identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
63E9E7D332BE5983A0C696635E26470F.text	63E9E7D332BE5983A0C696635E26470F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Touranella Attems 1937	<div><p>Genus  Touranella Attems, 1937</p><p>Touranella Attems, 1937: 231 (D).</p><p>Touranella – Attems 1938: 233 (D); Hoffman 1963: 591 (K); 1980: 172 (M); Jeekel 1968: 64 (M); Golovatch 1994: 187 (D, M); 2009 a: 6 (M, K); 2009 b: 120 (M); 2016: 139 (M, K), 2024: 408 (D); Nguyen and Sierwald 2013: 1179 (M); Golovatch and Semenyuk 2010: 125 (M); 2018: 16 (M); Nguyen et al. 2023: 171 (D, K).</p><p>Diagnosis.</p><p>Body small to medium-sized (~ 9–25 mm long, ~ 0.7–3.3 mm wide), with 20 rings. Paraterga poorly to moderately developed. Sternal lobe or cone (s) present between ♂ coxae 4. First pair of ♂ without femoral tubercles (= adenostyles). Transverse metatergal sulci distinct. Legs with neither modifications nor adenostyles. Gonopod with a long and subcylindrical coxite, slightly curved caudally; prefemoral part (= prefemorite) short, ~ 3–4 × shorter than acropodite; femoral part (= femorite) strongly reduced or very short compared to solenophore, solenomere mostly rod-shaped or subflagelliform, sheathed distally by solenophore; both lamina medialis and lamina lateralis well developed; femoral process (fp) present or absent.</p><p>Type species.</p><p>Touranella gracilis Attems, 1937, by original designation.</p><p>Other species included.</p><p>Tournaella cattiensis Golovatch &amp; Semenyuk, 2010,  T. champasak Nguyen, Sierwald &amp; Ware, 2023,  T. chenla sp. nov.,  T. jaegeri sp. nov.,  T. himalayaensis Golovatch, 1994,  T. hirsuta Golovatch, 2009,  T. logunovi Golovatch, 2024,  T. moniliformis Golovatch &amp; Semenyuk, 2018,  T. peculiaris Golovatch, 2009,  T. pilosa Golovatch, 2016,  T. srisonchaii sp. nov., and  T. trichosa Golovatch &amp; Semenyuk, 2018 .</p><p>Remarks.</p><p>This genus was originally described as monotypic, based on  Touranella gracilis Attems, 1937, which was distinguished by its markedly reduced gonopodal femoral part, the presence of a femoral process, and densely setose metaterga. The recognition of  T. himalayaensis further supported these diagnostic features. However, subsequent discoveries of seven additional species in Vietnam have expanded the generic diagnosis, revealing further key variations such as the absence of a gonofemoral process and the presence of smooth metaterga (Golovatch 2009 a, 2009 b, 2016, 2024; Golovatch and Semenyuk 2010, 2018). More recently,  T. champasak from Laos was described by Nguyen et al. (2023), which included a brief discussion of the genus and its geographic distribution. The latest contribution has described another new species from southern Vietnam (Golovatch 2024). Below a species catalogue of  Touranella is presented, arranged in chronological order.</p></div>	https://treatment.plazi.org/id/63E9E7D332BE5983A0C696635E26470F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Likhitrakarn, Natdanai;Golovatch, Sergei I.;Inkhavilay, Khamla;Panha, Somsak;Sutcharit, Chirasak	Likhitrakarn, Natdanai, Golovatch, Sergei I., Inkhavilay, Khamla, Panha, Somsak, Sutcharit, Chirasak (2025): Taxonomic review of the millipede genus Touranella Attems, 1937, with a redescription of the type species, T. gracilis Attems, 1937, and descriptions of three new species from Laos (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 1238: 183-208, DOI: 10.3897/zookeys.1238.147550
37F8CDDD72245A50A5074F65203BA911.text	37F8CDDD72245A50A5074F65203BA911.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Touranella cattiensis Golovatch & Semenyuk 2010	<div><p>Touranella cattiensis Golovatch &amp; Semenyuk, 2010</p><p>Touranella cattiensis Golovatch &amp; Semenyuk, 2010: 123 (D).</p><p>Touranella cattiensis – Semenyuk et al. 2011: 40 (M); Golovatch et al. 2011: 81 (M); Nguyen and Sierwald 2013: 1180 (L); Golovatch 2016: 139 (M, K); Golovatch and Semenyuk 2018: 14 (M); Nguyen et al. 2023: 170 (M, K).</p><p>Remarks.</p><p>This species was initially documented in Nam Cat Tien National Park, 11°26'48.2"N, 107°26'26.2"E, 137 m a. s. l., inhabiting deciduous tropical forests dominated by  Dipterocarpus species along the riverside of Dongnai River in Dongnai Province, Vietnam. Apparently, this species is endemic to Vietnam (Golovatch and Semenyuk 2010).</p></div>	https://treatment.plazi.org/id/37F8CDDD72245A50A5074F65203BA911	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Likhitrakarn, Natdanai;Golovatch, Sergei I.;Inkhavilay, Khamla;Panha, Somsak;Sutcharit, Chirasak	Likhitrakarn, Natdanai, Golovatch, Sergei I., Inkhavilay, Khamla, Panha, Somsak, Sutcharit, Chirasak (2025): Taxonomic review of the millipede genus Touranella Attems, 1937, with a redescription of the type species, T. gracilis Attems, 1937, and descriptions of three new species from Laos (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 1238: 183-208, DOI: 10.3897/zookeys.1238.147550
BA312F6C380B50CD988FA029E1D45575.text	BA312F6C380B50CD988FA029E1D45575.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Touranella champasak Nguyen, Sierwald & Ware 2023	<div><p>Touranella champasak Nguyen, Sierwald &amp; Ware, 2023</p><p>Touranella champasak Nguyen et al., 2023: 171 (D, K).</p><p>Remarks.</p><p>This species has recently been described from Ban Thongvay (= Xekatam), 15°14.288'N, 106°31.891'E, 1,095 m a. s. l., located on the Bolaven Plateau, Champasak Province, Laos. This species represented the first record of the genus  Touranella from Laos, also reporting another species apparently endemic to Laos (Nguyen et al. 2023).</p></div>	https://treatment.plazi.org/id/BA312F6C380B50CD988FA029E1D45575	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Likhitrakarn, Natdanai;Golovatch, Sergei I.;Inkhavilay, Khamla;Panha, Somsak;Sutcharit, Chirasak	Likhitrakarn, Natdanai, Golovatch, Sergei I., Inkhavilay, Khamla, Panha, Somsak, Sutcharit, Chirasak (2025): Taxonomic review of the millipede genus Touranella Attems, 1937, with a redescription of the type species, T. gracilis Attems, 1937, and descriptions of three new species from Laos (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 1238: 183-208, DOI: 10.3897/zookeys.1238.147550
97B290B7A1395D3EBDE6D5AA21F7EFF7.text	97B290B7A1395D3EBDE6D5AA21F7EFF7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Touranella chenla Likhitrakarn 2025	<div><p>Touranella chenla Likhitrakarn sp. nov.</p><p>Figs 4, 5, 6</p><p>Type material.</p><p>Holotype • ♂ (CUMZ - PD 0031), Laos, Champasak Province, Pakse District, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.827446&amp;materialsCitation.latitude=14.848027" title="Search Plazi for locations around (long 105.827446/lat 14.848027)">Vat Phou</a>, 107 m, 14°50'52.9"N, 105°49'38.8"E, 23.7.2013, leg. Chirasak Sutcharit  .</p><p>Diagnosis.</p><p>The new species seems to be particularly similar to  T. champasak from Bolaven Plateau, Laos, especially in gonopodal structure. Both species share a strongly reduced gonopodal femorite devoid of a femoral process and a distally slightly twisted solenophore (Figs 5, 6). However, the new species differs from  T. champasak by having a long, acute process d located around the mid-length of a more clearly erect and thicker solenophore (Figs 5, 6). Additionally, pleurosternal carinae are complete crests with sharp caudal denticles extending beyond the rear tergal margin only on rings 2 and 3, reduced to a rounded caudal crest on rings 4 and 5, and absent thereafter (Fig. 4 C, E) (vs absent on rings 18–19). Moreover, ♂ tarsal brushes are present until ring 8 (vs until ring 16).</p><p>Description.</p><p>Length 15.2 mm, width of midbody pro- and metazona 1.1 and 1.3 mm, respectively (♂). Coloration of live animals dark brown (Fig. 4 A); legs pale orange, venter and a few basal podomeres pale brown to yellow brown. Coloration in alcohol after 11 years of preservation faded to pale brown; antennae and epiproct pale brown to pallid, venter and podomeres pallid (Fig. 4 B – J).</p><p>Clypeolabral region and vertex sparsely setose, epicranial suture distinct. Antennae long (Fig. 4 A), reaching until ring 5 when stretched dorsally. In width, ring 4 &lt;3 &lt;2 &lt;collum &lt;ring 5 &lt;6 &lt;7 &lt;head &lt;8–17, thereafter body gently and gradually tapering towards telson. Collum with three transverse rows of setae: 3 + 3 anterior, 2 + 2 intermediate, and 3 + 3 posterior; caudal corner very narrowly rounded, slightly declined ventrally, not produced past rear tergal margin (Fig. 4 A – C).</p><p>Tegument smooth and shining, prozona finely shagreened, metaterga nearly smooth, faintly rugulose and leathery (Fig. 4 B – F, H). Postcollum metaterga each with two transverse rows of setae: 2 + 2 anterior and 2 + 2 posterior, traceable at least as insertion points when setae broken off. Tergal setae simple, slender, short, ~ 1 / 3 metatergal length (Fig. 4 F). Axial line well visible on metazona, traceable also on prozona.</p><p>Paraterga weak (Fig. 4 A – F, H), lying at ~ 1 / 2 midbody height, slightly upturned, anterior edge rounded and narrowly bordered; caudal corner very narrowly rounded on rings 2–4, not produced past rear tergal margin; almost missing on following rings; pore-bearing rings with evident lateral bulges (Fig. 4 B, D). Ozopores evident, lateral, each lying in an ovoid groove at ~ 1 / 3 metatergal length in front of posterior edge of metaterga (Fig. 4 C, E, H). Transverse metatergal sulci complete on rings 5–16, incomplete on ring 17, very narrow, shallow, not reaching the bases of paraterga, at most faintly beaded at bottom (Fig. 4 B – F, H). Stricture between pro- and metazona wide, deep, clearly ribbed at bottom down to base of paraterga (Fig. 4 B – F, H). Pleurosternal carinae complete crests with a very sharp median tooth on rings 2 and 3, reduced to a rounded caudal crest on rings 4 and 5, thereafter missing (Fig. 4 C, E, H).</p><p>Epiproct (Fig. 4 F – H) conical, flattened dorsoventrally, with two small, rounded, apical papillae; tip subtruncate; lateral pre-apical papillae very small, lying close to tip. Hypoproct roundly subtriangular, setigerous knobs at caudal edge small and well-separated (Fig. 4 G).</p><p>Sterna sparsely setose, without modifications; an entire, high, rounded, linguiform, setose sternal lobe between ♂ coxae 4 (Fig. 4 I, J). Legs long, midbody legs 1.2–1.4 × as long as body height, prefemora without modifications, ♂ tarsal brushes present until ring 8.</p><p>Gonopods simple, slim and suberect (Figs 5, 6). Coxite slightly curved caudad, densely setose distodorsally (Figs 5 A, B, 6 A, B, D). Prefemoral part (= prefemorite, pfe) as usual, densely setose, ~ 3.5 × shorter than acropodite (femoral + postfemoral parts) (Figs 5 A, B, 6 A – C). Femorite (fe) short, with a strong, long, flagelliform solenomere (sl) twisted distad and with a strong, oblique, lateral sulcus demarcating a postfemoral part (Figs 5 A, B, 6 A – C). Solenophore (sph) long, slightly twisted distad, sheathing most of solenomere (sl), slightly curved caudad (Figs 5 A, B, 6 A – D). Solenophore consisting of a well-developed lamina lateralis (ll) and a smaller lamina medialis (lm) (Figs 5 B, 6 B, C). Tip of lamina medialis subtruncate (Figs 5 B, 6 B), tip of lamina lateralis subrectangular with a long, slender, curved, nearly pointed process d (d), this rising distal to lamina lateralis (Figs 5 A, B, 6 A – D). Both supporting a long and flagelliform solenomere.</p><p>Distribution.</p><p>Known only from the type locality, apparently endemic to the southern part of Laos.</p><p>Etymology.</p><p>The species epithet chenla refers to the ancient kingdom of Chenla, a powerful kingdom that thrived from approximately 550 to 802 AD across the present-day territories of Thailand, Laos, Cambodia, and Vietnam. The naming also reflects the historical connection between the Chenla era and the construction of the Vat Phou temple complex in Champasak Province, Laos, where the species was discovered (Wikipedia 2025 a, b). This new species so named pays tribute to the rich cultural and historical heritage of the region.</p><p>Remarks.</p><p>The newly described species was discovered in the vicinity of Vat Phou, Champasak Province, Laos. This ruined Khmer Hindu temple complex, recognized as a UNESCO World Heritage Site, is situated amidst the lush landscapes of Southeast Asia, providing an intriguing backdrop for biodiversity exploration.</p></div>	https://treatment.plazi.org/id/97B290B7A1395D3EBDE6D5AA21F7EFF7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Likhitrakarn, Natdanai;Golovatch, Sergei I.;Inkhavilay, Khamla;Panha, Somsak;Sutcharit, Chirasak	Likhitrakarn, Natdanai, Golovatch, Sergei I., Inkhavilay, Khamla, Panha, Somsak, Sutcharit, Chirasak (2025): Taxonomic review of the millipede genus Touranella Attems, 1937, with a redescription of the type species, T. gracilis Attems, 1937, and descriptions of three new species from Laos (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 1238: 183-208, DOI: 10.3897/zookeys.1238.147550
AF25E3555C7B5FBDAFF73435864ECA48.text	AF25E3555C7B5FBDAFF73435864ECA48.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Touranella gracilis Attems 1937	<div><p>Touranella gracilis Attems, 1937</p><p>Figs 2, 3</p><p>Touranella gracilis Attems 1937: 231 (D).</p><p>Touranella gracilis – Attems 1938: 233 (D); Jeekel 1968: 64 (M); Golovatch 1983: 182 (M); 1994: 186 (D, M); 2009 a: 6 (M, K); 2009 b: 120 (M); 2016: 139 (M, K); Enghoff et al. (2004): 40 (L); Golovatch and Semenyuk 2010: 125 (M); 2018: 16 (M); Nguyen and Sierwald 2013: 1180 (L); Nguyen et al. 2023: 170 (M, K).</p><p>Type material examined.</p><p>Holotype. • ♂ (NHMW-3531), Vietnam, Danang Province,  Lienchieu, 1931, leg. C. Dawydoff.  Paratype. • ♀ (NHMW-3531), same locality, together with holotype .</p><p>Redescription.</p><p>Length 12.5 (♂) or 14.4 mm (♀), width of midbody pro- and metazona 0.73 and 0.97 mm (♂) or 1.17 and 1.47 mm (♀), respectively.</p><p>Coloration of alcohol material after long-term preservation dark red brown (Fig. 2 A – G, J – L) with pale castaneous brown paraterga and epiproct, yellowish antennae, venter, and legs (Fig. 2 A – L) (vs body color almost black with pale yellowish antennae and legs, as given in the descriptions by Attems 1937).</p><p>Clypeolabral region and vertex sparsely setose, epicranial suture distinct. Antennae rather short, reaching to body ring 3 (♂) or ring 2 (♀) when stretched dorsally. In width, ring 2 &lt;3 &lt;collum &lt;head &lt;ring 4 &lt;5–16 (♂, ♀), thereafter body gently and gradually tapering towards telson. Collum with abundant, small, setiferous knobs; anterior edge broadly rounded, narrowly bordered, fused to callus; caudal corner very broadly rounded, not produced past the rear tergal margin; lateral edge with two-minute incisions, slightly declined ventrad (Fig. 2 A, B, J).</p><p>Tegument smooth and shining, prozona very finely shagreened, metaterga roughly granulate-tuberculate and shagreened; surface below paraterga finely microgranulate (Fig. 2 A – F, J – L). Postcollum metaterga with abundant, small, irregular, setiferous knobs. Tergal setae long, slender, ~ 1 / 2 length of metaterga. Axial line rather faint but traceable both on pro- and metaterga. Paraterga strongly developed (Fig. 2 A – F, J – L), lying low (at ~ 1 / 2–1 / 3 midbody height); anterior edge narrowly rounded and narrowly bordered, fused to callus; caudal corner narrowly rounded to nearly pointed, not produced past the rear tergal margin; lateral edge with three evident, lateral, setigerous incisions on poreless calluses and two strong ones (anterior) on pore-bearing calluses. Calluses on paraterga delimited by a sulcus only on dorsally. Ozopores evident, lateral, each lying in an ovoid groove at ~ 1 / 4 in front of caudal corner. Transverse metatergal sulci usually distinct (Fig. 2 A, C, F, K, L), incomplete on ring 4, complete on metaterga 5–18, line-shaped, very deep, not reaching bases of paraterga, faintly ribbed at bottom. Stricture between pro- and metazona wide, line-shaped, rather deep, evidently ribbed at bottom down to base of paraterga (Fig. 2 A – F, K, L). Pleurosternal carinae with an anterior narrowly rounded crest on ring 2, thereafter missing.</p><p>Epiproct (Fig. 2 E, G) conical, flattened dorsoventrally, with two-minute apical papillae; tip subtruncate; pre-apical papillae absent. Hypoproct subtriangular, setiferous knobs at caudal edge well-separated and evident (Fig. 2 G).</p><p>Sterna sparsely setose, without modifications; an entire, short, linguiform, setose sternal lobe between ♂ coxae 4 (Fig. 2 H, I). A paramedian pair of evident tubercles in front of gonopods aperture. Legs moderately long and slender, slightly incrassate in ♂, midbody legs ~ 1.3–1.4 (♂) or 1.0–1.1 × (♀) as long as body height, prefemora without modifications, ♂ tarsal brushes absent.</p><p>Gonopods simple and stout (Fig. 3 A, B); coxa slightly curving caudally, densely setose distodorsally. Prefemoral part (pfe) densely setose, ~ 1 / 4 the length of acropodite (femoral part + postfemoral part) (Fig. 3 A, B). Femoral part (fe) strongly reduced, with a prominent, simple, lance-shaped lateral prefemoral process (fp) (Fig. 3 B) and a fully medial, strong, long, flagelliform solenomere (sl) (Fig. 3 A, B). Solenophore (sph) suberect, slightly twisted distally, strongly developed, sheathing most of solenomere (sl), with a lateral shoulder (u) at the base, distally bearing a pointed process (e) and a narrow unciform lamona (g) (Fig. 3 A, B).</p><p>Distribution.</p><p>It is known only from the type locality and appears to be endemic to Vietnam.</p><p>Remarks.</p><p>Although Golovatch (1994) examined and illustrated the gonopodal structure of this species, nearly all other morphological features remained ignored. The present study provides a comprehensive description of the species which appears to be endemic to Vietnam.</p></div>	https://treatment.plazi.org/id/AF25E3555C7B5FBDAFF73435864ECA48	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Likhitrakarn, Natdanai;Golovatch, Sergei I.;Inkhavilay, Khamla;Panha, Somsak;Sutcharit, Chirasak	Likhitrakarn, Natdanai, Golovatch, Sergei I., Inkhavilay, Khamla, Panha, Somsak, Sutcharit, Chirasak (2025): Taxonomic review of the millipede genus Touranella Attems, 1937, with a redescription of the type species, T. gracilis Attems, 1937, and descriptions of three new species from Laos (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 1238: 183-208, DOI: 10.3897/zookeys.1238.147550
6833D6E629DD5BE1A70EC7110CE07A75.text	6833D6E629DD5BE1A70EC7110CE07A75.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Touranella himalayaensis Golovatch 1994	<div><p>Touranella himalayaensis Golovatch, 1994</p><p>Touranella himalayaensis Golovatch, 1994: 186 (D).</p><p>Touranella himalayaensis – Golovatch 2009 a: 6 (M, K); 2009 b: 120 (M); 2016: 139 (M, K); Golovatch and Semenyuk 2010: 125 (M); 2018: 16 (M); Nguyen and Sierwald 2013: 1180 (L); Nguyen et al. 2023: 170 (M, K).</p><p>Remarks.</p><p>The original description is based on specimens collected from Paniporua and Dhorpar Kharka, 2,300 –2,700 m a. s. l., in mixed broadleaved or  Rhododendron -  Lithocarpus forests in Panchthar District, Nepal. The species is considered endemic to Nepal (Golovatch 1994).</p></div>	https://treatment.plazi.org/id/6833D6E629DD5BE1A70EC7110CE07A75	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Likhitrakarn, Natdanai;Golovatch, Sergei I.;Inkhavilay, Khamla;Panha, Somsak;Sutcharit, Chirasak	Likhitrakarn, Natdanai, Golovatch, Sergei I., Inkhavilay, Khamla, Panha, Somsak, Sutcharit, Chirasak (2025): Taxonomic review of the millipede genus Touranella Attems, 1937, with a redescription of the type species, T. gracilis Attems, 1937, and descriptions of three new species from Laos (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 1238: 183-208, DOI: 10.3897/zookeys.1238.147550
9066EEE767CD5401A3E3610205B03437.text	9066EEE767CD5401A3E3610205B03437.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Touranella hirsuta Golovatch 2009	<div><p>Touranella hirsuta Golovatch, 2009</p><p>Touranella hirsuta Golovatch, 2009 b: 119 (D, M).</p><p>Touranella hirsuta – Golovatch and Semenyuk 2010: 123 (M); 2018: 16 (M); Nguyen and Sierwald 2013: 1180 (L); Golovatch 2016: 139 (M, K), 2024: 408 (D, R); Nguyen et al. 2023: 170 (M, K).</p><p>Remarks.</p><p>This species was also described from Nui Ba Nature Reserve, 12°10'N, 108°40'E, 1,400 –1,900 m a. s. l., near Lang Lanh, Bi Doup, Lam Dong Province, Vietnam. Apparently, the species is endemic to Vietnam (Golovatch 2009 b, 2024).</p></div>	https://treatment.plazi.org/id/9066EEE767CD5401A3E3610205B03437	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Likhitrakarn, Natdanai;Golovatch, Sergei I.;Inkhavilay, Khamla;Panha, Somsak;Sutcharit, Chirasak	Likhitrakarn, Natdanai, Golovatch, Sergei I., Inkhavilay, Khamla, Panha, Somsak, Sutcharit, Chirasak (2025): Taxonomic review of the millipede genus Touranella Attems, 1937, with a redescription of the type species, T. gracilis Attems, 1937, and descriptions of three new species from Laos (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 1238: 183-208, DOI: 10.3897/zookeys.1238.147550
2763A5D63EF95FD1A24F10D263463331.text	2763A5D63EF95FD1A24F10D263463331.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Touranella jaegeri Likhitrakarn 2025	<div><p>Touranella jaegeri Likhitrakarn sp. nov.</p><p>Figs 10, 11, 12</p><p>Type material.</p><p>Holotype • ♂ (SMF -SM-01), Laos, Bolikhamsay Province, Lak Sao, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=104.81275&amp;materialsCitation.latitude=18.22114" title="Search Plazi for locations around (long 104.81275/lat 18.22114)">Tham Mankhone</a>, 501 m, 18°13'16.1"N, 104°48'45.9"E, 21. 7. 2016, leg. P. Jäger  .  Paratypes: • 1 ♂ (SMF -SM-01), same locality, together with holotype .</p><p>Diagnosis.</p><p>The new species closely resembles both  T. chenla sp. nov. and  T. champasak, especially in its moniliform body with significantly reduced paraterga (Fig. 4 A – H, 7 A – H). However, it differs from  T. chenla sp. nov. by possessing a broader and better expanded apical process d, which arises proximal to the lamina lateralis (Figs 11, 12) (vs a slender, long, curved, apical process d that rises distal to the lamina lateralis (Figs 5, 6)), coupled with a larger size, measuring 21.4–22.6 mm long (vs smaller, 15.2 mm long). In comparison to  T. champasak, this species has pleurosternal carinae with complete crests and sharp caudal teeth on rings 2–4, and absent from ring 5 (vs present until ring 17 and absent thereafter). ♂ legs are 1.3–1.5 × as long as midbody height (vs longer, 1.7–1.8 ×), and ♂ tarsal brushes present until ring 10 (vs until ring 16).</p><p>Description.</p><p>Length 21.4–22.6 (♂), width of midbody pro- and metazona 2.6–2.8 and 3.2–3.3 mm (♂), respectively. Coloration of live animals dark brown-orange (Fig. 10 A), with a contrasting longitudinal pale yellow-orange stripe, this being narrow on prozona, but characteristically bead-shaped and broadened on metazona; paraterga, epiproct, and venter pale orange to pale yellow; head and antennae dark brown; legs contrasting pale orange, a few basal podomeres pale yellow to pallid. Coloration in alcohol after eight years of preservation faded to dark brown; antennae dark brown and increasingly faded distally; paraterga, venter, and epiproct pale brown to pallid; and podomeres pallid, basal podomeres of legs pallid, increasingly dark brown distally (Fig. 10 B – J).</p><p>Clypeolabral region and vertex sparsely setose, epicranial suture distinct. Antennae long, reaching until ring 5 when stretched dorsally (Fig. 10 A). In width, ring 3 = 4 &lt;collum &lt;ring 2 &lt;5 &lt;6–17 (♂); thereafter body gently and gradually tapering towards telson. Collum with three transverse rows of setae: 3 + 3 anterior, 2 + 2 intermediate, and 4 + 4 posterior; caudal corner very narrowly rounded, paraterga slightly declined ventrally, not produced past rear tergal margin (Fig. 10 A, B).</p><p>Tegument smooth and shining, prozona finely shagreened, metaterga nearly smooth, faintly rugulose and leathery (Fig. 10 B – F, H) Postcollum metaterga each with two transverse rows of setae: 2 + 2 in anterior and 3 + 3 in posterior row, the latter traceable as insertion points when setae broken off. Tergal setae simple, short, slender, ~ 1 / 5 metatergal length. Axial line visible on metazona, traceable also on prozona. Paraterga weak (Fig. 10 A – F, H), slightly upturned posteriorly, lying at ~ 1 / 2 midbody height, lateral edge without incisions. Paraterga 2 subhorizontal, broad in dorsal view, thin in lateral view, anterior edge well-developed, slightly rounded and oblique laterally, caudal tip rounded, slightly produced past rear tergal margin (Fig. 10 B, C).</p><p>Paraterga 3 and 4 with evident lateral bulges, caudal corner very broadly rounded, not produced past rear tergal margin (Fig. 10 B, C). Following rings with small lateral bulges, not produced past rear tergal margin, broader on pore-bearing rings (Fig. 10 B, D, F). Calluses on paraterga delimited by a sulcus both dorsally and ventrally. Ozopores evident, lateral, each lying in an ovoid groove at ~ 1 / 3 metatergal length in front of posterior edge of metaterga (Fig. 10 E, H). Transverse metatergal sulci usually distinct (Fig. 10 B – F, H), incomplete on ring 17, complete on rings 5–16, shallow, not reaching the bases of paraterga, at most faintly beaded at bottom. Stricture between pro- and metazona wide and rather deep, ribbed at bottom down to base of paraterga (Fig. 10 B – E). Pleurosternal carinae complete crests with sharp caudal teeth on rings 2–4, thereafter missing (Fig. 10 C, E, H).</p><p>Epiproct (Fig. 10 F – H) conical, flattened dorsoventrally, tip subtruncate, subapical lateral papillae small, but visible, lying close to tip. Hypoproct roundly subtrapeziform, small setigerous knobs at caudal edge small and well-separated (Fig. 10 G).</p><p>Sterna sparsely setose, without modifications (Fig. 10 G); an entire, high, rounded, linguiform, setose, sternal lobe between ♂ coxae 4 (Fig. 10 I, J). A paramedian pair of evident tubercles in front of gonopodal aperture. Legs long and slender, midbody ones ~ 1.3–1.5 as long as body height; prefemora without modifications; ♂ tarsal brushes present until ring 10 (♂).</p><p>Gonopods relatively simple and suberect (Figs 11, 12). Coxite slightly curved caudally, rather densely setose distodorsally (Fig. 11 A, B). Prefemoral part (pfe) densely setose, ~ 1 / 4 as long as acropodite (femoral + postfemoral parts) (Fig. 11 A – C). Femorite (fe) strongly reduced, without femoral processes, bearing a medial, strong, long, flagelliform solenomere (sl), twisted distally, and with an oblique lateral sulcus defining a postfemoral part (Fig. 11 A, B). Solenophore (sph) long, rather slender, slightly curved caudally, sheathing most of solenomere (sl), directed caudally, with a lateral shoulder (u) at base (Figs 11 C, D, 12 A, D, E). Solenophore consisting of a well-developed, strongly twisted lamina lateralis (ll) and a smaller, slightly twisted lamina medialis (lm) (Fig. 11 A – D). Tip of the lamina lateralis (ll) split into two broad, expanded apical laminae, a median lamina with a small denticle, and halfway bearing a broad, expanded apical lamina (Figs 11 A, D, 12 C – F), subtruncate tip process d rising proximal to lamina lateralis (Fig. 11 A, B).</p><p>Distribution.</p><p>Known only from the type locality, apparently endemic to the central part of Laos.</p><p>Etymology.</p><p>To honor Dr. Peter Jäger, a renowned arachnologist from the Senckenberg Research Institute in Frankfurt, Germany, who collected the type specimens of this species.</p><p>Remarks.</p><p>The new species was discovered during a survey undertaken as part of the Laos Biodiversity Survey project.</p><p>Key to the currently known species of  Touranella (chiefly based on ♂), modified after Nguyen et al. (2023)</p></div>	https://treatment.plazi.org/id/2763A5D63EF95FD1A24F10D263463331	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Likhitrakarn, Natdanai;Golovatch, Sergei I.;Inkhavilay, Khamla;Panha, Somsak;Sutcharit, Chirasak	Likhitrakarn, Natdanai, Golovatch, Sergei I., Inkhavilay, Khamla, Panha, Somsak, Sutcharit, Chirasak (2025): Taxonomic review of the millipede genus Touranella Attems, 1937, with a redescription of the type species, T. gracilis Attems, 1937, and descriptions of three new species from Laos (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 1238: 183-208, DOI: 10.3897/zookeys.1238.147550
44F8C92BEC5E541486DC964EB5C3C948.text	44F8C92BEC5E541486DC964EB5C3C948.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Touranella logunovi Golovatch 2024	<div><p>Touranella logunovi Golovatch, 2024</p><p>Touranella logunovi Golovatch, 2024: 408 (D).</p><p>Remarks.</p><p>This species was described from Lam Dong Province, Durong District, Bidoup – Nui Ba National Park, near field station, 12°10'38.47"N, 108°40'49.28"E, 1450–1500 m a. s. l., mid-montane mixed tropical forest. Apparently, the species is endemic to Vietnam (Golovatch 2024).</p></div>	https://treatment.plazi.org/id/44F8C92BEC5E541486DC964EB5C3C948	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Likhitrakarn, Natdanai;Golovatch, Sergei I.;Inkhavilay, Khamla;Panha, Somsak;Sutcharit, Chirasak	Likhitrakarn, Natdanai, Golovatch, Sergei I., Inkhavilay, Khamla, Panha, Somsak, Sutcharit, Chirasak (2025): Taxonomic review of the millipede genus Touranella Attems, 1937, with a redescription of the type species, T. gracilis Attems, 1937, and descriptions of three new species from Laos (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 1238: 183-208, DOI: 10.3897/zookeys.1238.147550
B2F1D131B0E654D1B99ACAAD94BB3E36.text	B2F1D131B0E654D1B99ACAAD94BB3E36.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Touranella moniliformis Golovatch & Semenyuk 2018	<div><p>Touranella moniliformis Golovatch &amp; Semenyuk, 2018</p><p>Touranella moniliformis Golovatch &amp; Semenyuk, 2018: 13 (D).</p><p>Touranella moniliformis – Nguyen et al. 2023: 170 (M, K).</p><p>Remarks.</p><p>The original description of the species recorded from Cat Tien National Park, 11°26'N, 107°21'E, 180 m a. s. l., within the monsoon broadleaved lowland tropical forest of Dong Nai Province, Vietnam. Apparently, endemic to Vietnam (Golovatch and Semenyuk 2018).</p></div>	https://treatment.plazi.org/id/B2F1D131B0E654D1B99ACAAD94BB3E36	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Likhitrakarn, Natdanai;Golovatch, Sergei I.;Inkhavilay, Khamla;Panha, Somsak;Sutcharit, Chirasak	Likhitrakarn, Natdanai, Golovatch, Sergei I., Inkhavilay, Khamla, Panha, Somsak, Sutcharit, Chirasak (2025): Taxonomic review of the millipede genus Touranella Attems, 1937, with a redescription of the type species, T. gracilis Attems, 1937, and descriptions of three new species from Laos (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 1238: 183-208, DOI: 10.3897/zookeys.1238.147550
0E4B974486D5547997E7C5209605ED31.text	0E4B974486D5547997E7C5209605ED31.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Touranella peculiaris Golovatch 2009	<div><p>Touranella peculiaris Golovatch, 2009</p><p>Touranella peculiaris Golovatch, 2009 a: 4 (D, K).</p><p>Touranella peculiaris – Golovatch 2009 b: 120 (M); 2016: 132 (R); Golovatch and Semenyuk 2010: 125 (M); 2018: 16 (M); Nguyen and Sierwald 2013: 1180 (L); Golovatch 2016: 139 (M, K); Nguyen et al. 2023: 170 (M, K).</p><p>Remarks.</p><p>This species was first documented in Nui Ba Nature Reserve, 12°10'N, 108°40'E, 1,400 –1,900 m a. s. l., near Lang Lanh, Bi Doup, Lam Dong Province, Vietnam. It is an endemic species confined to Vietnam (Golovatch 2009 a, 2016).</p></div>	https://treatment.plazi.org/id/0E4B974486D5547997E7C5209605ED31	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Likhitrakarn, Natdanai;Golovatch, Sergei I.;Inkhavilay, Khamla;Panha, Somsak;Sutcharit, Chirasak	Likhitrakarn, Natdanai, Golovatch, Sergei I., Inkhavilay, Khamla, Panha, Somsak, Sutcharit, Chirasak (2025): Taxonomic review of the millipede genus Touranella Attems, 1937, with a redescription of the type species, T. gracilis Attems, 1937, and descriptions of three new species from Laos (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 1238: 183-208, DOI: 10.3897/zookeys.1238.147550
F3D3222F30105A7FA6311366FC47E396.text	F3D3222F30105A7FA6311366FC47E396.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Touranella pilosa Golovatch 2016	<div><p>Touranella pilosa Golovatch, 2016</p><p>Touranella pilosa Golovatch, 2016: 136 (D, K).</p><p>Touranella pilosa – Golovatch and Semenyuk 2018: 16 (M); Nguyen et al. 2023: 170 (M, K).</p><p>Remarks.</p><p>This species was originally described from Sankhua Sabha District, Nepal, 2,600 –2,800 m a. s. l., inhabits  Quercus semecarpifolia,  Rhododendron scrub forests. Endemic to Nepal (Golovatch 2016).</p></div>	https://treatment.plazi.org/id/F3D3222F30105A7FA6311366FC47E396	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Likhitrakarn, Natdanai;Golovatch, Sergei I.;Inkhavilay, Khamla;Panha, Somsak;Sutcharit, Chirasak	Likhitrakarn, Natdanai, Golovatch, Sergei I., Inkhavilay, Khamla, Panha, Somsak, Sutcharit, Chirasak (2025): Taxonomic review of the millipede genus Touranella Attems, 1937, with a redescription of the type species, T. gracilis Attems, 1937, and descriptions of three new species from Laos (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 1238: 183-208, DOI: 10.3897/zookeys.1238.147550
6075D95BDFBC598ABF27C5A3BAC7785F.text	6075D95BDFBC598ABF27C5A3BAC7785F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Touranella srisonchaii Likhitrakarn 2025	<div><p>Touranella srisonchaii Likhitrakarn sp. nov.</p><p>Figs 7, 8, 9</p><p>Type material.</p><p>Holotype • ♂ (CUMZ -PD 0032), Laos, Champasak Province, Paksong District, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=106.23893&amp;materialsCitation.latitude=15.180667" title="Search Plazi for locations around (long 106.23893/lat 15.180667)">Phu Thevada Hotel</a>, 106 m, 15°10'50.4"N, 106°14'20.17"E, 24.2.2017, leg. R. Srisonchai  .</p><p>Paratypes: • 25 ♂, 42 ♀ (CUMZ -PD 0032), same locality, together with holotype, leg. R. Srisonchai.</p><p>Diagnosis.</p><p>Morphologically, in gonopodal structure this new species seems to be especially similar to both  T. chenla sp. nov. and  T. champasak, but it is distinguished by the uniform black coloration lacking a cingulate pattern (Fig. 7 A – H). It differs from  T. champasak by having process d at the mid-length of the solenophore (Figs 8, 9) (vs absent); a more erect and thicker solenophore (Figs 8, 9) (vs slender and curved).  T. srisonchaii sp. nov. differs from  T. chenla sp. nov. in having ♂ tarsal brushes present until ring 8 (vs until ring 14) and a twisted solenophore with a laminate and subtruncate process d (Figs 8, 9) (vs a suberect gonopodal solenophore with a slender and acute process d, Figs 5, 6).</p><p>Description.</p><p>Length 14.8–21.0 (♂) or 14.6–19.8 mm (♀), width of midbody pro- and metazona 1.5–2.3 and 1.8–2.2 mm (♂) or 1.8–2.4 and 2.2–2.6 mm (♀), respectively. Coloration of live animals mostly dark or blackish (Fig. 7 A); head, antennae, paraterga, and epiproct slightly lighter; venter dark brown, legs pale yellowish. Coloration in alcohol after seven years of preservation faded to dark brown or blackish; head, antennae, paraterga, and epiproct dark brown to pale brown; venter pale brown; legs pale yellowish to pallid (Fig. 7 B – J).</p><p>Clypeolabral region and vertex sparsely setose, epicranial suture distinct. Antennae moderately long (Fig. 7 A, B), reaching until body ring 4 (♂, ♀) when stretched dorsally. Head in width &lt;collum &lt;ring 3 &lt;4 &lt;2 &lt;5–16 (♂, ♀); thereafter body gently and gradually tapering towards telson. Collum with three transverse rows of setae: 3 + 3 anterior, 2 + 2 intermediate, and 4 + 4 posterior; caudal corner very narrowly rounded, slightly declined ventrally, not produced past rear tergal margin (Fig. 7 B, C).</p><p>Tegument smooth and shining, prozona finely shagreened, metaterga finely rugulose (Fig. 7 A – F, H); surface below paraterga finely microgranulate (Fig. 7 C, E, F). Postcollum metaterga each with two transverse rows of setae: 3 + 3 anterior and 4 + 4 posterior, nearly always abraded, but still traceable as insertion points. Tergal setae simple, slender, short, ~ 1 / 5 metatergal length. Axial line well visible on metazona, traceable also on prozona.</p><p>Paraterga moderately developed (Fig. 7 A – F, H), slightly upturned, lying at ~ 1 / 2 midbody height. Paraterga 2 subhorizontal, broad in dorsal view, thin in lateral view; shoulders well-developed, slightly rounded and oblique laterally; caudal tip rounded, slightly produced past rear tergal margin (Fig. 7 B, C). Following rings with evident lateral bulges, not produced past rear tergal margin, broader on pore-bearing rings (Fig. 7 B, D, F). Ozopores evident, lateral, each lying in an ovoid groove at ~ 1 / 3 metatergal length in front of posterior edge of metaterga (Fig. 7 C, E, H).</p><p>Transverse metatergal sulci usually distinct (Fig. 7 B – F, H), incomplete on rings 4 and 19, complete on metaterga 5–17 (♂, ♀), shallow, not reaching the bases of paraterga, at most faintly beaded at bottom. Stricture between pro- and metazona narrow, beaded at bottom down to base of paraterga (Fig. 7 B – E). Pleurosternal carinae complete crests with a sharp caudal tooth on ring 2, reduced to a rounded caudal crest on rings 3 and 4, thereafter missing (♂, ♀) (Fig. 7 C, E, H).</p><p>Epiproct (Fig. 7 F – H) conical, flattened dorsoventrally, with two small, rounded, apical papillae; tip rounded; lateral pre-apical papillae very small, lying close to tip. Hypoproct roundly subtriangular, setigerous knobs at caudal edge small and well-separated (Fig. 7 G).</p><p>Sterna sparsely setose, without modifications (Fig. 7 G); an entire, rather short, rounded, linguiform, setose, sternal lobe between ♂ coxae 4 (Fig. 7 I, J). A paramedian pair of evident tubercles in front of gonopodal aperture. Legs moderately long and slender, midbody ones ~ 1.1–1.3 (♂) or 0.9–1.1 × (♀) as long as body height, prefemora without modifications, ♂ tarsal brushes present until ring 14.</p><p>Gonopods relatively simple and suberect (Figs 8, 9). Coxite slightly curved caudally, rather densely setose distodorsally (Figs 8 A, B, 9 A, B). Prefemoral part (pfe) densely setose, ~ 1 / 3 as long as acropodite (femoral + postfemoral parts) (Figs 8 A, B, 9 A, B). Femorite (fe) rather short, with a medial, strong, long, flagelliform solenomere (sl), strongly twisted distally, and with an oblique lateral sulcus defining a postfemoral part (Figs 8 A, B, 9 A, B). Solenophore (sph) long, stout, suberect, with a clear lateral shoulder, sheathing most of solenomere (sl) (Figs 8 A, B, 9 A – D). Lamina lateralis (ll) well-developed, strongly twisted, and lamina medialis (lm) suberect (Figs 8 B, 9 D, E). Tip of lamina lateralis a broad, expanded, apical lamina with three small denticles (Figs 8 A, B, 9 A – G), at halfway bearing a large, long, slightly curved, subtruncate tip process d, this rising distal to lamina lateralis (Figs 8 A, B, 9 A – G).</p><p>Distribution.</p><p>Known only form the type locality, apparently endemic to the southern part of Laos.</p><p>Etymology.</p><p>To honor Dr. Ruttapon Srisonchai, diplopodologist at the Faculty of Science of Khon Kaen University, who has not only contributed to the study of millipede taxonomy in Thailand, but also collected the type series of this new species.</p><p>Remarks.</p><p>The new species was discovered near Phu Thevada Hotel in the evening following a rainfall, after a full day of collecting. The area surrounding the hotel is a small pine forest situated on a low hill. The rainfall prompted millipedes and land snails to emerge on the ground and tree trunks, facilitating the collection of a significant number of samples. In total, 67 specimens were taken, revealing a male to female ratio of 1: 1.68.</p></div>	https://treatment.plazi.org/id/6075D95BDFBC598ABF27C5A3BAC7785F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Likhitrakarn, Natdanai;Golovatch, Sergei I.;Inkhavilay, Khamla;Panha, Somsak;Sutcharit, Chirasak	Likhitrakarn, Natdanai, Golovatch, Sergei I., Inkhavilay, Khamla, Panha, Somsak, Sutcharit, Chirasak (2025): Taxonomic review of the millipede genus Touranella Attems, 1937, with a redescription of the type species, T. gracilis Attems, 1937, and descriptions of three new species from Laos (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 1238: 183-208, DOI: 10.3897/zookeys.1238.147550
E0F02CA9A6745FB98F1E37F52E65DDF8.text	E0F02CA9A6745FB98F1E37F52E65DDF8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Touranella trichosa Golovatch & Semenyuk 2018	<div><p>Touranella trichosa Golovatch &amp; Semenyuk, 2018</p><p>Touranella trichosa Golovatch &amp; Semenyuk, 2018: 16 (D).</p><p>Touranella trichosa – Nguyen et al. 2023: 170 (M, K).</p><p>Remarks.</p><p>This species was originally described from Kon Ka Kinh National Park, Gia Lai Province, Vietnam. Apparently, endemic to Vietnam (Golovatch and Semenyuk 2018).</p></div>	https://treatment.plazi.org/id/E0F02CA9A6745FB98F1E37F52E65DDF8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Likhitrakarn, Natdanai;Golovatch, Sergei I.;Inkhavilay, Khamla;Panha, Somsak;Sutcharit, Chirasak	Likhitrakarn, Natdanai, Golovatch, Sergei I., Inkhavilay, Khamla, Panha, Somsak, Sutcharit, Chirasak (2025): Taxonomic review of the millipede genus Touranella Attems, 1937, with a redescription of the type species, T. gracilis Attems, 1937, and descriptions of three new species from Laos (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 1238: 183-208, DOI: 10.3897/zookeys.1238.147550
