, ending at midleaf; lamina smooth, upper and median laminal cells rhomboidal, 20 – 35 (– 40) u 8 – 11 µm, with length / width ratio 2 – 4: 1 and cell length, µm / leaf length, mm ratio 27.0 – 53.3; apical cell ca. 110 µm long; alar cells quadrate to short-rectangular, forming weakly delimited rectangular goup 3 – 4 cells wide and 6 – 9 cells along leaf margin. Autoicous, sporophytes rare. Perigonia bud-like, sometimes numerous. Perichaetial leaves 0.4 – 0.5 mm long and 0.3 – 0.4 mm wide, with oblong base and gradually attenuate narrow acumen; costa ending below apex; margins with longer, stronger recurved teeth in upper part. Sporophytes single in perichaetium. Seta 3 – 4 mm long, straight, flexuose when dry, yellow. Capsule ovate, with short neck, 0.35 – 0.45 mm long and 0.3 – 0.4 mm wide, often flared at mouth with age, smooth, pale brown, concolorous at rim or slightly darker; annulus absent. Peristome single, consisting of 16 exostome teeth fused in pair, light-brown or brown, obtuse, ca. 125 µm long; outer surface papillose-striolate, inner surface with less prominent ornamentation, in distal parts of teeth covered with low vertical ridges. Spores (10 –) 15 – 17 µm, papillose. Operculum low conic, mammillate or with short straight or, rarely, moderately long oblique beak.	en	Ignatova, Elena A., Kuznetsova, Oxana I., Milyutina, Irina A., Fedosov, Vladimir E., Ignatov, Michael S. (2017): The genus Fabronia (Fabroniaceae, Bryophyta) in Russia. Arctoa 26 (1): 11-34, DOI: 10.15298/arctoa.26.02, URL: https://doi.org/10.15298/arctoa.26.02
03BDA235A4776F67599DA71A89E1FD6B.taxon	materials_examined	Selected specimens examined: RUSSIA: CAUCASUS: Dagestan Republic: Gunib District: Gunib, 1550 m alt., 22. V. 2009, Ignatov & Ignatova 09 - 672 & 09 – 766 (MW); North Ossetia / Alania: Suargom Gorge, 800 m alt., 1. XII. 2014, Ukrainskaya 15945 (LE); Kabardino-Balkaria: Chereksky Disrict, Verkhnaya Balkaria Settl., 1100 – 1150 m alt., 28. VIII. 2005, Ignatov et al. 05 - 1825 & 05 - 1880 (MHA); Elbrussky District, Bylym, Baksan River valley, 1000 m alt., 30. VII. 2004, Ignatov et al. s. n. (MHA); Karachaevo-Cherkessia: Teberda Nature Reserve, Dzhemagat Creek valley, 1550 m alt., 22. VII. 1994, Onipchenko 115 / 94 (MW); Aksaut Gorge, 5. VI. 1989, Ukrainskaya 14444 (LE); Daut River valley, 4. VIII. 1993, Ukrainskaya 14443 (LE). URALS: Bashkortostan: Bashkirsky Nature Reserve, Kaga River mouth, 2. IX. 1945, Selivanova-Gorodkova s. n. (MHA); Chelyabinsk Province: Satka District, Ai River, 300 m alt., 6. VIII. 2011, Ibatullin 36 (MHA); Perm Territory: Kishertsky District, Kamen Vostryj, 2. VII. 1999, Bezgodov 268 (MW); ASIAN RUSSIA: Altai Republic: Ulagan District: Chulyshman River valley, 1950 m alt., 2. VIII. 1993, Ignatov 36 / 95 (MHA); Chulcha River valley, 1000 m alt., 11. VII. 1991, Ignatov 9 / 12 (MHA); northern shore of Teletzkoe Lake, Yailyu, 460 m alt., 1. VI. 1989, Ignatov 0 / 1054 (MHA); west of Yailyu, Izvestkovaya Mt., 650 m alt., 12. VI. 1989, Ignatov 0 / 669 (MHA); Artyshtu River 0.3 km of its mouth, 600 m alt., 20. IX. 1989, Zolotukhin s. n. (MHA); Kukol stow in the valley of Karakem River, 23. VI. 1989, Ignatov 0 / 1055 (MHA); Sebalino District: Katun River valley near Ust-Sema, 380 m alt., 28. VII. 1991, Ignatov & Ignatova 24 / 163 (MHA); Chemal District: Chemal Creek 3 rm upstream mouth, 450 m alt., 12. VII. 1993, Ignatov & Ignatova 34 / 63 (MHA); Irkutsk Province, Irkutsk surroundings, 10. VII. 1932, Kostin s. n. (LE); Baikal Lake, Maloe More, Yadyrtui Cape, 19. VII. 1987, Bardunov s. n. (LE); Zabaikalsky Territory: Dul’durga District, Alkhanai National Park, Ilya River, 8. VII. 2006, Afonina 1606 (LE); Kyra District: Sokhondinsky Biosphere Reserve: Agutsa River, 1098 m alt., 18. VII. 2010, Afonina A 3710 (LE); Enda River, 1320 m alt., 12. VII. 2010, Afonina A 1910 (LE); Bukukun River, 1646 m alt., Czernyadjeva 42 - 11 (LE); Ulota District: Ingoda River ner Leninskiy Settlement, 6. VIII. 2011, Afonina 0611 (LE); Krasnochikoisky District: Zhindokon Creek, 963 m alt., 8. VIII. 2011, Afonina 1411 (LE); Nerchinsk District: Nerchinsko-Kuenginsky Rangem Marikta River, 15. VII. 2012, Czernyadjeva 7 - 12 (LE); Aleksandrovsky Zavod District, Malaya Borzya River, 817 m alt., 27. VII. 2012, Afonina 4412 (LE); Tungokochin District, Ul’durga River, 618 m alt., 16. VII. 2012, Afonina 1112 (LE); Chita District, Aratsa River, 964 m alt., 13. VII. 2012, Afonina 0312 (LE); Republic of Buryatia: Selenga River near Novoselenginsk Settlement, 4. VII. 2007, Afonina 00707 (LE); Amurskaya Province: Selemdzhinsky District: Norskaya Hill, 220 m alt., 21. VI. 2011, Bezgodov 391 (MHA); Nora River, 240 m alt., 15. VII. 2010, Bezgodov 460 (MHA); Khabarovsk Territory: Verkhnebureinsky District: Chegdomyn, 450 m alt., Ignatov 97 - 284 (MHA); Sovgavan District: Botchinsky State Reserve: Spokoiny Creek, 290 m alt., 22. VIII. 2013, Ignatov & Ignatova 13 - 1035 (MHA); Mulpa River, 245 m alt., 14. VIII. 2013, Ignatov & Ignatova 13 - 494 (MHA); Primorsky Territory: Partizank District: Chandolaz Mt., 250 m alt., 8. IX. 2013, Ignatov et al. 13 - 1840 (MHA); Shkotovo District: Ussurijsky Nature Reserve, Koryavaya Pad’, Zmeinaya Mt., 331 m alt., 15. X. 2008, Ignatov 08 - 299 (MHA); Republic of Sakha / Yakutia: Khangalassky District: Lenskie Pillars National Park, Lena River downstream Labyja Creek mouth, 150 m alt., 19. VIII. 2000, Ignatov 00 - 164 (MHA); Verkhoyansk District, 30 rm W of Tabalakh Settl., Tuostakh Creek, 5. VIII. 2009, Isakova s. n. (MHA). ASIA: GEORGIA: Borzhom, pr. fl. Kura, VIII. 1877, A. H. & V. F. Brotherus s. n. (LE); Imeretia, pr. fl. Rion, 6. VI. 1877, A. H. & V. F. Brotherus s. n. (LE); Ossetia, Balta, ad. fl. Terek, 19. V. 1881, A. H. & V. F. Brotherus s. n. (LE). AZERBAIJAN: Zakatalsky District, Kebeloba Settl., 14. VI. 1969, Lyubarskaya s. n. (LE); Lenkoran, Bilyasar Mountains, 28. V. 1937, Topachevsky s. n. (LE). MONGOLIA: Khentii Province, 33 km NW of Dadal, Bukukun River valley, 4. VIII. 2006, Afonina s. n. (LE); Bayan-Hongor Province, Ikh-Bogd Mt., 2000 m alt., 29. VI. 2001, Ignatov 01 - 479 (MHA); Omnogovi Province, Khuren khanyn khep mountain ridge, 2000 m alt., 25. VI. 2001, Ignatov 01 - 456 (MHA). CHINA: Inner Mongolia, NE of Huhhot, Yingshan Mountains, Manhan Mt., 1650 m alt., 24. X. 2008, Ignatov 08 - 511 (MHA). EUROPE: AUSTRIA: Carinthia: Drautal, Danielsberg, ca. 900 m alt., 14. VI. 2003, Köckinger 15056 (MW, ex Hb. H. Köckinger); Lavanttal, Waldensteiner Graben E of Twimberg, 700 m alt., 12. VI. 2004, Köckinger 15057 (MW, ex Hb. H. Köckinger); SWITZERLAND: Ticino, Cugnasco-Gerra, Sciarana, 350 m alt., 9. IV. 2005, Schnyder 2005070 (ex Herbarium Norbert Schnyder). HUNGARY: Ktyptogamae exsiccatae # 1892, col. Lojka (LE). ITALY: Südtirol, Meran, no date, Zickendrath s. n. (LE). NORTH AMERICA: U. S. A.: Ohio: Adams Co., Spring Glen National Area, 215 m alt., 22. V. 2006, Buck 50414 (NY); Oklahoma: Payne Co., Stillwater Creek, 22. III. 1942, Ikenberry 829 (LE); Missouri: Shannon Co., Jam-Up Creek, 28. XI. 1961, Redfearn 9484 (LE).	en	Ignatova, Elena A., Kuznetsova, Oxana I., Milyutina, Irina A., Fedosov, Vladimir E., Ignatov, Michael S. (2017): The genus Fabronia (Fabroniaceae, Bryophyta) in Russia. Arctoa 26 (1): 11-34, DOI: 10.15298/arctoa.26.02, URL: https://doi.org/10.15298/arctoa.26.02
03BDA235A4776F67599DA71A89E1FD6B.taxon	discussion	Differentiation. Distinctive characters of F. ciliaris include ovate, abruptly acuminate leaves, regularly denticulate margins with small marginal teeth, short laminal cells 20 – 45 x 18 – 11 µm with length / width ratio 2 – 4: 1 and short, ovate capsules. Its difference from F. altaica, F. major, F. pusilla and F. rostrata are discussed in comments to them.	en	Ignatova, Elena A., Kuznetsova, Oxana I., Milyutina, Irina A., Fedosov, Vladimir E., Ignatov, Michael S. (2017): The genus Fabronia (Fabroniaceae, Bryophyta) in Russia. Arctoa 26 (1): 11-34, DOI: 10.15298/arctoa.26.02, URL: https://doi.org/10.15298/arctoa.26.02
03BDA235A4686F665A3DA64A8E1FFC72.taxon	materials_examined	Lectotype (selected here): specimen from herbarium of G. Hampe, BM 001007509 (Fig. 14).	en	Ignatova, Elena A., Kuznetsova, Oxana I., Milyutina, Irina A., Fedosov, Vladimir E., Ignatov, Michael S. (2017): The genus Fabronia (Fabroniaceae, Bryophyta) in Russia. Arctoa 26 (1): 11-34, DOI: 10.15298/arctoa.26.02, URL: https://doi.org/10.15298/arctoa.26.02
03BDA235A4686F665A3DA64A8E1FFC72.taxon	discussion	Comments on the lectotypification: there are two specimens of Fabronia major in BM, which obviously belong to the same species (Fig. 14). Collection from the Hampe herbarium is preferred by the following reasons, suggested by Len Ellis: (1) it has several sporophytes (in Schimper’ herbarium only one or two); (2) specimen from Hampe’ herbarium is generally in a better physical condition; (3) specimen from Schimper’ herbarium includes fragments of a different pleurocarp with leaves with conspicous single costa (superficially similar to Lescurea saxicola), while the specimen from Hampe herbarium has no admixtures.	en	Ignatova, Elena A., Kuznetsova, Oxana I., Milyutina, Irina A., Fedosov, Vladimir E., Ignatov, Michael S. (2017): The genus Fabronia (Fabroniaceae, Bryophyta) in Russia. Arctoa 26 (1): 11-34, DOI: 10.15298/arctoa.26.02, URL: https://doi.org/10.15298/arctoa.26.02
03BDA235A4686F665A3DA64A8E1FFC72.taxon	description	Illustrations: Figs. 13 – 14, 5: 12 – 16, 6: 11 – 14, 8: 13 – 15 and 11: 8. Plants small, yellow-green, green or grayish-green, glossy, in soft, dense patches. Stems creeping or ascending, 5 – 10 mm long, irregularly branching; branches 2 – 4 mm long, straight or arcuate, densely foliate. Stem and branch leaves similar, loosely appressed when dry, straight, occasionally secund, erect-spreading when moist, lanceolate, gradually long acuminate, 0.6 – 1.0 (– 1.3) u 0.2 – 0.3 (– 0.4) mm; leaf margins dentate or ciliate, marginal teeth mainly unicellular (occasionally with additional short cell at base, long (25 – 35 µm, occasionally to 50 µm long); costa slender, smooth, ending at midleaf; lamina smooth, upper and median laminal cells elongate-rhomboidal, (36 –) 50 – 70 (– 85) u 8 – 11 µm, with length / width ratio 4 – 8: 1 and cell length, µm / leaf length, mm ratio 42.8 – 91.2; apical leaf cell to 175 µm long; alar cells quadrate to short-rectangular, forming weakly delimited rectangular goup 3 – 4 cells wide and 8 – 13 cells along leaf margin. Autoicous, sporophytes infrequent. Perigonia bud-like. Perichaetial leaves 0.5 – 0.7 mm long and ca. 0.2 mm wide, with oblong base and gradually attenuate narrow acumen; costa ending above midleaf; margins with stronger dentate in upper part than stem leaves. Sporophytes single in perichaetium. Seta 3 – 4 mm long, straight, flexuose when dry, yellow. Capsule short cylindrical, with short neck, 0.7 – 0.8 mm long and 0.4 – 0.5 mm wide, occasionally flared at mouth with age, smooth, light brown, with dark-brown rim; annulus absent. Peristome single, consisting of 16 exostome teeth fused in pair, dark-brown, obtuse, ca. 125 µm long, with few round perforations in distal part; outer surface papillose-striolate, inner surface with less prominent ornamentation, with low vertical ridges. Spores 14 – 17 µm, brownish, papillose. Operculum low conic or almost flat, with very short, narrow, straight beak.	en	Ignatova, Elena A., Kuznetsova, Oxana I., Milyutina, Irina A., Fedosov, Vladimir E., Ignatov, Michael S. (2017): The genus Fabronia (Fabroniaceae, Bryophyta) in Russia. Arctoa 26 (1): 11-34, DOI: 10.15298/arctoa.26.02, URL: https://doi.org/10.15298/arctoa.26.02
03BDA235A4686F665A3DA64A8E1FFC72.taxon	materials_examined	Specimens examined: RUSSIA: Dagestan Republic: Gunib District, Gunib Settlement surroundings: 1370 m alt., 19. V. 2009, Ignatov & Ignatova 09 - 254 (MHA); 1600 m alt., 21. V. 2009, Ignatov & Ukrainskaya 09 - 472 (MHA); Republic of Severnaya Ossetia / Alania: North Ossetian Nature Reserve, Tsei River valley, 25. VII. 1979, L. I. Abramova s. n. (MW); Fiagdon Gorge, 1275 m alt., 13. VII. 2015, Ukrainskaya 16660 (LE); Nizhny Lars, 800 m alt., 1. VII. 2014, Ukrainskaya 15938 (LE). AUSTRIA: Styria: Stadl an der Mur, 1100 m alt., 11. X. 1993, Köckinger s. n. (MW, ex Hb. H. Köckinger); bei St. Michael, 600 m alt., 7. VII. 1884, Breidler s. n. (LE); Upper Austria: Donautal, Schlögen, 350 m alt., 2. X. 2013, Köckinger 14991 (MW, ex Herb. H. Köckinger); Carinthia, Lavant valley, between Twimberg and Waldenstein, 650 m alt., 18. IV. 2001, Köckinger 14992 (MW, ex Hb. H. Köckinger). SWITZERLAND: Zürich, 418 m alt., 29. III. 2015, Kiebacher 781 (MW, ex Herb. T. Kiebacher); Ticino: Muralto, am Lago Maggiore, 196 m alt., 11. X. 2002, Hofmann s. n. (MW, ex Herb. H. Hofmann); Vogomo, 510 m alt, 7. VII. 2001, Hofmann s. n. (MW, ex Herb. H. Hofmann); Monte Carasso, Pedemonte, 395 m alt., 9. IV. 2005, Schnyder 2005052 (MW, ex Herb. Norbert Schnyder); Grisons, Bregaglia, 890 m alt., 12. X. 2016, Hofmann s. n. (MW, ex Herb. H. Hofmann). ITALY: Piedmont, Lepontine-Verbano Alps, Cicogna, 740 – 750 m alt., 24. VII. 2012, Kučera 15135 (CBFS). Differentiation. Fabronia major was taken for both F. pusilla and F. ciliaris, since it considered as a variety of the former and a synonym of the latter. It resembles F. pusilla in long marginal teeth which, however, are unicellular, like in F. ciliaris. At the same time, it can be readily distinguished from both these species by narrow, lanceolate vs. ovate leaves and much longer, elongate-rhomboidal laminal cells with length / width ratio 4 – 8: 1 vs. 2 – 4: 1. It resembles F. altaica in leaf and capsule shape, but that species has slightly shorter cells, 4 – 6: 1, and shorter marginal teeth. There is some similarity in leaf shape, cell length and strong serration of leaf margins between F. major and South and Central American F. macroblepharis Schwägr., but the latter species has even longer marginal teeth which are often 2 – 3 - celled.	en	Ignatova, Elena A., Kuznetsova, Oxana I., Milyutina, Irina A., Fedosov, Vladimir E., Ignatov, Michael S. (2017): The genus Fabronia (Fabroniaceae, Bryophyta) in Russia. Arctoa 26 (1): 11-34, DOI: 10.15298/arctoa.26.02, URL: https://doi.org/10.15298/arctoa.26.02
03BDA235A46C6F6659C3A3D68F80F839.taxon	materials_examined	Type. Europe: Toscana, Raddi s. n.	en	Ignatova, Elena A., Kuznetsova, Oxana I., Milyutina, Irina A., Fedosov, Vladimir E., Ignatov, Michael S. (2017): The genus Fabronia (Fabroniaceae, Bryophyta) in Russia. Arctoa 26 (1): 11-34, DOI: 10.15298/arctoa.26.02, URL: https://doi.org/10.15298/arctoa.26.02
03BDA235A46C6F6659C3A3D68F80F839.taxon	description	Illustrations: Figs. 9: 1 – 6, 10: 1 – 5 & 11: 11 – 13; see also Ignatov & Ignatova (2004), Fig. 430 on page 648. Plants very small, soft, yellowish green, glossy, in small dense tufts. Stems creeping, ascending at apex, irregularly branched; branches erect, 1 – 2 mm long, densely foliate. Leaves loosely appressed when dry, often with recurved piliferous acumen, occasionally slightly secund, spreading when moist, ovate, abruptly long acuminate, 0.50 – 0.80 u 0.15 – 0.35 mm; leaf margins coarsely dentate to ciliate, marginal teeth unequal, large 3 – 5 - celled teeth alter with small unicellular teeth; costa slender, smooth, ending at mid-leaf; lamina smooth, upper and median laminal cells rhomboidal, 25 – 30 (– 45) u 9 – 11 µm, with length / width ratio 3 – 4: 1 and cell length, µm / leaf length, mm ratio 31.8 – 49.6; apical leaf cell to 110 µm long; alar cells quadrate to short-rectangular, forming weakly delimited rectangular goup 3 – 4 cells wide and 6 – 11 cells along leaf margin. Autoicous, sporophytes infrequent. Perigonia bud-like. Perichaetial leaves ca. 0.5 mm long and 0.2 mm wide, with oblong base and gradually attenuate narrow acumen; costa ending at midleaf; margins moderately dentate in upper part. Sporophytes single in perichaetium. Seta 2 – 3 mm long, straight, yellow. Capsule ovate, with short neck, 0.4 – 0.6 mm long and 0.3 – 0.4 mm wide, smooth, light brown, concolorous at rim; annulus absent. Peristome single, consisting of 16 exostome teeth fused in pair, light brown, obtuse, ca. 75 µm long; outer surface papillose-striolate, inner surface with less prominent ornamentation, with low vertical ridges. Spores 12 – 14 µm, brownish, papillose. Operculum low conic, with short oblique beak or mammillate.	en	Ignatova, Elena A., Kuznetsova, Oxana I., Milyutina, Irina A., Fedosov, Vladimir E., Ignatov, Michael S. (2017): The genus Fabronia (Fabroniaceae, Bryophyta) in Russia. Arctoa 26 (1): 11-34, DOI: 10.15298/arctoa.26.02, URL: https://doi.org/10.15298/arctoa.26.02
03BDA235A46C6F6659C3A3D68F80F839.taxon	materials_examined	Specimens examined: RUSSIA: EUROPEAN RUSSIA: Rostov Province: Rostov-na-Donu, Kamenka Settlement, park of the health resort “ Rostovsky ”, 16. X. 2000, Sereda s. n. (MHA); ASIAN RUSSIA: Republic of Buryatia: NE shore of Baikal Lake, Svyatoi Nos peninsula, 450 m alt, 26. VI. 1956, Bardunov s. n. (LE). EUROPE: SPAIN: Granada, Sierra Nevada, Güejar Sierra, Vereda de la Estrella, 145 m alt., 21. V. 2009, Guerra 30176 (LE); Andalucia, Grazalema, Sierra del Endrinal, 350 m alt., 3. V. 2015, Kučera 17500 (CBFS). ITALY: South Tyrol: Etschtal, Castel Feder S of Auer, 350 m alt., 29. X. 1989 Köckinger s. n. (MW, ex Herb H. Köckinger); prope Bozen, 260 m alt., Sauter, Flora Exsiccata Austro-Hungarica # 717 (LE). SWITZERLAND: Ticino, Lugano, 280 m alt., 11. II. 2003, Schnyder 2003001 (MW, ex Herb. Norbert Schnyder). AUSTRIA: Styria orientalis, Hartberg, 450 m alt., X. 1942, Baumgartner, Crypt. Exs. ed. a Museo Hist Natur. Vindobonensi # 892 (LE). ASIA: TADJIKISTAN: Darvaz Ridge, Pyandzh River basin, Egit Village, Surkh-Dara gorge, 1130 m alt., 9. VII. 1964, Mamatkulov 5218 (LE). KYRGYZSTAN: Kyrgyz Range, Dzhirly-Kaindy stow, 1900 m alt., 18. VI. 1965, Rakhmatullina s. n. (LE). NORTH AMERICA: U. S. A.: Colorado, Boulder Co., Nordseite des Boulder Canyon, 2150 m alt., 30. III. 1957, Weber, Crypt. Exs. ed. a Museo Hist Natur. Vindobonensi # 4476 (LE); California: Mariposa Co., Elephant Rock, ca. 975 m alt., 2. XII. 2006, Shevock 29285 (NY); San Benito Co., ca. 470 m alt., 12. II. 2005, Shevock 26293 (NY); Monterey Co., San Antonio Reservoir County Recreation Area, ca. 240 m alt., 3. I. 2009, Schevock 32581 (NY); Oregon: Wheeler Co., John Day River basin, 655 m alt., 23. III. 2008, Shevock 31600 (NY); Wasco Co., Colombia River Gorge National Scenic Area, 45 m alt., 27. III. 2008, Shevock 31659 (NY).	en	Ignatova, Elena A., Kuznetsova, Oxana I., Milyutina, Irina A., Fedosov, Vladimir E., Ignatov, Michael S. (2017): The genus Fabronia (Fabroniaceae, Bryophyta) in Russia. Arctoa 26 (1): 11-34, DOI: 10.15298/arctoa.26.02, URL: https://doi.org/10.15298/arctoa.26.02
03BDA235A46C6F6659C3A3D68F80F839.taxon	discussion	Differentiation. ‘ Typical’ specimens of Fabronia pusilla are easily recognized due to large, multicellular teeth at leaf margins. However, some specimens with smaller, 2 – 3 - celled teeth may be referred either to F. pusilla or F. ciliaris (as it is proved by DNA data) and cannot be identified with confidence. Such specimens were observed mainly in Central Europe where both species co-occur. Maybe slightly smaller spores of F. pusilla, 12 – 14 vs. 15 – 17 µm in F. ciliaris, can help in these difficult cases.	en	Ignatova, Elena A., Kuznetsova, Oxana I., Milyutina, Irina A., Fedosov, Vladimir E., Ignatov, Michael S. (2017): The genus Fabronia (Fabroniaceae, Bryophyta) in Russia. Arctoa 26 (1): 11-34, DOI: 10.15298/arctoa.26.02, URL: https://doi.org/10.15298/arctoa.26.02
03BDA235A46D6F67599DA0B08E45F8C6.taxon	materials_examined	Type. China: Yunnan, Mekong, 28 ° 11 ’ N, Handel-Mazzetti 8018 (holotype H).	en	Ignatova, Elena A., Kuznetsova, Oxana I., Milyutina, Irina A., Fedosov, Vladimir E., Ignatov, Michael S. (2017): The genus Fabronia (Fabroniaceae, Bryophyta) in Russia. Arctoa 26 (1): 11-34, DOI: 10.15298/arctoa.26.02, URL: https://doi.org/10.15298/arctoa.26.02
03BDA235A46D6F67599DA0B08E45F8C6.taxon	description	Illustrations: Figs. 9: 11; 10: 9 and 11: 20; see also Dudov et al., 2015. Plants small, in flat, loose or moderately compact mats, light green or grayish, glossy. Stems 10 – 15 cm, prostrate, irregularly branched, evenly foliate. Leaves appressed and imbricate when dry, erectopatent to spreading when wet, 0.6 – 1.0 (– 1.2) u 0.4 – 0.5 mm, ovate-lanceolate, gradually narrowed into a long and narrow acumen, slightly concave; margins plane, finely serrulate; costa slender, reaching 0.4 – 0.6 of leaf length; laminal cells rhomboidal, thin-walled, smooth, 30 – 60 u 6 – 10 µm, with length / width ratio 2 – 4: 1 and cell length, µm / leaf length, mm ratio 19.5 – 33.7; apical leaf cell to 110 µm long; alar cells differentiated, quadrate, in 3 – 4 rows at margin. Autoicous. Seta 2 – 3 mm. Capsule ovoid, 0.6 – 0.9 mm long. Operculum almost flat, with rather long oblique beak. Peristome totally reduced. Spores 10 – 15 µm.	en	Ignatova, Elena A., Kuznetsova, Oxana I., Milyutina, Irina A., Fedosov, Vladimir E., Ignatov, Michael S. (2017): The genus Fabronia (Fabroniaceae, Bryophyta) in Russia. Arctoa 26 (1): 11-34, DOI: 10.15298/arctoa.26.02, URL: https://doi.org/10.15298/arctoa.26.02
03BDA235A46D6F67599DA0B08E45F8C6.taxon	materials_examined	Specimens examined: RUSSIA: Zabaikalsky Territory, Nerchinsko-Zavodskiy District, near Nerchinsky Zavod settlement, 658 m alt., 25. VII. 2012, Afonina 3912 (LE); Amurskaya Province: Zeisky Disctrict, Zeya Nature Reserve, Tukhuringra range: Gilyiskiy bay of Zeya reservoir, right shore, at 0.3 km to W from cordon “ Medvezhiy ” 628 m alt., 2. VIII. 2013, Dudov & Kotel’nikova # 2013 _ Br _ 0021; Izvestkoviy bay of Zeya reservoir, right shore near Izvestkoviy stream estruary 421 m alt., 4. VIII. 2013. Dudov & Kotel’nikova 2013 _ Br _ 0336 (MW).	en	Ignatova, Elena A., Kuznetsova, Oxana I., Milyutina, Irina A., Fedosov, Vladimir E., Ignatov, Michael S. (2017): The genus Fabronia (Fabroniaceae, Bryophyta) in Russia. Arctoa 26 (1): 11-34, DOI: 10.15298/arctoa.26.02, URL: https://doi.org/10.15298/arctoa.26.02
03BDA235A46D6F67599DA0B08E45F8C6.taxon	discussion	Differentiation. In the description of this species V. F. Brotherus (1929) noted such characteristic features of the species as finely serrulate leaf margins and operculum with rather long oblique beak. These characters, as well as the absence of peristome, allow its separation from both F. ciliata and F. pusilla which possess much stronger serrate leaf margins (especially the latter species), mammillate operculum and well-developed, albeit fragile single peristome of 16 teeth, strongly reflexed in dry state and appressed to the urn of opened capsules. Even if partly brocken, teeth remains are apparent enough to indicate peristome presence in these species.	en	Ignatova, Elena A., Kuznetsova, Oxana I., Milyutina, Irina A., Fedosov, Vladimir E., Ignatov, Michael S. (2017): The genus Fabronia (Fabroniaceae, Bryophyta) in Russia. Arctoa 26 (1): 11-34, DOI: 10.15298/arctoa.26.02, URL: https://doi.org/10.15298/arctoa.26.02
