taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
2E7E87E5125C835BCC96FF590B8FEE50.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15219073/files/figure.png	https://doi.org/10.5281/zenodo.15219073	FIGURE 4. Luciobarbus subquincunciatus, not preserved, 500 mm SL; Türkiye: Tigris at Cizre, 37.4041, 42.0709.	FIGURE 4. Luciobarbus subquincunciatus, not preserved, 500 mm SL; Türkiye: Tigris at Cizre, 37.4041, 42.0709.	2025-04-08	Freyhof, Jörg;Segherloo, Iraj Hashemzadeh;Vatandoust, Saber;Abdollahi-Mousavi, Seyed Eisa;Normandeau, Eric;Geiger, Matthias F.;Yoğurtçuoğlu, Baran		Zenodo	biologists	Freyhof, Jörg;Segherloo, Iraj Hashemzadeh;Vatandoust, Saber;Abdollahi-Mousavi, Seyed Eisa;Normandeau, Eric;Geiger, Matthias F.;Yoğurtçuoğlu, Baran			
2E7E87E5125D8370CC96FE7A0AFAE84C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15219067/files/figure.png	https://doi.org/10.5281/zenodo.15219067	FIGURE 1. Maximum Likelihood (ML) phylogram reconstructed for a common 503 bp segment of COI. Numbers along the branches are bootstrap support values for the ML tree (before slash) and Bayesian posterior probabilities (after slash). Only bootstrap values over 50 and posterior probabilities over 0.95 for branches which we discuss are presented. Red arrows denote the branches to which the bootstrap values and posterior probabilities correspond.Absolute support values of 100/1 are denoted with a star. Red circles represent individuals that align with L. esocinus morphologically. Brown (A1) and grey (A2) vertical bars in Clade A identify two sub-clades with 70% and 53% support values. Each individual is represented by species, drainage/basin, country, and GenBank accession number/BOLD sequence ID (denoted with black circles).	FIGURE 1. Maximum Likelihood (ML) phylogram reconstructed for a common 503 bp segment of COI. Numbers along the branches are bootstrap support values for the ML tree (before slash) and Bayesian posterior probabilities (after slash). Only bootstrap values over 50 and posterior probabilities over 0.95 for branches which we discuss are presented. Red arrows denote the branches to which the bootstrap values and posterior probabilities correspond.Absolute support values of 100/1 are denoted with a star. Red circles represent individuals that align with L. esocinus morphologically. Brown (A1) and grey (A2) vertical bars in Clade A identify two sub-clades with 70% and 53% support values. Each individual is represented by species, drainage/basin, country, and GenBank accession number/BOLD sequence ID (denoted with black circles).	2025-04-08	Freyhof, Jörg;Segherloo, Iraj Hashemzadeh;Vatandoust, Saber;Abdollahi-Mousavi, Seyed Eisa;Normandeau, Eric;Geiger, Matthias F.;Yoğurtçuoğlu, Baran		Zenodo	biologists	Freyhof, Jörg;Segherloo, Iraj Hashemzadeh;Vatandoust, Saber;Abdollahi-Mousavi, Seyed Eisa;Normandeau, Eric;Geiger, Matthias F.;Yoğurtçuoğlu, Baran			
2E7E87E5125D8370CC96FE7A0AFAE84C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15219075/files/figure.png	https://doi.org/10.5281/zenodo.15219075	FIGURE 5. Barbus rajanorum, NMW 54494, holotype, 190 mm SL; Syria: Aleppo. Modified from Heckel (1943).	FIGURE 5. Barbus rajanorum, NMW 54494, holotype, 190 mm SL; Syria: Aleppo. Modified from Heckel (1943).	2025-04-08	Freyhof, Jörg;Segherloo, Iraj Hashemzadeh;Vatandoust, Saber;Abdollahi-Mousavi, Seyed Eisa;Normandeau, Eric;Geiger, Matthias F.;Yoğurtçuoğlu, Baran		Zenodo	biologists	Freyhof, Jörg;Segherloo, Iraj Hashemzadeh;Vatandoust, Saber;Abdollahi-Mousavi, Seyed Eisa;Normandeau, Eric;Geiger, Matthias F.;Yoğurtçuoğlu, Baran			
2E7E87E5125D8370CC96FE7A0AFAE84C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15219077/files/figure.png	https://doi.org/10.5281/zenodo.15219077	FIGURE 6. Barbus rajanorum, NMW 54494, holotype, 190 mm SL; Syria: Aleppo.	FIGURE 6. Barbus rajanorum, NMW 54494, holotype, 190 mm SL; Syria: Aleppo.	2025-04-08	Freyhof, Jörg;Segherloo, Iraj Hashemzadeh;Vatandoust, Saber;Abdollahi-Mousavi, Seyed Eisa;Normandeau, Eric;Geiger, Matthias F.;Yoğurtçuoğlu, Baran		Zenodo	biologists	Freyhof, Jörg;Segherloo, Iraj Hashemzadeh;Vatandoust, Saber;Abdollahi-Mousavi, Seyed Eisa;Normandeau, Eric;Geiger, Matthias F.;Yoğurtçuoğlu, Baran			
2E7E87E5125D8370CC96FE7A0AFAE84C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15219079/files/figure.png	https://doi.org/10.5281/zenodo.15219079	FIGURE 7. FSJF 2929, 135 mm SL; hybrid between Luciobarbus schejch (male) and Capoeta damascina (female), with mtDNA of C. damascina; Türkiye: Euphrates drainage.	FIGURE 7. FSJF 2929, 135 mm SL; hybrid between Luciobarbus schejch (male) and Capoeta damascina (female), with mtDNA of C. damascina; Türkiye: Euphrates drainage.	2025-04-08	Freyhof, Jörg;Segherloo, Iraj Hashemzadeh;Vatandoust, Saber;Abdollahi-Mousavi, Seyed Eisa;Normandeau, Eric;Geiger, Matthias F.;Yoğurtçuoğlu, Baran		Zenodo	biologists	Freyhof, Jörg;Segherloo, Iraj Hashemzadeh;Vatandoust, Saber;Abdollahi-Mousavi, Seyed Eisa;Normandeau, Eric;Geiger, Matthias F.;Yoğurtçuoğlu, Baran			
2E7E87E5125D8370CC96FE7A0AFAE84C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15219081/files/figure.png	https://doi.org/10.5281/zenodo.15219081	FIGURE 8. Luciobarbus schejch, syntype, and an individual identified as L. mystaceus; Iraq: Mosul. Modified from Heckel (1943).	FIGURE 8. Luciobarbus schejch, syntype, and an individual identified as L. mystaceus; Iraq: Mosul. Modified from Heckel (1943).	2025-04-08	Freyhof, Jörg;Segherloo, Iraj Hashemzadeh;Vatandoust, Saber;Abdollahi-Mousavi, Seyed Eisa;Normandeau, Eric;Geiger, Matthias F.;Yoğurtçuoğlu, Baran		Zenodo	biologists	Freyhof, Jörg;Segherloo, Iraj Hashemzadeh;Vatandoust, Saber;Abdollahi-Mousavi, Seyed Eisa;Normandeau, Eric;Geiger, Matthias F.;Yoğurtçuoğlu, Baran			
2E7E87E5125D8370CC96FE7A0AFAE84C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15219083/files/figure.png	https://doi.org/10.5281/zenodo.15219083	FIGURE 9. Luciobarbus schejch, identified as L. mystaceus by Heckel (1843), NMW 54384/1, 233 mm; 54382/2, 260 mm SL; Iraq: Mosul.	FIGURE 9. Luciobarbus schejch, identified as L. mystaceus by Heckel (1843), NMW 54384/1, 233 mm; 54382/2, 260 mm SL; Iraq: Mosul.	2025-04-08	Freyhof, Jörg;Segherloo, Iraj Hashemzadeh;Vatandoust, Saber;Abdollahi-Mousavi, Seyed Eisa;Normandeau, Eric;Geiger, Matthias F.;Yoğurtçuoğlu, Baran		Zenodo	biologists	Freyhof, Jörg;Segherloo, Iraj Hashemzadeh;Vatandoust, Saber;Abdollahi-Mousavi, Seyed Eisa;Normandeau, Eric;Geiger, Matthias F.;Yoğurtçuoğlu, Baran			
2E7E87E5125D8370CC96FE7A0AFAE84C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15219085/files/figure.png	https://doi.org/10.5281/zenodo.15219085	FIGURE 10. Luciobarbus schejch, identified as L. mystaceus by Heckel (1843), NMW 54385/1, 157 mm; 54385/2, 142 mm SL; Syria: Aleppo.	FIGURE 10. Luciobarbus schejch, identified as L. mystaceus by Heckel (1843), NMW 54385/1, 157 mm; 54385/2, 142 mm SL; Syria: Aleppo.	2025-04-08	Freyhof, Jörg;Segherloo, Iraj Hashemzadeh;Vatandoust, Saber;Abdollahi-Mousavi, Seyed Eisa;Normandeau, Eric;Geiger, Matthias F.;Yoğurtçuoğlu, Baran		Zenodo	biologists	Freyhof, Jörg;Segherloo, Iraj Hashemzadeh;Vatandoust, Saber;Abdollahi-Mousavi, Seyed Eisa;Normandeau, Eric;Geiger, Matthias F.;Yoğurtçuoğlu, Baran			
2E7E87E5125D8370CC96FE7A0AFAE84C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15219087/files/figure.png	https://doi.org/10.5281/zenodo.15219087	FIGURE 11. NMW 6596, Barbus barbulus, syntype, about 125 mm SL; Iran: Qarah Aqaj, Mond drainage.	FIGURE 11. NMW 6596, Barbus barbulus, syntype, about 125 mm SL; Iran: Qarah Aqaj, Mond drainage.	2025-04-08	Freyhof, Jörg;Segherloo, Iraj Hashemzadeh;Vatandoust, Saber;Abdollahi-Mousavi, Seyed Eisa;Normandeau, Eric;Geiger, Matthias F.;Yoğurtçuoğlu, Baran		Zenodo	biologists	Freyhof, Jörg;Segherloo, Iraj Hashemzadeh;Vatandoust, Saber;Abdollahi-Mousavi, Seyed Eisa;Normandeau, Eric;Geiger, Matthias F.;Yoğurtçuoğlu, Baran			
2E7E87E5125D8370CC96FE7A0AFAE84C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15219089/files/figure.png	https://doi.org/10.5281/zenodo.15219089	FIGURE 12. Luciobarbus schejch, syntype, NMW 50399, 138 mm SL; Iraq: Mosul.	FIGURE 12. Luciobarbus schejch, syntype, NMW 50399, 138 mm SL; Iraq: Mosul.	2025-04-08	Freyhof, Jörg;Segherloo, Iraj Hashemzadeh;Vatandoust, Saber;Abdollahi-Mousavi, Seyed Eisa;Normandeau, Eric;Geiger, Matthias F.;Yoğurtçuoğlu, Baran		Zenodo	biologists	Freyhof, Jörg;Segherloo, Iraj Hashemzadeh;Vatandoust, Saber;Abdollahi-Mousavi, Seyed Eisa;Normandeau, Eric;Geiger, Matthias F.;Yoğurtçuoğlu, Baran			
2E7E87E5125D8370CC96FE7A0AFAE84C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15219091/files/figure.png	https://doi.org/10.5281/zenodo.15219091	FIGURE 13. Luciobarbus schejch, syntypes, NMW 54520, 269 mm SL; NMW 54520, 275 mm SL; Iraq: Mosul.	FIGURE 13. Luciobarbus schejch, syntypes, NMW 54520, 269 mm SL; NMW 54520, 275 mm SL; Iraq: Mosul.	2025-04-08	Freyhof, Jörg;Segherloo, Iraj Hashemzadeh;Vatandoust, Saber;Abdollahi-Mousavi, Seyed Eisa;Normandeau, Eric;Geiger, Matthias F.;Yoğurtçuoğlu, Baran		Zenodo	biologists	Freyhof, Jörg;Segherloo, Iraj Hashemzadeh;Vatandoust, Saber;Abdollahi-Mousavi, Seyed Eisa;Normandeau, Eric;Geiger, Matthias F.;Yoğurtçuoğlu, Baran			
2E7E87E5125D8370CC96FE7A0AFAE84C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15219093/files/figure.png	https://doi.org/10.5281/zenodo.15219093	FIGURE 14. Luciobarbus schejch, from top: FSJF 3682, 116 mm SL; Türkiye: stream Ambar; FSJF 2931, individual with mtDNA within COI clade A, 121 mm SL; Türkiye: stream Merziman; FSJF 3387, 143 mm SL, Iraq: Tabin; FSJF 2861, 195 mm, individual with mtDNA in COI clade A; Türkiye: Tigris at Hasankeyf; not preserved; 196 mm SL; Türkiye: Murat, 38.8681, 40.7926. No molecular data is available if not mentioned.	FIGURE 14. Luciobarbus schejch, from top: FSJF 3682, 116 mm SL; Türkiye: stream Ambar; FSJF 2931, individual with mtDNA within COI clade A, 121 mm SL; Türkiye: stream Merziman; FSJF 3387, 143 mm SL, Iraq: Tabin; FSJF 2861, 195 mm, individual with mtDNA in COI clade A; Türkiye: Tigris at Hasankeyf; not preserved; 196 mm SL; Türkiye: Murat, 38.8681, 40.7926. No molecular data is available if not mentioned.	2025-04-08	Freyhof, Jörg;Segherloo, Iraj Hashemzadeh;Vatandoust, Saber;Abdollahi-Mousavi, Seyed Eisa;Normandeau, Eric;Geiger, Matthias F.;Yoğurtçuoğlu, Baran		Zenodo	biologists	Freyhof, Jörg;Segherloo, Iraj Hashemzadeh;Vatandoust, Saber;Abdollahi-Mousavi, Seyed Eisa;Normandeau, Eric;Geiger, Matthias F.;Yoğurtçuoğlu, Baran			
2E7E87E5125D8370CC96FE7A0AFAE84C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15219095/files/figure.png	https://doi.org/10.5281/zenodo.15219095	FIGURE 15. Luciobarbus schejch, VPFC Horalazim 1396.11; 331 mm SL; 229 mm SL; Iran: Hor Alazim wetland.	FIGURE 15. Luciobarbus schejch, VPFC Horalazim 1396.11; 331 mm SL; 229 mm SL; Iran: Hor Alazim wetland.	2025-04-08	Freyhof, Jörg;Segherloo, Iraj Hashemzadeh;Vatandoust, Saber;Abdollahi-Mousavi, Seyed Eisa;Normandeau, Eric;Geiger, Matthias F.;Yoğurtçuoğlu, Baran		Zenodo	biologists	Freyhof, Jörg;Segherloo, Iraj Hashemzadeh;Vatandoust, Saber;Abdollahi-Mousavi, Seyed Eisa;Normandeau, Eric;Geiger, Matthias F.;Yoğurtçuoğlu, Baran			
2E7E87E5125D8370CC96FE7A0AFAE84C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15219097/files/figure.png	https://doi.org/10.5281/zenodo.15219097	FIGURE 16. Luciobarbus schejch, from top, left column: AJRPC Kut 1397.8; 259 mm SL; AJRPC Kut 1397.8; 201 mm; Iraq: Tigris at Kut; VPFC Shoshtar 1396.11; 310 mm SL, Iran: Karun. Right column: AJRPC Kut 1397.8; 178 mm SL, Iraq: Tigris at Kut; VPFC Asiaban 1396.11; 298 mm SL; Iran: Karkheh; VPFC Keshavarz 1396.11; about 400 mm SL; Iran: Dez. No molecular data is available for these fish.	FIGURE 16. Luciobarbus schejch, from top, left column: AJRPC Kut 1397.8; 259 mm SL; AJRPC Kut 1397.8; 201 mm; Iraq: Tigris at Kut; VPFC Shoshtar 1396.11; 310 mm SL, Iran: Karun. Right column: AJRPC Kut 1397.8; 178 mm SL, Iraq: Tigris at Kut; VPFC Asiaban 1396.11; 298 mm SL; Iran: Karkheh; VPFC Keshavarz 1396.11; about 400 mm SL; Iran: Dez. No molecular data is available for these fish.	2025-04-08	Freyhof, Jörg;Segherloo, Iraj Hashemzadeh;Vatandoust, Saber;Abdollahi-Mousavi, Seyed Eisa;Normandeau, Eric;Geiger, Matthias F.;Yoğurtçuoğlu, Baran		Zenodo	biologists	Freyhof, Jörg;Segherloo, Iraj Hashemzadeh;Vatandoust, Saber;Abdollahi-Mousavi, Seyed Eisa;Normandeau, Eric;Geiger, Matthias F.;Yoğurtçuoğlu, Baran			
2E7E87E5125D8370CC96FE7A0AFAE84C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15219105/files/figure.png	https://doi.org/10.5281/zenodo.15219105	FIGURE 20. Luciobarbus schejch, VPFC Sosangerd-Horalazim 2023.05 (FSJF-DNA 3769, COI clade A), 226 mm SL; Individual superficially similar to L. kersin; Iran: Fish market in Ahwaz, lower Karun.	FIGURE 20. Luciobarbus schejch, VPFC Sosangerd-Horalazim 2023.05 (FSJF-DNA 3769, COI clade A), 226 mm SL; Individual superficially similar to L. kersin; Iran: Fish market in Ahwaz, lower Karun.	2025-04-08	Freyhof, Jörg;Segherloo, Iraj Hashemzadeh;Vatandoust, Saber;Abdollahi-Mousavi, Seyed Eisa;Normandeau, Eric;Geiger, Matthias F.;Yoğurtçuoğlu, Baran		Zenodo	biologists	Freyhof, Jörg;Segherloo, Iraj Hashemzadeh;Vatandoust, Saber;Abdollahi-Mousavi, Seyed Eisa;Normandeau, Eric;Geiger, Matthias F.;Yoğurtçuoğlu, Baran			
2E7E87E5125D8370CC96FE7A0AFAE84C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15219107/files/figure.png	https://doi.org/10.5281/zenodo.15219107	FIGURE 21. Luciobarbus schejch, VPFC Horalazim 1392.11, 252 mm SL; individual similar to fish identified as L. kersin by Khaefi et al. (2018); Iran: Horalazim wetland. No molecular data is available.	FIGURE 21. Luciobarbus schejch, VPFC Horalazim 1392.11, 252 mm SL; individual similar to fish identified as L. kersin by Khaefi et al. (2018); Iran: Horalazim wetland. No molecular data is available.	2025-04-08	Freyhof, Jörg;Segherloo, Iraj Hashemzadeh;Vatandoust, Saber;Abdollahi-Mousavi, Seyed Eisa;Normandeau, Eric;Geiger, Matthias F.;Yoğurtçuoğlu, Baran		Zenodo	biologists	Freyhof, Jörg;Segherloo, Iraj Hashemzadeh;Vatandoust, Saber;Abdollahi-Mousavi, Seyed Eisa;Normandeau, Eric;Geiger, Matthias F.;Yoğurtçuoğlu, Baran			
2E7E87E5125D8370CC96FE7A0AFAE84C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15219127/files/figure.png	https://doi.org/10.5281/zenodo.15219127	FIGURE 31. Luciobarbus schejch, individuals with mtDNA in COI clade B and “pure” genome-wide SNP data of L. schejch, from top, VPFC Sosangerd-Horalazim 2023.05 (FSJF-DNA 3756), 375 mm SL; 15 total gill rakers and 57 total lateral-line scales; Iran: Karkheh; VPFC Gotvand 2023.05 (FSJF-DNA 3755), 326 mm SL; 12 total gill rakers and 57 total lateral-line scales; Iran: Karun; VPFC DezShoaybiye 2023.05 (FSJF-DNA 3745), 426 mm SL; 15 total gill rakers and 55 total lateral-line scales; Iran: Karun.	FIGURE 31. Luciobarbus schejch, individuals with mtDNA in COI clade B and “pure” genome-wide SNP data of L. schejch, from top, VPFC Sosangerd-Horalazim 2023.05 (FSJF-DNA 3756), 375 mm SL; 15 total gill rakers and 57 total lateral-line scales; Iran: Karkheh; VPFC Gotvand 2023.05 (FSJF-DNA 3755), 326 mm SL; 12 total gill rakers and 57 total lateral-line scales; Iran: Karun; VPFC DezShoaybiye 2023.05 (FSJF-DNA 3745), 426 mm SL; 15 total gill rakers and 55 total lateral-line scales; Iran: Karun.	2025-04-08	Freyhof, Jörg;Segherloo, Iraj Hashemzadeh;Vatandoust, Saber;Abdollahi-Mousavi, Seyed Eisa;Normandeau, Eric;Geiger, Matthias F.;Yoğurtçuoğlu, Baran		Zenodo	biologists	Freyhof, Jörg;Segherloo, Iraj Hashemzadeh;Vatandoust, Saber;Abdollahi-Mousavi, Seyed Eisa;Normandeau, Eric;Geiger, Matthias F.;Yoğurtçuoğlu, Baran			
2E7E87E5125D8370CC96FE7A0AFAE84C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15219129/files/figure.png	https://doi.org/10.5281/zenodo.15219129	FIGURE 32. Luciobarbus schejch, individuals with mtDNA in COI clade A, from top, left column: VPFC Shadegan 2023.05 (FSJF-DNA 3780), 257 mm SL; Iran: Jarrahi; VPFC Sosangerd-Horalazim 2023.05 (FSJF-DNA 3761), 301 mm SL; Iran: Karkheh. Right column: VPFC AhvazBazar 2023.05 (FSJF-DNA 3750), 384 mm SL; Iran: Karun; VPFC Dez7tape 2023.05 (FSJF-DNA 3736), 365 mm SL; Iran: Karun.	FIGURE 32. Luciobarbus schejch, individuals with mtDNA in COI clade A, from top, left column: VPFC Shadegan 2023.05 (FSJF-DNA 3780), 257 mm SL; Iran: Jarrahi; VPFC Sosangerd-Horalazim 2023.05 (FSJF-DNA 3761), 301 mm SL; Iran: Karkheh. Right column: VPFC AhvazBazar 2023.05 (FSJF-DNA 3750), 384 mm SL; Iran: Karun; VPFC Dez7tape 2023.05 (FSJF-DNA 3736), 365 mm SL; Iran: Karun.	2025-04-08	Freyhof, Jörg;Segherloo, Iraj Hashemzadeh;Vatandoust, Saber;Abdollahi-Mousavi, Seyed Eisa;Normandeau, Eric;Geiger, Matthias F.;Yoğurtçuoğlu, Baran		Zenodo	biologists	Freyhof, Jörg;Segherloo, Iraj Hashemzadeh;Vatandoust, Saber;Abdollahi-Mousavi, Seyed Eisa;Normandeau, Eric;Geiger, Matthias F.;Yoğurtçuoğlu, Baran			
2E7E87E5125D8370CC96FE7A0AFAE84C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15219131/files/figure.png	https://doi.org/10.5281/zenodo.15219131	FIGURE 33. Luciobarbus schejch, individuals with mtDNA in COI clade B, from top, left column:VPFC Sosangerd-Horalazim 2023.05 (FSJF-DNA 3760), 285 mm SL; Iran: Karkheh; VPFC Shadegan 2023.05 (FSJF-DNA 3774), 351 mm SL; Iran: Jarrahi. Right column: VPFC AhvazBazar 2023.05 (FSJF-DNA 3768), 174 mm SL; Iran: Karun; VPFC Shadegan 2023.05 (FSJF-DNA 3776), 217 mm SL; Iran: Jarrahi.	FIGURE 33. Luciobarbus schejch, individuals with mtDNA in COI clade B, from top, left column:VPFC Sosangerd-Horalazim 2023.05 (FSJF-DNA 3760), 285 mm SL; Iran: Karkheh; VPFC Shadegan 2023.05 (FSJF-DNA 3774), 351 mm SL; Iran: Jarrahi. Right column: VPFC AhvazBazar 2023.05 (FSJF-DNA 3768), 174 mm SL; Iran: Karun; VPFC Shadegan 2023.05 (FSJF-DNA 3776), 217 mm SL; Iran: Jarrahi.	2025-04-08	Freyhof, Jörg;Segherloo, Iraj Hashemzadeh;Vatandoust, Saber;Abdollahi-Mousavi, Seyed Eisa;Normandeau, Eric;Geiger, Matthias F.;Yoğurtçuoğlu, Baran		Zenodo	biologists	Freyhof, Jörg;Segherloo, Iraj Hashemzadeh;Vatandoust, Saber;Abdollahi-Mousavi, Seyed Eisa;Normandeau, Eric;Geiger, Matthias F.;Yoğurtçuoğlu, Baran			
2E7E87E5125D8370CC96FE7A0AFAE84C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15219133/files/figure.png	https://doi.org/10.5281/zenodo.15219133	FIGURE 34. Luciobarbus schejch, juvenile individuals with mtDNA in COI clade A. FSJF 3953, 67 mm SL; Iraq: Great Zab; FSJF 3382, 104 mm SL; Iraq: Lesser Zab.	FIGURE 34. Luciobarbus schejch, juvenile individuals with mtDNA in COI clade A. FSJF 3953, 67 mm SL; Iraq: Great Zab; FSJF 3382, 104 mm SL; Iraq: Lesser Zab.	2025-04-08	Freyhof, Jörg;Segherloo, Iraj Hashemzadeh;Vatandoust, Saber;Abdollahi-Mousavi, Seyed Eisa;Normandeau, Eric;Geiger, Matthias F.;Yoğurtçuoğlu, Baran		Zenodo	biologists	Freyhof, Jörg;Segherloo, Iraj Hashemzadeh;Vatandoust, Saber;Abdollahi-Mousavi, Seyed Eisa;Normandeau, Eric;Geiger, Matthias F.;Yoğurtçuoğlu, Baran			
2E7E87E5125D8370CC96FE7A0AFAE84C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15219135/files/figure.png	https://doi.org/10.5281/zenodo.15219135	FIGURE 35. Luciobarbus schejch, voucher of FSJF-DNA 2243, not preserved, individual with mtDNA in COI clade A; about 400 mm SL; Iraq: Ducan reservoir.	FIGURE 35. Luciobarbus schejch, voucher of FSJF-DNA 2243, not preserved, individual with mtDNA in COI clade A; about 400 mm SL; Iraq: Ducan reservoir.	2025-04-08	Freyhof, Jörg;Segherloo, Iraj Hashemzadeh;Vatandoust, Saber;Abdollahi-Mousavi, Seyed Eisa;Normandeau, Eric;Geiger, Matthias F.;Yoğurtçuoğlu, Baran		Zenodo	biologists	Freyhof, Jörg;Segherloo, Iraj Hashemzadeh;Vatandoust, Saber;Abdollahi-Mousavi, Seyed Eisa;Normandeau, Eric;Geiger, Matthias F.;Yoğurtçuoğlu, Baran			
2E7E87E5125D8370CC96FE7A0AFAE84C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15219137/files/figure.png	https://doi.org/10.5281/zenodo.15219137	FIGURE 36. Luciobarbus schejch, FSJF 3818, 290 mm SL; individual with mtDNA in COI clade B; Iraq: Ducan reservoir.	FIGURE 36. Luciobarbus schejch, FSJF 3818, 290 mm SL; individual with mtDNA in COI clade B; Iraq: Ducan reservoir.	2025-04-08	Freyhof, Jörg;Segherloo, Iraj Hashemzadeh;Vatandoust, Saber;Abdollahi-Mousavi, Seyed Eisa;Normandeau, Eric;Geiger, Matthias F.;Yoğurtçuoğlu, Baran		Zenodo	biologists	Freyhof, Jörg;Segherloo, Iraj Hashemzadeh;Vatandoust, Saber;Abdollahi-Mousavi, Seyed Eisa;Normandeau, Eric;Geiger, Matthias F.;Yoğurtçuoğlu, Baran			
2E7E87E5125D8370CC96FE7A0AFAE84C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15219099/files/figure.png	https://doi.org/10.5281/zenodo.15219099	FIGURE 17. Barbus kersin, NMW 54215, syntype, 163 mm SL; Syria: Aleppo. Modified from Heckel (1943).	FIGURE 17. Barbus kersin, NMW 54215, syntype, 163 mm SL; Syria: Aleppo. Modified from Heckel (1943).	2025-04-08	Freyhof, Jörg;Segherloo, Iraj Hashemzadeh;Vatandoust, Saber;Abdollahi-Mousavi, Seyed Eisa;Normandeau, Eric;Geiger, Matthias F.;Yoğurtçuoğlu, Baran		Zenodo	biologists	Freyhof, Jörg;Segherloo, Iraj Hashemzadeh;Vatandoust, Saber;Abdollahi-Mousavi, Seyed Eisa;Normandeau, Eric;Geiger, Matthias F.;Yoğurtçuoğlu, Baran			
2E7E87E5125D8370CC96FE7A0AFAE84C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15219103/files/figure.png	https://doi.org/10.5281/zenodo.15219103	FIGURE 19. Luciobarbus pectoralis, not preserved, about 100 mm SL; Syria: Orontes at Mashr’a al Bouz, 35.9508 36.3958; FSJF 2329; 160 mm SL; Türkiye: Seyhan drainage; FSJF 2697, 190 mm SL; Syria: Qattina Lake.	FIGURE 19. Luciobarbus pectoralis, not preserved, about 100 mm SL; Syria: Orontes at Mashr’a al Bouz, 35.9508 36.3958; FSJF 2329; 160 mm SL; Türkiye: Seyhan drainage; FSJF 2697, 190 mm SL; Syria: Qattina Lake.	2025-04-08	Freyhof, Jörg;Segherloo, Iraj Hashemzadeh;Vatandoust, Saber;Abdollahi-Mousavi, Seyed Eisa;Normandeau, Eric;Geiger, Matthias F.;Yoğurtçuoğlu, Baran		Zenodo	biologists	Freyhof, Jörg;Segherloo, Iraj Hashemzadeh;Vatandoust, Saber;Abdollahi-Mousavi, Seyed Eisa;Normandeau, Eric;Geiger, Matthias F.;Yoğurtçuoğlu, Baran			
2E7E87E5125D8370CC96FE7A0AFAE84C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15219101/files/figure.png	https://doi.org/10.5281/zenodo.15219101	FIGURE 18. Luciobarbus kersin, from top: ZMB 3237, syntype, 143 mm SL; NMW 54215, syntype, 163 mm SL; Syria: Aleppo.	FIGURE 18. Luciobarbus kersin, from top: ZMB 3237, syntype, 143 mm SL; NMW 54215, syntype, 163 mm SL; Syria: Aleppo.	2025-04-08	Freyhof, Jörg;Segherloo, Iraj Hashemzadeh;Vatandoust, Saber;Abdollahi-Mousavi, Seyed Eisa;Normandeau, Eric;Geiger, Matthias F.;Yoğurtçuoğlu, Baran		Zenodo	biologists	Freyhof, Jörg;Segherloo, Iraj Hashemzadeh;Vatandoust, Saber;Abdollahi-Mousavi, Seyed Eisa;Normandeau, Eric;Geiger, Matthias F.;Yoğurtçuoğlu, Baran			
2E7E87E5125D8370CC96FE7A0AFAE84C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15219109/files/figure.png	https://doi.org/10.5281/zenodo.15219109	FIGURE 22. Luciobarbus xanthopterus, NMW 54786, 340 mm SL; individual with snout length 1.9 times in postorbital length; Luciobarbus esocinus, NMW 54092, 321 mm SL; Iraq: Mosul. Modified from Heckel (1943: Plate IV).	FIGURE 22. Luciobarbus xanthopterus, NMW 54786, 340 mm SL; individual with snout length 1.9 times in postorbital length; Luciobarbus esocinus, NMW 54092, 321 mm SL; Iraq: Mosul. Modified from Heckel (1943: Plate IV).	2025-04-08	Freyhof, Jörg;Segherloo, Iraj Hashemzadeh;Vatandoust, Saber;Abdollahi-Mousavi, Seyed Eisa;Normandeau, Eric;Geiger, Matthias F.;Yoğurtçuoğlu, Baran		Zenodo	biologists	Freyhof, Jörg;Segherloo, Iraj Hashemzadeh;Vatandoust, Saber;Abdollahi-Mousavi, Seyed Eisa;Normandeau, Eric;Geiger, Matthias F.;Yoğurtçuoğlu, Baran			
2E7E87E5125D8370CC96FE7A0AFAE84C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15219111/files/figure.png	https://doi.org/10.5281/zenodo.15219111	FIGURE 23. Luciobarbus xanthopterus, NMW 54841/1, 216 mm SL; snout length 1.8 times in postorbital length; NMW 54786, lectotype, 340 mm SL; snout length 1.9 times in postorbital length Iraq: Mosul.	FIGURE 23. Luciobarbus xanthopterus, NMW 54841/1, 216 mm SL; snout length 1.8 times in postorbital length; NMW 54786, lectotype, 340 mm SL; snout length 1.9 times in postorbital length Iraq: Mosul.	2025-04-08	Freyhof, Jörg;Segherloo, Iraj Hashemzadeh;Vatandoust, Saber;Abdollahi-Mousavi, Seyed Eisa;Normandeau, Eric;Geiger, Matthias F.;Yoğurtçuoğlu, Baran		Zenodo	biologists	Freyhof, Jörg;Segherloo, Iraj Hashemzadeh;Vatandoust, Saber;Abdollahi-Mousavi, Seyed Eisa;Normandeau, Eric;Geiger, Matthias F.;Yoğurtçuoğlu, Baran			
2E7E87E5125D8370CC96FE7A0AFAE84C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15219115/files/figure.png	https://doi.org/10.5281/zenodo.15219115	FIGURE 25. Luciobarbus schejch with meristic counts in the overlapping range of L. esocinus and L. schejch. VPFC Dez7tape 2023.05; 357 mmm SL; 12 gill rakers, 59 total lateral-line scales, snout/postorbital 2.1; Iran: Karun; VPFC AhvazBazar 2023.05; 250 mmm SL; 14 gill rakers, 60 total lateral-line scales, snout/postorbital 1.8; Iran: Karun; VPFC AhvazBazar 2023.05¸ 211 mmm SL; 14 gill rakers, 61 total lateral-line scales, snout/postorbital ratio 2.4; Iran: Karun.	FIGURE 25. Luciobarbus schejch with meristic counts in the overlapping range of L. esocinus and L. schejch. VPFC Dez7tape 2023.05; 357 mmm SL; 12 gill rakers, 59 total lateral-line scales, snout/postorbital 2.1; Iran: Karun; VPFC AhvazBazar 2023.05; 250 mmm SL; 14 gill rakers, 60 total lateral-line scales, snout/postorbital 1.8; Iran: Karun; VPFC AhvazBazar 2023.05¸ 211 mmm SL; 14 gill rakers, 61 total lateral-line scales, snout/postorbital ratio 2.4; Iran: Karun.	2025-04-08	Freyhof, Jörg;Segherloo, Iraj Hashemzadeh;Vatandoust, Saber;Abdollahi-Mousavi, Seyed Eisa;Normandeau, Eric;Geiger, Matthias F.;Yoğurtçuoğlu, Baran		Zenodo	biologists	Freyhof, Jörg;Segherloo, Iraj Hashemzadeh;Vatandoust, Saber;Abdollahi-Mousavi, Seyed Eisa;Normandeau, Eric;Geiger, Matthias F.;Yoğurtçuoğlu, Baran			
2E7E87E5125D8370CC96FE7A0AFAE84C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15219117/files/figure.png	https://doi.org/10.5281/zenodo.15219117	FIGURE 26. Potential hybrids between L. esocinus and L. schejch, not preserved and not sequenced. From top: about 600 mm SL, 60 total lateral-line scales; Iraq: Dukan reservoir; about 500 mm SL, 55 total lateral-line scales; Iraq: Bagdad; photo by Omar F. Al-Sheikhly; about 450 mm SL, 60 total lateral-line scales; Iraq: Shatt-al Arab at Basra; photo by Atheer Ali.	FIGURE 26. Potential hybrids between L. esocinus and L. schejch, not preserved and not sequenced. From top: about 600 mm SL, 60 total lateral-line scales; Iraq: Dukan reservoir; about 500 mm SL, 55 total lateral-line scales; Iraq: Bagdad; photo by Omar F. Al-Sheikhly; about 450 mm SL, 60 total lateral-line scales; Iraq: Shatt-al Arab at Basra; photo by Atheer Ali.	2025-04-08	Freyhof, Jörg;Segherloo, Iraj Hashemzadeh;Vatandoust, Saber;Abdollahi-Mousavi, Seyed Eisa;Normandeau, Eric;Geiger, Matthias F.;Yoğurtçuoğlu, Baran		Zenodo	biologists	Freyhof, Jörg;Segherloo, Iraj Hashemzadeh;Vatandoust, Saber;Abdollahi-Mousavi, Seyed Eisa;Normandeau, Eric;Geiger, Matthias F.;Yoğurtçuoğlu, Baran			
2E7E87E5125D8370CC96FE7A0AFAE84C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15219119/files/figure.png	https://doi.org/10.5281/zenodo.15219119	FIGURE 27. Hybrid between L. esocinus and L. schejch, VPFC AhvazBazar 2023.05 (FSJF-DNA 3772), 257 mm SL; individual with mtDNA in COI clade A; Iran: Karun. SNP data place this individual close to, but not within, L. esocinus (compare with Fig. 28). Individual with 11 gill rakers, 58 total lateral-line scales, and snout length 1.7 times in postorbital length; Iran: Fish market in Ahwaz, lower Karun.	FIGURE 27. Hybrid between L. esocinus and L. schejch, VPFC AhvazBazar 2023.05 (FSJF-DNA 3772), 257 mm SL; individual with mtDNA in COI clade A; Iran: Karun. SNP data place this individual close to, but not within, L. esocinus (compare with Fig. 28). Individual with 11 gill rakers, 58 total lateral-line scales, and snout length 1.7 times in postorbital length; Iran: Fish market in Ahwaz, lower Karun.	2025-04-08	Freyhof, Jörg;Segherloo, Iraj Hashemzadeh;Vatandoust, Saber;Abdollahi-Mousavi, Seyed Eisa;Normandeau, Eric;Geiger, Matthias F.;Yoğurtçuoğlu, Baran		Zenodo	biologists	Freyhof, Jörg;Segherloo, Iraj Hashemzadeh;Vatandoust, Saber;Abdollahi-Mousavi, Seyed Eisa;Normandeau, Eric;Geiger, Matthias F.;Yoğurtçuoğlu, Baran			
2E7E87E5125D8370CC96FE7A0AFAE84C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15219121/files/figure.png	https://doi.org/10.5281/zenodo.15219121	FIGURE 28. Luciobarbus esocinus, FSJF 3819, juvenile, 220 mm SL; Iraq: Ducan Reservoir.	FIGURE 28. Luciobarbus esocinus, FSJF 3819, juvenile, 220 mm SL; Iraq: Ducan Reservoir.	2025-04-08	Freyhof, Jörg;Segherloo, Iraj Hashemzadeh;Vatandoust, Saber;Abdollahi-Mousavi, Seyed Eisa;Normandeau, Eric;Geiger, Matthias F.;Yoğurtçuoğlu, Baran		Zenodo	biologists	Freyhof, Jörg;Segherloo, Iraj Hashemzadeh;Vatandoust, Saber;Abdollahi-Mousavi, Seyed Eisa;Normandeau, Eric;Geiger, Matthias F.;Yoğurtçuoğlu, Baran			
2E7E87E5125D8370CC96FE7A0AFAE84C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15219123/files/figure.png	https://doi.org/10.5281/zenodo.15219123	FIGURE 29. Luciobarbus esocinus, VPFC Shadegan 1392.11, 285 mm SL; Iran: Shadegan wetland. Individual with 59 total lateral-line scales and 9 gill rakers. No molecular data.	FIGURE 29. Luciobarbus esocinus, VPFC Shadegan 1392.11, 285 mm SL; Iran: Shadegan wetland. Individual with 59 total lateral-line scales and 9 gill rakers. No molecular data.	2025-04-08	Freyhof, Jörg;Segherloo, Iraj Hashemzadeh;Vatandoust, Saber;Abdollahi-Mousavi, Seyed Eisa;Normandeau, Eric;Geiger, Matthias F.;Yoğurtçuoğlu, Baran		Zenodo	biologists	Freyhof, Jörg;Segherloo, Iraj Hashemzadeh;Vatandoust, Saber;Abdollahi-Mousavi, Seyed Eisa;Normandeau, Eric;Geiger, Matthias F.;Yoğurtçuoğlu, Baran			
2E7E87E5125D8370CC96FE7A0AFAE84C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15219125/files/figure.png	https://doi.org/10.5281/zenodo.15219125	FIGURE 30. Luciobarbus esocinus, VPFC AhvazBazar 2023.05 (FSJF-DNA 3773, COI clade A), 207 mm SL; individual with 9 gill rakers, 65 total lateral-line scales, and snout length 2.0 times in postorbital length; Iran: Fish market in Ahwaz, lower Karun.	FIGURE 30. Luciobarbus esocinus, VPFC AhvazBazar 2023.05 (FSJF-DNA 3773, COI clade A), 207 mm SL; individual with 9 gill rakers, 65 total lateral-line scales, and snout length 2.0 times in postorbital length; Iran: Fish market in Ahwaz, lower Karun.	2025-04-08	Freyhof, Jörg;Segherloo, Iraj Hashemzadeh;Vatandoust, Saber;Abdollahi-Mousavi, Seyed Eisa;Normandeau, Eric;Geiger, Matthias F.;Yoğurtçuoğlu, Baran		Zenodo	biologists	Freyhof, Jörg;Segherloo, Iraj Hashemzadeh;Vatandoust, Saber;Abdollahi-Mousavi, Seyed Eisa;Normandeau, Eric;Geiger, Matthias F.;Yoğurtçuoğlu, Baran			
2E7E87E5125D8370CC96FE7A0AFAE84C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15219071/files/figure.png	https://doi.org/10.5281/zenodo.15219071	FIGURE 3. Distribution of L. esocinus and L. schejch individuals along the first four principal component axes representing 54.7% of SNP variation: (a) PCI-II, (b) PCI-III, and (c) PCI-IV.The PCA graphs are combined with admixture results. Pie graphs corresponding to panel d (admixture graph) show genomic clusters of each individual. Red pie graphs denote L. esocinus; the red pie graph with a black outline is individual VPFC AhvazBazar 2023.05 / FSJF-DNA 3772 (Fig. 27), which is morphologically in the overlap of meristic counts between L. esocinus and L. schejch (11 total gill rakers, 58 total lateral-line scales) but shares a similar genome with L. esocinus; blue pie graphs represent L. schejch. A and B denote the mtDNA clade of each individual.	FIGURE 3. Distribution of L. esocinus and L. schejch individuals along the first four principal component axes representing 54.7% of SNP variation: (a) PCI-II, (b) PCI-III, and (c) PCI-IV.The PCA graphs are combined with admixture results. Pie graphs corresponding to panel d (admixture graph) show genomic clusters of each individual. Red pie graphs denote L. esocinus; the red pie graph with a black outline is individual VPFC AhvazBazar 2023.05 / FSJF-DNA 3772 (Fig. 27), which is morphologically in the overlap of meristic counts between L. esocinus and L. schejch (11 total gill rakers, 58 total lateral-line scales) but shares a similar genome with L. esocinus; blue pie graphs represent L. schejch. A and B denote the mtDNA clade of each individual.	2025-04-08	Freyhof, Jörg;Segherloo, Iraj Hashemzadeh;Vatandoust, Saber;Abdollahi-Mousavi, Seyed Eisa;Normandeau, Eric;Geiger, Matthias F.;Yoğurtçuoğlu, Baran		Zenodo	biologists	Freyhof, Jörg;Segherloo, Iraj Hashemzadeh;Vatandoust, Saber;Abdollahi-Mousavi, Seyed Eisa;Normandeau, Eric;Geiger, Matthias F.;Yoğurtçuoğlu, Baran			
