taxonID	type	description	language	source
350BA544F2575C63809A4BE99B4BF1DD.taxon	description	Figs 1, 2, 3 A, B	en	Wood, Kenneth R., Lorence, David H., Wagner, Warren L., Fawcett, Susan (2025): Geniostoma imadae (Loganiaceae), a new single-island endemic species from Kaua‘i, Hawaiian Islands. PhytoKeys 257: 191-207, DOI: 10.3897/phytokeys.257.154236
350BA544F2575C63809A4BE99B4BF1DD.taxon	diagnosis	Diagnosis. Geniostoma imadae is morphologically most similar to G. degeneri (Sherff) Byng & Christenh. from which it differs by its combination of leaves abaxially glabrous (vs. hirtellous), peduncle length (15 –) 20 – 75 mm long (vs. sessile), and capsule length 10 – 15 mm long with no keel (vs. 20 – 30 mm long with keel).	en	Wood, Kenneth R., Lorence, David H., Wagner, Warren L., Fawcett, Susan (2025): Geniostoma imadae (Loganiaceae), a new single-island endemic species from Kaua‘i, Hawaiian Islands. PhytoKeys 257: 191-207, DOI: 10.3897/phytokeys.257.154236
350BA544F2575C63809A4BE99B4BF1DD.taxon	description	Description. Shrubs or small trees, 1 – 3.5 m tall; trunk 2 – 5 cm diameter near base, bark gray to gray-brown; stems terete, lateral branches dichotomously branched, young and old stems glabrous. Leaves opposite, pinnately nerved, upper surface medium green, lower surface pale green; blade 3 – 12 (– 14) × 1.2 – 4.5 (– 5.5) cm, coriaceous, elliptic to elliptic-obovate, young and mature leaves glabrous on upper and lower surface, margins entire, apex apiculate to obtuse, base cuneate to acuminate, petioles (1 –) 2 – 15 (– 18) mm long; stipules interpetiolar, glabrous, completely connate, forming a truncate sheath 4 – 8 (– 11) mm long, usually splitting with age, adnate to the petiole at base. Flowers functionally unisexual, plants dioecious, inflorescence in pendulous, open paniculate cymes, flowers 3 – 8 in pistillate individuals, 3 – 10 in staminate plants, peduncles glabrous, recurved, (15 –) 20 – 40 mm long, elongating to 75 mm in fruit, pedicels glabrous, 10 – 40 mm long, elongating to 50 mm in fruit, bracts and bracteoles linear subulate, 2 – 5.5 × 0.1 – 0.8 (– 2.5) mm; calyx lobes 5 (– 6), connate near base, imbricate, linear-lanceolate or ovate-lanceolate, glabrous, apex acuminate, 5 – 9 × 0.8 – 3 mm, 5 – 7 nerved, margin hyaline; corolla salverform, 5 (– 6) - lobed, orange or yellow, fleshy, glabrous externally, inner tube pilose below throat, staminate and pistillate flowers 12 – 22 mm long, the tube 8 – 12 × 1 – 4 (– 5.5) mm, sparsely pilose within becoming densely pilose near throat, the lobes reflexed at anthesis, linear, 9 – 11 mm long, apex acuminate to acute; anthers 5 (– 6), adnate to corolla tube, dorsifixed, slightly exerted; staminate flowers with anthers 2.1 – 4 × 0.8 – 1.2 mm, reduced ovary glabrous 4.0 × 0.9 mm, pistil 12.5 – 18 mm long, stigma cylindrical and branched 5 – 14 mm long, style 2 – 4 mm long; pistillate flowers with anthers reduced, 1.2 – 1.8 × 0.4 – 0.8 mm, ovary glabrous 5.0 × 2.5 mm, pistil 12 – 12.5 mm long, stigma cylindrical and branched 5 mm long, style 2 mm long. Capsules green, light brown at maturity, ovoid-ellipsoid, apex acuminate, 10 – 15 mm long, 2 - valved, valves transversely wrinkled, not keeled, apex with beak, 2 – 4 mm long. Seeds ovoid-ellipsoid, brown, 1.2 – 1.5 × 0.7 – 0.8 mm, embedded in orange pulp.	en	Wood, Kenneth R., Lorence, David H., Wagner, Warren L., Fawcett, Susan (2025): Geniostoma imadae (Loganiaceae), a new single-island endemic species from Kaua‘i, Hawaiian Islands. PhytoKeys 257: 191-207, DOI: 10.3897/phytokeys.257.154236
350BA544F2575C63809A4BE99B4BF1DD.taxon	etymology	Etymology. Geniostoma is derived from the Greek geneion, beard, and stoma, mouth, referring to the hairs on the inner rim of the corolla (Smith and Stone 1962; Lorence and Wagner 2020). The epithet of this new species recognizes Clyde Imada, Research Specialist at the Bishop Museum. It is with great respect and admiration for his many contributions to Hawaiian botany that we name this species in his honor.	en	Wood, Kenneth R., Lorence, David H., Wagner, Warren L., Fawcett, Susan (2025): Geniostoma imadae (Loganiaceae), a new single-island endemic species from Kaua‘i, Hawaiian Islands. PhytoKeys 257: 191-207, DOI: 10.3897/phytokeys.257.154236
350BA544F2575C63809A4BE99B4BF1DD.taxon	distribution	Distribution and ecology. Geniostoma imadae is endemic to the volcanic island of Kaua‘i, where it is has been documented along the central northern ridges, slopes and riparian valleys of Wainiha, Lumaha‘i and Hanalei and extends south along the islands central eastern windward regions of ‘ Ili‘ili‘ula, ‘ Iole, Kamo‘oloa, Waiahi, and Wahiawa (Figs 3 C, 5). We estimate ca. 800 to 1250 individuals occur in lowland to montane wet forests ranging in elevation between 600 – 1234 m and dominated by trees of Metrosideros Banks ex Gaertn. (Myrtaceae) and Cheirodendron Nutt. ex Seem. (Araliaceae). Greater numbers of G. imadae are found along forested stream banks but also inhabit upper forested slopes that extend up to dividing ridges and border expansive patches of matting ferns such as Dicranopteris Bernh. and Diplopterygium (Diels) Nakai (Gleicheniaceae). Associated genera of trees and shrubs include Polyscias J. R. Forst. & G. Forst. (Araliaceae); Pritchardia Seem. & H. Wendl. (Arecaceae); Dubautia Gaudich. (Asteraceae); Cyanea Gaudich. (Campanulaceae), Perrottetia Kunth (Dipentodontaceae); Antidesma L., Euphorbia L. (Euphorbiaceae); Hydrangea Gronov. (Hydrangeaceae); Geniostoma (Loganiaceae); Myrsine L. (Primulaceae); Syzygium Gaertn. (Myrtaceae); Bobea Gaudich., Coprosma J. R. Forst. & G. Forst., Kadua Cham. & Schltdl., Psychotria L. (Rubiaceae); Melicope J. R. Forst. & G. Forst. (Rutaceae); and Pipturus Wedd., Touchardia Gaudich. (Urticaceae). Genera of sedges and grasses include Carex L., Cyperus L., Machaerina Vahl (Cyperaceae); Eragrostis Wolf, Panicum L. (Poaceae); herbs and sub-shrubs include Bidens L. (Asteraceae); Vaccinium L. (Ericaceae); Cyrtandra J. R. Forst. & G. Forst. (Gesneriaceae); and Freycinetia Gaudich. (Pandanaceae). Genera of ferns include Asplenium L., Hymenasplenium Hayata (Aspleniaceae); Deparia Hook. & Grev., Diplazium Sw. (Athyriaceae); Sadleria Kaulf. (Blechnaceae); Cibotium Kaulf. (Cibotiaceae); Microlepia C. Presl (Dennstaedtiaceae); Ctenitis (C. Chr.) C. Chr., Dryopteris Adans. (Dryopteridaceae); Hoiokula S. E. Fawc. & A. R. Sm., and Menisciopsis (Holttum) S. E. Fawc. & A. R. Sm. (Thelypteridaceae). There is also a diverse association of terrestrial and epiphytic lichens and bryophytes. The holotype of Geniostoma imadae was collected in the immediate area of the recently discovered and described species Melicope iolensis K. R. Wood, Lorence & W. L. Wagner (i. e., ‘ Iole Valley, Wood et al. 2024), attesting to the floristic diversity of the region and the importance of ongoing botanical inventories and conservation.	en	Wood, Kenneth R., Lorence, David H., Wagner, Warren L., Fawcett, Susan (2025): Geniostoma imadae (Loganiaceae), a new single-island endemic species from Kaua‘i, Hawaiian Islands. PhytoKeys 257: 191-207, DOI: 10.3897/phytokeys.257.154236
