taxonID	type	description	language	source
0B2E630B885AC07E8AE00636FB34F91C.taxon	type_taxon	Type species: Pintalia lateralis Stål, 1862 by original designation.	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B885AC07E8AE00636FB34F91C.taxon	diagnosis	Amended diagnosis. (Adapted after Muir 1934, Kramer 1983.) Vertex trapezoidal (Fig. 1), with two compartments (apical and basal) and two carinae (apical transverse and subapical carina), apical transverse carina usually visible in dorsal view; basal compartment larger than apical compartment. Frons (Fig. 2 A), with median carina present. Rostrum reaching or surpassing the hind coxae. Lateral ocelli, and frontal ocellus present well developed. Pronotum short (Fig. 1), in dorsal view narrower medially. Mesonotum with three carinae (Fig. 1), planate in lateral view, usually distally inclined in lateral view. Tegmina (Fig. 12) tectiform, in resting position usually surpassing approx. 1 / 3 of body length with distal margins usually touching each other, and apical cells and veins usually elongated and parallel (between RA and icu veins), usually with 12 – 13 distal veins, with RP quadrifid, sometimes with punctual anastomosis between RA + RP 1; rm- 1 and mcu- 1 usually near the first MP fork; fork of MP 1 + 2 usually trifid and MP 3 + 4 usually bifid; ScP + RA and RP fork starting more proximally to the basal cell, than the fork in Cua 1 and Cua 2 that occurs more distally, in some species they occur at the same level; C 1 and C 5 cells usually more elongated; C 1 cell always larger than C 5 cell. Hind wing (Fig. 12 B – D) with RP bifid, MP usually trifid (petiolate anastomosis in MP 3 + 4 + CuA 1 distally forked – Y type) or bifid (complete anastomosis with distal fusion MP 3 + 4 + CuA 1 – I type). Posterior legs, hind tibia in most species with lateral spines, in some without any (see genus description below), usually with 6 apical teeth-grouped as follows (Fig. 3 B): 1 larger lateral (distal), 2 smaller together (medial), 1 larger (medial), and 2 (medium size) (internal).	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B885AC07E8AE00636FB34F91C.taxon	description	Morphology. Body length: ♀ 2.8 – 6.0 mm (n = 40). ♂ 2.6 – 5.3 mm (n = 42). Head (Figure 1). Vertex trapezoidal, apical transverse carina and subapical carina approx. the same size usually concave and convex (P. lateralis Stål, 1862) or straight, almost straight, straight bent forward; apical transverse carina distinct in some species, elongated medially and touching the median carina of the frons (Y-shape, in frontal view uniting forward of the eyes) (P. ustulata Stål, 1862; P. inornata Stål, 1862); angle formed by the caudal margin of the vertex usually concave (e. g., P. lateralis) or triangular (e. g., P. fasciatipennis Stål, 1862); lateral carinae elevated (except P. fasciatipennis); interspecific variation observed in the basal emargination (median carina), absent or inconspicuous (P. lateralis; P. obscuripennis Stål, 1862; P. pictipennis Stål, 1862; P. fraterna Stål, 1862), weakly developed (P. inornata), or well developed (P. fasciatipennis; P. consobrina Stål, 1862; P. ustulata). Frons, longer than wide; up to 2.0 times wider medially than apically (e. g., P. lateralis); trapezoidal apically, with apical region usually above the compound eyes (except P. ustulata; P. inornata) and straight or concave apically (apical transverse carina frontally); median carina not reaching or rarely reaching the frontal ocellus (except P. consobrina), short in some species, touching to curved apical transverse carina medially (Y-shape in frontal view); lateral carinae of the frons usually elevated (except P. fasciatipennis). Antenna with short scape and semi-globular pedicel, slightly surpassing the margins of the frons when in frontal view. Thorax (Figure 1). Pronotum tricarinate, with median carina absent or inconspicuous; submedian carinae moderately developed (e. g., P. lateralis), often not continuous dorsally (weakly developed); dorsally shorter than length in the middle between the eyes (e. g., P. lateralis); caudal margin rounded or straight / obtuse (e. g., P. lateralis). Mesonotum dorsally with three carinae (e. g., P. lateralis), which are usually narrow near the pronotum and wide near the abdomen (except P. inornata); in some species region between carinae lighter than on the lateral area near the tegulae (P. lateralis and little in P. consobrina). Tegmina (Forewings) with RP vein quadrifid (e. g., P. lateralis), in some species with punctual anastomosis between RA + RP 1 (P. consobrina; P. pictipennis); tubercles occur throughout all veins of the tegmina, often more evident in some species; can be evanescent (P. lateralis; P. pictipennis; P. fraterna), double (P. consobrina), simple (P. fasciatipennis; P. obscuripennis; P. proxima Stål, 1862 and P. ustulata), or yet absent or inconspicuous (P. inornata); distal margin totally curved (except P. ustulata); Cells C 1 b, C 2, C 3 a, C 3 and C 4 (central prenodal cells), stacked together right behind the center of the tegmina (e. g., P. lateralis); C 1 and C 5 elongated (lateral prenodal cells) (e. g., P. lateralis), C 1 a little closer to bc than C 5 (except, P. ustulata), or C 5 very short (P. inornata) or approx. the same size as C 1 (P. ustulata); usually 12 apical cells (nodal) and 7 subapical cells (prenodal) (except P. fasciatipennis and P. obscuripennis). Hind wing with RP bifid (e. g., P. lateralis), MP trifid usually with petiolate anastomosis in MP 3 + 4 + CuA 1 distally forked, Y – type (P. lateralis, P. ustulata, P. fraterna) or with complete anastomosis with distal MP 3 + 4 + CuA 1 fusion, I-type (possibly P. pictipennis). Legs. Hind tibia with 6 apical teeth; usually with 1 – 7 lateral spines, tiny near base, or unarmed in some species (P. inornata; P. obscuripennis; P. ustulata). 1 st tarsomere with 7 apical teeth of approx. the same size or, with 1 – 2 lateral teeth larger and 2 apical teeth moderately large in the middle. 2 nd tarsomere with 7 – 9 apical teeth, rarely 7 and rarely 9, all with approx. the same size, or with 1 – 2 lateral larger teeth and intermediate teeth gradually smaller medially or of uniform size, usually 1 – 3 platellae (Pintalia painensis - group, Pintalia constellaris - group) or with 3 very thin setae (Pintalia pictipennis - group). Male terminalia. Pygofer with or without processes between dorsal and distal margins, asymmetric in some species; ventromedian process triangular, crooked in some species (e. g., P. magnaepiprocti sp. nov.). Anal tube (Fig. 14), longer than wider, in some species distally developed, bent approx. at between 45 ° – 90 ° with epiproct and paraproct small (Pintalia painensis - group); or elongated but not developed distally, with epiproct and paraproct small (Pintalia constellaris - group); or moderately short and wide not developed distally, with paraproct large developed distally, and in some species with epiproct large developed distally (Pintalia pictipennis - group). Genital styles, in lateral view long and well expanded apically (Pintalia pictipennis - group, Pintalia constellaris - group), or long with the apex slightly or moderately expanded (Pintalia painensis - group); in ventral view, touching medially and forming the basal opening, moderately large, laterally concave, and apically triangular (Pintalia pictipennisgroup) or small and diamond-shaped (Pintalia constellaris - group and Pintalia painensis - group). Aedeagus with shaft tubular, ventral ridge developed and flagellum without spines (Pintalia pictipennis - group), or tubular usually with three conspicuous spines at the apex of the aedeagal shaft, without ventral ridge and flagellum spineless (Pintalia constellaris- group); or tubular and wider apically near to flagellum, ventral ridge absent, usually flagellum with spines and phallotreme laterally (Pintalia painensis - group). Female terminalia. Anal tube not developed distally (bent) ending together with epiproct and paraproct; usually elongated, tubular, or cylindrical dorsally with lateral margins moderately developed (P. lateralis; Pintalia painensis - group), or slightly longer than wide and flattened ventrally (P. obscuripennis; P. consobrina; P. ustulata) and in parts Pintalia pictipennis - group, present only P. serratilis sp. nov. and P. stali sp. nov. Epiproct and paraproct flattened, and small; epiproct triangular in dorsal view, with two rounded margins bent down laterally; paraproct rounded in dorsal view, sometimes weakly elongated. Gonocoxae VIII truncated, with a rounded laterally, in caudal view usually small and ovoid. Ovipositor sabre-shaped, in lateral view curved dorsally in some specimens surpassing anal tube.	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B885AC07E8AE00636FB34F91C.taxon	distribution	Distribution. Neotropical region. South America, Brazil (Stål 1862; Muir 1934), Bolivia (Fennah 1945), Chile (Coquimbo) (Campodonico 2021), Ecuador (Muir 1934; Fennah 1945), Guiana Francesca (Fennah 1945), Guyana (Muir 1934; Caldwell 1944), Peru (Fennah 1945), Trinidad and Tobago (Fennah 1945), Venezuela (Lethierry 1890). North America, United States of America (Kramer 1983; Bartlett et al. 2014; Van Duzee 1908; Caldwell 1944; Kramer 1983; Caldwell and Martorell 1951), Cuba (Bartlett et al. 2014), Guadeloupe (Fabricius 1803; Stål 1862), Cayman Islands (Fennah 1971), Martinique (Fabricius 1803; Stål 1862), Mexico (Caldwell 1944; Bartlett et al. 2014). Central America, Costa Rica (Caldwell 1944; Kunz and Holzinger 2022), Dominica (Fennah 1948), Granada (Fennah 1948), Guatemala (Caldwell 1944; Bartlett et al. 2014), Haiti (Uhler 1901), Nicaragua (Bartlett et al. 2014), Panama (Metcalf 1938), Saint Lucia (Fennah 1948), Saint Vincent and the Grenadines (Uhler 1895; Fennah 1948).	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B885AC07E8AE00636FB34F91C.taxon	discussion	Remarks. Pintalia is a large genus with very varied morphology and lacks an in-depth review of the species that make up the genus. All species here treated, including P. lateralis, have the vertex divided transversely into anterior and posterior compartments by the subapical carina. However, in P. inornata and P. ustulata the apical transverse carina is medially elongated and, as a result, it exhibits a larger apical compartment than in other species. Most species present the vertex with elevated carinae in dorsal view; however, in P. consobrina Stål, 1862 the subapical carina is not foliate, and in P. fasciatipennis Stål, 1862 neither of the two carinae (apical transverse and subapical) are elevated, which is uncommon for the genus. Furthermore, P. inornata, P. consobrina, P. fasciatipennis and P. ustulata are the only studied species with developed median carina in the basal compartment of the vertex (very distinct). In the three species-groups herein proposed, the transverse apical carina can be concave in frontal view as in Pintalia pictipennis- group, or straight as in the Pintalia painensis- group and the Pintalia constellaris- group. In dorsal view, it may be the same size or larger than the subapical carina as observed for the Pintalia pictipennisgroup or of the same size or smaller than the subapical carina as in the Pintalia painensis- group and Pintalia constellaris- group. In frontal view, there appear to be two main frontoclypeal suture patterns. Some species exhibit a semicircular frontoclypeal suture bent upwards as in P. lateralis and the Pintalia pictipennis- group; however, in other species, it is angularly bent upwards as in P. fraterna and Pintalia painensis- group. We did not find any significant variations in the thorax (pronotum and mesonotum) among the species reported here, except for P. inornata, which dorsally exhibits almost straight and parallel carinae on mesonotum vs. narrow near the pronotum and wider distally, which is observed in most species (e. g. P. lateralis). All species reported here have lateral spines on the posterior tibia, except for P. inornata; P. obscuripennis and P. ustulata (very distinct). Although this appears to be a convenient character, we were not successful in associating this feature with other features relevant to species groupings. Furthermore, it was not possible to accurately assess the apical teeth of the hind tibia of the species described by Stål, 1862. On the other hand, we observed thin setae (Pintalia pictipennisgroup) and platellae (Pintalia painensis- group and Pintalia constellaris- group) on the new species herein described, and this character is normally used for subgeneric-grouping in Cixiidae. However, it can be in some species absent or in a smaller number on one of the legs. Although the tegmina can present different anastomosis and crossveins in some species, it usually exhibits stable cell patterns, with 12 apical (nodal) cells and 7 subapical (prenodal) cells (Fig. 12 – 13). However, in some species, there may be an asymmetry of cells and veins between the right and left tegmina. For most species, the apical margin is smoothly curved between RA and icu, except in P. ustulata. Here, the Pintalia pictipennis - group exhibits punctual anastomosis in RA + RP 1 veins, vs. separate by crossvein in Pintalia constellaris - group and Pintalia painensis - group). The shape of the basal margin of the pterostigma can also be useful for identification, since in the Pintalia pictipennis- group it is small and distally triangular, while in the Pintalia constellaris- group it is large and sub-rectangular weakly angled distally, and in the Pintalia painensis- group it is large and sub-rectangular, weakly, or well angled distally. Although there are conspicuous spotting patterns in some species and - groups, some patterns can be observed in species of the three groups herein proposed, such as an inclined transverse spot in the center of the tegmina (most species), two semicircular spots in pcc (most species), and another three semicircular or rounded spots between ScP + RA and A 1. It was only possible to examine the posterior wings of P. lateralis (type), P. ustulata and P. fraterna. These species have a petiolate anastomosis in MP 3 + CuA 1 distally forked as Pintalia painensis- group (Fig. 12 B) and Pintalia constellaris- group (Fig. 12 C) (I-Type). However, Pintalia pictipennis- group exhibits complete anastomosis with distal MP 3 + 4 + CuA 1 fusion (Fig. 12 D) (Y-type) (Emeljanov 2002, Le Cesne 2022).	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B885AC07E8AE00636FB34F91C.taxon	diagnosis	The groups that we propose were based on the morphology of the new species presented. Although they were useful, we did not use the groups formed by Fennah (1945), as the species treated by him could not be fully evaluated before the construction of this work. Phylogenetic analyses will be necessary to identify whether the synapomorphies we indicate are useful for the proposal of subgenera or new genera in Pintalia. Here, we suggest the main characters for the proposal of new species and consequent allocation in groups of Pintalia (Table 1): • Vertex with two transverse carinae (apical and subapical) and two compartments (apical and basal) present (1) or absent (0) (AP). • Vertex with apical transverse carina and subapical carina elevated (2), not elevated (1), or only apical transverse carinae elevated (0) (CV). • Apical transverse carina in frontal view concave medially (2), straight (1), medially elongated (0) (AT). • Median carina of the basal compartment developed (1) or undeveloped (0) (MC). • Frons with lateral carinae elevated (1) or no elevated (0) (FR). • Frontoclypeal suture semicircular bent forwards (1) or straight bent forwards (0) (FS). • Median carina touching the frontal ocellus (1) or evanescent near the frontal ocellus (0) (FM). • Tegmina with RA + RP 1 with punctual anastomosis (1) or anastomosis absent (0) (TR). • ScP + R well forward of fork CuA (3), slightly (2), in the same level (1), behind (0) (TV). • Hind wing with a petiolate anastomosis in MP 3 + 4 + CuA 1 distally forked, Y-type (1) or with complete anastomosis with distal fusion between MP 3 + 4 + CuA 1, I-type (0) (WA). • Females with anal tube long (1) or short (0) (FA). • Hind tibia with thin setae (1) or platellae (0) on the 2 nd, tarsomere (PS). • Lateral spines in hind tibia present (1) or absent (0) (LS). • Anal tube distally developed (1), undeveloped (0) (AS). • Male anal tube in dorsal view with narrowing (lesser width) just near the base (1) or not narrowing (0) (AD). • Paraproct distally developed (1), undeveloped (0) (PP). • Epiproct distally developed (1), undeveloped (0) (EP). • Ridge on the aedeagal shaft present (1), absent (0) (VR). • Flagellum spine-like (1) or flagellum without spines (0) (FS). • Pygofer with medioventral process cone shaped laterally, wide, in caudal view (1) or without cone-shaped process laterally (0) (PY). • Shape and size of the genital styles, well large (1), or moderately large (0) (GS). • Shape and size of the basal opening of the genital styles, larger and laterally concave (1) or small in a diamond shape (0) (OG). • Shape of the aedeagal shaft, tubular (1) tubular and apically larger (0) (AF). • Pregenital segment evanescent in the middle (2), moderately evanescent (1), not evanescent (0) (PG).	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B885AC07E8AE00636FB34F91C.taxon	discussion	Remarks. This key does not include the species described by Stål (1862) and Muir (1934) as it was not possible to evaluate the aedeagus of these species, and this structure is essential for species-level recognition in the genus Pintalia. Species such as P. pictipennis and P. stali sp. nov., P. constellaris and P. speciosa sp. nov., and all species of the Pintalia painensis - group highlight the need to use genitalia for identification at the species level, until it is possible to better understand what an intraspecific or interspecific variation is in the genus. With this publication we hope to find male specimens of the species described by Stål (1862) to add them to the key. The supplementary description of the species described by Muir (1934) may be useful to clarify the morphology of the male terminalia and thus also add them to the key. However, it has not yet been possible to evaluate all holotypes and paratypes deposited at BPBM and BMNH respectively.	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B8854C07C8AE003FDFDFFFD2C.taxon	diagnosis	Diagnosis. This group can be distinguished mainly by characters of the male terminalia; however, some other characters are also useful for identification. Vertex small, with transverse apical carina slightly larger than the subapical carina; apical compartment very small; caudal margin well triangular or concave in some species; basal compartment with small basal emargination; in frontal view usually exhibits apical transverse carina (anterior region of the frons apically) concave, rarely almost straight. Frons approx. 1.5 – 2.3 times wider in the largest width than at the apex; frontoclypeal suture usually semicircular convex, slightly straight / inclined laterally. Tegmina (forewings) with pt small and distally triangular; RA bifid with punctual anastomosis RA + RP 1; fork of ScP + RA and RP slightly ahead or on the same level of fork CuA 1 and CuA 2; rm- 1 and mcu- 1 usually together, in some species proximally, very close or distally to the first MP fork; in some species crossveins in greater number in one of the tegmina (asymmetry). Hind wing with RP bifid, MP trifid with complete anastomosis in distal fusion of MP 3 + 4 + CuA 1 (I-type). In ventral view, males exhibit segment viii separated medially. Posterior legs, hind tibia with 3 – 7 lateral spines, usually with three spines larger, spines closest to the femur well small; 6 apical teeth as provide for the genus; 1 st 1 st tarsomere with 7 apical teeth; 2 nd tarsomere with 8 – 9 apical teeth and 3 very fine setae. Male terminalia. Pygofer bilaterally asymmetrical in some species; in lateral view dorsocaudal margin totally convex, without processes medially; in ventral view, ventromedian process triangular usually as wide as long. Anal tube short and wide, in some species larger and conspicuous distally; paraproct well developed distally, usually paraproct more than 2.0 times larger than epiproct; in dorsal view epiproct larger medially, developed distally in some species. Genital styles Genital styles in lateral view, thin at the base and very wide apically, approx. 3.0 times wider apically than basally; excavated ventrally, with ventral margins bent to the ventral region; in ventral view, basal opening between the genital styles large, laterally concave and apically triangular. Aedeagus usually with aedeagal shaft wide or slightly wider at base than apically; with 2 – 3 spines apically, in most species with one right lateral spine bifid; ventral ridge developed. Flagellum without spines, rarely exhibit a spine-like process apically. Female terminalia. Anal tube short, in dorsal view usually slightly longer than wide, flattened near base with apex truncated or rounded; in lateral view with lateral margin slightly projected and straight, usually flattened ventrally or slightly rounded; epiproct and paraproct thin and short. Gonocoxae VIII truncate, in caudal view rounded, in some species smaller apically, in lateral view usually with lateral margin surpassing the middle moderately or reaching the base; in caudal view usually rounded / swelling laterally and excavated out in the middle. Ovipositor large, in lateral view, sabre-shaped, curved upwards dorsally, in some species surpassing the anal tube.	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B8854C07C8AE003FDFDFFFD2C.taxon	distribution	Distribution. BRA. Rio de Janeiro and Minas Gerais State (Fig. 77).	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B8854C07C8AE003FDFDFFFD2C.taxon	discussion	Notes. Due to the conditions of the material, it was not possible to accurately evaluate all the characters proposed in the diagnosis for the species P. propria and P. pictipennis. Although Fennah (1945) created the P. propria - group to house this unique species, to avoid future conflicts, due to the shape of the male terminalia it will be allocated in the Pintalia pictipennis - group.	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B8854C07C8AE003FDFDFFFD2C.taxon	description	Morphology. Body length: ♂ 3.5 – 5.0 mm (n = 5). ♀ 4.0 – 5.5 mm (n = 4). Head. Vertex with rectangular apical compartment, in frontal view small / narrow laterally (P. minuta sp. nov., P. minima sp. nov., P. magnaepiprocti sp. nov.), or apical compartment large, in dorsal view with forward bent carinae (P. serratilis sp. nov.) or with straight apical transverse carina and concave subapical carina (P. stali sp. nov., P. pictipennis Stål 1862). Frons width larger below the scapus, approx. 2.1 – 2.3 times wider in the largest width than at the apex (P. minuta sp. nov., P. minima sp. nov., P. magnaepiprocti sp. nov.) or on the same level as the scapus, approx. 1.5 – 1.7 times wider in the largest width than at the apex (P. serratilis sp. nov., P. stali sp. nov., P. pictipennis); anterior region of the frons usually concave apically or almost straight (P. propria). Thorax. Pronotum narrow, median carina evanescent or inconspicuous; submedian carina moderately developed, but evanescent behind the eyes. Mesonotum with distance between lateral carinae slightly larger distally than near the pronotum (P. serratilis sp. nov., P. stali sp. nov.) moderately larger (P. minuta sp. nov., P. minima sp. nov.), or considerably larger (P. magnaepiprocti sp. nov.). Tegmina (forewings) mostly hyaline with a few light gray spots, and with three moderately elongated spots occurs between RP (+ MA) and A 2. (P. minuta sp. nov., P. minima sp. nov., P. magnaepiprocti sp. nov., P. propria); dark tegmina with two conspicuous hyaline spots in Cua 1 and Cua 2 (P. serratilis sp. nov.); tegmina with prenodal region mostly hyaline, and nodal region with dark spots between the postclaval cell and radial area with 5 conspicuous longitudinal spots (P. stali sp. nov., P. pictipennis). Posterior legs. Hind tibia with three lateral spines (P. serratilis sp. nov.), four lateral spines (P. magnaepiprocti sp. nov., P. propria), six lateral spines (P. minuta sp. nov., P. stali sp. nov.) or seven lateral spines (P. minima sp. nov.). 1 st tarsomere with 7 apical teeth, approx. the same size (P. minuta sp. nov., P. minima sp. nov., P. magnaepiprocti sp. nov.) with one or the two externals slightly larger and 1 – 2 spines of the middle larger (P. serratilis sp. nov., P. stali sp. nov.). 2 nd tarsomere with 8 apical teeth (P. minuta sp. nov., P. minima sp. nov., P. magnaepiprocti sp. nov., P. stali sp. nov., P. propria) or rarely with 9 apical teeth (P. serratilis sp. nov.), and usually with 3 thin setae. Male terminalia. Pygofer in symmetry bilaterally (P. stali sp. nov.), slightly asymmetrical (P. minuta sp. nov., P. serratilis sp. nov.), or very asymmetrical (P. minima sp. nov., P. magnaepiprocti sp. nov.). Anal tube flattened, dorsally rounded and distinct distally, without processes (P. minuta sp. nov., P. minima sp. nov., P. magnaepiprocti sp. nov., P. pictipennis), or tubular, moderately tubular and distally with processes in the ventral region (P. stali sp. nov., P. serratilis sp. nov. P. propria). Genital styles, usually thin at the base and very wide apically; in lateral view approx. 3.0 times wider apically, in some species moderately wide apically and very curved distally (sub angulated) (P. propria); basal opening of the genital styles usually large (except in P. serratilis sp. nov.). Aedeagus with bifid spines in right lateral of (P. minuta sp. nov., P. minima sp. nov., P. serratilis sp. nov.) or without bifid spines (P. magnaepiprocti sp. nov., P. stali sp. nov., P. propria); ventral ridge small weakly developed (P. minuta sp. nov.), or moderately developed and curved (P. minima sp. nov., P. magnaepiprocti sp. nov.), or serrated (P. serratilis sp. nov.), armed with a pair of large spines, on each side of the base of the aedeagal shaft (P. propria) or two pairs of spines (P. stali sp. nov.). Female terminalia. Anal tube short, distally rounded and ovipositor surpassing the anal tube in lateral view (P. minuta sp. nov., P. minima sp. nov.); or tubular dorsally, apically truncated and ovipositor not extending beyond anal tube in lateral view (P. stali sp. nov., P. serratilis sp. nov.). Gonocoxae VIII short, with lateral carinae slightly developed laterally (P. minuta sp. nov., P. minima sp. nov.); or with lateral carinae moderately developed laterally (P. stali sp. nov., P. serratilis sp. nov.).	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B8856C07D8AE007D8FEEFF811.taxon	diagnosis	Diagnosis. This group can be distinguished mainly by characters of the male terminalia; however, some other characters are also useful for identification. Vertex large, with apical transverse carina smaller than the subapical carina; apical compartment wide; caudal margin concave and slightly triangular medially; basal emargination small, or inconspicuous; in frontal view usually exhibits apical transverse carina straight. Frons approx. 1.5 times wider in the largest width than at the apex; frontoclypeal suture usually semicircular convex or straight bent dorsally, flattened apically. Tegmina (forewings) with pt large and sub rectangular weakly angled distally; with C 5 ’ large and quadratic; fork of ScP + RA and RP ahead of fork CuA 1 and CuA 2; rm- 1 and mcu- 1 usually together or very close to the first MP fork, in some species mcu- 1 proximally to first MP fork. Hind wing with RP bifid, MP trifid with petiolate anastomosis in MP 3 + 4 + CuA 1 distally forked (Y-type), CuA 1 and CuA 2 bifid. In ventral view, males exhibit segment viii slightly to moderately evanescent medially. Hind tibia usually exhibits 2 – 3 lateral spines, with spine near the femur very small, 6 apical teeth as provide for the genus. 1 st tarsomere with 7 apical teeth. 2 nd tarsomere with 7 – 8 apical teeth, usually with 3 platellae, rarely without platellae. Male terminalia. Pygofer bilaterally symmetrical; in lateral view, dorsocaudal margin with sharp processes medially; dorsal margin concave, caudal margin convex; in caudal view, processes wider than the rest of the distal margins (vs. thin in Pintalia painensis- group); ventromedian process triangular. Anal tube moderately long, not developed distally (bent); in lateral view, almost straight dorsally; in ventral view concave medially with a small longitudinal ridge medially; epiproct and paraproct short, in dorsal view the epiproct triangular, with lateral margin near to the base larger and bent down; paraproct rounded, narrow laterally and slightly larger than the epiproct distally. Genital styles thin at the base and moderately wide or very wide apically; excavated ventrally latero-basally with margins curved towards the ventral region; dorsal margins exhibit a conspicuous triangular process medially; in ventral view, basal opening between the genital styles shaped like a small diamond-shaped, with rounded lateral angles. Aedeagus usually with shaft as wide as or slightly wider at base than apically; exhibit 3 – 5 spines, usually three spines (1 st, 2 nd, 3 rd) occur at the apex of the aedeagal shaft near to flagellum; without ventral ridge. Flagellum without spines, in some species with a spine-like process laterally. Female terminalia. Anal tube moderately large or well developed, in dorsal view usually swollen medially or apically, slightly rounded, with lateral margin slightly developed or moderately developed; usually flattened ventrally or slightly excavated; epiproct and paraproct thin and short as to the males. Gonocoxae VIII truncated, in lateral view usually with lateral carinae short not reaching the middle of the segment. Ovipositor large, in lateral view, sabre-shaped, curved upwards dorsally, caudally surpassing anal tube.	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B8856C07D8AE007D8FEEFF811.taxon	distribution	Distribution. NIC. BOL. CR. ECU. BRA, Minas Gerais, Mato Grosso, Bahia, Rio de Janeiro, São Paulo and Paraná state (Fig. 77).	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B8856C07D8AE007D8FEEFF811.taxon	discussion	Notes. In South America, P. speciosa sp. nov. is the second species (Pintalia constellaris (Walker, 1858) = Pintalia quadrimaculata Fennah, 1945, by Fennah, 1947) reported to share the following character states: tegmina dark yellow with one black spot with four hyaline spots near to the center of the tegmina; C 3 and C 5 cells elongated and rounded, C 4 narrow. This character is widely observed in South American species, currently with most 40 records on iNaturalist, distributed in mainly southeastern Brazil, southern Peru, Costa Rica and Ecuador. In addition, P. constellaris and P. speciosa sp. nov. exhibit the pygofer with robust process laterally, apex of the aedeagal shaft display three spines moderately large, and medially with two spines in opposite sides, as P. fennahi sp. nov. P. constellaris and P. speciosa sp. nov. show minimal differences but can be differentiated by genital styles and the size, shape, and position of the aedeagus spines, as described here in morphology. The type of P. ecuadorensis was evaluated by photographs provided by BPBM. The aedeagus of P. ecuadorensis is more distinct among the species of the group, and this type requires a supplementary description. We believe that with advances in the knowledge of planthoppers in Brazil, Pintalia constellaris - group will be a case like the Australian genus Chidaea Emeljanov, 2000, where we have several species with small variations of the same aedeagal spines (Löcker and Holzinger, 2019) (See Remarks P. speciosa sp. nov).	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B8856C07D8AE007D8FEEFF811.taxon	description	Morphology. Body length: 4.0 – 4.4 ♂ mm (n = 4). 5.0 – 6.4 mm ♀ (n = 4). Head. Vertex with apical compartment wide laterally (P. constellaris, P. ecuadorensis, P. speciosa sp. nov.), or distinct apical compartment, weakly excavated, with distinct apical transverse carina evanescent and elongated medially (P. fennahi sp. nov.); subapical carina straight or weakly straight (P. constellaris, P. ecuadorensis) or bent medially (P. speciosa sp. nov., P. fennahi sp. nov.). Frons apically swollen and curved (P. fennahi sp. nov.) or trapezoidal straight (P. constellaris, P. speciosa sp. nov., P. ecuadorensis); approx. 1.5 times wider in the largest width than at the apex; frontoclypeal suture rounded (P. speciosa sp. nov., P. ecuadorensis) or bent dorsally (P. fennahi sp. nov.) Thorax. Pronotum narrow, inconspicuous media carina and moderately developed but evanescent submedian carina behind the eyes. Mesonotum with distance between lateral carinae and slightly larger distally than near the pronotum (P. fennahi sp. nov.), or moderately larger (P. speciosa sp. nov., P. constellaris, P. ecuadorensis). Tegmina with few hyaline parts, largely dark yellow (P. constellaris, P. speciosa sp. nov.) or dark brown (P. fennahi sp. nov.) spots; C 5 cell wide near at fork cua 1 - cua 2 (P. constellaris, P. speciosa sp. nov.) short / common (P. fennahi sp. nov.), C 3 cell wide and rounded (P. speciosa sp. nov.). Posterior legs, hind tibia with 3 lateral spines; 1 st tarsomere 7 apical teeth, being 2 lateral larger and 2 teeth moderately large in the middle; 2 nd tarsomere with 7 apical teeth (P. speciosa sp. nov.) or 8 apical teeth (P. fennahi sp. nov., P. ecuadorensis) usually the two outer ones larger, and the middle ones with approx. the same size, usually with 3 platellae. Male terminalia. Pygofer bilaterally symmetric, wider apically (P. constellaris, P. speciosa sp. nov.), wider medially (P. fennahi sp. nov.) or wider ventrally (P. ecuadorensis), cone-shaped process rounded apically (P. speciosa sp. nov., P. ecuadorensis, P. constellaris) or with cone-shaped process sharp apically (P. fennahi sp. nov.). Anal tube moderately large, in dorsal view slightly elevated medially and in lateral view with lateral margins moderately prominent, and ventrally very slightly prominent (P. speciosa sp. nov., P. constellaris), moderately flattened dorsoventrally with lateral margins weakly prominent (P. fennahi sp. nov.), or tubular dorsally and ventrally with a third basal tubular and two third excavated (P. ecuadorensis Muir 1934). Genital styles thin at the base wide distally, apically in the expanded region approx. 3.0 times more long than wide (P. speciosa sp. nov.), approx. 2.0 times more long than wide (P. constellaris), or very rounded ventrally approx. 1.5 times more long than wide (P. fennahi sp. nov.). Aedeagus, aedeagal shaft with three apical spines (P. constellaris, P. speciosa sp. nov., P. fennahi sp. nov.), ventrally arranged apical spines (P. fennahi sp. nov.), laterally arranged apical spines (P. constellaris, P. speciosa sp. nov.), the middle spine (b) is the longest of the apical spines (P. constellaris), or the shortest (P. speciosa sp. nov.), distinct spines of the aedeagus without three spines from apex of the shaft, and with conspicuous larger spine ventrally (P. ecuadorensis). Female terminalia. Anal tube moderately large, medially elevated dorsally and distally truncated, in lateral view with ovipositor moderately surpassing the anal tube (P. constellaris, P. speciosa sp. nov.). Gonocoxae VIII moderately large, in lateral view with lateral carinae not reaching the middle of the segment (P. constellaris, P. speciosa sp. nov., P. fennahi sp. nov.); in caudal view larger and swollen apically (P. constellaris or medially (P. fennahi sp. nov.).	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B8850C07B8AE005B5FECDFC68.taxon	diagnosis	Diagnosis. This group can be distinguished mainly by characters of the male terminalia. However, some other characters are also useful for identification. Vertex moderately large, with apical transverse carina smaller than the subapical carina; apical compartment large, wider than long, moderately long medially; caudal margin triangular, slightly concave laterally with a small basal emargination; in frontal view with apical transverse carina straight. Frons approx. 1.9 – 2.4 times wider in the largest width than at the apically; frontoclypeal suture bent upwards medially, laterally straight. Tegmina (forewings) with pt large and sub rectangular, weakly, or well angled distally; fork of ScP + RA and RP moderately closer to bc than fork CuA 1 and CuA 2; rm- 1 proximally, together, or distally to first MP fork; mcu- 1 distally to first MP fork. Hind wing with RP bifid, MP trifid with petiolate anastomosis in MP 3 + 4 + CuA 1 distally forked (Y-type), CuA 1 and CuA 2 bifid. In ventral view, males with segment viii, complete, or separated medially. Hind tibia with 3 – 4 lateral spines, spines closest to the femur well small; 6 apical teeth as provided for the genus. 1 st tarsomere with 7 apical teeth. 2 nd tarsomere with 8 apical teeth, and with 1 – 3 platellae, in some species absent or in a smaller number on one of the legs (asymmetric bilaterally). Male terminalia. Pygofer bilaterally symmetrical; in lateral view, usually without lateral process, when present, laterally flattened (thin) vs. broad in Pintalia constellaris - group; dorsal margin concave, caudal margin usually straight, in some species slightly irregular. in ventral view, ventromedian process triangular, usually as wide as long. Anal tube long, tubular dorsally and excavated ventrally, developed and flattened distally with apex truncate and irregular, usually straight laterally and slightly asymmetrical in most species; in dorsal view, occurs a narrowing near the base but gradually elongates again to the apex; epiproct and paraproct short; in dorsal view epiproct triangular, and laterally with two small bent down margin; paraproct rounded, narrow laterally and slightly larger than the epiproct distally. Genital styles moderately or weakly wide distally, excavated ventrally and with small bent margins to the ventral region; basal opening between the genital styles, in diamond-shaped usually moderately small. Aedeagus display 5 – 7 spines; aedeagal shaft tubular, and usually wider apically near to flagellum; aedeagal shaft with 3 – 4 spines; usually a long or wider spine occurs laterally on the apex of the aedeagal shaft; ventral ridge absent. Flagellum usually with 2 – 6 spines in right laterally. Female terminalia. Anal tube long, in dorsal view rounded medially, with apex truncated; in lateral view with lateral margin slightly projected and straight, ventrally produced two third basal and a third apical excavated and apically produced again; epiproct and paraproct thin and short; epiproct triangular in dorsal view with lateral margin near to the base slightly developed and bent down; paraproct in dorsal view rounded. Gonocoxae VIII truncated, usually with rounded lateral carinae and irregular; in caudal view, it shows straight carinas moderately distant from the margin, closer to the genital segment, and distant medially or close to the ovipositor. Ovipositor large, in lateral view, sabre-shaped, curved upwards dorsally surpassing the anal tube. Species here assigned to this group: P. painensis sp. nov., P. lundi sp. nov., P. montalvaniensis sp. nov., P. dorensis sp. nov., P. muiri sp. nov.	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B8850C07B8AE005B5FECDFC68.taxon	distribution	Distribution. BRA. Minas Gerais State (Fig. 77). Notes. Fennah (1945) grouped some species due to the anal tube developed distally, bent down in approx. 90 ° (e. g., P. bicaudata- group), bent in approx. 45 ° (e. g., P. albolineata- group), and to the shape of the genital styles. Here we have grouped all the new species with distally developed anal tube, with spines on the flagellum and genital styles weakly or moderately wide distally. A review of the species described by Muir 1934, and Fennah 1945 is indicated to properly allocate them to groups or new genera. Some species deposited in ISLA may have a morphology very similar to the morphology described for Pintalia painensis- group, however the flagellum does not present spines. Morphology. Body length: 3.9 – 5.3 ♂ mm (n = 18). 5.0 – 5.8 mm ♀ (n = 6). Head. Vertex with apical compartment Square and large, in dorsal view with apical transverse carina usually almost straight and slightly irregular, totally developed (P. painensis sp. nov., P. lundi sp. nov., P. montalvaniensis sp. nov.), or slightly evanescent laterally (P. dorensis sp. nov., P. muiri sp. nov.); the subapical carina can be concave (P. dorensis sp. nov., P. muiri sp. nov.) or bent medially (P. painensis sp. nov., P. lundi sp. nov., P. montalvaniensis sp. nov.); approx. 1.9 – 2.4 times wider in the largest width than apically. Thorax. Pronotum narrow, inconspicuous median carina and moderately developed but evanescent submedian carina behind the eyes. Tegmina (forewings) mostly hyaline with a dark brown spots, with four rounded spots in nodal region (P. painensis sp. nov., P. montalvaniensis sp. nov.) or with one irregular spot (P. dorensis sp. nov., P. muiri sp. nov.) or two irregular spots (P. lundi sp. nov.); rm- 1 ahead to first fork MP (P. painensis sp. nov.) together (P. dorensis sp. nov., P. lundi sp. nov., P. muiri sp. nov.) or distally (P. montalvaniensis sp. nov.) and mcu- 1 occurred distally to first MP fork (P. dorensis sp. nov., P. lundi sp. nov., P. muiri sp. nov., P. montalvaniensis sp. nov.). Posterior legs, hind tibia with three lateral spines (P. dorensis sp. nov., P. lundi sp. nov.) or four lateral spines (P. painensis sp. nov., P. muiri sp. nov., P. montalvaniensis sp. nov.); 1 st tarsomere with two apical teeth external slightly larger, and two medially with approx. the same size; 2 nd tarsomere with 1 – 2 lateral apical teeth well larger (P. painensis sp. nov., P. dorensis sp. nov., P. lundi sp. nov.) or slightly larger (P. muiri sp. nov., P. montalvaniensis sp. nov.) and with 1 – 3 platellae. Male terminalia. Pygofer bilaterally symmetric, laterally with a developed process, rounded and thin (P. dorensis sp. nov.), very large and rounded (P. muiri sp. nov.) or moderately rounded (P. painensis sp. nov., P. lundi sp. nov., P. montalvaniensis sp. nov.). Anal tube in dorsal view, very narrow near to the base (P. dorensis sp. nov., P. painensis sp. nov.) or slightly narrow (P. lundi sp. nov., P. montalvaniensis sp. nov., P. muiri sp. nov.). Genital styles moderately wide distally (P. dorensis sp. nov., P. montalvaniensis sp. nov.) or weakly wide distally (P. painensis sp. nov., P. lundi sp. nov., P. muiri sp. nov.); basal opening of styles usually moderately small and diamond-shaped (P. painensis sp. nov., P. lundi sp. nov., P. dorensis sp. nov., P. montalvaniensis sp. nov.) or weakly distinct, slightly rounded laterally and triangular elongated apically (P. muiri sp. nov.). Aedeagus with a large spine on the right side of the apex of aedeagal shaft (P. painensis sp. nov., P. lundi sp. nov., P. muiri sp. nov., P. dorensis sp. nov.) or a large spine on the left lateral of the apex of aedeagal shaft (P. montalvaniensis sp. nov.); in left lateral view with one spine (P. painensis sp. nov., P. lundi sp. nov., P. muiri sp. nov., P. montalvaniensis sp. nov.) or two spines (P. dorensis sp. nov.); in right lateral view with one short spine (P. lundi sp. nov., P. montalvaniensis sp. nov., P. dorensis sp. nov.) or two short spines (P. painensis sp. nov.). Flagellum with two spines (P. muiri sp. nov., P. lundi sp. nov.), one of them bifid (P. lundi sp. nov.), with three spines (P. painensis sp. nov., P. dorensis sp. nov.) or four spines (P. montalvaniensis sp. nov.). Female terminalia. Female terminalia very similar between species with little change in size of the anal tube and gonocoxae VIII.	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B884CC0648AE0017CFEBEFE98.taxon	description	(Figs. 1 A, 2 A – D, 3 A – B, 11 A, 12 A)	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B884CC0648AE0017CFEBEFE98.taxon	materials_examined	Type material. Holotype ♀. Brazil (BRA); (NHRS-GULI 000075710).	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B884CC0648AE0017CFEBEFE98.taxon	description	Supplementary description. Coloration (preserved specimen). Body mostly deep brown (56), contrasting with some regions of light orange-yellow (70) on the vertex, frons and legs, pale region between the carinae of the mesonotum in dark brown (55). In lateral view with rounded light orange-yellow spot above the eyes and gena. Tegmina hyaline with large conspicuous spot in deep yellowish-brown (75), forming a triangular despigmented area of the Rp 2.1, going to C 5 cell and approximate of the Scp vein. Body length. Female (examined): 5.2 mm (n = 1); 6.0 mm (Stål, 1862). Head. Vertex (Figs. 2 B – C): apical compartment 4.0 to 4.3 times wider than medially long; caudal margin of the vertex and subapical carina concave; apical transverse carina also concave, with opening towards the frons. Frons (Fig. 2 A): approx. 1.4 times longer than maximally wide; approx. 2.1 times wider at the height of the antennae than at the anterior region of the frons; median carina moderately developed; thin lateral carinae; anterior region of the frons trapezoidal, straight apically (apical transverse carina). Frontoclypeal suture: semicircular, bent upwards, slightly straight laterally. Post and anteclypeus: median carina well developed. Thorax. Pronotum (Fig. 2 B – D): submedian carinae weakly developed; hind margin obtuse; light coloration in the region between the eyes (like at vertex). Mesonotum (Fig. 2 B – D): tricarinate; darker lateral margins than between the carinae; flat in lateral view with distal region slightly inclined. Tegmina (forewings) (Fig. 2 D, 11 A, 12 A): length 8.4 mm; approx. 2.5 times longer than wide; hyaline / opaque with conspicuous dark spot in y-shape; fork of ScP + RA and RP ahead of forks CuA 1 and CuA 2; C 5 with two crossveins; r-m 1 occurring proximally of the first MP fork; m-cu 1 occurs together with the MP fork; RP quadrifid; MP 1 + 2 trifid; MP 3 + MP 4 bifid; tubercles evanescent, more evident between the cubital cell and postclaval margin, and some in ScP + RA and RP; 12 apical cells; 7 – 8 subapical cells. Hind wing with RP bifid, MP trifid with petiolate anastomosis in MP 3 + 4 + CuA 1 distally forked (Y- type), CuA 1 and CuA 2 bifid. Posterior legs. Hind tibia (Fig. 3 A – B): 4 lateral spines and 6 apical teeth. It was not possible to observe the apical teeth of the 1 st and 2 nd tarsomere. Female terminalia (Figs. 2 D; 3 A). Anal tube elongated; curved dorsally and excavated ventrally. Gonocoxae VIII in caudal view ovoid with lateral margins curved. Ovipositor long, in lateral view, sabre-shaped, curved upwards dorsally.	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B884CC0648AE0017CFEBEFE98.taxon	diagnosis	Diagnosis. Pintalia lateralis Stål, 1862 can be distinguished from other species of the genus Pintalia by the totally concave carinae of the vertex. Coloration of the frons, and vertex pale (except laterally) and conspicuous dark “ y ” - shaped spot on the tegmina.	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B884CC0648AE0017CFEBEFE98.taxon	distribution	Distribution. The only location reported by Stål (1862) was “ Brazil ” (Type locality) although the species described by Stål (1862) were presumed to be from the vicinity of Rio Janeiro.	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B884CC0648AE0017CFEBEFE98.taxon	discussion	Notes. As with the other species redescribed here, it was not possible to assess the apical teeth of the 1 st and 2 nd tarsomere.	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B884EC0638AE00220FF4DF817.taxon	description	(Figs. 1 B, 4 A – D, 11 B)	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B884EC0638AE00220FF4DF817.taxon	materials_examined	Type material. Holotype ♀. Brazil (BRA). (NHRS-GULI 000075711).	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B884EC0638AE00220FF4DF817.taxon	description	Supplementary description. Coloration (preserved specimen). Body mostly dark brown (59), contrasting with some regions of strong brown (55) on the pronotum medially, tegulae, mesonotum distally and legs. Tegmina hyaline with spots in strong brown (55), with a conspicuous rounded spot in radial area. Body length. Female (examined): 4.3 mm (n = 1); 4.5 mm (Stål, 1862). Head. Vertex (Fig. 4 B – C): distinct, subapical carina recorded on the disk; median carina developed in the basal compartment and evanescent in the apical compartment; raised apical transverse carina; curved caudal margin; basal compartment slightly larger than the apical compartment. Frons (Fig. 4 A): in frontal view approx. 1.2 times longer than maximally wide; approx. 2.4 times wider medially than apically; median carina well developed; thick lateral carinae between the largest width (near the scapus) and the anterior region of the frons; anterior region of the frons trapezoidal, curved in the middle, and straight apically. Frontoclypeal suture: arched medially and slightly concave laterally, bypassing the ocellus. Post-and anteclypeus: median carina well developed and rounded in the middle. Thorax. Pronotum (Figs. 4 B – D): submedian carina developed; median carina developed; hind margin obtusely angled or rectangular; coloration a little lighter in the region between the eyes. Mesonotum (Figs. 4 B – D): tricarinate, little darker lateral margins than between the carinae; flat in lateral view with distal region slightly inclined. Tegmina (forewings) (Fig. 4 D, 11 B): length 5.9 mm; approx. 2.6 times longer than wide; transverse dark spot in the center of the tegmina with curved lateral elongation towards the anterior cubital margin; two dark spots with " C " - shaped form in prenodal cells, and two dark spots in the postcostal cell; double tubercles occurring throughout the tegmina; fork of ScP + RA and RP, ahead of forks CuA 1 and CuA 2; C 5 with one traversal vein; r-m 1 together to the first MP fork; mcu 1 distally to r-m 1, together from MP 3 vein; RA bifid with punctual anastomosis RA + RP 1; RP quadrifid; MP 1 + 2 trifid; MP 3 + MP 4 bifid; 12 apical cells; 7 subapical cells. Posterior legs. Hind tibia: 4 lateral spines, and 6 apical teeth (see remarks). Female terminalia. Anal tube slightly longer than wide; flattened ventrally. Gonocoxae VIII, in caudal view appears ovoid with lateral margins rounded. Ovipositor in lateral view, sabre-shaped, curved upwards dorsally, just surpassing (or not surpassing) the anal tube.	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B884EC0638AE00220FF4DF817.taxon	diagnosis	Diagnosis. Pintalia consobrina can be distinguished from other species of the genus Pintalia mainly by the distinct vertex with subapical carina engraved into disc instead of elevated, and transverse dark spot in the center of the tegmina with curved lateral elongation towards the cubital margin.	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B884EC0638AE00220FF4DF817.taxon	distribution	Distribution. The only location reported by Stål (1862) is Brazil (type locality).	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B884EC0638AE00220FF4DF817.taxon	discussion	Notes. A female specimen that is very similar to P. consobrina was found in the ISLA collection. The specimen was collected during the dry season (20. x. 2011) (ISLA 100960) in the MP- 008 cave, in Itabirito municipality (MG), a region strongly impacted by mining. This species is probably a subtroglophile due to the low occurrence recorded in these cavities so far, and perhaps came to lay eggs on the roots present in the cave (subtroglophile habits with subterranean eggs and larva). However, we still cannot rule out the hypothesis that it is a eutroglophile species, and it is part of an underground population living in the cave or the MSS (more eutroglophile habits). However, cave MP- 008 also houses the species Ferricixius davidi Hoch and Ferreira, 2012 and during its discovery, several exhaustive collections were carried out in the cavity and its surroundings, and the only specimen of the P. consobrina type found was the one reported here. The referred specimen exhibits three lateral spines on the posterior tibia, eight apical teeth on the first posterior tarsomere and nine apical teeth on the 2 nd tarsomere posterior, five of them with platellae.	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B884AC0618AE005B5FE56FCD8.taxon	description	(Figs. 1 C, 5 A – D, 11 C)	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B884AC0618AE005B5FE56FCD8.taxon	materials_examined	Type material. Holotype ♀. Brazil (BRA); (NHRS-GULI 000075712).	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B884AC0618AE005B5FE56FCD8.taxon	description	Supplementary description. Coloration (preserved specimen). Body mostly brownish black (65), contrasting with some regions of pale orange yellow (73) on the head, pronotum medially, tegulae and legs. Tegmina mostly hyaline, strong yellowish brown (74), with transversal spots in dark yellowish brown (78), some regions in pale orange yellow (73) on costal margin, radial area, and some veins. Body length. Female (examined): 4.1 mm (n = 1); 4.3 mm (Stål, 1862). Head. Vertex (Fig. 5 B – C): distinct; carinae weakly or not elevated. Apical compartment 4.8 to 5.0 times wider than medially long; apical transverse carina and subapical carina arched; basal compartment with well-developed median carina. Frons (Fig. 5 A): 1.1 times longer than maximally wide; approx. 1.7 times wider medially than apically; lateral carinae weakly elevated; median carina well developed; anterior region of the frons trapezoidal wide, straight apically. Frontoclypeal suture: bent upwards, moderately semicircular, slightly straight laterally and below the frontal ocellus. Post and anteclypeus: wide apically, well rounded medially with median carina moderately developed. Thorax. Pronotum (Figs. 5 B – D): submedian carinae weakly developed; hind margin obtusely angulated or rectangular; light coloration in the region between the eyes (as at the vertex). Mesonotum (Figs. 5 B – D): tricarinate; uniformly dark; flat in lateral view with distal region slightly inclined. Tegmina (forewings) (Figs. 5 D, 11 C): length 5.3 mm; hyaline / opaque with two transverse dark spots, one near the tegulae and the other in the center of tegmina; postcostal cell with light longitudinal spot next to the costal vein and four transverse dark spots close to the pterostigma; radial area with two light spots near of the distal margin; distinct veins and cells; Cua 1 and Cua 2 fork starting more proximally to the basal cell than the fork in ScP + RA and RP that occurs more distally; C 5 with 5 crossveins; C 4 with 4 crossveins, occurring at approx. the same height as the crossveins of C 5; C 3 b very elongated; mcu- 1 closer to bc than rm- 1; rm- 1 occurs posteriorly of the first fork of MP; RA bifid with punctual anastomosis RA + RP 1; RP quadrifid; MP 1 + 2 bifid; MP 3 + 4 bifid; simple tubercles in all veins; 12 apical cells (small); 7 – 13 subapical cells (elongated). Posterior Legs. Hind tibia: 2 lateral spines and 6 apical teeth.	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B884AC0618AE005B5FE56FCD8.taxon	diagnosis	Diagnosis. Pintalia fasciatipennis can be distinguished from other species of the genus Pintalia mainly by the distinct frons, slightly longer than wide, with non-elevated lateral carinae, vertex clear in dorsal view, nodal line near the distal margin and two transverse dark spots on the tegmina, one near the tegulae and the other in the center of tegmina.	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B884AC0618AE005B5FE56FCD8.taxon	distribution	Distribution. The only location reported by Stål (1862) is Brazil (type locality).	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B884AC0618AE005B5FE56FCD8.taxon	discussion	Remarks. It was difficult to observe some of the veins and cells in the images, since they are in light blue in Fig. 11 C, so the information about the pattern of veins and cells of the tegmina of P. fasciatipennis provided here must be interpreted with care.	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B884BC0618AE0068CFA8AF8C8.taxon	description	(Figs. 1 D, 6 A – D, 11 D)	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B884BC0618AE0068CFA8AF8C8.taxon	materials_examined	Type material Holotype ♀. Brazil (RJ). (NHRS-GULI 000075713).	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B884BC0618AE0068CFA8AF8C8.taxon	description	Supplementary description. Coloration (preserved specimen). Body mostly deep orange yellow (69), contrasting with some regions of light orange yellow (70) on the legs. Tegmina hyaline, with defuse spots in dark orange-yellow (72). Body length. Female (examined): 4.6 mm (n = 1); 5.5 mm (Stål, 1862). Head. Vertex (Fig. 6 B – C): apical compartment 2.6 to 2.8 times wider than medially long; apical transverse carina slightly concave, almost straight; subapical carina and caudal margin well concave. Frons (Fig. 6 A): 1.5 times longer than maximally wide, approx. 1.8 times wider medially than apically; median carina moderately or weakly developed; thin lateral carinae; anterior region of the frons trapezoidal, and straight apically. Frontoclypeal suture: straight laterally and arched medially. Post and anteclypeus: with median carina moderately developed. Thorax. Pronotum (Figs. 6 B – C). short dorsally; submedian carina moderately developed; hind margin obtusely angulate or rectangular. Mesonotum (Figs. 6 B – D): tricarinate, flat in lateral view. Tegmina (forewings) (Fig. 6 D, 11 D): length 7.9 mm; approx. 2.9 times longer than wide; hyaline / opaque with blurred spots, a thin dark spot inclined occurs in the center connecting the pt and icu; fork of ScP + RA and RP, ahead of forks CuA 1 and CuA 2; r-m 1 inclined, and emerges in first MP fork, proximally of mcu 1; mcu 1 occurs distally of the first MP fork; RP quadrifid; MP 1 + 2 trifid; MP 3 + MP 4 bifid; tubercles evanescent, more evident between the cubital cell and postclaval margin, and some in ScP + RA and RP (+ MA); 12 apical cells; 7 subapical cells. Hind wing with RP bifid, MP trifid with petiolate anastomosis in MP 3 + 4 + CuA 1 distally forked (Y-type), CuA 1 and CuA 2 bifid. Posterior legs. Hind tibia: 1 lateral spine and 6 apical teeth.	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B884BC0618AE0068CFA8AF8C8.taxon	diagnosis	Diagnosis. Pintalia fraterna can be distinguished from other species of the Pintalia genus by the predominantly deep yellowish-brown color in dorsal view, anterior compartment of the vertex “ perfect ” rectangle when mostly trapezoidal in the other species of the genus.	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B884BC0618AE0068CFA8AF8C8.taxon	distribution	Distribution. The only location reported by Stål (1862) is Brazil: Rio de Janeiro State (type locality).	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B8844C06C8AE000B7FD9AFEE0.taxon	description	(Figs. 1 E, 7 A – D, 11 E)	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B8844C06C8AE000B7FD9AFEE0.taxon	materials_examined	Type material. Holotype ♀. Brazil (RJ). (NHRS-GULI 000075714).	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B8844C06C8AE000B7FD9AFEE0.taxon	description	Supplementary description. Coloration (preserved specimen). Body mostly dark brown (59), contrasting with some regions of light orange-yellow (70) on the pronotum, frons laterally, and some regions of the legs. Forelegs light orange-yellow (70), with dark brown stripes (59). Tegmina transparent and smoked, with dark brown (59) veins, without spots. Body length. Female: 5.0 mm (n = 1) (Stål, 1862). Head. Vertex (7 B – C): distinct; apical compartment 1.2 to 1.4 times wider than medially long; apical transverse carina elongated medially, more visible in frontal view; subapical carina straight; concave caudal margin; median carina occurs in little more than 1 / 2 of basal compartment. Frons (Fig. 7 A): approx. 1.0 times longer than maximally wide; approx. 1.4 times wider medially than apically; median carina moderately developed above the level of the antennae; apically touching the apical transverse carina, evanescent in the basal half near the median ocellus; thin lateral carinae; anterior region of the frons distinct, at same compound eye level. Frontoclypeal suture: semicircular, very curved, bent upwards. Post- and anteclypeus: with median carina well developed, starting at about half the length of anteclypeus. Thorax. Pronotum (Figs. 7 B – D): submedian carinae weakly developed; hind margin obtusely angled or rectangular; lighter coloration at the caudal margin and between the eyes. Mesonotum (Figs. 7 B – D): tricarinate, in dorsal view almost straight and parallel carinae, very flat in lateral view. Tegmina (forewings) (Figs. 7 D, 11 E): length 7.1 mm; approx. 2.5 times longer than wide; transparent and smoked, without spots; C 1 b exhibit a crossvein; C 5 small; fork of ScP + RA and RP, approx. 2.0 times closer to the basal cell than forks CuA 1 and CuA 2; r-m 1 occurs from the first MP fork; mcu 1 occurs distally r-m 1 and laterally to MP 3 + 4; RP quadrifid, RP 2 + 3 with petiolate anastomosis distally forked; MP 1 + 2 trifid with petiolate anastomosis in MP 1.1 + MP 1.2; MP 3 + 4 bifid; simple or inconspicuous tubercles; 12 – 13 apical cells; 8 subapical cells. Posterior legs. Hind tibia: without lateral spines and with 6 apical spines.	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B8844C06C8AE000B7FD9AFEE0.taxon	diagnosis	Diagnosis. P. inornata can be distinguished from other Pintalia species mainly by the distinct vertex, and in frontal view with median carina of the frons joined to the anterior margin of the vertex in a " y " shape, and petiolate ramifications in the distal veins of RP and MP.	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B8844C06C8AE000B7FD9AFEE0.taxon	distribution	Distribution. The only location reported by Stål (1862) is Brazil and Rio de Janeiro State (Type location).	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B8844C06C8AE000B7FD9AFEE0.taxon	discussion	Remarks. Emeljanov (2007) described the genus Proclytus Emeljanov, 2007 to allocate a single species from French Guiana, Proclytus guianensis Emeljanov, 2007. P. guianensis exhibits the vertex like P. consobrina with the apical transverse carina elongated medially, hind tibia without lateral spines. Some specimens identified with Proclytus in the ISLA collection, in addition to the corresponding characters of the head, wings and male terminalia with Proclytus guianensis, also exhibit dark stripes on the forelegs like P. consobrina, which unfortunately cannot yet have the male genitalia verified.	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B8846C06D8AE00494FAD2FCBC.taxon	description	(Figs. 1 F, 8 A – D, 11 F)	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B8846C06D8AE00494FAD2FCBC.taxon	materials_examined	Type material. Holotype ♀. Brazil (RJ); (NHRS-GULI 000075715).	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B8846C06D8AE00494FAD2FCBC.taxon	description	Supplementary description. Coloration (preserved specimen). Body mostly deep orange yellow (69), contrasting with some regions of moderate orange yellow (71) on the vertex and legs. Tegmina hyaline, deep orange yellow (69) and strong brown (55) on margins, without evident spots, clearer costal margin at the level of the pterostigma. Body length. Female (examined): 4.0 mm (n = 1); 4.3 mm (Stål, 1862). Head. Vertex (Fig. 8 B – C): apical compartment 2.5 to 2.8 times wider than medially long; apical transverse carina almost straight; subapical carina slightly concave; caudal margin concave. Frons (Fig. 8 A): 1.1 times longer than maximally wide; approx. 1.6 times wider medially than apically; median carina moderately or weakly developed, apically y-shaped forming a triangular region apically along with the apical transverse carina; thin lateral carinae; anterior region of the frons trapezoidal, and straight apically. Frontoclypeal suture: semicircular, bent upwards. Post and anteclypeus: median carina well developed. Thorax. Pronotum (Figs. 8 B – D): short, with submedian carina moderately developed; hind margin obtusely angulate. Mesonotum (Figs. 8 B – D): tricarinate; flat in lateral view with inclined distal region. Tegmina (forewings) (Figs. 8 D, 11 F): length 7.2 mm; approx. 2.5 times longer than wide; uniform color, no contrasting dark spots; fork of ScP + RA and RP, little ahead of forks CuA 1 and CuA 2; parallel r-m 1 and mcu- 1, occurring distally of first MP fork; RP quadrifid; MP 1 + 2 trifid; MP 3 + MP 4 trifid; simple tubercles, in all veins; 12 – 13 apical cells; 7 subapical cells. Posterior legs. Hind tibia: without lateral spines and 6 apical spines. Female terminalia. Anal tube small, wider than long, and cylindrical. Gonocoxae VIII truncate, in caudal view with lateral margin almost straight and moderately elevated. Ovipositor in lateral view, sabre-shaped, curved upwards dorsally.	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B8846C06D8AE00494FAD2FCBC.taxon	diagnosis	Diagnosis. Pintalia obscuripennis can be distinguished from other species of the genus Pintalia by uniform coloration (varying little), and opaque tegmina without conspicuous dark spots, and Cua 1 trifid.	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B8846C06D8AE00494FAD2FCBC.taxon	distribution	Distribution. The only location reported by Stål (1862) is Brazil and Rio de Janeiro State (type locality).	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B8847C06D8AE006A8FAD2F8C8.taxon	description	(Figs. 1 H, 9 A – D; 11 H)	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B8847C06D8AE006A8FAD2F8C8.taxon	materials_examined	Type material. Holotype ♂. Brazil (RJ); (NHRS-GULI 000075717).	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B8847C06D8AE006A8FAD2F8C8.taxon	description	Supplementary description. Coloration (preserved specimen). Body mostly dark brown (59), contrasting with some regions of light orange-yellow (70) on the vertex, pronotum, tegulae and legs. Tegmina hyaline with stitches over the veins and a large spot starting at the costal margin and covering the mostly distal region in dark brown (59). Body length. Male (examined): 3.6 mm (n = 1). Female 4.5 mm (n = 1) (Stål, 1862). Head. Vertex (Figs. 9 B – C): apical compartment 2.8 to 3.0 times wider than medially long; apical transverse carina slightly concave, almost straight; subapical carina slightly arched, slightly concave, and asymmetrical; concave caudal margin with small basal emargination. Frons (Fig. 9 A): approx. 1.2 times longer than maximally wide; approx. 1.5 times wider medially than apically; median carina moderately developed, evanescent basally, slightly y-shaped apically; thin lateral carinae; anterior region of the frons trapezoidal, almost straight apically. Frontoclypeal suture: straight bent upwards. Post-and anteclypeus: median carina well developed. Thorax. Pronotum (Figs. 9 B – D): submedian carina moderately developed and hind margin obtusely angled or rectangular. Mesonotum (Figs. 9 B – D): tricarinate; lateral margins darker than between the carinae; flat in lateral view with distal region inclined. Tegmina (forewings) (Figs. 9 D, 11 H): length 6.0 mm; approx. 2.8 times longer than wide; hyaline with a conspicuous spot that covers all the nodal cells and narrows in the prenodal area, until reaching the costal margin; ScP + RA and RP, ahead of forks CuA 1 and CuA 2; r-m 1 and mcu- 1 occurring distally of first MP fork; m-cu 1 occurs proximally r-m 1; RP quadrifid; MP 1 + 2 trifid; MP 3 + MP 4 bifid; simple tubercles throughout all the veins; 12 apical cells; 7 – 8 subapical cells. Posterior legs. Hind tibia: 1 lateral spine and 6 apical teeth.	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B8847C06D8AE006A8FAD2F8C8.taxon	diagnosis	Diagnosis. P. proxima can be distinguished from other species of the Pintalia genus by the tegmina with radial cells distinct, in the transverse direction, with a conspicuous spot that covers all the nodal cells and narrows before to MP fork, until reaching the costal margin.	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B8847C06D8AE006A8FAD2F8C8.taxon	distribution	Distribution. The only location reported by Stål (1862) is Brazil and Rio de Janeiro State (type locality).	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B8840C0688AE00307FAD2FC04.taxon	description	(Figs. 1 I, 10 A – D, 11 I)	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B8840C0688AE00307FAD2FC04.taxon	materials_examined	Type material. Holotype ♀. Brazil (RJ); (NHRS-GULI 000075718).	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B8840C0688AE00307FAD2FC04.taxon	description	Supplementary description. Coloration (preserved specimen). Body mostly dark brown (59), contrasting with some regions of strong brown (55) on the vertex, postclypeus, and legs. Tegmina hyaline with a moderate brown (58) coloration covering the terminal cells, from the pterostigma region to the posterior margin. Deep reddish orange (36) mostly at the base of the antenna pedicels and around (or under) the tegulae. Body length. Female (examined): 3.9 mm (n = 1); 4.0 mm (n = 1) (Stål, 1862). Head. Vertex (Figs. 10 B – C): apical compartment 1.5 to 1.7 times wider than medially long; apical transverse carina and subapical carina medially elongated; apical transverse carina more visible in frontal view, forming a sharp angle in its middle where the frontal median carina is joining; concave caudal margin with small recess medially; basal compartment with median carina reaching the submedian carina. Frons (Figs. 10 A): 1.1 times longer than maximally wide; approx. 1.5 times wider medially than apically; median carina moderately or weakly developed, touching the apical transverse carina; thin lateral carinae and sinuous in frontal view; anterior region of the frons below the eyes; very low sunk apically. Frontoclypeal suture: semicircular (shallow-open), bent upwards. Post- and anteclypeus: distinct, slightly longer than wide and tubular basally. Thorax. Pronotum (Figs. 10 B – D): short, with submedian carina weakly developed and hind margin obtuse. Mesonotum (Fig. 10 B – D): tricarinate, slightly darker between carinae than laterally; flat in lateral view. Tegmina (forewings) (Figs. 10 E, 11 I): length 5.4 mm; approx. 2.5 times longer than wide; hyaline, uniform coloration without conspicuous spots; fork of ScP + RA and RP, at the same height or slightly forward of forks CuA 1 and CuA 2; r-m 1 occurring proximally of the first MP fork; m-cu 1 occurs distally of the first MP fork; RP quadrifid; MP 1 + 2 trifid; MP 3 + MP 4 bifid; simple tubercles in all veins; 12 apical cells; 7 subapical cells; all the nodal cells and narrows before to MP fork, until reaching the costal margin. Hind wing with RP bifid, MP trifid with petiolate anastomosis in MP 3 + 4 + CuA 1 distally forked (Y-type), CuA 1 and CuA 2 bifid. Posterior legs. Hind tibia: without lateral spines and 6 apical spines. Female terminalia. Anal tube moderately small, approx. 1.2 – 1.4 times longer than wide; in dorsal view elevated medially and with two “ grooves ” laterally. Gonocoxae VIII, in caudal view rounded slightly larger apically, with lateral margins rounded and elevated. Ovipositor in lateral view, sabre-shaped, curved upwards dorsally.	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B8840C0688AE00307FAD2FC04.taxon	diagnosis	Diagnosis. P. ustulata can be distinguished from other Pintalia species by the tegmina with by the darkened apical cells and the remaining hyaline tegmina pale. Apical transverse carina and submedian carina medially elongated and apical transverse carina better visible in frontal or dorsofrontal view of the head. In lateral view, head with fully convex margins, from the vertex to the postclypeus.	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B8840C0688AE00307FAD2FC04.taxon	distribution	Distribution. The only location reported by Stål (1862) is Brazil and Rio de Janeiro State (type locality).	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B8842C0578AE00138FC35FD2C.taxon	description	(Figs. 1 G, 11 G, 14 A – D)	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B8842C0578AE00138FC35FD2C.taxon	materials_examined	Type material. Holotype ♂. Brazil (RJ); (NHRS-GULI 000075716).	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B8842C0578AE00138FC35FD2C.taxon	description	Supplementary description. Coloration (preserved specimen). Body mostly dark brown (59), contrasting with some regions of pale orange-yellow (73) on the vertex, frons laterally, pronotum, tegulae and legs. Pronotum light color as at vertex. Mesonotum uniformly dark in color. Tegmina hyaline with transverse spots in strong yellowish brown (74), medially with a large and diffuse spot covering the entire region near the postclaval margin and anterior cubital area. Body length. Male (examined): 3.0 mm (n = 1); 3.3 mm (Stål, 1862). Head. Vertex (Figs. 14 B – C): apical compartment 4.0 to 5.0 times wider than medially long; apical transverse carina and subapical carina concave; caudal margin straight. Frons (Fig. 14 A): 1.4 times longer than maximally wide; approx. 1.9 times wider medially than apically; median carina well developed; anterior region of the frons trapezoidal, concave apically. Frontoclypeal suture: semicircular, bent upwards, slightly straight laterally. Post- and anteclypeus: with median carina well developed. Thorax. Pronotum (Figs. 14 B – D): submedian carina weakly developed; hind margin obtusely angled or rectangular. Mesonotum (Figs. 14 B – D): tricarinate; flat in lateral view with distal region slightly elevated. Tegmina (forewings) (Fig. 14 D, 11 G): length 4.8 mm; approx. 2.3 times longer than wide; hyaline / opaque with dark spots; anterior cubital area and Postclaval area totally dark with a triangular hyaline spot between CuA 1 and the A 2; costal / radial half predominantly hyaline with 6 dark stripes (as follows: 2 small between SC and C; 1 long and curved starting in vein C and ending on the opposite side, in A 2 near CuP - 1 very thin starts in pt and ends near the fork of RP 2 and RP 3; 1 short but wide occurring between RP 3 and the radial margin; 1 thin, starts at the radial margin and ends in RP 4); fork of ScP + RA and RP, ahead of forks CuA 1 and CuA 2; r-m 1 occurring proximally first MP fork and mcu- 1; m-cu 1 occurs at the firs MP fork; RA bifid with punctual anastomosis RA + RP 1; RP quadrifid; MP 1 + 2 trifid; MP 3 + MP 4 bifid; C 1 b well small or inconspicuous; with small tubercles little evident on veins; 12 apical cells; 7 subapical cells. Posterior legs. Hind tibia: 2 lateral spines and 6 apical teeth. Male terminalia. Pygofer: in lateral view, without processes, dorsal margin straight / slanted, distal margin concave; ventromedian process triangular. Anal tube elongated and flattened dorsoventrally without ventral process distally; paraproct developed and flattened dorsoventrally with distal margin concave; epiproct short. Genital styles wider apically and slender near base.	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B8842C0578AE00138FC35FD2C.taxon	diagnosis	Diagnosis. Pintalia pictipennis can be distinguished from other species of the genus Pintalia by the darker frons in the middle than on the developed lateral carinae, spots pattern of the tegmina, and anal tube long and dorsoventrally flattened without process lateroventral distally, and paraproct developed flattened dorsoventrally.	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B8842C0578AE00138FC35FD2C.taxon	distribution	Distribution. The only location reported by Stål (1862) is Brazil and Rio de Janeiro State (type locality).	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B8842C0578AE00138FC35FD2C.taxon	discussion	Note: The holotype is male instead of female as stated by Stål.	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B887DC0528AE007CDFAF3FEE0.taxon	description	https: // zoobank. org / urn: lsid: zoobank. org: act: 54441 BD 3 - 9 EC 5 - 4 B 76 - 9 A 56 - 35 B 9 A 79 EFD 0 D (Figs. 15 A – F; 16 A – G; 17 A – H; 18 A – D; 19)	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B887DC0528AE007CDFAF3FEE0.taxon	materials_examined	Type material. Holotype: Male Brazil, MG. Itabirito municipality, MP 8 Cave, (UTM 619795 W, 7764761 S, 23 K), 29. iii. 2012, (Carste et al.) (ISLA 100935). Holotype condition: not dissected, stored in an individual vial in ethanol 70 %. Paratypes. Same data as holotype, except for 1 ♀ (ISLA 100936); 1 ♂ (dissected) (ISLA 100937); 3 ♀ (ISLA 100938); 1 ♂ and 2 ♀, MP 7 Cave, 20. x. 2011, (Carste et al.) (ISLA 100939); 2 ♀, 20. x. 2011, (Carste et al.) (ISLA 100940). Additional material examined. Brazil, MG: Mariana municipality, 2 ♀ FZL- 0006 Cave, (UTM 665069 W, 7754333 S, 23 K), 27. ix. 2018, (Carmo T. O) (ISLA 100941) and 14. ii. 2019 (ISLA 100942); 1 ♀ FZLM 5 - 0007 Cave, (UTM 665265 W, 7756203 S, 23 K), 12. ii. 2019, (Carmo T. O) (ISLA 100943); 1 ♀ FZL- 0009 Cave, (UTM 665210 W, 7753768, 23 K), 14. ii. 2019, (Carmo T. O) (ISLA 100944); 1 ♂ FZL- 0010 Cave, (UTM 665210 W, 7753756 S, 23 K), 14. ii. 2019, (dissected), (Carmo T. O) (ISLA 100945); 1 ♂ FN- 0005 Cave, (UTM 663582 W, 7763205 S, 23 K), 23. xii. 2020, (Spelayon et al.) (ISLA 100946); 1 ♂ FN- 0025 Cave, (UTM 662636 W, 7763761 S, 23 K), 24. IX - 30. x. 2020, (Spelayon et al.) (ISLA 83490); 1 ♂ FN- 0012 Cave, (UTM 663142 W, 7764734 S, 23 K), 24. IX- 9. x. 2020, (Spelayon et al.) (ISLA 83537); 1 ♂ GOGO _ 030 Cave, (UTM 662662 W, 7749239 S, 23 K), 13 - 16. i. 2014, (Spelayon et al.) (ISLA 46347); 1 ♂ GOGO _ 032 Cave, (UTM 662166 W, 7749320 S, 23 K), 13 - 6. i. 2014, (Spelayon et al.) (ISLA 46345).	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B887DC0528AE007CDFAF3FEE0.taxon	description	Description. Coloration (specimen preserved in 70 % ethanol). Body generally strong yellow orange (68), near carinae and abdomen dorsally in deep yellowish brown (75), contrasting with regions of brilliant yellow orange (67) on mesonotum, and light-yellow orange (70) on legs and abdomen ventrally. Tegmina hyaline with very light gray (93) diffuse spots. Body length. Male. 3.1 – 4.0 mm (n = 9); Female. 3.8 – 4.0 (n = 8). Head. Vertex (Figs. 15 A, C): approximately 2.0 times wider (0.4) than long (0.2); apical compartment small and laterally narrow, approx. 2.5 wider (0.2) than medially long (0.08); apical transverse carina (0.253) slightly larger than the subapical carina (0.228); angle formed by the caudal margin concave. Frons (Fig. 15 B): approx. 1.3 times longer (0.8) than wide (0.6), and approx. 3.0 times wider medially than apically (0.2); anterior region of the frons narrows laterally and weakly concave apically. Frontoclypeal suture, convex, evanescent close to the lateral carinae. Postclypeus (Fig. 15 B): compressed laterally, median carina moderately developed. Anteclypeus (Fig. 15 B): median carina well developed from the second half distally of suturing anterior to the clypeus. Rostrum reaching or surpassing slightly the middle of the abdomen. Thorax. Pronotum (Fig. 15 A, C): submedian carinae weakly developed, evanescent behind the eyes; hind margin obtusely angled. Mesonotum (Fig. 15 A): Median carina well-developed, but evanescent distally; lateral carinae strongly developed. Tegmina (forewings) (Figs. 15 D): length 5.6 mm; hyaline with light gray spots, three spots on the basal cells distinguishable as follows: one on the Pcu + A 1 fork, between CuP and A 2; another spot occurs on C 5, between CuP and MP; a third larger and curved spot occurs over mc, rc and C 1, between Cua 1 and ScP + RA; three cross spot occur in pcc; r-m 1 and mcu- 1, occurring distally to the first MP fork; m-cu 1 occurs proximally the first MP fork; petiole in RP 2.2 + RP 2.3 weakly developed; simple tubercles in all veins; 12 apical cells; 7 subapical cells. Posterior legs. Hind tibia (Fig. 15 E): approximately 2.1 mm; with 6 lateral spines, 3 closest to the femur being small. 1 st tarsomere (Fig. 15 F): 7 apical teeth of approx. the same size, one lateral larger. 2 nd tarsomere (Fig. 15 F): 8 apical teeth, the outer 2 larger and the middle ones smaller; 3 thin setae, one separated by apical teeth without setae. Male terminalia. Pygofer (Figs. 16 A – C, G): bilaterally symmetrical; in lateral view medially larger; in ventral view, ventromedian process triangular, as wide as tall. Anal tube (Figs. 16 A – C; 17 A – B): weakly asymmetrical bilaterally; in lateral view, dorsal margin with two convexities, and one smaller concavity apically; ventral margin moderately curved; in dorsal view, wider at the distal margin than at the base; thick distal margin; epiproct triangular moderately developed, flattened dorsoventrally; paraproct long, approx. 3.0 times longer than epiproct, flattened dorsoventrally, wider below epiproct. Genital styles (Figs. 16 A – C, G; 17 C – E): in lateral view, wide in the apical region and narrow near the base; rounded distal margin apically. Aedeagus (Figs. 16 D – F; 17 F – H) tubular, asymmetrical. Shaft of the aedeagus with three spines; in right lateral view, with two movable spines, 1 st spine long and bifid (a), occurs apically on the shaft, towards to ventral region, with fork distant of the base; 2 nd spine well elongated (b), occurs apically towards below the flagellum; in left lateral view, 3 rd spine conspicuous (c), semicircular near the apex of the shaft. Aedeagal shaft with four ventral ridges, three rounded and one distinct; in right lateral, rounded ridge larger, and distinct ridge sharp (triangular), occurs medially on the shaft; in left lateral a moderately small and rounded ridge occurs; in ventral view with a small almost straight ridge. Flagellum, thin and without spines, wider, and apically curved. Female terminalia. Anal tube (Figs. 18 A, C): in lateral view, rounded and larger distally with lateral margin well developed; in dorsal view, basally excavated and apically swollen; in ventral view, medially carinated and laterally excavated; epiproct and paraproct flattened dorsoventrally and not developed as in the males; in dorsal view, paraproct weakly wide, apically straight, and laterally rounded. Gonocoxae VIII (Figs. 18 A – B): truncated, in caudal view smaller apically; in lateral view, lateral carinae evanescent near to the base. Ovipositor (Figs. 18 A-B): in lateral view, sabre-shaped, curved upwards dorsally, surpassing the anal tube.	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B887DC0528AE007CDFAF3FEE0.taxon	etymology	Etymology. The specific epithet minuta comes from Latin and means small or diminutive and refers to the reduced size of P. minuta sp. nov.	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B887DC0528AE007CDFAF3FEE0.taxon	diagnosis	Diagnosis. P. minuta sp. nov. can be distinguished from other species of the genus Pintalia mainly by components of the male terminalia, as aedeagus with a conspicuous spine, well curved - " C " shape- laterally on the apex of shaft of the aedeagus and three small ventral ridges medially.	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B887DC0528AE007CDFAF3FEE0.taxon	distribution	Distribution. BRA, MG; Itabirito municipality MP 8 cave, (Type locality), and Mariana municipality.	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B8879C05C8AE003F7FB95FC20.taxon	description	https: // zoobank. org / urn: lsid: zoobank. org: act: 353 DB 61 B- 3 B 06 - 4 AA 5 - A 5 C 4 - CA 8259 CB 29 FA (Figs. 20 A – F; 21 A – G; 22 A – H; 23 A – D; 24)	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B8879C05C8AE003F7FB95FC20.taxon	materials_examined	Type material. Holotype: Male Brazil, MG. Igarapé municipality, SAZ- 0001 - Cave, (UTM 575027 W, 7776185 S, 23 K), 24. vii. 2018, (Alves, J. P) (ISLA 100947). Holotype condition: not dissected, stored individually in a vial with ethanol 70 %. Paratypes. Same data as male holotype, except for 1 ♀ (ISLA 100948), 1 ♂ (dissected), 1 ♀ and 5 nymphs (ISLA 100949). Additional material examined. Brazil, MG: São Joaquim de Bicas, municipality, 2 ♂ 11 - cave, and 1 nymph (UTM 575687 W, 7776492 S, 23 K), 16. i. 2018, (Alves, J. P) (ISLA 100950); 1 ♀ 13 - Cave, (UTM 575602 W, 7776349 S, 23 K), 16. i. 2018, (Alves, J. P) (ISLA 100951); 1 ♀ 16 - Cave, (UTM 575654 W, 7776271 S, 23 K), 15. i. 2018, (Alves, J. P) (ISLA 100952); 2 ♀ 1 ♂ and 2 nymphs, 22 - Cave (UTM, 575821 W, 7776137 S, 23 K), 17. i. 2018, (Alves, J. P) (ISLA 100953); Doresopólis municipality, 1 ♂ 34 - Cave, (UTM 410938 W, 7750122 S, 23 K) 24. i. 2019, (ISLA 100954). Conceição do Mato dentro municipality, 1 ♂, css- 009 cave, (UTM 667151 W, 7905478 S, 23 K), 20 - 29. xi. 2018, (Carste et. al) (ISLA 100955); 1 ♂, SERP _ 0126 cave (UTM 762010 W, 7888377 S, 23 K), 23. xi. 2013 (Spelayon et al.) (ISLA 100956).	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B8879C05C8AE003F7FB95FC20.taxon	description	Description. Coloration (specimen preserved in 70 % ethanol). As in P. minuta sp. nov. except for pterostigma region and apical margin of the tegmina is clearer in pale orange yellow (73). Body length. Male. 2.9 (n = 2); Female. 4.0 – 4.5 (n = 2). Head. Vertex (Fig. 20 A, C): approximately 3.0 times wider (0.3) than long (0.1); apical compartment small and laterally narrow, approx. 2.2 times wider (0.2) than medially long (0.09); apical transverse carina (0.286) weakly larger than the subapical carina (0.251); angle formed by the caudal margin well concave and irregular. Frons (Fig. 20 B): 1.4 times longer (0.7) than wide (0.5), approx. 2.5 times wider medially than apically (0.2); anterior region of the frons moderately large laterally and weakly concave apically. Frontoclypeal suture semicircular convex, weakly straight laterally. Postclypeus (Fig. 20 B): median carina moderately developed, slightly wider than the base of the anteclypeus; Anteclypeus (Fig. 20 B): median carina moderately developed. Rostrum reaching or surpassing the middle of the abdomen. Thorax. Pronotum (Figs. 20 A – B): submedian carinae weakly developed; hind margin asymmetrical, obtusely angled, or little rectangular. Mesonotum (Fig. 20 A): median carina weakly developed and evanescent distally; lateral carinae well developed. Tegmina (forewings) (Fig. 20 D): length 4.3 mm; hyaline with light gray spots, as in P. minuta sp. nov.; fork of ScP + RA and RP, moderately ahead of forks CuA 1 and CuA 2; r-m 1 and mcu- 1 very close, proximally of the first MP fork; simple tubercles in all veins; usually with punctual anastomosis in RP 2.2 + RP 2.3; 12 apical cells rarely 9; 7 subapical cells rarely 8. Posterior legs. Hind tibia (Fig. 20 E): approximately 1.9 mm; with 7 lateral spines, the 2 closest to the femur being small. 1 st tarsomere (Fig. 20 F): 7 apical teeth of approx. the same size, one larger laterally. 2 nd tarsomere (Fig. 20 F): 5 - 8 apical teeth (asymmetric in some specimens), the outer 2 larger and the middle ones smaller; 3 thin setae, one seta, separated by apical teeth without seta. Male terminalia. Pygofer (Figs. 21 A – C, G): bilaterally asymmetric; in right lateral view, dorsocaudal margin angulated medially; dorsal margin concave; caudal margin convex; in left lateral view, dorsal and caudal margin completely linear, without angular process medially; in ventral view, ventromedian process triangular, slightly crooked / inclined to right lateral. Anal tube (Figs. 21 A – C; 22 A – B): bilaterally asymmetric; in dorsal view oblique, tilted to the right laterally and rounded medially; in lateral view, dorsal margin with two convexities, and one concavity apically; ventral margin weakly curved; in dorsal view, wider at the distal margin than at the base; epiproct in lateral view, flat dorsoventrally, in dorsal view slightly triangular with lateral margin weakly rounded; paraproct long and asymmetrical, flattened dorsoventrally, approx. 3.0 times larger than epiproct. Genital styles (Figs. 21 A – C, G; 22 C – E): bilaterally asymmetric; right genital style in right lateral view wider and rounded, left genital style in left lateral view elongated laterally. Aedeagus (Figs. 21 D – F; 22 F – H): tubular, asymmetrical. Shaft of the aedeagus with three movable spines; in right lateral view with two spines, 1 st spine bifid and elongated (a), with bifurcation moderately near the base; 2 nd spine moderately elongated and curved (b), occurs apically on the shaft, not reaching the middle of the flagellum; in left lateral view, 3 rd spine elongated and slightly curved (c), occurs near the apex of the shaft and surpassing the middle of the shaft; ventral ridge developed in right lateral on the shaft, with small rigid process dorsally. Flagellum, thin and without spines. Female terminalia (Figs. 23 A – C). As in P. minuta sp. nov.	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B8879C05C8AE003F7FB95FC20.taxon	etymology	Etymology. The specific epithet minima come from the Latin minimum and means the smallest of all and refers to the reduced size of males of P. minima sp. nov., the smallest among its congeners described here.	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B8879C05C8AE003F7FB95FC20.taxon	diagnosis	Diagnosis. P. minima sp. nov. can be distinguished from other species of the genus Pintalia by the spines of the aedeagus exhibiting the fork of bifid spine moderately distant to the base, and with the spine “ b ” moderately elongated and curved, and in left lateral with a single spine occurs on the apex of the shaft elongated surpassing the middle of the shaft.	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B8879C05C8AE003F7FB95FC20.taxon	distribution	Distribution. BRA, MG; Igarapé municipality, SAZ- 0001 cave, (Type locality).	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B8879C05C8AE003F7FB95FC20.taxon	discussion	Remarks. Specimens from additional material examined are very similar to the type-series, however with some differences. In 11 - cave the male specimens are smaller, approx. 2.7 mm, tegmina’s exhibit complete anastomosis in RA + RP 1, and sometimes between RP 4 + Cua 1; the aedeagus are like the type-series but much smaller. In cave 22 the male individual found as in the type-series. In 34 - cave the male specimen displays approx. 3.7 mm and the left lateral spine (c) large and robust facing the dorsal region of the aedeagus, and the bifid spine (a) exhibits the bifurcation moderately distant from the base. Individual females were found in caves 13 and 16, not associated with males but very similar to females of the type series. Two other specimens from two caves were removed from the material examined, as they exhibited a moderately distinct aedeagus. The two specimens were found in Conceição do Mato Dentro municipally and the individual collected in css- 009 cave exhibits on the right side of the aedeagal stem a spine moderately elongated and straight (b), and on the left side (c) a spine moderately small and curved, not surpassing the middle of the aedeagal shaft; and the individual found in Serp _ 0126 cave, exhibits the spines (b) (c) small and straight, and the bifid spine (a) with the bifurcation very close to the base. Although these variations can be interpreted as troglomorphisms in more specific cases and with more information, here, the number of samples per cavity is very small, which makes larger taxonomic advances for this species difficult. Here we propose that this species can be interpreted as eutroglophilic (Sket 2008), as it presents populations in different caves with different stages of redcuction of tegmina and body length. This classification, however, is tentative as we lack information on the biology and ecology of the species, on which Sket´s (2008) criteria are based.	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B8876C0588AE006D4FBA1F830.taxon	description	https: // zoobank. org / urn: lsid: zoobank. org: act: CAF 23604 - E 8 FC- 4508 - A 6 B 5 - 7 DF 4 D 98 D 6 CD 6 (Figs. 25 A – F; 26 A – G; 27 A – H; 28)	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B8876C0588AE006D4FBA1F830.taxon	materials_examined	Type material. Holotype: Male Brazil, MG. Presidente Olegário municipality, Lapa Vereda da Palha cave (UTM 380963 W, 7981204 S, 23 K), 13. x. 2010, (Ferreira, R. L.) (ISLA 100957). Holotype condition: not dissected, stored individually vials with ethanol 70 %. Paratypes. Same data as holotype, except for 1 ♂ (ISLA 100958) (dissected); 1 ♂ (ISLA 2761).	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B8876C0588AE006D4FBA1F830.taxon	description	Description. Coloration (specimen preserved in 70 % ethanol). As in P. minuta sp. nov., body generally strong orange yellow (68), contrasting with regions in deep yellowish brown (75) of the carinae, and brilliant orange yellow (67) between the lateral carinae of mesonotum. Head, legs, and abdomen ventrally in Light orange yellow (70). Tegmina hyaline, with very light spots in yellowish gray (93), pterostigma region is clearer in pale orange yellow (73). Body length. Male. 3.4 – 3.7 mm (n = 3). Head. Vertex (Fig. 25 A, C): approximately 2.0 times wider (0.4) than long (0.2); apical compartment small and laterally narrow, approx. 2.5 times wider (0.2) than medially long (0.08); apical transverse carina (0.279) slightly larger than the subapical carina (0.242); the angle formed by the caudal margin usually concave. Frons (Fig. 25 B): 1.5 times longer (0.9) than wide (0.6), approx. 3.0 times wider medially than apically (0.2); anterior region of the frons, narrow laterally and moderately concave apically. Frontoclypeal suture semicircular, well bent upwards in the middle and straight laterally. Postclypeus (Fig. 25 B): median carina weakly developed. Anteclypeus (Fig. 25 B): median carina moderately developed, more carinated from the second half distally suture anterior to the clypeus. Rostrum in ventral view surpassing the middle of the abdomen, almost reaching the base of the pygofer. Thorax. Pronotum (Figs. 25 A, C): submedian carina weakly developed, well evanescent behind the eyes; hind margin obtusely angled. Mesonotum (Fig. 25 A): median carina weakly developed or evanescent; lateral carinae strongly developed. Tegmina (forewings) (Figs. 25 D): length 6.2 mm; the tegmina spots in parts as in P. minuta sp. nov. and P. minima sp. nov.; fork of ScP + RA and RP, ahead of forks CuA 1 and CuA 2; r-m 1 occurring distally mcu- 1 and together the first MP fork; m-cu 1 occurs proximally the first MP fork; simple tubercles in all veins; punctual anastomosis RP 2.2 + RP 2.3; 12 apical cells; 7 subapical cells. Posterior legs. Hind tibia (Fig. 25 E): approximately 2.3 mm; with 4 lateral spines, the two closest to the femur being small. 1 st tarsomere (Fig. 25 F): 7 apical teeth of approx. the same size, one lateral larger. 2 nd tarsomere (Fig. 25 F): 8 apical teeth, the outer 2 larger and the middle ones smaller; 3 thin setae, one separated by apical teeth without setae. Male terminalia. Pygofer (Figs. 26 A – C, G): bilaterally asymmetric; in right lateral view, dorsocaudal margin angulated medially; dorsal margin concave; caudal margin straight; in left lateral view, dorsal and caudal margin completely linear, without angular process medially; in ventral view, ventromedian process triangular, rounded apically, well crooked to right lateral. Anal tube (Figs. 26 A – C; 27 A – C): bilaterally asymmetric; " crooked " to right lateral; ventrally excavated and elevated dorsally; in lateral view, larger medially than at the distal margin; distal margin thin and flattened dorsoventrally; in caudal view, epiproct long, tubular and pointed at the distal margin; in right lateral view, paraproct long, quadratic and flattened dorsoventrally, bent towards the base of the anal tube ventrally; distal margin triangular with a small process medially. Genital styles (Figs. 26 A – C, G; 27 D – F): bilaterally asymmetric; in right lateral view, right genital style with apical portion large and very rounded; in left lateral view, left genital style with apical portion smaller, slightly angulate. Aedeagus (Figs. 26 D – F; 27 G – I) tubular, asymmetrical. Shaft of the aedeagus exhibit three movable spines elongated and thin; in right lateral view with two long spines of approximately the same size, 1 st spine almost straight (a) occurs apically on the shaft; 2 nd spine curved (b), occurs apically on the shaft near the first spine and towards to flagellum; in left lateral view, 3 rd spine long and curved (c), occurs apically on the shaft, reaching the base of the shaft dorsally; ventral ridge large and without processes, occurs dorsally near base of shaft. Flagellum, narrow and without spines.	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B8876C0588AE006D4FBA1F830.taxon	etymology	Etymology. The specific epithet magnaepiprocti comes from the junction of the Latin magnus meaning large and epiproct structure of the anal tube, highly developed in males, thus magnaepiprocti would mean large epiproct, referring to the hypertrophic epiproct of this species.	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B8876C0588AE006D4FBA1F830.taxon	diagnosis	Diagnosis. P. magnaepiprocti sp. nov. can be distinguished from other species of the genus Pintalia mainly by components of the male terminalia, which is very asymmetrical with a distinct anal tube, where the epiproct is cylindrical and long and cone-shaped, and the paraproct is quadratic / large, flattened dorsally and bent to ventral region.	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B8876C0588AE006D4FBA1F830.taxon	distribution	Distribution. BRA, MG; Presidente Olegário municipality, Lapa Vereda da Palha cave (Type locality).	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B8876C0588AE006D4FBA1F830.taxon	discussion	Remarks. Pintalia magnaepiprocti sp. nov. is known only from the type locality, Lapa Vereda da Palha cave, Presidente Olegário municipality, MG, BRA. The caves in the region are widely studied and the troglobiont status of some species has been discussed recently (e. g. Salvador et al. 2021, Ferreira et al. 2023, Salvador et al. 2024). P. magnaepiprocti does not present evident troglomorphisms, but its morphology is discrepant from the other species of the genus and the group in which it was placed. In addition to the presumed endemism, the most evident characteristic that may indicate adaptations to the subterranean environment in the species is the reduction of the pigmented region in the compound eyes, which perhaps indicates a reduction of functional ommatidia, although this is only speculation. The species also exhibits the unique characteristic of a highly developed epiproct, not observed in any other fulgoromorph species, evidencing its morphological isolation in the group. In some cases, it has been observed that in the case of troglomorphisms, behavior may precede morphology (Christiansen, 1965; Howarth, 2019). P. magnaepiprocti sp. nov. also presents a marked degree of bilateral asymmetry throughout the body, a condition still little explored in cave invertebrates. Sket (2008) stated that there is no biological justification for linking the term " troglobiont " only to the morphology of an organism or to the absence of superficial / epigean ancestors, as there are several factors to be considered in addition to troglomorphy, such as special adaptations, for example: ecology, behavior, life history, between others. Howarth and Moldovan (2018) detailed other characteristics in addition to morphology that may indicate troglomorphisms. Le Cesne et al. 2024 suggests that all species of Cixiidae found inside caves, without specimens collected in the epigean environment, even without sampling efforts in this environment, or evident troglomorphism, should be classified as troglobites due to their life habits. This approach is risky and implies severe consequences in the classification of species or even in conservation laws for troglobitic species in countries such as Brazil. In Brazil, species that have an enigmatic way of life, and that depend on the subterranean environment to complete their entire life cycle, are protected by law. Therefore, in this work only P. magnaepiprocti sp. nov. can be preliminarily classified as a troglobiont.	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B8873C0458AE005B5FBFAFE70.taxon	description	https: // zoobank. org / urn: lsid: zoobank. org: act: C 9 EF 4 B 20 - 9 CAA- 4 B 83 - A 46 F- 3 BA 5110 DEAE 9 (Figs. 29 A – F; 30 A – H; 31 A – I; 32 A – D; 33)	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B8873C0458AE005B5FBFAFE70.taxon	materials_examined	Type material. Holotype: Male Brazil, MG. Arcos municipality, BRAS _ 010 cave (UTM 441818 W 7746510 S, 23 K), 14 - 20. xii. 2018, (Spelayon et al.) (ISLA 70225). Holotype condition: not dissected, stored in an individual vial in ethanol 70 %. Paratypes. Same data as female holotype, except for 2 ♂ (1 ♂ dissected), 1 ♀ and 1 nymph (Spelayon et al.) (ISLA 100959); ♀ s 2 _ CRH _ 279 Cave, (UTM 439790 W, 7751428 S, 23 K), 06. xi- 13. xii. 2018, (Spelayon et al.) (ISLA 68546). Additional material examined. Brazil, MG: Iguatama municipality, 1 ♀ JAT 47 cave (UTM 413560 W 7757019 S, 23 K), 03. xii. 2022, (Zampaulo R. A; Reis A. S.) (ISLA 126050); 1 ♀ JAT 36 cave (UTM 414769 W 7756433 S, 23 K), 25. xi. 2022, (Zampaulo R. A; Reis A. S.) (ISLA 126051).	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B8873C0458AE005B5FBFAFE70.taxon	description	Description. Coloration (specimen preserved in 70 % ethanol). Body generally deep yellowish brown (75), contrasting with regions in dark brown (59) on carinae, extremities and abdomen dorsally, and strong yellowish brown (74) on head and parts of mesonotum. Tegmina with some hyaline regions and colors ranging from light yellowish brown (76) to dark yellowish brown (78) which takes place mainly in a dark spot with two hyaline dots between the postclaval margin and anterior cubital area. Body length. Male. 3.0 – 3.7 mm (n = 3); Female. 3.9 – 4.0 mm (n = 2). Head. Vertex (Figs. 29 A, C): approximately 3.0 times wider (0.3) than long (0.1); apical compartment narrows and medially elongated, approx. 2.8 times wider (0.2) than medially long (0.07); apical transverse carina (0.294) slightly larger than the subapical carina (0.262); angle formed by the caudal margin, triangular. Frons (Fig. 29 B): 1.2 times longer (0.6) than wide (0.5), approx. 1.6 times wider medially than apically (0.3); median carina evanescent; anterior region of the frons large laterally, and moderately concave apically. Frontoclypeal suture semicircular, convex .. Postclypeus (Fig. 29 B): median carina weakly developed. Anteclypeus (Fig. 29 B): median carina moderately developed. Rostrum in ventral view reaching the hind coxae. Thorax. Pronotum (Figs. 29 A, C): submedian carinae weakly developed; hind margin obtusely angled. Mesonotum (Fig. 29 A): median carina well developed but evanescent distally; lateral carinae strongly developed and slightly closer near the pronotum than distally. Tegmina (forewings) (Figs. 29 D): length 5.2 mm; tegmina with some hyaline regions and colors ranging from Light yellowish brown to Dark yellowish brown and two round and hyaline patches on the CuA 1 and CuA 2 veins in the C 5 cell, and darker in this region; in addition, the tegmina exhibit long bristles mainly in CuP, Pcu, A 1 and A 2. fork of ScP + RA and RP, ahead of forks CuA 1 and CuA 2; C 5 approx. half the size of C 1; mcu- 1 closest to the basal cell that r-m 1; r-m 1 and m-cu 1 occurs ahead the first MP fork; simple tubercles in all veins; petiole in RP 2.2 + RP 2.3 moderately developed; 12 apical cells; 7 subapical cells. Posterior legs. Hind tibia (Fig. 29 E): approximately 1.3 mm; with 3 lateral spines, the first one closest to the small femur. 1 st tarsomere (Fig. 29 F): 7 apical teeth the outer and inner lateral teeth the largest, and the intermediate teeth with approx. the same size. 2 nd tarsomere (Fig. 29 F): 9 apical teeth, the 2 external ones bigger, and the intermediate ones smaller; 3 thin setae, one separated by apical teeth without setae. Male terminalia. Pygofer (Figs. 30 A – C, H): bilaterally slightly asymmetric; in right lateral view, dorsocaudal margin totally convex, wider medially; in left lateral view, dorsal margin irregular with two small concavity and one convexity medially; caudal margin straight; in ventral view, ventromedian process triangular, slightly wider than long. Anal tube (Figs. 30 A – C; 31 A – B): short and bilaterally slightly asymmetric; in lateral view ventroapically with two acute processes moderately elongate; in dorsal view epiproct moderately developed, with distal margin straight; paraproct long, with distal margin rounded. Genital styles (Figs. 30 A – C, H; 31 C – D): slightly asymmetric; in right lateral view, slightly elongated apically; in left lateral view, rounded apically. Aedeagus (Figs. 30 D – G; 31 F – I) tubular, asymmetric. Shaft of the aedeagus with two movable spines, one bifid; in left lateral view, without spines; in right lateral view with two wide spines, 1 st spine short, apically curved and flattened at the base (b), occurs apically on the shaft; 2 nd spine slightly larger, tubular and bifid (a) occurs near the apex of the shaft below the first spine; ventral ridge dorsally developed, with two rows of teeth towards the right lateral, almost reaching half the length of the shaft. Flagellum thin, without spines, and with a spine-like process flattened apically. Female terminalia. Anal tube (Figs. 25 A, C): very short; moderately elevated dorsally with two grooves pronounced laterally; epiproct and paraproct flattened dorsoventrally; in dorsal view, epiproct triangular, and rounded apically. Gonocoxae VIII (Figs. 25 A, B): truncated, in caudal view larger medially; in lateral view with lateral carinae well developed, almost straight. Ovipositor (Figs. 32 A-B): in lateral view, sabre-shaped curved upwards dorsally, slightly surpassing or no surpassing the anal tube.	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B8873C0458AE005B5FBFAFE70.taxon	etymology	Etymology. The specific epithet serratilis comes from Latin and means serrated and refers to the serrated ventral ridge on aedeagal shaft of the males.	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B8873C0458AE005B5FBFAFE70.taxon	diagnosis	Diagnosis. P. serratilis sp. nov. can be distinguished from other Pintalia species by its typical wing pattern characterised by two round and hyaline patches on the CuA 1 and CuA 2 veins in the C 5 cell, and by components of the male terminalia, such as short anal tube and developed paraproct (large); exhibiting two rows of teeth dorsally on base of the aedeagal shaft, and two wider spines apically, one of which bifid.	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B8873C0458AE005B5FBFAFE70.taxon	distribution	Distribution. BRA, MG; Arcos municipality, BRAS _ 010 cave (Type location).	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B886FC0438AE00719FB3FFCD8.taxon	description	https: // zoobank. org / urn: lsid: zoobank. org: act: A 8 EED 3 C 9 - 052 B- 4 E 40 - 8 F 51 - 66952 AB 1 D 260 (Figs. 34 A – F; 35 A – G; 36 A – H; 37 A – D; 38)	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B886FC0438AE00719FB3FFCD8.taxon	materials_examined	Type material. Holotype: Male, Brazil, MG. Prados municipality, SJDR _ HOL _ 028 cave, (UTM 597088 W, 7657227 S, 23 K), 17 - 19. xi. 2014, (Spelayon et al.) (ISLA 40189). Holotype condition: elements of terminalia and left tegmina dissected and stored in individual vials with 70 % ethanol. Paratypes. Same data as holotype, except for 1 ♀ SJDR _ HOL _ 015 Cave, (UTM 596994 W, 7657161 S, 23 K), (Spelayon et al.) (ISLA 40192); 1 ♀ SJDR _ HOL _ 025 Cave, (UTM 597005 W, 7657260 S, 23 K), (Spelayon et al.) (ISLA 40196); 1 ♀ SJDR _ HOL _ 007 Cave, (UTM 596995, 7657216, 23 K), (Spelayon et al.) (ISLA 40183 _. 1 ♀ SJDR _ HOL _ 016 Cave, (UTM 597015 W, 7657150 S, 23 K), (Spelayon et al.) (ISLA 40193).	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B886FC0438AE00719FB3FFCD8.taxon	description	Description. Coloration (specimen preserved in 70 % ethanol). Body generally deep orange (50), contrasting with regions in deep orange (51) on carinae, mesonotum, frons medially and abdomen dorsally, and pale orange yellow (73) on vertex, frons laterally and abdomen ventrally. Tegmina hyaline, with diffuse spots from light grayish yellowish brown (79) to dark grayish yellowish brown (81) from medial to distal region. Body length. Male. 3.0 mm (n = 1); Female. 2.8 – 3.8 mm (n = 4). Head. Vertex (Fig. 34 A, C): approximately 1.5 times wider (0.3) than long (0.2); apical compartment approx. 3.3 times wider (0.2) than medially long (0.06); apical transverse carina (0.291) larger than the subapical carina (0.242); angle formed by the caudal margin well triangular. Frons (Fig. 34 B): 1.4 times longer (0.7) than wide (0.5), approx. 1.6 times wider medially than apically (0.3); median carina with two lines side by side that separate near the frontal ocelli and in the anterior region of the frons; anterior region of the frons large and moderately concave apically. Frontoclypeal suture convex, and straight / inclined laterally. Postclypeus (Fig. 34 B): median carina moderately developed. Anteclypeus (Fig. 34 B): median carina well developed. Rostrum in ventral view surpassing slightly the base of the abdomen. Thorax. Pronotum (Figs. 34 A, C): submedian carina weakly developed; hind margin obtusely angled. Mesonotum (Figs. 34 A): median carina weakly developed and evanescent distally; lateral carinae moderately developed. Tegmina (forewings) (Figs. 34 D): length 4.9 mm; hyaline with light gray to dark brown spots, the shape of the spots and veins in parts as in P. pictipennis Stål 1862, see Remarks; fork of ScP + RA and RP, approx. the same height or a little forward of forks CuA 1 and CuA 2; parallel r-m 1 and m-cu 1 occurs distally of the first MP fork; simple tubercles in all veins; short subapical cells (C 1 b, C 2, C 3 a, C 3 and C 4); petiole in RP 2.2 + RP 2.3 weakly developed; 12 apical cells; 7 subapical cells. Posterior legs. Hind tibia (Figs. 34 E): approximately 1.4 mm; with 6 lateral spines, the four closest to the femur being small or inconspicuous. 1 st tarsomere (Fig. 34 F): 7 apical teeth of approx. the same size, one being larger lateral. 2 nd tarsomere (Fig. 34 F): 8 apical teeth, the 2 external ones larger, and the intermediate ones smaller; 3 thin setae, one separated by apical teeth without setae. Male terminalia. Pygofer (Figs. 35 A – C, G): bilaterally symmetric; in lateral view, dorsocaudal margin totally convex and larger medially; in ventral view, ventromedian process triangular, as wide as tall. Anal tube (Figs. 35 A – C; 36 A – B): short and bilaterally symmetric; in lateral view expanded and rounded distally, with two flat and rectangular lateral processes ventrally, dorsally flattened near the base and rounded on the distal margin; epiproct in lateral view, moderately developed, dorsally elevated / swollen; in dorsal view rounded apically; paraproct long and rounded dorsally with curved distal margin; in caudal view approx. with the same wide on the base and to apex. Genital styles (Figs. 35 A – B, G; 36 C – E): bilaterally asymmetric, in left lateral view short and rounded apically; in right lateral view also rounded but slightly elongated apically. Aedeagus (Figs. 35 D – F; 36 F – H) tubular, asymmetrical. Shaft of the aedeagus with three spines, two movable and one rigid; in right lateral view, 1 st spine wide (a), occurs apically on the shaft of the aedeagus towards the ventral region of the flagellum; in left lateral view, 2 nd spine small (b), occurs apically on the shaft; 3 rd spine small and rigid, as wide as long (c), occurs below the middle of the shaft; ventral ridge developed dorsally near the base of the shaft, with four conspicuous spines, rigid and short, two closest to the base being larger. Flagellum thin and curved, slightly larger than the shaft of the aedeagus with a small process apically. Female terminalia. Anal tube (Figs. 37 A – C): almost as wide as it long and wider towards the base; in dorsal view rounded medially and with lateral margin well developed, but evanescent apically; flat ventrally; epiproct and paraproct weakly developed. Epiproct with elongated base, slightly triangular apically. Paraproct rounded in dorsal view. Gonocoxae VIII (Figs. 37 A – B): truncated, in caudal view with lateral margin well developed and rounded. Ovipositor (Figs. 37 A-B): in lateral view, sabre-shaped, curved upwards dorsally, slightly surpassing or no surpassing the anal tube.	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B886FC0438AE00719FB3FFCD8.taxon	etymology	Etymology. The specific epithet stali is a tribute to the author Carl Stål, who made major contributions to the knowledge of the Fulgoromorpha and described the genus Pintalia Stål, 1862.	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B886FC0438AE00719FB3FFCD8.taxon	diagnosis	Differential diagnosis. Pintalia stali sp. nov. can be distinguished from most species of the genus Pintalia mainly by characters of the male terminalia, the anal tube is rounded and expanded distally, with two conspicuous processes flattened in the lateral / ventral region. Aedeagus displays four not-movable spines dorsally on the base of aedeagal shaft. However, this species can be confused with P. pictipennis Stal, 1862, due to its external morphology, although it exhibits conspicuous characters that allow differentiation, for example: P. stali sp. nov displays on the clavus of the tegmina only a small circular spot over the Pcu + A 1 bifurcation, while P. pictipennis displays a large spot that covers almost the entire clavus, and the first bifurcation of ScP + R is slightly in front of the first bifurcation of CuA in P. stali sp. nov., while it is moderately in front in P. pictipennis. Furthermore, the anal tube is also useful to differentiate these species, in P. stali sp. nov. it is almost tubular and approx. 3 x wider distally with a pair of flattened processes in the ventrolateral region, while in P. pictipennis it is completely flattened dorsoventrally slightly curved medially, without processes in the ventrolateral region.	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B886FC0438AE00719FB3FFCD8.taxon	distribution	Distribution. BRA, MG; Prados municipality, SJDR _ HOL _ 028 cave (Type locality).	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B886BC04A8AE000FFFB8CFD74.taxon	description	(Figs. 39 A – F; 40 A – G; 41 A – H; 42 A – D; 43)	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B886BC04A8AE000FFFB8CFD74.taxon	materials_examined	Type material. Holotype: ♂ Brazil, AM. (NHMUK- 013588867). Additional material examined. Brazil, MG: Sete Lagoas municipality, 1 ♂ CGP- 0024 Cave, (UTM 574138 W, 7848650 S 23 K), 04 - 13. xi. 2021, (Candiani, D. F. & Guarda, D. D.) (ISLA 103995); Santa Barbara municipality, 1 ♂ Gand _ 0073 Cave, (UTM 640305 W, 7783515 S 23 K), 18. iv. 2017, (Rabelo et al.) (ISLA 126055); Iguatama municipality, 1 ♀ JAT 39 Cave, (UTM 414595 W 7756476 S 23 K), 27. xi. 2022, (Zampaulo R. A.; Reis A. S.) (ISLA 126052); 2 ♂ and 1 ♀ JAT 42 Cave, (UTM 414519 W 7756592 S 23 K), 27. xi. 2022, (Zampaulo R. A.; Reis A. S.) (ISLA 126053); 1 ♂ JAT 43 Cave, (UTM 414496 W 7756551 S 23 K), 27. xi. 2022, (Zampaulo R. A.; Reis A. S.) (ISLA 126054); Pains municipality, 1 ♂ Gruta da Torre caida cave, (UTM 436818 W 7746966 S 23 K), 22. xi. 1999, (Ferreira, R. L).	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B886BC04A8AE000FFFB8CFD74.taxon	description	Description. Coloration (specimen preserved in 70 % ethanol). Body generally strong orange yellow (68), contrasting with regions in strong yellowish brown (74) of the carinae, deep orange yellow (69) on mesonotum, frons, and abdomen dorsally and brilliant orange yellow (67) on legs and abdomen ventrally. Tegmina Brilliant yellow (83) with a distal area hyaline, spot in deep yellow (85) with four hyaline points on the cubital, median and C 5 cells. Body length. Male. 4.0 – 4.3 mm (n = 2). Head. Vertex (Fig. 39 A, C): laterally wider, approx. 2.5 times wider (0.5) than medially long (0.2); apical compartment narrows and laterally larger, approx. 4.0 times wider (0.4) than medially long (0.1); apical transverse carina (0.413) slightly smaller than the subapical carina (0.437); angle formed by the caudal margin, concave irregular. Frons (Fig. 39 B): 1.4 times longer (1.0) than wide (0.7), approx. 1.7 times wider medially than apically (0.4); anterior region of the frons trapezoidal, wide and straight apically. Frontoclypeal suture large, very convex. Postclypeus (Fig. 39 B): very carinated. Anteclypeus (Fig. 39 B): with a distinct median carina apically, rounded close to the frons. Rostrum in ventral view reaching the middle of the abdomen. Thorax. Pronotum (Figs. 30 A, C): submedian carinae weakly developed, evanescent behind the eyes; hind margin obtusely angled. Mesonotum (Fig. 30 A): median carina well developed but evanescent distally; lateral carinae strongly developed. Tegmina (Forewings) (Fig. 39 D): length 7.0 mm; dark yellow with a dark spot and four hyaline spots; dark spot between basal vein RP, hyaline spots over the veins MP, CuP, CuA 1 and CuA 2; r-m 1 and m-cu 1 occurs together before to the first MP fork; Cells C 3 and C 5, C 4 narrow; small and simple tubercles in all veins; petiole in RP 2.2 + RP 2.3 well developed; 12 apical cells; 7 subapical cells. Posterior legs. Hind tibia (Fig. 39 E): approximately 2.3 mm; 6 apical spines; 4 lateral spines, the first being at the base of the tarsomere difficult to visualize; 1 st tarsomere (Fig. 39 F): 7 apical teeth, the two outer teeth and the two middle teeth larger. 2 nd tarsomere (Fig. 39 F): 7 apical teeth, the two outer ones being larger, and in the middle with approx. the same size; 3 platellae, one separated by apical teeth without platellae. Male terminalia. Pygofer (Figs. 40 E – G): bilaterally symmetric. In lateral view, dorsocaudal margin with cone-shaped process tilted up laterally, rounded apically; in ventral view ventromedian process triangular, longer than wider. Anal tube (Figs. 40 A – B, H; 41 A – B): moderately short; in dorsal view slightly elevated medially with lateral margins slightly curved; in lateral view the lateral margins form one convex line. Genital styles (Figs. 40 A – B, D; 41 C – E): thin near to the base and distally approx. 3.5 times longer than wide with rounded distal margin; near of base with a small triangular process ventrally. Aedeagus (Figs. 40 E – G; 41 F – I): asymmetric tubular. Shaft of the aedeagus with five thin spines, four movable; in right lateral view with four spines, three large and curved spines (1 st, 2 nd, 3 rd) (a) (b) (c), occurs on the apex of the shaft, towards the base; spine (a) in S-shapped; spine (b) straight, biger the others; spine (C) in hook-shaped; 4 th spine (d) straight, inserted little above of the center on the aedeagal shaft towards the base; in left lateral view, 5 th spine small (e), not movable, occurs near the apex of the shaft towards to the flagellum. Flagellum with a spine-like process laterally.	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B886BC04A8AE000FFFB8CFD74.taxon	diagnosis	Diagnosis. Pintalia constellaris can be distinguished from other Pintalia species mainly by the spines apically on the aedeagal shaft towards the base of the shaft, with spine (b) being the largest of these.	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B886BC04A8AE000FFFB8CFD74.taxon	distribution	Distribution. BRA, AM, Ega (Tefé) (Type Locality); MG, Sete Lagoas municipality, Santa Barbara municipality, Iguatama municipality, Pains municipality; BOL, La Paz, Tumupasa (Fennah, 1945).	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B886BC04A8AE000FFFB8CFD74.taxon	discussion	Remarks. I was able to see images of the type deposited in the Natural History Museum, London (NHMUK) by Walker (1858). Despite the current state of conservation of the type, the specimen is identical to the specimen used in the (re) description, except for some small variations in the tegmina, such as the size of the spots and the spine (b) of the aedeagus, which is larger and curved in the specimen used by Walker (1858).	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B8860C0488AE00615FCCCF82D.taxon	description	https: // zoobank. org / urn: lsid: zoobank. org: act: 6 CFB 997 A-A 2 CB- 4905 - A 37 A-D 8 D 2881 E 5 C 7 A (Figs. 44 A – F; 45 A – G; 46 A – H; 47)	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B8860C0488AE00615FCCCF82D.taxon	materials_examined	Type material. Holotype: Male Brazil, BA. Iuiu municipality, Lapa do Honorato Cave, (UTM 651726 W, 8400447 S, 23 K), 19. x. 2021 (Ferreira R. L.) (ISLA 126057). Paratypes. 1 ♂ 19. x. 2021 (Ferreira R. L.) (ISLA 100963); 1 ♀ Iuiu municipality, Lapa do Honorato Cave, (UTM 651726 W, 8400447 S, 23 K), 12. ix. 2016, (Cardoso R. C) (ISLA 62687); 2 ♀ 12. ix. 2016 (Ferreira R. L) (ISLA 100962); 2 ♂ 19. x. 2021 (Ferreira R. L.) (ISLA 100963).	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B8860C0488AE00615FCCCF82D.taxon	description	Description. Coloration (specimen preserved in 70 % ethanol). As in P. constellaris (Walker, 1858) except for narrow dark spot in pcc, over the ScP vein bordering the base of the pterostigma. Body length. Male. 4.0 – 4.5 mm (n = 2). Head. Vertex (Fig. 44 A, C): laterally wider, approx. 2.0 times wider (0.4) than medially long (0.2); apical compartment narrows and laterally larger, approx. 4.0 times wider (0.4) than medially long (0.1); apical transverse carina (0.380) slightly smaller than the subapical carina (0.395); angle formed by the caudal margin, concave and slightly irregular. Frons (Fig. 44 B): as in P. constellaris. Frontoclypeal suture, moderately convex, laterally straight. Post and Anteclypeus (Fig. 44 B): very carinated. Rostrum in ventral view reaching the middle of the abdomen. Thorax. Pronotum (Figs. 44 A, C): submedian carinae well developed, slightly evanescent / irregular behind the eyes; hind margin obtusely angled. Mesonotum (Fig. 44 A): median carina well developed, evanescent distally; lateral carinae strongly developed. Tegmina (Forewings) (Fig. 44 D): length 6.6 mm; dark yellow with a dark spot and four hyaline spots; narrow dark spot in pcc, over the ScP vein bordering the base of the pterostigma; r-m 1 occurs together to the first MP fork; m-cu 1 occurs after of the first MP fork in Mp 3 + 4; Cells C 3 and C 5 slightly elongated and rounded; small and simple tubercles in all veins; petiole in RP 2.2 + RP 2.3 well developed; 12 apical cells; 7 subapical cells. Posterior legs. Hind tibia (Fig. 44 E): approximately 2.4 mm; with 3 spines, the first being at the base of the tarsomere difficult to visualize. 1 st tarsomere (Fig. 44 F): 7 apical teeth, the two outer teeth and the two middle teeth larger. 2 nd tarsomere (Fig. 44 F): 8 apical teeth, the two outer ones being larger, and in the middle with approx. the same size; without platellae. Male terminalia. Pygofer (Figs. 45 E – G): bilaterally symmetric. In lateral view, dorsocaudal margin with cone-shaped process tilted up laterally, rounded apically; in ventral view, ventromedian process triangular, longer than wide, rounded apically and slightly compressed basally. Anal tube (Figs. 45 A – B, H; 46 A – B): as in P. constellaris, moderately short; in dorsal view slightly elevated medially with lateral margins slightly curved. Genital styles (Figs. 45 A – B, D; 46 C – E): thin near to the base and distally approx. 2.0 times longer than wide with rounded and curved distal margin; near of base with a small triangular process ventrally. Aedeagus (Figs. 45 E – G; 46 F – H): asymmetric tubular. Shaft of the aedeagus with five spines, four movable; in right lateral view with four spines, three large and robust spines, slightly curved (1 st, 2 nd, 3 rd) (a) (b) (c), occurs on the apex of the shaft, towards the base; 4 th spine straight (d), is slghtly bent medially, inserted little above of the center on the aedeagal shaft towards the base; in left lateral view, 5 th spine small (e), not movable, occurs near the middle of the shaft towards to the flagellum, with a basal ridge in “ u ”. Flagellum with a spine-like process laterally.	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B8860C0488AE00615FCCCF82D.taxon	etymology	Etymology. The specific epithet speciosa means " beautiful " or " showy " and refers to the fact that the new species display bright colors.	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B8860C0488AE00615FCCCF82D.taxon	diagnosis	Diagnosis. Pintalia speciosa sp. nov. can be distinguished from other Pintalia species mainly by the three robust spines laterally at the apex of the aedeagal shaft towards the base of the shaft, and by the unique arrangement of the two other spines in the aedeagal shaft. Additionally, the genital styles wide and elongated; the pygofer larger than wide apically.	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B8860C0488AE00615FCCCF82D.taxon	distribution	Distribution. BRA, BA; Iuiu municipality, Lapa do Honorato Cave (Type locality).	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B8860C0488AE00615FCCCF82D.taxon	diagnosis	Diferential diagnosis. Pintalia speciosa sp. nov. exhibits differences to P. constellaris, which are: P. speciosa sp. nov., exhibit 8 apical teeth on 2 nd tarsomere, without platellae (vs. 7 apical teeth on 2 nd tarsomere, with three platellae P. constellaris). P. speciosa sp. nov., exhibits three lateral spines on hind tibia (vs. exhibit four lateral spines on hind tibia in P. constellaris) The ventromedian process of the pygofer is distinct, longer than wide and apically rounded in u-shapped in P. speciosa sp. nov (vs. triangular in P. constellaris). The basal opening of the genital styles is also diamond-shaped but is slightly more elongated in P. speciosa sp. nov (vs. large with lateral margins almost straight in P. constellaris). The aedeagal shaft exhibits three spines apically on both species, however they are larger and robust, with more distant bases than in P. speciosa sp. nov (vs. thin with bases well proximate P. constellaris). In P. speciosa sp. nov. the aedeagal spine (a) is almost straight (vs. in S-shapped in right lateral view in P. constellaris). P. speciosa sp. nov. with adeagal spine (e) near the middle of the shaft with curved ridged basally (vs. with adeagal spine (e) near the apex of the shaft without curved ridged basally).	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B881CC0348AE0013FFE98F853.taxon	description	https: // zoobank. org / urn: lsid: zoobank. org: act: 4 B 61 EA 3 B-D 701 - 4 DC 1 - A 397 - 0 C 48 CD 199 F 39 (Figs. 48 A – F; 49 A – G; 50 A – H; 51 A – D; 52)	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B881CC0348AE0013FFE98F853.taxon	materials_examined	Type material. Holotype: Male Brazil, MG: Prudente de Morais municipality, Icmat- 85 (A and B) Cave, (UTM 593122 W, 7841934 S 23 K), 12. xii. 2017, (ISLA 100964) (Ativo ambiental et al.). Holotype condition: dissected, stored in an individual vial in 70 % ethanol. Paratypes. same data as male holotype, except for 3 ♂ (ISLA 108333); 1 ♀ (ISLA 100965); 2 nymphs with same data of male holotype except for (ISLA 100966). Additional material examined. Brazil, MG: Santa Barbara municipality, 3 ♂ LS 005 cave, (UTM 639693 E, 7781816 N 23 K), 04 - 05. x. 2021, (ISLA 108333) (Ativo ambiental et al.).	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B881CC0348AE0013FFE98F853.taxon	description	Description. Coloration (specimen preserved in 70 % ethanol). Body generally strong brown (55), contrasting with regions in dark brown (59) of the carinae and frons apically, deep brown (56) on mesonotum and abdomen dorsally and brilliant orange yellow (67) on vertex, frons medially, legs, abdomen ventrally. Tegmina Light grayish yellowish brown (79) with transversal and rounded spots in dark yellowish brown (78). Body length. Male. 4.4 mm (n = 1). Female. 5.7 (n = 1). Head. Vertex (Fig. 48 A, C): approximately 3.0 times wider (0.6) than long (0.2); apical compartment laterally wider, approx. 5.0 times wider (0.5) than medially long (0.1); apical transverse carina (0.520) evanescent and slightly smaller than the subapical carina (0.555); subapical carina slightly elongated medially; angle formed by caudal margin, concave. Frons (Fig. 48 B): swollen apically; 1.5 times longer (1.1) than wide (0.7), and approx. 1.7 times wider medially than apically (0.4); in frontal view, anterior region of the frons curved frontally towards from the vertex; median carina weakly developed, almost disappearing, apically separating into two, inner region with coloration of the apical compartment; frontoclypeal suture well convex, laterally straight. Post-and anteclypeus (Fig. 48 B): with median carina well developed. Rostrum in ventral view surpassing the hind coxae. Thorax. Pronotum (Figs. 48 A, C): submedian carinae weakly developed, evanescent behind the eyes; hind margin obtusely angled. Mesonotum (Fig. 48 A): median carina well developed but evanescent or absent distally; lateral carinae strongly developed. Tegmina (Forewings) (Fig. 48 D): length 6.9 mm; dark brown with three dark spots in the prenodal region between the R and A 1 veins and two in pcc cell; r-m 1 occurs together to the first MP fork; m-cu 1 occurs distally of the first MP fork; Cells C 3 and C 5 common, not elongated (vs. P. speciosa sp. nov. and P. constellaris (Walker, 1858 )); C 1 b small; C 5 a cell present; simple tubercles in all veins; petiole in RP 2.2 + RP 2.3 moderately developed; 12 apical cells; 7 subapical cells. Posterior leg. Hind tibia (Fig. 48 E): approximately 2.3 mm; with 3 lateral spines. 1 st tarsomere (Fig. 48 F): 7 apical teeth, the two outer teeth and two teeth in the middle larger. 2 nd tarsomere (Fig. 48 F): 7 apical teeth, the two outer ones being larger, and in the middle with approx. the same size; 3 platellae, one separated by apical teeth without platellae. Male terminalia. Pygofer (Figs. 49 A – C, G): bilaterally symmetrical; in lateral view, dorsocaudal margin with cone-shaped process sharp apically; in ventral view, ventromedian process wider than tall. Anal tube (Figs. 49 A – C; 50 A – B): as in P. constellaris, moderately short and flattened dorsoventrally; lateral margins weakly developed. Genital styles (Figs. 49 A – B, G; 50 C – E): long and well expanded apically, curved distal margin; two very small triangular processes occur ventrally near to the base. Aedeagus (Figs. 49 D – F; 50 F – H): asymmetric tubular. Shaft of the aedeagus exhibit five spines, being three curved spines (1 st, 2 nd, 3 rd) (a) (b) (c) apically on the shaft towards the flagellum; 4 th, and 5 th, spines (d) (e) short and straight, occur on opposite sides, slightly above the middle of the aedeagal shaft. Flagellum small, and without spines. Female terminalia. Anal tube (Figs. 51 A, C): moderately long; in lateral view, narrow and flat ventrally with lateral margin moderately developed, in ventral view carinated medially; epiproct and paraproct flattened dorsoventrally; in dorsal view, paraproct very small and rounded. Gonocoxae VIII (Figs. 51 A – B): in ventral view swollen medially; in caudal view smaller apically; in lateral view, lateral carinae evanescent near to the middle. Ovipositor (Figs. 51 A – B): in lateral view, sabre-shaped, curved upwards dorsally, slightly surpassing or not surpassing anal tube.	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B881CC0348AE0013FFE98F853.taxon	etymology	Etymology. The specific epithet fennahi is a tribute to the late hemipterist Ronald G. Fennah, who made major contributions to the genus Pintalia Stål, 1862.	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B881CC0348AE0013FFE98F853.taxon	diagnosis	Diagnosis. Pintalia fennahi sp. nov. can be distinguished from the other species of the Pintalia by the swollen anterior region of the vertex, with the evanescent apical transversal carina, tegmina dark brown with three dark spots in the prenodal region between the R and A 1 veins and two in pcc cell; cells C 3 and C 5 common, not elongated (vs. P. speciosa sp. nov. and P. constellaris). In addition, the array of five aedeagus spines.	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B881CC0348AE0013FFE98F853.taxon	distribution	Distribution. BRA, MG; Prudente de mores municipality, Cave Icmat- 85 (Type Locality), and Santa Barbara municipality.	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B8818C03F8AE000DFFE8DFE0C.taxon	description	https: // zoobank. org / urn: lsid: zoobank. org: act: 317 AF 43 F-E 19 D- 45 FD- 8 DC 7 - 05138 BBDA 7 E 5 (Figs. 53 A – F; 54 A – G; 55 A – I; 56 A – D; 57)	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B8818C03F8AE000DFFE8DFE0C.taxon	materials_examined	Type material. Holotype: Male. Brazil, MG. Pains municipality, AGR 030 Cave, (UTM 429594 W, 7752810 N, 23 K), 12 - 17. xii. 2015, (Spelayon et al.) (ISLA 51218); Holotype condition: not dissected, stored in an individual vial in ethanol 70 %. Paratypes. Same data as male holotype, except for 1 ♂ (dissected), Faz. Baú, MIC _ 002 Cave (UTM 437656 W, 7738366 N, 23 K), 10 - 11. xii. 2014, (Spelayon et al.) (ISLA 43260); 1 ♂ Cave gruta dos canudos (UTM 437021 W, 7746967 N 23 K), 10. x. 2000, (Ferreira R. L) (ISLA 100970); 1 ♂ Lapa do Tamboril cave, (Ferreira R. L) (ISLA 100972); 1 ♀ Divina cave, 20. ix. 2003 (Ferreira R. L) (ISLA 100971). Additional material examined. Brazil, MG: Prudente de Morais municipality, 1 ♂ and 1 ♀ ICMAT _ 0089 Cave, (UTM 592775 W, 7841320 N), 25. ix. 2018, (Ativo ambiental et al.) (ISLA 100967); 2 ♂ and 2 ♀ ICMAT- 71 Cave (UTM 593096 W 7841912 N), 27. xi. 2017, (Ativo ambiental et al.) (ISLA 100968). 3 ♂ Lagoa santa municipality, cave Lapa da Várzea (UTM 610200 W, 7835522 N, 23 K), 06. x. 2012 (Ferreira R. L) (ISLA 3536). 1 ♂ and 1 ♀, Arcos municipality, Arco do Índio cave (UTM 437129 W, 7753132 S, 23 K), 29. i. 2006, (Ferreira R. L) (ISLA 100969). 6 ♂ and 3 ♀, Curvelo municipality, Lapa do Saco Curto cave (UTM 562577 W 7935990 N), 17. i. 2015, (Rabelo, L. M) (ISLA 75685).	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B8818C03F8AE000DFFE8DFE0C.taxon	description	Description. Coloration (specimen preserved in 70 % ethanol). body mainly strong yellowish brown (74), contrasting with regions in deep yellowish brown (75) of the carinae, strong orange yellow (68) between lateral carinae of the mesonotum and some small dots of the vertex and frons medially. Tegmina hyaline with diffuse spots from moderate yellow (87) to in dark yellow (88). Body length. Males. 4.3 – 4.7 mm (n = 4). Females. 5.4 – 5.5 mm (n = 2). Head. Vertex (Fig. 53 A, C): approximately 1.3 times wider (0.4) than long (0.3); apical compartment moderately large, approx. 3.0 times wider (0.3) than medially long (0.1); apical transverse carina (0.331) smaller than the subapical carina (0.374); angle formed by caudal margin, triangular, moderately concave medially. Frons (Fig. 53 B): 1.5 times longer (1.1) than wide (0.7), and approx. 2.3 times wider medially than apically (0.3). frontoclypeal suture, moderately bent upwards medially, straight laterally. Postclypeus (Fig. 53 B): median carina weakly developed or evanescent. Anteclypeus (Fig. 53 B): median carina moderately developed, sometimes evanescent near frontoclypeal suture. Rostrum in ventral view surpassing slightly the base of the abdomen. Thorax. Pronotum (Figs. 53 A, C): Submedian carinae well developed, but slightly evanescent behind the eyes; hind margin straight obtusely angled. Mesonotum (Fig. 53 A): median carina well developed but evanescent or absent distally; lateral carinae strongly developed. Tegmina (Forewings) (Fig. 53 D): length 7.3 mm; hyaline with brown spots, two curved spots in pcc, three also curved spots of increasing size between fork A 1 / A 2 and SCP + R, four round spots in the post nodal region between RP 1 and MP 3, transversal sloping spot in the center of the tegmina connecting the SCP + RA fork and the posterior cubital area; r-m 1 and mcu- 1 occurring together, and proximally to the first MP fork; simple tubercles in all veins more visible on the A 1 / A 2 fork; petiole in RP 2.2 + RP 2.3 moderately developed; 12 apical cells; 7 subapical cells. Posterior legs. Hind tibia (Fig. 53 E): approximately 2.5 mm; with 4 lateral spines, 2 closest to the femur being small. 1 st tarsomere (Fig. 53 F): 7 apical teeth of approx. the same size, two external lateral larger. 2 nd tarsomere (Fig. 53 F): 8 apical teeth, the two external ones large (one larger) and the middle ones gradually smaller; 3 large platellae, one separated by apical teeth, but sometimes without platellae in one of the legs. Male terminalia. Pygofer (Figs. 54 A – C, G): bilaterally symmetric; in lateral view, dorsocaudal margin without processes; caudal margin slightly convex basally and slightly concave apically; in ventral view, ventromedian process triangular almost wider than tall; anal tube (Figs. 54 A – C, G; 55 A – C): tubular and distally developed, bent in 90 ° approx; in dorsal view, well narrow near to the base and broad distally; in lateral view, wider near the base; distal region moderately large and slightly asymmetric; in ventral view, distal margin large on left lateral, and slightly narrow medially. Genital styles (Figs. 54 A – B, G; 55 D – F): in lateral view, not much wider in the apical region than the base; ventral margin straight; dorsocaudal margin convex apically; dorsal margin with a small triangular process occurring before the bent medially. Aedeagus (Figs. 54 D – F; 55 G – I): asymmetrical tubular, with seven spines, two movable (a, d). Shaft of the aedeagus with four spines; in right lateral view, 1 st spine stout, curved (movable) (a), occurs apically on the shaft; 2 nd spine small (b), occurs near the center of the shaft towards the flagellum; 3 rd spine small (c), occurs on ventral region of the shaft; 4 th spine small, slightly curved (d), occurs apically on the left lateral of the shaft. Flagellum with three spines, all on the right margin; 5 th spine small (e), occurring at the base of the flagellum; 6 th spine slightly larger (f), occurring at the base of the flagellum, more ventrally; 7 th spine large (g), occurs apically on the flagellum. Female terminalia. Anal tube (Figs. 56 A, C): long; in lateral with lateral margin weakly developed; in dorsal view, slightly rounded; epiproct and paraproct flattened dorsoventrally; in dorsal view, paraproct very small and rounded. Gonocoxae VIII (Figs. 56 A – B). Ovipositor (Figs. 56 A – C): in lateral view, sabre-shaped, curved upwards dorsally, surpassing the anal tube.	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B8818C03F8AE000DFFE8DFE0C.taxon	etymology	Etymology. The specific epithet painensis refers to the type locality, municipality of Pains (MG), recognized by the large number of caves within its limits.	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B8818C03F8AE000DFFE8DFE0C.taxon	diagnosis	Diagnosis. Pintalia painensis sp. nov. can be distinguished from other Pintalia species by the occurrence of a robust and curved spine at the apex of the aedeagal shaft and the unique arrangement of other six spines in the aedeagus.	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B8818C03F8AE000DFFE8DFE0C.taxon	distribution	Distribution. BRA, MG; Pains, AGR 030 Cave (Type locality), Prudende de Morais and Lagoa da Prata municipalities.	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B8816C03B8AE005B5FB95FC04.taxon	description	https: // zoobank. org / urn: lsid: zoobank. org: act: 1 D 6520 B 0 - 487 F- 47 BA- 9 EA 4 - E 38 F 6 B 211 C 01 (Figs. 58 A – F; 59 A – G; 60 A – I; 61 A – D; 62)	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B8816C03B8AE005B5FB95FC04.taxon	materials_examined	Type material. Holotype: Male Brazil, MG: Dores de Guanhães, DGN _ 02 Cave, (UTM 721790 W 780121 S, 23 K), 31. viii. 2018, (Rabelo et al.) (ISLA 66226). Holotype condition: not dissected, stored preserved in a vial with 70 % ethanol. Paratypes: Same data of the male holotype, except for 1 ♂ (ISLA 75690). Additional material examined. Brazil, MG: Rio Piracicaba, 1 ♂ MONL _ 0006 Cave, (UTM 685543 E 7791350 N, 23 K), 28. Ii. 2023, (Ativo Ambiental et al.) (ISLA 114425); Rio Piracicaba, 2 ♂ and 1 ♀ AGL _ 0011 Cave, (UTM 687972 E 7794674 N, 23 K), 03 - 04. iii. 2021, (Ativo Ambiental et al.) (ISLA 102668); Santa Barbara, 1 ♂ and 1 ♀ CAP 1 _ 0009 Cave, (UTM 646028 E 7773028 N, 23 K), 09. iii. 2022, (Ativo Ambiental et al.) (ISLA 106997); Santa Barbara, 1 ♂ CAP 1 _ 0010 Cave, (UTM 646048 E 7772983 N, 23 K), 16. iii. 2022, (Ativo Ambiental et al.) (ISLA 106992); Mariana, 1 ♂ and 1 nymph FN _ 0001 Cave, (UTM 663923 E 7752548 N, 23 K), 04. xii. 2020, (Spelayon et al.) (ISLA 83502); João Monlevade, 1 ♂ ARC- 006 Cave, (UTM 692486 E 7810300 N, 23 K), 28. viii- 03. ix. 2022, (Spelayon et al.) (ISLA 112841).	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B8816C03B8AE005B5FB95FC04.taxon	description	Description. Coloration (preserved specimen): Body generally strong orange yellow (68), contrasting with some regions in deep orange yellow (69) and deep orange yellow (56) of the carinae, brilliant orange yellow (67) between lateral carinae of the mesonotum and some points of the vertex and frons medially. Tegmina hyaline with diffuse spots of moderate yellow (87) to dark yellow (88), pterostigma region is clearer in pale orange yellow (73). Body length. Male. 4.1 – 5.3 mm (n = 2). Head. Vertex (Fig. 58 A, C): approximately 2.5 times wider (0.5) than long (0.2); apical compartment moderately large, approx. 4.0 times wider (0.4) than medially long (0.1); apical transverse carina (0.358) smaller than the subapical carina (0.408); angled formed by caudal margin, moderately triangular and slightly concave laterally. Frons (Fig. 58 B): 1.5 times longer (1.1) than wide (0.7), and approx. 2.3 times wider medially than apically (0.3). Frontoclypeal suture, well bent upwards medially, straight laterally. Postclypeus (Fig. 58 B): median carina developed or moderately developed. Anteclypeus (Fig. 58 B): median carina moderately developed. Rostrum in ventral view surpassing slightly the base of the abdomen. Thorax. Pronotum (Figs. 58 A, C): submedian carinae weakly developed slightly irregular and evanescent behind the eyes; hind margin obtuse or obtusely angled. Mesonotum (Fig. 58 A): median carina well developed but evanescent or absent distally; lateral carinae strongly developed. Tegmina (Forewings) (Fig. 58 D): length 6.5 mm; hyaline with brown spots, two curved spots in pcc, two also curved spots of increasing size between fork A 1 / A 2 and SCP + R, one large round spots in the post nodal region between RP 1 and MP 4, transversal sloping spot developed in the center of the tegmina connecting the SCP + RA fork and the posterior cubital area; rm- 1 occurring near the first fork MP fork; mcu- 1 occurring distally of the first fork MP fork; simple tubercles in all veins more visible on the A 1 / A 2 fork; petiole in RP 2.2 + RP 2.3 moderately developed; 12 apical cells; 7 subapical cells. Posterior legs. Hind tibia (Fig. 58 E): approximately 2.2 mm; with 3 spines laterally, two in the middle of the hind tibia usually smaller. 1 st tarsomere (Fig. 58 F): 7 apical teeth, two external lateral larger, and two in the middle slightly smaller. 2 nd tarsomere (Fig. 58 F): 8 apical teeth, the two external ones large (one larger) and the middle ones gradually smaller; 3 large platellae, one separated by apical teeth without platellae. Male terminalia. Pygofer (Figs. 59 A – C, G): bilaterally symmetric, in lateral view, dorsocaudal margin with moderately large and rounded process medially bent inward to genital capsule; caudal margin slightly convex basally; in ventral view, ventromedian process triangular almost wider than tall. Anal tube (Figs. 59 A – C, G; 60 A – C): tubular and distally developed, bent in 90 ° approx; in dorsal view, well narrow and asymmetrical near to the base and broad distally; in lateral view, wider near the base; distal region moderately large and slightly asymmetric; in ventral view, distal margin large on left lateral, and slightly narrow medially; epiproct and paraproct, short. Genital styles (Figs. 59 A – B, G; 60 D – F): in lateral view, moderately wider apically and narrow near the base; ventral margin straight; dorsocaudal margin convex apically; dorsal margin with a very small triangular region occurring above the bent medially. Aedeagus (Figs. 59 D – F; 60 G – I): asymmetric tubular, with seven spines, three of which are movables (a, c, d). Shaft of the aedeagus with four spines; in right lateral view, 1 st spine longer and curved (a), occurs apically on the shaft; 2 nd spine small (b), near the center of the shaft towards the base; in left lateral view, 3 rd spine slightly larger (c), occurs apically on the shaft, towards the ventral region; 4 th spine slightly larger (d), approx. the same size as the third spine, towards the flagellum. Flagellum with three spines, all on the right margin and distally the middle of the flagellum; 5 th spine short and curved (e), occurring at the middle of the flagellum; 6 th spine short and straight (f), occurs slightly dorsally; 7 th spine slightly longer and curved (g), occurs apically on the flagellum.	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B8816C03B8AE005B5FB95FC04.taxon	etymology	Etymology. The specific epithet dorensis refers to the type locality, municipality of Dores de Guanhães (MG).	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B8816C03B8AE005B5FB95FC04.taxon	diagnosis	Diagnosis. Pintalia dorensis sp. nov. can be distinguished from other Pintalia species by exhibiting three spines in the medial / apical region of the aedeagal flagellum in addition to four other spines and by the pygofer which displays rounded and curved process laterally.	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B8816C03B8AE005B5FB95FC04.taxon	distribution	Distribution. BRA, MG; Dores de Guanhães, DGN 02 cave (Type locality), Rio Piracicaba municipality, Santa Barbara municipallty, Mariana municipality, João Monlevade municipallty.	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B8811C0398AE006E5FA86FC04.taxon	description	https: // zoobank. org / urn: lsid: zoobank. org: act: 6 BED 959 A-FA 52 - 414 B-BE 23 - 632 EB 2364 EA 7 (Figs. 63 A – F; 64 A – G; 65 A – I; 66 A – D; 67)	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B8811C0398AE006E5FA86FC04.taxon	materials_examined	Type material. Holotype: Male Brazil, MG. Lagoa Santa municipality, Helictites Cave (UTM 609077 W, 7836837 N, 23 K), 11. x. 2011, (Ferreira R. L) (ISLA 100973). Holotype condition: not dissected, stored in an individual vial in ethanol 70 %. Paratypes. Same data as male holotype, except for 1 ♀ Lapa da Varzea Cave, (UTM 610200 W, 7835522 N, 23 K), 06. x. 2012, (Ferreira R. L.) (ISLA 100974); 1 ♂ (dissected), G. Tuneis Cave (UTM 609040 W, 7836792 N, 23 K), 10. x. 2011, (ISLA 100975); 2 ♂ and 2 ♀ (ISLA 3536) (Rabelo et al.). Additional material examined. Brazil, MG: Itacarambi municipality, 1 ♂ Lapa d´Água do João Ferreira Cave, (UTM 593308 W, 8340410 N, 23 K), 25. i. 2015 (ISLA 100976) (Rabelo et al.).	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B8811C0398AE006E5FA86FC04.taxon	description	Description. Coloration (preserved specimen): As in P. dorensis sp. nov. Body length. Male. 4.0 – 4.8 mm (n = 5). Female. 5.0 – 5.1 mm (n = 2). Head. Vertex (Fig. 63 A, C): approximately 1.3 times wider (0.4) than long (0.3); apical compartment moderately large, approx. 3.0 times wider (0.3) than medially long (0.1); apical transverse carina (0.338), smaller than the subapical carina (0.372); angled formed by caudal margin, well triangular, laterally concave. Frons (Fig. 63 B): 1.5 times longer (1.1) than wide (0.7); approx. 2.3 times wider medially than apically (0.3); in frontal view, anterior region of the frons trapezoidal, slightly elevated medially. Frontoclypeal suture, weakly bent upwards medially, straight laterally. Postclypeus (Fig. 63 B): median carina weakly developed. Anteclypeus (Fig. 63 B): median carina moderately developed sometimes weakly developed. Rostrum in ventral view surpassing slightly the base of the abdomen. Thorax. Pronotum (Figs. 63 A, C): submedian carinae developed, slightly irregular and evanescent behind the eyes; hind margin obtusely angled. Mesonotum (Fig. 63 A): median carina weakly developed, evanescent or absent distally; lateral carinae well developed. Tegmina (Forewings) (Fig. 63 D): length 6.6 mm; hyaline with brown spots, two curved spots in pcc, three also curved spots of increasing size between fork A 1 / A 2 and SCP + R, the two near fork A 1 / A 2 connected, inconspicuous spot in the post nodal region between RP 1 and MP 4, transversal spot weakly developed in the center of the tegmina connecting the SCP + RA fork and the posterior cubital area; rm- 1 occurring together the first fork MP; mcu- 1 occurring distally of the MP fork; simple tubercles in all veins, slightly more visible on the A 1 / A 2 fork; petiole in RP 2.2 + RP 2.3 moderately developed; 12 apical cells; 7 subapical cells. Rostrum in ventral view surpassing slightly the base of the abdomen. Posterior legs. Hind tibia (Fig. 63 E): approximately 2.2 mm; with 3 spines laterally, first at the base of the tarsomere tiny. 1 st tarsomere (Fig. 63 F): 7 apical teeth, two external lateral larger and two in the middle slightly smaller. 2 nd tarsomere (Fig. 63 F): 8 apical teeth, the two external ones bigger (one larger) and the middle ones gradually smaller; 3 large platellae, one separated by apical teeth without platellae. Male terminalia. Pygofer (Figs. 64 A – C, G): bilaterally symmetric, in lateral view, dorsocaudal margin without process; caudal margin, straight; in ventral view, ventromedian process triangular, wider than tall. Anal tube (Figs. 64 A – C, G; 65 A – C): tubular, distally developed, bent in 90 ° approx; in dorsal view, lateral margins slightly asymmetric, weakly narrow near base and broad distally; in lateral view, wider near the base; distal region moderately large and asymmetric; in ventral view distal margin weakly concave and slightly narrow medially; epiproct and paraproct short. Genital styles (Figs. 64 A – B, G; 65 D – F): in lateral view, not much wider in the apical region than narrow near the base; ventral margin straight, slightly concave above the bent medially; dorsocaudal margin convex apically; dorsal margin with a moderately triangular region occurring above the bent medially. Aedeagus (Figs. 64 D – F; 65 G – I): asymmetric tubular, with five spines, two movable (a, d) and one bifid (d). Shaft of the aedeagus with three spines; in right lateral view, 1 st spine longer and curved (a), occurs on the right apical region of the shaft; 2 nd spine smaller (b), slightly curved towards the base, occurs near the center of the shaft; in left lateral view, 3 rd spine small (c), occurs in the apical region of the aedeagal shaft towards the base. Flagellum with two spines, 4 th spine bifid and longer (d), occurs near of the base of the flagellum towards to ventral region; 5 th spine short (e), occurs on the apical region of the flagellum. Female terminalia. Anal tube (Figs. 66 A, C): long; in lateral with lateral margin weakly developed; in dorsal view, slightly rounded; epiproct and paraproct flattened dorsoventrally; in dorsal view, paraproct, very small and rounded. Gonocoxae VIII (Figs. 66 A – B). Ovipositor (Figs. 66 A-B): in lateral view, sabre-shaped, curved upwards dorsally, strongly surpassing the anal tube.	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B8811C0398AE006E5FA86FC04.taxon	etymology	Etymology. The specific epithet lundi honors the late Danish researcher Peter Wilhelm Lund. P. Lund had a great influence on the understanding of the Brazilian Pleistocene period from an infinity of fossils discovered in Brazilian caves, being considered the precursor of the Speleology and Paleontology sciences in Brazil.	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B8811C0398AE006E5FA86FC04.taxon	diagnosis	Diagnosis. Pintalia lundi sp. nov. can be distinguished from other species of Pintalia mainly by the occurrence of a bifid spine at the base of the flagellum, facing the ventral region transversally and by the other four spines of the aedeagus.	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B8811C0398AE006E5FA86FC04.taxon	distribution	Distribution. BRA, MG; Lagoa Santa and Itacarambi municipality, Helictites Cave (Type Locality).	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B880DC0228AE00327FBD9FEE0.taxon	description	https: // zoobank. org / urn: lsid: zoobank. org: act: 6 B 33 DB 89 - E 692 - 4 A 28 - 93 CC- 40 CE 04 F 707 BF (Figs. 68 A – F; 69 A – G; 70 A – I; 71 A – D; 72)	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B880DC0228AE00327FBD9FEE0.taxon	materials_examined	Type material. Holotype: Male Brazil, MG: Montalvânia municipality, Fosséis Cave (UTM 573822 W, 8417268 N, 23 K), 28. xi. 2016, (Ferreira R. L), (ISLA 75691). Holotype condition: not dissected, stored in an individual vial in ethanol 70 %. Paratypes. Same data as male holotype, except for 1 ♀ (ISLA 100977); 2 ♂, 1 ♀ and 2 nymph’s (ISLA 100978).	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B880DC0228AE00327FBD9FEE0.taxon	description	Description. Coloration (preserved specimen): As on the P. dorensis sp. nov. Body length. Male. 4.0 – 4.5 mm (n = 3). Female. 5.4 – 5.8 mm (n = 2). Head. Vertex (Fig. 68 A, C): approximately 1.6 times wider (0.5) than long (0.3); apical compartment moderately large, approx. 3.0 times wider (0.3) than medially long (0.1); apical transverse carina (0.316) smaller than the subapical carina (0.360); angle formed by caudal margin, triangular, moderately concave medially. Frons (Fig. 68 B): 1.5 times longer (1.2) than wide (0.8), and approx. 2.6 times wider medially than apically (0.3). Frontoclypeal suture, moderately bent upwards medially, straight laterally. Anteclypeus (Fig. 68 B): median carina well developed; Postclypeus (Fig. 68 B): median carina moderately developed. Rostrum in ventral view surpassing slightly the base of the abdomen. Thorax. Pronotum (Fig. 68 A, C): submedian carinae developed but slightly irregular and evanescent behind the eyes; hind margin obtusely angled. Mesonotum (Fig. 68 A): median carina well developed, but evanescent or absent distally; lateral carinae well developed. Tegmina (Forewings) (Fig. 68 D): length 7.2 mm; hyaline with brown spots, two curved spots in pcc, three also curved spots of increasing size between fork A 1 / A 2 and SCP + R, the two near fork A 1 / A 2 connected, four rounded spot in the post nodal region between RP 1 and MP 4, the spot in RP region larger, the transversal sloping spot weakly developed in the center of the tegmina connecting the SCP + RA fork and the posterior cubital area; rm- 1 and mcu- 1 occurring distally of the first MP fork; simple tubercles in all veins, slightly more visible on the A 1 / A 2 fork; with punctual anastomosis in RP 2.2 + RP 2.3; 12 apical cells; 7 subapical cells. Posterior leg. Hind tibia (Fig. 68 E): approximately 2.4 mm; with 4 spines laterally, the first at the base of the tarsomere being very small. 1 st tarsomere (Fig. 68 F): 7 apical teeth, approx. the same size, two in the middle slightly larger. 2 nd tarsomere (Fig. 68 F): 8 apical teeth, two external ones slightly large (one larger) and the middle ones gradually smaller; 1 - 2 platellae. Male terminalia. Pygofer (Figs. 69 A – C, G): bilaterally symmetric; in lateral view, dorsocaudal margin without process; dorsal margin large; caudal margin, straight; in ventral view, ventromedian process triangular, wider than tall. Anal tube (Figs. 69 A – C, G; 70 A – C): tubular and distally developed, bent in 90 ° approx; in dorsal view weakly narrow near base and wide distally; in lateral view slightly with small convexity near to the base; distal region moderately large and asymmetric large, slightly larger on left lateral than the right lateral; in ventral view, with distal margin well concave; epiproct and paraproct short. Genital styles (Figs. 69 A – B, G; 70 D – F): in lateral view, moderately wider apically and narrow near the base; ventral margin straight; dorsocaudal margin convex apically; dorsal margin with a moderately triangular region occurring above the bent medially. Aedeagus (Figs. 69 D – F; 70 G – I): asymmetric tubular, with seven spines, only one movable (c). Shaft of the aedeagus with three spines; in right lateral view with two small spines, 1 st spine small (a), inserted on the apex of the aedeagal shaft towards the ventral region; 2 nd spine smaller and no movable (b), occurs near the center of the shaft; in left lateral view, 3 rd spine movable, long and curved (c), occurs in the apex of the shaft. Flagellum with four spines, all on the right margin; 4 th spine small (d), occurs at the base of the flagellum; on the apical region of the flagellum occurs three spines, 5 th and 6 th large spines occur together (e) (f); 7 th spine small (g), occurs slightly above on the apical region. Female terminalia. Anal tube (Figs. 71 A, C): long; in lateral with lateral margin weakly developed; in dorsal view, slightly rounded; epiproct and paraproct flattened dorsoventrally; in dorsal view, paraproct very small and rounded. Gonocoxae VIII (Figs. 71 A – B). Ovipositor (Figs. 71 A – B): in lateral view, sabre-shaped, curved upwards dorsally, strongly surpassing the anal tube.	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B880DC0228AE00327FBD9FEE0.taxon	etymology	Etymology. The specific epithet montalvaniensis refers to the type locality, municipality of Montalvânia (MG).	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B880DC0228AE00327FBD9FEE0.taxon	diagnosis	Diagnosis. Pintalia montalvaniensis sp. nov. can be distinguished from other species of the genus Pintalia by the three spines at the apex of the antennal flagellum, the middle being larger and more robust, and by exhibiting a single long and movable spine on the left side of the aedeagal shaft.	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B880DC0228AE00327FBD9FEE0.taxon	distribution	Distribution. BRA, MG; Montalvânia municipality, Fosséis cave (Type Locality).	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B8809C02F8AE00331FDFBF9CF.taxon	description	https: // zoobank. org / urn: lsid: zoobank. org: act: 346 DEA 9 A- 7 B 12 - 4949 - 828 C- 52 A 838079 C 4 A (Figs. 73 A – F; 74 A – G; 75 A – I; 76)	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B8809C02F8AE00331FDFBF9CF.taxon	materials_examined	Type material. Holotype: Male Brazil, MG. Dores de Guanhães municipality, G. Energia Cave-SPT 002 (UTM 718204 W, 7896396 N, 23 K), 11 - 12. xii. 2015, (Spelayon et al.) (ISLA 45532). Holotype condition: not dissected, stored in an individual vial containing ethanol 70 %. Paratypes. Same data as male holotype, except for 1 ♂ Cave- 08, (UTM 718137 W, 7890794 N, 23 K) 30. i. - 03 - ii. 2017, (Spelayon et al.) (ISLA 52291); 1 ♂ DGN 02 Cave, (UTM 721790 W, 7890121 N, 23 K), 31. x. 2018, (Rabelo et al.) (ISLA 75684). Additional material examined. Brazil, MG: São Sebastião do Maranhão municipality, 1 ♂ Boa vista Cave, (UTM 764786 W, 8012307 N, 23 K), 11. i. 2018, (Rabelo et al.) (ISLA 100979).	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B8809C02F8AE00331FDFBF9CF.taxon	description	Description. Coloration (specimen preserved in 70 % ethanol). As in P. painensis sp. nov. Body length. Male. 3.9 – 4.3 mm (n = 4). Head. Vertex (Fig. 74 A, C): approximately 2.0 times wider (0.4) than long (0.2); apical compartment, moderately large, approx. 3.0 times wider (0.3) than medially long (0.1); apical transverse carina (0.319) smaller than the subapical carina (0.354); angled formed by caudal margin well triangular, slightly concave laterally. Frons (Fig. 74 B): 1.8 times longer (1.1) than wide (0.6), and approx. 2.0 times wider medially than apically (0.3). Frontoclypeal suture slightly concave and weakly bent upwards. Anteclypeus (Fig. 74 B): with median carina moderately developed; Postclypeus (Fig. 74 B): with median carina moderately developed. Rostrum in ventral view surpassing slightly the base of the abdomen. Thorax. Pronotum (Figs. 74 A, C): submedian carinae weakly developed, slightly evanescent and irregular behind the eyes; hind margin obtuse, sometimes obtusely angled. Mesonotum (Fig. 74 A): median carina well developed, evanescent or sometimes absent distally; lateral carinae moderately developed. Tegmina (Forewings) (Fig. 74 D): length 6.1 mm, hyaline with brown spots, two curved spots in pcc, two also curved spots of increasing size between fork A 1 / A 2 and SCP + R, one inconspicuous spot in the post nodal region between RP 1 and MP 1.1, transversal sloping spot in the center of the tegmina connecting the SCP + RA fork and the posterior cubital area; rm- 1 occurring together the first fork MP; mcu- 1 occurring distally of the first MP fork; simple tubercles in all veins more visible on the A 1 / A 2 fork; petiole in RP 2.2 + RP 2.3 weakly developed; 12 apical cells; with 7 subapical cells, rarely 8 in one of the tegmina (Fig. 74 D). Posterior leg. Hind tibia (Figs. 74 E): approximately 2.1 mm; with 4 spines laterally, the first at the base of the tibia being very small. 1 st tarsomere (Figs. 74 F): 7 apical teeth, approx. the same size, but two in the middle slightly larger. 2 nd tarsomere (Figs. 74 F): 8 apical teeth, the two external ones slightly large (one larger) and the middle ones gradually smaller; 3 large platellae one separated by apical teeth without platellae. Male terminalia. Pygofer (Figs. 74 A – C, G): bilaterally symmetric; in lateral view, dorsocaudal margin without process; caudal margin large and straight; in ventral view, ventromedian process triangular, longer than wide. Anal tube (Figs. 74 A – C, G; 75 A – C): tubular, distally moderately short and bent at approx. 45 °; in dorsal view, moderately narrow near base, and wider distally; in lateral view that wide at base as distally; distal region small and slightly asymmetric; in ventral view, distal margin slightly concave and smaller than lateral margins; epiproct and paraproct short. Genital styles (Figs. 74 A – B, G; 75 D – F): in lateral view, weakly wider apically and narrow near the base; ventral margin straight, slightly concave above the bent medially; dorsocaudal margin convex apically; dorsal margin with a weakly expanded region occurring above the bent medially. Aedeagus (Figs. 74 E – F; 75 G – I): asymmetric tubular, with six spines, two movables (a, d). Shaft of the aedeagus with four spines; in right lateral view occurs three spines, 1 st spine well long and curved (a), inserting on the apical region of the shaft reaching the base; 2 nd spine moderately long (b) occurs almost apically; 3 rd spine slightly smaller (c), occurs almost de middle of the shaft; in left lateral view, 4 th spine moderatelly long (d), occurs almost apically on the shaft. Flagellum with two spines, all on the right margin; 5 th spine small (e), occurs at the base of the flagellum; 6 th spine slightly larger and curved (f), occurs on the apical region of the flagellum.	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B8809C02F8AE00331FDFBF9CF.taxon	etymology	Etymology. The specific epithet muiri is a tribute to British hemipterist Frederick Muir, who made major contributions to the genus Pintalia Stål, 1862.	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B8809C02F8AE00331FDFBF9CF.taxon	diagnosis	Diagnosis. Pintalia muiri sp. nov. can be distinguished from other species of the genus Pintalia by presenting a very long spine at the apex of the aedeagal shaft almost reaching the base, and just below it with two non-movable spines with close origins but in opposite directions. In addition, the distal region of the anal tube is moderately short and bent at approx. 45 °.	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B8809C02F8AE00331FDFBF9CF.taxon	distribution	Distribution. BRA, MG; Dores de Guanhães municiapllty G. Energia Cave-SPT 002 (Type Locality), and São Sebastião do Maranhão municipality.	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
0B2E630B8809C02F8AE00331FDFBF9CF.taxon	discussion	Remarks. The holotype exhibits right tegmina with MP 1 bifid (MP 1.1 and MP 1.2) and MP 3 + 4 trifid (MP 2, MP 3 + MP 4) as Fig. 74 D. The spine (d) was broken in many specimens, as in figure 74 - F, in a few specimens it was found complete as in figure 75 - I.	en	SANTOS, JÚLIO CÉSAR DO CARMO VAZ, HOCH, HANNELORE, BARTLETT, CHARLES R., FERREIRA, RODRIGO LOPES (2025): Revision of the type series of the original species placed in the genus Pintalia Stål, 1862 (Hemiptera: Fulgoromorpha: Cixiidae), and description of 12 new species from Brazilian caves. Zootaxa 5678 (1): 1-96, DOI: 10.11646/zootaxa.5678.1.1, URL: https://doi.org/10.11646/zootaxa.5678.1.1
