identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
2E0687B1D661224790E7FD11FD2FFC93.text	2E0687B1D661224790E7FD11FD2FFC93.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Holochlora Stal 1873	<div><p>Genus Holochlora Stål, 1873</p><p>Holochlora Stål, 1873: 43 .</p><p>Type species: Holochlora venosa Stål, 1873</p></div>	https://treatment.plazi.org/id/2E0687B1D661224790E7FD11FD2FFC93	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Wu, Chao;Liu, Chun-Xiang	Wu, Chao, Liu, Chun-Xiang (2025): The genus Holochlora Stål (Orthoptera, Tettigoniidae) in Gaoligong Mountains, China. Zootaxa 5716 (3): 432-442, DOI: 10.11646/zootaxa.5716.3.9, URL: https://doi.org/10.11646/zootaxa.5716.3.9
2E0687B1D661224590E7FC0AFCDCFBDD.text	2E0687B1D661224590E7FC0AFCDCFBDD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Holochlora qinlangdanga Wu & Liu 2025	<div><p>Holochlora qinlangdanga Wu &amp; Liu sp. nov.</p><p>Chinese vernacular name: ȗDzṡũŝ</p><p>(Figs. 1A–M, 5A–B, 6)</p><p>Type Material. Holotype. ♂, China: Yunnan Province, Dulongjiang Township (üżä乡), <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=98.2775&amp;materialsCitation.latitude=27.6887" title="Search Plazi for locations around (long 98.2775/lat 27.6887)">Maku Village</a> (Ḷṽ "), Qin’Lang’Dang (ȗDzṡ), N 27.6887° E 98.2775°, 1248m, 2.IX.2023, Chao Wu leg. (IZCAS) . Paratypes. 5♂, same locality as holotype, 2–9.IX.2023 Chao Wu &amp; Chun-Xiang Liu leg. (IZCAS) . 2♂, same locality as holotype, 2–7.X.2024 Chao Wu leg. (BFU) . 105♂, same locality as holotype, 2–9.X.2024 Chao Wu, Hai-Dong Bian &amp; Chun-Xiang Liu leg. (IZCAS) . 1♀, same locality as holotype, 5.X.2024, Chao Wu leg. (IZCAS) . 1♀, same locality as holotype, 3.X.2024, Chun-Xiang Liu leg. (IZCAS) .</p><p>Description. Male.</p><p>Head. Ovoid, elongate; occiput smooth, dorsally round. Compound eyes large, rounded, anteriorly protruding. Antennae slender, long, flexible, longer than body. Pronotum. Pronotum gradually tilting and widened backwards, with anterior margin straight and posterior margin faintly convex; lateral lobes deeper than long, anterior margin straight, posterior margin slightly arched, ventral margin short and slightly curved. Legs. Each leg slender, straight. Fore tibia with inner tympanum conchate, significantly swollen, opening of the inner tympanum very narrow, slit (Fig. 1G); with outer tympanum large, droplet shaped, the medial sclerite completely black (Fig. 1H). Hind leg very long. Hind femur enlarged at basal half, and gradually narrowed towards apex, slightly swollen near knee; ventral margin with 5–6 external spines, about 3 spines on apical half clearly larger; the knee without black area, external lobes bispinose, tip of the lobes with sharp spine (Fig. 1I). Hind tibia longer than femur; dense small spines present on both dorsal margins, and small spines on ventral margins relatively sparse. Wings. Tegmina and hind wings fully developed. Leaf-like tegmen opaque, very long, about 6.7 times longer than pronotum, slightly narrow, tapering towards rounded apex; hind wing with apex distinctively projecting beyond tegmen. Stridulatory file on left tegmen stout, droplet shaped, and slightly narrower at anal end (Fig. 1K); about 3 mm long, with about 25 equally-spaced teeth, gradually smaller in size towards anal end (Fig. 1J). Hind wing wide, transparent except for tip, colorless, with greenish pale veins. Abdomen. Short, compressed. Tenth abdominal tergite long, thick, much longer than wide, deeply split into two bulbous lobes in dorsal view; posterior margin broadly rounded, with a wide, circular gap in middle; significantly curved in lateral view. Cercus hairy, strongly flattened, short; a spine at tip, sharp, strongly curved inward. Subgenital plate elongate, basally broad slightly gradually narrowing towards end, deeply divided at proximal third; apical lobes narrow, roundly emarginated, with short, conical styli. (Figs. 1A–C).</p><p>Coloration. Generally fresh green when alive, lustrous. Compound eyes bluish-green. Antennae yellowish brown, except for green base. Costal vein on tegmen base light yellow, with indistinct black anterior edge (Fig. 5A). Hind wings hyaline, with green exposed part. Tympanum on fore tibia black; large spines on ventral margin of hind femora brown, with black spots at base; each tibia having a small black area at base. Tenth abdominal tergum and subgenital plate yellowish-brown (Fig. 1M), with a black spot on each side of the base of subgenital plate; cerci yellow, with a black tip.</p><p>Female. Generally similar to male, but body comparatively large-sized and robust. Subgenital plate short, nearly pentagonal shape, longer than wide, with a small, elongated extension on each side; apex of subgenital plate with small V-shaped notch. Valvifer 1 large, oviform, longer than subgenital plate. Ovipositor long, curved upwards; valvulae 3 with a sharp apex, and about 25 large teeth on dorsal margin; valvulae 1 with about 15 small teeth on apex of ventral margin, gradually lengthening distad (Fig. 1D–F). Body color similar to male, ovipositor yellowish-brown, with black tip.</p><p>Measurements (length in mm). Body (head to wings end): ♂ 52.87–54.15, ♀ 77.02–79.50; pronotum: ♂ 7.55–8.02, ♀ 8.67–8.98; tegmen: ♂ 50.95–52.23, ♀ 61.78–62.34; fore femora: ♂ 7.79–8.34, ♀ 10.60–10.92; hind femora: ♂ 30.36–31.85, ♀ 37.82–38.93; hind tibiae: ♂ 35.92–37.03, ♀ 42.46–43.30; stridulatory file: ♂ 2.58–2.61; ovipositor: ♀ 12.82–19.21.</p><p>Biology. A significant number of male individuals were attracted to the light trap, but there were only two females. During nocturnal observations, adults were noted to be actively present on the upper surfaces of leaves on tall trees. Many eggs were dissected from female specimens, and the mature egg shells were hard, purple brown, flat, and narrowed at both ends (Fig. 1L).</p><p>Discussion. Body large-size and robust for the genus. The new species is generally similar to Holochlora fruhstorferi Carl, 1914 (from Than Moi, Tonkin, i.e. north Vietnam), but male tenth abdominal tergum of and stridulatory file under male left tegmen is distinctly different; in addition, the new species has not a black area at the tip of each femur.</p><p>Etymology. The new species is named after the type locality Qin’Lang’Dang, Yunnan Province.</p><p>Distribution. China: Northwest Yunnan, Dulong River Basin.</p></div>	https://treatment.plazi.org/id/2E0687B1D661224590E7FC0AFCDCFBDD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Wu, Chao;Liu, Chun-Xiang	Wu, Chao, Liu, Chun-Xiang (2025): The genus Holochlora Stål (Orthoptera, Tettigoniidae) in Gaoligong Mountains, China. Zootaxa 5716 (3): 432-442, DOI: 10.11646/zootaxa.5716.3.9, URL: https://doi.org/10.11646/zootaxa.5716.3.9
2E0687B1D663224390E7FB41FCDBFE1D.text	2E0687B1D663224390E7FB41FCDBFE1D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Holochlora fugonga Wu & Liu 2025	<div><p>Holochlora fugonga Wu &amp; Liu sp. nov.</p><p>Chinese vernacular name: ạīũŝ</p><p>(Figs. 2A–I, 6)</p><p>Type Material. Holotype. ♂, China: Yunnan Province, Fugong County (ạī县), Shi’Yue’Liang (eh亮), N 27.2658° E 98.8972°, 1400m, 4.X.2024, Chao Wu leg. (IZCAS) . Paratypes. 1♂, same data (IZCAS) .</p><p>Description.</p><p>Male.</p><p>Head. Ovoid, elongate; occiput smooth, dorsally round. Compound eyes large, rounded, anteriorly protruding. Antennae slender, long, flexible, longer than body. Pronotum. Pronotum gradually tilting and widened backwards, with faintly convex anterior margin straight and posterior margins; lateral lobes deeper than long, with straight anterior margin, slightly arched posterior margin, short and slightly curved ventral margin. Legs. Each leg slender, straight. Fore tibia with conchate, significantly swollen inner tympanum, opening of inner tympanum very narrow, slit (Fig. 2D); with large and droplet shaped outer tympanum, interior of the outer tympanum completely black (Fig. 2E). Hind leg very long. Hind femur enlarged at basal half, and gradually narrowed towards apex, slightly swollen near knee; ventral margin with 8–10 external spines, about 5 spines on apical half clearly larger; the knee with a small black marking, external lobes bispinose, tip of the lobes with sharp spine. Hind tibia longer than the femur; dense small spines are present on the internal and external margins of the tibia dorsal, and small spines on the ventral margins are relatively sparse (Fig. 2F). Wings. Tegmina and hind wings fully developed. Leaf-like tegmen opaque, about 6.7 times longer than pronotum, slightly narrow, tapering towards rounded apex; hind wing with apex distinctively projecting beyond tegmen. Stridulatory file on left tegmen stout, droplet shaped, and slightly narrower at anal end (Fig. 2G); about 2.5 mm long, with about 40 equally spaced teeth, gradually smaller in size towards both ends (Fig. 2H). Hind wing wide, transparent except for tip, colorless, with greenish pale veins. Abdomen. Short, compressed. The tenth abdominal tergite short and thick, the length is less than the width; deeply split into two bulbous lobes in dorsal view, posterior margin broadly rounded, with a wide, circular gap in the middle; significantly curved shape in lateral view, ventral with a small protuberance on apical margin. Cercus hairy, strongly flattened, short; with sharp spine at the tip, strongly curved inward. Subgenital plate elongate, basally broad and slightly narrowing gradually towards the end, deeply divided at the proximal third; apical lobes narrow, roundly emarginated between lobes, with short, conical styli. (Fig. 2A–C).</p><p>Coloration. Generally yellowish-green when alive, lustrous. Compound eyes bluish green.Antennae yellowish-brown, except for green base. Costal vein on tegmen base light yellow and with very indistinct black anterior edge. Hind wings hyaline, with green exposed part. The tympanum on fore tibia black; all spines on the hind femora and tibia are brown, with black spots at the base of all spines, form dense spots on hind leg; each femora having small black area at the tip, this is particularly evident on the fore and hind femora. Tenth abdominal tergum and subgenital plate yellowish-brown, with a black spot on each side of the base of subgenital plate; cerci yellow, with a black tip. (Fig. 2I).</p><p>Female. Unknown.</p><p>Measurements (length in mm). Body (head to wings end): 59.17–60.23; pronotum: 7.60–7.82; tegmen: 51.25– 52.44; fore femora: 7.76–7.94; hind femora: 31.48–32.20; hind tibiae: 35.77–37.04; stridulatory file: 2.29–2.85.</p><p>Etymology. The new species is named after the type locality Fugong County, Yunnan Province.</p><p>Discussion. Body large-sized and robust for the genus. The new species is generally similar to Holochlora fruhstorferi Carl, 1914, H. lancangensis Liu, Zheng &amp; Xi, 1991 and H. qinlangdanga sp. nov. . However, the posterior tibiae of this new species have small black spots at the base of all spines (including dorsal and ventral spines), making them easily distinguishable; the tenth abdominal tergite of males is also different from the latter three species.</p><p>Distribution. China, Northwest Yunnan, Nujiang River Basin.</p></div>	https://treatment.plazi.org/id/2E0687B1D663224390E7FB41FCDBFE1D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Wu, Chao;Liu, Chun-Xiang	Wu, Chao, Liu, Chun-Xiang (2025): The genus Holochlora Stål (Orthoptera, Tettigoniidae) in Gaoligong Mountains, China. Zootaxa 5716 (3): 432-442, DOI: 10.11646/zootaxa.5716.3.9, URL: https://doi.org/10.11646/zootaxa.5716.3.9
2E0687B1D665224390E7FD81FCDBF835.text	2E0687B1D665224390E7FD81FCDBF835.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Holochlora gaoligonga Wu & Liu 2025	<div><p>Holochlora gaoligonga Wu &amp; Liu sp. nov.</p><p>Chinese vernacular name: ḕṷīũŝ</p><p>(Figs. 3A–H, 6)</p><p>Type Material. Holotype. ♂, China: Yunnan Province, Baoshan City, Bai’Hua’Ling <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=98.8004&amp;materialsCitation.latitude=25.3042" title="Search Plazi for locations around (long 98.8004/lat 25.3042)">Village</a> (Ħĭĸ), N 25.3042° E 98.8004°, 1575m, 6–8.X.2024, Chao Wu leg. (IZCAS) . Paratypes. 3♂, same data (IZCAS) .</p><p>Description.</p><p>Male.</p><p>Head. Ovoid, elongate, occiput smooth, dorsally round. Compound eyes large, rounded, anteriorly protruding. Antennae slender, long and flexible, longer than the body. Pronotum. Pronotum gradually tilting and widened backwards, with anterior margin straight and posterior margin faintly convex, lateral lobes deeper than long, anterior margin straight, posterior margin slightly arched, ventral margin short and slightly curved. Legs. Each leg slender and straight. Fore tibia with inner tympanum conchate, significantly swollen, opening of the inner tympanum very narrow, slit (Fig. 3D); with outer tympanum large and droplet shaped, the medial sclerite completely black (Fig. 3E). Hind leg very long; hind femur enlarged at basal half, and gradually narrowed towards apex, slightly swollen near knee, ventral margin with 8–10 external spines; the end of femur without black area, external lobes bispinose, tip of the lobes with sharp spine. Hind tibia longer than the femur; dense small spines are present on both dorsal margins, with small spines on the ventral margin are relatively sparse. Wings. Tegmina and hind wings fully developed. tegmen opaque, very long, about 6 times longer than pronotum, narrow, tapering towards rounded apex; hind wing with apex distinctively projecting beyond tegmen (Fig. 3F). Stridulatory file on left tegmen stout, droplet shaped, with a prominent hump in the middle of the posterior margin, with about 35 equally spaced teeth, progressively smaller in size towards anal end (Fig. 3G). Hind wing wide, transparent except for tip, colorless, with greenish pale veins. Abdomen. Short, compressed. Tenth abdominal tergum short, the dorsal side is flat and not raised, cleft in basal half into two apical lobes: both lobes with interior spines on the ventral surface, in lateral view apical margin with a small spines. Cerci conical, hairy, with sharp spine at the tip, strongly curved inward. Subgenital plate extending far beyond tenth abdominal tergum, widest at base, gradually tapering, split in apical sixth into two subparallel thin sheet-like lobes, with short, conical styli. (Fig. 3A–C).</p><p>Coloration. The body generally monochromatic, primarily light green in color. Antennae yellowish-brown. All legs and spines yellowish green, except for the tympanum on the fore tibia is black. The tenth abdominal tergum and subgenital plate are also yellowish green. (Fig. 3H)</p><p>Female. Unknown.</p><p>Measurements (length in mm). Body (head to wings end): 38.48–40.50; pronotum: 5.25–6.03; tegmen: 32.72– 34.45; fore femora: 6.28–7.01; hind femora: 24.62–25.89; hind tibiae: 27.64–29.28; stridulatory file: 1.56–1.88.</p><p>Etymology. The new species is named after the type locality Gaoligong Mt.</p><p>Discussion. Small-sized for the genus Holochlora . Entirely green, except tympanum area on the fore tibia black. The new species is similar to Holochlora nigrotympana Ingrisch, 1990, this species is recorded from Thailand to southern China and is also a small sized species within the genus Holochlora . However, the tenth abdominal tergite of H. nigrotympana has a distinct hump on the dorsal side, while the tenth abdominal tergite of the new species has a flat dorsal side.</p><p>Distribution. China, Northwest Yunnan, Nujiang River Basin.</p></div>	https://treatment.plazi.org/id/2E0687B1D665224390E7FD81FCDBF835	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Wu, Chao;Liu, Chun-Xiang	Wu, Chao, Liu, Chun-Xiang (2025): The genus Holochlora Stål (Orthoptera, Tettigoniidae) in Gaoligong Mountains, China. Zootaxa 5716 (3): 432-442, DOI: 10.11646/zootaxa.5716.3.9, URL: https://doi.org/10.11646/zootaxa.5716.3.9
2E0687B1D666224E90E7F9D3FC74FEA9.text	2E0687B1D666224E90E7F9D3FC74FEA9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Holochlora nigrospinulosa Brunner von Wattenwyl 1893	<div><p>Holochlora nigrospinulosa Brunner von Wattenwyl, 1893</p><p>Chinese vernacular name: DZ刺ũŝ</p><p>(Figs. 4A–H, 6)</p><p>Holochlora nigro-spinulosa Brunner von Wattenwyl, 1893: 170 .</p><p>Holochlora nigrospinulosa, Kirby 1906: 431; Ingrisch 1990 [1989]: 105; Liu, Liu &amp; Kang 2008: 232; Kang, Liu &amp; Liu 2014: 218.</p><p>Type locality: Carin Chebà (Myanmar) .</p><p>Material examined: 2♂ China: Yunnan Province, Baoshan City, Bai’Hua’Ling <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=98.8004&amp;materialsCitation.latitude=25.3042" title="Search Plazi for locations around (long 98.8004/lat 25.3042)">Village</a> (Ħĭĸ), N 25.3042° E 98.8004°, 1575m, 6–7.X.2024, Chao Wu leg. (IZCAS) .</p><p>Redescription. Habitus typical for the genus Holochlora, body medium-sized. Entirely green, tympanum on the fore tibia black (Fig. 4D, E), with small black spots on the hind leg knees. The tegmina are scattered with sporadic small black dots, and with a small black spot at the base of the R-vein (Fig. 4H). The tenth abdominal tergite of the male is divided into two bulbous lobes, and each lobe has a spine on its ventral surface that is closely attached to the lobe and points towards the base of the tenth abdominal tergite (Fig. 4A–C). Cercus of male hairy, short, with sharp spine at the tip, strongly curved inward; Subgenital plate elongate, broad basally and slightly narrowing gradually towards the end, styli conical, short. Ingrisch (1990 [1989]) initially reported this species as being distributed in northern Thailand, whereas Liu et al. (2008) and Kang et al. (2014) subsequently documented its occurrence in southern Yunnan, China. Although we have not examined the type specimens or those from the type locality of this species, the individuals collected from the Gaoligong Mountain region correspond well with the supplementary descriptions provided by Ingrisch (1990 [1989]), Liu et al. (2008), and Kang et al. (2014), including the measurement data of the specimens. This constitutes the first confirmed record of this species in the Gaoligong Mountain area.</p><p>Measurements (length in mm). Body (head to wings end): 52.87–54.25; pronotum: 6.47–7.12; tegmen: 45.23– 47.15; fore femora: 8.20–9.68; hind femora: 34.02–36.97; hind tibiae: 40.22–42.76; stridulatory file: 2.24–2.57.</p><p>Distribution. Myanmar, Northern Thailand, and Southwestern China.</p></div>	https://treatment.plazi.org/id/2E0687B1D666224E90E7F9D3FC74FEA9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Wu, Chao;Liu, Chun-Xiang	Wu, Chao, Liu, Chun-Xiang (2025): The genus Holochlora Stål (Orthoptera, Tettigoniidae) in Gaoligong Mountains, China. Zootaxa 5716 (3): 432-442, DOI: 10.11646/zootaxa.5716.3.9, URL: https://doi.org/10.11646/zootaxa.5716.3.9
