identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
9F18B6F6A3555E5FB48868BB80DC570C.text	9F18B6F6A3555E5FB48868BB80DC570C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Caridina ngocson Do & Phan & von Rintelen & Le & von Rintelen 2025	<div><p>Caridina ngocson sp. nov.</p><p>Figs 2, 3, 4</p><p>Type material.</p><p>Holotype. • Adult male, cl 4.6 mm, IB -FS 007, Vietnam, Hoa Binh Province, Lac Son District, Ngoc Son Commune, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.30223&amp;materialsCitation.latitude=20.465479" title="Search Plazi for locations around (long 105.30223/lat 20.465479)">Van Village</a>, a small stream, 20°27'55.721"N, 105°18'8.027"E, 23 March 2017, collected by Van Tu Do . Paratypes. • 7 males, cl 3.8–5.3 mm, 4 females, cl 4.2–5.3 mm, 50 additional specimens, ZMB 30276, same data as holotype; • male, cl 4.4, female, cl 5 mm, IB -FS 008, Vietnam, Hoa Binh Province, Lac Son District, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.3732&amp;materialsCitation.latitude=20.40331" title="Search Plazi for locations around (long 105.3732/lat 20.40331)">Ngoc Son Commune</a>, a small stream, 20°24'11.92"N, 105°22'23.5"E, collected by Thi Yen Phan, 02 February 2023 .</p><p>Comparative material.</p><p>Caridina serrata Stimpson, 1860 . • 10 males, cl 3.1–4.7, 5 females, cl 4.0–4.6, ZMB 30306, Vietnam, Quang Nam Province, Hoi An City, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=108.52297&amp;materialsCitation.latitude=15.9430275" title="Search Plazi for locations around (long 108.52297/lat 15.9430275)">Cu Lao Cham Island</a>, a small stream running into the reservoir, 15°56'34.9"N 108°31'22.7"E, collected by Pham The Cuong, 10 May 2017.</p><p>Diagnosis.</p><p>Caridina ngocson sp. nov. can be distinguished from other members of the genus Caridina by a combination of the following characteristics: short, slender, and slightly downward-curving rostrum, reaching from beginning to middle of second segment of antennular peduncle (Fig. 2 A); stylocerite extending beyond distal end of basal antennular peduncle; rostral formula of 4–6 + 5 – 11 / 0 – 2; well-developed eyes (Fig. 2 B); kidney-shaped endopod of male first pleopod, with appendix interna slightly exceeding terminal margin of endopod by 0.23 of its length (Fig. 3 G, H); appendix masculina of male second pleopod slender, reaching approximately 0.43 × endopod length; appendix interna extending ~ 0.60 × length of appendix masculina (Fig. 3 I, J); and large eggs.</p><p>Description.</p><p>Cephalothorax and cephalic appendages. Carapace length 3.8–5.3 mm (median 4.4 mm, n = 12). Rostrum short and slender, straight, reaching to beginning to middle of second segment of antennular peduncle, 0.25–0.4 (median 0.33) × as long as carapace, rostral formula 4–6 + 5 – 11 / 0 – 2 (Fig. 2 A). Suborbital angle acute, completely fused with antennal spine; pterygostomian margin rounded, slightly introduced forward (Fig. 2 A). Eyes well developed with globular cornea, anterior end reaching to 0.8 × length of basal segment of antennular peduncle (Fig. 2 A). Antennular peduncle 0.47–0.58 (median 0.54) × as long as carapace; basal segment 1.98–2.39 (median 2.05) × as long as second segment, second segment 0.96–1.24 (median 1.05) × as long as third segment (Fig. 2 B). Stylocerite reaching to beginning or to 0.4 × length of second segment of antennular peduncle (Fig. 2 B). Scaphocerite ovate, reaching beyond distal end of antennular peduncle, 2.55–2.97 (median 2.55) × as long as wide (Fig. 2 A, C).</p><p>Abdominal somites, telson, and uropods. Sixth abdominal somite 0.36–0.4 (median 0.38) × length of carapace, 1.36–1.69 (median 1.45) × as long as fifth abdominal somite, 0.64–0.84 (median 0.73) × length of telson. Telson length 2.14–2.55 (median 2.27) × as long as proximal wide, distal margin triangular, terminating in a short median projection, with 4–6 pairs of dorsal spiniform setae and one pair of dorso-subdistal spiniform setae; distal end with three or four pairs of spiniform setae, lateral pair longer than intermediate pairs (Fig. 2 D, E). Preanal carina low, slightly bent backwards, with few setae, lacking a spine (Fig. 2 F). Uropodal diaeresis with 16–21 (median 20) movable spiniform setae, outermost longer than lateral angle (Fig. 2 G).</p><p>Mouthparts and branchiae. Incisor process of mandible ending in one row of 5–7 irregular teeth, molar process truncated (Fig. 2 H). Lower lacinia of maxillula broadly rounded, upper lacinia elongated, with several distinct teeth and setae on inner margin, palp stout with few simple setae at tip (Fig. 2 I). Upper endites of maxilla subdivided, palp short, scaphognathite tapering posteriorly, with numerous long, curved setae at posterior margin (Fig. 2 J). Distal end of palp of first maxilliped triangular, with a short projection; flagellum of the exopod very elongated, endopod high, reaching 0.8 × length of flagellum of exopod (Fig. 2 K, L). Podobranch of second maxilliped slightly reduced, with few finger-like projections (Fig. 2 M). Third maxilliped reaching to end of antennular peduncle, ending in single terminal claw, exopod reaching 0.6 × length of penultimate segment; ultimate segment longer than penultimate segment; epipod present on the coxa (Fig. 2 N). Branchial formula as typical for genus Paracaridina Liang, Guo &amp; Tang, 1999, five pairs of pleurobranchs well developed; two pairs of arthrobranchs on third maxillipeds, with second pair strongly reduced in size; one pair of podobranchs on second maxilliped slightly reduced, arthrobranch on first pereiopod absent (Liang et al. 1999).</p><p>Pereiopods. Epipods present on first fourth pereiopods. First pereiopod short, robust, reaching end of basal segment of antennular peduncle; chela 1.91–2.22 (median 2.02) × as long as wide, 1.22–1.44 (median 1.26) × length of carpus; tips of fingers rounded, without hook; dactylus 0.73–0.89 (median 0.83) × as long as palm; carpus excavated strongly anteriorly, 1.73–2.25 (median 1.89) × as long as wide; carpus 0.83–1.01 (median 0.95) × length of merus; merus 2.3–2.94 (median 2.67) × as long as wide, longer than ischium (Fig. 3 A). Second pereiopod long, slender, reaching beyond distal end of antennular peduncle; chela 2.19–2.62 (median 2.4) × as long as wide, 0.60–0.78 (median 0.69) × length of carpus; tips of fingers rounded, without hook; dactylus 1.04–1.4 (median 1.17) × as long as palm; carpus 5.0–5.69 (median 5.16) × as long as wide, 0.93–1.05 (median 1.0) × as long as merus; merus 4.81–5.58 (median 5.12) × as long as wide, longer than ischium (Fig. 3 B). Third pereiopod slender, reaching beyond distal end of antennular peduncle by its dactylus, terminating in one claw, with five accessory spiniform setae on flexor margin, dactylus 2.44–3.19 (median 2.82) × as long as wide (terminal claw and spiniform setae on flexor margin included), propodus 8.38–9.3 (median 8.79) × as long as wide, 4–4.64 (median 4.17) × as long as dactylus; carpus 3.41–4.41 (median 4.01) × as long as wide, 0.53–0.69 (median 0.62) × as long as propodus, 0.46–0.55 (median 0.52) × as long as merus; merus 5.33–6.68 (median 5.74) × as long as wide, bearing 3 strong, movable spiniform setae on posterior margin of outer surface; ischium with one small movable spiniform seta (Fig. 3 C, D). Fifth pereiopod slender, reaching to end of third segment of antennular peduncle, dactylus 3.27–3.67 (median 3.53) × as long as wide (terminal claw and spiniform setae on flexor margin included), terminating in one large claw, with 30–37 spiniform setae on flexor margin; propodus 9.82–12.06 (median 10.55) × as long as wide, 3.06–3.98 (median 3.54) × length of dactylus; carpus 3.45–4.39 (median 4.0) × as long as wide, 0.46–0.57 (median 0.48) × as long as propodus, 0.56–0.66 (median 0.61) × as long as merus; merus 5.4–6.21 (median 5.99) × as long as wide, bearing 3 strong, movable spiniform setae on posterior margin of outer surface, ischium without movable spiniform setae (Fig. 3 E, F).</p><p>Pleopods. Endopod of male first pleopod extending to 0.62 × exopod, elongated and kidney-shaped, 2.43–3.04 (median 2.91) × as long as proximal width, inner margin concave, outer margin slightly convex, rounded distally, long pappose setae on outer and distal margins, medium-length setae on inner margin; with appendix interna curved upwards, exceeding terminal margin of endopod by 0.51 its length (Fig. 3 G, H). Appendix masculina of male second pleopod slender, reaching to proximal 0.62 × endopod length, 7.38 × as long as distal width, stick-shaped, with some short spiniform setae on outer surface and some long spiniform setae on distal surface; appendix interna at the middle of appendix masculina, narrow, small, extending ~ 0.54 × length of appendix masculina (Fig. 3 I, J).</p><p>Coloration.</p><p>The body is grey to slightly yellowish in color, with many small reddish black spots. There are several small stripes formed by spots on the abdominal part (Fig. 4 A – C).</p><p>Reproductive biology.</p><p>One ovigerous female, ZMB 30276, cl 4.5 mm, developed eggs, with eyespots, 1.33 × 0.84 mm (15 eggs).</p><p>Etymology.</p><p>The new species is named after the type locality, Ngoc Son Commune. The name is used as a noun in apposition.</p><p>Habitat.</p><p>This new species was found in a stream with mixed sand, gravel, and rock substratum, and clear ﬂowing water from the forest (Fig. 5).</p><p>Distribution.</p><p>Our unpublished survey data throughout Vietnam indicate that this species is restricted to the Ngoc Son district of Hoa Binh province. The estimated area of occupancy is less than 500 km 2. The altitude of the habitat area ranges from 200 to 700 m. The large eggs are also clearly indicative of a landlocked life cycle.</p><p>Molecular phylogenetic results.</p><p>Caridina ngocson sp. nov. is well-supported as the sister species to C. cucphuongensis (Fig. 18). The minimum genetic divergence (p-distance) to C. cucphuongensis) is 6.2 % (COI) and 2.7 % (16 S), respectively (Suppl. materials 1, 2).</p><p>Remarks.</p><p>Caridina ngocson sp. nov. can be included in the C. serrata group based on a short rostrum, a long stylocerite that extends beyond the distal end of the basal antennular peduncle, the presence of dorsal teeth on the carapace, and the male pleopod endopod with a distinctive appendix interna (Cai and Ng 1999). However, C. ngocson sp. nov. differs from C. serrata in: the rostrum is longer (reaching to the beginning to middle of the second segment of antennular peduncle vs nearly reaching to or slightly exceeding the end of the basal segment of antennular peduncle); number of ventral teeth of the rostrum is smaller (0–2 vs 0–6); scaphocerite is stouter (2.6 vs 3.0 × as long as wide); endopod of male first pleopod is more slender (2.9 vs 2.5 × as long as proximal width); egg size is bigger (1.33 × 0.84 mm vs 0.9–1.0 × 0.7– 0.6 mm) (Fig. 2 A, C, G, H; cf. Cai and Ng 1999: fig. 2 A, C, K, J).</p><p>The new species somewhat resembles Caridina cucphuongensis Dang, 1980 in the shape of the male first endopod, the appendix masculine and appendix internal of the male second pleopod. However, C. ngocson sp. nov. can be separated from C. cucphuongensis by a longer rostrum, reaching to the beginning to middle of second segment of antennular peduncle (vs just overreaching eyes to beginning of second segment); higher number of teeth on the dorsal rostrum, armed with 4–6 teeth on carapace posterior to orbital margin and 5–11 teeth on rostrum anterior to orbital margin (vs 0–3 teeth on carapace posterior to orbital margin and 2–7 teeth on rostrum anterior to orbital margin); slender chela of second pereiopod, 2.4 × as long as wide (vs 2.78 × as long as wide); a longer stylocerite, reaching to beginning or to 0.4 × length of second segment of antennular peduncle (vs reaching to beginning of second segment of antennular peduncle); higher number of spiniform setae on flexor margin of the dactylus of fifth pereiopod, 30–37 (vs 24–52 spiniform setae on flexor margin) (Figs 2 A, B, 3 B, F; cf. Phan et al. 2021 b: figs 2 A, B, 3 B, F).</p><p>Caridina ngocson sp. nov. and the following two new species C. xuanlien sp. nov. and C. tanson, have the defining character of Paracaridina Liang, Guo &amp; Tang, 1999 because of their branchial formula (5 pairs of pleurobranchs well developed; 2 pairs of arthrobranchs on third maxillipeds, with second pair strongly reduced in size; 1 pair of podobranchs on second maxilliped slightly reduced, arthrobranch on first pereiopod absent). However, this character is not unique to Paracaridina (von Rintelen et al. 2008) and genetically the new species does not cluster with Paracaridina but – albeit with weak support – rather with Neocaridina . We have thus chosen the conservative approach of tentatively assigning it to Caridina given the clear need of a taxonomic revision at the genus level for these three genera (see Do et al. 2021 b).</p></div>	https://treatment.plazi.org/id/9F18B6F6A3555E5FB48868BB80DC570C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Do, Van Tu;Phan, Thi Yen;von Rintelen, Kristina;Le, Hung Anh;von Rintelen, Thomas	Do, Van Tu, Phan, Thi Yen, von Rintelen, Kristina, Le, Hung Anh, von Rintelen, Thomas (2025): Descriptions of four new species of atyid shrimp (Crustacea, Decapoda, Atyidae) in Vietnam. ZooKeys 1247: 151-186, DOI: 10.3897/zookeys.1247.148607, URL: https://zookeys.pensoft.net/article/148607/download/xml/
0EBC4DD566205EF681C05C4E0995CC2C.text	0EBC4DD566205EF681C05C4E0995CC2C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Caridina tamkim Do & Phan & von Rintelen & Le & von Rintelen 2025	<div><p>Caridina tamkim sp. nov.</p><p>Figs 14, 15, 16, 17</p><p>Type material.</p><p>Holotype. • Adult male, cl 4.8 mm, IB -FS 010, Vietnam, Cao Bang Province, Nguyen Binh District, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=106.02955&amp;materialsCitation.latitude=22.581448" title="Search Plazi for locations around (long 106.02955/lat 22.581448)">Tam Kim Commune</a>, a small stream, 22°34'53.207"N, 106°1'46.368"E, 08 October 2020, collected by Van Tu Do . Paratypes. • 14 females, cl 3.8–5.8 mm, 5 males, cl 4.0– 5.4 mm, ZMB 32924, same data as holotype; • male, cl 4.5 mm, IB -FS 010 _ 2, Vietnam, same location as holotype, 21 March 2024, collected by Van Tu Do; • two males, cl 4.4, 4.5 mm, female, cl 5.5 mm, 8 additional specimens, ZMB 32923, Vietnam, Nguyen Binh District, Tam Kim Commune, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=106.043304&amp;materialsCitation.latitude=22.593863" title="Search Plazi for locations around (long 106.043304/lat 22.593863)">Tran Hung Dao forest</a>, a well at the inflow of small stream, 22°35'37.902"N, 106°2'35.897"E, 08 October 2020, collected by Van Tu Do, Dang Van Dong ; • 2 males, cl 4.3, 4.5 mm, 2 females, cl 5.0, 5.5 mm, 7 additional specimens, ZMB 33814, Vietnam, Cao Bang Province, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.97559&amp;materialsCitation.latitude=22.63795" title="Search Plazi for locations around (long 105.97559/lat 22.63795)">Nguyen Binh District</a>, a small stream near the road, 22°38'16.626"N, 105°58'32.135"E, collected by Van Tu Do, Dang Van Dong, Nguyen Tong Cuong, 5 October 2020 ; • male, cl 4.7 mm, female, cl 4.4 mm, 4 additional specimens, ZMB 33793, Vietnam, Cao Bang Province, Nguyen Binh District, Quang Thanh Commune, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.92389&amp;materialsCitation.latitude=22.576477" title="Search Plazi for locations around (long 105.92389/lat 22.576477)">a small stream run out from Ong cave</a>, 22°34'35.316"N, 105°55'25.998"E, collected by Van Tu Do, 7 October 2020 ; • male, cl 5.4 mm, female, cl 5.2 mm, 10 additional specimens, ZMB 33788, Vietnam, Cao Bang Province, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.97592&amp;materialsCitation.latitude=22.639116" title="Search Plazi for locations around (long 105.97592/lat 22.639116)">Nguyen Binh District</a>, a small stream near the road, 22°38'20.82"N, 105°58'33.318"E, collected by Van Tu Do, 8 October 2020 .</p><p>Comparative material.</p><p>Caridina pacbo Do, von Rintelen &amp; Dang, 2020: • adult male, cl 4.2 mm, IB –FS 003 (Holotype), Vietnam, Cao Bang Province, Ha Quang District, Truong Ha Commune, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=106.042&amp;materialsCitation.latitude=22.983807" title="Search Plazi for locations around (long 106.042/lat 22.983807)">Pac Bo Village</a>, small stream in <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=106.042&amp;materialsCitation.latitude=22.983807" title="Search Plazi for locations around (long 106.042/lat 22.983807)">Khuoi Nam</a>, 22°59'1.7"N, 106°02'31.2"E, 25 May 2017, collected by Van Tu Do.</p><p>Diagnosis.</p><p>This new species is characterized by several morphological characters such as short and slender rostrum, reaching just to the end of the basal segment of antennular peduncle or sometimes to the middle of second segment, without or with a few small teeth (Fig. 14 A); medium stylocerite, reaching to the end of the basal segment or sometimes to beginning of second segment of the antennular peduncle (Fig. 14 B); dactylus of the first pereiopod is shorter than palm (Fig. 15 A); long, sub-rectangular endopod of the male first pleopod, extending to 0.87 of the length of the exopod, with short appendix interna, not or slightly exceeding the terminal margin of the endopod (Fig. 15 G, H); short and stick-shaped appendix masculina of male second pleopod, reaching to proximal 0.8 of the length of the endopod, with narrow and small appendix interna, extending ~ 0.6 of the length of appendix masculina (Fig. 15 I, J).</p><p>Description.</p><p>Cephalothorax and cephalic appendages. Carapace length 3.8–5.8 mm (median 4.6 mm, n = 20). Rostrum very short and slender, straight, mostly reaching near the end of basal segment of antennular peduncle, sometimes to middle of second segment, 0.1–0.2 (median 0.2) as long as carapace, teeth small, rostral formula 0–4 + 0 – 8 / 0 – 2 (Fig. 14 A). Suborbital angle acute, completely fused with antennal spine; pterygostomian margin rounded, slightly produced forward (Fig. 14 A). Eyes normally developed with globular cornea, eyestalk short, anterior end reaching to 0.6 of length of basal segment of antennular peduncle (Fig. 14 A). Antennular peduncle 0.37–0.44 (median 0.43) × as long as carapace; basal segment 1.6–2.2 (median 1.67) × as long as second segment, second segment 1.2–1.25 (median 1.25) × as long as third segment (Fig. 14 B). Stylocerite reaching to end of basal segment or sometime to beginning of second segment of antennular peduncle (Fig. 14 A, B). Scaphocerite ovate, reaching beyond distal end of antennular peduncle, 2.33–2.67 (median 2.5) × as long as wide (Fig. 14 A, C).</p><p>Abdominal somites, telson, and uropods. Sixth abdominal somite 0.3–0.42 (median 0.38) × length of carapace, 1.31–1.64 (median 1.42) × as long as fifth abdominal somite, 0.76–1.0 (median 0.89) × length of telson. Telson length 2.1–2.38 (median 2.25) × as long as proximal wide, distal margin triangular, terminating in a short median projection, with six pairs of dorsal spiniform setae and one pair of dorso-subdistal spiniform setae; distal end with three or four pairs of spiniform setae, lateral pair slightly shorter than intermediate pairs (Fig. 14 D, E). Preanal carina moderately high, slightly bent backwards, with few setae, lacking a spine (Fig. 14 F). Uropodal diaeresis with 17–20 (median 18) movable spiniform setae, outermost longer than lateral angle (Fig. 14 G).</p><p>Mouthparts and branchiae. Incisor process of mandible ending in one row of six or seven irregular teeth, molar process truncated (Fig. 14 H). Lower lacinia of maxillula broadly rounded, upper lacinia elongated, with several distinct teeth and setae on inner margin, palp stout with few simple setae at tip (Fig. 14 I). Upper endites of maxilla subdivided, palp short, scaphognathite tapering posteriorly, with numerous long, curved setae at posterior margin (Fig. 14 J). Distal end of palp of first maxilliped triangular, with a short projection; flagellum of the exopod very elongated, endopod high, reaching 0.9 of length of flagellum of exopod (Fig. 14 K, L). Podobranch of second maxilliped slightly reduced, with few finger-like projections (Fig. 14 M). Third maxilliped reaching near the end of antennular peduncle, ending in single terminal claw, exopod reaching 0.4 of length of penultimate segment; ultimate segment shorter penultimate segment; epipod present on the coxa (Fig. 14 N). Branchial formula as is typical for genus, five pairs of pleurobranchs well developed; three pairs of arthrobranchs, two on third maxillipeds, with second pair strongly reduced in size, one pair on first pereiopod; one pair of podobranchs on second maxillipeds slightly reduced.</p><p>Pereiopods. Epipods present on first to fourth pereiopods. First pereiopod short, robust, reaching beyond end of basal segment of antennular peduncle; chela 2.03–2.35 (median 2.11) × as long as wide, 1.37–1.5 (median 1.4) × length of carpus; tips of fingers rounded, with hook; dactylus shorter than palm, 0.82–0.9 (median 0.88) × as long as palm; carpus excavated strongly anteriorly, 1.5–1.75 (median 1.65) × as long as wide; carpus 0.8–1.0 (median 0.9) × length of merus; merus 2.39–2.67 (median 2.47) × as long as wide, longer than ischium (Fig. 15 A). Second pereiopod long, slender, reaching to distal end of antennular peduncle; chela 3.16–3.5 (median 3.28) × as long as wide, 0.64–0.71 (median 0.67) × length of carpus; tips of fingers rounded, without hook; dactylus 1.33–1.64 (median 1.52) × as long as palm; carpus 6.07–7.08 (median 6.57) × as long as wide, 1.02–1.12 (median 1.11) × as long as merus; merus 5.92–6.38 (median 6.33) × as long as wide, longer than ischium (Fig. 15 B). Third pereiopod slender, reaching beyond end of scaphocerite by its dactylus, terminating in one claw, with four or five accessory spiniform setae on flexor margin, dactylus 3.0–3.83 (median 3.64) × as long as wide (terminal claw and spiniform setae on flexor margin included), propodus 8.5–9.75 (median 9.0) × as long as wide, 3.48–3.950 (median 3.86) × as long as dactylus; carpus 4.69–4.83 (median 4.77) × as long as wide, 0.67–0.76 (median 0.72) × as long as propodus, 0.52–0.58 (median 0.53) × as long as merus; merus 5.63–6.59 (median 6.05) × as long as wide, bearing 3 strong, movable spiniform setae on posterior margin of outer surface; ischium without movable spiniform seta (Fig. 15 C, D). Fifth pereiopod slender, reaching to end of second segment of antennular peduncle, dactylus 2.67–3.38 (median 3.13) × as long as wide (terminal claw and spiniform setae on flexor margin included), terminating in one large claw, with 37–39 spiniform setae on flexor margin; propodus 11.25–13.14 (median 11.5) × as long as wide, 3.41–3.91 (median 3.75) × length of dactylus; carpus 4.73–5.0 (median 5.0) × as long as wide, 0.56–0.6 (median 0.58) × as long as propodus, 0.63–0.71 (median 0.67) × as long as merus; merus 5.5–6.33 (median 6.31) × as long as wide, bearing 3 strong, movable spiniform setae on posterior margin of outer surface, ischium without movable spiniform setae (Fig. 15 E, F).</p><p>Pleopods. Endopod of male first pleopod extending to 0.87 of exopod, sub-rectangular in shape, anterior part not folded backwards, 3.15–3.35 (median 3.19) × as long as proximal width, inner margin concave, outer margin slightly convex, rounded distally, long simple setae on outer and distal margins, medium-length setae on inner margin; with appendix interna curved upwards, not or slightly exceeding terminal margin of endopod by 0.23 of its length (Fig. 15 G, H). Appendix masculina of male second pleopod slender, reaching to proximal 0.8 of endopod length, 7.3 × as long as distal width, stick-shaped, with some spiniform setae on outer surface and on distal surface; appendix interna at the middle of appendix masculina, narrow, small, extending ~ 0.54 length of appendix masculina (Fig. 15 I, J).</p><p>Coloration.</p><p>The body is slightly yellowish to dark blue in color, with many black spots of irregular sizes (Fig. 16 A, B).</p><p>Reproductive biology.</p><p>Four ovigerous females, ZMB 33814, cl 5.0 mm, undeveloped eggs 1.1 × 0.7 mm, cl 5.5 mm, eggs with eyespots 1.1 mm × 0.8 mm.</p><p>Variation.</p><p>There are variations between the populations of the new species on the length of the rostrum, stylocerite, and number of teeth on rostrum. In general, compared to the specimens from the type locality and the nearby locality (ca 3 km, ZMB 32923) with the specimens from ZMB 33814, ZMB 33793, and ZMB 33788 they showed shorter rostrum (can reach only to the end of the basal segment of antennular peduncle vs reaching to the middle of the second segment of the antennular peduncle), lower number of teeth on the rostrum dorsal (up to 8 teeth vs up to 12 teeth) and shorter stylocerite (reaching to the end of the basal segment of the antennular peduncle vs reaching to the beginning of the second segment).</p><p>Etymology.</p><p>The new species is named after the type locality, Tam Kim Commune. The name is used as a noun in apposition.</p><p>Habitat.</p><p>This new species was found in small streams and pools running through the forest. The substratum includes sand, gravel, and bedrock in a water depth of 0.2–0.4 m (Fig. 17).</p><p>Distribution.</p><p>This species is only distributed in Nguyen Binh District, Cao Bang Province in the high elevation from 500 m to 800 m. The estimated area of occupancy is less than 1000 km 2.</p><p>Molecular phylogenetic results.</p><p>Caridina tamkim sp. nov. is well supported as the sister group a clade comprising C. namdat, C. pacbo and C. pseudoserrata . (Fig. 18). The minimum genetic divergence (p-distance) to a species within that clade ( C. namdat) is 9.5 % (COI) and 3.9 % (16 S), respectively (Suppl. materials 1, 2).</p><p>Remarks.</p><p>Caridina tamkim, new species, looks similar to C. pacbo in the shape of the rostrum and the male first pleopod (Do et al. 2020). However, it can be distinguished by a suite of characteristics: the lower number of teeth in the rostrum (0–8 + 0 – 4 / 0 – 2 vs 5–12 + 2 – 6 / 0 – 3); stylocerite extending (mostly reaching to end of basal segment, sometimes to the beginning of second segment vs reaching to the middle of the second segment of the antennular peduncle); slender merus of the first pereiopod (2.39–2.67 × as long as wide vs 2.75–3.8 × as long as wide); slender chela and merus of second pereiopod (3.16–3.5 × as long as wide vs 2.67–3.09 × as long as wide, 5.92–6.38 × as long as wide vs 5.0–5.86 × as long as wide, respectively); the distal margins of the endopod of the male first pleopod (slightly extended on both left and right sides vs not extended) (Figs 14 A, B, 15 A, B, G, H; cf. Do et al. 2020: figs 5 A, B, 6 A, B, G, H).</p></div>	https://treatment.plazi.org/id/0EBC4DD566205EF681C05C4E0995CC2C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Do, Van Tu;Phan, Thi Yen;von Rintelen, Kristina;Le, Hung Anh;von Rintelen, Thomas	Do, Van Tu, Phan, Thi Yen, von Rintelen, Kristina, Le, Hung Anh, von Rintelen, Thomas (2025): Descriptions of four new species of atyid shrimp (Crustacea, Decapoda, Atyidae) in Vietnam. ZooKeys 1247: 151-186, DOI: 10.3897/zookeys.1247.148607, URL: https://zookeys.pensoft.net/article/148607/download/xml/
EE5C738F20385400A0C0893854EE65A7.text	EE5C738F20385400A0C0893854EE65A7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Caridina tanson Do & Phan & von Rintelen & Le & von Rintelen 2025	<div><p>Caridina tanson sp. nov.</p><p>Figs 10, 11, 12</p><p>Type material.</p><p>Holotype. • Adult male, cl 4.5 mm, IB -FS 009, Vietnam, Phu Tho Province, Tan Son District, Xuan Son Commune, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=104.94465&amp;materialsCitation.latitude=21.14447" title="Search Plazi for locations around (long 104.94465/lat 21.14447)">Ban Lap Village</a>, a small stream, 21°8'40.09"N, 104°56'40.739"E, 28 July 2022, collected by Thi Yen Phan . Paratypes. • 6 females, cl 4.0–6.0 mm, 10 males, cl 4.0–6.0 mm, ZMB 32979, same data as holotype; • male, cl 4.5 mm, ZMB 30729, Vietnam, Phu Tho Province, Tan Son District, Dong Son Commune, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=104.87303&amp;materialsCitation.latitude=21.18761" title="Search Plazi for locations around (long 104.87303/lat 21.18761)">Than stream, site 2</a>, 21°11'15.4"N, 104°52'22.9"E, 29 August 2013, collected by Tran Anh Duc .</p><p>Comparative material.</p><p>Caridina pseudoserrata Dang &amp; Do, 2007, • adult male, cl 4.5 mm, ZMB 31570, Vietnam, Cao Bang Province, Phuc Hoa District, My Hung Commune, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=106.55091&amp;materialsCitation.latitude=22.486334" title="Search Plazi for locations around (long 106.55091/lat 22.486334)">Na Rieng village</a>, a stream runs inside <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=106.55091&amp;materialsCitation.latitude=22.486334" title="Search Plazi for locations around (long 106.55091/lat 22.486334)">Nguom Khuoi Khua Cave</a>, 22°29'10.8"N, 106°33'3.27"E, 27 May 2017, collected by Van Tu Do.</p><p>Diagnosis.</p><p>This new species is characterized by rostrum short, straight, slender, tapering towards tip, reaching from beginning to end of third segment of antennular peduncle (Fig. 10 A); stylocerite reaches from beginning to 0.2 × length of second segment of the antennular peduncle (Fig. 10 B); carpus of fifth pereiopod slender, 7.3 × as long as wide (Fig. 11 A); endopod of male first pleopod kidney-shaped, extending to 0.52 of exopod, 2.4 × as long as proximal width (Fig. 11 G, H).</p><p>Description.</p><p>Cephalothorax and cephalic appendages. Carapace length 4.0– 6.4 mm (median 4.5 mm, n = 11). Rostrum slender, straight, reaching to beginning, sometime to the end of the third segment of antennular peduncle to distal end of antennular peduncle, 0.45–0.75 (median 0.52) as long as carapace, rostral formula 4–6 + 9 – 14 / 2 – 5 (Fig. 10 A). Suborbital angle acute, completely fused with antennal spine; pterygostomian margin rounded, slightly produced forward (Fig. 10 A). Eyes well developed with globular cornea, anterior end reaching to 0.7 length of basal segment of antennular peduncle (Fig. 10 A). Antennular peduncle 0.46–0.67 (median 0.56) as long as carapace; basal segment 1.0–1.67 (median 1.25) × as long as second segment, second segment 1.14–2 (median 1.58) × as long as third segment (Fig. 10 B). Stylocerite reaching to beginning or to 0.2 of length of second segment of antennular peduncle (Fig. 10 B). Scaphocerite ovate, reaching beyond distal end of antennular peduncle, 3–3.81 (median 3.37) × as long as wide (Fig. 10 A, C).</p><p>Abdominal somites, telson, and uropods. Sixth abdominal somite 0.33–0.5 (median 0.4) length of carapace, 1.36–1.73 (median 1.5) × as long as fifth abdominal somite, 0.62–0.95 (median 0.76) length of telson. Telson length 2.0–2.89 (median 2.5) × as long as proximal wide, distal margin triangular, terminating in a short median projection, with 5–6 pairs of dorsal spiniform setae and one pair of dorso-subdistal spiniform setae; distal end with 4–5 pairs of spiniform setae, lateral pair longer than intermediate pairs (Fig. 10 D, E). Preanal carina low, slightly bent backwards, with few setae, lacking a spine (Fig. 10 F). Uropodal diaeresis with 16–21 (median 18) movable spiniform setae, outermost shorter than lateral angle (Fig. 10 G).</p><p>Mouthparts and branchiae. Incisor process of mandible ending in one row of 6–7 irregular teeth, molar process truncated (Fig. 10 H). Lower lacinia of maxillula broadly rounded, upper lacinia elongated, with several distinct teeth and setae on inner margin, palp stout with few simple setae at tip (Fig. 11 I). Upper endites of maxilla subdivided, palp short, scaphognathite tapering posteriorly, with numerous long, curved setae at posterior margin (Fig. 10 J). Distal end of palp of first maxilliped triangular, with a short projection; flagellum of the exopod very elongated, endopod high, reaching 0.8 length of flagellum of exopod (Fig. 10 K, L). Podobranch of second maxilliped slightly reduced, with few finger-like projections (Fig. 10 M). Third maxilliped reaching to end of antennular peduncle, ending in single terminal claw, exopod reaching 0.4 length of penultimate segment; ultimate segment slightly longer than penultimate segment; epipod present on the coxa (Fig. 10 N). Branchial formula as typical for Paracaridina genus, five pairs of pleurobranchs well developed; two pairs of arthrobranchs on third maxillipeds, with second pair strongly reduced in size; podobranch on second maxillipeds slightly reduced, arthrobranch on first pereiopod absent (Liang et al. 1999).</p><p>Pereiopods. Epipods present on first to fourth pereiopods. First pereiopod short, robust, reaching end of basal segment of antennular peduncle; chela 1.91–2.75 (median 2.28) × as long as wide, 1.17–1.69 (median 1.41) × length of carpus; tips of fingers rounded, without hook; dactylus 0.77–1.18 (median 0.97) × as long as palm; carpus excavated strongly anteriorly, 1.6–2.29 (median 2.0) × as long as wide; carpus 0.79–1.0 (median 0.84) × length of merus; merus 2.5–3.6 (median 3.0) × as long as wide, longer than ischium (Fig. 11 A). Second pereiopod long, slender, reaching beyond distal end of antennular peduncle; chela 2.11–3.63 (median 2.75) × as long as wide, 0.63–0.87 (median 0.76) × length of carpus; tips of fingers rounded, without hook; dactylus 0.92–1.4 (median 1.29) × as long as palm; carpus 4.6–6.0 (median 5.0) × as long as wide, 0.74–1.17 (median 1.0) × as long as merus; merus 4.8–7 (median 5.8) × as long as wide, longer than ischium (Fig. 11 B). Third pereiopod slender, reaching beyond distal end of antennular peduncle by its dactylus, terminating in one claw, with five to six accessory spiniform setae on flexor margin, dactylus 2.67–4.0 (median 3.0) × as long as wide (terminal claw and spiniform setae on flexor margin included), propodus 9.75–12 (median 10.13) × as long as wide, 3.89–5.13 (median 4.44) × as long as dactylus; carpus 4–5.5 (median 4.5) × as long as wide, 0.49–0.69 (median 0.6) × as long as propodus, 0.43–0.56 (median 0.47) × as long as merus; merus 5.63–7.86 (median 7.01) × as long as wide, bearing three strong, movable spiniform setae on posterior margin of outer surface; ischium with one small movable spiniform seta (Fig. 11 C, D). Fifth pereiopod slender, reaching to end of third segment of antennular peduncle, dactylus 3.67–7 (median 4.17) × as long as wide (terminal claw and spiniform setae on flexor margin included), terminating in one large claw, with 41–58 spiniform setae on flexor margin; propodus 10.75–14.67 (median 12.38) × as long as wide, 3.31–4.82 (median 4.01) × length of dactylus; carpus 4.17–6.25 (median 5) × as long as wide, 0.42–0.54 (median 0.48) × as long as propodus, 0.49–0.71 (median 0.58) × as long as merus; merus 6.5–8.6 (median 7.3) × as long as wide, bearing 3 strong, movable spiniform setae on posterior margin of outer surface, ischium without movable spiniform setae (Fig. 11 E, F).</p><p>Pleopods. Endopod of male first pleopod extending to 0.52 of exopod, elongated and kidney-shaped, 2.14–2.57 (median 2.43) × as long as proximal width, inner margin concave, outer margin slightly convex, rounded distally, long pappose setae on outer and distal margins, medium-length setae on inner margin; appendix interna long, exceeding terminal margin of endopod by 0.43 its length (Fig. 11 G, H). Appendix masculina of male second pleopod slender, reaching to proximal 0.63 endopod length, 6.73 × as long as distal width, stick-shaped, with some short spiniform setae on outer surface and some long spiniform setae on distal surface; appendix interna at the middle of appendix masculina, small, extending ~ 0.66 length of appendix masculina (Fig. 11 I, J).</p><p>Coloration.</p><p>The body is slightly yellowish to dark blue in color, with many small reddish spots and small lines of irregular sizes (Fig. 12 A, B).</p><p>Reproductive biology.</p><p>Ovigerous female, ZMB 32979, cl 6.0 mm, eggs with eyespots 1.1 × 0.7 mm.</p><p>Etymology.</p><p>The new species is named after the type locality, Tan Son District. The name is used as a noun in apposition.</p><p>Habitat.</p><p>This new species was found in small streams. The substratum includes sand, gravel, bedrock, rotten leaves, and organic mulch in a water depth of 0.2–0.5 m (Fig. 13).</p><p>Distribution.</p><p>This species has only been recorded in the highland streams of Tan Son District, Phu Tho Province at altitudes from 200 m to 500 m. The estimated area of occupancy is less than 1000 km 2.</p><p>Molecular phylogenetic results.</p><p>Caridina tanson sp. nov. is well supported as the sister group a clade comprising C. haivanensis, C. clinata and C. xuanlien sp. nov. (Fig. 18). The minimum genetic divergence (p-distance) to a species within that clade is 10.0 % (COI, C. haivanensis) and 4.1 % (16 S, C. clinata), respectively (Suppl. materials 1, 2).</p><p>Remarks.</p><p>Caridina tanson, new species, is similar to C. pseudoserrata, also from northeast Vietnam, in the shape of the rostrum and endopod of male first pleopod (Dang and Do 2007 a). The new species can be distinguished from C. pseudoserrata by a suite of characteristics: shorter stylocerite (reaching to the beginning or to 0.2 the length of the second segment of antennular peduncle vs reaching to 0.4–0.5 of second segment of antennular peduncle); shorter endopod of male first pleopod (2.4 × as long as proximal width vs 2.77 × as long as proximal width); longer appendix interna of endopod of male first pleopod (exceeding terminal margin of endopod by 0.43 its length vs not exceeding or just slightly exceeding the terminal margin of endopod) (Fig. 10 A, B, 11 G, H; cf. Dang and Do 2007 a: figs 1 A, 2 A, B).</p><p>Caridina tanson sp. nov. is also close to C. xuanlien sp. nov. in the shape of the rostrum and endopod of the male first pleopod. However, C. tanson can be distinguished from C. xuanlien sp. nov. by: there are more teeth on the rostrum, armed with 9–14 dorsal teeth on the rostrum anterior to the orbital margin and 2–5 ventral teeth (vs 3–9 dorsal teeth on the rostrum anterior to the orbital margin and 0–2 ventral teeth); the endopod of the male first pleopod is stouter, 2.43 × as long as proximal width (vs 2.79 × as long as proximal width); appendix interna of endopod of male first pleopod is longer, exceeding terminal margins of endopod by 0.43 of its length (vs exceeding terminal margins of endopod by 0.23 of its length); appendix masculina of male second pleopod is stouter, 6.73 × as long as distal width (vs 8.25 × as long as distal width).</p></div>	https://treatment.plazi.org/id/EE5C738F20385400A0C0893854EE65A7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Do, Van Tu;Phan, Thi Yen;von Rintelen, Kristina;Le, Hung Anh;von Rintelen, Thomas	Do, Van Tu, Phan, Thi Yen, von Rintelen, Kristina, Le, Hung Anh, von Rintelen, Thomas (2025): Descriptions of four new species of atyid shrimp (Crustacea, Decapoda, Atyidae) in Vietnam. ZooKeys 1247: 151-186, DOI: 10.3897/zookeys.1247.148607, URL: https://zookeys.pensoft.net/article/148607/download/xml/
A50CBB9C1CBD5F7390305EBBFC07B754.text	A50CBB9C1CBD5F7390305EBBFC07B754.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Caridina xuanlien Do & Phan & von Rintelen & Le & von Rintelen 2025	<div><p>Caridina xuanlien sp. nov.</p><p>Figs 6, 7, 8</p><p>Type material.</p><p>Holotype. • Adult male, cl 4.5 mm, IB -FS 008, Vietnam, Thanh Hoa Province, Thuong Xuan District, Yen Nhan Commune, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.12517&amp;materialsCitation.latitude=19.93497" title="Search Plazi for locations around (long 105.12517/lat 19.93497)">a small stream in Xuan Lien National Park</a>, 19°56'05.9"N, 105°07'30.6"E, 21 November 2021, collected by Van Tu Do . Paratypes. • 8 females, cl 4.0– 4.8 mm, 11 males, cl 3.6–4.7 mm, 49 additional specimens, ZMB 32948, same data as holotype; male, cl 4.5 mm, female, cl. 4.6 mm, 107 additional specimens, ZMB 32944, Vietnam, Thanh Hoa Province, Thuong Xuan District, Van Xuan Commune, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.15342&amp;materialsCitation.latitude=19.920834" title="Search Plazi for locations around (long 105.15342/lat 19.920834)">a small stream in Xuan Lien National Park</a>, 19°55'15.0"N, 105°09'12.3"E, 24 October 2021, collected by Van Tu Do .</p><p>Comparative material.</p><p>Caridina nguyeni Li &amp; Liang, 2002: • adult male, cl. 4.7 mm, ZMB 30280, Vietnam, Cao Bang Province, Ha Quang District, Truong Ha Commune, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=106.04825&amp;materialsCitation.latitude=22.984446" title="Search Plazi for locations around (long 106.04825/lat 22.984446)">Pac Bo village</a>, a small stream near <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=106.04825&amp;materialsCitation.latitude=22.984446" title="Search Plazi for locations around (long 106.04825/lat 22.984446)">Le Nin</a> stream, 22°59'4"N, 106°2'53.7"E, 25 May 2017, collected by Van Tu Do.</p><p>Diagnosis.</p><p>Caridina xuanlien sp. nov. is characterized by several morphological characters such as short rostrum, just reaching to beginning of the third segment of antennular peduncle; stylocerite reaching to beginning of second segment of antennular peduncle (Fig. 6 A, B); dactylus of first pereiopod longer than palm (Fig. 7 A); short, kidney-shaped endopod of male first pleopod, extending to 2.79 × exopod, with well-developed appendix interna, slightly exceeding terminal margin of endopod by 0.23 its length (Fig. 7 G, H); short and stick-shaped appendix masculina of male second pleopod, reaching to proximal 0.43 × endopod length, with narrow and small appendix interna, extending ~ 0.6 × length of appendix masculina (Fig. 7 I, J).</p><p>Description.</p><p>Cephalothorax and cephalic appendages. Carapace length 3.6–4.8 mm (median 4.3 mm, n = 20). Rostrum short and slender, slightly downward, reaching to beginning of the third segment of antennular peduncle, 0.1–0.2 (median 0.2) × as long as carapace, rostral formula 2–7 + 3 – 9 / 0 – 2 (Fig. 6 A). Suborbital angle acute, completely fused with antennal spine; pterygostomian margin rounded, slightly produced forward (Fig. 6 A). Eyes well developed with globular cornea, anterior end reaching to 0.7 × length of basal segment of antennular peduncle (Fig. 6 A). Antennular peduncle 0.6–0.74 (median 0.64) × as long as carapace; basal segment 1.63–2.0 (median 1.88) × as long as second segment, second segment 1.17–1.4 (median 1.33) × as long as third segment (Fig. 6 B). Stylocerite reaching to beginning of second segment of antennular peduncle (Fig. 6 B). Scaphocerite elongated ovate, reaching beyond distal end of antennular peduncle, 2.68–2.88 (median 2.86) × as long as wide (Fig. 6 A, C).</p><p>Abdominal somites, telson, and uropods. Sixth abdominal somite 0.47–0.5 (median 0.47) × length of carapace, 1.47–1.62 (median 1.58) × as long as fifth abdominal somite, 0.85–0.95 (median 0.91) × length of telson. Telson 2.4–3.25 (median 2.75) × as long as proximal wide, distal margin triangular, terminating in a short median projection, with five or six pairs of dorsal spiniform setae and one pair of dorso-subdistal spiniform setae; distal end with three or four pairs of spiniform setae, lateral pair shorter than intermediate pairs (Fig. 6 D, E). Preanal carina low,, with few setae, lacking a spine (Fig. 6 F). Uropodal diaeresis with 18–21 (median 21) movable spiniform setae, outermost shorter than lateral angle (Fig. 6 G).</p><p>Mouthparts and branchiae. Incisor process of mandible ending in one row of six or seven irregular teeth, molar process truncated (Fig. 6 H). Lower lacinia of maxillula broadly rounded, upper lacinia elongated, with a several distinct teeth and setae on inner margin, palp stout with few simple setae at tip (Fig. 6 I). Upper endites of maxilla subdivided, palp short, scaphognathite tapering posteriorly, with numerous long, curved setae at posterior margin (Fig. 6 J). Distal end of palp of first maxilliped triangular, with a short projection; flagellum of the exopod very elongated, endopod reaching 0.8 × length of flagellum of exopod (Fig. 6 K, L). Podobranch of second maxilliped slightly reduced, with few finger-like projections (Fig. 6 M). Third maxilliped reaching near the end of antennular peduncle, ending in single terminal claw, exopod reaching 0.3 × length of penultimate segment; ultimate segment slightly longer penultimate segment; epipod present on the coxa (Fig. 6 N). Branchial formula as typical for Paracaridina genus, five pairs of pleurobranchs well developed; two pairs of arthrobranchs on third maxillipeds, with second pair strongly reduced in size; one pair of podobranchs on second maxillipeds slightly reduced, arthrobranch on first pereiopod absent (Liang et al. 1999).</p><p>Pereiopods. Epipods present on first to fourth pereiopods. First pereiopod short, robust, reaching end of basal segment of antennular peduncle; chela 2.08–2.29 (median 2.12) × as long as wide, 1.21–1.47 (median 1.40) × length of carpus; tips of fingers rounded, without hook; dactylus longer than palm, 1.0–1.07 (median 1.03) × as long as palm; carpus excavated strongly anteriorly, 1.5–1.87 (median 1.73) × as long as, respectively; carpus 0.78–0.91 (median 0.84) × length of merus; merus 2.65–3.54 (median 3.33) × as long as wide, longer than ischium (Fig. 7 A). Second pereiopod long, slender, reaching to distal end of antennular peduncle; chela 2.65–3.0 (median 2.86) × as long as wide, 0.66–0.74 (median 0.73) × length of carpus; tips of fingers rounded, without hook; dactylus 1.42–1.58 (median 1.46) × as long as palm; carpus 5.25–6.21 (median 5.6) × as long as wide, 1.05–1.21 (median 1.16) × as long as merus; merus 5.46–6.0 (median 5.71) × as long as wide, longer than ischium (Fig. 7 B). Third pereiopod slender, reaching beyond distal end of antennular peduncle by its dactylus, terminating in one claw, with five accessory spiniform setae on flexor margin, dactylus 3.29–4 (median 3.57) × as long as wide (terminal claw and spiniform setae on flexor margin included), propodus 8.9–9.44 (median 9.0) × as long as wide, 3.24–4.0 (median 3.71) × as long as dactylus; carpus 4.2–4.92 (median 4.83) × as long as wide, 0.69–0.78 (median 0.72) × as long as propodus, 0.5–0.57 (median 0.56) × as long as merus; merus 5.75–6.82 (median 6.0) × as long as wide, bearing three strong, movable spiniform setae on posterior margin of outer surface; ischium with one small movable spiniform seta (Fig. 7 C, D). Fifth pereiopod slender, reaching to end of third segment of antennular peduncle, dactylus 3.33–3.75 (median 3.5) × as long as wide (terminal claw and spiniform setae on flexor margin included), terminating in one large claw, with 37–40 spiniform setae on flexor margin; propodus 10.88–13.13 (median 11.88) × as long as wide, 3.22–3.53 (median 3.33) × length of dactylus; carpus 4.15–4.36 (median 4.28) × as long as wide, 0.52–0.55 (median 0.55) × as long as propodus, 0.6–0.69 (median 0.62) × as long as merus; merus 5.83–7.17 (median 6.57) × as long as wide, bearing 3 strong, movable spiniform setae on posterior margin of outer surface, ischium without movable spiniform setae (Fig. 7 E, F).</p><p>Pleopods. Endopod of male first pleopod extending to 0.59 × exopod, kidney-shaped, anterior part not folded backwards, 2.75–3.05 (median 2.79) × as long as proximal width, inner margin concave, outer margin slightly convex, rounded distally, long pappose setae on outer and distal margins, medium-length setae on inner margin; with appendix interna curved upwards, slightly exceeding terminal margin of endopod by 0.23 of its length (Fig. 7 G, H). Appendix masculina of male second pleopod slender, reaching to proximal 0.6 × endopod length, 8.25 × as long as distal width, stick-shaped, with some short spiniform setae on outer surface and some long spiniform setae on distal surface; appendix interna at the middle of appendix masculina, narrow, small, extending ~ 0.60 length of appendix masculina (Fig. 7 I, J).</p><p>Coloration.</p><p>The body is white to slightly yellowish in color, with many small black spots and small lines of irregular sizes (Fig. 8).</p><p>Etymology.</p><p>The new species is named after the type locality, Xuan Lien National Park. The name is used as a noun in apposition.</p><p>Habitat.</p><p>This new species was found in the streams running in the forests. The substratum includes bedrock, stone, boulder, gravel, and sand in a water depth of 0.3–0.8 m (Fig. 9).</p><p>Distribution.</p><p>This species has been found only in the upland streams of Thuong Xuan District, Thanh Hoa Province, with elevations from 130 m to 600 m. The estimated area of occupancy is less than 1000 km 2.</p><p>Molecular phylogenetic results.</p><p>Caridina xuanlien sp. nov. is well supported as the sister species to C. clinata (Fig. 18). The minimum genetic divergence (p-distance) to C. clinata) is 4.0 % (COI) and 1.5 % (16 S), respectively (Suppl. materials 1, 2).</p><p>Remarks.</p><p>The new species is quite similar to C. nguyeni from Cao Bang province, northeast Vietnam, in the shape of the rostrum and endopod of male first pleopod (Li and Liang 2002). However, it can be separated by the number of dorsal teeth on the rostrum anterior to the orbital margin (3–9 vs 9–12), the number of ventral teeth of the rostrum (0–2 vs 1–4); the length of stylocerite (reaching to the beginning of the second segment of antennular peduncle vs reaching to beyond the middle to the end of second segment of the antennular peduncle); the appendix interna of the second male pleopod (~ 0.6 of the length of the appendix masculina vs ~ 0.4 length of appendix masculina); the uropodal diaeresis (with 18–21 movable spiniform setae vs with 12–18 movable spiniform setae) (Figs 6 A, B, 7 G, H; cf. Li and Liang 2002, figs 1 A, C, 2 C, B).</p><p>The new species resembles Caridina clinata Cai, Quynh &amp; Ng, 1999 in the shape of the endopod of male first pleopod, appendix masculine, and appendix interna of male second pleopod. However, C. xuanlien sp. nov. can be separated from C. clinata by a longer rostrum, reaching to the beginning of third segment of antennular peduncle (vs not reaching distal margin of basal segment of antennular peduncle); fewer number of teeth on the rostrum, armed with 3–9 dorsal teeth on anterior to the orbital margin and 0–2 ventral teeth (vs 10–16 dorsal teeth on anterior to the orbital margin and 2–5 ventral teeth); a longer stylocertite, reaching beginning of second segment of antennular peduncle (vs not reaching to distal margin of basal segment of antennular peduncle); slender endopod of male first pleopod, 2.75 × as long as proximal width (vs 2.5 × as long as proximal width); higher number of movable spiniform setae on uropodal diaresis, 18–21 (vs 14–17 movable spiniform setae); higher number of spiniform setae on flexor margin of the dactylus of fifth pereiopod, 37–40 (vs 45–49 spiniform setae on flexor margin) (Figs 6 A, B, G, 7 H, F; cf. Cai et al. 1999: figs 1 A, B, D, K, 2 F).</p></div>	https://treatment.plazi.org/id/A50CBB9C1CBD5F7390305EBBFC07B754	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Do, Van Tu;Phan, Thi Yen;von Rintelen, Kristina;Le, Hung Anh;von Rintelen, Thomas	Do, Van Tu, Phan, Thi Yen, von Rintelen, Kristina, Le, Hung Anh, von Rintelen, Thomas (2025): Descriptions of four new species of atyid shrimp (Crustacea, Decapoda, Atyidae) in Vietnam. ZooKeys 1247: 151-186, DOI: 10.3897/zookeys.1247.148607, URL: https://zookeys.pensoft.net/article/148607/download/xml/
