identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
3B7BDFCF2801595690A2A3719F6D4950.text	3B7BDFCF2801595690A2A3719F6D4950.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Exaesiopus therondi Lackner 2015	<div><p>Exaesiopus therondi Lackner, 2015</p><p>Figs 24–36, 37</p><p>Material examined.</p><p>♂, United Arab Emirates, FUJAIRAH [= Sharjah], Wadi al Helo, SW 1, KHOR KALBA, 24°53'833"N, 56°19'875"E, 25.ii.2007, G. Sama lgt., deposited in Penati collection at MSNG . A recently described species, the holotype of which was collected from the stomach of a Kentish plover [ Anarhynchus alexandrinus (Linnaeus, 1758), Aves: Charadriidae] in Afghanistan (Lackner 2015). This is the second known specimen (also male); new to United Arab Emirates. Its occurrence in the UAE suggests the species is widely spread, most likely present in Iran and Pakistan as well. For its better recognition, we show the morphological characteristics of the Arabian specimen (Figs 24–36).</p></div>	https://treatment.plazi.org/id/3B7BDFCF2801595690A2A3719F6D4950	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Zhang, Yejun;Lackner, Tomáš	Zhang, Yejun, Lackner, Tomáš (2025): Contribution to the Arabo-Saharan psammophilous Saprininae, with descriptions of two new species (Coleoptera, Histeridae, Saprininae, Hypocaccini). ZooKeys 1249: 289-301, DOI: 10.3897/zookeys.1249.160249
8A5C3593CF7C56D6A566D4A0C014389A.text	8A5C3593CF7C56D6A566D4A0C014389A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Neopachylopus penatii Zhang & Lackner 2025	<div><p>Neopachylopus penatii sp. nov.</p><p>Figs 20, 21, 37</p><p>Material examined.</p><p>• Holotype, female, side-mounted on triangular mounting card, ultimate right protarsomere and left metatarsomere broken off, with the following labels: “ female sign ” (printed); followed by: “ ALGERIE / Sidi Bel Abbés / collection le moult ” (printed); followed by: “ Muséum Paris / 1933 / Coll. Desbordes ” (light-turquoise label printed); followed by: “ Muséum national / Histoire nat. Paris / coll. générale ” (blue label, printed, black-framed); followed by: “ Neopachylopus / sp. nov. / det. Fabio Penati, 2022 ” (printed; black-framed label); followed by red, printed holotype label: “ Neopachylopus / penatii spec. nov. / Det. T. Lackner et Y. / Zhang 2025 ” (MNHN).</p><p>Description.</p><p>Body (Figs 20, 21) APW: 0.94 mm; PPW: 1.98 mm; EW: 2.11 mm; EL: 1.59 mm; PEL: 2.58 mm: oval, convex, pronotum dark brown, elytra lighter, castaneous; body appendages light brown, antennal club somewhat darker. Head: frontal disk slightly convex, glabrous; frontal stria almost straight, carinate, supraorbital and occipital striae fine, complete. Eyes flattened, almost invisible from above. Clypeus flattened, with several irregular, faint rugae; labrum typical for Hypocaccini, with two erect setae arising from each labral pit. Mandibles light brown, smooth, pointed apically, sub-apical tooth on left mandible large, perpendicular. Rest of visible mouthparts typical for the subfamily. Antennal club rather small, globular, without visible sensory structures; antennal scape with several long, thick yellow setae. Pronotum: pronotal hypomeron glabrous; marginal pronotal stria complete, carinate laterally, slightly weakened behind head. Pronotal disk weakly convergent anteriorly, apical angles blunt; disk glabrous, with several irregular punctures in ante-scutellar area. Elytra: elytral epipleuron glabrous, marginal epipleural stria fine, complete; marginal elytral stria fine, complete, continuous with complete apical elytral stria that is continuous with complete sutural elytral stria. Elytral humeri inconspicuous, humeral elytral stria very thin, present on basal elytral fifth; outer subhumeral stria present as a short median fragment. Dorsal elytral striae I – IV in punctures, of sub-equal length, reaching approximately to elytral third to half apically; stria IV basally connected with sutural elytral stria. Elytral disk on apical half (approximately) with dense, large punctures separated by 0.5–1.5 times their diameter; elytral intervals and flanks impunctate; punctures stopping short of elytral apex, leaving a narrow impunctate band. Propygidium: partly covered by elytra, with dense, deep punctures separated by less than their diameter; interspaces with microsculpture; pygidium triangular, medially convex, with sparser and finer punctures than those of propygidium. Prosternum: very compressed, knife-like, carinal prosternal striae between procoxae very approximate, thence running parallel, diverging and again convergent apically. Lateral prosternal striae short, carinate, attaining carinal prosternal striae at their mid-length; prosternal foveae tiny, rather deep; prosternal process concave. Mesoventrite: subquadrate, mesoventral disk with irregular punctures of unequal sizes; marginal mesoventral stria complete. Meso-metaventral suture with distinct punctate stria. Metaventrite: broad, short, slightly concave medially, intercoxal disk glabrous, near apical margin with a band of irregular punctures of various sizes. Lateral disk of metaventrite with large ocelloid shallow setigerous punctures, interspaces with alutaceous microsculpture. Lateral metaventral stria short, carinate, straight. Metepisternum with very dense, large setigerous punctures. First visible abdominal ventrite completely striate laterally; with irregular punctures of various sizes on basal fourth; rest of disk almost glabrous; along apical edge a row of tiny punctures appears. Legs: protibial apex with two prominent, rounded denticles, followed by a distant short tooth topped by similar denticle, followed by a dense row of six regular, elongate denticles diminishing in size in proximal direction. Protarsal spur short, inconspicuous, anterior protibial margin with three short, thin denticles; protibial groove deep, straight; protarsus thin, ultimate protarsomere shorter than two preceding together; claws short, thin, bent. Mesotibia thickened, outer margin with several rows of dense, thick set denticles on outer margin, growing in distal direction; mesotibial spur long, stout; mesotarsus thickened, each mesotarsomere with long, thick seta; mesotarsal claws thin, bent, shorter than half of ultimate mesotarsomere. Metatibia triangularly dilated and thickened, otherwise like mesotibia; metafemora thickened. Male unknown.</p><p>Distribution.</p><p>Known only from Sidi Bel Abbès, near Oran, north-western Algeria (Fig. 37). Possibly distributed also into Morocco.</p><p>Etymology.</p><p>Named after our colleague Fabio Penati (Morbegno, Italy), who first recognised this unique taxon.</p><p>Etymology.</p><p>Unknown.</p><p>Remarks.</p><p>The glabrous frontal disk, knife-like prosternal process and peculiar shape of protibia (protibial apex with two prominent rounded denticles, followed by a distant short tooth topped by similar denticle) clearly separate this taxon from members of genus Hypocaccus, which it superficially resembles most. The smooth, impunctate pronotum would suggest affinity to the subgenus Baeckmanniolus Reichardt, 1926 of the aforementioned genus. Yet by the above-mentioned characters, N. penatii sp. nov. cannot be classified within it. Hesitant to erect a new genus, we decided to include it in the heterogeneous Neopachylopus, pending further studies (see also Discussion).</p></div>	https://treatment.plazi.org/id/8A5C3593CF7C56D6A566D4A0C014389A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Zhang, Yejun;Lackner, Tomáš	Zhang, Yejun, Lackner, Tomáš (2025): Contribution to the Arabo-Saharan psammophilous Saprininae, with descriptions of two new species (Coleoptera, Histeridae, Saprininae, Hypocaccini). ZooKeys 1249: 289-301, DOI: 10.3897/zookeys.1249.160249
B4207EC0F4CF50EEA81844487E97FE42.text	B4207EC0F4CF50EEA81844487E97FE42.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Neopachylopus pharkidodes Zhang & Lackner 2025	<div><p>Neopachylopus pharkidodes sp. nov.</p><p>Figs 1, 2–7, 8–12, 13–19, 37</p><p>Material examined.</p><p>• Holotype, male, side-mounted on triangular mounting card, right protarsus and left metatarsus broken off, latter glued to the same mounting card as specimen, male genitalia extracted, dismembered and glued to a separate mounting card under the specimen, with the following labels: “ QATAR-Madinat Al Shamal / <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=51.35&amp;materialsCitation.latitude=26.066668" title="Search Plazi for locations around (long 51.35/lat 26.066668)">Al Ghariyah</a> 9.III.2003 / 26°04'N, 51°21'E G. Sama ” (printed); followed by: “ MUSEO GENOVA / ex. coll. G. Sama / (acquisto 2008) ” (printed); followed by yellow, pencil-written label “ 10-205 ” added by T. Lackner in 2010; followed by red, printed holotype label: “ Neopachylopus / pharkidodes spec. / nov. Det. T. Lackner / et Y. Zhang 2025 ”. The type is deposited in the Penati collection at MSNG . • Paratype female, with the following labels: “ S. Arabia: / Kamaran I. / 7-11 - 1903 / Dr. M. Cameron / B. M. 1928-109 ” (printed-written), followed by printed label “ Sea Weed ”, followed by a pencil-written label: “ Neopachylopus / sp. nov. 2009 / Det. T. Lackner ”, followed by yellow, pencil-written label: “ 09-048 ”, added by T. Lackner in 2009; followed by red, printed paratype label: “ Neopachylopus / pharkidodes spec. / nov. Det. T. Lackner / et Y. Zhang 2025 ” (MNHN) .</p><p>Description.</p><p>Body (Fig. 1): APW: 0.73–0.77 mm; PPW: 1.65–1.69 mm; EW: 1.85–1.91 mm; EL: 1.44–1.48 mm; PEL: 2.35–2.59 mm. Body oval, convex, reddish-brown, dorsal surface of head and pronotum fuscous; body appendages lighter. Head (Fig. 2): frons with microsculpture and indistinct microscopic punctures; medial area with several coarse and intersecting grooves. Frontal stria carinate and complete, anterior part feebly arcuate forward, supraorbital stria extending past posterior margin of eye. Epistoma coarse, carinately margined, depressed medially. Labrum transverse, anterior margin straight medially; setae inconspicuous (broken off?). Mandibles stout, curved, tips acute, left mandible with a distinct sub-apical tooth on inner margin, right mandible with an indistinct sub-apical tooth on inner margin. Mentum (Fig. 3) rectangular, anterior margin inwardly arcuate; eyes flattened, invisible from above. Antennal club (Fig. 4) globose and tomentose, sensory structures not examined. Pronotum (Fig. 1): sides nearly straight and slightly convergent forward in posterior two-thirds, strongly arcuate in anterior third; anterior margin emarginate and evenly arcuate; posterior margin nearly straight laterally, forming an indistinct angle medially. Marginal pronotal stria carinate, complete. Surface densely and coarsely impressed with longitudinal striae and rugae; occasionally intersecting and arcuate medially; a smooth, narrow band along margins present; ante-scutellar area with a single coarse and deep round puncture. Pronotal hypomeron glabrous. Scutellum small, triangular. Elytra (Fig. 1): relatively straight, humeri slightly projecting. Dorsal elytral stria I almost complete, slightly weakened and briefly interrupted on basal third in holotype; dorsal elytral stria II present in basal 2 / 3; dorsal elytral striae III – IV sub-equal in length, stopping short of elytral half apically; stria IV basally connected with complete sutural elytral stria, which is linked with complete, albeit somewhat weakened apical elytral stria, which, on the other hand is linked to complete and carinate marginal elytral stria. Humeral elytral stria smooth, deeply impressed in basal fourth; inner subhumeral stria present as short median fragment; outer subhumeral stria absent; all striae (except humeral one) feebly crenate. Elytral disk micro-sculptured and irregularly scattered with small punctures. Elytral epipleuron microscopically punctate, almost smooth; marginal epipleural stria deeply and completely impressed. Abdomen: propygidium and pygidium (Fig. 5) with scattered tiny shallow punctures. Prosternum (Fig. 6): anterior margin feebly protruding and slightly extending ventrally, posterior margin nearly straight. Prosternal process narrow, knife-like, carinal prosternal striae very approximate, running sub-parallel, anterior ends joined. Prosternal process with microsculpture and scattered small punctures near anterior margin and laterad of carinal striae; prosternal foveae small, well visible. Lateral prosternal striae short, carinate, convergent and attaining carinal prosternal striae at their apical two-thirds. Mesoventrite (Fig. 7): subtrapezoid; disk glabrous. Marginal mesoventral stria complete and straight laterally, absent anteriorly. Meso-metaventral suture well impressed, medially angulate. Intercoxal disk of metaventrite smooth, like mesoventrite; longitudinal suture deeply impressed, surface around it slightly depressed. Lateral metaventral stria rather short; lateral disk of metaventrite (Fig. 8) slightly depressed, with shallow setigerous punctures. Metepisternum (Fig. 8) with coarser and denser punctures bearing long amber setae. Intercoxal disk of first visible abdominal sternite slightly longer than metaventrite, smooth, incompletely striate laterally. Legs: profemur slightly slender; protibia (Figs 9, 10) broad and flat, outer margin with four teeth bearing blunt denticle, followed by another four tiny denticles entombed in outer protibial margin; all diminishing in size in proximal direction. Anterior face of protibia (Fig. 10) with rib-like structures. Mesofemora stout; mesotibia rather slender, outer margin with two rows of thickly set denticles; mesotarsus relatively thick, slightly longer than protarsus. Metafemora rather stout, sub-semicircular; metatibia (Figs 11, 12) dilated and thickened, outer margin with four rows of densely set short denticles; metatarsus like mesotarsus. Male genitalia: aedeagus (Figs 16, 17) basal piece short, ratio to parameres approximately 1: 4; parameres on their basal half (approximately) parallel-sided, thence slightly divergent and convergent again apically, apex blunt; aedeagus slightly curved from ventral view. Rest of male terminalia (Figs 13–15, 18, 19) typical for Hypocaccini; eighth sternite and tergite not fused laterally (Fig. 19); apex of eighth sternite with few short setae.</p><p>Female genitalia not studied.</p><p>Distribution.</p><p>Known from Qatar and Yemeni Island of Kamaran; these two localities are geographically widely separated (about 1430 km; Fig. 37); the lack of specimens from between these two points can most likely be attributed to inadequate sampling, thus the species is probably present along the entire coast of the southern Arabian Peninsula.</p><p>Etymology.</p><p>From the Greek word pharkis, meaning wrinkle, pharkidodes, wrinkled; named after the rugulose pronotal disk.</p><p>Etymology.</p><p>Collected under seaweed.</p><p>Remarks.</p><p>A morphologically unique taxon. The rugulose pronotal dorsum of N. pharkidodes sp. nov. is unmatched among all Saprininae taxa known to the senior author (see also Key to species and Discussion).</p></div>	https://treatment.plazi.org/id/B4207EC0F4CF50EEA81844487E97FE42	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Zhang, Yejun;Lackner, Tomáš	Zhang, Yejun, Lackner, Tomáš (2025): Contribution to the Arabo-Saharan psammophilous Saprininae, with descriptions of two new species (Coleoptera, Histeridae, Saprininae, Hypocaccini). ZooKeys 1249: 289-301, DOI: 10.3897/zookeys.1249.160249
