identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
345E87D7FFC4FFFBFF47FAFAFCB0FE51.text	345E87D7FFC4FFFBFF47FAFAFCB0FE51.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Seracamaurops (Seracamaurops) cvorovici Hlaváč & Brachat & Delić 2025	<div><p>Seracamaurops (Seracamaurops) cvorovici sp. nov.</p><p>Figs 1–3</p><p>Type material. HOLOTYPE: MONTENEGRO: ♂: one label “MONTENEGRO XB239 / Studena Pećina / 42.927034, 19.07300 / <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=19.073&amp;materialsCitation.latitude=42.927032" title="Search Plazi for locations around (long 19.073/lat 42.927032)">Mokro Polje</a>, Šavnik / 30.IX.2023 T. Delić lgt.“ [white, printed] (PCPH) . PARATYPES: MONTENEGRO: 3 ♂♂, same data as for holotype, 2 specimens bearing the following voucher codes: XB240 and XB250. (DBBF, NHMM, PCPH) .</p><p>Diagnosis. Body shiny, unicolorous dark reddish-brown; head capsule elongate; eyes completely atrophied with minuscule ocular spine; lacking ocular carinae; vertexal foveae large; antennae when bent backwards, almost reaching half of length of elytra; all antennomeres elongate; pronotum with shallow median impression and well-defined lateral foveae; lacking antebasal sulcus; elytra trapezoidal; abdomen about 1.2 times as long as elytra and wider than elytra; anterior part of first visible tergite 1 (IV) with one median and two lateral, subequal well-separated deep impressions; external stria diffuse with tergal margin, internal stria fine, very short, almost invisible. Aedeagus with right distal part of median lobe bird's head with a beak like, with more pre-distal setae; left distal part of median lobe with two slender, long, inward-oriented projections.</p><p>Description. Body (Fig. 1) shiny, unicolorous dark reddish-brown, smooth with long setae; length about 3.1 mm, maximum width 0.95 mm; whole body almost smooth, with only very sparse, almost invisible and unevenly distributed minuscule punctures.</p><p>Head capsule elongate, about 1.2 times as long as wide, as long as pronotum; lacking rostrum; frons with deep excavation between prominent antennal tubercles; eyes completely atrophied with minuscule ocular spine; lacking ocular carinae; vertexal foveae large, deep, well-separated by slightly convex surface; temples with long, erect setae, sharply arcuately convergent to posterior margin of head capsule, here width is about 0.35 of maximum width of head capsule; maxillary palpi small, palpomere 2 pedunculate, about as long as 4 and 3, triangular, smallest, palpomere 4 fusiform, 0.27 mm long, about 3.4 times as long as wide. Antennae about 1.5 mm long, when bent backwards, almost reaching half of length of elytra; all antennomeres elongate, scape about 1.4 times as long as pedicel, pedicel 1.5 times as long as 3, antennomeres 3, 4, 6 and 10 subequal in length, 5 twice as long as wide, 1.27 times as long as 4 and 6, antennomere 7 slightly longer than 6 and 1.5 times as long as 8 and 9, 9 as long as wide, 10 about 1.2 as long as wide, terminal antennomere longest, pointed apically, about 1.5 times as long as 10.</p><p>Pronotum about 1.2 times as long as wide, widest in middle; with shallow median impression and well-defined lateral foveae; lacking antebasal sulcus.</p><p>Elytra trapezoidal, with slightly rounded lateral margins; 1.15 times as long as wide, about 1.2 times as long as pronotum.</p><p>Abdomen about 1.2 times as long as elytra, wider than elytra; first visible tergite 1 (IV) trapezoidal, evenly divergent posteriad; clearly longer than rest of visible tergites, on anterior margin with one median and two lateral, subequal deep well separated impressions; external stria diffuse with tergal margin, internal stria fine, very short, almost invisible.</p><p>Legs long and slender, without any modification.</p><p>Aedeagus (Figs 2, 3) 0.63 mm long, about 2.5 times as long as wide, basal part subcircular, about 0.4 of length of aedeagus; in ventral view, right distal part of median lobe bird's head with a beak like, with more pre-distal setae; left distal part of median lobe with two slender, long, inward-oriented projections; diaphragm large, subcircular.</p><p>Sexual dimorphism. Female unknown.</p><p>Habitat. All specimens were found “floating” on the water surface of the syphon, at the end of the so-called “meandarski kanal” in Studena pećina, Morko polje, at the approximate distance of 350 m from the cave entrance.</p><p>Etymology. The origin of the name is patronymic, dedicated to Petar and Svetomir Čvorović, the locals who showed us the entrance to the cave and generously expressed their hospitality to the last author.</p><p>Remarks. Seracamaurops cvorovici is classified in the nominal subgenus having dorsal and ventral parts of the first visible tergite (IV) delimited by a well-defined lateral margin, and a pronotum lacking discal carinae, lateral spines and a median sulcus. Identification of species of this subgenus requires examination of the aedeagus, where the main distinguishing features are the shape of the left and right distal part of the median lobe. The aedeagus of S. cvorovici has similar structure as S. delici Lohaj, Pavićević &amp; Lakota, 2023, both having the right distal part of median lobe bird's head with a beak like, although the beak of S. delici is very long and slender with oval top of head, in S. cvorovici is short and the top of head is truncate; the left distal part of S. delici and S. cvorovici differs by length of external processes as well as by their orientation.</p></div>	https://treatment.plazi.org/id/345E87D7FFC4FFFBFF47FAFAFCB0FE51	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hlaváč, Peter;Brachat, Volker;Delić, Teo	Hlaváč, Peter, Brachat, Volker, Delić, Teo (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXIX. Further contribution to the taxonomy of the tribe Amauropini (Coleoptera: Staphylinidae: Pselaphinae) from the Balkan Peninsula. Zootaxa 5665 (1): 137-143, DOI: 10.11646/zootaxa.5665.1.10, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
345E87D7FFC6FFFCFF47FAE7FEAEFCE9.text	345E87D7FFC6FFFCFF47FAE7FEAEFCE9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protamaurops peristeri Karaman 1961	<div><p>Protamaurops peristeri Karaman, 1961, stat. resurr.</p><p>Fig. 5</p><p>Protamaurops peristeri Karaman, 1961: 159, 168. Type data: Na Peristeru, jedan mužjak [one male].</p><p>Protamaurops peristeri Karaman: Bekchiev &amp; Hlaváč, 2020: 376 (as synonym of P. winneguthi (Székessy, 1943)) . Protamaurops winneguthi Székessy: Besuchet, 1999: 48 (synonymy).</p><p>Material examined. NORTH MACEDONIA: 1♂: four labels “ Maced. Perister / Sv. Petka, 7.-14 / Dr. Rambousek “ [white, printed], “ Am. Roubal / purkyňei / R. Rous det., 1965“ [white, printed and handwritten], “Ex. Coll. R. Rous “ [white, printed], “Sammlung / Brachat Geretsried“ [yellow, printed] (PCVB) .</p><p>Description. Body length 2.62 mm, maximum width of elytra 0.77 mm; slightly shiny, reddish-brown, densely pubescent, covered with long golden setae; legs, antennae and maxillary palpi slightly lighter. Head capsule about as long as wide, eyes composed from about 10 ommatidia located around ocular spine; clypeus large, round; frons with large, deep excavation that is slightly prolonged anteriorly, forming angled, distal projection; vertex elevated, forming large hump delimited on sides by parallel carinae and with one well-defined, long medial carina; vertexal pits separated, adjacent to parallel carinae and distant from lateral margin of head capsule; surface between carinae almost glabrous; antennae 1.05 mm long, scape about 1.4 times as long wide, pedicel about 1.6 times as long as wide, antennomere 3 1.33 times as long as wide, 4 slightly shorter than 3, 5 1.5 times as long as wide, about 1.3 times as long as 4, antennomeres 3, 6, 7, 10 subequal in length, 9 1.15 times as long as wide, 10 transverse, 1.25 times as wide as long, terminal antennomere subtriangular, 2.5 times as long as 10. Pronotum about 1.15 times as long as wide, as long as head capsule, surface between lateral carinae flat, with deep median fovea, its posterior margin V-shaped, posterior angle of median fovea prolonged by short median carina reaching posterior margin of pronotum, together with V shaped margin of median fovea Y-shaped; lateral carinae long and weekly prominent, lacking triangular dent in posterior part; lateral foveae well defined, deep. Elytra slightly wider than long, at sides round, about 1.45 times as long as pronotum, widest in posterior part, shortly behind middle; with three antebasal foveae, two inner ones confluent, sutural stria entire, with short, humeral impression. Abdomen about as wide as elytra, first visible tergite 1 (IV) large, distance between lateral carina and median carina and distance between median carinae equal. Legs long and slender, simple, metatrochanter lacking medium spine. Aedeagus (Fig. 5) 0.48 mm long, about three times as long as its maximum width, distal part prolonged to long, pointed apex, with bunch of setae.</p><p>Remarks. Protamaurops peristeri, erroneously considered as a junior synonym of P. winneguthi, is instead a valid species, distinguishable from the latter by the following diagnostic characters: 1) body length about 2.6–2.7 mm, against 2.3–2.4 mm, 2) head capsule about as long as wide against head capsule 1.15 as wide as long, 3) antennomere 5 1.5 times as long as wide, against 1.25 times as long as wide and 4) lacking medium spine on metatrochanter, against spine on metatrochanter present.</p><p>Distribution. North Macedonia (so far known only by 2 male specimens, both known only from Baba planina Mts. Pelister).</p></div>	https://treatment.plazi.org/id/345E87D7FFC6FFFCFF47FAE7FEAEFCE9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hlaváč, Peter;Brachat, Volker;Delić, Teo	Hlaváč, Peter, Brachat, Volker, Delić, Teo (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXIX. Further contribution to the taxonomy of the tribe Amauropini (Coleoptera: Staphylinidae: Pselaphinae) from the Balkan Peninsula. Zootaxa 5665 (1): 137-143, DOI: 10.11646/zootaxa.5665.1.10, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
345E87D7FFC1FFFCFF47FCE5FF73F9D1.text	345E87D7FFC1FFFCFF47FCE5FF73F9D1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protamaurops macrophthalmus (Muller 1944)	<div><p>Protamaurops macrophthalmus (Müller, 1944)</p><p>Figs 4, 6</p><p>Material examined. ALBANIA: 2 ♂♂, 4 ♀♀: two label “ALBANIA, 27.X.2024 / 41°53'0,53''N, 730 m / 20°24'47,8''E, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=20.413277&amp;materialsCitation.latitude=41.883484" title="Search Plazi for locations around (long 20.413277/lat 41.883484)">2.2 km / N of Matranxhë vill.</a> / A. Šíma leg.“ [white, printed], “under stone“ [white, printed] (PCAS, PCPH) . 2 ♂♂, 8 ♀♀: AL – Albania mer. 15.V.2016 / 40°12'55.18''N, 19°34'12.74''E / <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=19.570206&amp;materialsCitation.latitude=40.21533" title="Search Plazi for locations around (long 19.570206/lat 40.21533)">Llogara NP</a>, sifting ( Abies, / Pinus, moss), A. Šíma leg. (PCAS, PCPH, PCVB) .</p><p>Distribution. Albania (Mali I Shejtit, Mali i Tomorrit, Maja e Korabit, Kukës and Korçë municipality, Llogara NP), Greece (Epirus, Mts. Vitsi) and North Macedonia (Šar planina).</p><p>Remarks. Both wings and legs polymorphy are well-known phenomena in Pselaphinae and represent one of the major issues in their taxonomy (Löbl, Kurbatov &amp; Nomura 1998). Differently, the eyes polymorphy is much rarer and it is only scarcely documented in the genus Tychobythinus Ganglbauer, 1896, tribe Bythinini (Holdhaus 1905; Orousset &amp;Dubault 1985).The later phenomenon is especially expressed in P.macrophthalmus, with males exhibiting strong eye dimorphism, resulting in easily differentiated morphological forms – macro and microphthalmous. The macrophthalmous form has very large, round and laterally protruding eyes with a short lateral spine located under the eye. Microphthalmous form, described as P. macrophthalmus beieri (J. Müller, 1944) and synonymized by Bekchiev &amp; Hlaváč (2020), has small eyes with about 3–5 ommatidia located under or around the longer ocular spine. Both forms have been recently collected in Albania, the macrophthalmous form near Matranxhë village and the microphthalmous from Llogara National Park. In both forms, only microphthalmous or blind females are known.</p></div>	https://treatment.plazi.org/id/345E87D7FFC1FFFCFF47FCE5FF73F9D1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hlaváč, Peter;Brachat, Volker;Delić, Teo	Hlaváč, Peter, Brachat, Volker, Delić, Teo (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXIX. Further contribution to the taxonomy of the tribe Amauropini (Coleoptera: Staphylinidae: Pselaphinae) from the Balkan Peninsula. Zootaxa 5665 (1): 137-143, DOI: 10.11646/zootaxa.5665.1.10, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
345E87D7FFC1FFFDFF47F9FCFDF8FD9D.text	345E87D7FFC1FFFDFF47F9FCFDF8FD9D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protamaurops Muller 1944	<div><p>Remaining taxonomic problem in Protamaurops</p><p>The taxonomy of P. winneguthi is problematic. Karaman (1961) described three new species, P. similis Karaman, 1961, P. peristeri Karaman, 1961 and P. vitolistensis (Karaman, 1954) . Later, all three species names were synonymized with P. winneguthi . First, Karaman (1967) synonymized P. similis based on the same structure of the aedeagus. The remaining two species, P. peristeri and P. vitolistensis, were synonymized by Besuchet (1999), however, without a clear taxonomical reasoning. Moreover, it seems that the synonymy was not based on the examination of the type material but solely reporting the number of ommatidia of P. winneguthi, P. peristeri and P. vitolistensis from the original descriptions. Thus, the synonymy of the latter two species names is doubtful and has to be additionally verified. Synonymy of these two species names becomes even more questionable if we consider original work of Karaman (1961). P. peristeri was originally described based on a single male from NP Pelister, most probably from its 2601 m highest peak (at the time known also as Perister) in Baba planina mountains, where it was collected by the author, Zora Karaman, herself. She placed the newly described species in a group of species characterized by well-developed round eyes with the ocular spine located either in the middle or under the eye, together with P. macrophthalmus, P. beieri (now synonym of P. macrophthalmus) and P. serbicus Karaman, 1961 . Differently, the second group of species, including P. bicarinatus, P. winneghuti, P. similis and P. vitolistensis, is characterized by the existence of rudimentary eyes, with some ommatidia around or above the eye. Due to the strong eye dimorphism in P. macrophthalmus, the separation of two groups of species based on the structure of eyes is not informal. Due to this, the status of P. bicarinatus (Reitter, 1918), known only by a single female holotype (Hlaváč, Baňař &amp; Koukalová, 2021), remains doubtful and needs to be confirmed by finding a male from the type locality. The discovery of one male specimen of P. peristeri from Pelister, collected by a well-known Czech specialist on Staphylinidae Jan Rambousek in 1914, in the collection of Volker Brachat, allowed us to correct the taxonomical status of P. peristeri (see the text above).</p></div>	https://treatment.plazi.org/id/345E87D7FFC1FFFDFF47F9FCFDF8FD9D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hlaváč, Peter;Brachat, Volker;Delić, Teo	Hlaváč, Peter, Brachat, Volker, Delić, Teo (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXIX. Further contribution to the taxonomy of the tribe Amauropini (Coleoptera: Staphylinidae: Pselaphinae) from the Balkan Peninsula. Zootaxa 5665 (1): 137-143, DOI: 10.11646/zootaxa.5665.1.10, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
345E87D7FFC0FFFDFF47FAFCFD43F9B7.text	345E87D7FFC0FFFDFF47FAFCFD43F9B7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Paramaurops apfelbecki (Ganglbauer 1896)	<div><p>Paramaurops apfelbecki (Ganglbauer, 1896)</p><p>Figs 7, 8</p><p>Material examined. CROATIA: 2♂♂: 2 labels “CROATIA, Lastovo / Jama na Lastovu / VII/1997 T. Radja “ [white, printed] (PCPH). First record for the island Lastovo .</p><p>Distribution. Bosna and Hercegovina, Croatia.</p></div>	https://treatment.plazi.org/id/345E87D7FFC0FFFDFF47FAFCFD43F9B7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hlaváč, Peter;Brachat, Volker;Delić, Teo	Hlaváč, Peter, Brachat, Volker, Delić, Teo (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXIX. Further contribution to the taxonomy of the tribe Amauropini (Coleoptera: Staphylinidae: Pselaphinae) from the Balkan Peninsula. Zootaxa 5665 (1): 137-143, DOI: 10.11646/zootaxa.5665.1.10, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
345E87D7FFC0FFFDFF47F911FE60F80D.text	345E87D7FFC0FFFDFF47F911FE60F80D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Paramaurops kaufmanni (Ganglbauer 1895)	<div><p>Paramaurops kaufmanni (Ganglbauer, 1895)</p><p>Figs 9, 10</p><p>Material examined. MONTENEGRO: 6♂♂, 7♀♀: two labels “Montenegro 725 m / 42°32'56''N, 18°39'37''E / Crkvice env. / A. Šíma leg., 3.XI.2023 “ [white, printed], “under deeply embedded / stones placed in grass / Quercus forest, 16°C / after heavy rains“ [white, printed] (PCAS, PCPH, PCRK) . 1♂: four labels “Dalmácia / leg. Dr. Fodor “ [white, printed], “Zelenika / Salina 927.V.2“ [white, handwritten], “ Amaurops / Kaufmanni Glb. / det. Székessy “ [white, printed and handwritten], “coll. / Dr. J. Fodor “ [white, printed] (HNHM) . 1♂: one label “CRNA GORA / Stoliv, 27.08.2014 / M. Stevanović leg.“ [white, printed] (PCPH) . New records.</p><p>Distribution. Montenegro.</p></div>	https://treatment.plazi.org/id/345E87D7FFC0FFFDFF47F911FE60F80D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hlaváč, Peter;Brachat, Volker;Delić, Teo	Hlaváč, Peter, Brachat, Volker, Delić, Teo (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXIX. Further contribution to the taxonomy of the tribe Amauropini (Coleoptera: Staphylinidae: Pselaphinae) from the Balkan Peninsula. Zootaxa 5665 (1): 137-143, DOI: 10.11646/zootaxa.5665.1.10, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
